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Syspira armasi species nova Figures 6, 9, 10 A–B, 11B Types. Male holotype (MNHNSD 09.1627) from Sabana Vieja, Sabaneta, Parque Nacional José del Carmen Ramírez (19°04'38.8"N– 71°11'37.8"W), 1876 m, Province San Juan, Dominican Republic, 19.XI.2014, C. Suriel, C. Marte & G. de los Santos. Paratypes: female (MNHNSD 09.1628) from Sabana Nueva (19°03'24.1"N– 71°09'52.9"W), 1877 m, 17.XI.2014, C. Suriel, C. Marte & G. de los Santos, other data as holotype; two females (MNHNSD 09.1630) from Sabana Vieja (19°04'42.8"N– 71°11'50.8"W), 1897 m, 10.I.2008, R. Rodríguez & G. de los Santos, other data as holotype; two males (MNHNSD 09.1629) from Sabana Vieja (19°04'42.8"N– 71°11'50.8"W), 1897 m, 10.I.2008, A. Sánchez-Ruiz & C. Suriel, other data as holotype. Other material examined. DOMINICAN REPUBLIC: Province San Juan: Sabaneta, Sabana Nueva, Parque Nacional José del Carmen Ramírez (19°03'24.1"N– 71°09'52.9"W), 1877 m, 17.XI.2014, C. Suriel, C. Marte & G. de los Santos, 2♀ (MNHNSD 09.1631). Sabaneta, Sabana Vieja, Parque Nacional José del Carmen Ramírez (19°04'42.8"N– 71°11'50.8"W), 1897 m, 10.I.2008, A. Sánchez-Ruiz & G. de los Santos, 1♀ (MNHNSD 09.1638). Sabaneta, Sabana Vieja, Parque Nacional José del Carmen Ramírez (19°04’42.8”N– 71°11’50.8”W), 1897 m, 10.I.2008, C. Suriel & G. de los Santos, 1♂ (MNHNSD 09.1639). Province Elías Piña: Sabana del Silencio, El Hoyazo, Hondo Valle, Parque Nacional Sierra de Neiba (18°39'8.73"N – 71°33'18.74"W), 2007 m, 19–22.VI.2015, A. Sánchez, C. Marte & G. de los Santos, 3♀ (MNHNSD 09.1620–09.1622). Etymology. The specific name is a patronym in honor of the Cuban arachnologist Luis F. de Armas. Diagnosis. Males of S. armasi sp. n. resembles those of S. medialuna by having a small median apophysis (Figs 5 B–C, 6B–C), but can be distinguished by having a smaller embolar base process (Fig. 6 A–C); smaller finger-shaped median apophysis (Fig. 6A, C) and a strongly curved apical projection of embolar base in ventral view (Fig. 6B). Females resemble those of S. bryantae sp. n. by having an atrial hood projected (Figs 6D, 7A), but can be distinguished by having it larger and slightly sclerotized, completely covering anterior edges of epigynal posterior plates (Fig. 6D). Description. Male (holotype MNHNSD 09.1627): carapace orange with the two scarcely distinguishable gray paramedian bands (Fig. 10B). Chelicerae orange. Labium and endites light orange; sternum yellow with orange edges. Legs orange, darker from tibiae to tarsus. Abdomen dorsally gray with white spots (Fig. 10B), ventrally yellow. Total length 10.4, carapace length 5.3, width 4.0. Eye diameters: AME 0.12, ALE 0.16, PME 0.18, PLE 0.18. Chelicerae 1.7 long, with 3 promarginal and 2 retromarginal teeth. Leg femur length: I—4.0; II—3.9; III—3.5; IV—4.5. Leg spination: I—tibia v2-2-2, p0-1-1, r0-1-1, d0; metatarsus v2-2-0, p1-2-0, r0-1-0, d0; II—tibia v2-2-2, p1-1-0, r1-1-0, d0, metatarsus v2-2-0, p1-2-1, r0-1-0, d0; III—tibia v2-2-2, p1-1-0, r1-1-0, d2-1-0, metatarsus v2-2- 1, p1-1-1, r1-1-1, d2-2-2; IV—tibia v2-2-2, p1-1-1, r1-1-0, d1-1-0, metatarsus v2-2-2, p1-1-1, r1-1-1, d2-2-2. Palp: cymbial groove half-moon shaped, distally closed, reaching ventral margin of cymbium; retrolateral tibial apophysis short and truncated distally; median apophysis small with bifurcate tip (Fig. 6B); apical projection of embolar base strongly curved and sharp at tip, resembling a bird head (Fig. 6B); embolar base covered by sub-quadrangular ventral small tegular process, directed proximally (Fig. 6 B–C). Female (paratype MNHNSD 09.1628): carapace orange, darker in the first anterior quarter, especially at the lateral eyes level; with two light gray paramedian bands (Fig. 10A). Chelicerae reddish brown. Labium, endites and sternum dark orange. Legs orange with metatarsus and tarsus dark orange. Palp orange, with tarsus dark orange. Abdomen ventrally orange, dorsally gray with white spots (Fig. 10A). Total length 12.1, carapace length 5.2, width 3.9. Eye diameters: AME 0.14, ALE 0.18, PME 0.20, PLE 0.20. Chelicerae 2.2 long, with teeth as in male. Leg femur length: I—3.8; II—3.4; III—3.2; IV—4.2. Palp: claw with five teeth (Fig. 6F); spination: patella p1; tibia p2–1–0, d1–1–0; tarsus v0–0–2 p2–1–0, r1–1–0, d1–0–0. Leg spination: I—tibia v2–2–2, p0, r0, d0; metatarsus v2–2–0, p0, r0, d0; II—tibia v1–1–2, p0, r0, d0, metatarsus v2–2–0, p0, r0, d0; III—tibia v2–2–2, p1–1–0, r1–1–0, d1–1– 0, metatarsus v2–2–1, p1–1–0, r1–1–0, d2–2–2; IV—tibia v2–2–2, p1–1–0, r1–1–0, d1–1–0, metatarsus v2–2–2, p1–1–1, r1–1–1, d2–2–2. Abdomen: length 6.4. Epigynum: atrial hood large and slightly sclerotizated, completely covering anterior edges of epigynal posterior plates (Fig. 6D); primary spermathecae coiled; secondary spermathecae projected internally; fertilization ducts small and C-shaped copulatory ducts (Fig. 6 D–E). Remarks. This species is known from the highland savannas and surrounding pine forests at the National Parks José del Carmen Ramírez and Sierra de Neiba. Distribution. Provinces of Elías Piña and San Juan, Dominican Republic (Fig. 9).

Syspira bryantae species nova Figures 7, 9, 10F, 11A Types. Female holotype (MNHNSD 09.1617) from Parque Nacional Armando Bermúdez (19°03'59.79"N– 71°02'5.70"W), 1790 m, Valle de Bao, Province Santiago, Dominican Republic, 11–12.VII.2015, C. Suriel, C. Marte & G. de los Santos. Paratypes: two females (MNHNSD 09.1618 and MNHNSD 09.1619), same data as holotype. Etymology. The specific name is a patronym in honor of the American arachnologist Elizabeth B. Bryant. Diagnosis. Females resemble those of S. armasi sp. n. by having an atrial hood projected (Figs 6D, 7A), but can be distinguished by having it smaller and heavily sclerotized, covering only part of anterior edges of epigynal posterior plates (Fig. 7 D–E). Description. Male: Unknown. Female (holotype MNHNSD 09.1617): carapace orange, with gray paramedian bands (Fig. 10F). Chelicerae red brown with its inner distal part orange. Labium, endites and sternum dark orange. Palp orange, claw with three teeth. Legs orange with metatarsus and tarsus dark orange. Abdomen dorsally gray with white spots (Fig. 10F), ventrally yellow. Total length 11.6, carapace length 5.4, width 3.8. Eye diameters: AME 0.16, ALE 0.18, PME 0.20, PLE 0.20. Chelicerae 2.2 long, with 3 promarginal and 2 retromarginal teeth. Leg femur length: I—3.9; II—3.8; III—3.3; IV—4.4. Palp: claw with three teeth (Fig. 7C); spination: patella p1; tibia p2-1-0, d0-1-1; tarsus v0-0-2, p2-1-0, r1-1-0, d0-1-0. Leg spination: I—tibia v2-2-2, p0, r0, d0, metatarsus v2-2-0, p0, r0, d0; II—tibia v1-1-2, p0, r0, d0, metatarsus v2-2-0, p0, r0, d0; III—tibia v1-2-1, p0-1-1, r1-1-0, d1-1-0, metatarsus v2-2-1, p1-1-1, r1-1-0, d2-2-2; IV—tibia v2-1-2, p1-1-0, r1-1-0, d0, metatarsus v2-2-2, p1-1-0, r1-1-1, d2-2-2. Epigynum: atrial hood projected, small and heavily sclerotized, just covering part of anterior edges of epigynal posterior plates (Fig. 7 D–E); primary spermathecae coiled; secondary spermathecae projected internally; fertilization ducts smaller than spermathecae and C-shaped copulatory ducts (Fig. 7 A–B). Distribution. Known only from the highland savanna found in the type locality (Fig. 9).

Syspira cimitarra Brescovit, S��nchez-Ruiz & Bonaldo Figures 1, 9 Syspira cimitarra Brescovit, S��nchez-Ruiz & Bonaldo, 2018: 58, figs 1, 2, 7. Male holotype and female paratype (IBSP 168411) and female paratype (MNHNSD 09.1539) from Parque Nacional Valle Nuevo, 18��41���11.9���N ��� 70��35���37.4���W, 2041 m, 15 km of Constanza, Province La Vega, Cibao Sur, Dominican Republic, 11.IV.2015, A. S��nchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province Barahona, Parque Nacional Sierra Mart��n Garc��a, Loma El Curro (18��22'48.8"N ��� 70��01'35.2"W), 1379 m, 30.V.2014, G. de los Santos, 1♂ (MNHNSD 09.1623). Diagnosis. Males of S. cimitarra are easily distinguished from other Hispaniola species by the retrolateral tibial apophysis elongated and curved at tip, and by the filiform apical projection of the embolar base (Fig. 1A ��� C; Brescovit et al. 2018: figs 1A ��� B, 2B ��� F). Females resemble those of Syspira jimmyi by the enlarged atrium (Fig. 1D ��� E), but can be distinguished by the dorsal median plate truncated at the distal margin, and wider epigynal posterior plates, exceeding in half the atrial lateral dorsal folds (Fig. 1D ��� E; Brescovit et al. 2018: figs 1C ��� D, 2I). Description. Male and female described by Brescovit et al. (2018). Remarks. Specimens from Parque Nacional Sierra Mart��n Garc��a, Barahona Province represent new distribution records for this species. Previously known from Provinces La Vega and Peravia (Brescovit et al. 2018), its distribution is now extended to the cloud forest remnants at the top of Loma El Curro mountain (Fig. 9), where it lives sympatrically with Syspira alayoni sp. n. Distribution. Provinces Barahona, La Vega and Peravia, Dominican Republic (Fig. 9)., Published as part of S��nchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on page 415, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Brescovit, A. D., Sanchez-Ruiz, A. & Bonaldo, A. B. (2018) On the spider genus Syspira Simon, 1895 (Araneae: Miturgidae) in the Caribbean: four new species from Dominican Republic. Zootaxa, 4370 (1), 57 - 76. https: // doi. org / 10.11646 / zootaxa. 4370.1.4"]}

Syspira medialuna Brescovit, Sánchez-Ruiz & Bonaldo Figures 5, 9 Syspira medialuna Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 62, figs 5–7. Male holotype (IBSP 168415) and two female paratypes (IBSP 168416 and MNHNSD 09.1541) from Reserva Científica Ébano Verde, 19°1’56.8”N– 70°32’30.9”W, 1029 m, Constanza, Province La Vega, Dominican Republic, 16–17.IV.2015, A. Sánchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province La Vega, Jarabacoa, Reserva Científica Ébano Verde, Paso Bajito, La Sal, Loma La Golondrina (19°03'55.9"N – 70°33'02.9"W), 1520 m, 17.V.2009, A. Mateo, R. Carreras & G. de los Santos, 1♂ (MNHNSD 09.1637). Diagnosis. Males of Syspira medialuna are easily distinguished from other Hispaniolan species of the agujas group by a fairly developed embolar base process (Fig. 5 A–C), and a spatula-shaped median apophysis in ventral view (Fig. 5B; Brescovit et al. 2018: fig. 5A–B). Females are distinguished from other Hispaniolan species by large epigynal posterior plates with quite high anterior edges, twice the size of the other known species (Fig. 5D; Brescovit et al. 2018: figs 5C, 6E). Description. Male and female described by Brescovit et al. (2018). Distribution. Province La Vega, Dominican Republic (Fig. 9).

Syspira agujas Brescovit, S��nchez-Ruiz & Bonaldo Figures 4, 9, 10D Syspira agujas Brescovit, S��nchez-Ruiz & Bonaldo, 2018: 63, figs 5, 6. Female holotype (IBSP 168408) and female paratype (IBSP 168410) from Via Las Agujas, 18��40���32.4���N��� 70��35���31.6���W, 1952 m, Province San Jos�� de Ocoa, Dominican Republic, 11.IV.2015, A. S��nchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province La Vega, Constanza, Parque Nacional Valle Nuevo: La Neverita (18��41'49.9"N��� 70��35'27.1"W), 2262 m, 17.X.2011, G. de los Santos, 1♂ (MNHNSD 09.1632); 1♀ (MNHNSD 09.1633). La Pir��mide (18��42'22.7"N��� 70��36'01.6"W), 2392 m, 18.X.2011, S. Carrero 1♂ (MNHN- SD 09.1634). La Pir��mide (18��42'24.7"N ��� 70��35'58.3"W), 2395 m, 19.I.2011, S. Carrero & G. de los Santos, 1♀ (MNHNSD 09.1635). Sabana Qu��liz (18��45'32.5"N��� 70��38'03.5"W), 2290 m, 25.VIII.2010, S. Carrero & G. de los Santos, 1♀, 1juv (MNHNSD 09.1636). Diagnosis. Males can be distinguished from other Hispaniolan species of the agujas group by a larger and heavily sclerotized median apophysis (Fig. 4B), apical projection on embolar base bowed and claw-shaped (Fig. 4 A���B). Females are distinguished from other Hispaniolan species of agujas group by the wide and straight anterior edges of epigynal posterior plates (Fig. 4D). Description. Male (MNHNSD 09.1632). carapace orange, without gray paramedian bands (Fig. 10D). Legs orange, with tibiae of legs I and II dark orange and metatarsus and tarsus brown (Fig. 10D); metatarsus and tarsus of legs III and IV dark orange. Chelicerae orange. Labium, endites, sternum and coxae orange with dark orange edges. Abdomen dorsally gray with white spots (Fig. 10D), ventrally yellow. Total length 10.9, carapace length 6.0, width 4.3. Eye diameters: AME 0.16, ALE 0.18, PME 0.20, PLE 0.18. Chelicerae 2.2 long, with 3 promarginal and 2 retromarginal teeth. Leg femur length: I���4.8; II���4.7; III���4.4; IV���5.4. Leg spination: I���tibia v2-2-2, p1-1-0, r1-1-0, d0; metatarsus v2-2-0, p1-2-0, r1-1-0, d0; II���tibia v2-2-2, p1-1-0, r1-1-0, d0, metatarsus v2-2-0, p1-2-0, r1- 1-0, d0; III���tibia v2-2-2, p1-1-0, r1-1-0, d1-1-0, metatarsus v2-2-1, p1-1-0, r1-1-0, d2-2-2; IV���tibia v2-2-2, p1- 1-0, r1-1-0, d0, metatarsus v2-2-2, p1-1-1, r1-1-1, d2-2-2. Palp: cymbial groove half-moon shaped, distally closed, reaching ventral margin of cymbium; retrolateral tibial apophysis short and truncated distally; median apophysis large, palette-shaped, projecting anteriorly and separated about 45�� from the embolus; apical projection of embolar base curved and sharp at tip, claw-shaped; embolar base covered by sub-quadrangular ventral tegular process, directed proximally (Fig. 4 A���C). Female (holotype, IBSP 168408). Described by Brescovit et al. (2018). Supplement: female papal claw with four teeth (Fig. 4F). Remarks. Specimens of both sexes from Parque Nacional Valle Nuevo, La Vega Province represent new distribution records for this species, previously known only from San Jos�� de Ocoa Province (Brescovit et al. 2018). All these specimens from La Vega were found in highland savannas above 2200 m. Variation. Examined females from La Vega Province have the palp claw with four teeth (Fig. 4F) while specimens from Las Agujas have only two. Distribution. Provinces of San Jos�� de Ocoa and La Vega, Dominican Republic (Fig. 9)., Published as part of S��nchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on pages 419-420, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Brescovit, A. D., Sanchez-Ruiz, A. & Bonaldo, A. B. (2018) On the spider genus Syspira Simon, 1895 (Araneae: Miturgidae) in the Caribbean: four new species from Dominican Republic. Zootaxa, 4370 (1), 57 - 76. https: // doi. org / 10.11646 / zootaxa. 4370.1.4"]}

The species of Syspira Simon from Hispaniola are revised and detailed documentation is provided for all known species from this island. Four new species (Syspira alayoni sp. n., S. armasi sp. n., S. barbacoa sp. n., and S. bryantae sp. n.), and the unknown male of Syspira agujas Brescovit, Sánchez-Ruiz & Bonaldo are described. New distribution records were found for three known species (S. agujas, S. cimitarra Brescovit, Sánchez-Ruiz & Bonaldo and S. medialuna Brescovit, Sánchez-Ruiz & Bonaldo). The Hispaniolan endemic species Teminius monticola (Bryant, 1948) is transferred to Syspira based on the revision of its holotype. A key for Hispaniola Syspira species is provided. These species can be separated into two distinct groups based on features of the male palpal morphology and female genitalia: the cimitarra group including S. cimitarra, S. jimmyi Brescovit, Sánchez-Ruiz & Bonaldo, S. alayoni and S. barbacoa sp. n., and the agujas group including S. agujas, S. medialuna, S. monticola comb. n., S. armasi sp. n. and S. bryantae sp. n.

Syspira cimitarra Brescovit, Sánchez-Ruiz & Bonaldo Figures 1, 9 Syspira cimitarra Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 58, figs 1, 2, 7. Male holotype and female paratype (IBSP 168411) and female paratype (MNHNSD 09.1539) from Parque Nacional Valle Nuevo, 18°41’11.9”N – 70°35’37.4”W, 2041 m, 15 km of Constanza, Province La Vega, Cibao Sur, Dominican Republic, 11.IV.2015, A. Sánchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province Barahona, Parque Nacional Sierra Martín García, Loma El Curro (18°22'48.8"N – 70°01'35.2"W), 1379 m, 30.V.2014, G. de los Santos, 1♂ (MNHNSD 09.1623). Diagnosis. Males of S. cimitarra are easily distinguished from other Hispaniola species by the retrolateral tibial apophysis elongated and curved at tip, and by the filiform apical projection of the embolar base (Fig. 1A – C; Brescovit et al. 2018: figs 1A – B, 2B – F). Females resemble those of Syspira jimmyi by the enlarged atrium (Fig. 1D – E), but can be distinguished by the dorsal median plate truncated at the distal margin, and wider epigynal posterior plates, exceeding in half the atrial lateral dorsal folds (Fig. 1D – E; Brescovit et al. 2018: figs 1C – D, 2I). Description. Male and female described by Brescovit et al. (2018). Remarks. Specimens from Parque Nacional Sierra Martín García, Barahona Province represent new distribution records for this species. Previously known from Provinces La Vega and Peravia (Brescovit et al. 2018), its distribution is now extended to the cloud forest remnants at the top of Loma El Curro mountain (Fig. 9), where it lives sympatrically with Syspira alayoni sp. n. Distribution. Provinces Barahona, La Vega and Peravia, Dominican Republic (Fig. 9).

Syspira medialuna Brescovit, S��nchez-Ruiz & Bonaldo Figures 5, 9 Syspira medialuna Brescovit, S��nchez-Ruiz & Bonaldo, 2018: 62, figs 5���7. Male holotype (IBSP 168415) and two female paratypes (IBSP 168416 and MNHNSD 09.1541) from Reserva Cient��fica ��bano Verde, 19��1���56.8���N��� 70��32���30.9���W, 1029 m, Constanza, Province La Vega, Dominican Republic, 16���17.IV.2015, A. S��nchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province La Vega, Jarabacoa, Reserva Cient��fica ��bano Verde, Paso Bajito, La Sal, Loma La Golondrina (19��03'55.9"N ��� 70��33'02.9"W), 1520 m, 17.V.2009, A. Mateo, R. Carreras & G. de los Santos, 1♂ (MNHNSD 09.1637). Diagnosis. Males of Syspira medialuna are easily distinguished from other Hispaniolan species of the agujas group by a fairly developed embolar base process (Fig. 5 A���C), and a spatula-shaped median apophysis in ventral view (Fig. 5B; Brescovit et al. 2018: fig. 5A���B). Females are distinguished from other Hispaniolan species by large epigynal posterior plates with quite high anterior edges, twice the size of the other known species (Fig. 5D; Brescovit et al. 2018: figs 5C, 6E). Description. Male and female described by Brescovit et al. (2018). Distribution. Province La Vega, Dominican Republic (Fig. 9)., Published as part of S��nchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on pages 420-421, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Brescovit, A. D., Sanchez-Ruiz, A. & Bonaldo, A. B. (2018) On the spider genus Syspira Simon, 1895 (Araneae: Miturgidae) in the Caribbean: four new species from Dominican Republic. Zootaxa, 4370 (1), 57 - 76. https: // doi. org / 10.11646 / zootaxa. 4370.1.4"]}

Key for Hispaniolan Syspira species 1. Male............................................................................................... 2 - Female............................................................................................. 6 2. Cymbial groove not reaching the ventral distal margin of cymbium (Figs 1C, 2C); RTA enlarged and curved (Figs 1B, 2B); apical projection of embolar base small and not curved and with lighter sclerotization (Figs 1A, 2A); process on embolar base inconspicuous (Figs 1A, 2B) (the cimitarra group ).......................................................... 3 - Cymbial groove distally closed, reaching the ventral margin of cymbium (Figs 4C, 5C, 6C); RTA short and truncated (Figs 4B, 5B, 6B); bigger and curved embolar base apical projection with heavy sclerotization (Figs 4B, 5B, 6B); process on embolar base developed (Figs 4A, 5A, 6A) (the agujas group )........................................................ 4 3. Median apophysis with finger-shaped tip (Fig. 1B; Brescovit et al. 2018: fig. 2C); RTA elongated and curved distally, with bifurcated projections unequal in size and with the longer projection striated at tip (Fig. 1C, Brescovit et al. 2018: fig. 2F)............................................................................................. S. cimitarra - Median apophysis with bifurcated tip (Fig. 2 A, Brescovit et al. 2018: fig. 4C); RTA shorter, sickle-shaped, with bifurcated projections almost equal in size (Fig. 2B – C, Brescovit et al. 2018: fig. 4F)................................. S. jimmyi 4. Median apophysis enlarged and heavily sclerotized (Fig. 4B, C); apical projection on embolar base bowed, claw-shaped (Fig. 4A, B)........................................................................................ S. agujas - Median apophysis small (Figs 5B – C, 6B – C); apical projection of embolar base not claw-shaped (Figs 5A – B, 6A – B)...... 5 5. Embolar base process fairly developed (Fig. 5A, C); spatula-shaped median apophysis in ventral view (Fig. 5B); apical projection of embolar base slightly curved in ventral view (Fig. 5B)........................................ S. medialuna - Embolar base process small (Fig. 6A – C); very small finger-shaped median apophysis (Fig. 6A, C); apical projection of embolar base strongly curved in ventral view (Fig. 6B).................................................. S. armasi sp. n. 6. Epigynal posterior plates (PP) medially fused at the inner borders; epigynal median plate (dmp) projecting dorsally from the atrium (Figs 1D – E, 2D – E, 3A – B, D – E) (the cimitarra group )................................................. 7 - Epigynal posterior plates (PP) not fused, touching (or not) the inner borders and lacking the epigynal dorsal median plate (dmp) (Figs 4D – E, 5D – E, 6D – E, 7A – B, 8D) (the agujas group ).................................................... 10 7. Anterior edges of epigynal posterior plates (PP) slightly straight and close together, posterior edges widely separated from each other (Fig. 3D – E)....................................................................... S. barbacoa sp. n. - Anterior edges of epigynal posterior plates (PP) rounded and elevated, and posterior edges fused (Figs 1D – E, 2D – E, 3A – B).................................................................................................... 8 8. Dorsal median plate (dmp) narrow, 2.5 times longer than wide, atrial hood (H) sclerotized (Fig. 3A, B)..... S. alayoni sp. n. - Dorsal median plate (dmp) wide, less than 1.5 times longer than wide, atrial hood (H) weakly sclerotized or inconspicuous (Figs 1D, 2D)............................................................................................. 9 9. Dorsal median plate (dmp) truncated at the distal margin (Fig. 1E), epigynal posterior plates (PP) wider, exceeding in half the atrial lateral dorsal folds (Fig. 1D)............................................................... S. cimitarra - Dorsal median plate (dmp) rounded at the distal margin, epigynal posterior plates (PP) narrower, barely exceeding the atrial lateral dorsal folds (Fig. 2D – E)................................................................... S. jimmyi 10. Anterior edges of epigynal posterior plates (PP) wide and straight (Fig. 4D)................................. S. agujas - Anterior edges of epigynal posterior plates (PP) relatively narrow and rounded (Figs 5D, 6D, 7A, 8D)................. 11 11. Atrial hood (H) dorsally projected from the atrium (Figs 6D, 7D – E)............................................ 12 - Atrial hood (H) not projected, remaining as a fold (Figs 5D, 8D)............................................... 13 12. Atrial hood (H) large and weakly sclerotized, completely covering anterior edges of epigynal posterior plates (Fig. 6D)............................................................................................ S. armasi sp. n. - Atrial hood (H) small and heavily sclerotized, covering only part of anterior edges of epigynal posterior plates (Fig. 7D – E)........................................................................................ S. bryantae sp. n. 13. Epigynal posterior plates (PP) large, with quite high anterior edges, twice the size seen in other species (Fig. 5D)................................................................................................... S. medialuna - Epigynal posterior plates (PP) smaller, with anterior edges similar in size to the other species (Fig. 8D).................................................................................................. S. monticola comb. nov., Published as part of Sánchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on pages 414-415, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Brescovit, A. D., Sanchez-Ruiz, A. & Bonaldo, A. B. (2018) On the spider genus Syspira Simon, 1895 (Araneae: Miturgidae) in the Caribbean: four new species from Dominican Republic. Zootaxa, 4370 (1), 57 - 76. https: // doi. org / 10.11646 / zootaxa. 4370.1.4"]}

Syspira monticola (Bryant, 1948) combinatio nova Figures 8, 9 Teminius monticolus Bryant, 1948: 411, fig. 94 (female holotype from Loma Rucilla, Armando Bermúdez National Park, Santiago Province, ~ 1500–2450 m, June, 1938, P. Darlington, deposited in MCZ, examined). Teminius monticola: Roewer, 1955: 407. Diagnosis. Females of S. monticola comb. n. resembles those of S. medialuna by having an atrial hood not projected (Figs 5D, 8D), but are readily recognized by having lower posterior plates and higher atrium (Fig. 8D). Description. Male unknown. Female described by Bryant (1948). Remarks. Collection efforts at the type and nearby localities, looking for new specimens, were unsuccessful. Vegetation in the type locality comprises a mixture of broadleaf and pine forest. Distribution. Know only from the type locality (Fig. 9)., Published as part of Sánchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on page 424, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Bryant, E. B. (1948) The spiders of Hispaniola. Bulletin of the Museum of Comparative Zoology, 100, 329 - 447.","Roewer, C. F. (1955) Katalog der Araneae von 1758 bis 1940, bzw. 1954. 2. Band, Abt. a (Lycosaeformia, Dionycha [excl. Salticiformia]). 2. Band. Abt. b (Salticiformia, Cribellata) (Synonyma-Verzeichnis, Gesamtindex). Institut royal des Sciences naturelles de Belgique, Bruxelles, 1751 pp."]}

Syspira jimmyi Brescovit, S��nchez-Ruiz & Bonaldo Figures 2, 9 Syspira jimmyi Brescovit, S��nchez-Ruiz & Bonaldo, 2018: 59, figs 3, 4, 7. Male holotype (IBSP 168419) and female paratype (IBSP 168420) from Parque Nacional Valle Nuevo, 18��41���11.9N ��� 70��35���37.4W, 2041 m, 15 km of Constanza, Province La Vega, Cibao Sur, Dominican Republic, 11.IV.2015, A. S��nchez-Ruiz & J. Cabra, examined. Diagnosis. Males of S. jimmyi are easily distinguished from other Hispaniolan species by the sickle-shaped RTA (Fig. 2C) and by the median apophysis with bifurcated tip (Fig. 2A; Brescovit et al. 2018: fig. 4C). Females resemble those of S. cimitarra by the enlarged atrium (Fig. 2D ��� E), but can be distinguished by the dorsal median plate rounded at the distal edge, epigynal posterior plates narrower, barely exceeding the atrial lateral dorsal folds (Fig. 2D ��� E, Brescovit et al. 2018: fig. 3C ��� D). Description. Male and female described by Brescovit et al. (2018). Distribution. Provinces La Vega, San Jos�� de Ocoa and Peravia, Dominican Republic (Fig. 9)., Published as part of S��nchez-Ruiz, Alexander, Santos, Gabriel De Los, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, The genus Syspira Simon, 1895 (Araneae: Miturgidae) from Hispaniola, with the description of four new species, pp. 413-431 in Zootaxa 4894 (3) on page 416, DOI: 10.11646/zootaxa.4894.3.7, http://zenodo.org/record/4315939, {"references":["Brescovit, A. D., Sanchez-Ruiz, A. & Bonaldo, A. B. (2018) On the spider genus Syspira Simon, 1895 (Araneae: Miturgidae) in the Caribbean: four new species from Dominican Republic. Zootaxa, 4370 (1), 57 - 76. https: // doi. org / 10.11646 / zootaxa. 4370.1.4"]}

Syspira agujas Brescovit, Sánchez-Ruiz & Bonaldo Figures 4, 9, 10D Syspira agujas Brescovit, Sánchez-Ruiz & Bonaldo, 2018: 63, figs 5, 6. Female holotype (IBSP 168408) and female paratype (IBSP 168410) from Via Las Agujas, 18°40’32.4”N– 70°35’31.6”W, 1952 m, Province San José de Ocoa, Dominican Republic, 11.IV.2015, A. Sánchez-Ruiz & J. Cabra, examined. Other material examined. DOMINICAN REPUBLIC: Province La Vega, Constanza, Parque Nacional Valle Nuevo: La Neverita (18°41'49.9"N– 70°35'27.1"W), 2262 m, 17.X.2011, G. de los Santos, 1♂ (MNHNSD 09.1632); 1♀ (MNHNSD 09.1633). La Pirámide (18°42'22.7"N– 70°36'01.6"W), 2392 m, 18.X.2011, S. Carrero 1♂ (MNHN- SD 09.1634). La Pirámide (18°42'24.7"N – 70°35'58.3"W), 2395 m, 19.I.2011, S. Carrero & G. de los Santos, 1♀ (MNHNSD 09.1635). Sabana Quéliz (18°45'32.5"N– 70°38'03.5"W), 2290 m, 25.VIII.2010, S. Carrero & G. de los Santos, 1♀, 1juv (MNHNSD 09.1636). Diagnosis. Males can be distinguished from other Hispaniolan species of the agujas group by a larger and heavily sclerotized median apophysis (Fig. 4B), apical projection on embolar base bowed and claw-shaped (Fig. 4 A–B). Females are distinguished from other Hispaniolan species of agujas group by the wide and straight anterior edges of epigynal posterior plates (Fig. 4D). Description. Male (MNHNSD 09.1632). carapace orange, without gray paramedian bands (Fig. 10D). Legs orange, with tibiae of legs I and II dark orange and metatarsus and tarsus brown (Fig. 10D); metatarsus and tarsus of legs III and IV dark orange. Chelicerae orange. Labium, endites, sternum and coxae orange with dark orange edges. Abdomen dorsally gray with white spots (Fig. 10D), ventrally yellow. Total length 10.9, carapace length 6.0, width 4.3. Eye diameters: AME 0.16, ALE 0.18, PME 0.20, PLE 0.18. Chelicerae 2.2 long, with 3 promarginal and 2 retromarginal teeth. Leg femur length: I—4.8; II—4.7; III—4.4; IV—5.4. Leg spination: I—tibia v2-2-2, p1-1-0, r1-1-0, d0; metatarsus v2-2-0, p1-2-0, r1-1-0, d0; II—tibia v2-2-2, p1-1-0, r1-1-0, d0, metatarsus v2-2-0, p1-2-0, r1- 1-0, d0; III—tibia v2-2-2, p1-1-0, r1-1-0, d1-1-0, metatarsus v2-2-1, p1-1-0, r1-1-0, d2-2-2; IV—tibia v2-2-2, p1- 1-0, r1-1-0, d0, metatarsus v2-2-2, p1-1-1, r1-1-1, d2-2-2. Palp: cymbial groove half-moon shaped, distally closed, reaching ventral margin of cymbium; retrolateral tibial apophysis short and truncated distally; median apophysis large, palette-shaped, projecting anteriorly and separated about 45° from the embolus; apical projection of embolar base curved and sharp at tip, claw-shaped; embolar base covered by sub-quadrangular ventral tegular process, directed proximally (Fig. 4 A–C). Female (holotype, IBSP 168408). Described by Brescovit et al. (2018). Supplement: female papal claw with four teeth (Fig. 4F). Remarks. Specimens of both sexes from Parque Nacional Valle Nuevo, La Vega Province represent new distribution records for this species, previously known only from San José de Ocoa Province (Brescovit et al. 2018). All these specimens from La Vega were found in highland savannas above 2200 m. Variation. Examined females from La Vega Province have the palp claw with four teeth (Fig. 4F) while specimens from Las Agujas have only two. Distribution. Provinces of San José de Ocoa and La Vega, Dominican Republic (Fig. 9).

MITURGIDAE Syspira spp.: Entypus aratus, Entypus unifasciatus californicus, Tachypompilus unicolor cerinus.

Kurczewski, Frank E., West, Rick C., Waichert, Cecilia, Kissane, Kelly C., Ubick, Darrell, Pitts, James P. (2020): New and unusual host records for North American and South American spider wasps (Hymenoptera: Pompilidae). Zootaxa 4891 (1): 1-112, DOI: https://doi.org/10.11646/zootaxa.4891.1.1

Cheiracanthium caudatum (Thorell, 1887) Eutittha caudata Thorell, 1887, p. 58 (description of immature ♀ (?) from MYANMAR: Yangon; Dr L. Fea leg. 1885). Cheiracanthium caudatum: Simon 1901, p. 67. Comments This species is known only from the female specimen collected in Yangon Division, Myanmar (= Burma) (Thorell 1887). Thorell expressed doubt about the maturity of the examined specimen. This is the only Eutittha species for which Simon (1901 p. 67) established a new combination. The absence of figures and descriptions of the epigyne make it difficult to define this species and solve the problem of its systematic position., Published as part of Esyunin, Sergei L. & Zamani, Alireza, 2020, ' Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae), pp. 1293-1323 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 54 (19 - 20) on page 1312, DOI: 10.1080/00222933.2020.1781950, http://zenodo.org/record/5028442, {"references":["Thorell T. 1887. Viaggio di L. Fea in Birmania e regioni vicine. II. Primo saggio sui ragni birmani. Anal Museo Civico Storia Nat Genova. 25: 5 - 417.","Simon E. 1901. On the Arachnida collected during the Skeat expedition to the Malay Peninsula. Proc Zool Soc London. 71 (1): 45 - 84. doi: 10.1111 / j. 1469 - 7998.1901. tb 08164. x"]}

Cheiracanthium incomptum (Thorell, 1891) Eutittha incomptum Thorell, 1891, p. 29 (description of immature ♀ from INDIA: Nicobar Islands; Th. Workman leg. 1845���1847). Chiracanthium (Sic!) incomptum: Bonnet 1956, p. 1058. Comments This species is known only from an immature female collected on Nicobar Islands (Thorell 1891). As in the previous case, a detailed study of the type is required to confirm the validity of this species., Published as part of Esyunin, Sergei L. & Zamani, Alireza, 2020, ' Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae), pp. 1293-1323 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 54 (19 - 20) on page 1313, DOI: 10.1080/00222933.2020.1781950, http://zenodo.org/record/5028442, {"references":["Thorell T. 1891. Spindlar fran Nikobarerna och andra delar af sodra Asien. Kongliga Svens Vetenskaps-Akademiens Handlingar. 24 (2): 1 - 149.","Bonnet P. 1956. Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome II. Systematique des araignees (Etude par ordre alphabetique) (2 me partie: C-F). Douladoure Toulouse. p. 919 - 1926."]}

Cheiracanthium tigbauaensis Barrion & Litsinger, 1995, stat. resurr. Cheiracanthium tigbauaensis Barrion & Litsinger, 1995, p. 161, figs 91a���f, 92a���d (description of holotype ♂ from PHILIPPINES: Panay Island (10��40 �� 29���N, 122��22 �� 39���E), A.T. Barrion leg. 3.VIII.1977, paratype ♂ and ♀ from same locality, A.T. Barrion leg. 9. XI.1978; paratype ♀ from PHILIPPINES: Luzon Island (16��02 �� 38���N, 120��29 �� 08���E), J.P. Bandong leg. 18.VIII.1979). Deeleman-Reinhold 2001, p. 228 (synonymy with C. insulanum; here rejected). Comments This species was originally described from Panay Island, Philippines (Barrion and Litsinger 1995) from both sexes. The conspecificity of the two species, C. adjacensoides and C. tigbauaensis, is not at all certain. The respective suggestion by Deeleman-Reinhold (2001, p. 231) is not reasonable. In the original description of C. tigbauaensis it is indicated that ���tegular apophysis moderately broad, squarely cut below before the tapering distal end��� (Barrion and Litsinger 1995, p. 164, fig. 91b, c), while TA of C. adjacensoides is unmodified, rounded at the apex (Song et al. 1990: fig 3)., Published as part of Esyunin, Sergei L. & Zamani, Alireza, 2020, ' Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae), pp. 1293-1323 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 54 (19 - 20) on page 1318, DOI: 10.1080/00222933.2020.1781950, http://zenodo.org/record/5028442, {"references":["Barrion AT, Litsinger JA. 1995. Riceland spiders of South and Southeast Asia. Wallingford (UK): CAB International; p. xix + 700.","Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill Leiden: Koninklijke Brill NV; p. 591.","Song DX, Chen XE, Hou JW. 1990. A new species of the genus Chiracanthium from China (Araneae: Clubionidae). Acta Zootaxonomica Sin. 15: 427 - 430."]}

Cheiracanthium longipes (Thorell, 1890) Eutittha longipes Thorell, 1890, p. 372 (description of ♀ from INDONESIA: Sumatra, Padang; Dr. O. Beccari leg. 1878). Chiracanthium (Sic!) longipes: Reimoser 1934, p. 15. Comments This species is known only from a female specimen collected in Sumatra (Thorell 1890). The absence of figures and descriptions of the epigyne make it difficult to define this species and solve the problem of its systematic position., Published as part of Esyunin, Sergei L. & Zamani, Alireza, 2020, ' Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae), pp. 1293-1323 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 54 (19 - 20) on page 1312, DOI: 10.1080/00222933.2020.1781950, http://zenodo.org/record/5028442, {"references":["Thorell T. 1890. Studi sui ragni Malesi e Papuani. IV, 1. Anal Museo Civico Storia Nat Genova. 28: 5 - 421.","Reimoser E. 1934. The spiders of Krakatau. Proc Zool Soc London. 104 (1): 13 - 18. doi: 10.1111 / j. 1469 - 7998.1934. tb 06215. x"]}

Esyunin, Sergei L., Zamani, Alireza (2020): 'Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae). Journal of Natural History (J. Nat. Hist.) 54 (19-20): 1293-1323, DOI: 10.1080/00222933.2020.1781950, URL: http://dx.doi.org/10.1080/00222933.2020.1781950

Cheiracanthium sadanai Tikader, 1976, stat. resurr. Cheiracanthium sadanai Tikader, 1976, p. 175, fig 1���4 (description of holotype ♀ and allotype ♂ from INDIA: Punjab, Ludhiana Agricultural University compound; Dr L. Sadana leg. 12.6.1887). Majumder and Tikader 1991: 90, figs. 197���202 (♂ ♀). Cheiracanthium melanostomum: Deeleman-Reinhold 2001, p. 231, p. 254 (synonymy of C. sadanai with C. melanostomum, even though the author herself was uncertain about this decision; here rejected). Comments This species was originally described from Punjab State, India (Tikader 1976) from both sexes. The conspecificity of the two species, C. melanostomum and C. sadanai, is not at all certain. The respective suggestion by Deeleman-Reinhold (2001, p. 231) is not reasonable. In the original description of C. sadanai it is indicated that ���cephalothorax, abdomen and legs light green��� (Tikader 1976, p. 176), while in the original description of C. melanostomum, it is indicated that the cephalothorax colouration ranges from testaceous to dark testaceous (Thorell 1895, p. 44). According to the redescription by Majumder and Tikader (1991), Indian specimens of C. melanostomum also have a colour pattern (���cephalothorax, legs and abdomen light brown���, cf. Majumder and Tikader 1991, p. 63) different from that of C. sadanai. Other potential distinguishing characteristics are indicated in the definitive key of Majumder and Tikader (1991, p. 54���56). However, currently, there is no publication that includes illustrations that allow an unambiguous definition of either C. melanostomum or C. sadanai according to the characters of the copulatory organs (all illustrations of these species in the relevant literature are too small and/or hard to interpret). Moreover, there is no indication that the types of C. sadanai had been re-examined by any of the subsequent authors (that either treated C. melanostomum or C. sadanai). Unfortunately, due to strict regulations in India, the type specimens were not available for this study either., Published as part of Esyunin, Sergei L. & Zamani, Alireza, 2020, ' Conundrum of esoterica': on the long-forgotten genus Eutittha Thorell, 1878, with new taxonomic considerations in Cheiracanthium C. L. Koch, 1839 (Araneae: Cheiracanthiidae), pp. 1293-1323 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 54 (19 - 20) on page 1318, DOI: 10.1080/00222933.2020.1781950, http://zenodo.org/record/5028442, {"references":["Tikader BK. 1976. Two new species of spiders of the genera Cheiracanthium Koch and Clubiona Latreille (family: Clubionidae) from India. J Bombay Nat Hist Soc. 73: 175 - 177.","Majumder SC, Tikader BK. 1991. Studies on some spiders of the family Clubionidae from India. Rec Zool Surv India, Occasional Pap. 102: 1 - 175.","Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill Leiden: Koninklijke Brill NV; p. 591.","Thorell T. 1895. Descriptive catalogue of the spiders of Burma. London; p. 1 - 406."]}

Four new species of the genusCheiracanthiumC.L. Koch, 1839 from Xishuangbanna, Yunnan Province, China are described:C. daofengYu & Li,sp. nov.(♂♀),C. duanbiYu & Li,sp. nov.(♂♀),C. gouYu & Li,sp. nov.(♂), andC. wuquanYu & Li,sp. nov.(♀). In addition,SinocanthiumYu & Li,gen. nov., is described with the type speciesS. shuangqiuYu & Li,sp. nov.A key to cheiracanthiid genera distributed in East and Southeast Asia is provided.

One new genus, Callipelis gen. n., and six new species belonging to five families are described as new to science: Paracedicus kasatkini sp. n. (♀, Desidae), Ajmonia rajaeii sp. n. (♂♀, Dictynidae), Devade naderii sp. n. (♂, Dictynidae), Callipelis deserticola sp. n. (♂, Gnaphosidae), Zora huseynovi sp. n. (♂, Miturgidae) and Diaea osmanii sp. n. (♂♀, Thomisidae). All new species are known exclusively from Iran. Three related genera, Cedicus Simon, 1875, Cedicoides Charitonov, 1946 and Paracedicus Fet, 1993 are removed from Cybaeidae and placed in Desidae. One family (Desidae) and three genera (Ajmonia Caporiacco, 1934, Devade Simon, 1884 and Zora C.L. Koch, 1847) are reported from Iran for the first time.http://www.zoobank.org/urn:lsid:zoobank.org:act:5B79AF9D-63CF-4233-97A5-E90FEAFAC5B3http://www.zoobank.org/urn:lsid:zoobank.org:act:0A36F32E-E53D-4BDB-B258-0305317EB860http://www.zoobank.org/urn:lsid:zoobank.org:act:0BDA0C55-8311-49DB-9CCF-3152B50738E9http://www.zoobank.org/urn:lsid:zoobank.org:a...

We present a phylogenetic analysis of spiders using a dataset of 932 spider species, representing 115 families (only the family Synaphridae is unrepresented), 700 known genera, and additional representatives of 26 unidentified or undescribed genera. Eleven genera of the orders Amblypygi, Palpigradi, Schizomida and Uropygi are included as outgroups. The dataset includes six markers from the mitochondrial (12S, 16S, COI) and nuclear (histone H3, 18S, 28S) genomes, and was analysed by multiple methods, including constrained analyses using a highly supported backbone tree from transcriptomic data. We recover most of the higher-level structure of the spider tree with good support, including Mesothelae, Opisthothelae, Mygalomorphae and Araneomorphae. Several of our analyses recover Hypochilidae and Filistatidae as sister groups, as suggested by previous transcriptomic analyses. The Synspermiata are robustly supported, and the families Trogloraptoridae and Caponiidae are found as sister to the Dysderoidea. Our results support the Lost Tracheae clade, including Pholcidae, Tetrablemmidae, Diguetidae, Plectreuridae and the family Pacullidae (restored status) separate from Tetrablemmidae. The Scytodoidea include Ochyroceratidae along with Sicariidae, Scytodidae, Drymusidae and Periegopidae; our results are inconclusive about the separation of these last two families. We did not recover monophyletic Austrochiloidea and Leptonetidae, but our data suggest that both groups are more closely related to the Cylindrical Gland Spigot clade rather than to Synspermiata. Palpimanoidea is not recovered by our analyses, but also not strongly contradicted. We find support for Entelegynae and Oecobioidea (Oecobiidae plus Hersiliidae), and ambiguous placement of cribellate orb-weavers, compatible with their non-monophyly. Nicodamoidea (Nicodamidae plus Megadictynidae) and Araneoidea composition and relationships are consistent with recent analyses. We did not obtain resolution for the titanoecoids (Titanoecidae and Phyxelididae), but the Retrolateral Tibial Apophysis clade is well supported. Penestomidae, and probably Homalonychidae, are part of Zodarioidea, although the latter family was set apart by recent transcriptomic analyses. Our data support a large group that we call the marronoid clade (including the families Amaurobiidae, Desidae, Dictynidae, Hahniidae, Stiphidiidae, Agelenidae and Toxopidae). The circumscription of most marronoid families is redefined here. Amaurobiidae include the Amaurobiinae and provisionally Macrobuninae. We transfer Malenellinae (Malenella, from Anyphaenidae), Chummidae (Chumma) (new syn.) and Tasmarubriinae (Tasmarubrius, Tasmabrochus and Teeatta, from Amphinectidae) to Macrobuninae. Cybaeidae are redefined to include Calymmaria, Cryphoeca, Ethobuella and Willisius (transferred from Hahniidae), and Blabomma and Yorima (transferred from Dictynidae). Cycloctenidae are redefined to include Orepukia (transferred from Agelenidae) and Pakeha and Paravoca (transferred from Amaurobiidae). Desidae are redefined to include five subfamilies: Amphinectinae, with Amphinecta, Mamoea, Maniho, Paramamoea and Rangitata (transferred from Amphinectidae); Ischaleinae, with Bakala and Manjala (transferred from Amaurobiidae) and Ischalea (transferred from Stiphidiidae); Metaltellinae, with Austmusia, Buyina, Calacadia, Cunnawarra, Jalkaraburra, Keera, Magua, Metaltella, Penaoola and Quemusia; Porteriinae (new rank), with Baiami, Cambridgea, Corasoides and Nanocambridgea (transferred from Stiphidiidae); and Desinae, with Desis, and provisionally Poaka (transferred from Amaurobiidae) and Barahna (transferred from Stiphidiidae). Argyroneta is transferred from Cybaeidae to Dictynidae. Cicurina is transferred from Dictynidae to Hahniidae. The genera Neoramia (from Agelenidae) and Aorangia, Marplesia and Neolana (from Amphinectidae) are transferred to Stiphidiidae. The family Toxopidae (restored status) includes two subfamilies: Myroinae, with Gasparia, Gohia, Hulua, Neomyro, Myro, Ommatauxesis and Otagoa (transferred from Desidae); and Toxopinae, with Midgee and Jamara, formerly Midgeeinae, new syn. (transferred from Amaurobiidae) and Hapona, Laestrygones, Lamina, Toxops and Toxopsoides (transferred from Desidae). We obtain a monophyletic Oval Calamistrum clade and Dionycha; Sparassidae, however, are not dionychans, but probably the sister group of those two clades. The composition of the Oval Calamistrum clade is confirmed (including Zoropsidae, Udubidae, Ctenidae, Oxyopidae, Senoculidae, Pisauridae, Trechaleidae, Lycosidae, Psechridae and Thomisidae), affirming previous findings on the uncertain relationships of the "ctenids" Ancylometes and Cupiennius, although a core group of Ctenidae are well supported. Our data were ambiguous as to the monophyly of Oxyopidae. In Dionycha, we found a first split of core Prodidomidae, excluding the Australian Molycriinae, which fall distantly from core prodidomids, among gnaphosoids. The rest of the dionychans form two main groups, Dionycha part A and part B. The former includes much of the Oblique Median Tapetum clade (Trochanteriidae, Gnaphosidae, Gallieniellidae, Phrurolithidae, Trachelidae, Gnaphosidae, Ammoxenidae, Lamponidae and the Molycriinae), and also Anyphaenidae and Clubionidae. Orthobula is transferred from Phrurolithidae to Trachelidae. Our data did not allow for complete resolution for the gnaphosoid families. Dionycha part B includes the families Salticidae, Eutichuridae, Miturgidae, Philodromidae, Viridasiidae, Selenopidae, Corinnidae and Xenoctenidae (new fam., including Xenoctenus, Paravulsor and Odo, transferred from Miturgidae, as well as Incasoctenus from Ctenidae). We confirm the inclusion of Zora (formerly Zoridae) within Miturgidae.

Trilikauskas, L.A. (2019): To the fauna of spiders (Arachnida: Aranei) of the "Leopard Land" National Park and the "Kedrovaya Pad" State Nature Reserve, Primorskii krai. Far Eastern Entomologist 392: 6-20, DOI: 10.25221/fee.392.2, URL: http://dx.doi.org/10.25221/fee.392.2

Thorell (1897) erected the monotypic genus Palicanus based on a single male specimen from Yangon (formerly Rangoon) in Myanmar (formerly Burma). Initially the genus was placed in Drassidae Sundevall, 1833, which later became a junior synonym of Gnaphosidae Pocock, 1898. The genus was subsequently transferred to Clubionidae Wagner, 1887 (Simon 1897) and finally to Miturgidae Simon, 1886 (Lehtinen 1967). Palicanus caudatus Thorell, 1897 is notable because of its mouse-grey appearance (Deeleman-Reinhold 2001) and is currently known to occur in China, Indonesia, Myanmar, Reunion Island and Seychelles (World Spider Catalog 2019). Gravely (1931) described Systaria barkudensis (Gravely, 1931), originally placed under Syrisca Simon, 1886, on the basis of specimens collected from the Indian state of Odisha. The species was redescribed once (Majumder & Tikader 1991) and later it was transferred to Systaria Simon, 1897 (Deeleman-Reinhold 2001). In this paper, we synonymise S. barkudensis with P. caudatus and provide supplementary information on the somatic morphology and male genitalia with detailed illustrations.

Spiders are thought to play a significant role in limiting pest outbreaks in agroecosystems such as vineyards, orchards and cotton. The diversity and impact of spiders in vegetable crops are less well understood, although there is evidence that predators may be important for suppression of lepidopteran pests in Brassica crops, particularly early in the season before parasitoids become established. Sampling was conducted in early season plantings of Brassicas in the Lockyer Valley (South East Queensland, Australia) in order to determine the most commonly occurring spider families. The most numerous were Theridiidae, which were more strongly associated with cauliflower and poorly associated with cabbage. The Lycosidae and Clubionidae/Miturgidae (formerly in the ‘catch-all’ family Clubionidae) also occurred commonly. Lycosidae (and to a lesser extent Salticidae) had above average abundance in Chinese cabbage and below average abundance in broccoli compared with average abundance for these spider families; Clubionidae/Miturgidae had above average abundance in cauliflower. Laboratory studies were then conducted to explore the predatory capacity of these three most commonly occurring spider families. All three were capable of feeding on larvae of the diamondback moth, Plutella xylostella (Linnaeus), and cabbage cluster caterpillar, Crocidolomia pavonana (Fabricius), under laboratory conditions. Theridiidae, which are thought to prey on small pests such as leafhoppers and aphids, were able to successfully attack larvae up to five times their body size. Predation rates varied from an average of 1.7 (SE = 0.47) (1.6 control corrected) larvae consumed over a 24 h period in the case of the Theridiidae, to 3.3 (SE = 0.60) larvae for the Clubionidae/Miturgidae.

To date, 24 spider families have been identified on Reunion Island. But published data on spider fauna mainly concerned natural habitats and highlands. Studies of predatory arthropods in agroecosystems provided the opportunity to conduct a thorough investigation of spider diversity. Seven species and families of spiders are newly recorded for Reunion Island : Creugas gulosus Thorell, 1878 (Corinnidae), Oedignatha scrobiculata Thorell, 1881 (Liocranidae), Ero aphana (Walckenaer, 1802) (Mimetidae), Palicanus caudatus Thorell, 1897 (Miturgidae), Oecobius marathaus Tikader, 1962 (Oecobiidae) and Orthobula impressa Simon, 1897 (Phrurolithidae) and Tropizodium peregrinum Jocqué & Churchill, 2005 (Zodariidae)., Sept espèces et familles d’Araignées nouvelles pour la Réunion (Araneae, Araneomorphae). Jusqu’à aujourd’hui, 24 familles d’Araignées avaient été recensées à l’île de la Réunion. Ces données publiées sur l’arachnofaune ont principalement concerné les habitats naturels et d’altitude. Des inventaires des arthropodes prédateurs dans les agroécosystèmes ont permis de compléter les connaissances sur la diversité des Araignées. Ainsi, sept espèces et familles d’Araignées nouvelles pour la Réunion ont été recensées : Creugas gulosus Thorell, 1878 (Corinnidae), Oedignatha scrobiculata Thorell, 1881 (Liocranidae), Ero aphana (Walckenaer, 1802) (Mimetidae), Palicanus caudatus Thorell, 1897 (Miturgidae), Oecobius marathaus Tikader, 1962 (Oecobiidae) and Orthobula impressa Simon, 1897 (Phrurolithidae) et Tropizodium peregrinum Jocqué & Churchill, 2005 (Zodariidae)., Jacquot Maxime, Derepas Brice, Deguine Jean-Philippe. Seven newly recorded species and families of spiders from Reunion Island (Malagasy region) (Araneae, Araneomorphae). In: Bulletin de la Société entomologique de France, volume 121 (4),2016. pp. 421-430.

Diversity of spider groups have received less research attention and there are limited published references for spiders from Peninsular Malaysia. The current survey was conducted to locate and identify foliage-dwelling spiders (Arachnida, Araneae) at five different sites in Peninsular Malaysia. Spider specimens were collected using manual search and sweep-netting between September 2012 and November 2012. A total of 92 morpho-species from 65 genera that belong to 15 families have been successfully recorded and identified. The greatest proportion of specimens captured (40%) were Foliage-runners (Clubionidae, Miturgidae, Oxyopidae, Pisauridae, Salticidae, Scytodidae, Thomisidae), followed by orb-weavers (Araniedae, Nephilidae, Tetragnathidae, Uloboridae) (36.5%), space-weavers (Pholcidae, Psechridae Theridiidae) (21.5%) and ground-dwellers (Sparassidae) (2.0%). Cluster analysis has revealed that the same habitat types share a more similar diversity composition compared to different habitat types, which indicates that spider assemblage composition was partly co-dependent on vegetation structure. However, no significant difference in spider assemblage composition was found between all the five sites which follows that these diurnal group of spiders are actually adaptable to various habitat types.

Eutichurus tropicus (L. Koch, 1866) Map 2 Cheiracanthium tropicum L. Koch, 1866: 239, pl. 10, fig. 155. Eutichurus tropicus; Bonaldo & Brescovit 1992: 738; Bonaldo 1994: 119, figs 46¯47, 108. Eutichurus brescoviti Bonaldo, 1994: 119, figs 43̄45, 108. Syn. n. Note. The present synonymy is justified by the fairly number of specimens from both sexes collected together in Cundinamarca. Diagnosis and description. See Bonaldo (1994: 119). Distribution. Central Colombia (Map 2). New Records. COLOMBIA. Cundinamarca: Mosquera, Mondoñedo, [Desierto de] Zabrinsky [04°40'30.00”N 74°16'05.00”W], 2600 m, 2 females and 3 females (ICN-Ar 2116), IV.2002, I. Martinez, G. Niño, G. Mora. A. Castañeda leg.; Laguna de [La] Herrera [04°41'29.00”N 74°16'36.00”W], 1 male (ICN-Ar 6472), 21.II.2010, D. Luna leg.; Boyacá, SFF Iguaque La Planada, 5°25’N, 73°27’W, 2850 m, 1 male (ICN-Ar 6476), 21.II–VII.2001, P. Reina leg.; San Antonio del Tequendama [04°36'59.00”N 74°02'17.75”W], 1 male (ICN-Ar 6477), II.1996, P.M. Chicaque leg., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ramírez, Martín J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 344-345, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Koch, L. (1866) Die Arachniden-Familie der Drassiden. Hefte 1 ‾ 6. Lotzbeck, Nurnberg, 304 pp.","Bonaldo, A. B. & Brescovit, A. D. (1992) As aranhas do genero Cheiracanthium C. L. Koch, 1839 na regiao neotropical (Araneae, Clubionidae). Revista Brasileira de Entomologia, 36, 731 ¯ 740.","Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus cumbia Bonaldo & Ram��rez, new species Figs 12 ��14, Map 1 Type material. Male holotype (AMNH) from Colombia, Ovalle leg. (Poorly preserved, dehydrated, abdomen missing, legs broken). Etymology. The specific name refers to a music stile typical from Colombia, the c��mbia. Diagnosis. Males of E. cumbia n. sp. resemble those of E. ravidus, E. itamaraju and E. nancyae n. sp. (Figs 58 ��60; Bonaldo 1994: figs 65, 70) by the flattened embolus, with distal prolateral process, but differ by the long MA, with the apex unfolded and prolaterally oriented, and by the RTA with dorsal prong shorter than the ventral prong (Figs 12, 14). Description. Male (Holotype). Colour in ethanol, dried specimen: carapace brownish red, thoracic groove brownish, chelicerae brownish red. Endites, labium and sternum brownish red. Legs orange. Chelicerae with 3 promarginal teeth, the second the largest, the last a denticle, and 4 retromarginal teeth, the first a denticle, separated by twice their width from second, second to fourth separated by their width. Tarsi I���II with moderate scopulae, tarsi III���IV and metatarsi III���IV with dense scopulae. Carapace 6.2 long, 4.9 wide. Eye diameters: AME 0.37, ALE 0.33, PME 0.24, PLE 0.31. Eye interdistances: AME���AME 0.28, AME���ALE 0.33, PME���PME 0.46, PME���PLE 0.68, AME���PME 0.32, ALE���PLE 0.15, clypeus AME 0.24, clypeus ALE 0.28. Leg and palp measurements: palp 6.7 (2.4, 1.0, 1.0, -, 2.3); leg I 20.12 (5.2, 2.5, 5.0, 5.4, 2.5); leg II 19.7 (5.3, 2.4, 4.7, 5.1, 2.2); leg III 15.08 (4.4, 2.0, 3.6, 4.1, 1.7); leg IV 19.9 (5.5, 2.2, 4.5, 5.6, 2.1). Leg formula I-IV-II-III. Spination: palpal femur: d 0-0-2; tibia p 1. Leg. I femur d 1-1-0, p 0-0-1, r 0; tibia d 0, p 0-0, r 0, v 0-2-0; metatarsus d 0, p 0, r 0, v 2-2 -1. II���femur d 0, p 0-0-d1, r 0; tibia d 0, p 0, r 0, v 0-p1-0; metatarsus d 0, p 0, r 0, v 0. III���femur d 1-1-0, p 0-0-d1, r 0; tibia d 0, p 1-1, r 0-1, v 0-p1-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV���femur d 1-1-0, p 0, r 0; tibia d 0, p 0-1, r 1-1, v p1-2-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Palp: Tibia slender, two-thirds the cymbium length; RTA large, bifid, divided at base, C-shaped; MA apically placed, apex unfolded, directed prolaterally, median prong absent; conductor hyaline; embolus inserted sub-basally, not fused to tegulum, flattened, with apical prolateral process (Figs 12 ��14). Female. Unknown. Distribution. Colombia (Map 1). Other material examined. None., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 329-330, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus nancyae Bonaldo & Saturnino, new species Figs 58 ��62, Map 1 Type material. Male holotype (MPEG 21914) from Brazil, Par��, Juruti, V��rzea do Piranha, 02��12'36.1"S 56��07'20.7"W, 22.II.2011, N. F. Lo Man Hung leg. Paratypes: female (MPEG 21915), same locality, 4.VIII.2010, N. Abrahim leg.; male (MPEG 21916), same locality, 25.II.2011, N.F. Lo Man Hung leg.; male (MPEG 21917), Bel��m, Universidade Federal do Par��, Mata do Betina, 01��28'S 48��27'W, 12.IV.2007, D. F. Candiani leg.; female (MPEG 21918), same data; female (MPEG 21919), same locality and collector, 10.IV.2007; Ilha de Cotijuba, [01��13'60���S 48��33'3���W], female (MPEG 4714), XI.2004, D. R. Santos-Souza leg.; female (MPEG 21920), Santar��m, Alter do Ch��o, [02��30'19.00��� S 54��57'8.00���W], 27.IV.2007, B. J.F. Silva leg.; male (MCN 25014), same locality, 26.I.1994, H. H��fer leg.; male (MPEG 21921), same locality, Fragmento S��o Lu��s, 2.VII.2007; male (MPEG 925), Melga��o, Esta����o Cient��fica Ferreira Penna, Floresta Nacional de Caxiuan��, 01��44'18.02"S 51��27'48.01"W, 14.VII.2002, C. Pereira leg.; female (MZSP 10734), Barreirinhas [02��32'53.00��� S 54��05'9.00���W], Rio Tapaj��s, 24.XI.1970, EPA leg. Etymology. The specific name is a patronym in honour of arachnologist Nancy Fran��a Lo Man Hung, one of the collectors of the types. Diagnosis. Males of E. nancyae n. sp. resemble those of E. ravidus and E. itamaraju by the wide, subrectangular embolus and by the simple MA without median prong (Bonaldo 1994: figs 65, 70). They differ from both species by the simple, undivided RTA, with small ventral, proximal lamellae (Figs 58, 60). Females share with those of E. itamaraju the epigynal anterior projections extremely reduced (Bonaldo 1994: fig. 73) but are promptly recognized by the median field wider than long and by the posterior plates medially constricted (Fig. 61). Description. Male (Holotype). Colour in ethanol: carapace pale yellow, thoracic groove mark darker. Chelicerae, endites and labium brownish red. Sternum yellow with brown margins. Legs and abdomen pale yellow. Chelicerae with 4 promarginal teeth, the third the largest, and 4 retromarginal teeth, contiguous. Tarsi I���IV and metatarsi III���IV with moderate scopulae. Total length 8.9. Carapace 4.0 long, 3.0 wide. Eye diameters: AME 0.25, ALE 0.25, PME 0.22, PLE 0.23. Eye interdistances: AME���AME 0.22, AME���ALE 0.23, PME���PME 0.35, PME��� PLE 0.33, AME���PME 0.21, ALE���PLE 0.04, clypeus AME 0.08, clypeus ALE 0.12. Leg and palp measurements: palp 4.8 (1.7, 0.6, 1.0, -, 1.5); leg I 17.8 (4.7, 1.6, 4.7, 5.0, 1.8); leg II 17.5 (4.7, 1.5, 4.6, 5.0, 1.7); leg III 13.6 (3.9, 1.4, 3.2, 3.7, 1.4); leg IV 16.3 (4.5, 1.4, 3.9, 4.9, 1.6). Leg formula I-II-IV-III. Spination: palpal femur: d 0-1-1, p 0- 0-1; tibia p 2-1. Legs. I femur d 1-1-0, p 0-d1-d2, r 0-d1-d1; tibia d 0, p 1-1, r 0, v 2 -p1-2-r1-p1-2 (v 2 p1-2r1-p12); metatarsus d 0, p 0, r 0, v 2 -r1-2-p1-1 (2-r12-p1ap1). II femur d 1-0-0, p 0-d1-d1, r 0-d1-d1; tibia d 0, p 1-1, r 0, v 2-2 -2; metatarsus d 0, p 0, r 0, v 2-2 -1. III���femur d 1-0-0, p 0-d1-d1, r 0-d1-d2; tibia d 0, p 1-1, r 1-1, v p1-2-2; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV���femur d 1-1-0, p 0-d2-d1, r 0-d1-d2; tibia d 0, p 1-3, r 1-0, v 2-2 -2-2-2; metatarsus d p1r1-p2-p1r1, p 0-0-1, r 0-0-1, v r1-2-2-2-r1-1 (r12-4r1-ap1). Palp: tibia slender, almost the same length of cymbium; RTA unbranched, relatively long and thin, with a basal, ventral lamellae. MA apically placed, without median prong; conductor hyaline; embolus inserted medially, wide, short, with large prolateral process and pars pendula (Figs 58 ��60). Female (Paratype, MPEG 21915). Colour in ethanol: dark yellow orange, thoracic groove mark orange. Chelicerae, endites and labium brownish red. Sternum orange with brown margins. Legs yellow, tibia and metatarsi orange. Abdomen pale yellow. Chelicerae with 5 promarginal teeth, the fourth the largest, and 4 retromarginal teeth, contiguous. Tarsi I���IV and metatarsi I���II with dense scopulae; metatarsi III���IV with moderate scopulae. Total length 14.0. Carapace 5.3 long, 4.0 wide. Eye diameters: AME 0.31, ALE 0.28, PME 0.25, PLE 0.30. Eye interdistances: AME���AME 0.39, AME���ALE 0.43, PME���PME 0.57, PME���PLE 0.56, AME���PME 0.36, ALE���PLE 0.13, clypeus AME 0.17, clypeus ALE 0.15. Leg and palp measurements: palp 5.6 (2.0, 0.7, 1.3, -, 1.6); leg I 18.4 (4.9, 2.0, 4.6, 5.1, 1.8); leg II 18.0 (4.9, 2.0, 4.5, 4.9, 1.7); leg III 14.2 (4.3, 1.7, 3.1, 3.7, 1.4); leg IV 17.8 (4.9, 1.7, 4.1, 5.3, 1.8). Leg formula I-II-IV-III. Spination: palp femur: d 0-1-1; tibia p 1-1, tarsus 1-1. Leg. I femur d 1-0-0, p 0-0-1, r 0; tibia d 0, p 0, r 0, v p1-2-0; metatarsus d 0, p 0, r 0, v 2 -0-1. II���femur d 1-0-0, p 0-d1-d1, r 0; tibia d 0, p 0, r 0, v 0-p1-0; metatarsus d 0, p 0, r 0, v 2 -p1-1. III���femur d 1-0-0, p 0-d1-d1, r 0-d1-d2; tibia d 0, p 1-1, r 1-1, v p1-p1-0; metatarsus d p1r1-r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV���femur d 1-0-0, p 0-d1-d1, r 0-0-d2; tibia d 0, p 1-1, r 1-1, v p1-2-r1; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Epigynum: median field wider than long; anterior projections extremely reduced, inconspicuous; posterior plates separated medially, both anterior and posterior margins constricted medially; primary spermathecae inconspicuous, with large fertilization ducts; copulatory ducts wide, globose; secondary spermathecae small, globose (Figs 61, 62). Distribution. Eastern Brazilian Amazonia (Map 1). Other material examined. BRAZIL. Par��: Juruti, v��rzea do Rio Amazonas, 02��24'20.7''S 56��26'00.3''W, 2 females (MPEG 21922; MPEG 21923), 18.IX.2002, A.B. Bonaldo leg.; 02��12'36.1"S 56��07'20.7"W, 1 male (MPEG 21924), 22.II.2011, N.F. Lo Man Hung leg.; 02��12'39.2" S 56��07'12.7" W, 1 male (MPEG 21925), 24.VIII.2011, N.C. Bastos leg.; Santar��m, Alter do Ch��o [02��30'19.00��� S 54��57'8.00���W], 1 female (MPEG 21926), 18.VI.2007, M.L.N. Sirotheau leg. MAP 1. Distribution records of species of Eutichurus., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 335-341, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus putus O. Pickard-Cambridge, 1898 Map 2 Eutichurus putus O. Pickard-Cambridge, 1898: 237, pl. 34, fig. 3; Bonaldo 1994: 133, figs 89��93, 109; Schmidt & Osmani 2012: 20, fig. 5. Eutichurus ecuadorensis Schmidt, 1971: 409, fig. 14; Bonaldo, 1994: 133 (Syn.). Diagnosis and description. See Bonaldo (1994: 133) Distribution. Peru, Panam��, Ecuador, Colombia, northern Brazil (Map 1). New record. PERU. Loreto: Rio Samiria [05��12���38���S, 74��35���36���W], 1 male (MUSM), 15.V.1990, Erwin et al. leg., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on page 342, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Pickard-Cambridge, O. (1898) Arachnida. Araneida. Biologia Centrali-Americana, Zoology, 1, 233 ¯ 288. https: // doi. org / 10.1111 / j. 1096 - 3642.1898. tb 03159. x","Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159.","Schmidt, G. & Osmani, J. (2012) Zur Stellung der Gattung Cheiracanthium C. L. Koch, 1839 im Systen (Araneae: Araneomorphae). Arthropoda Scientia, 2, 14 ¯ 20."]}

Eutichurus lizeri Mello-Leit��o, 1938 Map 2 Eutichurus lizeri Mello-Leit��o, 1938: 114, fig. 34; Bonaldo, 1994: 117, figs 38��42, 110. Eutichurus birabeni Mello-Leit��o, 1941: 175, fig. 65, pl. 10, fig. 46; Bonaldo, 1994: 117 (Syn.). Diagnosis and description. See Bonaldo (1994: 117). Distribution. South Bolivia, North and Central Argentina (Map 2). New record. ARGENTINA. Sunehal [probably El Sunchal 26��37'0.00'' S 65��03'60.0''W, Tucum��n Province], 1 male (MCZ), Lockerell leg., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 341-342, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Mello-Leitao, C. F. de (1938) Algunas aranas nuevas de la Argentina. Revista del Museo de La Plata, New Series, 1, 89 ¯ 118.","Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159.","Mello-Leitao, C. F. de (1941) Las aranas de Cordoba, La Rioja, Catamarca, Tucuman, Salta y Jujuy colectadas por los Profesores Biraben. Revista del Museo de La Plata, New Series, Zoology, 2, 99 ¯ 198."]}

Eutichurus murgai Bonaldo & Lise, new species Figs 4 ̄8, Map 1 Type material. Male holotype (MUSM-Ent 502099) from Peru, Cajamarca, Parque Nacional Cutervo, Puente del Suro, 6°12'10"S, 78°44'22"W, 2438 m, 22.V.2010, R. Pinto da Rocha, M. Alvarado & D. Silva leg.; two males and one female paratypes (MUSM-Ent 502099), same data as the holotype; male paratype (MUSM-Ent 501775) from Peru, Cajamarca, San Pedro, 6°16'51"S, 78°49'55"W, 2428 m, 23.V.2010. Etymology. The specific name is a patronym in honour of biologist and educator Salomón Vílchez Murga, who created the Cutervo National Park in 1961, the oldest national park in Peru. Diagnosis. Males of E. murgai n. sp. are similar to those of E. palatanga by the long, smooth, distally rounded MA median prong (Bonaldo 1994: figs 32̄34), differing by the RTA distally unbranched (Figs 4, 6). They also resemble males of E. zarate (Figs 9 ̄11) by the RTA distally wide, but can be distinguished by the unsclerotized, hyaline conductor (Fig. 4). Females resemble those of E. saylapampa by the epigynum with triangular anterior projections and those of E valderramai by the distally sclerotized anterior projections (Bonaldo 1994: figs 50, 52), but can be distinguished by the combined presence of anterior projections with acute apices and posterior margin nearly straight (Fig. 7). Description. Male (Holotype, MUSM 502099). Colour in ethanol: carapace brownish red, cephalic region darker, thoracic groove mark reddish, chelicerae brownish red. Endites and labium brownish red with black margins. Sternum orange with brown margins. Legs orange. Abdomen grey with some pale-yellow spots. Chelicerae with 3 promarginal teeth, the second the largest, 4 retromarginal teeth, separated by their width. Tarsi I– II with moderate scopulae, tarsi III–IV and metatarsi III–IV with dense scopulae. Total length 10.4. Carapace 4.9 long, 3.8 wide. Eye diameters: AME 0.23, ALE 0.31, PME 0.25, PLE 0.27. Eye interdistances: AME–AME 0.25, AME–ALE 0.29, PME–PME 0.31, PME–PLE 0.50, AME–PME 0.29, ALE–PLE 0.13, clypeus AME 0.16, clypeus ALE 0.20. Leg and palp measurements: palp 6.1 (2.3, 0.8, 1.0, -, 2.0); leg I 18.0 (4.7, 1.9, 4.5, 4.5, 2.4); leg II 16.4 (4.5, 1.7, 4.0, 4.2, 2.0); leg III 14.0 (3.8, 1.8, 3.1, 3.4, 1.9); leg IV 18.0 (5.0, 1.9, 4.2, 4.9, 2.0). Leg formula I=IV-II- III. Spination: palpal femur: d 0-0-2; tibia p 1. Leg. I femur d 1-1-0, p 0-0-1, r 0-d1-d1; tibia d 0, p 0-0, r 0, v 0-2-0; metatarsus d 0, p 0, r 0, v 2-2 -1. II—femur d 0, p 0-d1-d2, r 0-d1-d1; tibia d 0, p 0, r 0, v 0; metatarsus d 0, p 0, r 0, v 0. III—femur d 1-1-0, p 0-d1-d1, r 0-0-d2; tibia d 0, p 1-1, r 1-1, v p1-2-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV—femur d -1-0, p 0-d1-d1, r 0-0-d1; tibia d 0, p 1-0, r 1-1, v p1-2-0; metatarsus d p1r1-p1r1- p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Palp: tibia slender, nearly half the cymbial length; RTA single, strongly sclerotized, widened distally, tip projected ventrally; MA apically placed, median prong rounded apically; conductor hyaline, long; embolus not fused to tegulum, inserted basally, filiform, without processes (Figs 4 ¯6). Female (Paratype, MUSM 502099). Colour in ethanol: carapace brownish orange, thoracic groove reddish, chelicerae, endites and labium brownish red. Sternum yellow with black margins. Legs yellow. Abdomen brown. Chelicera with 3 promarginal teeth, the second the largest, the first a denticle, and with 4 retromarginal teeth closely placed. Tarsi I–IV and metatarsi I–II with dense scopulae; metatarsi III–IV with moderate scopulae. Total length 10.9. Carapace 5.3 long, 4.3 wide. Eye diameters: AME 0.24, ALE 0.34, PME 0.21, PLE 0.28. Eye interdistances: AME–AME 0.34, AME–ALE 0.30, PME–PME 0.37, PME–PLE 0.61, AME–PME 0.34, ALE–PLE 0.13, clypeus AME 0.20, clypeus ALE 0.22. Leg and palp measurements: palp 5.85 (2.0, 0.91, 1.3, -, 1.8); leg I 16.6 (4.6, 2.2, 3.8, 4.1, 1.9); leg II 15.3 (4.3, 2.0, 3.5, 3.7, 1.8); leg III 12.5 (3.7, 1.8, 2.6, 3.0, 1.4); leg IV 16.5 (4.7, 2.0, 3.7, 4.4, 1.7). Leg formula I-IV-II-III. Spination: palpal femur: d 0-0-2; tibia p 1. Leg. I femur d 1-1-0, p 0-0-1, r 0; tibia d 0, p 0-0, r 0, v 0-p1-0; metatarsus d 0, p 0, r 0, v 2-2 -1. II—femur d 0-1-1, p 0-0-1, r 0; tibia d 0, p 0, r 0, v 0-p1-0; metatarsus d 0, p 0, r 0, v 2-2 -1. III—femur d 1-1-0, p 0-d1-d1, r 0-0-d2; tibia d 0, p 1-1, r 1-1, v 0-p1-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV—femur d -1-1, p 0-0-d1, r 0-0-d1; tibia d 0, p 1-0, r 1-1, v p1-2-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Epigynum: Median field longer than wide; Anterior projections large in relation to posterior plates, with acute, sclerotized apices; posterior plates separated medially, with strongly divergent anterior margins; primary spermathecae large, secondary spermathecae small, tube-like, superposed to primary spermathecae; fertilization ducts inserted adjacently to insertion of secondary spermathecae (Figs 7, 8). Distribution. Northern Peru. Other material examined. None.

Eutichurus tequendama Bonaldo & Lise, new species Figs 20 ��22, Map 1 Type material. Male holotype (ICN-Ar 6475) from Colombia, Cundinamarca, San Antonio del Tequendama [04��36���59.21���N 74��21���08.00���W], Bosque El Ermita��o, 12.XII.1997, 2000 m, Santiago leg. Diagnosis. Males of E. tequendama resemble those of E. tropicus (L. Koch) and E. valderramai Bonaldo by the short, excavated RTA, the extensive, sclerotized, lowered prolatero-apical sector of tegulum and by the long median prong of MA (Bonaldo 1994: 43, 48). They differ by the absence of a longitudinal keel on the median prong of MA, which is apically bifid (Figs 20, 22). Etymology. The specific name is a noun in apposition taken from the type locality. Description. Male (Holotype). Colour in ethanol: carapace brownish red, thoracic groove mark darker. Chelicerae, endites and labium dark brownish red. Sternum brownish yellow, with brown margins. Legs orange. Abdomen dark gray. Chelicerae with 3 promarginal teeth, the second the largest, and 4 retromarginal teeth, separated by their length. Tarsi I���IV and metatarsi I���II with dense scopulae, and metatarsi III���IV with moderate scopulae. Total length 14.2. Carapace 7.0 long, 5.2 wide. Eye diameters: AME 0.36, ALE 0.35, PME 0.28, PLE 0.34. Eye interdistances: AME���AME 0.32, AME���ALE 0.43, PME���PME 0.50, PME���PLE 0.73, AME���PME 0.34, ALE���PLE 0.15, clypeus AME 0.25, clypeus ALE 0.32. Leg and palp measurements: palp 8.1 (2.8, 1.0, 1.3, -, 3.0); leg I 23.0 (6.0, 2.8, 5.5, 6.0, 2.7); leg II 22.1 (5.9, 2.7, 5.3, 5.7, 2.5); leg III 17.7 (5.0, 2.2, 4.0, 4.6, 1.9); leg IV 21. 3 (6.0, 2.5, 4.8, 5.8, 2.2). Leg formula I-IV-II-III. Spination: palp femur: d 1-1-0; tibia p 1-1. Leg. I femur d 1-1-0, p 0-0-1, r 0; tibia d 0, p 0, r 0, v p1-2-0; metatarsus d 0, p 0, r 0, v 2 -p1-1. II���femur d 1-1-0, p 0-0-d1, r 0; tibia d 0, p 0, r 0, v 0-2-0; metatarsus d 0, p 0, r 0, v 2 -p1-1. III���femur d 1-1-0, p 0-d1-d1, r 0-d1-d1; tibia d 0, p 1-1, r 1-1, v 0-2-2; metatarsus d p1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV���femur d 0-1-1, p 0-0-d1, r 0-0-d1; tibia d 0, p 0- 1, r 1-1, v 0-2-0; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Palp: tibia short, thin; RTA short, excavated, rectangular; MA sub-apically placed, median prong long, apically bifid; conductor hyaline; embolus inserted medially, not fused to tegulum, thick, with sub-apical prolateral prong and extensive pars pendula (Figs 20 ��22). Female. Unknown. Distribution. Central Colombia (Map 1). Other material examined. None., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 331-332, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus silvae Bonaldo, 1994 Map 2 Eutichurus silvae Bonaldo, 1994: 112, figs 10��11, 16��17, 22��25, 110. Diagnosis and description. See Bonaldo (1994: 112). Distribution. Ecuador, North and Central Peru (Map 2). New records. PERU. Cajamarca: San Benito [07��25'29.00���S 78��55'40.00���W], 1300 m, 1 female (MUSM-Ent 504408), 1.VII.1993, P. Hocking leg.; Lima, [Reserva Nacional Lomas de Lachay [11��21'30.00���S 77��22'10.00���W], 1 female (MUSM-Ent 504409), 27.IX.1998, K. Caro leg.; Jes��s Mar��a [12��04'30.00���S 77��02'36.00���W], 1 female (MUSM-Ent 504413), 27.II.2003, W. Paredes leg. MAP 2. Distribution records of species of Eutichurus., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on page 344, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus zarate Bonaldo, 1994 Figs 9 ��11, Map 1 Eutichurus zarate Bonaldo, 1994: 123, figs 54��55, 109. Female holotype from Peru, Lima, Bosque de Z��rate (Gigant��n), 2860 m, 20.IV.1980, I. Franke leg., [11��55'53.6"S 76��28'22.0"W], (MUSM VF5093, not re-examined). Diagnosis. Males of E. zarate resemble those of E. murgai (Figs 4, 6) by the wide RTA apex, but differ by the sclerotized conductor (Figs 9, 11). Females are promptly recognized by the excavated anterior projections of the epigynum, with digitiform apices, and by the laterally constricted posterior plates (Bonaldo 1994: fig. 54). Description. Male (MUSM 504416). Colour in ethanol: carapace orange, thoracic groove mark reddish. Chelicerae, endites and labium brownish red. Sternum yellow with brown margins. Legs yellow. Chelicerae with 3 promarginal teeth, the second the largest, and 4 retromarginal teeth separated by twice their width. Abdomen pale yellow. Tarsi I���IV with moderate scopulae, and metatarsi I���II with sparse scopulae. Total length 12.0. Carapace 5.4, long, 4.0 wide. Eye diameters: AME 0.24, ALE 0.27, PME 0.23, PLE 0.22. Eye interdistances: AME���AME 0.25, AME���ALE 0.21, PME���PME 0.31, PME���PLE 0.45, AME���PME 0.23, ALE���PLE 0.11, clypeus AME 0.17, clypeus ALE 0.19. Leg and palp measurements: palp 5.3 (2.3, 0.9, 1.1, -, 2.0); leg I 19.4 (5.1, 2.3, 5.0, 5.0, 2.0); leg II 18.1 (4.9, 2.1, 4.5, 4.7, 1.9); leg III 14.3 (4.1, 1.8, 3.2, 3.7, 1.5); leg IV 19 (5.3, 2.0, 4.5, 5.4, 1.8). Leg Formula I- IV-II-III. Spination: palpal femur: d 0-1-1; tibia p 1-1. Leg. I femur d 1-1-0, p 0-0-1, r 0; tibia d 0, p 0, r 0, v p1-2- 0; metatarsus d 0, p 0, r 0, v 2-2 -1. II���femur d 1-1-0, p 0-d1-d1, r 0; tibia d 0, p 0-0-1, r 0, v p1-2-0; metatarsus d 0, p 0, r 0, v 2-2 -1. III���femur d 1-1-0, p 0-d1-d2, r 0-d1-d1; tibia d 0, p 1-1, r 1-1, v 2-2 -0; metatarsus d p1r1-p2r1- p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV���femur d 0-1-1, p 0-0-d1, r 0-0-d1; tibia d 0, p 1-1, r 1-1, v 2-2 -0; metatarsus d p1r1-p1r1-p1r1, p 0-1-1, r 0-0-1, v 2-2 -1. Palp: Tibia strong, half the cymbium length; RTA single, large, distally wide, apex projected dorsally; tegulum with bifurcated projection near MA insertion; MA apically placed, median prong small, with rounded distal margin; conductor with hyaline base, distally a sclerotized lamella, gently curved retrolaterally; embolus inserted medially, not fused to tegulum, filiform, with narrow base, without processes (Figs 9 ��11). Female. Described by Bonaldo (1994: 123). Distribution. Central Peru (Map 1). New Records. PERU. Lima: Bosque de Z��rate [11��55'53.6"S 76��28'22.0"W], Gatero leg., 1 male (MUSM 504416); Province Huarochir��, Matucana [11��50'25.00"S 76��22'46.5"W], Marachanca, 1 female (UNALM 150), 17.IX.2001, M. Deza & I. Rivera leg. Distribution. Peru., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on pages 328-329, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159."]}

Eutichurus ibiuna Bonaldo, 1994 Figs 63��65, Map 2 Eutichurus ibiuna Bonaldo 1994: 125, figs 62��64, 110, male holotype from Brazil, S��o Paulo, Ibi��na (29o39���20" S 47o13���32" W), IBSP 16268, not re-examined; Laborda & Sim�� 2015: 591, figs 1��4. Diagnosis and description. See Bonaldo (1994: 125), male, and Laborda & Sim�� (2015: 591), female. Note. The females here attributed to E. ibiuna differ slightly from those described by Laborda & Sim�� (2015) from Uruguay, in the relative length of the anterior epigynal projections, longer in the specimens from Brazil. The epigynal median field of these specimens is longer than wide, while in the specimens from Uruguay it is nearly as long as wide (Figs 63��65; Laborda & Sim�� 2014: fig. 1D, E). These differences are here considered to be intraspecific. Distribution. South and Southwestern Brazil; Uruguay (Map 2). New Records. BRAZIL. S��o Paulo: ��guas de Santa B��rbara [22��49'50''S 49��16'4''W], 1 female (IBSP 4925), I.1989, I. Shimura leg.; Paran��: Pinh��o, Usina Hidrel��trica Segredo [currently Usina Hidrel��trica Governador Ney Aminthas de Barros Braga, 25��47'36''S 52��6'43''W], Ribeir��o Estreito, 1 female (MHCI), III.1992, R. Pinto da Rocha leg., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on page 341, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159.","Laborda, A. & Simo, M. (2015) Description of the female of Eutichurus ibiuna Bonaldo, 1994 (Araneae: Eutichuridae) with notes on natural history and sexual behavior. Zootaxa, 4021 (4), 591 ¯ 596. https: // doi. org / 10.11646 / zootaxa. 4021.4.9"]}

Eutichurus ravidus Simon, 1897 Map 2 Eutichurus ravidus Simon, 1897a: 501; Bonaldo 1994: 125, figs 62��64, 110. Philisca pallida Mello-Leit��o, 1943: 219, fig. 46; Bonaldo 1994: 125 (Syn.). Diagnosis and description. See Bonaldo (1994: 125). Distribution. South and southeast Brazil, Paraguay and northern Argentina (Map 2). New records. BRAZIL. Rio Grande do Sul: Viam��o, Fitot��cnica [30��02���10.00���S, 51��01���17.00���W], male (MCTP 5543), 23.IX.1994, A.A. Lise leg.; male (MCTP 5572), 7.X.1994; male (MCTP 7961), 7.VII.1995, L.A. Bertoncello & A.A. Lise leg.; male (MCTP 8148), 17.X.1995, A.A. Lise et al. leg.; Gua��ba [30��06���32.00���S, 51��19���25.00���W], Fazenda S��o Maximiliano, male (MCTP 6905), 24.X.1995; Charqueadas [29��57���44.00���S, 51��37���46.00���W], male (MCTP 11083), 17.II.2000, F. Diefenthaeler & M. Vieira leg.; Novo Cabrais, Parque Witeck [29��46���59.00���S, 52��58���18.00���W], male (MCTP 28336), R.G. Buss leg., 19.X.2007; male (MCTP 28434), 2.XI.2007., Published as part of Bonaldo, Alexandre B., Lise, Arno A., Ram��rez, Mart��n J. & Saturnino, Regiane, 2018, Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions, pp. 321-346 in Zootaxa 4382 (2) on page 342, DOI: 10.11646/zootaxa.4382.2.6, http://zenodo.org/record/1182079, {"references":["Bonaldo, A. B. (1994) A subfamilia Eutichurinae na regiao Neotropical, com a revisao do genero Eutichurus Simon, 1896 (Araneae, Miturgidae). Iheringia, 76, 101 ¯ 159.","Mello-Leitao, C. F. de (1943) Catalogo das aranhas do Rio Grande do Sul. Arquivos do Museu Nacional do Rio de Janeiro, 37, 147 ¯ 245."]}

Eutichurus paredesi Bonaldo & Saturnino, new species Figs 1 ̄3, Map 1 Type material. Male holotype (MUSM-Ent 504411) and male paratype (MUSM-Ent 511659) from Peru, Amazonas, Nueva Esperanza, Rodriguez de Mendoza Province, 6°24'27.5"S 77°27'31.4"W, 1493 m, 1.XII.2004, W. Paredes leg. Etymology. The specific name is a patronym in honour of arachnologist Williams Paredes, collector of the types. Diagnosis. Males of E. paredesi n. sp. resemble those of E. abiseo by the embolar base fused to the tegulum (Figs 1, 3; Bonaldo, 1994: figs 35¯37), but differ by the longer, filiform embolus and by the RTA excavated (Figs 1, 2). They also resemble those of E. madre Bonaldo by the wide, retrolaterally directed median prong of the MA (Fig. 23), but are readily recognized by the embolus not articulated, without processes. Description. Male (Holotype). Colour in ethanol: carapace dark orange, thoracic groove darker. Chelicerae, endites and labium brownish red. Sternum orange with brown margins. Legs orange. Chelicerae with 3 promarginal teeth, second largest, and 3 contiguous retromarginal teeth. Tarsi I–IV and metatarsi I–II with dense scopulae; metatarsi III–IV with sparse scopulae. Total length 10.4. Carapace 5.0 long, 3.6 wide. Eye diameters: AME 0.29, ALE 0.27, PME 0.22, PLE 0.24. Eye interdistances: AME–AME 0.26, AME–ALE 0.26, PME–PME 0.39, PME– PLE 0.09, AME–PME 0.24, ALE–PLE 0.44, clypeus AME 0.15, clypeus ALE 0.12. Legs and palp measurements: palp 5.5 (2.0, 0.7, 1.0, -, 1.8); leg I 18.7 (4.9, 1.9, 4.7, 5.2, 2.0); leg II 17.8 (4.7, 2.0, 4.4, 4.8, 1.9); leg III 13.7 (3.9, 1.7, 3.0, 3.6, 1.5); leg IV 17.8 (4.9, 1.9, 4.1, 5.2, 1.7). Leg formula I–II=IV–III. Spination: palpal femur: d 1-1-0; tibia p 1-1. Leg I femur d 1-1-0, p 0-0-1, r 0; tibia d 0, p 0, r 0, v 2-2 -2; metatarsus d 0, p 0, r 0, v 2-2 -1. II—femur d 1-1-0, p 0-d1-d1, r 0; tibia d 0, p 0, r 0, v 2-2 -2; metatarsus d 0, p 0, r 0, v 2-2 -1. III—femur d 1-1-0, p 0-d1-d1, r 0-d1-d2; tibia d 0, p 1-1, r 1-1, v 2-2 -2; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. IV—femur d 1-1-0, p 0-d1-d1, r 0-0-d1; tibia d 0, p 1-1, r 1-1, v p1-2-2; metatarsus d p1r1-p1r1-p1r1, p 0-0-1, r 0-0-1, v 2-2 -1. Palp: tibia slender, two-thirds the cymbium length; RTA short, distally wide, sub-rectangular; MA apically placed, with long median prong gently curved retrolaterally; conductor hyaline; embolus inserted basally, conical, filiform, without processes, posterior surface hyaline (Figs 1 ̄3). Female. Unknown. Distribution. Northern Peru (Map 1). Other material examined. None.