Search

Henosepilachna boisduvali (Mulsant) Epilachna boisduvali Mulsant, 1850: 765. Epilachna montrouzieri Fauvel, 1862: 174 ���synonymized by Korschefsky 1931: 32. Henosepilachna boisduvali: Li & Cook 1961: 47. This ladybug is a widely distributed species in South East Asia (from Taiwan, Vietnam, Philippines, Ryukyu Islands, Indonesia, West Timor), but also in Australia, Vanuatu, Fiji Islands and Samoa. In New Caledonia, it is recorded from both coasts of Grande Terre, but also on Loyalty Islands and Isle of Pines (Chazeau, pers. obs.). It may be an accidental introduction. This species is considered as a pest on several plants, especially on Cucurbitaceae (Abe & Matsuda 2000)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 314, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Fauvel, A. (1862) Coleopteres de la Nouvelle-Caledonie, recueillis par M. E. Deplanche, chirurgien de la marine imperiale (1858 - 59 - 60). Bulletin de la Societe Linneenne de Normandie, 7, 120 - 185.","Korschefsky, R. (1931) Pars 118: Coccinellidae I. In: Junk, W. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 224.","Li, C. S. & Cook, E. F. (1961) The Epilachninae of Taiwan (Col.: Coccinellidae). Pacific Insects, Honolulu, 3, 31 - 91.","Abe, M. & Matsuda, K. (2000) Feeding responses of four phytophagous lady beetle species (Coleoptera: Coccinellidae) to cucurbitacins and alkaloids. Applied Entomology and Zoology, 35, 257 - 264."]}

Orcus artensis Crotch Orcus artensis Crotch, 1874: 188. This endemic species is known from the type specimen and two specimens from Art Island, in the Belep Archipelago (Laczynski & Tomaszewska 2009). It is not a common species, but we report here some recent collections in the southern part of the main island ���Grande Terre��� (La Ouenghi). According to Laczynski & Tomaszewska (2009), the coloration of this species is unique within the genus and separates easily O. artensis from all its congeners. Its biology is unknown, but this Orcus appears to be a predator of hemipterous Sternorrhyncha in forested areas (Chazeau, pers. obs.). Material examined. NEW CALEDONIA: Pont de la Ouenghi, 22.xi. 1987 (J. Chazeau rec.) (ONNC). Literature records. NEW CALEDONIA: Coll. R.I.Sc.N.B., Nouvelle Caledonie, Ile d���Art, rec. Montrousier, ex coll. Fauvel, Coll. et det. A. Fauvel, Orcus artensis Crotch (1: ISNB); same and R.I.Sc.N.B. 17.479 (2: ISNB) (Laczynski & Tomaszewska 2009)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 303, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Laczynski, P. & Tomaszewska, W. (2009) Revision of the genus Orcus Mulsant (Coleoptera: Coccinellidae: Chilocorini). Annales Zoologici, 59, 585 - 611."]}

Apolinus lividigaster (Mulsant) Platyomus lividigaster Mulsant, 1853: 286. Scymnodes (Apolinus) lividigaster: Pope & Lawrence 1990: 244. Apolinus lividigaster: Slipinski 2007: 99. This species was originally described from Australia; it is also recorded from New Zealand and Hawaii. In New Caledonia, it was not recorded by Bielawski (1972) in his review of New Caledonian Scymnini. It has only been collected in Noum��a and Yat�� regions (southern Grande Terre). It is predominantly aphidiphagous, but is also able to use pollen when preys are scarce (Lundgren 2009). Based on this information, we consider it to be a recent arrival because of the lack of autochthonous aphids, whether by natural spread or accidental introduction. Material examined. NEW CALEDONIA: Noum��a, Anse Vata, ix. 1976 (A. Delobel rec.) (ONNC); Noum��a, Ouen Toro, 24.vi. 1978 (A. Delobel rec.) (ONNC); St Gabriel (Yat��), 0-200m, xi. 1986 (N.L.H. Krauss rec.) (ONNC); Farino, "Tend��a", 21.61928 ��S / 165.70907 ��E / 403m, 29.iv. 2004 (S. Caz��res rec.) (CXMNC); Ouv��a, Hynthot, 30.vi. 2008 (C. Mille rec.) (CXMNC); Thio, tribu de Koua, COL /ins/ 28 -14, 18.iii. 2013 (R-M. M'Boueri rec.) (CXMNC); Ouv��a, COL /ins/ 87 -14, 14.iii. 2014 (C. Mille rec.) (CXMNC); Bourail, Gouaro, 21.58113 ��S / 165.42045 ��E / 68m, ii. 2001 (S. Caz��res rec.) (CXMNC); Thio, Dothio, 21 �� 35.305 'S / 165 ��09.084'E / 18m, 04.v. 2006 (S. Caz��res rec.) (CXMNC); Noum��a, 15.xii. 1981 (R. Catala rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 317-318, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Mulsant, E. (1853) Supplement a la monographie des Coleopteres Trimeres Securipalpes. Annales de la Societe linneenne de Lyon, 1, 129 - 333.","Pope, R. D. & Lawrence, J. F. (1990) A review of Scymnodes Blackburn, with the description of a new Australian species and its larva (Coleoptera: Coccinellidae). Systematic Entomology, 15, 241 - 252.","Slipinski, A. (2007) Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their Biology and Classification. Australian Biological Resources Study, Canberra, xviii + 286 pp.","Bielawski, R. (1972) The Epilachninae (Coleoptera, Coccinellidae) of New Caledonia. Bulletin de l'Academie polonaise des sciences. Serie des sciences biologiques, 20, 557 - 561.","Lundgren, J. G. (2009) Nutritional aspects of non-prey foods in the life histories of predaceous Coccinellidae. Biological Control, 51, 294 - 305."]}

Harmonia octomaculata (Fabricius) Coccinella octomaculata Fabricius, 1781: 97 (not Thunberg, 1781: 131). Coccinella arcuata Fabricius 1787: 55. Harmonia arcuata: Mulsant 1850: 77. This species is widespread in Asia and also from Australia, Papua New Guinea, Vanuatu, Solomon Islands, Fiji Islands, Micronesia (Marianna, Marshall, Kiribati), Cook Islands and French Polynesia. It is also known from South Africa. According to this geographic range, we consider that this species was moved by anthropogenic activities. This species was recorded early by Montrouzier (1861) from Ile Art and Balade. In New Caledonia, this species is widespread on both coasts, valleys from the Central Range including also the remote Chesterfield Islands but not recorded from Loyalty. This is a polyphagous predator of Hemiptera (Cicadellidae, Aphididae [Rhopalosiphum maidis (Fitch)], Psyllidae [Heteropsylla cubana]). Material examined. NEW CALEDONIA: Bourail, 11.iv. 1933 (J. Chazeau rec.) (ONNC); Houa��lou, 15.iii. 1982 (J. Chazeau rec.) (ONNC); Iles Chesterfield, I. Loop, 2.x. 1977 (J. Kusser rec.) (ONNC); Iles Loyaut��, Mar�� (ONNC); La Foa, 2.x. 1979 (B. Sall��e rec.) (ONNC); Mt Dore, La Coul��e, 4.xii. 1979 (J. Gutierrez rec.) (ONNC); Mts Koghi, 12.x. 1958 (Y. Shibata rec.) (ONNC); Noum��a, i-iii. 1955 (ONNC); Noum��a, v. 1959 (ONNC); Noum��a, 11.iv. 1988 (J. Chazeau rec.) (ONNC); Pa��ta, 21.v. 1979 (B. Sall��e rec.) (ONNC); Pied du Col d'Amos, versant Est, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Poindimi��, 30.x. 1988 (J. Chazeau rec.) (ONNC); Port- Laguerre, Station Agricole, 3.iii. 1977 (J. Chazeau rec.) (ONNC); Port-Laguerre, Station Agricole, 3.iii. 1977 (J. Chazeau rec.) (ONNC); Port-Laguerre, ii-vi. 1977 (A. Delobel rec.) (ONNC); Port-Laguerre, 19.vii. 1986 (R. Toutain rec.) (ONNC); Pouembout, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Sud de Kaala-Gomen (6 km), 31.iii. 1986 (J. Chazeau rec.) (ONNC); Thio Ourou��, 5.i. 1982 (J. Chazeau rec.) (ONNC); Ile longue, Chesterfield islands, xi. 2012 (E. Bourguet et H. Jourdan rec.) (ONNC); La Foa, Pocquereux, 21.73205 ��S / 165.89936 ��E / 32m, 09.iii. 2007 (N. Degallier rec.) (CXMNC); La Foa, M��ar��, 16.i. 2007 (R-M. M'Boueri rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 07.i. 2002 (C. Mille rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 15.xii. 1999 (J-P. Kataoui rec.) (CXMNC); La Foa, Pocquereux, 21.89630 ��S / 165.81789 ��E / 28m, 20.viii. 2012 (R-M. M'Boueri rec.) (CXMNC); Pa��ta, Na��a, COL /ins/ 35 -14, 24.i. 2014 (R-M. M'Boueri & C. Mille rec.) (CXMNC); Koumac, Ti��baghi, 20 �� 28.074 'S / 164 �� 12.035 'E / 448m, 09.iii. 2013 (C. Mille rec.) (CXMNC); Boulouparis, Ouenghi, 21 �� 53 ' 51.61 "S / 166 ��06' 49.62 ������E / 7m, 10.ii. 2012 (C. Mille rec.) (CXMNC); Mar�� chez M. N. War��kaicane, 07.xi. 2000 (D. Paulaud rec.) (CXMNC); Boulouparis, Ouenghi, 21 �� 53.821 ���S / 166 ��06.656���E / 7m, 22.viii. 2012 (S. Caz��res & R-M. M'Boueri rec.) (CXMNC); Boulouparis, Ouenghi, 21 �� 53.801 ���S / 166 ��06.662���E / 7m, 26.ix. 2012 (S. Caz��res, R-M. M'Boueri, J. Brinon rec.) (CXMNC); Oua Tom, 4.iv. 1995 (M. Boulard rec.) (MNHN); Noum��a, 1904 (R. Bavay rec.) (MNHN); Magenta, 21.i. 1951 (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 307, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fabricius, J. C. (1781) Species Insectorum; Exhibentes Eorum Differentias Specificas, Synonyma, Auctorum, Loca Natalia, Metamorphosin Ediectis Observationibus, Descriptionibus. Vol. 1. Carol. Ernest. Bohnii, Hamburgi & Kilonii, VIII + 552 pp.","Thunberg, C. P. (1781) Dissertatio Entomologica Novas Insectorum Species, Sistens, Cujus Partem Primam. Joh. Edman, Upsaliae, 28 pp.","Fabricius, J. C. (1787) Mantissa Insectorum: Sistens Eorvm Species Nvper Detectas. Adiectis Synonymis, Observationibvs, Descriptionibvs, Emendationibvs. Vol 1. Proft, Hafniae, 382 pp.","Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306."]}

Scymnus micros Fauvel Scymnus micros Fauvel, 1903: 324 (pars). Endemic to New Caledonia (Noum��a). In his review, Bielawski (1973) assigned a lectotype for this species, which represents the only known specimen in collections (Col d���Amieu, dense rain forest). In Fauvel���s collection, this species was partially confused with Stethorus vagans (Blackburn) (Chazeau 1979). At the genus level, its position remains uncertain. Male genitalia and the heavily convex specimen habitus suggest a Sticholotidini. Its biology remains unknown., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 319, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409.","Chazeau, J. (1979) Mise au point sur le genre Stethorus en oceanie et description de deux especes nouvelles de Melanesie [Col., Coccinellidae]. Entomophaga, 24, 295 - 303."]}

Coccinella boletifera Fauvel Coccinella boletifera Fauvel, 1903: 325. Coccinella bolitifera: Korschefsky, 1931: 508 (misspelling). This species has been quoted as an endemic one (Fauvel 1903; Korschefsky 1931, as C. bolitifera), but it is only known from the type specimen, which has an unknown locality. This Coccinella seems to be a color variation of Coccinella transversalis Fabricius and therefore may be considered a synonym., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 321, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Korschefsky, R. (1931) Pars 118: Coccinellidae I. In: Junk, W. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 224."]}

Micraspis frenata (Erichson) Coccinella frenata Erichson, 1842: 239. Verania frenata: Mulsant 1850: 362. Coccinella bicruciata Montrouzier, 1861: 304 (pars)���synonymized by Fauvel 1903: 328. Alesia frenata: Masters 1888: 90. Micraspis frenata: Pope 1988: 679. This species was misidentified as Coelophora bicruciata by Montrouzier (1861). According to Fauvel (1903), who examined the type series of Coelophora bicruciata, it was a mixture of Coelophora inaequalis and Micraspis frenata specimens. This species is common in Australasia (Australia, Tasmania and Papua New Guinea). Iablokoff- Khnzorian (1982) treated the species as a junior synonym of Micraspis lineata (Thunberg, 1781). This Micraspis is very common over New Caledonia, mostly from open areas on the west coast, but also the Central Range, Isle of Pines, and Belep Islands (Art Island). Its trophic preferences are wide, being mostly a hemipteran predator (aphids, cicadellids, psyllids), but it is able to eat pollen or extrafloral nectaries when prey are scarce (Hawkeswood & Turner 2002; Lundgren 2009). It has been widely recorded in close association with the invasive psyllid, Heteropsylla cubana, in New Caledonia (Gatimel 2005). This species has not yet been widely promoted for biocontrol, despite high potential in agroecosystems (Chenaux et al. 2011). Material examined. NEW CALEDONIA: Bond��, 6.vi. 1975 (J. Gutierrez rec.) (ONNC); Bourail, N��m��ara, 10.x. 1958 (Y. Shibata rec.) (ONNC); Bourail, bords de la N��ra, 23.vii. 1988 (J. Chazeau rec.) (ONNC); Bourail, bords de la N��ra, 6.viii. 1992 (J. Chazeau rec.) (ONNC); Canala, 20.vi. 1989 (J. Chazeau, L. Bonnet de Larbogne, E. Bouy�� rec.) (ONNC); Col de la Pirogue, 16.i. 1987 (J. Chazeau rec.) (ONNC); Col d'Amieu, 23.vii. 1977 (J. Chazeau rec.) (ONNC); Col d'Amieu, 11.iii. 1982 (J. Chazeau rec.) (ONNC); Col de Cr��vecoeur, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Col de Koh, From La Foa to Canala 20.vi. 1989 (J. Chazeau, L. Bonnet de Larbogne, E. Bouy�� rec.) (ONNC); Col de Mu��o, 17.viii. 1992 (J. Chazeau rec.) (ONNC); Col de Nassirah, versant Thio, 27.vi. 1989 (J. Chazeau, L. Bonnet de Larbogne, E. Bouy�� rec.) (ONNC); Col de T��mala, 16.iv. 1986 (J. Chazeau rec.) (ONNC); Entre Moindou et Bourail, RT 1 pk 148, 6.viii. 1992 (J. Chazeau rec.) (ONNC); Entre Port Laguerre et Tontouta, 22.xi. 1987 (J. Chazeau rec.) (ONNC); Environs de Port-Laguerre, 29.i. 1978 (A. Delobel rec.) (ONNC); Environs de Houa��lou (6 km ouest), 23.ix. 1988 (J. Chazeau rec.) (ONNC); Isle of Pines, Grottes d'Oumagne, 22.viii. 1988 (J. Chazeau rec.) (ONNC); Kaala-Gomen, 22.ix. 1986 (J. Chazeau rec.) (ONNC); La Foa, 2.x. 1979 (B. Sall��e rec.) (ONNC); Negropo, iii. 1955 (ONNC); Ouaco, 22.vii. 1977 (J. Gutierrez rec.) (ONNC); Pindai, 9.ix. 1992 (E. & M. Schlinger rec.) (ONNC); Pindai, 13.ix. 1992 (D. Webb rec.) (ONNC); Pont de la Ouenghi, RT1, 1.xii. 1981 (J. Chazeau rec.) (ONNC); Pont de la Rivi��re du Cap, RT1, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Port-Laguerre, ii-vi. 1977 (A. Delobel rec.) (ONNC); Port-Laguerre, Station Agricole, 28.i. 1977 (J. Chazeau rec.) (ONNC); Port-Laguerre, Station Agricole, 3.iii. 1977 (J. Chazeau rec.) (ONNC); Pouembout, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Pouembout sud, 17.viii. 1992 (J. Chazeau rec.) (ONNC); Pouembout, Franco Plage, 19.iii. 1982 (J. Chazeau rec.) (ONNC); Poya, 16.iv. 1986 (J. Chazeau rec.) (ONNC); Poya, 16.iv. 1986 (J. Chazeau rec.) (ONNC); Route de Fat��naou��, environs confluent Ouagatch, 19.x. 1988 (J. Chazeau rec.) (ONNC); RT 1, environs limite provinciale (7 km sud), 17.viii. 1992 (ONNC); Sarram��a, x. 1969 (P. Cochereau rec.) (ONNC); Timbia (Pa��ta), 25.xii. 1992 (J. Chazeau rec.) (ONNC); Tontouta, 20.v. 1979 (B. Sall��e rec.) (ONNC); Vall��e du Diahot, vii. 1955 (ONNC); Ile longue, Chesterfield Islands, xi. 2012 (E. Bourguet et H. Jourdan rec.) (ONNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 20.iv. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 05.vi. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 07.viii. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, COL /ins/ 213 - 14, 10.2006 (J. Sudre rec.) (CXMNC); Thio, tribu de Koua, COL /ins/ 28 -14, 18.iii. 2013 (R-M. M'Boueri rec.) (CXMNC); Yat��, Prony, COL /ins/ 67 -14, 2006 (H. Jourdan rec.) (CXMNC); Bourail, Gouaro, 21.58113 ��S / 165.42045 ��E / 68m, ii. 2001 (S. Caz��res rec.) (CXMNC); Bourail, Gouaro, 21.58113 ��S / 165.42045 ��E / 68m, 26.iii. 2001 (S.Caz��res rec.) (CXMNC); Bourail, Gouaro chez M. Houarau, 04.x. 2004 (S.Caz��res rec.) (CXMNC); Lifou, 27.vii. 2002 (C. Mille rec.) (CXMNC); Boulouparis, Ouenghi, 21 �� 53 ' 51.61 "S / 166 ��06' 49.62 ������E / 7m, 10.ii. 2012 (C. Mille rec.) (CXMNC); Noum��a, Ouen Toro, 24.x. 1979 (J. Chazeau rec.) (MNHN); Noum��a, 1904 (R. Bavay rec.) (MNHN); Ouen Toro, 4.iv. 1995 (M. Boulard rec.) (MNHN); Noum��a, 1929 (Risbec rec.) (MNHN); Sanatorium du Col de la Pirogue, 10.i. 1951 (MNHN); Canala, 1869 (Delacour rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 308, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Erichson, W. F. (1842) Beitrag zur Insecten-Fauna von Vandiemensland, mit besonderer Berucksichtigung der geographischen Verbreitung der Insecten. Archiv fur Naturgeschichte, 8 (1), 239 - 240.","Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Masters, G. (1888) Catalogue of the described Coleoptera of Australia. Part VII. Proceedings of the Linnean Society of New South Wales, 2, 13 - 94.","Pope, R. D. (1988) A revision of the Australian Coccinellidae (Coleoptera). Part 1. Subfamily Coccinellinae. Invertebrate Systematics, 2, 633 - 735.","Iablokoff-Khnzorian, S. M. (1982) Les coccinelles, Coleopteres Coccinellidae. Tribu Coccinellini des regions Palearctiques et Orientale. Boubee, Paris, 568 pp.","Thunberg, C. P. (1781) Dissertatio Entomologica Novas Insectorum Species, Sistens, Cujus Partem Primam. Joh. Edman, Upsaliae, 28 pp.","Hawkeswood, T. J. & Turner, J. R. (2002) Observations on the adults of Micraspis frenata (Erichson, 1842) (Coleoptera: Coccinellidae) feeding on the pollen of native and non-native grasses (Poaceae) in eastern New South Wales, Australia. Journal of the Entomological Research Society, 4, 21 - 29.","Lundgren, J. G. (2009) Nutritional aspects of non-prey foods in the life histories of predaceous Coccinellidae. Biological Control, 51, 294 - 305.","Gatimel, B. (2005) Inventaire critique et perspectives des introductions d'especes realisees a des fins de lutte biologique en Nouvelle-Caledonie: applications aux cas de la mineuse des agrumes (Phyllocnistis citrella Stainton) et du psylle du fauxmimosa (Heteropsylla cubana Crawford). Master of Science (MSc), Ecology & Biodiversity Management, Universite Montpellier 2, 30 pp.","Chenaux, B., Costamagna, A. C., Bianchi, F. J. & Schellhorn, N. A. (2011) Functional response of two common Australian predators, Dicranolaius bellulus (Guerin? Meneville) (Coleoptera: Melyridae) and Micraspis frenata (Erichson) (Coleoptera: Coccinellidae), attacking Aphis gossypii Glover (Hemiptera: Aphididae). Australian Journal of Entomology, 50, 453 - 459."]}

Coelophora inaequalis (Fabricius) Coccinella inaequalis Fabricius, 1775: 80. Coccinella circularis Thunberg, 1820: 365 ���synonymized by Pope 1987: 54. Lemnia desolata Mulsant, 1850: 387 ���synonymized by Houston 1979: 49. Coccinella bicruciata Montrouzier, 1861: 304 (pars)���synonymized by Fauvel 1903: 328. Verania artensis Montrouzier, 1861: 305 ���synonymized by Chazeau 1978: 59. Coelophora ripponi Crotch, 1874: 156 ���synonymized by Iablokoff-Khnzorian 1979: 61. Coelophora mastersi Blackburn, 1892: 238 ���synonymized by Houston 1979: 49. Coelophora inaequalis: Fauvel 1903: 327. At first, this species was misidentified by Montrouzier who described it under two endemic species: Coccinella bicruciata and Verania artensis. According to Fauvel (1903), who examined the type series, Coelophora bicruciata (Montrouzier) was a mixture of Coelophora inaequalis and Micraspis frenata (Erichson) specimens. Verania artensis was subsequently synonymized by Chazeau (1978). Coelophora inaequalis is widespread from both coasts to high altitude (> 1,000m) on Grande Terre but also on Loyalty Islands and Isle of Pines. This species is widely distributed across Asia and the Pacific region (Australia, Papua New Guinea, Vanuatu, Solomon Islands, Fiji Islands, Hawaii, Micronesia (Bonin, Marianna, Caroline, Wake, Marshall, Kiribati), Cook Islands, Samoa, Wallis and Futuna, French Polynesia); but also has been recorded from eastern North America. This is a polyphagous predator of hemipterous insects (Aphididae, Coccomorpha) from natural habitats to agricultural lands and disturbed areas. According to its geographic range and feeding biology, humans have probably spread this species during the first settlements. It is a well-known and largely promoted biocontrol agent against aphids (Wang & Tsai 2001). Material examined. NEW CALEDONIA: Bac de la Oua����me, 17.iv. 1986 (J. Chazeau rec.) (ONNC); Bourail, 10.iii. 1986 (J. Chazeau rec.) (ONNC); from La Foa to Canala, ii. 1907 (Le Rat rec.) (ONNC); Goro, 12.viii. 1985 (J. Chazeau rec.) (ONNC); Hiengh��ne, 18.xi. 1958 (M. Ogata rec.) (ONNC); Houa��lou, 7.iii. 1982 (J. Chazeau rec.) (ONNC); Isle of Pines, airport, 24.viii. 1988 (J. Chazeau rec.) (ONNC); Iles Loyaut��, Lifou; Iles Loyaut��, Mar��; Koumac, 10.iii. 1986 (J. Chazeau rec.) (ONNC); Koumac, 26.viii. 1986 (J. Chazeau rec.) (ONNC); Koumac, 27.viii. 1986 (J. Chazeau rec.) (ONNC); Mt Do, 1000 m, 4.xi. 1992 (J. Chazeau & D. Webb rec.) (ONNC); Noum��a, Anse Vata, 25.ii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Anse Vata, 10.iii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Anse Vata, 3.x. 1977 (J. Chazeau rec.) (ONNC); Noum��a, IFO, 4.i. 1964; Noum��a, IFO, 13.i. 1965; Noum��a, OuenToro, vii. 1966 (J. Fainika rec.) (ONNC); Noum��a, Baie des Citrons, 27.vi. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Trianon, 20.iv. 1964 (J. Gutierrez rec.) (ONNC); Noum��a, vi, vii, xi. 1955; Noum��a, ix. 1964; Pouembout, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Pindai, 6.xi. 1992 (D. Webb rec.) (ONNC); La Foa-Canala Road, i. 1958; Thio, 27.vi. 1983 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Thio Ourou��, 5.i. 1982 (J. Chazeau rec.) (ONNC); Ile longue, Chesterfield islands, xi. 2012 (E. Bourguet et H. Jourdan rec.) (ONNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 10.ix. 2001 (S. Caz��res rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 20.iv. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 16.xi. 1997 (CXMNC); Bourail, Gouaro, 21 �� 36 ���S / 165 �� 28 ���E / 68m, 04.iv. 2001 (S. Caz��res rec.) (CXMNC); Farino, Tend��a, 21.61928 ��S / 165.70907 ��E / 403m, 29.iv. 2004 (S. Caz��res rec.) (CXMNC); Houa��lou, N��oua, 13.iv. 2013 (C. Mille rec.) (CXMNC); Pouembout chez M. Soury-Lavergne, COL /ins/ 255 -14, 01.viii. 2014 (C. Mille rec.) (CXMNC); Poya, Beaupr�� chez M. Mezdorf, 21 �� 205 ���S/ 165 ��09���E, 22.vi. 2001 (R. Guiglion rec.) (CXMNC); Mar��, La Roche, 16.iv. 2004 (C. Mille rec.) (CXMNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 305, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fabricius, J. C. (1775) Systema Entomologiae Sistens Insectorvm Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibvs, Observationibvs. Vol 1. Kortius, Lipsiae, 832 pp.","Thunberg, C. P. (1820) Coleoptera Capensia, antennarum clava solida et perfoliata, collecta, recensita et descripta. Memoires de l'Academie Imperiale des Sciences de Saint Petersbourg, 7, 362 - 372.","Pope, R. D. (1987) The Coccinellidae (Coleoptera) described by CP Thunberg. Insect Systematics & Evolution, 18, 51 - 66.","Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Houston, K. J. (1979) Mosaic dominance in the inheritance of the colour patterns of Coelophora inaequalis (F.) (Coleoptera: Coccinellidae). Australian Journal of Entomology, 18, 45 - 51.","Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Chazeau, J. (1978) Les especes du genre Coelophora en Nouvelle Caledonie et aux Nouvelles Hebrides (Coleopteres, Coccinellidae). Cahiers ORSTOM. Serie Biologie: Entomologie Agricole, 13, 59 - 69.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Iablokoff-Khnzorian, S. M. (1979) Genera der palaarktischen Coccinellini (Coleoptera, Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75.","Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia, 15, 207 - 261.","Wang, J. J. & Tsai, J. H. (2001) Development and functional response of Coelophora inaequalis (Coleoptera: Coccinellidae) feeding on brown citrus aphid, Toxoptera citricida (Homoptera: Aphididae). Agricultural and Forest Entomology, 3, 65 - 69."]}

Scymnus (Caledonus) angusticollis Fauvel Scymnus angusticollis Fauvel, 1903: 323. Scymnus (Caledonus) angusticollis: Bielawski 1973: 397. This species was redescribed by Bielawski (1973), who established an endemic subgenus Caledonus. It is known only from the west coast region (around Noum��a and Dumb��a���Chazeau pers. obs, and Col de Tonghou��). No information is available on its biology. Literature records. NEW CALEDONIA: Tonghou��, i (A. Fauvel rec.) (coll. A. Fauvel in IRSNB) (Bielawski 1973)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 319, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Rhyzobius pulchellus (Montrouzier) Epilachna pulchella Montrouzier, 1861: 306. Rhizobius submetallica Crotch, 1874: 298 ���synonymized by Tomaszewska 2010: 181. Rhizobius debilis Blackburn, 1888: 201 ���synonymized by Tomaszewska 2010: 181. Rhizobius lindi Blackburn, 1888: 202 ���synonymized by Tomaszewska 2010: 181. Rhizobius coerulens Blackburn, 1892: 256 ���synonymized by Tomaszewska 2010: 181. Rhizobius pulchellus Fauvel 1903: 321. This species was described from New Caledonia, and considered as an endemic species until a recent review by Tomaszewska (2010). Its range encompasses all Australian states. It is considered to be an autochthonous species in New Caledonia. The species description by Montrouzier was accompanied by an infrapaginal note, which has been attributed to his metropolitan correspondent: ���This insect arrived in bad condition, and then was unable to be identified���. Therefore, Bielawski (1973) established his review on specimens from Fauvel���s collection, in Brussels. This author made a redescription where he illustrated male genitalia, but he did not assign a lectotype. Tomaszewska (2010) established a lectotype and a paralectotype from the Fauvel collection (respectively a male from Canala County and a male from Boulari in Mont-Dore County). This species is recorded from the east and west coasts (Noum��a, Mont-Dore, Canala). Uncommon in the wild, we consider this species as a scale-feeding specialist. Literature records. NEW CALEDONIA: Kanala (Montrouzier rec.) (ISNB); Coul��e, Boulari (A. Fauvel rec.) (ISNB) (Tomaszewska 2010)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 312, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Tomaszewska, W. (2010) Rhyzobius Stephens, 1829 (Coleoptera, Coccinellidae), a revision of the world species. Fauna Mundi, Vol. 2, Polish Academy of Sciences, Warszawa, 475 pp.","Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia, 15, 207 - 261.","Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Genus Rhyzobius Stephens, 1829 Among the following species, only Rhyzobius fagus (Broun) can be considered as a common species. All species are suspected to feed on Diaspididae, especially from the tribe Aspidiotini., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 311, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Stephens, J. F. (1829) A Systematic Catalogue of British Insects. Bladwin and Cradock, London, 388 pp."]}

Nephus fijiensis (Sicard) Scymnus fijiensis Sicard, 1922: 353. Nephus fijiensis: Chazeau 1982: 181. Described from Fiji Islands, it is a Pacific-wide species [Papua New Guinea (including New Britain), French Polynesia, American Samoa, Western Samoa, Cook Islands)] but also in Asia (China, Vietnam). It was not recorded by Bielawski (1973). Its occurrence in the Noum��a area may indicate recent natural spread or an accidental introduction between 1970 and 1980. This scale and mealybug insect predator (Diaspidae, Pseudococcidae, Coccidae) is found on ornamental plants and in orchards. Material examined. NEW CALEDONIA: Noum��a, Anse Vata. (CPS), vi. 1966 (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 318, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Sicard, A. (1922) Descriptions de varietes, especes et genres nouveaux appartenant a la famille des Coccinellides. The Annals and Magazine of Natural History, 9 (9), 349 - 360.","Chazeau, J. (1982) Contribution a la connaissance des especes fidjiennes originellement classees dans le genre Scymnus, et description de deux Pseudoscymnus nouveaux de l'archipel (Col. Coccinellidae). Bulletin de la Societe Entomologique de France, 87, 180 - 187.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Scymnus (Scymnus) cochereaui Bielawski Scymnus (Scymnus) cochereaui Bielawski, 1973: 403. Endemic to New Caledonia, this species is only known from the type series (holotype and two paratypes, Noum��a, on coconut tree). No information is available on its biology and feeding ecology. Material examined. NEW CALEDONIA: Noum��a, on coconut, viii. 1968 (P. Cochereau rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 319, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Rodolia delobeli Chazeau Rodolia delobeli Chazeau, 1981: 57. This endemic species is only known from southern New Caledonia (Ouen-Toro, sclerophyllous forest, dense rain forest and ultramafic shrubs in Rivi��re Bleue, dense rain forest in La Thi Valley). It is an uncommon species and no information is available on its prey preferences. Material examined. NEW CALEDONIA: Noum��a, Ouen Toro, 20.x. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Ouen Toro, 24.vi. 1978 (A. Delobel rec.) (ONNC); For��t de la Thi, 27.iv. 1981 (J. Chazeau rec.) (ONNC); Rivi��re Bleue (parc. 7), for��t dense humide sur pente, 170m, 420. xii. 1985, 825. xii. 1986 (L. Bonnet de Larbogne & J. Chazeau rec.) (ONNC); Rivi��re Bleue, maquis sur cr��te, 310m, 25.xii. 19865. i. 1987, 5.20.i, 2031. i. 1987 (L. Bonnet de Larbogne, J. Chazeau, A.& S. Tillier rec.) (ONNC); Pinda��, 9.xi. 1982, for��t scl��rophylle (E. & M. Schlinger rec.) (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 316-317, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980

Coelophora quadrivittata Fauvel Coelophora quadrivittata Fauvel, 1903: 328. This is an endemic species from New Caledonia, known from both coasts and Loyalty Islands (Lifou). This taxon is highly polymorphic from a morphological point of view, which leads to some identification confusions with C. mulsanti. But C. quadrivittata is always smaller than C. mulsanti. Its biology has been well studied by Chazeau (1980, 1981a). Known preys are scale insects [Aspidiotus sp., Aspidiotus destructor Signoret, Coccus viridis (Green)] and psyllids (Heteropsylla cubana). It is a widespread species in open habitats, and attracts some interest for biocontrol programs against Coccomorpha (Chazeau 1981 a). Material examined. NEW CALEDONIA: Iles Loyaut��, Lifou, 11.ix. 1969 (P. Cochereau rec.) (ONNC); Mts Koghi, 10.x. 1958 (Y. Shibata rec.) (ONNC); Noum��a, i. 1955 (ONNC); Noum��a, Anse Vata, xi. 1976 (A. Delobel rec.) (ONNC); Noum��a, OuenToro, 27.x. 1977 (J. Fainika rec.) (ONNC); Noum��a, OuenToro, 24.vi. 1978 (A. Delobel rec.) (ONNC); Noum��a, Ile Nou (ONNC); Noum��a, OuenToro, 8.xii. 1977 (J. Fainika rec.) (ONNC); Noum��a, OuenToro, 15.xii. 1977 (J. Fainika rec.) (ONNC); Noum��a, Baie des Citrons, 27.vi. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Baie des Citrons, 6.vii. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Baie des Citrons, 27.vi. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Baie des Citrons, 6.vii. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Baie des Citrons, 8.x. 1986 (L. Bonnet de Larbogne rec.) (ONNC); Noum��a, Anse Vata, xi. 1978 (J. Chazeau rec.) (ONNC); Pa��ta, 28.ii. 1948 (ONNC); Ponerihouen, 1985 (R. Amice rec.) (ONNC); Port-Laguerre, 7 / 14.iv. 1977 (A. Delobel rec.) (ONNC); Port-Laguerre, 15.vi. 1977 (A. Delobel rec.) (ONNC); Pouembout, Franco Plage, 19.iii. 1982 (J. Chazeau rec.) (ONNC); St Gabriel (Thio), 12.vi. 1990 (L. Bonnet de Larbogne & J. Chazeau rec.) (ONNC); Timbia (Pa��ta), 25.xii. 1992 (J. Chazeau rec.) (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 306-307, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Chazeau, J. (1981 a) Donnees sur la biologie de Coelophora quadrivittata (Col.: Coccinellidae), predateur de Coccus viridis (Hom.: Coccidae) en Nouvelle-Caledonie. Entomophaga, 26, 301 - 312."]}

Scymnus (Pullus) unicolor (Montrouzier) Epilachna unicolor Montrouzier, 1861: 306. Epilachna emarginata: Montrouzier, 1864: 219. Scymnus unicolor: Fauvel 1903: 323. Scymnus (Pullus) unicolor: Korschefsky 1931: 149. Endemic to New Caledonia, this species has been recorded from both coasts (Noum��a, Canala), Isle of Pines and Lifou (Loyalty Islands) (Chazeau, pers. obs.). No information is available on its biology. Literature records. NEW CALEDONIA: Noum��a (Delauney rec.) (coll. Fleutiaux in MNHN); Isle of Pines (coll. A. Fauvel in IRSNB); Saint-Louis (Montrouzier rec.) (coll. A. Fauvel in IRSNB) (Bielawski 1973)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 319-320, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Montrouzier, P. (1864) Essai sur la faune entomologique de Kanala (Nouvelle-Caledonie) et description de quelques especes nouvelles ou peu connues. Annales de la Societe Linneenne de Lyon, 11, 46 - 257.","Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Korschefsky, R. (1931) Pars 118: Coccinellidae I. In: Junk, W. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 224.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Diomus caledoniensis (Bielawski) Scymnus (Diomus) caledoniensis Bielawski, 1973: 400. Diomus caledoniensis shows rough morphological similarities with Scymnus unicolor (Montrouzier). It has been described by Bielawski (1973) as an endemic species, from specimens which were mixed in collection with Scymnus unicolor Montrouzier. This taxon is known from southern Grande Terre (Mont-Dore area: Boulari, Saint- Louis, Mont-Dore and Rivi��re Bleue), including dense rain forest habitat. Its biology and diet are unknown. Material examined. NEW CALEDONIA: Coul��e Boulari, Mont Dore (Delauney rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 313, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Orcus chujoi Bielawski Orcus chujoi Bielawski, 1961: 3. This endemic species is recorded from the west coast (Poya) and the east coast (Pou��bo) and the Central Range (Sarram��a, Mt Aoupini��, For��t Plate, Mt Yambi, Mt Djiaouma). It is an uncommon species in the wild. This Orcus would be a predator of hemipterous prey in forested areas. According to Laczynski & Tomaszewska (2009), the pattern and coloration of the elytra and pronotum are unique within the genus and easily separates O. chujoi from all its congeners. Material examined. NEW CALEDONIA: Mt Djiaouma, 1020m, 18.x. 1988 (J. Chazeau rec.) (ONNC); Pou��bo (around), 23.xi. 1987 (J.P. Morin rec.) (ONNC); Poya, 22.v. 1958 (Y. Shibata rec.) (ONNC); Sarram��a, montain, 10.ii. 1968 (P. Cochereau rec.) (ONNC); Pon��rihouen, Aoupini��, 21.1038 ��S / 165.1707 ��E / 945m, 11.x. 2006 (C. Mille rec.) (CXMNC); Pon��rihouen, Aoupini��, 21.1038 ��S / 165.1707 ��E / 945m, 11.x. 2006 (S. Caz��res rec.) (CXMNC); Pon��rihouen, Aoupini��, 21 ��08' 9.46 ''S / 165 �� 19 ' 4.07 ''E / 389m, 25.iv. 2006 (C. Mille & J. Brinon rec.) (CXMNC); Ou��goa, Mandj��lia, 20.39683 ��S / 164.53218 ��E / 787m, 07.ii. 2005 (S. Caz��res & C. Mille rec.) (CXMNC); Ou��goa, Mandj��lia, 20.39683 ��S / 164.53218 ��E / 787m, 14.xii. 2009 (C. Mille, J. Brinon, J-P. Kataoui rec.) (CXMNC); Farino, "Tend��a", 21.61431 ��S / 165.70065 ��E / 408m, 23.vi. 2004 (S. Caz��res rec.) (CXMNC); Farino, "Tend��a", 29.iv. 2004 (S. Caz��res rec.) (CXMNC); Sarram��a, Col d'Amieu, 21.62418 ��S / 165.81045 ��E / 514m, 18.xi. 2004 (S. Caz��res & J. Brinon rec.) (CXMNC); Sarram��a, Col d'Amieu, 21 �� 34.684 ��S/ 165 �� 46.278 ��E/ 489m, 04.x. 2006 (S. Caz��res rec.) (CXMNC). Literature records. NEW CALEDONIA: Mt. Aoupini��, 700 m, 18.IV. 1996, M.S. Moulds (1: ANIC); For��t Plate, 600 m, 22.VI. 1996, M.S. Moulds, Orcus chujoi Biel., det. A. S. Ślipiński (1: ANIC); Yambi, NE, 500���700 m, 14.X. 1967, J. & M. Sedlacek, Collectors, Bishop (1: BPBM); 10km S. of Koh, 31.I. 1963 (1: BPBM) (Laczynski & Tomaszewska 2009)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 303, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Bielawski, R. (1961) Materialien zur Kenntnis Coccinellidae (Col.) Melanesien's. Niponius, 1 (13), 1 - 9.","Laczynski, P. & Tomaszewska, W. (2009) Revision of the genus Orcus Mulsant (Coleoptera: Coccinellidae: Chilocorini). Annales Zoologici, 59, 585 - 611."]}

Rhyzobius fagus (Broun) Scymnus fagus Broun, 1880: 648. Rhizobius satelles Blackburn, 1892: 255 ���synonymized by Tomaszewska 2010: 109. Rhizobius kingensis Lea, 1908: 206 ���synonymized by Tomaszewska 2010: 109. Rhizobius erythrogaster Lea, 1929: 241 ���synonymized by Tomaszewska 2010: 109. Rhyzobius nigrovatus Bielawski, 1973: 391 ���synonymized by Tomaszewska, 2010: 109. Some misidentifications of this species have led to false records in the literature from New Caledonia (Huffaker & Messenger 1976): Lindorus lophanthae (Blaisdell) and Rhyzobius pulchellus (Montrouzier). Bielawski (1973) made a new description under Rhyzobius nigrovatus (type locality: Noum��a). Tomaszewska (2010) redescribed this species and gave an account of its geographic range. This Australian species appears to have recently spread to New Caledonia and Vanuatu, where it is now quite common. In Vanuatu, it has been heavily used and promoted against the coconut scale Aspidiotus destructor Signoret (Hemiptera: Diaspididae) between 1960 and 1980 (Chazeau 1981 b). This efficient diaspidid feeder was also deliberately introduced to the Caroline Islands (Nafus & Schreiner 1989) and Wallis Island in 1965. It has been seen established in Wallis in 1981 (Gutierrez 1981), and also in South Africa (before 1988). Apart from Aspidiotus destructor Signoret, it has been seen to feed on Aspidiotus nerii Bouch��, Aulacaspis tubercularis Newstead, Pseudaulacaspis pentagona (Targioni-Tozzetti), Lepidosaphes beckii (Newman) (Cochereau 1965, 1969). Literature records. NEW CALEDONIA: Noum��a (A. Fauvel rec.) (ISNB); Noum��a, 0-100m, ii. 1980 (N.H.L. Krauss rec.) (BPBM); Noum��a, 0-100m, 3.iii. 1980 (N.H.L. Krauss rec.) (BPBM); Hills behind Noum��a, 15.viii. 1940 (F.X. Williams rec.) (BPBM); Noum��a, 26.vii. 1940 (F.X. Williams rec.) (BPBM); Noum��a, 24.vii. 1940 (F.X. Williams rec.) (BPBM); Noum��a, 13.viii. 1940 (F.X. Williams rec.) (BPBM); In Mts. above Ouaco, 20.x. 1958 (C.R. Joyce rec.) (BPBM); Koebele (BPBM); La Crouen, 16.iii. 1961 (J. Sedlacek rec.) (BPBM); Poindimi��, 26.xi. 1958 (C.R. Joyce rec.) (BPBM); Mt Mou, 1100 m, 9.iii. 1972 (J.L. Gressit rec.) (BPBM); Mokoue to Dothio, 150-500 m, 20-22.iii. 1968 (T.C. Maa rec.) (BPBM) (Tomaszewska 2010)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 311, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Broun, T. (1880) Manual of the New Zealand Coleoptera. Colonial Museum and Geological Survey Department, Wellington, iv + 651 pp.","Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia, 15, 207 - 261.","Tomaszewska, W. (2010) Rhyzobius Stephens, 1829 (Coleoptera, Coccinellidae), a revision of the world species. Fauna Mundi, Vol. 2, Polish Academy of Sciences, Warszawa, 475 pp.","Lea, A. M. (1908) The Coleoptera of King Island, Bass Strait. Proceedings of the Royal Society of Victoria (n. s.), 20 (2), 143 - 207.","Lea, A. M. (1929) Notes on some miscellaneous Coleoptera, with descriptions of new species. Part VII. Transactions of the Royal Society of South Australia, 53, 203 - 244.","Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409.","Huffaker, C. B. & Messenger, P. S. (1976) Theory and Practice of Biological Control. Academic Press, New York, 810 pp.","Chazeau, J. (1981 b) La lutte biologique contre la cochenille transparente du cocotier Temnaspidiotus destructor (Signoret) aux Nouvelles-Hebrides (Homoptere Diaspididae). Cahiers ORSTOM. Serie Biologie: Phytopathologie et Zoologie Appliquee, 44, 11 - 22.","Nafus, D. & Schreiner, I. (1989) Biological control activities in the Mariana Islands from 1911 to 1988. Micronesica, 22, 65 - 106.","Gutierrez, J. (1981) Catalogue des ravageurs de Wallis et Futuna. Rapport ORSTOM, Noumea, 26 pp.","Cochereau, P. (1965) Contre un ravageur du cocotier aux Nouvelles-Hebrides: controle biologique d' Aspidiotus destructor Signoret (Homoptera-Diaspinae) par Lindorus lophantae Baisd. (Coleoptera-Coccinellidae), ile Vate. Oleagineux, 20, 507 - 512.","Cochereau, P. (1969) Controle biologique d' Aspidiotus destructor Signoret (Homoptera Diaspinae) dans l'ile de Vate (Nouvelles Hebrides au moyen de Rhizobius pulchellus Montrouzier (Coleoptera Coccinellidae). Cahiers ORSTOM. serie Biologie, 8, 57 - 100."]}

Stethorus proximus Chazeau Stethorus proximus Chazeau, 1979: 296. Endemic to New Caledonia, it has been recorded from the west coast, where it seems restricted to sclerophyllous forest patches: Noum��a (Ouen Toro), Pa��ta (Mt Nondou��), Pinda�� peninsula. It has been seen to feed on Tetranychus neocaledonicus Andr�� by Chazeau (1979). Material examined. NEW CALEDONIA: Noum��a, Ouen-Toro, 23.vi. 1977 (J. Chazeau rec.) (MNHN); Pa��ta, Mt Nondou��, 110m, fogging on sclerophyllous forest, 28.iv. 1993 (J. Chazeau, E. Guilbert & L. Bonnet de Larbogne rec.) (MNHN), Pinda��, 30m, fogging on sclerophyllous forest, 06.x. 1992 (J. Chazeau, E. Guilbert & L. Bonnet de Larbogne rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 320, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Chazeau, J. (1979) Mise au point sur le genre Stethorus en oceanie et description de deux especes nouvelles de Melanesie [Col., Coccinellidae]. Entomophaga, 24, 295 - 303."]}

Rodolia cardinalis (Mulsant) Vedalia cardinalis Mulsant, 1850: 906. Novius cardinalis: Crotch 1874: 283. Eurodolia cardinalis: Weise 1895: 150. Rodolia (Macronovius) cardinalis: Weise 1905: 220. Macronovius cardinalis: Weise 1922: 104. Rodolia cardinalis: Korschefsky 1931: 101. This species was originally described from Australia. It is considered to be a recent introduction in New Caledonia and Cook Islands. It has been introduced successfully into Vanuatu, Fiji Islands, Micronesia (Guam), Hawaii, in the southern United States (California, Florida, Louisiana, South Carolina and Texas), southern Europe, North Africa, South Africa, India and Indonesia (Java). It was also accidentally introduced to New Zealand, where it is not widespread. The date of arrival in New Caledonia is unknown, but it probably dispersed from Vanuatu (introduced to there in 1964 by Cochereau from Fiji Islands). It has been regularly collected since its first reported occurrence in 1975 in Noum��a but remains uncommon. This species is a major biocontrol agent worldwide as it represents the world���s first biocontrol success after its introduction in California in citrus orchards to control Icerya purchasi Maskell outbreaks (Debach 1974). This is an aggressive predator against scale insects, especially Icerya spp. (Hemiptera, Monophlebidae). In Noum��a, it has been seen feeding on I. seychellarum (Westwood). Material examined. NEW CALEDONIA: Noum��a, Anse Vata, 11.vii. 1975 (J. Gutierrez rec.) (ONNC); Noum��a, 28.ix. 1977 (J. Fons rec.) (ONNC); Noum��a, Anse Vata, ix. 1978 (J. Chazeau rec.) (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 316, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Weise, J. (1905) Ueber Coccinelliden. Deutsche Entomologische Zeitschrift, 1905, 217 - 220.","Weise, J. (1922) Ueber einige amerikanische und australische, nach Sudfrankreich eingefuhrte Coccinelliden. Wiener Entomologische Zeitung, 39, 104.","Korschefsky, R. (1931) Pars 118: Coccinellidae I. In: Junk, W. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 224.","Debach, P. (1974) Biological Control by Natural Enemies. Cambridge University Press, London, 334 pp."]}

Olla v-nigrum (Mulsant) Harmonia V-nigrum Mulsant, 1866: 64. Coccinella V-nigrum: Crotch 1874: 109. Cycloneda abdominalis Crotch 1874: 163 ���synonymized by Gorham 1892: 172. Olla sobrina Casey, 1899: 95 ���synonymized by Chapin 1974: 64. Olla fenestralis Casey, 1899: 95 ���synonymized by Belicek 1976: 329. Olla plagiata Casey, 1899: 94 ���synonymized by Chapin 1974: 64. Olla minuta Casey, 1908: 406 ���synonymized by Casey 1924: 157. Olla v-nigrum: Timberlake 1943: 24. This American species has been introduced (from a Tahitian strain) to New Caledonia in 1987 to achieve biological control against Heteropsylla cubana (Hemiptera: Psyllidae), which was destroying Leucaena leucocephala (Lam.) de Wit, a legume used as fodder for extensive cattle breeding (Chazeau 1987; Chazeau et al. 1991; Disney & Chazeau 1990). This ladybug is now widespread all over New Caledonia, including Isle of Pines and Walpole Island. There are two color morphs which have been introduced, the paler one is the most widespread in New Caledonia (Gatimel 2005). This species is also recorded from North America (its native range), Hawaii and French Polynesia. It is known to feed on various prey sources (Chazeau et al. 1991): psyllids (e.g., Heteropsylla cubana), aphids [e.g., Cerataphis lataniae (Boisduval)] and cicadellids (e.g., Iassinae larvae). Material examined. NEW CALEDONIA: Bac de la Oua����me, rive Sud, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Bouloupari, iv-vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Bourail, route belv��d��re de la Roche Perc��e, iv-vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Bourail, bord de la N��ra, 12.iv. 1988 (L. Bonnet de Larbogne, J. Chazeau rec.) (ONNC); Bourail, Poe, Creek Sal��, 23.vii. 1988 (J. Chazeau rec.) (ONNC); Bourail, bord de la N��ra, 23.vii. 1988 (J. Chazeau rec.) (ONNC); Bourail, route belv��d��re de la Roche Perc��e, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Bourail, route belv��d��re de la Roche Perc��e, 30.iii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Bourail, route de Nessadiou, 30.iii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Bourail, Gouaro, 30.iii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Canala, 20.vi. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Col de Koh, Rte La Foa-Canala, 20.vi. 1989 (ONNC); Col d'Amos, versant Est, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Col de BirHakeim, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Col de la Pirogue, v. 1987 (J. Chazeau rec.) (ONNC); Col de la Pirogue, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Col des Mimosas, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Col de Nassirah, versant Boulouparis, 27.vi. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Col de Nassirah, versant Thio, 27.vi. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Col de Nekoro, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Col des Roussettes, 5.iv. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Col de T��mala, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Entre Tipinje et Vieux Touho, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Goro, xii. 1989 (L. Bonnet de Larbogne, J. Chazeau rec.) (ONNC); Goyetta, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Hiengh��ne Nord, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Hiengh��ne belv��d��re, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Houa��lou Nord, 5.iv. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Ile Walpole, 11-15.viii. 1988 (A. Renevier rec.) (ONNC); Kouaoua, 28.v. 1989 (J. Chazeau rec.) (ONNC); Koumac, 22.ix. 1988 (J. Chazeau rec.) (ONNC); La Foa, 5.vii. 1989 (L. Bonnet de Larbogne, E. Bouy�� rec.) (ONNC); Noum��a, Ile Nou, ix. x. 1987 (R. Amice, L. Bonnet de Larbogne, L. O. Brun rec.) (ONNC); Noum��a, Ile Nou, 11.iv. 1988 (J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, 20.v. 1988 (L. Bonnet de Larbogne, J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, iv, v, vi, vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, 11.i. 1992 (J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, 3.ii. 1992 (L. Bonnet de Larbogne, J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, 18.iii. 1992 (L. Bonnet de Larbogne, J. Chazeau rec.) (ONNC); Noum��a, marinas, 3.vii. 1988 (J. Chazeau rec.) (ONNC); Ouaco, iv-vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Pa��ta, route de Tiar��, iv-vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Pied du Col d'Amos, C��te Est, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Pinda��, bord de mer, 6.x. 1992 (J. Chazeau rec.) (ONNC); Pont de la Rivi��re du Cap, RT1, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Poindimi�� Sud, 23.ix. 1988 (J. Chazeau rec.) (ONNC); Port Bois��, wharf; 25.x. 1988 (J. Chazeau rec.) (ONNC); Pouembout, iv-vii. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); RT 1, pk 66, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Tao, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Timbia (Paita), 25.xii. 1992 (J. Chazeau rec.) (ONNC); Thio, 27.vi. 1989 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Vall��e de la Tontouta, Mine Liliane, 18.iii. 1988 (J. Danloux rec.) (ONNC); Voh, 20.ix. 1988 (J. Chazeau rec.) (ONNC); Ile Walpole, 11 / 15.viii. 1988 (A. Renevier rec.) (ONNC); Koumac, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Mt Do, 1015 m, 31.x. 1992 (J. Chazeau, E. & M. Schlinger rec.) (ONNC); Mt Do, 900-1000 m, 4.xii. 1992 (J. Chazeau & D. Webb rec.) (ONNC); Boulouparis, Mont Do, 21.75394 ��S / 161.99994 ��E / 1029m, 13.iv. 2006 (M. & P. Jolivet rec.) (CXMNC); Boulouparis, Mont Do, 07.xii. 2003 (C. Mille rec.) (CXMNC); Boulouparis, Mont Do, 21.75394 ��S / 161.99994 ��E / 1029m, 27.xi. 2012 (C. Mille rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 26.xi. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 07.i. 2002 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 21.73205 ��S / 165.89936 ��E / 32m, 09.iii. 2007 (N. Degallier rec.) (CXMNC); La Foa, 04.iii. 2004 (R-M. M'Boueri rec.) (CXMNC); La Foa, 24.vi. 2002 (J-P. Kataoui rec.) (CXMNC); Yat��, Prony, COL /ins/ 67 -14, 2006 (H. Jourdan rec.) (CXMNC); Farino, Tend��a, 21.61928 ��S / 165.70907 ��E / 403m, 29.iv. 2004 (S. Caz��res rec.) (CXMNC); Mar��, 21.xi. 2001 (J. Brinon rec.) (CXMNC); Bourail, Gouaro, 21.58113 ��S / 165.42045 ��E / 68m, i. 2001 (S. Caz��res rec.) (CXMNC); Bourail, Gouaro, 21.58113 ��S / 165.42045 ��E / 68m, 11.x. 1998 (S. Caz��res rec.) (CXMNC); Pa��ta, Mont Humboldt, 21.88277 ��S / 166.4125 ��E / 1350m, 15.ii. 2006 (S. Caz��res rec.) (CXMNC); Koumac, rivi��re Bambou, route de Pou��bo, 17.iii. 1995 (M. Boulard rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 309-310, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Mulsant, E. (1866) Monographie des Coccinellides. 1 re partie Coccinelliens. Savy & Deyrolle, Paris, 294 pp.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Gorham, H. S. (1892) Insecta, Coleoptera: Coccinellidae. Biologia Centrali-Americana, 7, 161 - 176.","Casey, T. L. (1899) A revision of the American Coccinellidae. Journal of the New York Entomological Society, 7, 71 - 169.","Chapin, J. B. (1974) The Coccinellidae of Louisiana (Insecta: Coleoptera). Louisiana Agricultural Experiment Station Bulletin, 682, 2 - 87.","Casey, T. L. (1908) Notes on the Coccinellidae. Canadian Entomologist, 40, 393 - 421.","Casey, T. L. (1924) Additions to the known Coleoptera of North America. Memoirs on the Coleoptera, 11, 1 - 347.","Timberlake, P. H. (1943) The Coccinellidae or Ladybeetles of the Koebele Collection. Part 1. Hawaiian Sugar Planter's Association, Honolulu, 67 pp.","Chazeau, J. (1987) Le psylle du faux-mimosa en Asie du Sud-Est et dans le Pacifique: etat du probleme et perspectives de lutte (Leucaena leucocephala (Lam.) de Wit- Heteropsylla cubana Crawford). Revue d'Elevage et de Medecine Veterinaire de Nouvelle Caledonie, 9, 23 - 27.","Disney, R. H. L. & Chazeau, J. (1990) The recognition and biology of Phalacrotophora quadrimaculata (Diptera: Phoridae) parasitising Olla v-nigrum (Coleoptera: Coccinellidae) used in attemps to control the Leucaena psyllid. Annales de parasitologie humaine et comparee, 65, 98 - 100.","Gatimel, B. (2005) Inventaire critique et perspectives des introductions d'especes realisees a des fins de lutte biologique en Nouvelle-Caledonie: applications aux cas de la mineuse des agrumes (Phyllocnistis citrella Stainton) et du psylle du fauxmimosa (Heteropsylla cubana Crawford). Master of Science (MSc), Ecology & Biodiversity Management, Universite Montpellier 2, 30 pp."]}

Coccinella transversalis Fabricius Coccinella transversalis Fabricius, 1781: 97. Coccinella repanda Thunberg, 1781: 18 ���synonymized by Mulsant 1850: 1022 (as Coccinella repanda Casstr��m). The synonymy, established by Mulsant (1850) (first reviewer who attributed the species authorship to Casstr��m) is currently accepted by recent authors (e.g. Pope 1987; Slipinski 2007). C. transversalis is a very widespread species, it has been recorded widely in the Asia-Pacific region, from India to Polynesia, including Australia, Tasmania, Papua New Guinea, New Zealand, Vanuatu, Solomon Islands, Fiji Islands, Micronesia, Cook Islands, Wallis & Futuna, and French Polynesia. In New Caledonia, it is a common species on both coasts, including high altitudes (greater than 1,000m), Isle of Pines, Belep Islands, Loyalty Islands, Surprise, Chesterfield and Walpole islands. This is a polyphagous predator of various Hemiptera (Aphididae, Psyllidae, Coccomorpha). Humans have promoted its spread and we consider it as introduced according to its biological traits (since aphids are all introduced in New Caledonia; CM & HJ, unpublished data) and distribution. When prey is scarce, it is also known to feed on pollen and extrafloral nectaries (Evans 2000) which are uncommon in the New Caledonian native flora. This species has attracted great interest in biocontrol especially against aphids (Aphis gossypii Glover) and against the pseudococcid Planococcus citri (Risso) (Waterhouse 1998). Material examined. NEW CALEDONIA: Bond��, vall��e du Diahot, vii. 1955 (ONNC); Bourail, 11.iv. 1988 (J. Chazeau rec.) (ONNC); Canala, 23.vii. 1977 (A. Delobel rec.) (ONNC); Col d'Amieu, Tribu de Couli, vi. 1955 (ONNC); Col de Cr��vecoeur, 22.ix. 1988 (J. Chazeau rec.) (ONNC); Col des Mimosas, RT1, 16.iv. 1986 (J. Chazeau rec.) (ONNC); Col des Roussettes, 16.iv. 1986 (J. Chazeau rec.) (ONNC); D��me de Thi��baghi, 18.iii. 1982 (J. Chazeau rec.) (ONNC); Dumb��a, chez Trichard, 9.ii. 1977 (J. Chazeau rec.) (ONNC); Environs de Galarino, 12.ii. 1966 (ONNC); Environs de Port-Laguerre, 29.i. 1978 (A. Delobel rec.) (ONNC); Hiengh��ne, 16.iii. 1982 (J. Chazeau rec.) (ONNC); Houa��lou, vi. 1955 (ONNC); Ile Surprise, 11.i. 1989 (P. Laboute rec.) (ONNC); Ile Walpole, 1115. viii. 1988 (A. Renevier rec.) (ONNC); Kaala-Gomen, 22.ix. 1986 (J. Chazeau rec.) (ONNC); Mandj��lia, ancienne scierie, ii. 1985 (J. Danloux rec.) (ONNC); Mt Do, 9.v. 1990 (J. Danloux rec.) (ONNC); Mt Do, 1015 m, 31.x. 1992 (J. Chazeau rec., E.& M. Schlinger rec.) (ONNC); Mt Mou, iii. 1956 (ONNC); Mt Mou, sentier captage tribu des Pirogues, 14.ii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, xii. 1954 (ONNC); Noum��a, iiv, vii, ix, xi, xii. 1955 (ONNC); Noum��a, i, vi. 1956 (ONNC); Noum��a, xii. 1957 (ONNC); Noum��a, ii. 1958 (ONNC); Noum��a, iii. 1959 (ONNC); Noum��a, ii. 1964 (ONNC); Noum��a, x. 1976 (A. Delobel) (ONNC); Noum��a, Anse Vata, 25.ii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Ile Nou, 11.iv. 1988 (J. Chazeau rec.) (ONNC); Noum��a, jardin Mus��e, 9.iii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Ouen Toro, 24.vi. 1978 (A. Delobel) (ONNC); Ponerihouen, 1985 (R. Amice rec.) (ONNC); Port-Laguerre, ii-v. 1977 (A. Delobel) (ONNC); Port-Laguerre, Station Agricole, 3.iii. 1977 (J. Chazeau rec.) (ONNC); Pouembout, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Poum, 4.xii. 1958 (Y. Shibata rec.) (ONNC); Poum, 18.iii. 1986 (J. Chazeau rec.) (ONNC); Poum, La Gologne, 18.iii. 1988 (J. Chazeau rec.) (ONNC); Poya, 16.iv. 1986 (J. Chazeau rec.) (ONNC); Thio, bord de rivi��re, 1.xi. 1976 (A. Delobel rec.) (ONNC); Thio Ourou��, 5.i. 1982 (J. Chazeau rec.) (ONNC); Vall��e de la Boghen, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Voh, Tribu de Keta, vi, viii. 1955 (ONNC); Ile Loop et il��ts Avon, Chesterfield islands, xi. 2012 (E. Bourguet et H. Jourdan rec.) (ONNC); La Foa, Pocquereux, 21 �� 44 'S / 165 �� 54 'E / 28m, 20.iv. 2001 (J. Brinon rec.) (CXMNC); Sarram��a, 21.66941 ��S / 165.81723 ��E / 3m, 21.xi. 2001 (J. Brinon rec.) (CXMNC); La Foa, 21 �� 43 ���S / 165 �� 49 ���E, 24.ii. 2004 (R-M. M'Boueri rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 'S / 165 �� 54 'E / 28m, 07.viii. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 'S / 165 �� 54 'E / 28m, 24.ii. 2004 (R-M. M'Boueri rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ' 124 ''S/ 165 �� 53 ' 852 ''E/ 38m, 19.ii. 2015 (M. Dierkens rec.) (CXMNC); Yat��, Prony, COL /ins/ 67 -14, 2006 (H. Jourdan rec.) (CXMNC); Boulouparis, Ouenghi, COL /ins/ 41 -14, 21�� 53 ' 44.62 "S / 166 ��06' 41.53 "E / 7m, 28.i. 2014 (C. Mille rec.) (CXMNC); Boulouparis, Ouenghi, COL /ins/ 41 -14, 28.i. 2014 (C. Mille rec.) (CXMNC); Ouv��a, T��outa-ounes, 20 �� 24 ' 40 ''S / 166 �� 36 ' 20 ''E, COL /ins/ 87 -14, 14.iii. 2014 (C. Mille rec.) (CXMNC); Pouembout, 21 ��08' 48.54 ''S / 164 �� 55 ' 25.31 ''/ 38m, COL /ins/ 220 -14, 15.vii. 2014 (S. Tron rec.) (CXMNC); Pouembout chez M. Soury-Lavergne, 11.vii. 2013 (C. Mille rec.) (CXMNC); Pouembout chez M. Soury-lavergne, COL /ins/ 255 -14, 01.viii. 2014 (C. Mille rec.) (CXMNC); La Foa, Pocquereux, 05.iii. 2004 (R-M. M'Boueri rec.) (CXMNC); Sarram��a, Col d'amieu, 26.x. 2005 (S. Caz��res & J-P. Kataoui rec.) (CXMNC); Boulouparis, Mont Do, 1029m, 03.xii. 2009 (CXMNC); Boulouparis, Ouenghi, 7m, 12.ix. 2012 (R-M. M'Boueri rec.) (CXMNC); Bourail, 1929 (Risbec rec.) (MNHN); Timbia, 1929 (Risbec rec.) (MNHN); Plum, 1928 (Pruvot rec.) (MNHN); Iles Loyaut��, Lifou, 1928 (Pruvot rec.) (MNHN); Noum��a, 4.i. 1951 (MNHN); Canala, 1869 (Delacour rec.) (MNHN)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on pages 304-305, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Fabricius, J. C. (1781) Species Insectorum; Exhibentes Eorum Differentias Specificas, Synonyma, Auctorum, Loca Natalia, Metamorphosin Ediectis Observationibus, Descriptionibus. Vol. 1. Carol. Ernest. Bohnii, Hamburgi & Kilonii, VIII + 552 pp.","Thunberg, C. P. (1781) Dissertatio Entomologica Novas Insectorum Species, Sistens, Cujus Partem Primam. Joh. Edman, Upsaliae, 28 pp.","Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Pope, R. D. (1987) The Coccinellidae (Coleoptera) described by CP Thunberg. Insect Systematics & Evolution, 18, 51 - 66.","Slipinski, A. (2007) Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their Biology and Classification. Australian Biological Resources Study, Canberra, xviii + 286 pp.","Evans, E. W. (2000) Egg production in response to combined alternative foods by the predator Coccinella transversalis. Entomologia Experimentalis et Applicata, 94, 141 - 147.","Waterhouse, D. F. (1998) Biological Control of Insect Pests: Southeast Asian Prospects (n ° 51). Australian Centre for International Agricultural Research, Canberra, Australia, 548 pp."]}

Nephus fauveli (Bielawski) Scymnus (Nephus) fauveli Bielawski, 1973: 406. Endemic to New Caledonia, this Nephus is only known from the type specimen (Noum��a) and few records which established its occurrence on both coasts of Grande Terre (Noum��a, Thio). Material examined. NEW CALEDONIA: Noum��a, Ouen Toro, 24.vi. 1978 (A. Delobel rec.) (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 318, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Bielawski, R. (1973) Rhyzobiini, Stethorini, Scymnini et Pharini (Coleoptera, Coccinellidae) de Nouvelle Caledonie. Annales Zoologici, 30, 387 - 409."]}

Henosepilachna sumbana (Bielawski) Epilachna vigintioctopunctata sumbana Bielawski, 1959: 147. Epilachna cucurbitae Richards, 1983: 37 ���synonymized by Jadwiszczak & Wegrzynowicz 2003: 173. Henosepilachna sumbana: Jadwiszczak & Wegrzynowicz 2003: 173. Nomenclature of Epilachnini has been highly debated during recent decades, and H. sumbana has not been cited under this name in the context of local census. But, it is highly probable that the ��� vigintioctopunctata ��� recorded in the region (i.e. Epilachna with 28 dots), which attacks Cucurbitaceae, belongs to this species. It was originally described from Indonesia, but this taxon was redescribed from Australia (Epilachna cucurbitae Richards, 1983, synonymized with Epilachna sumbana Bielawski, 1959: Jadwiszczak & Wegrzynowicz, 2003, as quoted by Slipinski 2007). It is also recorded widely in Asia and the Pacific region (Australia, Indonesia, West Timor, Philippines, Papua New Guinea, Solomon Islands, Vanuatu, Fiji Islands, Samoa and Wallis & Futuna). This species was not recorded by Bielawski (1972) and therefore must have been introduced accidentally to New Caledonia after Bielawski���s study. This ladybug was frequent around Noum��a and Dumb��a counties by the end of the 1970 s (Chazeau, pers. obs.), but today it is rarer. In New Caledonia, its host plants are cultivated or wild Cucurbitaceae, suggesting that this species belongs to the vigintioctopunctata species complex, whereas the two Australian species are specific to Solanaceae (Richards & Filewood 1990). Material examined. NEW CALEDONIA: Dumb��a, 9.ii. 1977 (J. Chazeau rec.) (ONNC)., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 315, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Bielawski, R. (1959) Coccinellidae (Coleopt.) von Sumba, Sumbawa, Flores, Timor und Bali. Verhandlungen der Naturforschenden Gesellschaft in Basel, 69, 145 - 166.","Richards, A. M. (1983) The Epilachna vigintioctopunctata complex (Coleoptera: Coccinellidae). International Journal of Entomology, 25, 11 - 41.","Jadwiszczak, A. S. & Wegrzynowicz, P. (2003) World catalogue of Coccinellidae Part I - Epilachninae. Mantis, Olsztyn, 264 pp.","Slipinski, A. (2007) Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their Biology and Classification. Australian Biological Resources Study, Canberra, xviii + 286 pp.","Bielawski, R. (1972) The Epilachninae (Coleoptera, Coccinellidae) of New Caledonia. Bulletin de l'Academie polonaise des sciences. Serie des sciences biologiques, 20, 557 - 561.","Richards, A. M. & Filewood, L. W. (1990) Feeding behaviour and food preferences of the pest species comprising the Epilachna vigintioctopunctata (F.) complex (Col., Coccinellidae). Journal of Applied Entomology, 110, 501 - 515."]}

Henosepilachna urvillei (Montrouzier) Epilachna urvillei Montrouzier, 1861: 305. Epilachna moultoni Crotch, 1874: 89 ���synonymized by Korschefsky 1933: 236. Henosepilachna urvillei: Bielawski 1972: 559. This species was described from Lifou (type specimen) by Montrouzier. Bielawski (1963) added details to the original description. This ladybug is widespread on Grande Terre, Loyalty Islands and Isle of Pines (Chazeau, pers. obs.). It appears widely distributed in Melanesia, occurring in Papua New Guinea, Solomon and Fiji islands. Li (1993) stated that only one specimen is recorded from Australia, where this species may not be established., Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 316, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980, {"references":["Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Korschefsky, R. (1933) Synonymische und andere Bemerkungen uber Crotch'sche Coccinelliden-Typen. Proceedings of the Royal Entomological Society of London. Series B, 2, 236 - 237.","Bielawski, R. (1972) The Epilachninae (Coleoptera, Coccinellidae) of New Caledonia. Bulletin de l'Academie polonaise des sciences. Serie des sciences biologiques, 20, 557 - 561.","Bielawski, R. (1963) Monographie der Epilachninae (Coleoptera, Coccinellidae) der Australischen Region. Annales Zoologici, 21, 295 - 461.","Li, C. S. (1993) Review of the Australian Epilachninae (Coleoptera: Coccinellidae). Australian Journal of Entomology, 32, 209 - 224."]}

Genus Agnotecous Saussure, 1878 Type species: Agnotecous tapinopus Saussure, 1878 Diagnosis. Among Lebinthini genera, Agnotecous species are characterised by stout global shape, and male and female FWs and genitalia. Size small to large for subfamily, generally larger than Lebinthus species. Colouration little contrasted, yellowish brown to dark brown, more or less mottled with brown, without distinctive marks. Head globular, dorsum flat; fastigium very wide. Eyes little prominent. Pronotum distinctively long and wide; dorsal disc slightly wider than long, trapezoidal, straight posteriorly. FWs short in males and females, covering less (males) or much less (females) than half abdomen, and tightly set on body; hindwings lacking in both sexes. Other general traits: TI, TII laterally compressed. TI with two small tympana, inner slit like, outer oval; membrane of outer tympanum transversally plicate in dorsal half; TI with three apical spurs; outer dorsal spur missing. TII with four apical spurs, inner longest. TIII with three inner and three outer apical spurs, median longest on each side; median outer spur as long as other outer spurs and obviously broader; four pairs of subapical spurs, inner spurs distinctively curved and longer; TIII serrulate over whole length and slightly furrowed dorsally. Tarsomeres III- 1 with two rows of dorsal spines in addition to apical spines, inner row with one or few spines., Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 19-20, DOI: 10.5281/zenodo.196996, {"references":["Saussure, H. de (1878) Melanges orthopterologiques. VIe fascicule. Gryllides (2 eme partie). Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 25, 369 - 702."]}

Agnotecous nekando Robillard n. sp. (Figs 1 B, F���H) Type material. Holotype male: New Caledonia. Grande Terre, Mt. Nekando, 27.V. 1914 (1918 ��� 87), P.D. Montague, New Caledonia Exped (BMNH). Type locality. New Caledonia, Grande Terre, Mount Nekando. Etymology. Named after the type locality. Distribution. New Caledonia, Grande Terre, Province Sud, Mont Nekando. Diagnosis. Species small, close to A. chopardi and A. humboldti n. sp. from which it differs by the shape of the pseudepiphallus in male genitalia. Description. Size small for the genus. Colouration little contrasted, mostly light brown. Head dorsum with 6 brown longitudinal bands. Fastigium setose, brown with dark spots. Scapes light brown with dark brown patterns anteriorly, antennae dark brown with yellow rings. Cheeks black dorsally, dark brown ventrally. Face homogeneously brown, the dorsal limit with fastigium black. Mouthparts and palpi dark brown. Pronotum: Dorsal disk almost homogeneously brown. Lateral lobes black. Legs: FI, FII yellow brown with faint dark brown spots; TI, TII yellow brown with 2 dark brown rings; tarsomeres yellow brown, their distal part black. Legs III almost homogeneously brown; tarsomeres III- 1 dark brown with a yellow brown ring. TIII serrulation: 6 inner and 10 outer spines above spurs; 7 inner and 9 outer spines between spurs. Tarsomeres III- 1: 1 spine on dorsal inner edges; 3 spines on dorsal outer edges; 1 outer lateral spine. Abdomen and cerci almost homogeneously brown. Male: FW not reaching abdomen mid length. Dorsal field clearly shorter than lateral field. FW colouration: Cells and veins homogeneously brown, except MP, dark brown, and a yellowish sclerotized area around 1 A and posterior to the plectrum; lateral field homogeneously brown. FW venation (Fig. 1 B): CuP absent; diagonal strong; harp triangular, flat, with 2 complete transverse veins, the anterior one faint; c 1 cell wide, partly crossed by a transverse vein; D alignment and mirror (d 1) little differentiated. Apical field restricted to one cell in E alignment (e 1). Lateral field: Latero-dorsal angle made by MP; distal part of MP weak, fusion with MA not visible; R bifurcated once at mid length; ventral part of lateral field with 7 longitudinal veins. Male genitalia (Figs 1 F���H): Very similar to that of A. chopardi. Dorsal face of pseudepiphallic sclerite with lateral crests forming a dorsal furrow; median process wider than in A. chopardi and A. humboldti n sp., spoon-shaped, with 4 wide apical teeth. Pseudepiphallic apex with ventral crest very close together. Female: unkwown. Juvenile: unkwown. Measurements. See Table 2. Habitat and life history traits. unknown. Behaviour. unknown. Calling song. unknown. TABLE 2. Measurements of Agnotecous nekando n. sp. PronL PronW FWL FWW FIIIL FIIIW TIIIL OL Male holotype 3.2 4.1 3.5 2.3 11.4 3.9 9.8, Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 22-23, DOI: 10.5281/zenodo.196996

Agnotecous pinsula Robillard n. sp. (Figs 6���7) Type material. Holotype male: New Caledonia. ��le des Pins, sentier reliant Baie d���Oro et Upi, for��t s��che littorale sur calcaire, 22 �� 35 ' 21.7 ''S 167 �� 31 ' 19.4 ''E, 06.V. 2009, T. Robillard (TR 219), nuit, liti��re, enregistrement appel Take 43 (MNHN-ENSIF 1327). Allotype female: same locality, collector and date as HT, 1 �� (TR 215), nuit, liti��re (MNHN-ENSIF 111). Paratypes (43): same locality, collector and date as HT, 13 (TR 209), jour, liti��re, comportement furtif et chant de cour (MNHN-ENSIF 2625); 13 (TR 218), nuit, liti��re sur sentier, enregistrement appel Take 35 (MNHN-ENSIF 2624); same locality and date as HT, R. Nattier, 13 (RN 187), nuit, liti��re (MNHN-ENSIF 2626); 13 (RN 188), nuit, liti��re, enregistrement appel File 67 (MNHN- ENSIF 2627). Type locality. New Caledonia, ��le des Pins, sentier reliant Baie d���Oro et Upi, 22 �� 35 ' 21.7 ''S 167 �� 31 ' 19.4 ''E. Etymology. Named after the type locality. Other material examined. New Caledonia. ��le des Pins, sentier reliant Baie d���Oro et Upi, for��t s��che littorale sur calcaire, 22 �� 35 ' 21.7 ''S 167 �� 31 ' 19.4 ''E, 06.V. 2009, T. Robillard, 1 juvenile (TR 214), nuit, liti��re. Distribution. New Caledonia, ��le des Pins. Diagnosis. Species small, very close to A. brachypterus from which it differs by the shape the pseudepiphallus in male genitalia and by the calling song, male and female FW venation, female copulatory papilla and general colouration similar to A. brachypterus. Description. Size small for the genus. Colouration contrasted mostly dark brown and orange brown. Head dorsum with 6 brown longitudinal bands more or less pronounced; dorsal eye margins yellow. Fastigium setose, quite homogeneously brown. Scapes yellow brown with dark brown patterns anteriorly, antennae yellow basally then black with yellow rings. Cheeks black, except a yellow band posterior to eyes and 2 ventral yellow lines and one yellow spot below the eyes. Face dark brown to black, with a median yellow line, sometimes very faint. Mouthparts yellow or whitish, more or less mottled with black. Palpi yellow with black rings. Pronotum: Dorsal disk homogeneously orange brown to dark brown, sometimes with darker spots, the posterior margin dark brown. Lateral lobes black. Legs I, II generally orange brown, femora mottled with dark brown, tibiae with dark brown or black rings; distal parts of tarsomeres I- 1 and I- 3 dark brown. FIII, TIII almost homogeneous orange brown, distal part of tarsomeres III- 1 and III- 3 and dark brown. TIII serrrulation: 4���6 (m= 4.8, n= 6) inner and 7���11 (m= 9, n= 6) outer spines above spurs; 2���5 (m= 4.5, n= 6) inner and 5���7 (m= 6.2, n= 6) outer spines between spurs. Tarsomeres III- 1: 3���4 (m= 3.3, n= 6) spines on dorsal outer edges; 1 (n= 6) outer lateral spine. Abdomen homogeneously dark brown dorsally; sternites yellowish brown. Cerci brown with thin dark brown rings. Male: FW setose, not reaching abdomen mid length. Dorsal field shorter than lateral field. FW colouration: Cells and veins homogeneously brown, a dark area around basal parts of veins 3 A and 4 A; lateral field with orange longitudinal veins. FW venation (Fig. 6 A) very close to that of A. brachypterus: CuP absent; harp triangular, raised at posterior angle, crossed by one incomplete faint transverse vein; CuA weak basally, stronger apically; c 1 cell not crossed by a transverse vein, its posterior limit unclear; D alignment and mirror (d 1) well differentiated, but mirror not rounded. Apical field restricted to a few cells in E alignment. Stridulatory file: 142���162 teeth (m= 152, n= 2). Lateral field: latero-dorsal angle made by MP; distal part of MP weak, fused to MA; R bifurcated once, strong until distal third, then faint and fused apically with MA; ventral part of lateral field with 5 longitudinal veins. Male genitalia (Figs 6 F���H): Close to that of A. yahoue Otte, 1987 and A. brachypterus. Pseudepiphallus widened preapically, apex with a short median process, but not in the same plane as the rest of the sclerite as in A. brachypterus and A. yahoue. Pseudepiphallic lophi forming lateral crests ventrally. Female: FWs very short and not overlapping (Fig. 6 B), posterior margin sinuate. Venation very close to that of A. brachypterus, with 5 main dorsal longitudinal veins, the second external one faint and bifurcate apically, and 3 parallel longitudinal veins in lateral field. Ovipositor slightly shorter than FIII. Female genitalia (Figs 6 C���E): Copulatory papilla triangular; plicate ventrally anterior to apex; basal sclerite with a median dorsal expansion. Juvenile: Same contrasted colour pattern as adults. Variation: Fore and median femora homogeneously brown in male paratype TR 209. Measurements. See Table 6. Habitat and life history traits. A. pinsula lives discreetly in forested areas in the leaf litter. Males call at night from the top of the leaf litter and mating occurs at night and day within the leaf litter. Behaviour. Mating behaviour not observed. Calling song (Fig. 7): At 19.7 ��C, the calling song of A. pinsula consists of a succession of echemes of 3 �� 0.1 s emitted every 20.5 �� 1.3 s, with a duty cycle of 6.7 %; each echeme is made of 128 �� 5 syllables, with the following characteristics: syllable duration= 14.5 �� 2.8 ms; syllable period = 24 �� 3 ms; syllable duty cycle = 60.6 %; the dominant frequency is 17 �� 1.7 kHz and corresponds to the second frequency peak of the song. The field work in New Caledonia was done in collaboration with Philippe Grandcolas (CNRS), Herv�� Jourdan (IRD) and Franck Muller and was funded by the ANR project BIONEOCAL (P. Grandcolas). We are grateful to Direction du D��veloppement Economique et de l���Environnement, Province Nord and Province Sud, for permitting field work in New Caledonia. We thank Simon Poulain (CNRS) and Guy Lecorvec (MNHN) for preparation of the specimens. We also thank Geoff Monteith (QMF) for loan of New Caledonian material from the Queensland Museum, and Judith Marshall and George Beccaloni (BMNH), for their help during the study of Eneopterinae crickets in the Natural History Museum, London, funded by the SYNTHESYS European program (GB-TAF- 531); and G.B. Monteith (QMF) for loan of specimens from the collections of Brisbane., Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 31-34, DOI: 10.5281/zenodo.196996, {"references":["Otte, D., Alexander, R. D. & Cade, W. (1987) The crickets of New Caledonia (Gryllidae). Proceedings of the Academy of Natural Science of Philadelphia, 139, 375 - 457."]}

Agnotecous minoris Robillard n. sp. (Fig. 4) Type material. Holotype male: New Caledonia. Grande Terre, Base du Mont Mou, for��t sur pente 22 ��04��� 30 ������S 165 �� 19 ��� 52 ������E, 390 m (Pt 29), (TR 266), 16.V. 2008, jour, liti��re, T. Robillard & F. Muller (MNHN-ENSIF 1389). Type locality. New Caledonia, Grande Terre, Base du Mont Mou. Etymology. Named after the small size of the species. Distribution. New Caledonia, Grande Terre, Province Sud, Mont Mou. Diagnosis. Species small, close to A. brachypterus from which it differs by its smaller size, details of colouration and by the shape of the pseudepiphallus in male genitalia. Description. Size very small for the genus. Colouration contrasted, quite dark with lighter areas on dorsum. Head dorsum with 6 brown longitudinal bands without yellow brown background; dorsal eye margins yellow. Fastigium yellow brown with median black spots. Scapes yellow brown with dark brown patterns anteriorly, antennae light brown basally then dark brown with yellow rings. Cheeks black dorsally, dark brown ventrally, with a whitish spot bellow eyes. Face homogeneously dark brown, the dorsal limit with fastigium black. Mouthparts yellow mottled with black, brown and whitish. Palpi black with a yellow brown ring on the 5 th segment. Pronotum: Dorsal disk cream except posterior margin dark brown. Lateral lobes black. FI, FII black with few yellow brown spots; knees yellow brown. TI, TII black with a faint yellowish ring; tarsomeres yellow brown, their distal part black. Legs III almost homogeneously brown. TIII serrulation: 5 inner and 9 outer spines above spurs; 5 inner and 7 outer spines between spurs. Tarsomeres III- 1: 1 spine on dorsal inner edges, 3 spines on dorsal outer edges; 1 outer lateral spine. Abdomen homogeneously dark brown dorsally; lateral parts of tergites mottled with yellow brown; sternites homogeneously brown. Cerci yellow brown with dark brown spots. Male: FW not reaching abdomen mid length. Dorsal field shorter than lateral field. FW colouration: Cells and veins homogeneously brown, a yellow brown sclerotized area around 1 A and posterior to the plectrum; lateral field with orange longitudinal veins and dark brown transverse veins; few whitish cells between R and its bifurcations. FW venation (Fig. 4 A): CuP absent, harp triangular, flat, without complete transverse vein; c 1 cell wide, not crossed by a transverse vein; D alignment and mirror (d 1) well differentiated, but mirror not rounded. Apical field restricted to a few cells in E alignment (e 1 -e 3) and one cell in F alignment. Lateral field: latero-dorsal angle made by MP; distal part of MP weak, fused to MA; R bifurcated twice, strong until distal third, then faint and fused apically with MA; ventral part of lateral field with 6 longitudinal veins. Male genitalia (Figs 4 B, C): Close to that of A. brachypterus Pseudepiphallus not widened preapically, apex with a short, flat median process. Pseudepiphallic lophi forming lateral crests ventrally. Female: unkwown. Juvenile: unkwown. Measurements. See Table 4. Habitat and life history traits. A. minoris lives in forested areas in the leaf litter. Behaviour. Mating behaviour not observed. Calling song. unknown. PronL PronW FWL FWW FIIIL FIIIW TIIIL OL Male holotype 2.7 3.9 3.3 2.9 10.4 3.6 8.9, Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 27-28, DOI: 10.5281/zenodo.196996

Agnotecous petchekara Desutter-Grandcolas n. sp. (Fig. 5) Type material. Holotype. New Caledonia. 11473, 21�� 34 ���S 166 ��07���E, Col de Petchekara, S. end, 13, 22.XI. 2003 ��� 28.I. 2004, G. Monteith, flight intercept. Trap (MNHN-ENSIF 2681). Allotype: Same locality, collector and date as holotype, 1 ��, 11473 (MNHN-ENSIF 2682). Paratypes (33, 3��). Same locality, collector and date as holotype: 13, 11473, (MNHN-ENSIF 2683), 13, 2��, 11473 (QMF); 13, 11474 (QMF), 1 ��, 11474 (MNHN-ENSIF 2684). Type locality. New Caledonia, col de Petchekara (21 �� 34 ���S 166 ��07���E). Etymology. Named after the type locality. Distribution. New Caledonia, Grande Terre, Province Sud, col de Petchekara. Diagnosis. Stout and large species, close to A. azurensis Desutter-Grandcolas, 2006 by its male genitalia and FW venation. General colouration light brown, with legs I and II spotted yellow and brown. Cheeks light yellow along lower margin with a large black rounded spot behind the eye. Male FW dorsal field well shorter than lateral field; venation with few, large cells. Male genitalia twice as large as those of A. azurensis; distal margin of pseudepiphallic sclerite straight or slightly concave, and without dorsal lobes; pseudepiphallus almost flat in lateral view, the lophi only slightly raised dorsally; pseudepiphallic sclerite disto-lateral margins almost parallel, not strongly divergent as in A. azurensis. Female: FWs not overlapping, separated by a distance smaller than their own length, the dorsal field margin sinuated laterally; ovipositor longer than FIII. Description. Size big, body stout. Head dorsum light brown with variable yellow and dark brown lines, and many short golden setae; a short dark, transverse line between the fastigium basis and the eyes, extended along the inner margins of the eyes to the occiput; a short and thick yellow line behind each eye between this dark line and the cheek; 3 additional short and wide dark lines on the occiput; fastigium with a dark line behind median ocellus, yellow laterally between the median ocellus and each lateral ocellus. Median ocellus circled with dark brown. Face dark brown, shining, with sparse golden setae, except in a large area below each eye, covered with many short setae and not shining; a yellow spot or thin line under median ocellus; 4 additional yellowish spots, 2 above tentorium insertion points and 2 above the epistemal suture; a large yellow spot under the lower angle of each eye. Cheeks light yellow along their lower margins; with a wide black brown fleck along the distal margins of the eyes; a thin and concave yellow line within the dark fleck. Maxillary palpi largely yellow; joint 3 brown apically and distally; joint 4 yellow only laterally; joint 5 brown basally, ventrally and apically, only a dorsal fleck and the sides yellow. Scapes yellowish with brown spots on anterior and posterior sides. Antennae brown with small yellow rings made of 1 or 2 articles, their bases lighter. Pronotum dorsal disc light brown, including anterior and posterior margins, mottled with brown; a very faint medio-longitudinal light line. Lateral lobes dark brown, including the margins. Coxae I black brown with a large yellow spot. Coxae II yellow, the outer spine and 2 spots along the basal margin black. FI, FII yellow with brown spots, including a row of brown spots on the ventral margins; brown longitudinal lines on cephalic and caudal sides. TI, TII dark brown with two yellow rings, one distal and one at mid length, and a basal, dorsal yellow spot or ring. FIII with many regular oblique dark lines separated with yellow; ventral margins with brown spots. TIII brown, the spurs yellowish; spurs and spines with brown apex. TIII serrulation: 4���7 (m= 5.7, n= 8) inner and 9���14 (m= 11.1, n= 8) outer spines above spurs; 5���8 (m= 6.6, n= 8) inner and 7���9 (m= 8.3, n= 8) outer spines between spurs. Tarsomeres III- 1: 3���7 (m= 4.5, n= 8) spines on dorsal outer edge; 1���2 (mean= 1.3, n= 8) spine on inner edge, in addition to apical spines; 1 outer lateral spine at one fourth of tarsomere length. Tarsi brown, the bases of tarsomeres 1 and 3 yellow. Tergites light brown, with a short medio-longitudinal yellow line in distal half. Cerci yellow mottled with brown. Male: FW proportion and venation as in A. azurensis, with the dorsal field well shorter than the lateral field (Desutter-Grandcolas & Robillard 2006: Fig. 2 B). FW venation (Fig. 5 A): CuA 1 and CuA 2 fused at base; mirror smaller than c 2 cell; chords conspicuous; harp with only one transverse vein not reaching CuA; CuA very faint compared to media veins. Apical field restricted to few alignments and cells. Stridulatory file with 170 teeth (n= 1). FW lateral field not reaching tergite 3 distal margin on body dorsum. Lateral field venation: R bifurcated most often once, rarely twice, over its whole length; 6���7 additional longitudinal, parallel veins (mean 6.1, n= 4). FW yellowish brown, the veins brown, except yellowish file and part of anal veins between the file and the chords; a yellowish spot at the base of MP and CuA; lateral field light brown, the longitudinal veins ochre brown, the transverse veins brown. Subgenital plate long, low in lateral view and acuminate; glandular area on dorsal margin mid length highly protruding. Male genitalia: Large. Very similar to that of A. brachypterus (see Desutter-Grandcolas & Robillard 2006: Fig. 6 F���H) and A. azurensis (see Desutter-Grandcolas & Robillard 2006: Fig. 6 C), but the distal margin of pseudepiphallic sclerite only sightly concave with almost straight middle part (contra A. azurensis), not bisinuated (contra A. brachypterus) (Figs 5 C, D); anterior margin of the arc only slightly bisinuate (it is strongly bisinuate in both A. azurensis and A. brachypterus). Female: FWs short, separated by a distance smaller than their own length; venation made of strong longitudinal parallel veins: 5 ��� 7 veins in dorsal field, in addition to a fainter, irregularly bifurcated vein along the limit with lateral field, and 3 ��� 4 longitudinal parallel veins in the lateral field, in addition to the vein separating the dorsal and lateral fields (Fig. 5 B); very faint transverse veins on dorsal field; as in A. brachypterus, dorsal field sinuated close to lateral field, and lateral field slightly acuminate dorso-distally. FW colouration light brown, the veins brown or yellowish brown. Tergites abundantly covered with short brown and white setae; light brown mottled with brown, darker along distal margin. Subgenital plate light brown with a median brown fleck. Ovipositor longer than FIII, much longer than in A. brachypterus (see above). Female genitalia: Copulatory papilla having the shape of a low sclerotized half-ring and a more or less elongated and hardly sclerotized distal lobe where the spermathecal duct opens (Fig. 5 E���H). Juvenile: unkwown. Measurements. See Table 5. Habitat and life history traits. Specimens collected in interception traps in the lower layers of vegetation and near leaf litter in the forest (G. Monteith pers. comm.), so Agnotecous petchekara is likely to live in the leaf litter within forest. Behaviour. unknown. Calling song. unknown. PronL PronW FWL FWW FIIIL FIIIW TIIIL OL Male holotype 3.7 6 - 2.8 13 4.9 10.1 - Males (n= 3) 3.6���4.1 5.6���6.3 4���4.7 3.1���3.2 12.4���14.2 4.6���5.3 9. 4 ��� 1 0. 9 - (Mean) (3.9) (6.1) (4.4) 3 (13.5) (5) (10.4) - Female allotype 4.2 6.5 1.7 - 14.3 5.6 11.4 17.2 Females (n= 3) 3.9���4.4 5. 9 ��� 6. 6 1. 6 ��� 1. 9 - 1 3. 9 ��� 1 5 5. 1 ��� 5. 3 1 0. 7 ��� 1 2. 4 1 5. 9 ��� 1 8. 9 (Mean) (4.1) (6.3) (1.7) - (14.3) (5.2) (11.3) (17.1), Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 29-31, DOI: 10.5281/zenodo.196996, {"references":["Desutter-Grandcolas, L. & Robillard, T. (2006) Phylogenetic systematics and evolution of Agnotecous in New Caledonia (Orthoptera: Grylloidea, Eneopteridae). Systematic Entomology, 31, 65 - 92."]}

Agnotecous brachypterus pocquensis Desutter-Grandcolas n. ssp. (Fig. 2) Type material. Holotype: New Caledonia. Station fruiti��re de Pocquereux, for��t galerie en bord de rivi��re, 21 �� 44 ���03"S 165 �� 53 ��� 51 "E, 37 m (Pt 19), 13 (TR 115), liti��re, accouplement film�� DV-NC 1, enregistrement appel TR take 70, 7.V. 2008, T. Robillard & F. Muller (MNHN-ENSIF 2651). Allotype: same locality, collector and date as HT, nuit 1 �� (TR 116), liti��re (MNHN-ENSIF 2649), male holotype and female allotype observed mating. Paratypes (153 6 ��): same locality, collector and date as HT, 13 (TR 118), chant sur fleur morte dans liti��re (��chant. bot. / carnet), enregistrement appel TR take 71 (MNHN-ENSIF 2662); 13 (FM 157) (MNHN- ENSIF 2663); 13 (TR 92), liti��re (MNHN-ENSIF 2666); 1 3 (TR 107), liti��re, 1 �� (TR 108), enregistrement appel/cour TR take 64, 65 (MNHN-ENSIF2665, 2650); 13 (TR 111), liti��re, enregistrement appel TR take 68 (MNHN-ENSIF 2667); 13 (TR 109), liti��re, enregistrement appel TR take 66 (MNHN-ENSIF 2664); 13 (FM 120), 1 �� (FM 119), liti��re, en couple, film��s et photographi��s (MNHN-ENSIF2669, 2651); 13 (TR 94), liti��re (MNHN-ENSIF 2668); 1 �� (TR 112), liti��re (MNHN-ENSIF 2653); 1 �� (TR 117), sur tronc h= 2.5 m, photo TR (MNHN-ENSIF 2652). Same locality as the holotype, 2.V. 2009, nuit, T. Robillard: 23 (TR 239, 241), 2 �� (TR 233, 236), liti��re; 13 (TR 171), liti��re, enregistrement appel Take 22 (MNHN-ENSIF2670, 2671, 2654, 2655, 2672); 13 (TR 172), enregistrement appel Take 23; 13 (TR 173), enregistrement appel Take 24 (MNHN-ENSIF2673, 2674). Same locality as the holotype, 2.V. 2009, nuit, R. Nattier: 13 (RN 162), liti��re, enregistrement appel file 47; 13 (RN 163), liti��re, enregistrement appel file 50 (MNHN-ENSIF 2675-2676). Type locality. New Caledonia. Station fruiti��re de Pocquereux, for��t galerie en bord de rivi��re, 21 �� 44 ���03"S 165 �� 53 ��� 51 "E, 37 m. Other material examined. New Caledonia. Same locality, collector and date as HT, 13, souche ��levage F0 male n�� 1, enregistrement appel (MNHN-ENSIF 2679); 13, nuit, F0 ��levage; 13 (TR 114), liti��re, enregistrement appel (?) TR take 70 (MNHN-ENSIF 2680); 3 ��, souche ��levage F0; 1 �� (TR 93), liti��re 1 3, 2 juveniles (F0 ��levage), 4 juveniles (TR 88-91), liti��re. Same locality as the holotype, 2.V. 2009, nuit, liti��re, T. Robillard: 4 �� (TR 234, TR 235, TR 237, TR 240). Distribution. New Caledonia, Grande Terre, Province Sud, Station fruiti��re de Pocquereux, for��t galerie en bord de rivi��re. Diagnosis. Very close to the specimens of A. brachypterus Gorochov, 1986 from Sarramea, from which it differs by the shape of pseudepiphallus apex in male genitalia (more narrow with less divergent lateral margins, and shorter disto-lateral angles, Fig. 2 C), a higher number of stridulatory teeth in males, smaller and largely separated FWs and a shorter ovipositor in females. Description. Size variable. Colouration close to that of nominal A. brachypterus, but usually less contrasted: wide yellow line behind the eyes present, but occiput darker, with fewer and thinner yellow lines; head dorsum dark brown, the outer triangles of the fastigium between median and each lateral ocellus lighter. Face dark brown, with a variable thin, light line under median ocellus and two small yellowish spots close to epistemal suture. Cheeks dark brown, variously mottled with yellow. Mid third of the eyes black brown, the upper third lighter with a darker line close to upper margin, and the lower third brownish separated from the black brown band by a lighter line. Maxillary palpi dark brown, with a dorso-lateral yellow spot on distal half of 5 th joint (in A. b. brachypterus they are entirely dark brown, the 5 th joint being lighter on distal half). FI, FII black with a more or less conspicuous yellow ring, subapical on FI, distal and wider on FII (in A. b. brachypterus, only the distal part of FII is more or less yellowish). TIII serrulation: 5���9 inner (m= 6.7, n= 10) and 9���13 outer (m= 11.1, n= 10) spines above spurs; 5���7 inner (m= 5.3, n= 10) and 6���9 outer (m= 7.5, n= 10) spines between spurs. Tarsomeres III- 1: 1 spine (n= 5) on inner edges and 3���5 spines (m= 3.8, n= 5) on dorsal outer edges in males, in addition to apical spines (females: 1���3 spines (m= 1.7, n= 5) on inner edges and 3���4 spines (m= 3.9, n= 5) on dorsal outer edges); 1���2 outer lateral spines at one fourth of tarsomere length. TI black brown to black with a conspicuous yellow ring at mid length; their base and apex more or less mottled with yellow. TII black brown to black, with a transverse yellow spot on dorsal side close to the knee, and a variable yellow, distal ring (sometimes absent). FIII light brown, darker on dorsal inner and outer halves; inner and outer ventral margins light yellow with a row of brown spots. TIII light brown with a dorsal yellow spot near the knee. Tarsi I, II: tarsomeres 1 and 3 black brown with yellow base, tarsomeres 2 black brown. Tarsi III: tarsomeres black brown, marked with yellow at base or at mid length. Spurs yellow with black apex. Suranal plate and dorsal part of cerci basis yellowish, the ventral part of cerci basis brown; cerci otherwise yellowish with brown spots. Male: FW proportion and venation as in A. b. brachypterus, with the dorsal field only slightly shorter than the lateral field (Desutter-Grandcolas & Robillard 2006: Fig. 2 C), the dorsal field not reaching tergite 3 distal margin. FW venation (Fig. 2 A): CuA 1 and CuA 2 fused at base; mirror and c 2 cell almost equal in size; chords conspicuous; harp with two transverse veins fused at mid length. Apical field restricted to a few cells in E alignment. Stridulatory file with 160 ��� 183 teeth (mean 169, n= 5). Lateral field venation: R bifurcated twice over its whole length; 4���7 additional longitudinal, parallel veins (mean 5.4, n= 8). FW brown, the file and plectrum somewhat yellowish; lateral field light yellow, the longitudinal veins light ochre, the transverse veins brown. Subgenital plate very long and quite low in lateral view; glandular area on dorsal margin mid length highly protruding. Male genitalia: Very similar to that of A. b. brachypterus (see Desutter-Grandcolas & Robillard 2006: Fig. 6 F���H), especially for the distal margin of pseudepiphallic sclerite (bisinuated, with one median and two lateral processes almost equal in length), but characterized by the smaller width of pseudepiphallic apex, not as divergent as in A. b. brachypterus, the shape of pseudepiphallic sclerite seen in dorsal view (lateral margins more strongly convergent in distal half, and the relative size of pseudepiphallic distal lobes (lateral ones somewhat shorter) (Figs 2 C, D). Female: FWs shorter than in A. brachypterus, and separated by a distance greater than their own length; venation made of strong longitudinal parallel veins, with 4 ��� 5 veins in dorsal field, in addition to a fainter, irregularly bifurcated vein along the limit with lateral field, and 3 ��� 4 longitudinal parallel veins in the lateral field (Fig. 2 B); very faint transverse veins on dorsal field. FW colouration more or less whitish brown, the veins brown, the vein separating the dorsal and the lateral field yellowish. Tergites abundantly covered with short setae; brownish mottled with black and yellow; a short longitudinal, yellow line along tergite mid line. Subgenital plate light yellow with a longitudinal, median brown line growing wider toward posterior margin (Fig. 2 E). Ovipositor slightly shorter than FIII; much shorter than in A. brachypterus (10.5���12.3, m= 11.1, n= 5, against 12.1���13.3, m= 12.7, n= 4 in A. brachypterus from Sarramea, and 12.8 for the female type according to Gorochov, 1986). Female genitalia: Copulatory papilla very similar to that of A. brachypterus (compare Figs 2 F, G with Desutter-Grandcolas & Robillard 2006, Fig. 3 R). Measurements. See Table 3. Variation. Head dorsum variously marbled with yellowish brown, but sometimes entirely dark brown except for yellow lines on occiput; yellow patterns on the face variously surrounded, from homogeneous dark brown to yellowish brown. In darker specimens, cheeks black brown hardly mottled with yellow under the eyes; in lighter specimens, cheeks largely bordered with yellow on lower margin. Some specimens present an additional yellow spot on TII at about mid length dorsally. In male FW lateral field sometimes brown over part of its basal third, the veins being always light ochre over their whole length; R rarely bifurcated only once over its whole length. Colouration of female subgenital plate not always conspicuous (as in specimens TR 117 and TR 112). According to the general colouration of the specimens, FIII variously coloured, from light yellowish brown to dark brown with light yellow ventral part; in the same way, lateral parts of the fastigium between the median and the lateral ocelli more or less of lighter colouration (note: all the specimens collected in 2009 are very dark in colouration, without yellow ring on TII). PronL PronW FWL FWW FIIIL FIIIW TIIIL OL Male holotype 2.9 4.7 3.6 2.9 11.2 4 8.7 Males (n= 4) 2.8���3.1 4.5���5.1 3.1���3.5 2.8���3 10.5���11.9 3.4���4.1 8.4���10.1 (Mean) (2.9) (4.8) (3.2) (2.9) (10.8) (3.7) (8.9) Female allotype 3.2 5.2 0.7 - 12.3 4.3 10.5 11.5 Females (n= 4) 2.7���3.1 4.5���5.3 0.4 ��� 0.9 - 10.5���12.8 3.2���4.6 8.6���10.5 10.5���12.3 (Mean) (2.9) (5) (0.8) - (11.9) (4) (9.7) (11.1) Habitat and life history traits. A. brachypterus pocquensis lives in secondary forest, in the leaf litter. Males emit their calling song at night, perched on prominent position in the leaf litter and sometimes on tree trunks. Behaviour. Mating behaviour briefly observed in the field and includes multiple copulations and intercopulatory songs (T. Robillard & F. Muller, pers. obs.). Calling song (Fig. 3): At 22 ��C, the calling song of A. brachypterus pocquensis (TR 111: MNHN- ENSIF 2267) consists of a succession of echemes of 2.3 �� 0.4 s emitted every 12.9 �� 0.7 s, with a duty cycle of 18.8 %; each echeme is made of 84 �� 15 syllables, with the following characteristics: syllable duration= 12.8 �� 1.7 ms; syllable period = 27.6 �� 1.5 ms; syllable duty cycle = 46.9 %; the dominant frequency is 19.2 �� 1.8 kHz and corresponds to the second frequency peak of the song. Remark. Within A. brachypterus, several specimens could not be attributed to one subspecies of the other. One male from the Station fruiti��re de Pocquereux (TR 110, liti��re, enregistrement appel TR take 67, T. Robillard & F. Muller) agrees A. b. pocquensis n. ssp. for colouration, but its genitalia are similar to nominal A. brachypterus from Sarramea. Another male from the same locality (TR 113, liti��re, h= 5 cm, enregistrement appel TR take 69) agrees A. b. pocquensis n. ssp. for colouration and shape of pseudepiphallus, but the apex of pseudepiphallus is similar to A. brachypterus., Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 23-27, DOI: 10.5281/zenodo.196996, {"references":["Gorochov, A. V. (1986) New and little known crickets (Orthoptera, Gryllidae) from Australia and Oceania. Revue d'Entomologie de l'URSS, 65, 692 - 708 (in ru).","Desutter-Grandcolas, L. & Robillard, T. (2006) Phylogenetic systematics and evolution of Agnotecous in New Caledonia (Orthoptera: Grylloidea, Eneopteridae). Systematic Entomology, 31, 65 - 92."]}

International audience