95 results on '"Subcritical Hopf bifurcation"'
Search Results
2. The impact of harvesting on the evolutionary dynamics of prey species in a prey-predator systems.
- Author
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Bandyopadhyay, Richik and Chattopadhyay, Joydev
- Abstract
Matsuda and Abrams (Theor Popul Biol 45(1):76–91, 1994) initiated the exploration of self-extinction in species through evolution, focusing on the advantageous position of mutants near the extinction boundary in a prey-predator system with evolving foraging traits. Previous models lacked theoretical investigation into the long-term effects of harvesting. In our model, we introduce constant-effort prey and predator harvesting, along with individual logistic growth of predators. The model reveals two distinct evolutionary outcomes: (i) Evolutionary suicide, marked by a saddle-node bifurcation, where prey extinction results from the invasion of a lower forager mutant; and (ii) Evolutionary reversal, characterized by a subcritical Hopf bifurcation, leading to cyclic prey evolution. Employing an innovative approach based on Gröbner basis computation, we identify various bifurcation manifolds, including fold, transcritical, cusp, Hopf, and Bogdanov-Takens bifurcations. These contrasting scenarios emerge from variations in harvesting parameters while keeping other factors constant, rendering the model an intriguing subject of study. [ABSTRACT FROM AUTHOR]
- Published
- 2024
- Full Text
- View/download PDF
3. From deterministic to stochastic: limits of extracting bifurcation diagrams for noisy bistable oscillators with the control-based continuation method
- Author
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Sykora, Henrik T. and Beregi, Sandor
- Published
- 2024
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4. Empirical Modeling of Subcritical Hopf Bifurcation of Thermoacoustic Stirling Engine.
- Author
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Lai, Chuan-Heng and Hsu, Shu-Han
- Subjects
STIRLING engines ,HOPF bifurcations ,TRANSIENTS (Dynamics) ,REGENERATORS ,THERMOACOUSTIC heat engines ,OSCILLATIONS - Abstract
This study models the subcritical Hopf bifurcation in thermoacoustic Stirling engines using the Stuart–Landau model, highlighting the role of nonlinear dynamics. By inducing self-sustained oscillations and measuring pressure fluctuations across different temperature gradients imposed on the regenerator, we reveal the engine's transition to a nonlinear domain, characterized by heightened oscillation amplitudes and unique periodic patterns. Interpreted Landau constants and growth rates illuminate the stabilizing effects of nonlinear dynamics, demonstrating the Stuart–Landau model's applicability in thermoacoustic engine analysis. Our research confirms that this empirically refined model reliably describes oscillation amplitudes and transient phenomena, contributing valuable perspectives for advancing thermoacoustic engine design and operational understanding. [ABSTRACT FROM AUTHOR]
- Published
- 2024
- Full Text
- View/download PDF
5. Empirical Modeling of Subcritical Hopf Bifurcation of Thermoacoustic Stirling Engine
- Author
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Chuan-Heng Lai and Shu-Han Hsu
- Subjects
thermoacoustic Stirling engine ,Stuart–Landau model ,subcritical Hopf Bifurcation ,Motor vehicles. Aeronautics. Astronautics ,TL1-4050 - Abstract
This study models the subcritical Hopf bifurcation in thermoacoustic Stirling engines using the Stuart–Landau model, highlighting the role of nonlinear dynamics. By inducing self-sustained oscillations and measuring pressure fluctuations across different temperature gradients imposed on the regenerator, we reveal the engine’s transition to a nonlinear domain, characterized by heightened oscillation amplitudes and unique periodic patterns. Interpreted Landau constants and growth rates illuminate the stabilizing effects of nonlinear dynamics, demonstrating the Stuart–Landau model’s applicability in thermoacoustic engine analysis. Our research confirms that this empirically refined model reliably describes oscillation amplitudes and transient phenomena, contributing valuable perspectives for advancing thermoacoustic engine design and operational understanding.
- Published
- 2024
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6. HOPF BIFURCATIONS IN DYNAMICAL SYSTEMS VIA ALGEBRAIC TOPOLOGICAL METHOD.
- Author
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Jawarneh, Ibrahim and Altawallbeh, Zuhier
- Subjects
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HOPF bifurcations , *DYNAMICAL systems , *DIFFERENTIAL equations , *LIMIT cycles - Abstract
A nonlinear phenomenon in nature is often modeled by a system of differential equations with parameters. The bifurcation occurs when a parameter varies in such systems, causing a qualitative change in its solution. In this paper, we study one of the most exciting bifurcations, which is Hopf bifurcation. We use tools from algebraic topology to analyze and reveal supercritical and subcritical Hopf bifurcations. [ABSTRACT FROM AUTHOR]
- Published
- 2023
- Full Text
- View/download PDF
7. Inhibitory autapse with time delay induces mixed-mode oscillations related to unstable dynamical behaviors near subcritical Hopf bifurcation
- Author
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Li Li and Zhiguo Zhao
- Subjects
inhibitory autapse ,time delay ,mixed-mode oscillations ,subcritical hopf bifurcation ,neural firing ,Mathematics ,QA1-939 ,Applied mathematics. Quantitative methods ,T57-57.97 - Abstract
Mixed-mode oscillations (MMOs) consisting of spikes alternating with a series of subthreshold oscillations have been observed in various neurons related to some physiological functions. In the present paper, inhibitory-autapse-induced MMOs are simulated by using the Hodgkin-Huxley neuron model, and the underlying dynamical mechanism is identified to be related to dynamics of unstable behaviors near subcritical Hopf bifurcation. For the monostable spiking, a delayed inhibitory current pulse activated by a spike can suppress the phase trajectory corresponding to depolarization phase of the next spike to the unstable focus nearby or the neighborhood outside of unstable limit cycle, respectively. Then the trajectory rotates multiple cycles away and converges to the stable limit cycle, resulting in an evolution process of membrane potential from small-amplitude subthreshold oscillations to a large-amplitude spike, i.e., MMOs. For the spiking coexisting with the resting state, inhibitory autapse induces MMOs and resting state from the spiking. The difference in the MMOs from those induced by the excitatory autapse is identified. The result presents the underlying nonlinear mechanisms of inhibitory autapse to suppress the neuronal firing and reveals the potential role to control the neuronal firing patterns near subcritical Hopf bifurcation.
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- 2022
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8. Effect of background noise characteristics on early warning indicators of thermoacoustic instability.
- Author
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Vishnoi, Neha and Kabiraj, Lipika
- Subjects
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HOPF bifurcations , *GAS turbines , *TIME series analysis , *COMBUSTION chambers , *NOISE , *WHITE noise - Abstract
In this work, we investigate the effects of background noise characteristics — specifically noise color (or correlation time) and intensity — arising from velocity-coupled and additive noise sources, on the early warning indicators (EWIs) of thermoacoustic instability. We employ a nonlinear reduced-order combustion dynamics model for our investigation. In the absence of noise, the nonlinear model undergoes a subcritical Hopf bifurcation, and our focus lies within the linearly stable region of the system (subthreshold region). The studied class of EWIs encompasses those derived from time series (variance), spectral analysis (coherence factor), Hurst exponent, and nonlinear methods (permutation entropy). We find that when the background noise is purely multiplicative, the trends in EWIs are primarily influenced by noise characteristics rather than the control parameter. Further, the EWIs cannot be estimated at low noise levels for large correlation times. In case of purely additive noise driven system, the coherence factor and variance are reliable EWIs across all range of investigated noise characteristics. The Hurst exponent can serve as effective EWI when the system features large noise correlation times, while permutation entropy is effective only when the system features small noise correlation time, i.e., where white noise assumption is acceptable. When the background noise includes contributions from both multiplicative and additive sources, coherence factor and variance emerges as the most reliable EWIs. These results provide insights for selecting appropriate EWIs to be employed in practical systems, considering potential variations in noise characteristics with simultaneous changes in combustor operating conditions. Novelty and significance This study conclusively shows that background noise (multiplicative and additive) characteristics – noise correlation time (color) and intensity – can significantly change the trends in early warning indicators (EWIs) for predicting the impending thermoacoustic oscillations. In gas turbine combustors, where thermoacoustic instability poses a critical challenge, inherent noise dynamics undergo variations with changing operating conditions and combustor designs. The development of effective EWIs to foresee the onset of thermoacoustic instability is crucial for preventing potential damage and ensuring the reliable operation of combustion systems. Previous works on stochastic dynamics of combustors simplify noise with the additive white noise assumption. Thus, our results demonstrated on a nonlinear combustion dynamics model advance the state-of-the-art with respect to the early prediction and control of thermoacoustic instability. The results provide valuable insights for selecting appropriate EWIs for engine monitoring in the absence of information on noise properties and their variation with operating conditions. This practical information is of direct relevance and interest to any gas turbine manufacturer/user. [ABSTRACT FROM AUTHOR]
- Published
- 2024
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9. Bistability Resulting from Rebound Firing
- Author
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Börgers, Christoph, Antman, S.S., Editor-in-chief, Bell, J., Series editor, Keller, J., Series editor, Greengard, L., Editor-in-chief, Holmes, P.J., Editor-in-chief, Kohn, R., Series editor, Newton, P., Series editor, Peskin, C., Series editor, Pego, R., Series editor, Ryzhik, L., Series editor, Singer, A., Series editor, Stevens, A., Series editor, Stuart, A., Section editor, Witelski, T., Series editor, Wright, S., Series editor, and Börgers, Christoph
- Published
- 2017
- Full Text
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10. Bursting
- Author
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Börgers, Christoph, Antman, S.S., Editor-in-chief, Bell, J., Series editor, Keller, J., Series editor, Greengard, L., Editor-in-chief, Holmes, P.J., Editor-in-chief, Kohn, R., Series editor, Newton, P., Series editor, Peskin, C., Series editor, Pego, R., Series editor, Ryzhik, L., Series editor, Singer, A., Series editor, Stevens, A., Series editor, Stuart, A., Section editor, Witelski, T., Series editor, Wright, S., Series editor, and Börgers, Christoph
- Published
- 2017
- Full Text
- View/download PDF
11. Different dynamics of repetitive neural spiking induced by inhibitory and excitatory autapses near subcritical Hopf bifurcation.
- Author
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Zhao, Zhiguo, Li, Li, Gu, Huaguang, and Gao, Yu
- Abstract
Based on the post-inhibitory rebound (PIR) spike induced by inhibitory current pulse, in the present paper, a novel counterintuitive phenomenon that the inhibitory autapse with time delay can induce the resting state changed to stable spiking pattern is identified near subcritical Hopf bifurcation of Hodgkin–Huxley model. The delayed inhibitory autaptic current pulse induced by the preceded action potential can induce the preceding PIR spike via the hyperpolarization, rebound, and depolarization processes, which is compared with spiking induced by excitatory autapse via only a depolarization process. The threshold of inhibitory or excitatory autaptic conductance to induce spiking with increasing time delay, and the threshold curve of inhibitory or excitatory pulse current to evoke a spike exhibit damping oscillations can be well interpreted with the damping dynamics of focus near subcritical Hopf bifurcation. However, due to PIR mechanism, the threshold conductance of inhibitory autapse is stronger than that of excitatory autapse, and the spiking period for inhibitory autapse, which is composed of time delay and durations of the other three processes, is longer than the one for excitatory autapse, which is composed of time delay and duration of only a depolarization process. Therefore, a linear correlation between spiking period and time delay is identified, which shows that autapse can modulate the spike timing related to temporal coding. The results present a novel viewpoint and a potential function that inhibitory autapse can facilitate spiking like the excitatory autapse and provide effective measures to modulate neuronal spiking pattern, which is related to subcritical Hopf bifurcation. [ABSTRACT FROM AUTHOR]
- Published
- 2020
- Full Text
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12. Oscillations
- Author
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Milton, John, Ohira, Toru, Milton, John, and Ohira, Toru
- Published
- 2014
- Full Text
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13. Evolutionary Suicide of Prey: Matsuda and Abrams' Model Revisited.
- Author
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Vitale, Caterina and Kisdi, Eva
- Subjects
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SUICIDE , *PREDATION , *PREY availability , *LIMIT cycles , *POPULATION dynamics , *HOPF bifurcations , *BIOLOGICAL extinction - Abstract
Under the threat of predation, a species of prey can evolve to its own extinction. Matsuda and Abrams (Theor Popul Biol 45:76–91, 1994a) found the earliest example of evolutionary suicide by demonstrating that the foraging effort of prey can evolve until its population dynamics cross a fold bifurcation, whereupon the prey crashes to extinction. We extend this model in three directions. First, we use critical function analysis to show that extinction cannot happen via increasing foraging effort. Second, we extend the model to non-equilibrium systems and demonstrate evolutionary suicide at a fold bifurcation of limit cycles. Third, we relax a crucial assumption of the original model. To find evolutionary suicide, Matsuda and Abrams assumed a generalist predator, whose population size is fixed independently of the focal prey. We embed the original model into a three-species community of the focal prey, the predator and an alternative prey that can support the predator also alone, and investigate the effect of increasingly strong coupling between the focal prey and the predator's population dynamics. Our three-species model exhibits (1) evolutionary suicide via a subcritical Hopf bifurcation and (2) indirect evolutionary suicide, where the evolution of the focal prey first makes the community open to the invasion of the alternative prey, which in turn makes evolutionary suicide of the focal prey possible. These new phenomena highlight the importance of studying evolution in a broader community context. [ABSTRACT FROM AUTHOR]
- Published
- 2019
- Full Text
- View/download PDF
14. Cyclic prey evolution with cannibalistic predators.
- Author
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Lehtinen, Sami O. and Geritz, Stefan A.H.
- Subjects
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PREY availability , *PREDATION , *BIOLOGICAL evolution , *REMOTELY piloted vehicles , *PREDATORY animals - Abstract
• We derive an ecological predator-prey model from individual-level processes with cannibalistic predators and timid prey. • Bifurcation analysis reveals ecological bistability between equilibrium and periodic population attractors. • We investigate the evolution of timidity of the prey along each ecological attractor, obtaining ten qualitatively different evolutionary scenarios. • Ecological bistability plays a central role in the evolutionary scenarios, which involve abrupt switching between alternative ecological attractors through catastrophic bifurcations. • We find that the end result can be an evolutionary cycle of the level of timidity, driven by ecological attractor switching. We investigate the evolution of timidity in a prey species whose predator has cannibalistic tendencies. The ecological model is derived from individual-level processes, in which the prey seeks refuge after detecting a predator, and the predator cannibalises on the conspecific juveniles. Bifurcation analysis of the model reveals ecological bistability between equilibrium and periodic attractors. Using the framework of adaptive dynamics, we classify ten qualitatively different evolutionary scenarios induced by the ecological bistability. These scenarios include ecological attractor switching through catastrophic bifurcations, which can reverse the direction of evolution. We show that such reversals often result in evolutionary cycling of the level of timidity. In the absence of cannibalism, the model never exhibits ecological bistability nor evolutionary cycling. We conclude that cannibalistic predator behaviour can completely change both the ecological dynamics and the evolution of prey. [ABSTRACT FROM AUTHOR]
- Published
- 2019
- Full Text
- View/download PDF
15. Subcritical Hopf and saddle-node bifurcations in hunting motion caused by cubic and quintic nonlinearities: experimental identification of nonlinearities in a roller rig.
- Author
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Wei, Weiyan and Yabuno, H.
- Abstract
Railway vehicles suffer from hunting motion, even when traveling below the critical speed obtained by linear analysis, due to the nonlinear characteristics of the wheel system. Nonlinear characteristics in Hopf bifurcations can be characterized as subcritical or supercritical, depending on whether the cubic nonlinearity is softening or hardening, respectively. In a system with softening cubic nonlinearity, third-order nonlinear analysis cannot detect nontrivial stable steady-state oscillations because they are affected by quintic nonlinearity. Therefore, in such a system, it is necessary to apply fifth-order nonlinear analysis to a system model in which quintic nonlinearity is taken into account. In this study, we investigated the cubic and quintic nonlinear phenomena in hunting motion with a roller rig that is widely used for hunting motion research. Previous experimental studies using a roller rig were restricted to the linear stability and the cubic nonlinear stability. We clarified that roller rig experiments can observe the hysteresis phenomenon and the existence of subcritical Hopf and saddle-node bifurcations, indicating that not only the cubic but also the quintic nonlinearity of the wheel system plays an important role. In addition, we obtained the normal form governing the nonlinear dynamics. We developed an experimental identification method to obtain the coefficients of the normal form. The validity of our method was confirmed by comparing the bifurcation diagrams obtained from the experimental time history and the normal form whose coefficients were experimentally identified using the proposed method. [ABSTRACT FROM AUTHOR]
- Published
- 2019
- Full Text
- View/download PDF
16. Dynamical principles underlying song degradation in birdsong neural circuit
- Author
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Zang, Jie and Liu, Shenquan
- Published
- 2021
- Full Text
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17. Subcritical Hopf Bifurcation and Stochastic Resonance of Electrical Activities in Neuron under Electromagnetic Induction
- Author
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Yu-Xuan Fu, Yan-Mei Kang, and Yong Xie
- Subjects
electromagnetic induction ,subcritical Hopf bifurcation ,stochastic resonance ,weak signal detection ,improved FitzHugh-Nagumo model ,Neurosciences. Biological psychiatry. Neuropsychiatry ,RC321-571 - Abstract
The FitzHugh–Nagumo model is improved to consider the effect of the electromagnetic induction on single neuron. On the basis of investigating the Hopf bifurcation behavior of the improved model, stochastic resonance in the stochastic version is captured near the bifurcation point. It is revealed that a weak harmonic oscillation in the electromagnetic disturbance can be amplified through stochastic resonance, and it is the cooperative effect of random transition between the resting state and the large amplitude oscillating state that results in the resonant phenomenon. Using the noise dependence of the mean of interburst intervals, we essentially suggest a biologically feasible clue for detecting weak signal by means of neuron model with subcritical Hopf bifurcation. These observations should be helpful in understanding the influence of the magnetic field to neural electrical activity.
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- 2018
- Full Text
- View/download PDF
18. Counterexample
- Author
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Flunkert, Valentin and Flunkert, Valentin
- Published
- 2011
- Full Text
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19. Introduction
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Hövel, Philipp and Hövel, Philipp
- Published
- 2010
- Full Text
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20. Bistable emergence of oscillations in growing Bacillus subtilis biofilms.
- Author
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Martinez-Corral, Rosa, Jintao Liu, Süel, Gürol M., and Garcia-Ojalvo, Jordi
- Subjects
- *
BACILLUS subtilis , *BIOFILMS , *BIOLOGICAL systems , *CELLS , *PLANT nutrients - Abstract
Biofilm communities of Bacillus subtilis bacteria have recently been shown to exhibit collective growth-rate oscillations mediated by electrochemical signaling to cope with nutrient starvation. These oscillations emerge once the colony reaches a large enough number of cells. However, it remains unclear whether the amplitude of the oscillations, and thus their effectiveness, builds up over time gradually or if they can emerge instantly with a nonzero amplitude. Here we address this question by combining microfluidics-based time-lapse microscopy experiments with a minimal theoretical description of the system in the form of a delay-differential equation model. Analytical and numerical methods reveal that oscillations arise through a subcritical Hopf bifurcation, which enables instant high-amplitude oscillations. Consequently, the model predicts a bistable regime where an oscillating and a nonoscillating attractor coexist in phase space. We experimentally validate this prediction by showing that oscillations can be triggered by perturbing the media conditions, provided the biofilm size lies within an appropriate range. The model also predicts that the minimum size at which oscillations start decreases with stress, a fact that we also verify experimentally. Taken together, our results show that collective oscillations in cell populations can emerge suddenly with nonzero amplitude via a discontinuous transition. [ABSTRACT FROM AUTHOR]
- Published
- 2018
- Full Text
- View/download PDF
21. Dynamic Analysis of a Bistable Bi-Local Active Memristor and Its Associated Oscillator System.
- Author
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Chang, Hui, Wang, Zhen, Li, Yuxia, and Chen, Guanrong
- Subjects
- *
MEMRISTORS , *MATHEMATICAL symmetry , *HOPF bifurcations , *NONLINEAR theories , *CHAOS theory - Abstract
This paper proposes a new type of memristor with two distinct stable pinched hysteresis loops and twin symmetrical local activity domains, named as a bistable bi-local active memristor. A detailed and comprehensive analysis of the memristor and its associated oscillator system is carried out to verify its dynamic behaviors based on nonlinear circuit theory and Hopf bifurcation theory. The local-activity domains and the edge-of-chaos domains of the memristor, which are both symmetric with respect to the origin, are confirmed by utilizing the mathematical cogent theory. Finally, the subcritical Hopf bifurcation phenomenon is identified in the subcritical Hopf bifurcation region of the memristor. [ABSTRACT FROM AUTHOR]
- Published
- 2018
- Full Text
- View/download PDF
22. Nonlinear control of stick-slip oscillations by normal force modulation.
- Author
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Nath, Jyayasi and Chatterjee, S.
- Subjects
- *
STICK-slip response , *VIBRATION (Mechanics) , *NONLINEAR control theory , *HOPF bifurcations , *POLYNOMIALS - Abstract
The present paper investigates the efficacy of controlling friction induced vibration by normal load modulation. Friction-induced self-excited vibration, attributed to the low-velocity drooping characteristics of friction (Stribeck effect), is modelled by a mass-on-belt model where the normal force of the mass is being modulated based on the acceleration feedback followed by a second order filtering. Polynomial model is employed to study the friction phenomenon between the mass and the belt. The pole crossover design (to ensure faster transient and greater relative stability) is implemented to optimize the filter parameters with an independent choice of the belt velocity and control gain. These sets of optimized parameter values are then used to construct local stability boundaries in the plane of control parameters. Numerical simulations in a MATLAB SIMULINK model and bifurcation diagrams obtained in AUTO (while using belt velocity as the bifurcation parameter) indicate that a significantly small-amplitude limit cycle resulting from a supercritical Hopf bifurcation stabilizes the extreme low velocity region at higher values of the control gain. With the increase of the control gain the subcritical nature of Hopf bifurcation changes to a supercritical one. The efficacy of this optimization (based on numerical results) in the delicate low velocity region is also enclosed. [ABSTRACT FROM AUTHOR]
- Published
- 2018
- Full Text
- View/download PDF
23. Subcritical Hopf Bifurcation and Stochastic Resonance of Electrical Activities in Neuron under Electromagnetic Induction.
- Author
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Fu, Yu-Xuan, Kang, Yan-Mei, and Xie, Yong
- Subjects
NEURAL stimulation ,ELECTROMAGNETIC induction ,STOCHASTIC resonance ,HOPF bifurcations ,HARMONIC oscillators - Abstract
The FitzHugh–Nagumo model is improved to consider the effect of the electromagnetic induction on single neuron. On the basis of investigating the Hopf bifurcation behavior of the improved model, stochastic resonance in the stochastic version is captured near the bifurcation point. It is revealed that a weak harmonic oscillation in the electromagnetic disturbance can be amplified through stochastic resonance, and it is the cooperative effect of random transition between the resting state and the large amplitude oscillating state that results in the resonant phenomenon. Using the noise dependence of the mean of interburst intervals, we essentially suggest a biologically feasible clue for detecting weak signal by means of neuron model with subcritical Hopf bifurcation. These observations should be helpful in understanding the influence of the magnetic field to neural electrical activity. [ABSTRACT FROM AUTHOR]
- Published
- 2018
- Full Text
- View/download PDF
24. Shear instabilities in Taylor-Couette flow
- Author
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Meseguer, A., Mellibovsky, F., Marques, F., Avila, M., and Eckhardt, Bruno, editor
- Published
- 2009
- Full Text
- View/download PDF
25. Complex Features in Lotka-Volterra Systems with Behavioral Adaptation
- Author
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Tebaldi, Claudio, Lacitignola, Deborah, Minai, Ali, editor, Braha, Dan, editor, and Bar-Yam, Yaneer, editor
- Published
- 2008
- Full Text
- View/download PDF
26. Monitoring and Control of Bifurcations Using Probe Signals
- Author
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Hassouneh, Munther A., Yaghoobi, Hassan, Abed, Eyad H., Thoma, M., editor, Morari, M., editor, Colonius, Fritz, editor, and Grüne, Lars, editor
- Published
- 2002
- Full Text
- View/download PDF
27. Summary and Outlook
- Author
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Hövel, Philipp and Hövel, Philipp
- Published
- 2010
- Full Text
- View/download PDF
28. Scenario of the Birth of Hidden Attractors in the Chua Circuit.
- Author
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Stankevich, Nataliya V., Kuznetsov, Nikolay V., Leonov, Gennady A., and Chua, Leon O.
- Subjects
- *
CHAOS theory , *DYNAMICAL systems , *MATHEMATICAL symmetry , *EQUILIBRIUM , *HOPF bifurcations , *PARAMETER estimation - Abstract
Recently it was shown that in the dynamical model of Chua circuit both the classical self-excited and hidden chaotic attractors can be found. In this paper the dynamics of the Chua circuit is revisited. The scenario of the chaotic dynamics development and the birth of self-excited and hidden attractors is studied. A pitchfork bifurcation is shown in which a pair of symmetric attractors coexist and merge into one symmetric attractor through an attractor-merging bifurcation and a splitting of a single attractor into two attractors. The scenario relating the subcritical Hopf bifurcation near equilibrium points and the birth of hidden attractors is discussed. [ABSTRACT FROM AUTHOR]
- Published
- 2017
- Full Text
- View/download PDF
29. Chaotic Oscillation via Edge of Chaos Criteria.
- Author
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Itoh, Makoto and Chua, Leon
- Subjects
- *
CHAOS theory , *PHASE transitions , *POINCARE series , *COORDINATE transformations , *BIFURCATION theory , *NONLINEAR dynamical systems - Abstract
In this paper, we show that nonlinear dynamical systems which satisfy the edge of chaos criteria can bifurcate from a stable equilibrium point regime to a chaotic regime by periodic forcing. That is, the edge of chaos criteria can be exploited to engineer a phase transition from ordered to chaotic behavior. The frequency of the periodic forcing can be derived from this criteria. In order to generate a periodic or a chaotic oscillation, we have to tune the amplitude of the periodic forcing. For example, we engineer chaotic oscillations in the generalized Duffing oscillator, the FitzHugh-Nagumo model, the Hodgkin-Huxley model, and the Morris-Lecar model. Although forced oscillators can exhibit chaotic oscillations even if the edge of chaos criteria is not satisfied, our computer simulations show that forced oscillators satisfying the edge of chaos criteria can exhibit highly complex chaotic behaviors, such as folding loci, strong spiral dynamics, or tight compressing dynamics. In order to view these behaviors, we used high-dimensional Poincaré maps and coordinate transformations. We also show that interesting nonlinear dynamical systems can be synthesized by applying the edge of chaos criteria. They are globally stable without forcing, that is, all trajectories converge to an asymptotically-stable equilibrium point. However, if we apply a forcing signal, then the dynamical systems can oscillate chaotically. Furthermore, the average power delivered from the forced signal is not dissipated by chaotic oscillations, but on the contrary, energy can be generated via chaotic oscillations, powered by locally-active circuit elements inside the one-port circuit connected across a current source. [ABSTRACT FROM AUTHOR]
- Published
- 2017
- Full Text
- View/download PDF
30. Bifurcation analysis of the simplified models of boiling water reactor and identification of global stability boundary.
- Author
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Pandey, Vikas and Singh, Suneet
- Subjects
- *
BIFURCATION theory , *BOILING water reactors , *NUCLEAR models , *STABILITY theory , *BOUNDARY value problems , *HOPF bifurcations - Abstract
Nonlinear stability study of the neutron coupled thermal hydraulics instability has been carried out by several researchers for boiling water reactors (BWRs). The focus of these studies has been to identify subcritical and supercritical Hopf bifurcations. Supercritical Hopf bifurcation are soft or safe due to the fact that stable limit cycles arise in linearly unstable region; linear and global stability boundaries are same for this bifurcation. It is well known that the subcritical bifurcations can be considered as hard or dangerous due to the fact that unstable limit cycles (nonlinear phenomena) exist in the (linearly) stable region. The linear stability leads to a stable equilibrium in such regions, only for infinitesimally small perturbations. However, finite perturbations lead to instability due to the presence of unstable limit cycles. Therefore, it is evident that the linear stability analysis is not sufficient to understand the exact stability characteristics of BWRs. However, the effect of these bifurcations on the stability boundaries has been rarely discussed. In the present work, the identification of global stability boundary is demonstrated using simplified models. Here, five different models with different thermal hydraulics feedback have been investigated. In comparison to the earlier works, current models also include the impact of adding the rate of change in temperature on void reactivity as well as effect of void reactivity on rate of change of temperature. Using the bifurcation analysis of these models the globally stable region in the parameter space has been identified. The globally stable region has only stable solutions and does not have even unstable limit cycles. Hence, the system is stable irrespective of the size of the perturbation in these regions. [ABSTRACT FROM AUTHOR]
- Published
- 2017
- Full Text
- View/download PDF
31. Third-Order Memristive Morris-Lecar Model of Barnacle Muscle Fiber.
- Author
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Rajamani, Vetriveeran, Sah, Maheshwar PD., Mannan, Zubaer Ibna, Kim, Hyongsuk, and Chua, Leon
- Subjects
- *
POTASSIUM ions , *CALCIUM ions , *BARNACLES , *LIMIT cycles , *STABLE equilibrium (Physics) , *HOPF bifurcations - Abstract
This paper presents a detailed analysis of various oscillatory behaviors observed in relation to the calcium and potassium ions in the third-order Morris-Lecar model of giant barnacle muscle fiber. Since, both the calcium and potassium ions exhibit all of the characteristics of memristor fingerprints, we claim that the time-varying calcium and potassium ions in the third-order Morris-Lecar model are actually time-invariant calcium and potassium memristors in the third-order memristive Morris-Lecar model. We confirmed the existence of a small unstable limit cycle oscillation in both the second-order and the third-order Morris-Lecar model by numerically calculating the basin of attraction of the asymptotically stable equilibrium point associated with two subcritical Hopf bifurcation points. We also describe a comprehensive analysis of the generation of oscillations in third-order memristive Morris-Lecar model via small-signal circuit analysis and a subcritical Hopf bifurcation phenomenon. [ABSTRACT FROM AUTHOR]
- Published
- 2017
- Full Text
- View/download PDF
32. Population Dynamics of Tribolium
- Author
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Desharnais, Robert A., Usher, Michael B., editor, DeAngelis, D. L., editor, Manly, B. F. J., editor, Tuljapurkar, Shripad, editor, and Caswell, Hal, editor
- Published
- 1997
- Full Text
- View/download PDF
33. Tangential acceleration feedback control of friction induced vibration.
- Author
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Nath, Jyayasi and Chatterjee, S.
- Subjects
- *
TANGENTIAL acceleration (Physics) , *FEEDBACK control systems , *FRICTION , *VIBRATION (Mechanics) , *AIRDROP , *COMPUTER simulation - Abstract
Tangential control action is studied on a phenomenological mass-on-belt model exhibiting friction-induced self-excited vibration attributed to the low-velocity drooping characteristics of friction which is also known as Stribeck effect. The friction phenomenon is modelled by the exponential model. Linear stability analysis is carried out near the equilibrium point and local stability boundary is delineated in the plane of control parameters. The system is observed to undergo a Hopf bifurcation as the eigenvalues determined from the linear stability analysis are found to cross the imaginary axis transversally from RHS s-plane to LHS s-plane or vice-versa as one varies the control parameters, namely non-dimensional belt velocity and the control gain. A nonlinear stability analysis by the method of Averaging reveals the subcritical nature of the Hopf bifurcation. Thus, a global stability boundary is constructed so that any choice of control parameters from the globally stable region leads to a stable equilibrium. Numerical simulations in a MATLAB SIMULINK model and bifurcation diagrams obtained in AUTO validate these analytically obtained results. Pole crossover design is implemented to optimize the filter parameters with an independent choice of belt velocity and control gain. The efficacy of this optimization (based on numerical results) in the delicate low velocity region is also enclosed. [ABSTRACT FROM AUTHOR]
- Published
- 2016
- Full Text
- View/download PDF
34. Dynamics and control in a novel hyperchaotic system
- Author
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Matouk, A. E.
- Published
- 2019
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35. Multiple Bifurcation of Free-Convection Flow between Vertical Parallel Plates
- Author
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Kropp, M., Busse, F. H., Hoffmann, K.-H., editor, Mittelmann, H. D., editor, Todd, J., editor, Seydel, R., editor, Schneider, F. W., editor, Küpper, T., editor, and Troger, H., editor
- Published
- 1991
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36. Snaking bifurcations in a self-excited oscillator chain with cyclic symmetry.
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Papangelo, A., Grolet, A., Salles, L., Hoffmann, N., and Ciavarella, M.
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- *
BIFURCATION theory , *NONLINEAR oscillators , *MATHEMATICAL symmetry , *HOPF bifurcations , *LAW of large numbers - Abstract
Snaking bifurcations in a chain of mechanical oscillators are studied. The individual oscillators are weakly nonlinear and subject to self-excitation and subcritical Hopf-bifurcations with some parameter ranges yielding bistability. When the oscillators are coupled to their neighbours, snaking bifurcations result, corresponding to localised vibration states. The snaking patterns do seem to be more complex than in previously studied continuous systems, comprising a plethora of isolated branches and also a large number of similar but not identical states, originating from the weak coupling of the phases of the individual oscillators. [ABSTRACT FROM AUTHOR]
- Published
- 2017
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37. Cyclic prey evolution with cannibalistic predators
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Sami O. Lehtinen, Stefanus A.H. Geritz, and Department of Mathematics and Statistics
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DYNAMICS ,0301 basic medicine ,Statistics and Probability ,Food Chain ,Bistability ,Biology ,Models, Biological ,General Biochemistry, Genetics and Molecular Biology ,Predation ,03 medical and health sciences ,0302 clinical medicine ,BISTABILITY ,FOOD ,Attractor ,Animals ,Cannibalism ,Shyness ,PERCH PERCA-FLUVIATILIS ,CATASTROPHIC SHIFTS ,Predator ,Ecosystem ,Adaptive dynamics ,Extinction ,General Immunology and Microbiology ,Ecology ,Applied Mathematics ,Ecological bistability ,General Medicine ,Biological Evolution ,MODEL ,Supercritical Hopf bifurcation ,EXTINCTION ,SIZE ,030104 developmental biology ,Bifurcation analysis ,Fold bifurcation of periodic orbits ,Predatory Behavior ,Modeling and Simulation ,1181 Ecology, evolutionary biology ,Subcritical Hopf bifurcation ,GROWTH ,Social ecological model ,General Agricultural and Biological Sciences ,030217 neurology & neurosurgery - Abstract
We investigate the evolution of timidity in a prey species whose predator has cannibalistic tendencies. The ecological model is derived from individual-level processes, in which the prey seeks refuge after detecting a predator, and the predator cannibalises on the conspecific juveniles. Bifurcation analysis of the model reveals ecological bistability between equilibrium and periodic attractors. Using the framework of adaptive dynamics, we classify ten qualitatively different evolutionary scenarios induced by the ecological bistability. These scenarios include ecological attractor switching through catastrophic bifurcations, which can reverse the direction of evolution. We show that such reversals often result in evolutionary cycling of the level of timidity. In the absence of cannibalism, the model never exhibits ecological bistability nor evolutionary cycling. We conclude that cannibalistic predator behaviour can completely change both the ecological dynamics and the evolution of prey. (C) 2019 The Authors. Published by Elsevier Ltd.
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- 2019
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38. Mathematical modeling of the population dynamics of a distinct interactions type system with local dispersal.
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Aliyu, Murtala Bello and Mohd, Mohd Hafiz
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COEXISTENCE of species ,BIOTIC communities ,POPULATION dynamics ,ECOSYSTEMS ,ECOLOGICAL models ,PARTIAL differential equations - Abstract
• Symmetric and asymmetric dispersal affects the community structure of complex ecological system. • Moderate and intense predation strength provides buffer against extinction in a multiple interactions type model. • Short predator handling time support stable coexistence of species in this ecological system. • Long predator handling time disrupt the persistence of species in this multi-species system. • The complexity of distinct biotic interactions leads to an unstable limit cycle. Distinct biotic interactions in multi-species communities are a ubiquitous force in the natural ecosystem, and this force is an essential determinant of community stability and species coexistence outcomes. We conduct numerical simulations and bifurcation analysis of partial differential equations to gain better understanding and ecological insights into how predation (a), predator handling time (h), and local dispersal affect multi-species community dynamics. This system consists of resource-mutualist-exploiter-competitor interactions and local dispersal. From the inspection of our numerical simulations and co-dimension one bifurcation analysis findings, we discover several critical values that correspond to transcritical bifurcation, subcritical and supercritical Hopf bifurcations. This occurs as we vary the bifurcation parameters a and h in this complex ecological system under symmetric and asymmetric dispersal scenarios. Furthermore, the interplay between these local bifurcation points results in an exciting co-dimension two bifurcations, i.e., Bogdanov-Takens and cusp bifurcation points, respectively, which act as the synchronization points in this complex ecological system. From an ecological viewpoint, we find that (i) the effect of the no-dispersal scenario supports the maintenance of species biodiversity when the predation strength is moderate; (ii) symmetric dispersal induces both subcritical and supercritical Hopf bifurcation and support species diversity for moderate predation strength; and (iii) asymmetric dispersal promotes species diversity as it simplifies the bifurcation changes in dynamics by eliminating the subcritical bifurcations that trigger uncertainty, and this dispersal mechanism mediates species coexistence outcomes. Fundamentally, stable limit cycles have been reported as predator handling time varies in some ecological models; however, we observed in our bifurcation analysis the emergence of the unstable limit cycle as predator handling time changes. We discover that intense predator handling time destabilizes this complex ecological community. In general, our results demonstrate the influential roles of predation, predator handling time, and local dispersal in determining this system's coexistence dynamics. This knowledge provides a better understanding of species conservation and biological control management. [ABSTRACT FROM AUTHOR]
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- 2022
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39. Theoretical analysis of three-dimensional bifurcated flow inside a diagonally lid-driven cavity.
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Feldman, Yuri
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- *
NUMERICAL analysis , *FINITE element method , *REYNOLDS number , *BIFURCATION theory , *HELICITY of nuclear particles - Abstract
The instability mechanism of fully three-dimensional, highly separated, shear-driven confined flow inside a diagonally lid-driven cavity was investigated. The analysis was conducted on 100 and 200 stretched grids by a series of direct numerical simulations utilizing a standard second-order accuracy finite volume code, openFoam. The observed oscillatory instability was found to set in via a subcritical symmetry breaking Hopf bifurcation. Critical values of the Reynolds number Re = 2320 and the non-dimensional angular oscillating frequency $${\omega_{\rm cr}=0.249}$$ for the transition from steady to oscillatory flow were accurately determined. An oscillatory regime of the bifurcated flow was analyzed in depth, revealing and characterizing the spontaneous symmetry breaking mechanism. Characteristic spatial patterns of the base flow and the main flow harmonic were determined for the velocity, vorticity and helicity fields. Lagrangian particle tracers were utilized to visualize the mixing phenomenon of the flow from both sides of the diagonal symmetry plane. [ABSTRACT FROM AUTHOR]
- Published
- 2015
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40. EXISTENCE OF MULTIPLE LIMIT CYCLES IN A PREDATOR-PREY MODEL WITH arctan(ax) AS FUNCTIONAL RESPONSE.
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GUNOG SEO and WOLKOWICZ, GAIL S. K.
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LIMIT cycles , *MATHEMATICAL models , *SIMULATION methods & models , *MATHEMATICAL analysis , *MATHEMATICS - Abstract
We consider a Gause type predator-prey system with functional response given by ϑ(x) = arctan(ax), where a > 0, and give a counterexample to the criterion given in Attili and Mallak [Commun. Math. Anal. 1:33-40(2006)] for the nonexistence of limit cycles. When this criterion is satisfied, instead this system can have a locally asymptotically stable coexistence equilibrium surrounded by at least two limit cycles. [ABSTRACT FROM AUTHOR]
- Published
- 2015
41. Summary and Conclusions
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Flunkert, Valentin and Flunkert, Valentin
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- 2011
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42. Influence of system parameters on the hysteresis characteristics of a horizontal Rijke tube.
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Gopalakrishnan, E. A. and Sujith, R. I.
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- *
ELECTROMAGNETIC induction , *HYSTERESIS , *ELASTICITY , *RIJKE tube resonators , *HEATING - Abstract
The influence of system parameters such as heater power, heater location and mass flow rate on the hysteresis characteristics of a horizontal Rijke tube is presented in this paper. It is observed that a hysteresis zone is present for all the mass flow rates considered in the present study. A power law relation is established between the non-dimensional hysteresis width and the Strouhal number, defined as the ratio between convective time scale and acoustic time scale. The transition to instability in a horizontal Rijke tube is found to be subcritical in all the experiments performed in this study. When heater location is chosen as the control parameter, period-2 oscillations are found for specific values of mass flow rate and heater power. [ABSTRACT FROM AUTHOR]
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- 2014
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43. NEURONS ARE POISED NEAR THE EDGE OF CHAOS.
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CHUA, LEON, SBITNEV, VALERY, and KIM, HYONGSUK
- Subjects
- *
CHAOS theory , *NEURONS , *EIGENVALUES , *JACOBIAN matrices , *NONLINEAR differential equations , *ACTION potentials , *HOPF bifurcations - Abstract
This paper shows the action potential (spikes) generated from the Hodgkin-Huxley equations emerges near the edge of chaos consisting of a tiny subset of the locally active regime of the HH equations. The main result proves that the eigenvalues of the 4 × 4 Jacobian matrix associated with the mathematically intractable system of four nonlinear differential equations are identical to the zeros of a scalar complexity function from complexity theory. Moreover, we show the loci of a pair of complex-conjugate zeros migrate continuously as a function of an externally applied DC current excitation emulating the net synaptic excitation current input to the neuron. In particular, the pair of complex-conjugate zeros move from a subcritical Hopf bifurcation point at low excitation current to a super-critical Hopf bifurcation point at high excitation current. The spikes are generated as the excitation current approaches the vicinity of the edge of chaos, which leads to the onset of the subcritical Hopf bifurcation regime. It follows from this in-depth qualitative analysis that local activity is the origin of spikes. [ABSTRACT FROM AUTHOR]
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- 2012
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44. Time-delayed feedback control of unstable periodic orbits near a subcritical Hopf bifurcation
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Brown, G., Postlethwaite, C.M., and Silber, M.
- Subjects
- *
TIME delay systems , *FEEDBACK control systems , *PERIODIC functions , *HOPF algebras , *BIFURCATION theory , *NORMAL forms (Mathematics) , *MANIFOLDS (Mathematics) , *DELAY differential equations - Abstract
Abstract: We show that the Pyragas delayed feedback control can stabilize an unstable periodic orbit (UPO) that arises from a generic subcritical Hopf bifurcation of a stable equilibrium in an -dimensional dynamical system. This extends results of Fiedler et al. [B. Fiedler, V. Flunkert, M. Georgi, P. Hövel, E. Schöll, Refuting the odd-number limitation of time-delayed feedback control, Phys. Rev. Lett. 98(11) (2007) 114101], who demonstrated that such a feedback control can stabilize the UPO associated with a two-dimensional subcritical Hopf normal form. The Pyragas feedback requires an appropriate choice of a feedback gain matrix for stabilization, as well as knowledge of the period of the targeted UPO. We apply feedback in the directions tangent to the two-dimensional center manifold. We parameterize the feedback gain by a modulus and a phase angle, and give explicit formulae for choosing these two parameters given the period of the UPO in a neighborhood of the bifurcation point. We show, first heuristically, and then rigorously by a center manifold reduction for delay differential equations, that the stabilization mechanism involves a highly degenerate Hopf bifurcation problem that is induced by the time-delayed feedback. When the feedback gain modulus reaches a threshold for stabilization, both of the genericity assumptions associated with a two-dimensional Hopf bifurcation are violated: the eigenvalues of the linearized problem do not cross the imaginary axis as the bifurcation parameter is varied, and the real part of the cubic coefficient of the normal form vanishes. Our analysis reveals two qualitatively distinct cases when the degenerate bifurcation is unfolded in a two-parameter plane. In each case, the Pyragas-type feedback successfully stabilizes the branch of small-amplitude UPOs in a neighborhood of the original bifurcation point, provided that the feedback phase angle satisfies a certain restriction. [Copyright &y& Elsevier]
- Published
- 2011
- Full Text
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45. Analysis of periodic solutions in an eco-epidemiological model with saturation incidence and latency delay.
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Bhattacharyya, R. and Mukhopadhyay, B.
- Abstract
Abstract: In the present work, a mathematical model of predator–prey ecological interaction with infected prey is investigated. A saturation incidence function is used to model the behavioral change of the susceptible individuals when their number increases or due to the crowding effect of the infected individuals [V. Capasso, G. Serio, A generalization of the Kermack–McKendrick deterministic epidemic model, Math. Biosci. 42 (1978) 41–61]. Stability criteria for the infection-free and the endemic equilibria are deduced in terms of system parameters. The basic model is then modified to incorporate a time delay, describing a latency period. Stability and bifurcation analysis of the resulting delay differential equation model is carried out and ranges of the delay inducing stability and as well as instability for the system are found. Finally, a stability analysis of the bifurcating solutions is performed and the criteria for subcritical and supercritical Hopf bifurcation derived. The existence of a delay interval that preserves the stability of periodic orbits is demonstrated. The analysis emphasizes the importance of differential predation and a latency period in controlling disease dynamics. [Copyright &y& Elsevier]
- Published
- 2010
- Full Text
- View/download PDF
46. ON THE STUDY OF DELAY FEEDBACK CONTROL AND ADAPTIVE SYNCHRONIZATION NEAR SUB-CRITICAL HOPF BIFURCATION.
- Author
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Ghosh, Dibakar, Saha, Papri, and Chowdhury, A. Roy
- Subjects
- *
BIFURCATION theory , *SYNCHRONIZATION , *NONLINEAR systems , *COMBINATORIAL dynamics , *CHAOS theory - Abstract
The effect of delay feedback control and adaptive synchronization is studied near sub-critical Hopf bifurcation of a nonlinear dynamical system. Previously, these methods targeted the nonlinear systems near their chaotic regime but it is shown here that they are equally applicable near the branch of unstable solutions. The system is first analyzed from the view point of bifurcation, and the existence of Hopf bifurcation is established through normal form analysis. Hopf bifurcation can be either sub-critical or super-critical, and in the former case, unstable periodic orbits are formed. Our aim is to control them through a delay feedback approach so that the system stabilizes to its nearest stable periodic orbit. At the vicinity of the sub-critical Hopf point, adaptive synchronization is studied and the effect of the coupling parameter and the speed factor is analyzed in detail. Adaptive synchronization is also studied when the system is in the chaotic regime. [ABSTRACT FROM AUTHOR]
- Published
- 2008
- Full Text
- View/download PDF
47. Insight into the intrinsic time-varying aerodynamic properties of a truss girder undergoing a flutter with subcritical Hopf bifurcation.
- Author
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Wu, Bo, Shen, Huoming, Liao, Haili, Wang, Qi, Zhang, Yan, and Li, Zhiguo
- Subjects
- *
HOPF bifurcations , *PSYCHOLOGICAL feedback , *AERODYNAMIC load , *LIMIT cycles , *TRUSSES , *ATRIAL flutter , *BRIDGE design & construction , *AERODYNAMICS of buildings - Abstract
In this study, the investigation on the intrinsic time-varying nonlinear aerodynamic properties and actual energy feedback mechanism of limit cycle oscillation (LCO) and subcritical Hopf bifurcation, which is of great importance of the flutter design of bridges, but once rarely be conducted, was comprehensively carried out. The response characteristics of the nonlinear flutter were experimentally investigated. The dynamic mechanism of the subcritical Hopf bifurcation was firstly introduced in terms of the equivalent modal damping ratio as a function of amplitude. Further, a modified nonlinear self-excited force model was proposed to investigate the intrinsic time-varying characteristics of the aerodynamic properties and the real energy feedback mechanism of the subcritical Hopf bifurcation. Based on the model, the characteristics of nonlinear self-excited forces and nonlinear aerodynamic damping coefficients, as well as their contribution to the energy exchange behaviors, were studied. Subsequently, the energy feedback mechanisms of LCO and subcritical Hopf bifurcation were qualitatively discussed in detail considering the hysteresis loop of nonlinear self-excited forces (NSEF), which highlighted the important role of the linear self-excited-moment (including damping and stiffness) in the generation of a subcritical Hopf bifurcation, and the higher-order force components on the generation of stable LCO. Finally, the driving mechanisms of LCO and subcritical Hopf bifurcation were also quantitatively explained via an energy budget analysis, wherein the total work applied by structural and aerodynamic forces was presented as a function of amplitude and time. • A nonlinear self-excited force model is proposed for the calculation of subcritical Hopf bifurcation of a truss girder. • The contribution of force component to the LCO and subcritical Hopf bifurcation was discussed. • The dynamic mechanisms of LCO and subcritical Hopf bifurcation were qualitatively and quantitatively explained. [ABSTRACT FROM AUTHOR]
- Published
- 2022
- Full Text
- View/download PDF
48. Hopf bifurcation of an oscillator with quadratic and cubic nonlinearities and with delayed velocity feedback.
- Author
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Huailei, Wang, Zaihua, Wang, and Haiyan, Hu
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- 2004
- Full Text
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49. Evolutionary Suicide of Prey : Matsuda and Abrams' Model Revisited
- Author
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Caterina Vitale, Éva Kisdi, Department of Mathematics and Statistics, and University of Helsinki, Department of Mathematics and Statistics
- Subjects
0301 basic medicine ,Evolutionary suicide ,Foraging effort ,Population Dynamics ,Poison control ,Saddle-node bifurcation ,Predation ,COEXISTENCE ,0302 clinical medicine ,111 Mathematics ,Fold bifurcation of limit cycles ,General Environmental Science ,education.field_of_study ,PREDATION ,CONSTRUCTION ,Predator-prey model ,General Neuroscience ,Population size ,TRADE-OFF GEOMETRIES ,RESIDENT-INVADER DYNAMICS ,Biological Evolution ,DERIVATION ,Computational Theory and Mathematics ,030220 oncology & carcinogenesis ,Subcritical Hopf bifurcation ,General Agricultural and Biological Sciences ,ADAPTIVE DYNAMICS ,Food Chain ,General Mathematics ,education ,Immunology ,Population ,Foraging ,FUNCTIONAL-RESPONSE ,Biology ,Extinction, Biological ,Models, Biological ,General Biochemistry, Genetics and Molecular Biology ,03 medical and health sciences ,Animals ,Selection, Genetic ,Ecosystem ,Population Density ,Pharmacology ,Extinction ,Mathematical Concepts ,030104 developmental biology ,Evolutionary biology ,Predatory Behavior ,ATTRACTOR ,SELF-EXTINCTION - Abstract
Under the threat of predation, a species of prey can evolve to its own extinction. Matsuda and Abrams (Theor Popul Biol 45:76-91, 1994a) found the earliest example of evolutionary suicide by demonstrating that the foraging effort of prey can evolve until its population dynamics cross a fold bifurcation, whereupon the prey crashes to extinction. We extend this model in three directions. First, we use critical function analysis to show that extinction cannot happen via increasing foraging effort. Second, we extend the model to non-equilibrium systems and demonstrate evolutionary suicide at a fold bifurcation of limit cycles. Third, we relax a crucial assumption of the original model. To find evolutionary suicide, Matsuda and Abrams assumed a generalist predator, whose population size is fixed independently of the focal prey. We embed the original model into a three-species community of the focal prey, the predator and an alternative prey that can support the predator also alone, and investigate the effect of increasingly strong coupling between the focal prey and the predator's population dynamics. Our three-species model exhibits (1) evolutionary suicide via a subcritical Hopf bifurcation and (2) indirect evolutionary suicide, where the evolution of the focal prey first makes the community open to the invasion of the alternative prey, which in turn makes evolutionary suicide of the focal prey possible. These new phenomena highlight the importance of studying evolution in a broader community context.
- Published
- 2019
50. Bistable emergence of oscillations in growing Bacillus subtilis biofilms
- Author
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Rosa Martinez-Corral, Jordi Garcia-Ojalvo, Jintao Liu, and Gürol M. Süel
- Subjects
0301 basic medicine ,Biofilm growth ,Bistability ,Bacillus subtilis ,Biological oscillations ,01 natural sciences ,Quantitative Biology::Cell Behavior ,Delay-induced oscillations ,03 medical and health sciences ,symbols.namesake ,Delayed negative feedback ,0103 physical sciences ,Attractor ,010306 general physics ,Hopf bifurcation ,Physics ,Multidisciplinary ,biology ,Biofilm ,Mechanics ,biology.organism_classification ,Nutrient starvation ,030104 developmental biology ,Amplitude ,Phase space ,Subcritical Hopf bifurcation ,symbols - Abstract
Biofilm communities of Bacillus subtilis bacteria have recently been shown to exhibit collective growth-rate oscillations mediated by electrochemical signaling to cope with nutrient starvation. These oscillations emerge once the colony reaches a large enough number of cells. However, it remains unclear whether the amplitude of the oscillations, and thus their effectiveness, builds up over time gradually or if they can emerge instantly with a nonzero amplitude. Here we address this question by combining microfluidics-based time-lapse microscopy experiments with a minimal theoretical description of the system in the form of a delay-differential equation model. Analytical and numerical methods reveal that oscillations arise through a subcritical Hopf bifurcation, which enables instant high-amplitude oscillations. Consequently, the model predicts a bistable regime where an oscillating and a nonoscillating attractor coexist in phase space. We experimentally validate this prediction by showing that oscillations can be triggered by perturbing the media conditions, provided the biofilm size lies within an appropriate range. The model also predicts that the minimum size at which oscillations start decreases with stress, a fact that we also verify experimentally. Taken together, our results show that collective oscillations in cell populations can emerge suddenly with nonzero amplitude via a discontinuous transition. This work was supported by the Spanish Ministry of Economy and Competitiveness and Fondo Europeo de Desarrollo Regional (Project FIS2015-66503-C3-1-P) and by the Generalitat de Catalunya (Project 2017 SGR 1054). R.M.-C. acknowledges financial support from La Caixa Foundation. J.G.-O. acknowledges support from the Institució Catalana de Recerca i Estudis Avançats Academia programme and from the “María de Maeztu” Programme for Units of Excellence in Research and Development (Spanish Ministry of Economy and Competitiveness, MDM-2014-0370). J.L. acknowledges support from the J.L. acknowledges support from the Tsinghua–Peking Center for Life Sciences and the Thousand Talents Program of China. G.M.S. acknowledges support for this research from the San Diego Center for Systems Biology (NIH Grant P50 GM085764), the National Institute of General Medical Sciences (Grant R01 GM121888 to G.M.S.), the Defense Advanced Research Projects Agency (Grant HR0011-16-2-0035 to G.M.S.), and the Howard Hughes Medical Institute–Simons Foundation Faculty Scholars program (G.M.S.).
- Published
- 2018
- Full Text
- View/download PDF
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