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Natural disturbance‐based management (NDBM) is hypothesized to maintain managed forest ecosystem integrity by reducing differences between natural and managed forests. The effectiveness of this approach often entails local comparisons of species composition or diversity for a variety of biota from managed and unmanaged forests. Understory vegetation is regularly the focus of such comparison because of its importance in nutrient cycling, forest regeneration, and for wildlife. However, larger scale comparisons between regions with distinct species assemblages may require a trait‐based approach to better understand understory responses to disturbance. We compared the long‐term effects of retention harvesting on understory vegetation in two large experimental study sites located in eastern and western regions of the Canadian boreal forest. These sites included the Sylviculture en Aménagement Forestier Ecosystémique (SAFE) experiment and the Ecosystem Management Emulating Natural Disturbance (EMEND) experiment, located in the eastern and western regions of Canada, respectively. EMEND and SAFE share common boreal understory species but have distinct tree communities, soils, and climate. Both experiments were designed to evaluate how increasing tree retention after harvest affects biodiversity. Here, we examined taxonomic richness, functional diversity, and functional composition (using community trait mean values) of understory plant communities, and also examine intraspecific trait variability (ITV) for five species common and abundant in both experiments. We observed the limited impacts of retention level on richness, functional diversity, and functional composition of understory plants 20 years postharvest. However, ITV of leaf morphological traits varied between retention levels within each experiment, depending on the species identity. Common species had different functional responses to retention level, showing species‐specific reactions to environmental variation. Our result suggests that understory plant communities in the boreal forest achieve resilience to disturbance both in terms of interspecific and intraspecific functional trait diversity. Such diversity may be key to maintaining understory biodiversity in the face of future disturbances and environmental change. Our results reveal the significance of ITV in plant communities for understanding responses to forest harvesting and the importance of choosing appropriate traits when studying species responses to the environment. [ABSTRACT FROM AUTHOR]

The most rapid climate warming is occurring in northern, permafrost environments. Peatlands of these regions are particularly sensitive to climate warming, with the high ground ice content of peat plateaux resulting in the formation of collapse scars as ground temperatures warm.To quantify the rates of permafrost thaw and associated vegetation changes, we sampled the plant and lichen communities in transects spanning actively thawing collapse scar margins at sites from mid‐Boreal to Low Subarctic conditions, within Canada's Northwest Territories. Seventeen transects were sampled in 2007/08 and 14 of these were resampled 10 years later.The rate of lateral permafrost thaw of collapse scar margins ranged from −6 to 63 cm year−1 (mean: 22.0 cm ± 4.7 cm year−1 (SE)); variability was high and no trends with respect to latitude or temperature gradients were detected.Plant communities displayed a clear gradient from lichen‐ and ericaceous shrub‐dominated peat plateaux, to collapse scars primarily characterized by Sphagnum mosses and graminoids. Both space‐for‐time (distance from collapsing margin) and direct measurement of 10‐year changes showed a successional sequence following permafrost thaw; floating mat communities characterized by Sphagnum riparium or S. balticum proceeded to lawn communities of S. angustifolium and, finally, to hummock communities with Mylia anomala or S. fuscum. This successional sequence was associated with increased water table depth and lower soil water content in plant communities farther from the actively collapsing front, illustrating that peat growth above the water table was driving plant community successional changes.The most rapid plant community succession occurred in recently thawed environments as peat growth propelled the ground surface above the water table while the slowest succession occurred in the collapse scar hummock communities located farthest from the actively collapsing peat plateau margin.Synthesis. In just 10 years, significant vegetation change was detected, in association with both permafrost thaw and subsequent plant community succession. These changes occurred across a broad climatic and latitudinal gradient, from mid‐Boreal to Low Subarctic and have implications for wildlife, global C cycle and indigenous communities who depend on this landscape for harvesting, spiritual and cultural practices. [ABSTRACT FROM AUTHOR]

In peatlands, microtopography strongly affects understory plant communities. Disturbance can result in a loss of microtopographic variation, primarily through the loss of hummocks. To address this, mounding treatments can be used to restore microtopography. We examined the effects of mounding on the understory vegetation on seismic lines in wooded fens. Seismic lines are deforested linear corridors (~3 to 8 m wide) created for oil and gas exploration. Our objectives were to compare the recovery of understory communities on unmounded and mounded seismic lines and determine how recovery varies with microtopographic position. Recovery was evident in the unmounded seismic lines, with higher shrub and total understory cover at the 'tops' of the small, natural hummocks than at lower microtopographic positions--much like the trends in adjacent treed fens. In contrast, mounding treatments that artificially created hummocks on seismic lines significantly changed understory communities. Mounded seismic lines had higher forb cover, much lower bryophyte cover, less variation along the microtopographic gradient, and community composition less similar to that of the reference sites than unmounded seismic lines due to higher abundance of marsh-associated species. Our results suggest that mounding narrow seismic lines can be detrimental to the recovery of the understory communities in treed peatlands. Key words: moderate-rich fens, linear disturbances, mounding, restoration, understory vegetation. Dans les tourbieres, la microtopographie influence fortement les communautes vegetales de sous-etage. Une perturbation peut entrainer une baisse de variation de la microtopographie, surtout par la perte de monticules. Pour resoudre ce probleme, la creation de monticules peut etre utilisee pour restaurer la microtopographie. Nous avons etudie les effets de la creation de monticules sur la vegetation de sous-etage dans des lignes sismiques traversant des tourbieres boisees. Les lignes sismiques sont des couloirs lineaires deboises (environ 3 a 8 m de large), crees pour l'exploration petroliere et gaziere. Nos objectifs etaient de comparer le retablissement des communautes de sous-etage dans des lignes sismiques avec et sans creation de monticules, et de determiner comment le retablissement varie en fonction de la position microtopographique. Le retablissement etait manifeste dans les lignes sismiques sans creation de monticules, ou le recouvrement d'arbustes et de vegetation totale sur le dessus de petits monticules naturels etait plus grand qu'aux positions microtopographiques inferieures, ce qui tendait aussi a etre le cas dans les tourbieres boisees adjacentes. En revanche, la creation artificielle de monticules dans les lignes sismiques a considerablement change les communautes de plantes de sous-etage. Les lignes sismiques avec des monticules artificiels avaient un recouvrement de plantes herbacees plus eleve, un recouvrement de bryophytes beaucoup plus faible, moins de variation le long du gradient microtopographique, et une composition de la vegetation moins semblable a celle des stations de reference que celle des lignes sismiques non traitees, en raison d'une plus grande abondance d'especes associees aux marais. Nos resultats indiquent que la creation de monticules dans les lignes sismiques etroites peut nuire au retablissement des communautes de plantes de sous-etage dans les tourbieres boisees. [Traduit par la Redaction] Mots-cles: tourbieres moderement riches, perturbations lineaires, creation de monticules, restauration, vegetation de sous-etage., Introduction Forests across the world are experiencing increases in the levels of human disturbance that are influencing their ecological function and biodiversity (Haddad et al. 2015). In regions with oil [...]

Regionally fitted allometric equations for individual trees and root-to-shoot ratio values are normally used to estimate local aboveground and belowground forest biomass, respectively. However, uncertainties arising from such applications are poorly understood. We developed equations for both aboveground and belowground biomass using destructive sampling for three dominant upland boreal tree species in northwestern Alberta, Canada. Compared with our equations, the diameter-based national equations derived for use across Canada underestimated aboveground biomass for Picea glauca (Moench) Voss but gave reasonable estimates for Populus balsamifera L. and Populus tremuloides Michx. The national equations based on both tree diameter and height overestimated aboveground biomass for the Populus species but underestimated it for Picea glauca in our study area. The approach of root-to-shoot ratio proposed by the Intergovernmental Panel on Climate Change (IPCC) overestimated belowground biomass by 16%-41%, depending on forest cover type, in comparison with our values estimated directly on site, with the greatest bias in deciduous-dominated stands. When the general allometric equations for aboveground biomass and the root-toshoot ratio for belowground biomass were combined to estimate stand biomass, overestimation could be as high as 18% in our study area. The results of our study support the development of improved regional allometric equations for more accurate local-scale estimations. Incorporating intraspecific variation of important traits such as tree taper may be especially helpful. Key words: EMEND (Ecosystem Management Emulating Natural Disturbance), intraspecific trait variation, root-to-shoot ratio, tree taper, wood density. On utilise generalement des equations allometriques ajustees a l'echelle regionale pour les arbres individuels et des valeurs du rapport racines-tiges pour estimer respectivement la biomasse forestiere locale, aerienne et souterraine. Cependant, les incertitudes decoulant de telles applications sont mal comprises. Nous avons developpe des equations de biomasse aerienne et souterraine en utilisant un echantillonnage destructif pour trois especes d'arbres boreales dominantes dans les hautes terres du nord-ouest de l'Alberta, au Canada. Comparativement a nos equations, les equations nationales basees sur le diametre developpees pour etre utilisees partout au Canada sous-estimaient la biomasse aerienne de Picea glauca (Moench) Voss, mais donnaient des estimations raisonnables pour Populus balsamifera L. et Populus tremuloides Michx. Les equations nationales basees sur le diametre et la hauteur des arbres surestimaient la biomasse aerienne des especes de Populus L., mais la sous-estimaient pour Picea glauca dans notre zone d'etude. L'approche du rapport racines-tiges, proposee parle Groupe d'experts intergouvernemental sur l'evolution du climat (GIEC), a surestime la biomasse souterraine de 16 a 41% selon le type de couvert forestier par rapport a nos valeurs estimees directement sur le site, le biais le plus important etant observe dans les peuplements domines par les feuillus. En combinant les equations allometriques generales pour la biomasse aerienne et le rapport racines-tiges pour la biomasse souterraine pour estimer la biomasse du peuplement, la surestimation pouvait atteindre 18% dans notre zone d'etude. Les resultats de notre etude justifient le developpement d'equations allometriques regionales ameliorees pour des estimations plus precises a l'echelle locale. L'incorporation des variations intraspecifiques des attributs importants, tels que le defilement de la tige des arbres, peut etre particulierement utile. [Traduit par la Redaction] Mots-cles : amenagement ecosystemique, variations intraspecifiques des attributs, rapport racines-tiges, defilement de la tige, densite du bois., Introduction Increasing pressure to reduce net carbon dioxide (CO2) emissions in light of climate change is spurring development of global strategies such as carbon markets that encourage carbon offsets through [...]

The recent expansion of mountain pine beetle (MPB, Dendroctonus ponderosae) from its native range in western North America into northern boreal pine forests in Alberta, Canada has resulted in conditions for tree regeneration that are dramatically different from those after wildfire, the predominant natural disturbance in these forests. We assessed natural regeneration post-MPB for northern boreal lodgepole pine sites in Alberta, Canada via intensive surveys of small plots at 33 severely attacked pine stands and using data from 205 permanent sample plots representing various site types and levels of MPB mortality. We used model selection to identify factors explaining regeneration. Overall, pine regeneration was very poor 6–9 years post-MPB; only 42% of the 33 intensively surveyed plots and only 9% of the 205 permanent plots had pine seedlings. This poor regeneration is attributed to high levels of cone serotiny in these populations, unsuitable regeneration microsites due to undisturbed litter or feathermoss layers, and competition from the residual canopy and understory vegetation. Other species (aspen, birch, poplar, and black and white spruce) were found on most sites, either as post-attack regeneration or regeneration established in advance. Without intervention, many of these stands will likely transition away from pine, to broadleaf and other conifer species. [ABSTRACT FROM AUTHOR]

The first issue of the Canadian Journal of Forest Research (CJFR) was published in March 1971 by the National Research Council of Canada with J.L. Farrar, Faculty of Forestry at [...]

Wildfire is the predominant natural disturbance in the boreal forests of western Canada. Natural disturbance based forest management involves the use of retention harvesting to retain stand structural diversity after harvest; however, unlike fire, this partial harvesting technique does not cause combustion of the forest floor. Application of prescribed burning to areas treated with retention harvesting might emulate the influence of wildfires more effectively than harvesting alone. We compared understory vascular plant diversity, abundance, and composition between forest stands subjected to dispersed retention harvesting (10% retention) with and without prescribed burning 1, 6, and 11 or 12 years after burning. Untreated forest was included as a reference. Research was conducted in conifer-dominated, mixedwood, and deciduous-dominated boreal forest stands in northwestern Alberta, Canada. In deciduous-dominated stands, burned areas of retention harvested stands had higher species richness and greater cover than did unburned areas. In all three forest cover types, effects of harvest with and without burn on species richness, cover, and composition were still evident a decade after disturbance. Fire-adapted species benefited most from the prescribed burn treatment. The combination of prescribed burning with retention harvesting can be considered a useful option in forest management that aims to emulate natural disturbance. Key words: prescribed burning, retention harvesting, fire, understory vascular plants, boreal forest. Les perturbations qui dominent dans les forets boreales de l'ouest du Canada sont causees par les feux de foret. L'amenagement forestier qui s'inspire des perturbations naturelles inclut l'utilisation de la coupe avec retention pour conserver la diversite structurale a la suite de la recolte. Cependant, cette methode de coupe partielle n'entraine pas la combustion de la couverture morte comme c'est le cas avec le feu. Le recours au brulage dirige dans les zones soumises a une coupe avec retention pourrait imiter l'effet des feux de foret plus efficacement que la coupe seule. Nous avons compare la composition, l'abondance et la diversite des plantes vasculaires en sous-etage dans des peuplements forestiers soumis a une coupe avec retention des arbres disperses (10 % de retention) avec et sans brulage dirige 1, 6 et 11 ou 12 ans apres le brulage. Une foret non traitee a ete incluse comme reference. Cette etude a ete realisee dans le nord-ouest de l'Alberta, au Canada, dans des peuplements de foret boreale domines soit par des resineux soit par des feuillus et dans des peuplements mixtes. Dans les peuplements domines par des feuillus soumis a une coupe avec retention, la richesse en especes et la couverture etaient plus grandes dans les zones brulees que dans les zones non brulees. Dans les trois types de couvert forestier, les effets de la recolte avec et sans brulage sur la richesse en especes, la couverture et la composition etaient encore evidents 10 ans apres la perturbation. Les especes adaptees au feu ont le plus beneficie du brulage dirige. La combinaison du brulage dirige et de la coupe avec retention peut etre consideree comme une option utile pour l'amenagement forestier qui cherche a imiter les perturbations naturelles. [Traduit par la Redaction] Mots-cles: brulage dirige, coupe avec retention, feu, sous-etage, plantes vasculaires, foret boreale., Introduction Wildfire is the predominant stand-replacing natural disturbance in the boreal forest and it drives the region's vegetation dynamics (Johnson 1992; Payette 1992). The understory vegetation layer hosts most of [...]

The spatial effects of vegetation control on early tree growth were investigated in central Alberta, Canada, for four years after the establishment of hybrid poplar plantations including the two clones Walker (Populus deltoides x (P. laurifolia x P. nigra)) and its progeny Okanese (Walker x (P. laurifolia x P. nigra)). Tree survival and growth, herbaceous vegetation cover, soil nutrient availability, moisture, temperature, and light availability were assessed. Tree growth in the first two years after establishment was improved through selective in-row vegetation control close (within 50 cm) to trees for both aboveground (mechanical) and above- and below-ground (chemical) control. This was associated with increased light availability for trees. In contrast, growth in the third and fourth years benefitted from control of aboveground vegetation within 140 cm of the stem, and this was associated with increased nutrient availability. These findings suggest that the effects of neighbouring vegetation on trees shift from aboveground competition near the tree stem to belowground competition further (>50 cm) away; thus between-row vegetation control is more important starting in the third year after establishment. Okanese outperformed Walker poplar across all treatments and was more responsive to vegetation control, reflecting its superior performance, higher plasticity, and greater potential for short-rotation intensive-culture plantations. Key words: hybrid poplar, tree growth, aboveground and belowground competition, resource availability, vegetation management. Les effets de nature spatiale de la maitrise de la vegetation sur la croissance precoce des arbres ont ete etudies dans le centre de l'Alberta, au Canada, pendant quatre ans suivant l'etablissement de plantations de peupliers hybrides incluant les deux clones Walker (Populus deltoides x (P. laurifolia x P. nigra)) et sa descendance Okanese (Walker x (P. laurifolia x P. nigra)). La survie et la croissance des arbres, le couvert de vegetation herbacee, la disponibilite des nutriments dans le sol, l'humidite, la temperature, et la disponibilite de la lumiere ont ete evalues. Durant les deux premieres annees suivant l'etablissement, la croissance des arbres a ete amelioree par la maitrise selective de la vegetation dans les rangs pres (a moins de 50 cm) des arbres tant dans le cas de la maitrise aerienne (mecanique) qu'aerienne et souterraine (chimique). Cela etait associe a plus grande disponibilite de la lumiere pour les arbres. Par contre, durant les troisieme et quatrieme annees la croissance a beneficie de la maitrise aerienne de la vegetation en dedans de 140 cm de la tige et cela etait associe a une plus grande disponibilite des nutriments. Ces resultats indiquent que les effets de la vegetation avoisinante passent de la competition aerienne pres du tronc des arbres a la competition souterraine plus loin (>50 cm). Par consequent, la maitrise de la vegetation entre les rangs est plus importante a partir de la troisieme annee suivant l'etablissement. Le clone Okanese a ete plus performant que le clone Walker peu importe le traitement et reagissait mieux a la maitrise de la vegetation, refletant sa performance superieure, sa plus grande plasticite et son potentiel plus eleve pour les plantations a culture intensive et courte rotation. [Traduit par la Redaction] Mots-cles: peuplier hybride, croissance des arbres, competition aerienne et souterraine, disponibilite des ressources, gestion de la vegetation., Introduction Plantations of fast-growing hybrid poplar trees play a significant role in wood and fibre production and as a supplier of biomass for energy, alternative fodder sources, and ecosystem services [...]

Given a seed source, the quality of available substrates is a key factor in determining the success of white spruce (Picea glauca (Moench) Voss) natural regeneration. We examined the influence of substrate and competing vegetation on survival and growth of natural regeneration of white spruce up to 4 years following harvesting in deciduous-dominated upland boreal mixedwood sites. Feather moss, thick soil surface organic layers, litter, and solid wood were poor substrates for establishment. Early successional mosses establishing on mineral soil, thin organics, and rotten wood were generally favourable microsites but were not highly available on postharvest sites. Mineral soil substrates were not as suitable as expected, likely because on a postlogged site, they are associated with unfavourable environmental characteristics (e.g., low nutrient availability, exposure). There was some evidence that survival and growth of seedlings were improved by surrounding vegetation in the first years, but heavy competing vegetation had a negative impact on older seedlings. Burial by aspen litter greatly increased seedling mortality, especially when combined with a brief period of submergence due to heavy spring snowmelt. The results provide insight into conditions under which natural regeneration could be an option for establishing white spruce following harvesting of deciduous-dominated boreal mixedwood forests. Key words: natural regeneration, white spruce, substrate, seedling survival, competing vegetation, forest management. Pour une source de graines donnee, la qualite des substrats disponibles est un facteur cle pour le succes de la regeneration naturelle d'epinette blanche (Picea glauca (Moench) Voss). Nous avons etudie l'influence du substrat et de la vegetation concurrente sur la survie et la croissance de la regeneration naturelle d'epinette blanche jusqu'a 4 ans apres la coupe de peuplements mixtes boreaux domines par des especes decidues et etablis sur des stations bien drainees. Les mousses hypnacees, les couches epaisses de matiere organique sur le sol, la litiere et le bois massif constituaient de mauvais substrats pour l'etablissement des semis. Les mousses de debut de succession etablies sur le sol mineral, les couches minces de matiere organique sur le sol et le bois decompose constituaient generalement des microsites favorables, mais etaient peu disponibles sur les stations recoltees. Les substrats formes de sol mineral n'etaient pas aussi appropries que prevu, probablement parce que sur une station qui a ete recoltee, ils sont associes a des caracteristiques environnementales defavorables (p. ex.: faible disponibilite des nutriments, exposition). La survie et la croissance des semis ont montre des signes d'amelioration lies a la presence de vegetation environnante au cours des premieres annees, mais une forte competition exercee par cette vegetation a eu un impact negatif sur les plus vieux semis. L'ensevelissement des semis par la litiere de peuplier a considerablement augmente leur mortalite, particulierement lorsqu'il etait jumele a une breve periode d'immersion causee par une importante fonte des neiges au printemps. Les resultats donnent un apercu des conditions dans lesquelles la regeneration naturelle pourrait constituer une option pour l'etablissement de l'epinette blanche apres la recolte des forets mixtes boreales dominees par des especes decidues. [Traduit par la Redaction] Mots-cles: regeneration naturelle, epinette blanche, substrat, survie des semis, vegetation concurrente, amenagement forestier., Introduction White spruce (Picea glauca (Moench) Voss) is an economically and ecologically important conifer that is widely distributed across the boreal forests of North America. On upland mixedwood sites that [...]

Abstract: Distylous species typically experience self-incompatibility with one morph often having partial selfcompatibility Small populations may therefore experience greater rates of selfing/intramorph crosses leading to skewed morph ratios and reduced [...]

Treed peatlands are dominated by bryophytes, particularly Sphagnum spp., which create the characteristic hummock‐hollow microtopography. This, in turn, shapes the distribution of bryophyte communities. Disturbances can lead to a loss of this microtopographic variation, impacting the bryophyte community. Seismic lines are deforested linear disturbances (approximately 3–8 m wide) created during oil and gas exploration and very extensive in Alberta, Canada. Loss of microtopographic variation leads to slow tree recovery on seismic lines in treed peatlands. To facilitate tree establishment, mounding is being applied to restore the microtopography on seismic lines. We compared bryophyte communities on unmounded and mounded seismic lines (18 years after seismic line creation, 3 years after mounding), and in adjacent treed fens as the reference, to evaluate how recovery varies with microtopographic position (i.e., top of hummock, slope or side of the hummock, and level ground). We found that recovery on the unmounded seismic lines was underway 18 years after seismic lines were created, while mounding redisturbed the sites and thus set back recovery, at least in the short term (3 years post‐mounding). In unmounded seismic lines, Sphagnum cover was similar to that of reference treatments at all microtopographic positions. In contrast, mounded seismic lines had lower Sphagnum cover and higher cover of true mosses (sans feather mosses) than reference and unmounded treatments. Overall, these results show mounding sets back the recovery of bryophyte communities—the very organisms responsible for the microtopographic variation we are trying to establish. Further monitoring is needed to understand how this changes over time. [ABSTRACT FROM AUTHOR]

Site moisture is an important component of the forest landscape for maintaining biodiversity, including forest‐floor bryophytes. However, little is known about its role in shaping understorey responses to harvesting.We investigated the influence of site wetness, determined using a remotely sensed, topographic depth‐to‐water (DTW) index, on responses of bryophyte cover, richness, diversity and composition to variable retention harvesting (comparing: 2% [clear‐cut], 20% and 50% dispersed green tree retention and uncut controls [100% retention]) in three boreal forest cover‐types (broadleaf, mixed and conifer forests) in western Canada. The DTW index provides an approximation of DTW at or below the soil surface and was derived from wet‐areas mapping based on discrete Airborne Laser Scanning data acquired over an experimentally harvested landscape located in north‐western Alberta, Canada.The effectiveness of leaving retention (vs. clear‐cutting) for conserving bryophyte communities depended on site wetness, as indicated by DTW, with the specifics varying among forest types. In broadleaf forests, bryophyte cover and richness were generally low and not much affected by harvesting but drier sites had higher richness and a few more unique species. In mixed and conifer forests, leaving retention (vs. clear‐cutting) on wetter (vs. drier) sites was more effective for conserving bryophyte cover, wetter sites had higher total species richness and more species were exclusive to wetter sites.Synthesis and applications. Site wetness, as indicated using the remotely sensed topographic site wetness index "depth‐to‐water" mediates bryophyte responses to variable‐retention harvests. Specifically, our results suggested that in conifer and mixed forests it would be more beneficial to target wetter sites for retention patches or dispersed retention whereas in broadleaf forests there might be a slight advantage to targeting drier sites. Our study demonstrates that this tool could be used to inform management decisions around leaving dispersed or patch retention. Site wetness, as indicated using the remotely sensed topographic site wetness index "depth‐to‐water" mediates bryophyte responses to variable‐retention harvests. Specifically, our results suggested that in conifer and mixed forests it would be more beneficial to target wetter sites for retention patches or dispersed retention whereas in broadleaf forests there might be a slight advantage to targeting drier sites. Our study demonstrates that this tool could be used to inform management decisions around leaving dispersed or patch retention. [ABSTRACT FROM AUTHOR]