Coelophora inaequalis Fabricius

Coelophora inaequalis (Fabricius) Coccinella inaequalis Fabricius, 1775: 80. Coccinella circularis Thunberg, 1820: 365 ���synonymized by Pope 1987: 54. Lemnia desolata Mulsant, 1850: 387 ���synonymized by Houston 1979: 49. Coccinella bicruciata Montrouzier, 1861: 304 (pars)���synonymized by Fauvel 1903: 328. Verania artensis Montrouzier, 1861: 305 ���synonymized by Chazeau 1978: 59. Coelophora ripponi Crotch, 1874: 156 ���synonymized by Iablokoff-Khnzorian 1979: 61. Coelophora mastersi Blackburn, 1892: 238 ���synonymized by Houston 1979: 49. Coelophora inaequalis: Fauvel 1903: 327. At first, this species was misidentified by Montrouzier who described it under two endemic species: Coccinella bicruciata and Verania artensis. According to Fauvel (1903), who examined the type series, Coelophora bicruciata (Montrouzier) was a mixture of Coelophora inaequalis and Micraspis frenata (Erichson) specimens. Verania artensis was subsequently synonymized by Chazeau (1978). Coelophora inaequalis is widespread from both coasts to high altitude (> 1,000m) on Grande Terre but also on Loyalty Islands and Isle of Pines. This species is widely distributed across Asia and the Pacific region (Australia, Papua New Guinea, Vanuatu, Solomon Islands, Fiji Islands, Hawaii, Micronesia (Bonin, Marianna, Caroline, Wake, Marshall, Kiribati), Cook Islands, Samoa, Wallis and Futuna, French Polynesia); but also has been recorded from eastern North America. This is a polyphagous predator of hemipterous insects (Aphididae, Coccomorpha) from natural habitats to agricultural lands and disturbed areas. According to its geographic range and feeding biology, humans have probably spread this species during the first settlements. It is a well-known and largely promoted biocontrol agent against aphids (Wang & Tsai 2001). Material examined. NEW CALEDONIA: Bac de la Oua����me, 17.iv. 1986 (J. Chazeau rec.) (ONNC); Bourail, 10.iii. 1986 (J. Chazeau rec.) (ONNC); from La Foa to Canala, ii. 1907 (Le Rat rec.) (ONNC); Goro, 12.viii. 1985 (J. Chazeau rec.) (ONNC); Hiengh��ne, 18.xi. 1958 (M. Ogata rec.) (ONNC); Houa��lou, 7.iii. 1982 (J. Chazeau rec.) (ONNC); Isle of Pines, airport, 24.viii. 1988 (J. Chazeau rec.) (ONNC); Iles Loyaut��, Lifou; Iles Loyaut��, Mar��; Koumac, 10.iii. 1986 (J. Chazeau rec.) (ONNC); Koumac, 26.viii. 1986 (J. Chazeau rec.) (ONNC); Koumac, 27.viii. 1986 (J. Chazeau rec.) (ONNC); Mt Do, 1000 m, 4.xi. 1992 (J. Chazeau & D. Webb rec.) (ONNC); Noum��a, Anse Vata, 25.ii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Anse Vata, 10.iii. 1977 (J. Chazeau rec.) (ONNC); Noum��a, Anse Vata, 3.x. 1977 (J. Chazeau rec.) (ONNC); Noum��a, IFO, 4.i. 1964; Noum��a, IFO, 13.i. 1965; Noum��a, OuenToro, vii. 1966 (J. Fainika rec.) (ONNC); Noum��a, Baie des Citrons, 27.vi. 1978 (J. Chazeau rec.) (ONNC); Noum��a, Trianon, 20.iv. 1964 (J. Gutierrez rec.) (ONNC); Noum��a, vi, vii, xi. 1955; Noum��a, ix. 1964; Pouembout, 17.ix. 1975 (J. Gutierrez rec.) (ONNC); Pindai, 6.xi. 1992 (D. Webb rec.) (ONNC); La Foa-Canala Road, i. 1958; Thio, 27.vi. 1983 (L. Bonnet de Larbogne, E. Bouy��, J. Chazeau rec.) (ONNC); Thio Ourou��, 5.i. 1982 (J. Chazeau rec.) (ONNC); Ile longue, Chesterfield islands, xi. 2012 (E. Bourguet et H. Jourdan rec.) (ONNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 10.ix. 2001 (S. Caz��res rec.) (CXMNC); La Foa, Pocquereux, 21 �� 44 ���S / 165 �� 54 ���E / 28m, 20.iv. 2001 (J. Brinon rec.) (CXMNC); La Foa, Pocquereux, 16.xi. 1997 (CXMNC); Bourail, Gouaro, 21 �� 36 ���S / 165 �� 28 ���E / 68m, 04.iv. 2001 (S. Caz��res rec.) (CXMNC); Farino, Tend��a, 21.61928 ��S / 165.70907 ��E / 403m, 29.iv. 2004 (S. Caz��res rec.) (CXMNC); Houa��lou, N��oua, 13.iv. 2013 (C. Mille rec.) (CXMNC); Pouembout chez M. Soury-Lavergne, COL /ins/ 255 -14, 01.viii. 2014 (C. Mille rec.) (CXMNC); Poya, Beaupr�� chez M. Mezdorf, 21 �� 205 ���S/ 165 ��09���E, 22.vi. 2001 (R. Guiglion rec.) (CXMNC); Mar��, La Roche, 16.iv. 2004 (C. Mille rec.) (CXMNC).
Published as part of Nattier, Romain, Jourdan, Herv��, Mille, Christian & Chazeau, Jean, 2015, An annotated checklist of the Coccinellidae (Coleoptera) from New Caledonia, pp. 301-331 in Zootaxa 4058 (3) on page 305, DOI: 10.11646/zootaxa.4058.3.1, http://zenodo.org/record/236980
{"references":["Fabricius, J. C. (1775) Systema Entomologiae Sistens Insectorvm Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibvs, Observationibvs. Vol 1. Kortius, Lipsiae, 832 pp.","Thunberg, C. P. (1820) Coleoptera Capensia, antennarum clava solida et perfoliata, collecta, recensita et descripta. Memoires de l'Academie Imperiale des Sciences de Saint Petersbourg, 7, 362 - 372.","Pope, R. D. (1987) The Coccinellidae (Coleoptera) described by CP Thunberg. Insect Systematics & Evolution, 18, 51 - 66.","Mulsant, E. (1850) Species des coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, 2, XV - 1104.","Houston, K. J. (1979) Mosaic dominance in the inheritance of the colour patterns of Coelophora inaequalis (F.) (Coleoptera: Coccinellidae). Australian Journal of Entomology, 18, 45 - 51.","Montrouzier, P. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des Iles des Pins, Art, Lifu, etc. Coleopteres (Fin). Annales de la Societe entomologique de France, 1, 265 - 306.","Fauvel, A. (1903) Faune analytique des Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie, 22, 203 - 379.","Chazeau, J. (1978) Les especes du genre Coelophora en Nouvelle Caledonie et aux Nouvelles Hebrides (Coleopteres, Coccinellidae). Cahiers ORSTOM. Serie Biologie: Entomologie Agricole, 13, 59 - 69.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, xv + 311 pp.","Iablokoff-Khnzorian, S. M. (1979) Genera der palaarktischen Coccinellini (Coleoptera, Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75.","Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia, 15, 207 - 261.","Wang, J. J. & Tsai, J. H. (2001) Development and functional response of Coelophora inaequalis (Coleoptera: Coccinellidae) feeding on brown citrus aphid, Toxoptera citricida (Homoptera: Aphididae). Agricultural and Forest Entomology, 3, 65 - 69."]}