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The fauna of the click-beetles of the Sikhote-Alin Nature Reserve and adjacent areas is listed and analyzed. The current checklist of the Elateridae of this territory comprises 77 species of 36 genera, 11 tribes, and 5 subfamilies. Several species are recorded for the first time in the following territories: 54 species in the fauna of the Sikhote-Alin Nature Reserve and adjacent areas, 7 species in Primorskii Territory, 5 in the mainland Far East of Russia. Two speceis are recorded by us in the Sikhote-Alin Nature Reserve and adjacent areas only from published data without examination of the material; the doubtful records of 4 species require confirmation and are not included in the checklist. Notes on the distribution and taxonomy and new data on the morphology, including the genital characters, of a number of the species are also given. The following new synonymy is established: Cardiophorus inflatus Candeze, 1882 = Cardiophorus heydeni Buysson, 1915, syn. n. The following new combination is proposed: Pseudocrepidophorus laetus (Candeze, 1879), comb. n. (ex Crepidophorus).

Females of two click beetle species, Cardiophorus tenebrosus and C. edwardsi (Coleoptera: Elateridae), produce methyl (3R,6E)-2,3-dihydrofarnesoate as their sex pheromone. We had serendipitously discovered that males of both species were also strongly attracted to (R)-fuscumol acetate ((E)-6,10-dimethylundeca-5,9-dien-2-yl acetate), a known longhorned beetle (Coleoptera: Cerambycidae) pheromone, due to its structural similarities to the click beetle pheromone. To further investigate the specificity of the responses of Cardiophorus males, additional analogs with different chain lengths and structural relationships compared to the natural pheromone were synthesized and tested. In field and electroantennogram bioassays, only fuscumol propionate ((E)-6,10-dimethylundeca-5,9-dien-2-yl propionate) elicited strong responses from Cardiophorus males, indicating that they were able to distinguish chain length and spatial relationships between the structural elements. In field trials, C. tenebrosus males were attracted equally to the analog and their natural pheromone, but the pheromone elicited stronger antennal responses from males. In contrast, traps baited with fuscumol propionate caught approximately 26 times as many C. edwardsi males compared to traps baited with the natural pheromone, although the analog elicited significantly smaller antennal responses from C. edwardsi males. Thus, in terms of behavioral responses, fuscumol propionate appears to be acting as a hyperactive pheromone mimic, a phenomenon which has rarely been observed in insect semiochemistry.

Cardiophorus magnanii Platia, Furlan & Gudenzi, 2002 (Figs 11–12, Map 8) Material examined (HMIM). Ardabil province: 5 exs: " ARDABIL, Daryâcheh-e Neour [Neor lake], [N37°59' E48°33'], 2250 m, 1.VII.1997, [leg.] Bar./Mof.". Azarbaijan-e Gharbi province: 2 exs.: Mirabad Protected Area, Sardasht to Mirabad rd., nr. Nalas, N36°20'14.6", E45°25'05.8", 1357 m, 27. V.2016, leg. H. Nasserzadeh. Zanjan province: 1 ex.: Sorkhabad Protected Area, Mari to Golchin rd., ca. 8 km NE Golchin, Solodarreh, N37°04'17.5", E48°27'36.6", 1898 m, 22. V.2016, leg. H. Nasserzadeh. Climate types. Sub-humid and semi-arid, cold winters and warm summers., Published as part of Nasserzadeh, Hiva, 2021, A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n., pp. 338-368 in Zootaxa 4941 (3) on pages 362-363, DOI: 10.11646/zootaxa.4941.3.2, http://zenodo.org/record/4595227, {"references":["Platia, G., Furlan, L. & Gudenzi, I. (2002) descrizione di sei nuove species di Elateridi dell' Iran (Insecta, Coleoptera, Elateridae). Il Naturalista Valtellinese: Atti del museo civico di storia naturale di Morbegno, 13, 65 - 77."]}

Cardiophorus vestigialis Erichson, 1840 (Map 8) Material examined (HMIM). Azarbaijan-e Gharbi province: 3 exs.: Sardasht, Mirabad to Alootan rd., ca. 3 km W Mirabad, N36°22'53.6", E45°20'15.6", 1650 m, 27.V.2016, leg. H. Nasserzadeh. Ardabil province: 5 exs.: " ARDABIL, Daryâcheh-e Neour [Neor lake], [N37°59' E48°33'], 2250 m, 1.VII.1997, [leg.] Bar./Mof.". Climate types. Sub-humid, cold winters and mild to warm summers., Published as part of Nasserzadeh, Hiva, 2021, A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n., pp. 338-368 in Zootaxa 4941 (3) on page 363, DOI: 10.11646/zootaxa.4941.3.2, http://zenodo.org/record/4595227

The Chinese species of Phorocardius Fleutiaux, 1931 have been studied and six species are described as new: P. alterlineatus Ruan & Douglas, sp. nov.; P. flavistriolatus Ruan & Douglas, sp. nov.; P. minutus Ruan & Douglas, sp. nov.; P. rufiposterus Ruan & Douglas, sp. nov.; P. yunnanensis Ruan & Douglas, sp. nov.; and P. zhiweii Ruan, Douglas & Qiu, sp. nov. Lectotypes are designated for Cardiophorus comptus Candèze, 1860, Cardiophorus contemptus Candèze, 1860, Phorocardius magnus Fleutiaux, 1931, and Cardiophorus manuleatus Candèze, 1888. The holotype is identified for Cardiophorus yanagiharae Miwa, 1927. Phorocardius florentini (Fleutiaux, 1895) and P. manuleatus (Candèze, 1888) are newly reported from China; P. comptus (Candèze, 1860) is excluded from the Chinese fauna. A key to the 11 Phorocardius species known from China is given. Phorocardius is newly recorded from deep within the Palearctic Region. The procoxal cavities of P. rufiposterus Ruan & Douglas, sp. nov. are closed, which is different from all other species of Phorocardius. An annotated checklist of the 21 Phorocardius species of the world is provided. Additionally, Phorocardius contemptus (Candèze, 1860), comb. nov. is transferred from Cardiophorus to Phorocardius; four species are transferred from Phorocardius to Displatynychus: Displatynychus bombycinus (Candèze, 1895), comb. nov., Displatynychus pakistanicus (Platia & Ahmed, 2016), comb. nov., Displatynychus sobrinus (Laporte, 1840), comb. nov., and Displatynychus tibialis (Platia & Ahmed, 2016), comb. nov.

Cardiophorus carinatus Mathison and Douglas sp. nov. (Coleoptera: Elateridae) is described from the Lake Wales Ridge ecosystem of Florida, USA. In addition, C. robustus LeConte, 1853, is rediscovered in North America from Massachusetts, Maine, and Rhode Island.

Ninty-five species of Elateridae, from 30 genera and five subfamilies are documented from Turkey based on specimens collected throughout the country. Ampedus sanguineus (Linnaeus, 1758), A. sinuatus Germar, 1844; Elater ferrugineus Linnaeus, 1758; Agriotes acuminatus (Stephens, 1830); A. brevis Candeze, 1863; A. turcicus Candeze, 1863, Lacon punctatus (Herbst, 1779) and Idotarmonides anatolicus Agajev, 1985 are recorded for the first time in the Eastern Anatolian Region. Those of Cardiophorus cyanipennis Mulsant and Wachanru, 1852 and Mulsanteus guillebeaui (Mulsant and Godart, 1853) are recorded from the southeastern Anatolia region. New distribution records of the most of the species are evaluated. So that, each species show different distribution pattern; most of species are sparsely collected, the rest of examined species have moderate or widespread distributions. Some species were recorded from one or two provinces, even a single sample. They might be classified as endangered species and sho...

Cardiophorus convexithorax Desbrochers des Loges, 1869 (Fig. 9) Cardiophorus convexithorax Desbrochers des Loges, 1869: 32 [= 1870b: 99]. Material examined. ��� LEBANON, Mount Lebanon Range, Arsoun, Pinus forest, swept & beaten, 33��51���8.03���N, 35��41���44.36���E, 7.V.2017, leg. A. Kot��n, P. Nemes & T. N��meth (No. 42)��� (4, HNHM); same but ��� from Tomicus pheromone trap, X.2016, leg. N. Nemer ��� (4, HNHM); ��� LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Tannourine Cedars Nat. Reserve, swept, 34��12���34���N, 35��55���45���E, 1750 m, 6.V.2017, leg. A. Kot��n, P. Nemes & T. N��meth (No. 40)��� (1, HNHM) same but ��� 1.V. (No. 30)��� (1, HNHM); same but ��� 10.V. (No. 47)��� (2, HNHM); same but ��� pitfall trap, 3- 11.V. (No.56)��� (1, HNHM); ��� LEBANON, Northern gov., Bcharre env., Quadisha Valley, swept & beaten, 34��14���57���N, 35��58���34���E, 970 m, 5.V.2017, leg. A. Kot��n, P. Nemes, T. N��meth & C. Tanios (No. 38)��� (2, HNHM); ��� LEBANON, Northern gov., Bcharre env., 1 km E Ariz, Horsh Arz el-Rab, ancient Cedrus forest, swept & singled, 34��14���33���N, 36��2���59���E, 1900 m, 22.V.2015, leg. A. M��rkus, T. N��meth, M. Rehayem & W. Yammine ��� (1, HNHM); ��� LEBANON, Bekaa Valley, Taanayel, university farm, 33��47���33.63���N, 35��52���2.41���E, 878 m, 8.V.2017, leg. A. Kot��n, P. Nemes & T. N��meth (No. 43)��� (2, HNHM); ��� LEBANON, Mount Lebanon Range, Kfar Hay, Mar Youanna Maroun, macchia, swept & hand-collected, 34��14���47.69���N, 35��45���1.02���E, 322 m, 3.V.2017, leg. M. Boustani, A. Kot��n, P. Nemes & T. N��meth (No. 34)��� (7, HNHM); ��� LEBANON, Bekass valley, Haouch Al Nabi, 18.IV.2018, leg. W. Yammine ��� (3, HNHM); ��� Leban ��� (2, HNHM); ��� LEBANON, Northern gov., Tannourine El Tahta, macchia, beaten, 34��12���46.84���N, 35��52���26.40���E, 960 m, 3.V.2017, leg. M. Boustani, A. Kot��n, P. Nemes & T. N��meth (No. 33)��� (2, HNHM); same but ���beaten, 10.V.2017 ��� (1, HNHM). Published data. ��� Lebanon: El Arz (Bcherri), m 1950, 1.V.1997, D. Gianasso ���; ��� Beqaa Valley, Zahle Kfar Dabach, 9.V.2000, G. Sama ���; ��� Nabh Safa, 1000 m, V.1966, G. Fagel ���; ��� Jbail prov., Qartaba, 8.V. 2000, P. Rapuzzi ��� (Platia 2018: 78) [without number of specimens and place of depository]. Distribution. N: Egypt. A: Israel, Syria (Cate 2007), Jordan, Lebanon (Platia 2018)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 132, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Desbrochers des Loges, J. (1869) [La description d'une assez grand nombre d'especes nouvelles d'Europe et confins]. Petites Nouvelles Entomologiques, 1 (8), 29 - 30, 33 - 34, 45 - 46.","Desbrochers des Loges, J. (1870 b) Descriptions de coleopteres nouveaux d'Europe et confins. L'Abeille, Memoires d'Entomologie, 7, 97 - 135. [1869 - 1870]","Platia, G. (2018) New species and new records of click beetles from the palearctic region (Coleoptera, Elateridae). Boletin de la Sociedad Entomologica Aragonesa, 62, 71 - 82.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus maculicollis Reiche & Saulcy, 1857 (Fig. 11) Cardiophorus maculicollis Reiche & Saulcy, 1857: 420. Material examined. “ LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Tannourine Cedars Nat. Reserve, swept, 34°12’34”N, 35°55’45”E, 1750 m, 23.V.2015, leg. O. Akiki, M. Boustani, A. Márkus, N. Nemer, T. Németh, M. Rehayem & W. Yammine” (2, HNHM); “ LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Fuvar, stream valley, canopy netting, 34°12’22”N, 35°55’17”E, 1460 m, 17.VI.2016, leg. A. Kotán, P. Nemes & T. Németh ” (1, HNHM); “ LEBANON, Northern gov., Bcharre env., Quadisha Valley, swept & beaten, 34°14’57”N, 35°58’34”E, 970 m, 5.V.2017, leg. A. Kotán, P. Nemes, T. Németh & C. Tanios (No. 38)” (4, HNHM); LEBANON, Batroun distr., Tannourine, Wadi Tannourine, 35.195070 N, 35.892637 E, 1166 m alt., 21.V.2012, leg. C. Cocquempot ” (1, PCNN). Distribution. E: Greece. A: Turkey (Cate 2007). New for Lebanon.

Cardiophorus sp. (Fig. 48) Material examined. ��� Lebanon, Salima (2), Pinus pinea ecosyst. 07/06/1997, Masri, Routier, Nemer, ref12C, IIE 23988 ��� (1, PCNN). Remarks. The only available specimen is a male with broken legs and antennae. It probably represents a new species with atrophied wings similar to C. schuberti Platia & Gudenzi, 2002. More material should be examined to make a conclusion about its status., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on pages 135-136, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194

Cardiophorus miniaticollis Cand��ze, 1860 Cardiophorus miniaticollis Cand��ze, 1860: 122. Distribution. E: Armenia, Greece. A: Iran, Lebanon, Syria, Turkey (Cate 2007), Jordan (Platia 2010a)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 134, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Candeze, E. C. A. (1860) Monographie des elaterides. III. Memoires de Ia Societe Royale des Sciences de Liege, 15, 1 - 512, 5 pls.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2010 a) New species and chorological notes of click beetles from Palearctic Region, especially from the Middle East (Coleoptera, Elateridae). Boletin de la Sociedad Entomologica Aragonesa, 46, 23 - 49."]}

Cardiophorus tenellus Reiche & Saulcy, 1857 (Fig. 46) Cardiophorus tenellus Reiche et Saulcy, 1857: 421. Material examined. ��� LEBANON, Northern gov., Ehden, Horsh Ehden Natural Reserve, swept & singled, 34��18���33���N, 35��59���14���E, 1525 m, 21.V.2015, leg. M. Boustani, A. M��rkus & T. N��meth ��� (1, HNHM). Published data. Original description. Type locality: ��� Beyrouth [Lebanon]��� (Reiche & Saulcy 1857: 422). Distribution. N: Egypt. A: Israel, Jordan, Lebanon, Syria (Cate 2007)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 135, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Reiche, L. J. & Saulcy, F. de (1857) Especes nouvelles ou peu connus de coleopteres, recueillies par M. F. de Saulcy. Membre de l'Institut, dans son voyage en orient, et decrites par MM. Annales de la Societe entomologique de France, 3, 385 - 422.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus sacratus Erichson, 1840 Cardiophorus sacratus Erichson, 1840: 284. Cardiophorus nigricornis Baudi di Selve, 1859: 342. Cardiophorus augustalisi Pic, 1913: 169. Cardiophorus tokatensis Pic, 1913: 169. Distribution. E: Azerbaijan, Armenia, Georgia, South European territory of Russia. N: Egypt, Libya. A: Cyprus, Iraq, Iran, Israel, Jordan, Lebanon, Syria, Turkey, Turkmenistan (Cate 2007), Usbekistan (Platia 2010b)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 134, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Erichson, W. F. (1840) Entomographien, untersuchungen in dem Gebiete der Entomologie, mit besonderer Benutzung der Konigl. Sammlung zu Berlin. Erstes Heft. F. H. Morin, Berlin, 180 pp., 2 pls. https: // doi. org / 10.5962 / bhl. title. 47509","Baudi di Selve, F. (1859) Kleine Mitteilungen. Berliner Entomologische Zeitschrift, 3, 321 - 342.","Pic, M. (1913) Notes diverses, descriptions et diagnoses (Suite). L'Echange, Revue Linneenne, 29, 169 - 171.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2010 b) Descriptions of fourteen new species of click beetles from the Palearctic Region, with chorological notes (Insecta Coleoptera Elateridae). Quaderno di Studi e Notizie di Storia Naturale della Romagna, 30, 103 - 130."]}

Cardiophorus megathorax Faldermann, 1835 Cardiophorus megathorax Faldermann, 1835: 171. Cardiophorus antennalis Desbrochers des Loges, 1875: 39. Distribution. E: Azerbaijan, Georgia, Greece (Samos). A: Iran, Israel, Jordan, Lebanon, Syria, Turkey (Cate 2007)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 134, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Faldermann, F. (1835) Addimenta entomologica ad faunam rossicam in itineribus Jussu Imperatoris Augustissimi annis 1827 - 1831 a Cl. Menetries et Szovitz susceptis collecta, in lucem edita. Nouveaux Memoires de Ia Societe Imperiale des Naturalistes de Moscou, Series 2, 4, 1 - 310, 10 pls.","Desbrochers des Loges, J. (1875) Description d'un Throscus et de quelques elaterides nouveaux. Opuscules Entomologiques, Coleopteres, 1, 37 - 56. [1874 - 1785]","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus syriacus (Linnaeus, 1758) (Fig. 12) Elater syriacus Linnaeus, 1758: 404. Cardiophorus pictus Faldermann, 1835: 170. Material examined. “ LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Tannourine Cedars Nat. Reserve, canopy netting, 34°12’34”N, 35°55’45”E, 1750 m, 16.VI.2016, leg. A. Kotán, P. Nemes & T. Németh ” (1, HNHM). Distribution. E: Azerbaijan, Armenia, Georgia. A: Iran, Syria, Turkey, Turkmenistan (Cate 2007). New for Lebanon., Published as part of Németh, Tamás, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 134, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp. https: // doi. org / 10.5962 / bhl. title. 542","Faldermann, F. (1835) Addimenta entomologica ad faunam rossicam in itineribus Jussu Imperatoris Augustissimi annis 1827 - 1831 a Cl. Menetries et Szovitz susceptis collecta, in lucem edita. Nouveaux Memoires de Ia Societe Imperiale des Naturalistes de Moscou, Series 2, 4, 1 - 310, 10 pls.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus maculicollis Reiche & Saulcy, 1857 (Fig. 11) Cardiophorus maculicollis Reiche & Saulcy, 1857: 420. Material examined. ��� LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Tannourine Cedars Nat. Reserve, swept, 34��12���34���N, 35��55���45���E, 1750 m, 23.V.2015, leg. O. Akiki, M. Boustani, A. M��rkus, N. Nemer, T. N��meth, M. Rehayem & W. Yammine��� (2, HNHM); ��� LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Fuvar, stream valley, canopy netting, 34��12���22���N, 35��55���17���E, 1460 m, 17.VI.2016, leg. A. Kot��n, P. Nemes & T. N��meth ��� (1, HNHM); ��� LEBANON, Northern gov., Bcharre env., Quadisha Valley, swept & beaten, 34��14���57���N, 35��58���34���E, 970 m, 5.V.2017, leg. A. Kot��n, P. Nemes, T. N��meth & C. Tanios (No. 38)��� (4, HNHM); LEBANON, Batroun distr., Tannourine, Wadi Tannourine, 35.195070 N, 35.892637 E, 1166 m alt., 21.V.2012, leg. C. Cocquempot ��� (1, PCNN). Distribution. E: Greece. A: Turkey (Cate 2007). New for Lebanon., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 134, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Reiche, L. J. & Saulcy, F. de (1857) Especes nouvelles ou peu connus de coleopteres, recueillies par M. F. de Saulcy. Membre de l'Institut, dans son voyage en orient, et decrites par MM. Annales de la Societe entomologique de France, 3, 385 - 422.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus vestigialis Erichson, 1840 (Fig. 47) Cardiophorus vestigialis Erichson, 1840: 293. Cardiophorus curtulus Mulsant & Guillebeau, 1855b: 317. Cardiophorus atripes Buysson, 1902c: 425. Cardiophorus captiosus Normand, 1941: 333. Cardiophorus hipponensis Desbrochers des Loges, 1870b: 102. Cardiophorus infimus Buysson, 1906: 471. Cardiophorus laetipes Buysson, 1902c: 425. Cardiophorus maurus Desbrochers des Loges, 1870a: 71. Cardiophorus mauritanicus Desbrochers des Loges, 1869: 34 [= 1870b: 103] [homonym]. Cardiophorus nijazi Dolin & Agajev, 1983: 31. Cardiophorus neotericus Buysson, 1902a: 228. Cardiophorus persicus Schwarz, 1892: 384. Cardiophorus ragusae Ragusa, 1911: 194. Cardiophorus rubidiventris Buysson, 1919: 116. Material examined. ��� LEBANON, Northern gov., Tannourine env., 2 km N Harissa, Tannourine Cedars Nat. Reserve, canopy netting, 34��12���34���N, 35��55���45���E, 1750 m, 23.V.2015, leg. O. Akiki, M. Boustani, A. M��rkus, N. Nemer, T. N��meth, M. Rehayem & W. Yammine��� (1, HNHM); same but ��� 16.VI.2016, leg. A. Kot��m, P. Nemes & T. N��meth (No. 1) (2, HNHM); same but ��� 18.VI.2016, leg. A. Kot��n, P. Nemes & T. N��meth (No. 6) (2, HNHM); same but ��� 21.VI.2016, leg. P. Nemes & T. N��meth (No. 14)��� (1, HNHM); same but ��� 10.V.2017, leg. A. Kot��n, P. Nemes & T. N��meth (No. 47)��� (1, HNHM); same but ��� 25.VII.2018, leg. P. Nemes & T. N��meth (No. 62)��� (1, HNHM); ��� LEBANON, Mount Lebanon Range, Arsoun, Pinus forest, swept & beaten, 33��51���8.03���N, 35��41���44.36���E, 7.V.2017, leg. A. Kot��n, P. Nemes & T. N��meth (No. 42)��� (1, HNHM); ��� LEBANON, Northern gov., Bcharre env., Quadisha Valley, swept & beaten, 34��14���57���N, 35��58���34���E, 970 m, 5.V.2017, leg. A. Kot��n, P. Nemes, T. N��meth & C. Tanios (No. 38)��� (1, HNHM). Published data. ���Akkar, Fnaideq, m 1200-1600, 14.VI.1999, G. Magnani���; ���Zgharta, Horsh Edhen Nat. Res., m 1350���1500, 8.18. VI.1999, G. Magnani���; ���Byblos, Laqlouq, 1300���1400 m, 25.VI.2009, Padovani & Malmusi���; ���caza Bcharre, Horch Ehden, 1400���1600 m, 19-26.VI.2009, Padovani & Malmusi���; ���caza Aakkar, Ain Yaaqoub, 750 m, 20���24.VI.2009, L. Saltini, at light��� (PCPG, MCSN, PCMM) (Platia 2010a: 39). Distribution. E: Azerbaijan, Albania, Armenia, Austria, Belgium, Bosnia & Herzegovina, Bulgaria, Croatia, Central European territory of Russia, Czech Republic, Finland, France, Georgia, Germany, Great Britain, Greece, Hungary, Italy, Moldavia, Northern European territory of Russia, Norway, Poland, Portugal, Romania, Serbia and Montenegro, Slovakia, Slovenia, Spain, Sweden, South European territory of Russia, Switzerland, Ukraine. N: Algeria, Egypt, Libya, Morocco, Tunesia. A: Cyprus, East Siberia & Far East of Russia, Iran, Kazakhstan, Mongolia, Turkmenistan, Turkey, (Cate 2007), Jordan, Lebanon, Syria (Platia 2010a)., Published as part of N��meth, Tam��s, 2019, Elateridae (Coleoptera) of Lebanon, pp. 127-159 in Zootaxa 4614 (1) on page 135, DOI: 10.11646/zootaxa.4614.1.6, http://zenodo.org/record/3241194, {"references":["Erichson, W. F. (1840) Entomographien, untersuchungen in dem Gebiete der Entomologie, mit besonderer Benutzung der Konigl. Sammlung zu Berlin. Erstes Heft. F. H. Morin, Berlin, 180 pp., 2 pls. https: // doi. org / 10.5962 / bhl. title. 47509","Mulsant, E. & Guillebeau, F. (1855 b) Description de deux nouvelles especes delaterides. Annales de Ia Societe Linneenne de Lyon, Series 2, 2, 317 - 320.","Buysson, H. (1902 c) Remarques sur quelques elaterides du genre Cardiophorus et descriptions despeces nouvelles. Annales de Ia Societe Entomologique de France, 71, 422 - 439.","Normand, Η. (1941) Remarques sur les especes Nord-Africaines du groupe du Cardiophorus rufipes Goeze. Bulletin de Ia Societe dHistoire Naturelle de l'Afrique du Nord, 32, 331 - 334.","Desbrochers des Loges, J. (1870 b) Descriptions de coleopteres nouveaux d'Europe et confins. L'Abeille, Memoires d'Entomologie, 7, 97 - 135. [1869 - 1870]","Buysson, H. (1906) Coleopteres. Vol. 5. Elateridae. In: Fauvel, C. Α. Α. (Ed.), Faune gallo-rhenane ou description des insectes qui habitent la France, la Belgique, la Hollande, les Provinces Rhenanes et le Valais. Revue d'Entomologie, 25 (Supplement), pp. 417 - 494.","Desbrochers des Loges, J. (1870 a) Description de coleopteres nouveaux algeriens et remarques diverses. Bulletin de l'Academie d'Hippone, 9, 68 - 82.","Desbrochers des Loges, J. (1869) [La description d'une assez grand nombre d'especes nouvelles d'Europe et confins]. Petites Nouvelles Entomologiques, 1 (8), 29 - 30, 33 - 34, 45 - 46.","Dolin, V. G. & Agajev, B. I. (1983) Novye vidy zhukov-shchelkunov (Coleoptera, Elateridae) iz Azerbaydzhanskoy SSR [New click-beetle species (Coleoptera, Elateridae) from the Azerbaidjan SSR]. Vestnik Zoologii, 1983 (6), 31 - 34.","Buysson, H. (1902 a) Notes sur le Cardiophorus argiolus Gene, var. (Col.). Bulletin de la Societe Entomologique de France, 1902, 228.","Ragusa, E. (1911) Coleotteri nuovi o poco conosciuti della Sicilia. Rivista Coleotterologica Italiana, 3, 190 - 197.","Buysson, H. (1919) Contributions a la faune du Maroc et du Nord de l'Afrique. Elaterides Col. (2 e Note). Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord, 10, 114 - 116.","Platia, G. (2010 a) New species and chorological notes of click beetles from Palearctic Region, especially from the Middle East (Coleoptera, Elateridae). Boletin de la Sociedad Entomologica Aragonesa, 46, 23 - 49.","Cate, P. (2007) Elateridae. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209."]}

The prior genus-level classification of Cardiophorinae had never been assessed phylogenetically, and not revised since 1906. A phylogeny for Cardiophorinae and Negastriinae is inferred by Bayesian analyses of 163 adult morphological characters to revise the generic classification. Parsimony analysis is also performed to assess the sensitivity of the Bayesian results to the choice of optimality criterion. Bayesian hypothesis testing rejected monophyly for: Negastriinae; Cardiophorinae (but monophyletic after addition of four taxa); Cardiophorini; cardiophorine genera Aphricus LeConte, 1853; Aptopus Eschscholtz, 1829; Cardiophorus Eschscholtz, 1829; Cardiotarsus Eschscholtz, 1836; Paracardiophorus Schwarz, 1895; Phorocardius Fleutiaux, 1931; Dicronychus sensu Platia, 1994; Dicronychus sensu Méquignon, 1931; Craspedostethus sensu Schwarz, 1906 (i.e., including Tropidiplus Fleutiaux, 1903); Paracardiophorus sensu Cobos, 1970, although well-supported alternative classifications were available for only some. Based on taxonomic interpretation of phylogenetic results: Nyctorini is syn. n. of Cardiophorini; Globothorax Fleutiaux, 1891 (Physodactylinae), Margogastrius Schwarz, 1903 (Physodactylinae), and Pachyelater Lesne, 1897 (Dendrometrinae) are transferred to Cardiophorinae. The following changes are proposed for cardiophorine genera: Aptopus Eschscholtz, 1829 is redefined to exclude Horistonotus-like species; Coptostethus Wollaston, 1854 is subgenus of Cardiophorus; Dicronychus Brullé, 1832 and Diocarphus Fleutiaux, 1947, Metacardiophorus Gurjeva, 1966, Platynychus Motschulsky, 1858, and Zygocardiophorus Iablokoff-Khnzorian and Mardjanian, 1981 are placed at genus rank; Paracardiophorus Schwarz, 1895 is redefined based on North American and Eurasian species only; Horistonotus Candèze, 1860 redefined to include species with multiple apices on each side of their tarsal claws; Patriciella Van Zwaluwenburg, 1953 is syn. n. of Aphricus LeConte, 1853; Teslasena Fleutiaux, 1892 (Physodactylinae) is syn. n. of Globothorax Fleutiaux, 1891. The following new genera are described: Austrocardiophorus (type species: Cardiophorus humeralis Fairmaire and Germain, 1860); Chileaphricus (type species: Aphricus chilensis Fleutiaux, 1940); Floridelater (type species: Coptostethus americanus Horn, 1871, transferred from Negastriinae to Cardiophorinae). Paradicronychus (nomen nudum), is syn. n. of Cardiophorus Eschscholtz, 1829. Generic reassignments to make Cardiodontulus, Cardiophorus, Cardiotarsus, Paracardiophorus consistent with phylogenetically revised genus concepts resulted in 84 new combinations. Lectotypes are designated for 29 type species to fix generic concepts: Anelastes femoralis Lucas, 1857; Aphricus chilensis Fleutiaux, 1940; Athous argentatus Abeille de Perrin, 1894; Cardiophorus adjutor Candèze, 1875; Cardiophorus florentini Fleutiaux, 1895; Cardiophorus inflatus Candèze, 1882; Cardiophorus luridipes Candèze, 1860; Cardiophorus mirabilis Candèze, 1860; Cardiophorus musculus Erichson, 1840; Cardiotarsus capensis Candèze, 1860; Cardiotarsus vitalisi Fleutiaux, 1918; Craspedostethus rufiventris Schwarz, 1898; Elater cinereus Herbst, 1784; Elater minutissimus Germar, 1817; Elater sputator Linnaeus, 1758; Elater thoracicus Fabricius, 1801; Eniconyx pullatus Horn, 1884; Esthesopus castaneus Eschscholtz, 1829; Gastrimargus schneideri Schwarz, 1902; Globothorax chevrolati Fleutiaux, 1891; Horistonotus flavidus Candèze, 1860; Horistonotus simplex LeConte, 1863; Lesnelater madagascariensis Fleutiaux, 1935; Oedostethus femoralis LeConte, 1853; Phorocardius solitarius Fleutiaux, 1931; Platynychus indicus Motschulsky, 1858; Platynychus mixtus Fleutiaux, 1931; Triplonychus acuminatus Candèze, 1860; Tropidiplus tellinii Fleutiaux, 1903. A key to genera and diagnoses are provided for all genera and subgenera. A bibliographic synonymy includes references for all taxonomic changes to genera and new species through 2015.

Cardiophorus nigratissimus Buysson, 1891 (Map 3) Material examined (HMIM). Lorestan Province: 4 exs.: 8 km SW Khorramabad, next by the Shourab park, 33��26'44"N 48��14'01"E, 1144 m, 4.V.2008, leg. Nas. / Nem. / Montreuil; 2 exs.: Mamulan to Poldokhtar road, 7 km W Mamulan, 33��20'50"N 47��55'50"E, 910 m, 4.V.2008, leg. Nas. / Nem. / Montreuil. Hamedan Province: 46 exs.: 5 km S Hamedan, 34��43'48"N 48��29'33"E, 2322 m, 5.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 1 ex.: Ganjnameh to Tarik-e Darreh road, 34��45'09"N 48��26'21"E, 2318 m, 9.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 47 exs.: Goltappeh to Alisadr cave, 20 km S Alisadr cave, 35��12'39"N 48��13'57"E, 2257 m, 6.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 1 ex.: Goltappeh to Kabudarahang, 5 km W Kabudarahang, 35��13'20"N 48��25'57"E, 1907 m, 6.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 3 exs.: Hamedan to Malayer (or Jowkar) road, 7 km N Jowkar, 34��32'09"N 48��45'11"E, 1807 m, 10.VI.2006, Moghaddam, Nasserzadeh, Hajesmaelian; 22 exs.: Hamedan to Razan road, 2 km S Razan, Khamigan village, 35��21'10"N 49��01'24"E, 1844 m, 8.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 53 ex.: Kabudarahang, 35��14'07"N 48��35'44"E, 1755 m, 6.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 79 exs.: Kabudarahang to Goltappe road, 5 km W Kuhin, 35��11'52"N 48��16'33"E, 2335 m, 6.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 5 exs.: Malayer to Nahavand, 2 km E Kortilabad, 34��18'15"N 48��40'42"E, 1710 m, 31.V.2007, leg. Nasserzadeh, Zahiri, Montreuil; 15 exs.: Nahavand to Firuzan road, 5 km E Firuzan, Kahriz Salidin, 34��19'41"N 48��10'44"E, 1540 m, 1.VI.2007, leg. Nasserzadeh, Zahiri, Montreuil; 29 exs.: Razan to Damagh road, 15 km W Razan, 35��25'40"N 48��52'58"E, 1944 m, 8.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 52 exs.: S Hamedan, Oshtoran to Karzan road, 1 km NW Karzan, 34��36'24"N 48��19'43"E, 1910 m, 2.VI.2007, leg. Nasserzadeh, Zahiri, Montreuil. Ilam Province: 1 ex.: Chovar, Tang-e Dalab, 33��42'35"N 46��17'22"E, 1200 m, 15.V.1997, leg. Pazuki / Borumand; 11 exs.: Dehloran, Abgarm, 32��43'12"N 47��17'49"E, 290 m, 21.IV.2007, leg. Nasserzadeh; 22 exs.: Malekshahi to Ilam road, 12 km NW Malekshahi, 33��31'29"N 46��29'21"E, 1469 m, 7.V.2008, leg. Nas. / Nem. / Montreuil; 2 ex.: Sarableh to Zarneh road, 15 km NW Sarableh, Soltangholi-e Sofla village, 33��50'55"N 46��27'43"E, 1176 m, 9.V.2008, leg. Nas. / Nem. / Montreuil. Kermanshah Province: 3 exs.: Kerend to Sarpol-e Zahab road, 35 km W Kerend, Dizeh, 34��24'34"N 46��02'23"E, 1170 m, 24.V.2007, leg. Nasserzadeh, Zahiri, Montreuil; 4 exs.: Paveh, Shamshir to Bayangan road, 4 km W Shamshir, 34��58'54"N 46��23'47"E, 1754 m, 3.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 13 exs.: Kangavar to Hamedan road, 80 km SE Hamedan, 34��27'59"N 47��55'49"E, 1474 m, 4.VI.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian; 5 exs.: Songhor to Kermanshah road, 5 km S Myanrahan, 34��33'36"N 47��25'01"E, 1365 m, 29.V.2006, leg. Moghaddam, Nasserzadeh, Hajesmaelian. Climate types. Semi-arid, cool to cold winter and warm to very warm summer., Published as part of Nasserzadeh, Hiva, Platia, Giuseppe, Parsi, Farzaneh & Serri, Sayeh, 2018, Three new species, Elathous zagrosensis sp. n., Zorochros dilatatus sp. n., and Dicronychus iranicus sp. n., and noteworthy records of click beetles from western Iran (Coleoptera: Elateridae), pp. 517-536 in Zootaxa 4394 (4) on page 533, DOI: 10.11646/zootaxa.4394.4.3, http://zenodo.org/record/1199964

Cardiophorus bellus * Platia & Gudenzi, 2000 (Map 3) Material examined (HMIM). Kermanshah Province: 1 ex.: Javanrood to Tazehabad road, 13 km W Javanrood, Aliabad, 34��47'57"N 46��23'54"E, 1680 m, 27.V.2007, leg. Nasserzadeh, Zahiri, Montreuil. Remarks. distribution in Turkey (Cate, 2007). New species for Iran. Climate types. Semi-arid, cool winter and warm summer., Published as part of Nasserzadeh, Hiva, Platia, Giuseppe, Parsi, Farzaneh & Serri, Sayeh, 2018, Three new species, Elathous zagrosensis sp. n., Zorochros dilatatus sp. n., and Dicronychus iranicus sp. n., and noteworthy records of click beetles from western Iran (Coleoptera: Elateridae), pp. 517-536 in Zootaxa 4394 (4) on page 533, DOI: 10.11646/zootaxa.4394.4.3, http://zenodo.org/record/1199964

To date, all known or suspected pheromones of click beetles (Coleoptera: Elateridae) have been identified solely from species native to Europe and Asia; reports of identifications from North American species dating from the 1970s have since proven to be incorrect. While conducting bioassays of pheromones of a longhorned beetle (Coleoptera: Cerambycidae), we serendipitously discovered that males of Cardiophorus tenebrosus L. and Cardiophorus edwardsi Horn were specifically attracted to the cerambycid pheromone fuscumol acetate, (E)-6,10-dimethylundeca-5,9-dien-2-yl acetate, suggesting that this compound might also be a sex pheromone for the two Cardiophorus species. Further field bioassays and electrophysiological assays with the enantiomers of fuscumol acetate determined that males were specifically attracted by the (R)-enantiomer. However, subsequent analyses of extracts of volatiles from female C. tenebrosus and C. edwardsi showed that the females actually produced a different compound, which was identified as (3R,6E)-3,7,11-trimethyl-6,10-dodecadienoic acid methyl ester (methyl (3R,6E)-2,3-dihydrofarnesoate). In field trials, both the racemate and the (R)-enantiomer of the pheromone attracted similar numbers of male beetles, suggesting that the (S)-enantiomer was not interfering with responses to the insect-produced (R)-enantiomer. This report constitutes the first conclusive identification of sex pheromones for any North American click beetle species. Possible reasons for the strong and specific attraction of males to fuscumol acetate, which is markedly different in structure to the actual pheromone, are discussed.

Cardiophorus goezei Sanchez-Ruiz, 1996 Cardiophorus ruflpes (Goeze, 1777). Distribution in Iran. Mazandaran (Mardjanian et al. 2008), Iran (no locality cited) (Cate 2007). General distribution. ? Algeria,? Armenia, Austria,? Azerbaijan, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic,? Egypt, France,? Georgia, Germany, Hungary,? Iran, Italy,? Kazakhstan, Latvia, Macedonia, Moldavia,? Morocco, Poland, Portugal, Romania,? Russia, Slovakia, Spain, Sweden, Switzerland,? Tunisia,? Turkey,? Turkmenistan (Cate 2007). Comments. Presence of this species in Iran is doubtful and needs confirmation. Additionally, the presence in the countries with question mark is dubious in opinion of G.P., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Sanchez-Ruiz, A. (1996) Catalogo bibliografico de las especies de la familia Elateridae (Coleoptera) de la Peninsula Iberica e Islas Baleares. Documentos Fauna Iberica, 2. In: Ramos, M. A. (Ed.), Museo Nacional de Ciencias Naturales, Consejo Superior de Investigaciones Cientificas. Madrid, 265 pp.","Mardjanian, M. A., Barimani, H. V. & Kalashian, M. Yu. (2008) Materials to Fauna of click-beetles (Coleoptera, Elateridae) of Iran. Biological Journal of Armenia, 3 (60), 68 - 70.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus pruinosus Buysson, 1902 Distribution in Iran. "Perse" (no locality cited) (Buysson 1902), Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). General distribution. Endemic to Iran (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Buysson, H. du. (1902) Remarques sur quelques Elaterides du Genre Cardiophorus descriptions d'especes nouvelles. Annales de la Societe Entomolgique de France, 71, 422 - 439.","Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus sacratus Erichson, 1840 Distribution in Iran. Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). New materials. Semnan (9 km E Shamirzad) (Platia coll.). General distribution. Armenia, Azerbaijan, Cyprus, Egypt, Georgia, Iran, Iraq, Israel, Jordan, Lebanon, Russia, Syria, Turkey, Turkmenistan (Cate 2007), Uzbekistan (Platia 2010)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus carnosus Platia & Gudenzi, 2002 Distribution in Iran. Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). New materials. West Azarbaijan (Pass 25 km NNW Orumiye) (Platia coll.). General distribution. Iran, Iraq, Turkey (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus persianus Cate, Platia & Schimmel, 2002 Distribution in Iran. Sistan & Baluchestan (Kuh-e Taftan, Tamandan) (Cate et al. 2002), Iran (no locality cited) (Cate 2007; Platia 2013 a). General distribution. Endemic to Iran (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus magnanii Platia, Furlan & Gudenzi, 2002 Distribution in Iran. Fars (above Qalat vill.) (Platia 2013 a), Kordestan (13 km S Saqqez) (Platia et al. 2002), Iran (no locality cited) (Cate 2007). New materials. Qazvin (10 km NE of Ziaran), West Azarbaijan (26 km W Mahabad, Types; Ghasemlo S Orumiyeh), Zanjan (Zangan-Passo N Sorkheh Disaj) (Platia coll.). General distribution. Endemic to Iran (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Platia, G., Furlan, L. & Gudenzi, I. (2002) Descrizione di sei nuove specie di Elateridi dell'Iran (Insecta, Coleoptera, Elateridae). IL Naturalista Valtellinese - Atti del Museo civico di Storia Naturale di Morbegno, 13, 65 - 77.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus picinus Platia & Gudenzi, 2002 Distribution in Iran. Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). General distribution. Iran, Turkey (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus vestigialis Erichson, 1840 Distribution in Iran. Golestan (Bagheri et al. 2004 a), Kerman (Cate et al. 2002), Mazandaran (Cate et al. 2002; Platia 2013 a), Tehran (Platia 2013 a), Iran (no locality cited) (Jakobson 1913; Platia & Gudenzi 2000 a; Khanjani 2006; Cate 2007; Zapata & S��nchez-Ruiz 2014). New materials. Ardabil (Sareyn), East Azarbaijan (Passo N Khalkhal), Isfahan, Kordestan, Mazandaran, Tehran, Semnan (9 km E Shahmirzad), West Azarbaijan (Platia coll.). General distribution. Albania, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Cyprus, Czech Republic, Egypt, Finland, France, Georgia, Germany, Great Britain, Greece, Hungary, Iran, Italy, Kazakhstan, Moldavia, Mongolia, Netherlands, Norway, Poland, Portugal, Romania, Russia, Serbia- Montenegro, Slovakia, Slovenia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine (Cate 2007), Lebanon, Syria, Jordan (Platia 2010)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 249, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Bagheri, M. R., Rajabi, Gh. R., Ardebili, Zh., Haghshenas, A. R., Hosseini Nia, A. & Poorghaz, A. (2004 a) Faunistic study of click beetles (Col.: Elateridae) in Iran. Proceedings of the 16 th Iranian Plant Protection Congress, pp. 129.","Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Jakobson, G. G. (1913) Beetles of Russia and Western Europe. St. Petersburg, 1024 pp.","Platia, G. & Gudenzi, I. (2000 a) Revisione dei Cardiophorus della Turchia. I. Le specie bicolori. Lambillionea, 100 (4), 601 - 612.","Khanjani, M. (2006) Vegetable pests in Iran. Bu-Ali Sina University, No. 205, 467 pp.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Zapata, J. L. & Sanchez-Ruiz, A. (2014) Nuevas aportaciones al catalogo de la familia Elateridae (Coleoptera) en la Peninsula Iberica e Islas Baleares, II. Arquivos Entomoloxicos, 10, 129 - 166."]}

Cardiophorus megathorax (Faldermann, 1835) Distribution in Iran. East Azarbaijan, Khuzestan, West Azarbaijan (Cate et al. 2002), Iran (no locality cited) (Mardjanian 1987; Agaev 1988; Platia & Gudenzi 2000 a; Cate 2007; Platia 2013 a). General distribution. Armenia, Azerbaijan, Georgia, Greece, Iran, Israel, Jordan, Lebanon, Syria, Turkey (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Mardjanian, M. A. (1987) Fauna Armyanskoi SSR. Nasekomye Zhestkokryle. Shchelkuny (Elateridae). Izdatelstvo Akademii Nauk Armanskoi SSR. Erevan, 204 pp.","Agaev, B. I. (1988) Zoogeograficheskaya analiz fauny zhelkov shchelkunov (Coleoptera, Elateridae) vostochnogo Zakavkaz'ya. Zoologicheskii Zhurnal, 67 (4), 539 - 547.","Platia, G. & Gudenzi, I. (2000 a) Revisione dei Cardiophorus della Turchia. I. Le specie bicolori. Lambillionea, 100 (4), 601 - 612.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus hauseri Schwarz, 1900 Distribution in Iran. Hormozgan (Bandar-Abbas) (Cate et al. 2002), Iran (no locality cited) (Dolin & Atamuradov 1994; Cate 2007; Platia 2013 a). General distribution. Afghanistan, China, Iran, Kyrgyzstan, Kazakhstan, Turkmenistan Uzbekistan (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Dolin, V. G. & Atamuradov, Kh. I. (1994) Zhuki-shchelkuny (Elateridae) Turkmenistana. - Click-beetles (Elateridae) of Turkmenistan, Institut zoologii NAN Ukrainy, Kiev, 178 pp.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus gramineus (Scopoli, 1763) Distribution in Iran. Kerman (Jiroft) (Samin et al. 2016). General distribution. Armenia, Austria, Azerbaijan, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czzech Republic, France, Germany, Georgia, Hungary, Italy, Latvia, Luxemburg, Moldavia, Netherlands, Palestine, Poland, Portugal, Romania, Russia, Serbia-Montenegro, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Samin, N., Hava, J. & Kubisz, D. (2016) A contribution to the knowledge of some families of Coleoptera (Insecta) from Iran. Arquivos Entomoloxicos, 15, 29 - 38.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus impressiventris Schwarz, 1900 Distribution in Iran. Mazandaran (N��meth & Platia 2014), Iran (no locality cited) (Nemeth & Platia 2014). General distribution. Iran (Nemeth & Platia 2014), Turkey (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Nemeth, T. & Platia, G. (2014) On some Palaearctic click beetles deposited in the Hungarian Natural History Museum, 2 (Coleoptera: Elateridae). Zootaxa, 3841 (4), 451 - 490. http: // dx. doi. org / 10.11646 / zootaxa. 3841.4.1","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus bioculatus Platia, 2010 Distribution in Iran. Kuhgiloyeh & Boyerahmad (Yasuj, 10 km SE Sepidar) (Platia 2010), Iran (no locality cited) (Platia 2013 a). General distribution. Endemic to Iran (Platia 2013 a)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 245, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus bogatschevi Dolin, 1985 Distribution in Iran. Ardabil (Platia 2013 a), Kermanshah, northern Khorasan, Razavi Khorasan (Cate et al. 2002), Iran (no locality cited) (Cate 2007). New materials. Golestan (Platia coll.). General distribution. Azerbaijan, Iran, Kazakhstan, Turkmenistan (Cate 2007), Uzbekistan (Platia 2011), Russia (South European Territory) (Prosvirov 2013)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on pages 245-246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Prosvirov, A. S. (2013) To the knowledge of the fauna of click-beetles (Coleoptera: Elateridae) of south of European Russia and the Caucasus. Caucasian Entomological Bulletin, 9 (2), 253 - 259."]}

Cardiophorus khnzoriani Mardjanian & Barimani, 2011 Distribution in Iran. Mazandaran (Poshte-Koh) (Mardjanian & Barimani 2011), Iran (no locality cited) (Platia 2013 a). General distribution. Endemic to Iran (Platia 2013 a)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Mardjanian, M. A. & Barimani, H. V. (2011) New species of click-beetles of genus Cardiophorus Eschscholtz, 1829 (Coleoptera, Elateridae, Cardiophorinae) from Mazandaran (Iran). National Academy of Sciences of Armenia Reports, 111 (4 Entomology), 396 - 403.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus rotundicollis Frivaldszky, 1845 Distribution in Iran. East Azarbaijan (Ghahari et al. 2012), Isfahan (Farhangian et al. 2013). General distribution. Greece, Turkey (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Ghahari, H., Makhan, D. & Hawkeswood, T. J. (2012) A faunistic survey on some families of Coleoptera from cotton fields of northern Iran. Calodema, 203, 1 - 7.","Farhangian, S., Sakenin Chelav, H. & Monem, R. (2013) A faunistic study on Elateridae (Coleoptera: Elateroidea) from Isfahan, Iran. Calodema, 264, 1 - 3.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus trimaculatus Schwarz, 1894 Distribution in Iran. " Persien " (no locality cited) (Schwarz 1894), Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). General distribution. Endemic to Iran (Cate 2007). Cardiophorus varandi Mardjanian & Barimani, 2011 Distribution in Iran. Mazandaran (Sari: Alikola) (Mardjanian & Barimani 2011), Iran (no locality cited) (Platia 2013 a). General distribution. Endemic to Iran (Platia 2013 a)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on pages 248-249, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Schwarz, O. (1894) Neue Elateriden. Deutsche Entomologische Zeitschrift, 38 (1), 145 - 150.","Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Mardjanian, M. A. & Barimani, H. V. (2011) New species of click-beetles of genus Cardiophorus Eschscholtz, 1829 (Coleoptera, Elateridae, Cardiophorinae) from Mazandaran (Iran). National Academy of Sciences of Armenia Reports, 111 (4 Entomology), 396 - 403."]}

Cardiophorus kryzhanovskyi Dolin & Chantladze, 1980 Distribution in Iran. Golestan (Platia 2008), Iran (no locality cited) (Platia 2013 a). General distribution. Armenia, Azerbaijan, Georgia, Russia (Cate 2007), Iran (Platia 2008)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus syriacoides Platia & Cate, 2001 Distribution in Iran. Golestan (Platia & Cate 2001; Cate 2007; Cate et al. 2002). General distribution. Armenia, Azerbaijan, Iran, Syria, Tajikistan, Turkmenistan (Cate 2007).

Cardiophorus gianassoi Platia, 2010 Distribution in Iran. Kuhguiloyeh & Boyerahmad (Dena Reserve, Sisakht env.) (Platia 2010). General distribution. Endemic to Iran (Platia 2010)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 246, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211

Cardiophorus maculatus Cate, Platia & Schimmel, 2002 Distribution in Iran. Sistan & Baluchestan (26 km NW of Deh Parbid), Southern Khorasan (13 km N Birjand) (Cate et al. 2002), Iran (no locality cited) (Cate 2007; Platia 2013 a). General distribution. Endemic to Iran (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus terminasiani Mardjanian & Barimani, 2011 Distribution in Iran. Mazandaran (Poshte-Koh) (Mardjanian & Barimani 2011), Iran (no locality) (Platia 2013 a). General distribution. Endemic to Iran (Platia 2013 a)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Mardjanian, M. A. & Barimani, H. V. (2011) New species of click-beetles of genus Cardiophorus Eschscholtz, 1829 (Coleoptera, Elateridae, Cardiophorinae) from Mazandaran (Iran). National Academy of Sciences of Armenia Reports, 111 (4 Entomology), 396 - 403.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus klimenkoi Platia, 2008 Distribution in Iran. Kuhgiloyeh & Boyerahmad (Yasuj Area, 10 km SE Sepidar) (Platia 2008), Iran (no locality cited) (Platia 2013 a). New materials. Fars (Kuhha-ye-Zagros; Sepidan), Isfahan (Pass ca. 55 km SW Semirom) (Platia coll.). General distribution. Endemic to Iran (Platia 2013 a)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus miniaticollis Cand��ze, 1860 Distribution in Iran. Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a). General distribution. Armenia, Greece, Iran, Lebanon, Syria, Turkey, " Palestine " (Cate 2007), Jordan (Platia 2010)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 247, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402."]}

Cardiophorus syriacus (Linnaeus, 1758) Distribution in Iran. Golestan (Cate 2007). General distribution. Armenia, Azerbaijan, Georgia, Iran, Syria, Turkey, Turkmenistan (Cate 2007)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209."]}

Cardiophorus ruficruris (Brull��, 1832) Distribution in Iran. Iran (no locality cited) (Cate et al. 2002; Cate 2007; Platia 2013 a; N��meth & Platia 2014). General distribution. Armenia, Cyprus, Greece, Iran, Italy, Spain, Morocco, Turkey (Cate 2007), Macedonia (Nemeth & Platia 2014)., Published as part of Platia, Giuseppe & Ghahari, Hassan, 2016, An annotated checklist of click-beetles (Coleoptera, Elateridae) from Iran, pp. 239-275 in Zootaxa 4137 (2) on page 248, DOI: 10.11646/zootaxa.4137.2.6, http://zenodo.org/record/255211, {"references":["Cate, P. C., Platia, G. & Schimmel, R. (2002) New species and records of click beetles (Coleoptera: Elateridae) from Iran, with a checklist of known species. Folia Heyrovskyana, 10 (1), 25 - 68.","Cate, P. G. (2007) Family Elateridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209.","Platia, G. (2013 a) Click-beetles collected in Iran by the expeditions of the Naturkundemuseum Erfurt with description of two new species (Insecta: Coleoptera: Elateridae). Vernate, 32, 391 - 402.","Nemeth, T. & Platia, G. (2014) On some Palaearctic click beetles deposited in the Hungarian Natural History Museum, 2 (Coleoptera: Elateridae). Zootaxa, 3841 (4), 451 - 490. http: // dx. doi. org / 10.11646 / zootaxa. 3841.4.1"]}