Search

Teucrium montanum is a species widely distributed in Europe, occurring also in North Africa and Asia Minor. Because of its wide distribution and high morphological variability, many taxa related to T. montanum have been described. In this paper we discuss and lectotypify the following six names of taxa described or reported for the Balkan Peninsula: T. helianthemoides, T. montanum var. hirsutum, T. montanum var. modestum, T. montanum var. parnassicum, T. pannonicum, and T. skorpilii.

Teucrium montanum var. parnassicum Čelakovski (1883: 153) ��� Teucrium parnassicum (Čelak.) Wettst. (1889: 376). Lectotype (designated here):��� GREECE. Central Greece, Parnass [In m. Parnassi], above Arachova [reg. inferiori supra Rachova], 914.4 m [alt. 3000���], July 1855, Guicciardi J. 393 (S-G 6069!, scan seen; image of the lectotype available at https://herbarium. nrm.se/specimens/S-G-6069, isolectotype JE00011347!, scan seen; image of the isolectotype available at http://131.130.131.10/ herbaria/jacq-viewer/viewer.html?rft_id=je_00011347&identifiers=je_00011347; isolectotype P04283824!, scan seen; image of the isolectotype available at http://coldb.mnhn.fr/catalognumber/mnhn/p/p04283824; isolectotype L2880110!, scan seen; image of the isolectotype available at https://bioportal.naturalis.nl/specimen/L.2880110). Notes:��� Čelakovsk�� (1883) described Teucrium montanum var. parnassicum based on a gathering from Mount Parnassus collected by Guicciardi in July 1855. Čelakovsk�� noted that the plant from Parnassus is smaller, densely hairy, with narrow leaves, with smaller calyx and corolla and shorter calyx teeth than the typical T. montanum. According to Čelakovsk�� (1883), when considering the overall variability of the present forms, there are not enough differences between the typical T. montanum and his newly described variety, and he therefore described it as var. parnassicum. Subsequently, this vaiety was raised to a species level by Wettstein (1889). The original material was collected also from Aglassan-Dagh (Turkey). Wettstein noted that specimen completely correspond to those collected by Guicciardi from Parnassus. The original material of T. montanum var. parnassicum collected by Guicciardi is deposited at the herbarium S, and it fits with the description provided in the protologue. Hence, we here designate the specimen S-G 6069 as the lectotype of the name T. montanum var. parnassicum. The lectotypification of E. Bertels on the label of the specimen selected here as lectotype is not effective publication (Art. 7.10)., Published as part of Zbilji��, Milo��, Laku��i��, Dmitar, Stevanoski, Ivana & Kuzmanovi��, Nevena, 2022, Lectotypification of names related to Teucrium montanum L. (Lamiaceae) reported for the Balkan peninsula, pp. 198-204 in Phytotaxa 530 (2) on page 201, DOI: 10.11646/phytotaxa.530.2.6, http://zenodo.org/record/5832740, {"references":["Celakovsky, L. (1883) Ueber einige Arten der Gattung Teucrium. Botanisches Centralblatt; referierendes Organ fur das Gesamtgebiet der Botanik 14 (4): 151 - 155.","Wettstein, R. von (1889) Beitrag zur Flora des Orientes. Bearbeitung der von Dr. A. Heider im Jahre 1885 in Pisidien und Pamphylien gesammelten Pflanzen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch naturwissenschaftliche Classe Abt. 1 98 (1): 348 - 398."]}

Teucrium helianthemoides Adamović (1904: 136) ��� Teucrium montanum subsp. helianthemoides (Adamović) Baden (1991: 74). Lectotype (designated here):��� GREECE. Thessaly: Olympus [In Olympo thessalo], June 1902, Kindl s.n. (WU 0073104!, scan seen; image of the lectotype available at http://131.130.131.10/djatoka/jacq-viewer/viewer.html?rft_id=wu_0073104&identifiers=wu_ 0073104). Notes:��� Adamović (1904) described Teucrium helianthemoides on the basis of the material previously collected by Kindl at Olympus in June 1902. The specimens were collected from alpine rocky grasslands. Indumentum density, the presence of two leaf forms, size of the terminal heads, the length of the pedicle, and the structure of the calyx were noted as the main differences from the closely related T. montanum. The illustration is published in the protologue together with the description. Subsequently, Baden (1991) proposed a new status and gave a new combination��� T. montanum subsp. helianthemoides. The main difference between the two Greek forms is the characteristics of the indumentum: T. montanum ssp. montanum possesses greyish-pubescent stems with reflexed hairs, appressed-puberulent on the upper side and greyish-tomentose on lower sides of the leaves, and appressed-puberulent to subglabrous calyces, whereas T. montanum ssp. helianthemoides possesses white-pilose to velutinous stems with patent hairs, patent-pilose to the velutinous upper side and tomentose to lanate lower sides of the leaves, and patent-pilose calyces (Baden 1991). The original material was found in herbarium WU and it fits with the description provided in the protologue. We here designate the specimen deposited at WU (WU 0073104) as the lectotype of the name T. helianthemoides., Published as part of Zbilji��, Milo��, Laku��i��, Dmitar, Stevanoski, Ivana & Kuzmanovi��, Nevena, 2022, Lectotypification of names related to Teucrium montanum L. (Lamiaceae) reported for the Balkan peninsula, pp. 198-204 in Phytotaxa 530 (2) on page 199, DOI: 10.11646/phytotaxa.530.2.6, http://zenodo.org/record/5832740, {"references":["Adamovic, L. (1904) Flora von Macedonien und Altserbien. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematischnaturwissenschaftliche Klasse 74: 115 - 150.","Baden, C. (1991) Teucrium L. In: Strid, A. & Tan, K. (Eds.) Mountain flora of Greece, Vol. 2. Edinburgh University Press, Edinburgh, pp. 69 - 75."]}

Teucrium pannonicum Kerner (1863: 384) ��� Teucrium montanum var. pannonicum (A.Kern.) Nyman (1881: 566) ��� Teucrium montanum subsp. pannonicum (A.Kern.) Domin (1936: 193) Lectotype (designated here):��� ROMANIA. Bihor, mount Bihor [Bontos k��], Beius [Belenyes], Kerner s.n. (WU 0070075!, scan seen; image of the lectotype available at: http://131.130.131.10/djatoka/jacq-viewer/viewer.html?rft_id=wu_0070075&identifiers=wu_ 0070075). Notes:��� Kerner has described Teucrium pannonicum on the basis of material collected in Transylvania (Romania) on the southern slopes of the Mount Bihor (��� Botnos k�����) near Beius (��� Belenyes ���). He noted that T. pannonicum differs from the typical T. montanum by the size of the plant, the presence of woolly hairs on the stem, broader leaves densely covered with soft woolly hairs on both sides, and toothed calyxes with setaceous narrow tips. Subsequently, Nyman (1881) included T. pannonicum in T. montanum s.l. while Domin (1936) proposed a new status for the species Kerner described, at the subspecies level ��� T. montanum subsp. pannonicum. The original specimen is deposited in the herbarium WU and it fits with the description provided in the protologue. We here designate the specimen WU 0070075 as the lectotype of the name Teucrium pannonicum., Published as part of Zbilji��, Milo��, Laku��i��, Dmitar, Stevanoski, Ivana & Kuzmanovi��, Nevena, 2022, Lectotypification of names related to Teucrium montanum L. (Lamiaceae) reported for the Balkan peninsula, pp. 198-204 in Phytotaxa 530 (2) on pages 201-202, DOI: 10.11646/phytotaxa.530.2.6, http://zenodo.org/record/5832740, {"references":["Kerner, A. (1863) Descriptiones plantarum novarum florae hungaricae et transsilvanicae. Oesterreichische Botanische Zeitschrift 13 (10): 384 - 385. https: // doi. org / 10.1007 / BF 01678940","Domin, K. (1936) Plantarum Cechoslovakiae enumeratio, species vasculares indigenas et introductas exhibens. Vytiskla Statni Tiskarna v Praze, Praha. Preslia 13 - 15: 1 - 305."]}

Teucrium montanum var. modestum Heldreich (1898: 1464) Lectotype (designated here):��� GREECE. Peloponnese: Taygetos, Megala Zonaria [M. Taygetus: l. d. Megala Zonaria], in the alpine region [in regione alpina], 1828.8 m [6000���], August 1897, Heldreich 1464 (WU0073103! scan seen; image of the lectotype available at http://131.130.131.10/djatoka/jacq-viewer/viewer.html?rft_id=wu_0073103&identifiers=wu_0073103; isolectotype JE00011339!, scan seen; image of the isolectotype available at http://131.130.131.10/herbaria/jacq-viewer/viewer.html?rft_id=je_ 00011339&identifiers=je_00011339; isolectotype K000881466!, scan seen; image of the isolectotype available at http://www.kew. org/herbcatimg/489415.jpg). Notes:��� Heldreich (1898) described Teucrium montanum var. modestum on the basis of the material collected in Greece, from the alpine region of Mount Taygetos. He noted on the label that the plants from Taygetos differ from T. montanum var. parnassicum by the smaller size of leaves and corolla, and capitulum with fewer flowers. The original material of T. montanum var. modestum collected by Heldreich is deposited at multiple herbaria ��� JE, K, WU. We here designate the specimen WU 073103 as the lectotype of the name Teucrium montanum var. modestum as it fits best the description. The same specimen was cited as syntype in the paper of Reich et al. (2021). The name with diagnosis appears in printed label of Heldreich��s Herbarium Graecum normale., Published as part of Zbilji��, Milo��, Laku��i��, Dmitar, Stevanoski, Ivana & Kuzmanovi��, Nevena, 2022, Lectotypification of names related to Teucrium montanum L. (Lamiaceae) reported for the Balkan peninsula, pp. 198-204 in Phytotaxa 530 (2) on page 201, DOI: 10.11646/phytotaxa.530.2.6, http://zenodo.org/record/5832740, {"references":["Heldreich, T. H. H. von (1898) Herbarium graecum normale 15. No. 1401 - 1500."]}

Teucrium montanum var. hirsutum Boissier (1879: 819) Lectotype (designated here):��� GREECE. Central Greece: Euboea, Dirfi [Mons Dirphys Euboeae], in the higher zone [in reg. superiori], 1219.2 - 1676.4 m [alt. 4000���-5500���], 5 August 1868, Heldreich 2757 (G00741484!), Figure 1. Notes:��� The name Teucrium montanum var. hirsutum was proposed by Boissier (1879) to describe plants from Greece (Olympus Thessaly and Dirfi Euboea) collected by Heldreich and plants from Turkey (mount Spil above Manisa) collected by Balansa. The localities from Greece were originally cited as ���Olympo Thessalo et monte Delphi Eubeae���, the second locality refers to Mt Dirfis on the island of Euboea (Evvia). The name ��� Monte Delphi Eubeae��� is frequently used by Boissier and his contemporaries, and has sometimes been misinterpreted by later authors who incorrectly believed he was referring to the archaeological site of Delphi at the foot of Mt Parnassos (A. Strid pers. comm.). The presence of the hispid indumentum on the stems, leaves and calyxes is the most important diagnostic feature. The original material is preserved at the herbarium G, and it fits with the description provided in the protologue. We here designate the specimen G00741484 collected in Greece as the lectotype of the name Teucrium montanum var. hirsutum., Published as part of Zbilji��, Milo��, Laku��i��, Dmitar, Stevanoski, Ivana & Kuzmanovi��, Nevena, 2022, Lectotypification of names related to Teucrium montanum L. (Lamiaceae) reported for the Balkan peninsula, pp. 198-204 in Phytotaxa 530 (2) on pages 200-201, DOI: 10.11646/phytotaxa.530.2.6, http://zenodo.org/record/5832740, {"references":["Boissier, E. (1879) Flora Orientalis: sive, Enumeratio plantarum in Oriente a Graecia et Aegypto ad Indiae fines hucusque observatarum, Vol. 4. Genevae et Basileae, Apud H. Georg, Lugduni: Apud H. Georg, 1288 pp."]}