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Hodophilus pseudoglabripes, a new species with a strong earthy odor, was found in forests dominated by Castanopsis (Fagaceae) in northern Thailand. This species is introduced with microscopic and macroscopic descriptions and colour photographs. Phylogenetic affinities were determined based on a combination of internal transcribed spacers (ITS) and large subunit (nrLSU) of the nuclear ribosomal DNA (rDNA). The new species was placed in Hodophilus, within an independent clade comprising the other two species from Asia, H. glabripes and H. indicus. This is the first report of the genus Hodophilus in Thailand. An updated key to all members of the genus Hodophilus is also provided.

Phallus aureus is described as a new species from China and is characterized by an obovate, rugose, squamulose on the immature basidiomata surface, pseudostipe yellowish, grey volva, and pale yellow at the base becoming light yellow upwards. The new species is phylogenetically located in an independent lineage. A full description, colour photographs, illustrations, and a phylogenetic tree to show the position of new species are provided.

Hodophilus pseudoglabripes S. M Tang & S.H. Li, sp. nov. (Figs. 2, 3) MycoBank: MB838312 Etymology:—‘pseudoglabripes’ refers to its similarity to H. glabripes. Holotype:— THAILAND, Chiang Mai Province, Mae Taeng District, Pa Pae Sub-district, 19°14.66’N, 98°38.61’E, 850 m a.s.l., 14 August 2020, collected by Song-Ming Tang (MFLU21-0005, holotype!; HKAS110665 isotype!). Diagnosis:— Hodophilus pseudoglabripes is similar to H. glabripes Ming Zhang, C.Q. Wang & T.H. Li, but can be distinguished by its pileus slightly striated near-margin and flocculose with age, white to the greyish yellow stipe, as well as its smaller basidiomata. Description: Basidiomata small to medium-sized. Pileus 11–25 mm diam., hemispherical at first, then convex to plano-convex, often with a slight depression at the center, surface smooth when young, becoming slightly striate near margin and flocculose with age, when wet translucently striated halfway to the disc, hygrophanous, white (1A1) to yellowish white (1A2) when young, later greyish yellow (1B4) to light orange (5A5), when dry uniformly pale at center. Pileus context up to 1.5 mm thick, white (1A1). Lamellae number 16–20, white (1A1) when young, light orange (5A5) to greyish orange (5B5) when old, 3–4 mm wide, lamellullae frequent, in (1–) 2 tiers. Stipe 34–64 × 2–3 mm, surface smooth, white (1A1), central, usually flexuous, hollow, and narrowing downwards, becoming greyish yellow (2B3) near the base, hollow when mature. Odor strong earthy. Basidiospores (103/5/2) (4.0–) 4.3–6.2 (–6.3) × (3.4–) 3.5–5.5 (–5.6) μm, av. 5.2–4.6 μm, Q (length/width) = 1.00–1.44 (–1.57), av. Q=1.14, globose to subglobose, smooth, thin-walled. Basidia 4–spored, (25.0–) 33.0–48.0 × (5.2–) 5.8–7.4 μm, av. 40.1 × 6.5 μm, clavate, hyaline. Basidioles cylindrical to narrowly clavate, obtuse, (28.6–) 30.2–44.3 (–45.2) × (3.8–) 4.2–6.3 (–6.7) μm. Pleurocystidia absent. Pileipellis a hymeniderm to epithelium, hyphal terminations vertically arranged, composed of cells arranged in one or two ranks, terminal cells near the pileus margin obovoid, obpyriform, subglobose, clavate, thin-walled, (18–) 21–51 (–67) × (–15) 16–26 (–34) μm, av. 35 × 20 μm; subterminal cells usually distinctly narrower, cylindrical, some inflated, broadly clavate, ventricose or branched, (18–) 20–46 (–50) × (6–) 8–19 (–23) μm, av. 29–13 μm. Terminal cells near the pileus center similar in shape and size to those near the pileus margin, but usually narrower, (15–) 17–34 (–40) × (–5) 7–14 (–20) μm, av. 23–11 μm, subterminal cells similar to those near the pileus margin, (6–) 7–23 (–34) × (1–) 2–5 (–10) μm, av. 16–6 μm. Caulocystidia without pigments, thin walled, usually densely clustered, flexuous or spirally coiled, clavate and regular, (37–) 33–48 (–50) × (4.7–) 5.8–7.4 (–8.9) μm. Clamp connections absent in all parts. Ecology and Distribution: Solitary, on soil, in a forest dominated by Castanopsis. Known only from the type locality in Chiang Mai Province, Northern Thailand. Additional specimen examined: Thailand, Chiang Mai Province, Wat Pah Pa Deng, Mae Taeng District, 14 Aug 2020, Feng-Ming Yu (MFLU21-0006); same location, 5 September 2020, Hong-Wei Shen (HKAS110665); Chiang Mai Province, Mae Sae Village, Pa Pae Sub-district, Mae Taeng District, 6 September 2020, Hong-Wei Shen (HKAS110666)., Published as part of Tang, Songming, Lv, Tong, He, Jun, Yu, Fengming, Luo, Hongmei & Li, Shuhong, 2023, Hodophilus pseudoglabripes (Clavariaceae, Agaricales), a new species from Thailand, pp. 219-230 in Phytotaxa 597 (3) on pages 224-226, DOI: 10.11646/phytotaxa.597.3.3, http://zenodo.org/record/7958565

Key to known Hodophilus species 1 Pileus white to yellow, vivid yellow, yellow brown, orange grey......................................................................................................2 1 Pileus greyish brown, light brown, coffee brown, hair brown, dark brown, greyish brown to brownish orange............................10 2 Stipe vivid yellow to brown...............................................................................................................................................................3 2 Stipe white to yellowish, greyish orange............................................................................................................................................5 3 Stipe brown, without any yellow colour............................................................................................................... H. hymenocephalus 3 Stipe have more or less yellow colour................................................................................................................................................4 4 Stipe colour soon changes and becomes partly to almost completely brown with age..................................................... H. praecox 4 Stipe vivid yellow and remains so even in mature basidiomata....................................................................................... H. micaceus 5 Lamellae number>20..........................................................................................................................................................................6 5 Lamellae number≤20..........................................................................................................................................................................9 6 Caulocystidia often irregularly inflated, nodulose, lobate and twisted..................................................................... H. albofloccipes 6 Caulocystidia clavate, occasionally subcapitate or obpyriform, obtuse.............................................................................................7 7 Basidiomata with a slight yam bean odor...........................................................................................................................................8 7 Basidiomata with an unpleasant, mercaptan-like odor........................................................................................... H. tenuicystidiatus 8 Pileus 15–50 mm diam.; pileipellis cells obpyrifrom, subglobous, ellipsoid, sphaero-pedunculate or broadly clavate.. H. glabripes 8 Pileus 11–25 mm diam.; pileipellis cells obpyrifrom, subglobose....................................................................... H. pseudoglabripes 9 Naphthalene odour; lamella number≤18; caulocystidia clavate or obpyriform...............................................................................11 9 Indistinct odour; lamella number=16–22; caulocystidia irregularly nodulose-lobate, mostly obtuse...................... H. phaeoxanthus 10 Pileus 6–18 mm; stipe 25–40 × 1.0–2.0 mm; basidiospores broadly ellipsoid, Q m =1.17–1.36......................................... H. pallidus 10 Pileus 2–10 mm; stipe 9–30 × 0.8–1.5 mm; basidiospores narrowly ellipsoid, Q m =1.22-1.40............................................ H. smithii 11 Basidiomata with distinct naphthalene odours.................................................................................................................................12 11 Basidiomata with weak naphthalene odours....................................................................................................................................14 12 Pileus 4–11 mm diam.; pileus colour dark blond to eye brown and stipe colour bronze to brownish orange..................... H. hesleri 12 Some pileus more than 11 mm diam.; pileus colour and stipe colour grey brown..........................................................................13 13 Lamellae number 18–26; basidiospores 5.1–5.9 × 4.2–4.7 μm, Q m =1.20–1.36.................................................................. H. foetens 13 Lamellae number 15–20; basidiospores 5.0–5.7 × 3.9–4.5 μm, Q m =1.20–1.36............................................................ H. subfoetens 14 Stipe without dots............................................................................................................................................................ H. rugulosus 14 Stipe have dots..................................................................................................................................................................................15 15 Stipe with distinct dark dots.............................................................................................................................................................16 15 Stipe never with dark dots................................................................................................................................................................17 16 Pileus 7–14 mm diam.; basidiospores 4.6–5.4 × 3.5–4.2 μm, Q m =1.21-1.39; caulocystidia 18.0–33.5 × 5.5–11.5 μm............................................................................................................................................................................................................ H. atropunctus 16 Pileus 5–22 mm diam.; basidiospores 4.8–5.4 × 3.9–4.5 μm, Q m = 1.15–1.28; caulocystidia 20.5–38.5 × 6.5–13.5 μm.......................................................................................................................................................................................................... H. variabilipes 17 Strong odour; lamellae colour pallid to pale orange........................................................................................................................18 17 Weak or indistinct odour; lamellae colour orange grey, brown, or grey brown...............................................................................20 18 Lamellae pale orange; basidiospores 4.0–5.0 × 3.0–5.0 μm, Q m =1.0–1.66........................................................................ H. indicus 18 Lamellae pale drab to darker with age.............................................................................................................................................19 19 Caulocystidia on average wider than 8 μm..................................................................................................................... H. peckianus 19 Caulocystidia on average narrower than 7 μm............................................................................................................. H. paupertinus 20 Stipe colour changing with age........................................................................................................................................................21 20 Stipe colour unchanging with age....................................................................................................................................................23 21 Stipe grey brown to dark brown with age................................................................................................................... H. cambriensis 21 Stipe yellow or light yellow to greyish yellow or dark grey............................................................................................................22 22 Basidiospores 4.1–4.7 × 3.3–3.8 μm............................................................................................................................. H. stramineus 22 Basidiospores 4.8–5.5 × 3.8–4.4 μm................................................................................................................................. H. anatinus 23 Stipe yellow; basidiospores 4.9–5.6 × 3.9–4.3 μm................................................................................................... H. subfuscescens 23 Stipe dark brown to greyish brown; basidiospores shorter or longer...............................................................................................24 24 Stipe 25–35 mm; 22–24 lamellae; basidiospores 4.4–4.9 μm long.............................................................................. H. carpathicus 24 Stipe 18–25 mm; 11–19 lamellae; basidiospores 6.3–6.5 μm long............................................................................. H. decurrentior, Published as part of Tang, Songming, Lv, Tong, He, Jun, Yu, Fengming, Luo, Hongmei & Li, Shuhong, 2023, Hodophilus pseudoglabripes (Clavariaceae, Agaricales), a new species from Thailand, pp. 219-230 in Phytotaxa 597 (3) on pages 227-228, DOI: 10.11646/phytotaxa.597.3.3, http://zenodo.org/record/7958565

Ganodermataceae was introduced by Donk (1948) which belongs to Polyporales and the latest studies indicated that it is a monophyletic group (Costa-Rezende et al. 2020). Currently, eleven genera viz. Amauroderma Murril, Amaurodermellus Costa-Rezende, Cristataspora Costa-Rezende, Foraminispora Robledo, Costa-Rezende & Drechsler-Santos, Furtadoa Costa-Rezende, Robledo & Drechsler-Santos, Ganoderma P. Karst., Haddowia Steyaert, Humphreya Steyaert, Magoderna (Murrill) Steyaert, Sanguinoderma Y.F. Sun, D.H. Costa & B.K. Cui and Tomophagus Murrill are accepted in Ganodermataceae and supported by morphology and phylogeny (Steyaert 1972; Furtado 1981; C orner 1983; Zhao and Zhang 2000; Ryvarden 2004; Thametal 2012; Costa-Rezende et al. 2017; Costa-Rezende et al. 2020; Sun et al. 2020). Ganoderma P. Karst (Ganodermataceae, Polyporales) was introduced to accommodate a laccate and stipitate fungus, Ganodermalucidum (Curtis) P. Karst (Karsten 1881). Ganoderma is characterized by double-walled basidiospores with inter-wall protuberances (Karsten 1881; Moncalvo and Ryvarden 1997). There are 462 records in the Index Fungorum (http://www.Indexfungorum.org/; accessed date: 7 October 2021) and 506 records in MycoBank (http://www.mycobank.org/; accessed date: 7 October 2021). Ganoderma is one of the most taxonomically scrutinized genera among the Ganodermataceae and even in Polyporales (Richter et al. 2015; Costa-Rezende et al. 2020). Most Ganoderma species are wood decomposers, found in all temperate and tropical regions (Pilotti et al. 2004; Cao et al. 2012; Zhou et al. 2015). Ganoderma has long been regarded as one of the most important medicinal fungi in the world (Paterson 2006); they have been used as medicine for over two millennia in China (Dai et al. 2009). Several Ganoderma species are known to be prolific sources of highly active bioactive compounds, especially polysaccharides, protein, sterols, and triterpenoids (Ahmadi and Riazipour 2007; Chan et al. 2007). These compounds are known to possess extensive therapeutic properties, such as antioxidant, antitumor, and antiviral agents, and improve sleep function (De Silva et al. 2013). Species diversity of Ganoderma is abundant in China and more than 30 species have been described (Zhao and Zhang 2000; Wang et al. 2009; Cao et al. 2012; Li et al. 2015; Xing et al. 2016; Hapuarachchi et al. 2018; Liu et al. 2019; He et al. 2019; Wu et al. 2020). Yunnan province is considered as one of the hot-spots for studying biodiversity of polypores, and some new Ganoderma species have been described (Zhao 1989; Wang et Wu 2010; Cao and Yuan 2013). During our investigation into the diversity of Ganoderma in Yunnan province, several specimens of Ganoderma were collected from central and southern Yunnan. Phylogenetic analysis showed that the seven collections formed two distinct lineages and can be recognized as new species, hence two new species, namely G.dianzhongense and G.esculentum are introduced based on morphology and phylogeny.

Phallus aureus S.M. Tang & S.H. Li, sp. nov. (FIGURE 3, 4) MycoBank: MB 844523 Etymology:—the epithet ‘ aureus ’ refers to light yellow indusium. Holotype:— CHINA, Yunnan Province, Zhaotong City, Xiaocaoba Town, 27°14.36’N, 103°52.61’E, 2000 m a.s.l., 14 October 2021, collected by Ao-cheng Ce (L7010, holotype!). Diagnosis:— Phallus aureus is distinguished from other Phullus species by an obovate, rugose, squamulose on the immature basidiomata surface, pseudostipe yellowish, grey volva, and pale yellow at the base becoming light yellow upwards. Description: Immature basidiomata globose to subglobose, 70 × 67 mm, yellowish white (2A2) to pale yellow (2A3), clear rugose on the surface, attached to the substrate by white (2A1) to yellowish grey (2B2) rhizomorphs. Exoperidium membranous; endoperidium gelatinous, hyaline. Expanded basidiomata up to 220 mm high when fresh. Receptacle 49–73 mm high, 43–72 mm broad, campanulate, pale yellow (2A3) to yellowish white (2A2), rugose. Gleba olive brown (4F5-6), mucilaginous. Pseudostipe subcylindrical, constricted at apex, enlarged downwards, 140–218 mm high when mature, 18–22/30–28/ 39–51 mm broad (apex/middle/base), pale yellow (2A3) at the base, upward becoming light yellow (3A5), spongiform, hollow. Volva globose or slightly obovate, 68–81 mm high, 50–69 mm broad, rugose, light grey (1C1) to grey (1D1). Indusium well-developed, almost touching ground, orange yellow (4B8), 113–142 mm in length, attached to the apex of pseudostipe, with polygonal to irregular meshes; meshes 2–21 mm wide, 1–4 mm thick. Rhizomorphs simple, white (2A1), 2–4 mm thick, about 35 mm long. Odour and taste unknown. Basidiospores (2.2–) 2.5–3.7 (–3.9) × (1.2–) 1.3–2.0 (–2.3) µm, Q=(1.6–) 1.8–2.0, cylindrical to long ellipsoid, hyaline, thin-walled, smooth under light microscope. Hyphae of pseudostipe and indusium thin-walled, pseudoparenchymatic, consisting of globose to subglobose or irregularly globose cells up to 21–45 µm in diam. Hyphae of volva tubular, 2–8 µm wide, thin walled, smooth. Hyphae of rhizomorphs filamentous, up to 8.0 µm in diam, thin-walled, smooth, septate, rarely branched. Habitat and distribution: Solitary or scattered on soil with decaying litter under Fargesia spathacea forest. So far known only from Yunnan Province, China. Season: July to August. Additional specimen examined: China. Yunnan Province, Zhaotong city, Xiaocaoba county, Ao Chengce 10 July (L7005); same location, Tang Songming 10 July (L7006); same location, Ao Chengce 14 August (L7011); Wang li 14 August (L7012).

Agriotypus maae sp. nov. Figs 3A, 4 Material examined. Holotype. ♀, China, Shaanxi prov., Qinling, Tiantaishan, 34.2°N, 106.8°E, 2000 m, 1998.VI.8, Du Yuzhou, No. 983376 (ZJUH). Paratypes: 1♀, China, Shaanxi prov., Zhouzhi Houzhenzi, 1990.V.24, Du Yuzhou, No. 984474 (TARI); 1♀, China, Shaanxi prov., Zhouzhi Houzhenzi, 1990.V.24, Du Yuzhou, No. 984475 (ZJUH); 1♀, China, Shaanxi prov., Foping Longcaoping, 33.5°N, 107.9°E, 1998.VI.4, Ma Yun, No. 981986 (ZJUH). Diagnosis. This new species belongs to the Agriotypus himalensis species group, and it runs to A. maculiceps Chao in Bennett’s key (2001), but differs in frons with longitudinal carina (the latter without); pronotum finely punctate-rugose (the latter reticulate-rugose); scutellum entirely black (the latter pale in apical half), spine of scutellum produced more or less 85° dorsoposteriorly (the latter 35-45°). It differentiated from other new species proposed in this study by the combination characters: ♀ vertex near eyes and mesoscutum medio-posteriorly with large reddish brown patch; frons with a longitudinal carina; mesoscutum mostly glabrous; spine of scutellum produced 85° dorsoposteriorly. Description. Female. Body length 7.4 mm; fore wing length 5.8 mm. Head. Head width 0.8 × its median length. Antennae 23-segmented, 0.5 × length of forewing. Temple behind eyes distinct narrowed in dorsal view. Transverse diameter of eye 1.0 × temple in dorsal view. Frons convex medially, with longitudinal carina. Antennal scrobe more or less shallow. Ocelli in triangle with base equal to its sides. POL: OD: OOL = 12: 11: 24. Face convex medially with coarse rugosity and dense pubescence. The area between antennal socket with a glabrous circular tubercle. Clypeus nearly pentagonal, almost equal to its maximum width, roundly convex, flat and rugose with punctuation in basal 0.6; thin in apical 0.4, without distinct rugosity, bluntly rounded on apical margin; summit of covexity of clypeus smoothly rounded in lateral view. Distance between tentorial pits 1.3 × length between a dorsal tentorial pit and eye. Malar space 1.15 × basal width of mandible. Occipital carina fine, but complete. Mesosoma. Pronotum with fine epomia, dorsally curved angulate posteriorly; lateral of pronotum glabrous, reticulate in dorsal half, dense pubescent in ventral half. Mesoscutum mostly glabrous, shiny, with fine punctulation and rugosity partly; median lobe medially and lateral lobe mostly smooth; notaulus narrow, more or less crenulated, meeting in apical 0.3. Scutellum long triangle-shaped, 2.5 × its basal width, 1.2 × length of propodeum; basal half of scutellum with lateral carinae and reticulation-rugosity, convex medially, lateral margin of scutellum paralleled in basal 2/3, tapering towards to apex, spine-like apical half, smooth; in lateral view scutellum roundly curved medially, spine-like in apical 0.4, distinctly up curved, apex tapered, produced 85° dorsoposteriorly; ecepicnemial carina strong, extending to ventral-anterior of mesopleuron; mesopleural sulcus complete; sternaulus strong. Propodeum punctate-reticulate; lateral longitudinal carinae straight and strong and paralleled with lateromedian longitudinal carinae. Wings. Fore wing with vein 1cu-a distad of M&RS, abscissa of vein M between 2rs-m and 2m-cu 0.8 × 2rs-m; hind wing with vein CU & cu-a interrupted by distal abscissa of CU at lower 0.15. Metasoma. T1 3.5 × its apical width, 1.5 × length of propodeum; T1 finely reticulate-punctate, spiracles situated in basal 0.3, lateral margin of petiole behind spiracles parallel; lateromedian carinae and dorsolateral carinae complete and strong, reaching to apex. T2 convex on basomedian area, with a pair of short carinae medially and a lateral carina before spiracle. T2 and following tergite densely punctulate. Ovipositor sheath 2.2 × length of hind basitarsus. Colour. Black, vertex near eyes and mesoscutum medio-posteriorly with large reddish brown patch. Hypopygium yellowish brown apically. Leg black. Fore wing with three dark brown longitudinal bands: basal band on basal 0.1-0.5 of forewing, but with a wide hyaline fascia occupying middle of cell R; substigmal band below stigma and distal band in distal of forewing; hind wing nearly hyaline. Variation. Body length 7.0-7.4 mm; fore wing length 5.6-5.8 mm. Antennae 25-segmented. Transverse diameter of eye 0.9 × temple in dorsal view. Scutellum 2.0 × its basal width, 1.4 × length of propodeum. T1 4.1 × its apical width, 2.2 × length of propodeum. Male. Unknown. Host. Unknown. Distribution. China (Shaanxi). Etymology. This species is named in honour of Mrs Ma Yun, who collected many materials of Agriotypus and provided a lot of help in our research work.

Agriotypus yangae sp. nov. Figs 6B, 8 Material examined. Holotype. ♀, China, Shaanxi prov., Qinling, Tiantaishan, 34.2°N, 106.8°E, 1998.VI.9, Sun Changhai, No. 985922 (ZJUH). Diagnosis. This new species belongs to the Agriotypus himalensis species group, and it runs to A. chaoi Bennett in Bennett’s key (2001), but differs in ♀ scutellum funnel-shaped (the latter triangle-shaped); spine of scutellum black (the latter light brown apical half); mesoscutum punctate and pubescent (the latter sparsely pubescent and impunctate postero-medially). It differentiates from other new species proposed in this study by the combination characters: ♀ frons without a longitudinal carina; mesoscutum and scutellum completely black, without pale spots; mesoscutum pubescent; scutellum funnel-shaped; length of scutellum 1.8 × its basal width; spine of scutellum short, distinctly obliquely upward, rounded apically; T1 3.4 × its apical width. Description. Female. Body length 7.2 mm; fore wing length 6.0 mm. Head. Head width equal to its median length. Antennae 26-segmented, 0.7 × length of forewing. Temple behind eyes roundly narrowed in dorsal view. Transverse diameter of eye 1.2 × temple in dorsal view. Frons convex medially, with shallow depression near middle ocelli, without longitudinal carina. Antennal scrobe deep, with a weak carina on its lateral margin parallel to inner orbit of eye. Ocelli medium-sized, in triangle with base equal to its sides. POL: OD: OOL = 15: 13: 22. Face distinctly convex medially with coarse rugosity and dense pubescence. Area between antennal socket with a glabrous tubercle. Clypeus nearly pentagonal, equal to its maximum width, roundly convex, more or less flat medially and rugose with punctuation in basal half; thin apically, finely strigate, slightly rounded on apical margin; summit of covexity of clypeus smoothly rounded in lateral view. Distance between dorsal tentorial pits 1.7 × length between a tentorial pit and eye. Malar space 1.6 × basal width of mandible. Occipital carina complete. Mesosoma. Pronotum with long and strong epomia, portion dorsal to pronotal furrow curved sigmoidally; lateral of pronotum with more than seven fine carinae dorso-posteriorly. Mesoscutum punctate-rugose; notaulus deep, meeting in apical 0.2. Scutellum funnel-shaped, 1.8 × its basal width, 1.1 × length of propodeum; basal half of scutellum with lateral carinae and punctuation-rugosity, convex medially, lateral margin of scutellum paralleled in basal 2/3, tapering towards to apex in apical 1/3; in lateral view scutellum less roundly curved medially, distinctly obliquely upward, apex more or less rounded, produced 10° dorsoposteriorly. Mesopleuron and metapleuron finely punctuate and pubescent; ecepicnemial carina strong, only extending to ventral-anterior of mesopleuron; mesopleural sulcus complete; sternaulus strong. Propodeum punctate-rugose; lateromedian longitudinal carinae weakly convergent posteriorly; lateral longitudinal carinae straight, complete and paralleled with lateromedian longitudinal carinae. Wings. Fore wing with vein 1cu-a distad of M&RS, abscissa of vein M between 2rs-m and 2cu-m 1.25 × 2rs-m; hind wing with vein CU & cu-a intercepted by distal abscissa of CU interrupted at lower 0.15. Metasoma. T1 3.4 × its apical width, 1.65 × length of propodeum; T1 finely irregularly rugose-striate, transverse rugose before spiracles, spiracles situated in basal 0.3, widest at level of spiracles; lateromedian carinae and dorsolateral carinae complete and strong, reaching to apex. T2 with a pair of short carinae medially and a lateral carina before spiracle; T2 densely finely strigose. T3 and following tergite densely punctulate. Ovipositor sheath 0.8 × length of hind basitarsus. Colour. Black. Apical of four tergites, sternites and ovipositor reddish brown. Leg black. Fore wing faintly infuscate, with three slightly fuscous longitudinal bands: basal band near cu-a; substigmal band below stigma and distal band in distal 0.25 of forewing; hind wing more or less hyaline. Male. Unknown. Host. Unknown. Distribution. China (Shaanxi). Etymology. This species is named in honour of Dr Yang Lianfang, the famous entomologist who studied the taxonomy of Trichoptera in China., Published as part of Tang, Pu, He, Jun-hua & Chen, Xue-xin, 2022, Five new species of Agriotypus Curtis, 1832 (Hymenoptera, Ichneumonidae, Agriotypinae) from China, pp. 1-22 in Journal of Hymenoptera Research 90 on page 1, DOI: 10.3897/jhr.90.79244

Agriotypus dui sp. nov. Figs 1, 2 Material examined. Holotype. ♀, China, Shaanxi prov., Qinling, Jialingjiang, 34.2°N, 106.8°E, 2000 m, 1998.VI.8, Du Yuzhou, No. 983341 (ZJUH). Paratypes: China: 30♀♀4♂♂, same data with holotype, Nos. 983342-983375 (ZJUH, 2♀♀ No. 983368-983369 kept in TARI); 2♀♀4♂♂, same location and time data with holotype, Ma Yun, Nos. 982946-982947 (ZJUH); 4♀♀, Shaanxi prov., Qinling, Tiantaishan, 1998.VI.9, Sun Changhai, Nos. 985918-985921 (ZJUH); 2♀♀, Shaanxi prov., Zhouzhi, Houzhenzi, 1300 m, 1998.VI.2-3, Ma Yun, Nos.981235-981236 (ZJUH); 1♀, Shaanxi prov., Foping, Longcaoping, 33.5°N, 107.9°E, 1998.VI.4, Ma Yun, No. 981985 (ZJUH); 4♀♀1♂, Shaanxi prov., Lantian, Tangyu, 34.1°N 109.3°E, 1998.VI.1, Ma Yun, Nos. 981217-981221 (ZJUH); 1♀, Shaanxi prov., Fengxian, Jialingjiang, 33.9°N, 106.5°E, 1998.VI.10, Sun Changhai, No. 985865 (ZJUH); 2♂♂, Shaanxi prov., Ningshan, Xunyangzhen, 33.3°N, 108.3°E, 1998.VI.6, Ma Yun, Nos. 982885-9828856 (ZJUH); 23♂♂, Shaanxi prov., Ningshan, Xunyangba, 2011.V.6, Chen Huayan, Nos. 201100192-201100214 (ZJUH). Diagnosis. This new species belongs to the Agriotypus himalensis species group, and it runs to A. himalensis Mason in Bennett’s key (2001), but differs in ♂ lateral of pronotum without longitudinal rugosity dorso-posteriorly (the latter with longitudinal rugosity); ♀ length of spine of scutellum 0.9 × length of propodeum, produced 50° dorsoposteriorly (the latter 0.6-0.7 ×, more or less 35°); ♂ claspers in lateral view parallel dorso-ventrally, 1.2 × its apical width (the latter slightly enlarged, 2.4 × its apical width); legs black (the latter at least partly pale yellowish-brown). It differentiates from other new species proposed in this study by the combination characters: ♀ frons without a longitudinal carina; mesoscutum and scutellum completely black, without pale spots; mesoscutum mostly pubescent; scutellum long triangle-shaped, tapered apically; length of scutellum 2.5 × its basal width; spine of scutellum produced 50° dorsoposteriorly; length of T1 4.1-4.2 × apical width; ♂scutellum more or less parallel in basal 0.25; spine of scutellum produced 50° dorsoposteriorly; length of T1 6.5 × its apical width; paramere 1.2 × its apical width. Description. Female. Body length 6.2 mm; fore wing length 5.0 mm. Head. Head width 1.0 × its median length. Antennae 29-segmented, 0.6 × length of forewing. Temple behind eyes slightly roundly narrowed in dorsal view. Transverse diameter of eye 1.2 × temple in dorsal view. Frons convex medially, without longitudinal carina. Antennal scrobe deep, with a weak carina on its lateral margin parallel to inner orbit of eye. Ocelli medium-sized, in triangle with base equal to its sides. POL: OD: OOL = 10: 12: 16. Face convex medially with coarse rugosity and dense pubescence. Area between antennal socket with a glabrous short longitudinal tubercle. Clypeus nearly pentagonal, 1.0 × its maximum width, roundly convex, flat and rugose with punctuation in basal half; thin apically, finely strigate, slightly rounded on apical margin; summit of covexity of clypeus smoothly rounded in lateral view. Distance between dorsal tentorial pits 2.5 × length between a tentorial pit and eye. Malar space equal to basal width of mandible. Occipital carina fine, but complete. Mesosoma. Pronotum with long and strong epomia, portion dorsal to pronotal furrow curved sigmoidally; lateral of pronotum with more than 10 fine carinae dorso-posteriorly. Mesoscutum densely punctulate; notaulus distinct, meeting in apical 0.2. Scutellum long triangle-shaped, 2.5 × its basal width, 1.4 × length of propodeum; basal half of scutellum with lateral carinae and punctuation-rugosity, convex medially, tapering towards to apex, spine-like apical half, smooth; in lateral view scutellum roundly curved medially, distinctly up curved, apex slightly obliquely truncate, produced 50° dorsoposteriorly. Mesopleuron and metapleuron finely punctuate and pubescent; ecepicnemial carina strong, extending to ventral-anterior of mesopleuron; mesopleural sulcus complete; sternaulus strong. Propodeum finely coriaceous-punctate; lateromedian longitudinal carinae weakly convergent posteriorly; lateral longitudinal carinae straight, complete and paralleled with lateromedian longitudinal carinae. Wings. Fore wing with vein 1cu-a just distad of M&RS, abscissa of vein M between 2rs-m and 2cu-m 1.5 × 2rs-m; hind wing with vein CU & cu-a intercepted by distal abscissa of CU at lower 0.15. Metasoma. T1 4.1 × its apical width, 1.9 × length of propodeum; T1 finely punctulate-reticulate, spiracles situated in basal 0.3, lateral margin of T1 behind spiracles parallel; lateromedian carinae and dorsolateral carinae complete and strong, reaching to apex. T2 convex on basomedian area, with fainly carina before spiracle. T2 and following tergite densely punctulate. Ovipositor sheath 0.9 × length of hind basitarsus. Colour. Black. Pronotum ventrally, tip of spine of scutellum, apical of metasomal tergite reddish brown. Fore wing with three fuscous longitudinal bands: basal band on basal 0.1-0.5 of forewing, but with a wide hyaline spot occupying middle of cell R; substigmal band below stigma and distal band in distal 0.25 of forewing; hind wing only apical margin slightly infuscate. Variation. Body length 5.6-6.6 mm; fore wing length 4.8-5.2 mm. Head width 1.1 × its median length. Antennae 30-segmented. Scutellum 2.8 × its basal width; Fore wing with 1cu-a slightly distad of M&RS, abscissa of vein M between 2rs-m and 2cu-m 1.0-2.0 × 2rs-m; T1 3.6-3.9 × its apical width, 1.6 × length of propodeum. Body color dark reddish brown. Male. Body length 6.2-6.8 mm; fore wing length 5.4-5.7 mm. Antennae 39-40-segmented. POL: OD: OOL = 13: 12: 20. Lateral of pronotum without longitudinal rugosity dorso-posteriorly. Scutellum long triangle-shaped, 1.15-1.2 × length of propodeum; lateral carinae in basal 0.25 of scutellum more or less parallel, in apical 0.75 lineally narrowed; basal 0.45-0.5 of scutellum convex medially, with punctuation-rugosity; apical 0.5-0.55 of scutellum spine-like. lateromedian longitudinal carinae of propodeum more or less parallel, distinctly convergent in apical half. Fore wing with vein 1cu-a just distad of M&RS, abscissa of vein M between 2rs-m and 2cu-m slightly longer than 2rs-m; hind wing with CU & cu-a interrupted at lower 0.2. T1 6.5-6.8 × its apical width, 2.4-2.5 × length of propodeum; lateral margin of petiole more or less parallel. Paramere short, 1.2 × its apical width, apically bluntly round, more or less truncate. Fore wing faintly infuscate, only with more or less fuscous bands apically and below stigma. Otherwise similar to female. Host. Unknown. Distribution. China (Shaanxi). Etymology. This species is named in honour of Dr Du Yuzhou, the collector of the type specimen., Published as part of Tang, Pu, He, Jun-hua & Chen, Xue-xin, 2022, Five new species of Agriotypus Curtis, 1832 (Hymenoptera, Ichneumonidae, Agriotypinae) from China, pp. 1-22 in Journal of Hymenoptera Research 90 on page 1, DOI: 10.3897/jhr.90.79244

Bethylus ningxicus Wang, He & Chen sp. nov. Fig. 7 Description. Holotype (Fig. 7a). Male. Body length 4.36 mm. Forewing length 1.94 mm. Color. Body dark castaneous to black, metasoma castaneous. Mandible yellow. Antenna yellow. Legs castaneous, tarsi and protibia yellow. Forewing hyaline tinged with yellow; veins and pterostigma light yellowish-brown. Pubescence. Body with sparse setae, denser in propleuron and mesopectus ventrally and legs. Antenna with dense appressed setae. Forewing with short dense setae. Head (Fig. 7 b-e). Head slightly longer than wide, LH 1.04 × WH. Mandible with four teeth; upper most one with apex straight. Median clypeal lobe with apex straight; median clypeal carina slightly extending posterad into frons. Malar space narrow. Antennomeres II–VI in ratio of 1.26:1.0:1.04:1.15:1.0 in length and respectively 2.40, 1.90, 1.93, 2.0, 1.82 × width; antennal scrobal carina absent. Frons and vertex coriaceous with shallow sparse punctures. WF 1.36 × LE; LE 1.18 × DEV. Anterior ocellus far away from eye; POL1.69 × AOL; OOL 2.01 × WOT; DPV 0.97 × DAO. Vertex crest outcurved; sides of head posterior to eyes converging posteriorly. Gena coriaceous; ventral area coriaceous with punctures less than frons. Mesosoma (Fig. 7f). Pronotum coriaceous, dorsal pronotal area shorter than wide with shallow punctures. Mesoscutum coriaceous with shallow punctures; mesoscutum 0.86 × mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with shallow punctures; mesoscutellar pits nearly oval. Metapostnotum depressing and rugulose, median ridge weakly coriaceous; dorsal area of propodeum rugulose, depressing near lateral marginal carina; lateral marginal carina complete, basal half weak; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with three pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Prosternum coriaceous with distinct median longitudinal sulcus. Mesopectus coriaceous with shallow punctures; mesopleural pit and fovea weak. Forewing (Fig. 7g). Rs+M 2 v vein extremely short; 2r-rs&Rs 2 v almost straight with apex abruptly curved. Metasoma. Metasomal terga shiny, weakly coriaceous basally; median longitudinal sulcus of first metasomal tergum short not exceeding first metasomal spiracle. Metasomal sterna weakly coriaceous; first metasomal sternum coriaceous with median longitudinal carina; hypopygium bilobate with median notch deep and narrow (Fig. 7h). Genitalia (Fig. 7i–j): ventral arm of paramere with long setae, apex of dorsal arm obtuse; margin of cuspis distinctly rounded in ventral view; digitus sickle-shaped, papillate dorsally; aedeagus bottle-shaped. Female. Unknown. Variation. Body length: 2.38–3.20 mm. Length of forewing: 1.72–2.13 mm. LH 1.04–1.10 × WH; WF 1.36– 1.53 × LE; LE 1.01–1.18 × DEV; POL 1.05–1.69 × AOL; OOL 2.01–2.25 × WOT; DPV 0.97–1.71 × DAO. Type material. Holotype ♂ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3– 14.vii.2009, Huayan Chen, No. 202016006. Paratypes: ♂ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3–14.vii.2009, Huayan Chen, No. 202016011; ♂ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3–14.vii.2009, Huayan Chen, No. 202016012; ♂ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3–14.vii.2009, Huayan Chen, No. 202016013. Distribution. China (Ningxia). Etymology. This species is named based on the type specimen locality. Diagnosis. This species can be distinguished from other species of this genus by having median clypeal lobe weak (Fig. 7b), median notch of hypopygium deep and narrow (Fig. 7h), and apical margin of cuspis rounded., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on pages 373-375, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728

Nine new species as well as a known species of Bethylus Latreille (Hymenoptera, Bethylidae) from China are described and illustrated: B. crassicapitis sp. nov., B. convexus sp. nov., B. gansensis sp. nov., B. glabricarinatus sp. nov., B. hunanensis sp. nov., B. incurvus sp. nov., B. ningxicus sp. nov., B. prolatus sp. nov., B. quadraticapitis sp. nov., and B. sinensis Xu, He & Terayama. A key to the known Chinese species is provided.

Bethylus crassicapitis Wang, He & Chen sp. nov. Fig. 2 Description. Holotype (Fig. 2a). Female. Body length 4.38 mm. Forewing length 2.66 mm. Color. Body black, metasoma dark castaneous. Mandible castaneous. Antenna yellow. Legs yellowish-brown, coxae and femora castaneous. Forewing hyaline tinged with yellow; veins and pterostigma light castaneous. Pubescence. Body with sparse setae, denser in head, propleuron, mesopectus ventrally and legs. Antenna with dense appressed setae. Forewing with short dense setae. Head (Fig. 2 b-e). Head slightly longer than wide, LH 1.09 × WH. Mandible with four teeth; upper most with apex rounded. Median clypeal lobe developed with apex rounded; median carina developed, slightly extending posterad into frons. Antennomeres II–VI in ratio of 1.11:1.11:1.09:1.0:1.0 in length and respectively 2.21, 2.28, 2.25, 2.0, 1.95, 2.02 × width; antennal scrobal carina absent. Frons and vertex coriaceous with shallow sparse punctures separate 1.0–4.0 × their own diameter. WF 1.46 × LE; LE 1.08 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL 0.94 × AOL; OOL 2.07 × WOT; DPV 1.17 × DAO. Vertex crest slightly outcurved; sides of head posterior to eyes converging posteriorly. Malar space narrow, malar line between mandible and eye absent. Gena coriaceous. Mesosoma (Fig. 2 f-g). Pronotum coriaceous; dorsal pronotal area shorter than wide. Mesoscutum coriaceous with shallow sparse punctures; mesoscutum 0.82 × mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with few punctures; mesoscutellar pits elongated. Metapostnotum depressed and rugulose, median ridge weakly coriaceous; dorsal area of propodeum rugulose, depressing and obliquely rugose near lateral marginal carina of metapectal-propodeal disc; lateral marginal carina of metapectal-propodeal disc complete; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with three pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Mesopectus coriaceous with shallow punctures; mesopleural pit and fovea weak. Prosternum coriaceous with weak longitudinal sulcus medially. Forewing (Fig. 2h). Rs+M 2 v extremely short; 2r-rs&Rs 2 v slightly rounded with apex rounded curved. Metasoma. Metasomal terga shiny, weakly coriaceous basally; longitudinal sulcus of first metasomal tergum distinctly exceeding first metasomal spiracle. Metasomal sterna weakly coriaceous. Male. Unknown. Variation. Body length: 3.62–4.38 mm. Forewing length: 2.18–2.66 mm. LH 1.09–1.12 × WH; WF 1.56–1.66 × LE; POL 0.94–1.01 × AOL; OOL 2.07–2.43 × WOT; DPV 1.17–1.38 × DAO. Length of mesoscutum 0.82–0.93 × length of mesoscutellum. Type material. Holotype ♀ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3– 14.vii.2009, Huayan Chen, No. 202016004. Paratypes: ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3–14.vii.2009, Huayan Chen, No. 202016001; ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3–14.vii.2009, Huayan Chen, No. 202016007. Distribution. China (Ningxia). Etymology. The specific name “ crassicapitis ” is a compound of “ crass ” (thick) and “ caput ” (head) which refers to the thicken head of this species. Diagnosis. This species can be distinguished from other species of this genus by having ocellar triangle elongate, head thickened in lateral view, and median clypeal carina distinctly protuberant in lateral view.

Bethylus Latreille, 1802 Bethylus Latreille, 1802: 315. Type species: Omalus fuscicornis Jurine, 1807. Anoxus Thomson, 1862: 452. Type species: Anoxus boops Thomson, 1862. Synonymy by Polaszek & Krombein 1994. Anoxys Dalla Torre, 1898. Unjustified emendation to Anoxus by Dalla Torre (1898, 5: 550) Perisemus Förster, 1856: 95-96. Type species: Bethylus triareolatus Förster, 1856. Synonymy by Kieffer 1905 (in Kieffer & Marshall 1904 –1906). Episemus Thompson, 1862: 452. Type species: Epysemus variabilis Thompson, 1862 [1861]. Synonymy by Richards 1939. Digoniozus Kieffer, 1905. Type species: Perisemus oregonensis Ashmead, 1893. Synonymy by Evans 1962. Diagnosis. Clypeus short, usually not strongly angulated medially; antenna with 10 flagellomeres; notauli absent; parapsidal signum usually present; lateral marginal carina of metapectal-propodeal disc present; metapostnotal median carina and metapostnotal lateral carina absent; prestigma of forewing absent; Rs+M 2 v usually short as stub; second radial 1 cell of forewing (2R1 2 c) open apically; posterior margin of hypopygium bilobate; paramere of male genitalia distinctly divided into two parts, ventral and dorsal arms. Biology. Most of the host records known of Bethylus species are lepidopterous larvae, e.g., Agonoxenidae, Tortricidae, Zygaenidae, Cosmopterigidae, Coleophoridae, and Gelechiidae (Evans 1964; Richards 1939). Richards (1939) reported that Bethylus amoenus Fouts, 1928 attacks the larvae of both Lepidoptera and Coleoptera. Distribution. Palaearctic, Nearctic and Oriental Regions. Key to the Chinese Bethylus species 1. Median clypeal lobe with apex rounded................................................................... 2 - Median clypeal lobe with apex truncate................................................................... 7 2. Frontal angle of ocellar triangle obtuse (Fig. 3b)............................................ B. gansensis sp. nov. - Frontal angle of ocellar triangle acute (Fig. 1b)............................................................. 3 3. LH at least 1.24 × width; POL at least 3.80 × DAO.......................................................... 4 - LH less than 1.20 × width; POL less than 3.20 × DAO....................................................... 5 4. Eye distinctly protuberant (Fig. 1b); median clypeal lobe long; median clypeal carina rounded in lateral view (Fig. 1e)........................................................................................ B. convexus sp. nov. - Eye not distinctly protuberant (Fig. 10b); median clypeal lobe short; median clypeal carina almost arched in right angle in lateral view (Fig. 10e)................................................... B. sinensis Xu, He & Terayama, 2002 5. Distance of posterior ocellus to vertex crest more than 2.80 × DAO; median clypeal lobe slightly protuberant (Fig. 4b)................................................................................... B. glabricarinatus sp. nov. - Distance of posterior ocellus to vertex crest less than 1.50 × DAO; median clypeal lobe distinctly protuberant (Fig. 6b).... 6 6. Head thickened in lateral view (Fig. 2e); median clypeal carina distinctly protuberant in lateral view (Fig. 2e)............................................................................................ B. crassicapitis sp. nov. - Head not thickened in lateral view (Fig. 6e); median clypeal carina incurved in lateral view (Fig. 6e)... B. incurvus sp. nov. 7. DPV more than 2.40 × DAO....................................................... B. quadraticapitis sp. nov. - DPV less than 1.50 × DAO............................................................................. 8 8. Apex of cuspis narrow (Fig. 5k)....................................................... B. hunanensis sp. nov. - Apex of cuspis rounded (Fig. 8k)........................................................................ 9 9. Median depression of hypopygium broad (Fig. 8i); apical margin of cuspis strongly echinate.......... B. prolatus sp. nov. - Median depression of hypopygium narrow and deep (Fig. 7h); apical margin of cuspis not echinate (Fig. 7j)................................................................................................. B. ningxicus sp. nov., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on pages 362-363, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728, {"references":["Latreille, P. A. (1802) Histoire naturelle, genera et particuliere des Crustaces et des insects. Vol. 3. Dufort, Paris, 348 pp.","Jurine, L. (1807) Nouvelle methode de classer les hymenopteres et les dipteres. J. J. Paschaud, Geneva, 319 pp. https: // doi. org / 10.5962 / bhl. title. 60886","Thomson, C. G. (1862 [1861]) Sveriges Proctotruper (Fortsattning). Svenska Vetenskapsakademien. Stockholm. Ofversigt at K. Akademiens. Forhandlingar, 18, 451 - 453.","Polaszek, A. & Krombein, K. V. (1994) The genera of Bethylinae (Hymenoptera: Bethylidae). Journal of Hymenoptera Research, 3, 91 - 105.","Dalla Torre, C. G. D. (1898) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Chalcididae et Proctotrupidae. Vol. 5. Suptibus Gulleimi Engelmann, Lipsiae, 598 pp.","Forster, A. (1856) Hymenopterologische Studien II: Chalcidiae und Proctotrupii. Vol. 2. Ernst ter Meer, Aachen, 152 pp.","Kieffer, J. J. & Marshall, T. A. (1904 - 1906) Proctotrypides. In: Andre, E. (Ed.), Species des Hymenopteres d'Europe & d'Algerie. Tome IX. Librairie Scientifique A. Herman, Paris, pp. 1 - 55, 21 pls.","Richards, O. W. (1939). The British Bethylidae (S. L.) (Hymenoptera). Ecological Entomology, 89 (8), 185 - 344. https: // doi. org / 10.1111 / j. 1365 - 2311.1939. tb 00740. x","Ashmead, W. H. (1893) A monograph of the North American Proctotrypidae. Bulletin of the United States National Museum, 45, 1 - 472. https: // doi. org / 10.5479 / si. 03629236.45.1","Evans, H. E. (1962) The genus Bethylus in North America (Hymenoptera: Bethylidae). Breviora, 150, 1 - 12.","Evans, H. E. (1964) A synopsis of the American Bethylidae (Hymenoptera, Aculeata). Bulletin of the Museum of Comparative Zoology, 132 (1), 1 - 222.","Fouts, R. M. (1928) Notes on the Bethylinae with descriptions of one new Cuban and twelve new North American species (Hym.). Proceedings of the Entomological Society of Washington, 30, 121 - 132.","Xu, Z., He, J. & Terayama, M. (2002) The genus Bethylus Latreille, 1802 from China with description of a new species. Acta Entomologica Sinica, 45, 112 - 114."]}

Bethylus glabricarinatus Wang, He & Chen sp. nov. Fig. 4 Description. Holotype (Fig. 4a). Female. Body length 4.85 mm. Forewing length 2.91 mm. Color. Body black, metasoma dark castaneous. Mandible orange. Antenna yellow to yellowish-brown. Legs dark castaneous, tibiae and tarsi yellow. Forewing hyaline tinged with yellow; veins light castaneous to castaneous; pterostigma castaneous. Pubescence. Body with sparse setae, denser in propleuron, mesopectus ventrally and legs. Antenna with dense appressed setae. Forewing with short dense setae. Head (Fig. 4 b-e). Head longer than wide, LH 1.07 × WH. Mandible with four teeth; upper most with apex straight. Median clypeal lobe with apex rounded; median clypeal carina slightly extending posterad into frons. Malar space narrow. Antennomeres II-VI in ratio of 0.95:0.83:0.93:0.89:1.0 in length and respectively 2.11, 1.82, 1.89, 1.85, 2.18 × width; antennal scrobal carina absent. Frons and vertex coriaceous with dense punctures separate less than 3.0 × their diameter. WF 1.75 × LE; LE 0.84 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL 0.94 × AOL; OOL 2.54 × WOT; DPV 3.20 × DAO. Vertex crest straight; sides of head posterior to eyes converging posteriorly. Gena coriaceous; ventral area convex with punctures less than frons. Mesosoma (Fig. 4 f-g). Pronotum coriaceous; dorsal pronotal area shorter than wide with shallow punctures. Mesoscutum coriaceous with punctures separate less than 3.0 × their own diameter, 0.90 × mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with punctures; mesoscutellar pits elongate. Metapostnotum rugulose, median ridge weakly coriaceous; dorsal area of propodeum coriaceous; lateral marginal carina of metapectal-propodeal disc very weak; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with two pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Mesopectus coriaceous with punctures; mesopleural pit and fovea present. Forewing (Fig. 4h). Rs+M 2 v vein short, almost as long as width of Rs 2 v; 2r-rs&Rs 2 v almost straight with apex abruptly curved. Metasoma. Metasomal terga shiny with shallow tiny punctures; longitudinal sulcus of first metasomal tergum present, almost 0.5 × length of first metasomal tergum. Metasomal sterna shiny with shallow tiny punctures. Male. Unknown. Variation. Body length: 3.90–4.85 mm. Length of forewing: 2.61–2.98 mm. LH 1.05–1.09 × WH; WF 1.61– 1.83 × LE; LE 3.53–4.15 × DEV; POL 0.94–1.05 × AOL; OOL 2.28–2.54 × WOT; DPV 2.82–3.20 × DAO. Type material. Holotype ♀ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3– 14.vii.2009, Huayan Chen, No. 202016010. Paratypes: ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3–14.vii.2009, Huayan Chen, No. 202016018; ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3–14.vii.2009, Huayan Chen, No. 202016019. Distribution. China (Ningxia). Etymology. The specific name “ glabricarinatus ” compounded with two Latin words: “ glabr ” (smooth) and “ carina ” (keel) refers to the area alongside median clypeal carina shiny and slightly elevated. Diagnosis. This species can be distinguished from other species of this genus by having LH less than 1.10× WH, median clypeal carina weak, and DPV at least 2.82 × DAO., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on pages 368-370, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728

Bethylus hunanensis Wang, He & Chen sp. nov. Fig. 5 Description. Holotype (Fig.5a). Male. Body length 2.92 mm. Forewing length 1.67 mm. Color. Head castaneous; mandible castaneous, teeth light castaneous; antenna light castaneous. Mesosoma castaneous; legs light castaneous, tarsi pale castaneous. Metasoma light castaneous. Forewing hyaline; veins and pterostigma light castaneous. Pubescence. Head with sparse setae. Antenna with dense short white setae. Mesosoma ventrally and metasoma with sparse setae. Forewing with dense short setae, radial cell and costal cell of forewing with sparse setae. Head (Fig. 5 b-e). Head longer than wide, LH 1.15 × WH. Mandible with four apical teeth. Median clypeal lobe weak with apex truncate; median clypeal carina slightly extending posterad into frons. Antennomeres II–VI in ratio of 1.13:0.97:0.87:1.01:1.0 in length and 1.90, 2.03, 1.55, 1.89, 1.82 × width respectively; antennal scrobal carina absent. Frons coriaceous with shallow punctures separate at least 1.50 × their diameter. WF 1.43 × LE. LE 1.02 × DEV. Frontal angle of ocellar triangle acute; POL 1.59 × AOL; OOL 1.98 × WOT; DPV 0.80 × DAO. Vertex coriaceous with shallow sparse punctures; vertex crest slightly outcurved. Sides of head posterior to eyes slightly converging posteriorly. Occipital carina absent. Malar space narrow, malar line between mandible and eye absent. Gena coriaceous with punctures separate at least 1.50 × their diameter. Mesosoma (Fig. 5 f-g). Pronotum coriaceous, posterior margin almost straight. Mesoscutum coriaceous with few punctures; parapsidal signum complete. Mesoscutellum coriaceous with few punctures. Metapectal-propodeal disc imbricate, median ridge shiny; lateral marginal carina of metapectal-propodeal disc present, basal one third absent; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with three pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Mesopectus coriaceous with shallow punctures; mesopleural fovea absent; mesopleural pit present. Forewing (Fig. 5h). Rs+M 2 v 0.20 × M 2 v; Rs 2 v 1.94 × M 2 v; 2r-rs&Rs 2 v arched in obtuse angle apically. Metasoma. Metasomal terga shiny with shallow tiny punctures; longitudinal sulcus of first metasomal tergum short and not exceeding first metasomal spiracle. Metasomal sterna shiny; basal half of first metasomal sternum irregularly rugulose with weak median longitudinal carina, apical half coriaceous; hypopygium bilobate with median notch broad (Fig. 5i). Genitalia (Fig. 5j, k): ventral arm of paramere with long setae, apex of dorsal arm acute; margin of cuspis distinctly outcurved in ventral view; digitus sickle-shaped and papillate dorsally; aedeagus bottle-shaped. Female. Unknown. Type material. Holotype ♂ (ZJUH), China, Hunan Province, Shimen Country, Huping Mountain, 110°40′35.9544″E, 30°0′34″N, 2000m, 29.vi.1987, Guangchun Lei, No. 20044537. Distribution. China (Hunan). Etymology. This species is named based on the type specimen locality. Diagnosis. This species can be distinguished from other species of this genus by having LH 1.15× WH, median notch of hypopygium broad (Fig. 5i), and apex of cuspis outcurved (Fig. 5k).

Bethylus prolatus Wang, He & Chen sp. nov. Fig. 8 Description. Holotype (Fig. 8a). Male. Body length 3.54 mm. Forewing length 2.09 mm. Color. Body dark castaneous to black, metasoma castaneous. Mandible yellow. Antenna yellow. Legs yellow, coxae and femora castaneous. Forewing hyaline tinged with light castaneous; veins and pterostigma light castaneous. Pubescence. Body with sparse setae, denser in head, propleuron, mesopectus ventrally and legs. Antenna with dense appressed setae. Forewing with short dense setae. Head (Fig. 8b–e). Head slightly longer than wide, LH 1.07 × WH. Mandible with four teeth; upper most one with apex straight. Median clypeal lobe developed with apex straight; median clypeal carina slightly extending posterad into frons. Antennomeres II–VI in ratio of 1.32:1.18:1.08:1.04:1.0 in length and respectively 2.37, 2.37, 2.11, 1.92, 1.80 × width; antennal scrobal carina absent. Frons and vertex coriaceous with shallow sparse punctures separate 1.0–4.0 × their own diameter. WF 1.51 × LE; LE 1.05 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL1.07 × AOL; OOL 2.17 × WOT; DPV 0.93 × DAO. Vertex crest straight; sides of head posterior to eyes converging posteriorly. Malar space narrow, malar line between mandible and eye absent. Gena coriaceous; ventral area coriaceous with punctures less than frons. Mesosoma (Fig. 8 f-g). Pronotum coriaceous; dorsal pronotal area shorter than wide with shallow punctures. Mesoscutum coriaceous with shallow punctures; mesoscutum 0.86 × mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with shallow punctures; mesoscutellar pits oblong. Metapostnotum rugulose, median ridge weakly coriaceous; dorsal area of propodeum rugulose, obliquely rugose near lateral marginal carina; lateral marginal carina of metapectal-propodeal disc complete; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with three pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Prosternum coriaceous. Mesopectus coriaceous with shallow punctures; mesopleural pit and fovea weak. Forewing (Fig. 8h). Rs+M 2 v extremely short; 2r-rs&Rs 2 v slightly rounded with apex curved. Metasoma. Metasomal terga shiny, weakly coriaceous basally; median longitudinal carina of first metasomal tergum exceeding first metasomal spiracle, about one third as long as first metasomal tergum. Metasomal sterna coriaceous; first metasomal sternum rugulose, basal half elevated medially, median longitudinal carina absent; hypopygium bilobate with median notch deep and narrow (Fig. 8i). Genitalia (Fig. 8 j-k): ventral arm of paramere with long setae, apex of dorsal arm obtuse; margin of cuspis distinctly rounded in ventral view; digitus sickle-shaped and papillate dorsally; aedeagus bottle-shaped. Female. Unknown. Variation. Body length: 3.45–4.36 mm. Length of forewing: 1.94–2.33 mm. LH 1.04–1.07 × WH; WF 1.51– 1.64 × LE; LE 1.02–1.09 × DEV; POL 1.02–1.07 × AOL; OOL 2.10–2.23 × WOT; DPV 0.93–1.14 × DAO. Type material. Holotype ♂ (ZJUH), China, Hebei Province, Zhangjiakou, Xiaowutai Mountain, 20– 23.viii.2005, Jingxian Liu, No. 200609401. Paratypes: ♂ (ZJUH), Ningxia, Jingyuan, Hongxia Forest Farm, 106°20′14″E, 35°30′15″N, 1–7.vii.2008, Minjie Yao, No. 200808904; ♂ (ZJUH), Ningxia, Jingyuan, Hongxia Forest Farm, 106°20′14″E, 35°30′15″N, 1–7.vii.2008, Minjie Yao, No. 200808906. Distribution. China (Hebei, Ningxia). Etymology. The specific name “ prolatus “refers to the extended clypeus of this species. Diagnosis. This species can be distinguished from other species of this genus by having its head almost square (Fig. 8b), apex of median clypeal lobe truncate (Fig. 8b), notch of hypopygium broad (Fig. 8i), and cuspis echinate (Fig. 8k)., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on pages 375-377, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728

Bethylus quadraticapitis Wang, He & Chen sp. nov. Fig. 9 Description. Holotype (Fig. 9a). Male. Body length 3.51 mm. Forewing length 2.42 mm. Color. Body black, metasoma castaneous. Mandible yellow. Antenna yellow. Legs castaneous, trochanters, tarsi and protibia yellow. Forewing hyaline tinged with yellow; veins light castaneous; pterostigma castaneous. Pubescence. Body with sparse setae, denser in propleuron, mesopectus ventrally and legs. Antenna with dense appressed setae. Forewing with short dense setae. Head (Fig. 9b–e). Head slightly longer than wide, LH 1.04 × WH. Mandible with four teeth; upper most one with apex straight. Median clypeal lobe with apex straight; median clypeal carina slightly extending posterad into frons. Antennomeres II–VI in ratio of 1.13: 0.91: 1.07: 1.06:1.0 in length and respectively 2.45, 1.92, 2.13, 2.07, 2.26 × width; antennal scrobal carina absent. Frons and vertex coriaceous with shallow sparse punctures. WF 1.68 × LE; LE 0.92 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL 1.12 × AOL; OOL 2.16 × WOT; DPV 2.47 × DAO. Vertex crest straight; sides of head posterior to eyes almost parallel. Malar space narrow, malar line between mandible and eye absent. Gena coriaceous, ventral area coriaceous and elevated with punctures less than frons. Mesosoma (Fig. 9 f-g). Pronotum coriaceous; dorsal pronotal area shorter than wide with shallow punctures. Mesoscutum coriaceous with shallow punctures; mesoscutum 0.96 × mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with shallow punctures; mesoscutellar pits almost oval. Metapostnotum rugulose, median ridge weakly coriaceous; dorsal area of propodeum rugulose, obliquely rugose near lateral marginal carina; lateral marginal carina of metapectal-propodeal disc weak basally; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with two pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Mesopectus coriaceous with shallow punctures; mesopleural pit and fovea weak. Forewing (Fig. 9h). Rs+M 2 v short, longer than the width of Rs 2 v; 2r-rs&Rs 2 v with apex rounded curved. Metasoma. Metasomal terga shiny, weakly coriaceous basally; median longitudinal carina of first metasomal tergum not exceeding first metasomal spiracle. Metasomal sterna weakly coriaceous; basal half of first metasomal sternum rugulose, apical half coriaceous, median longitudinal carina present at basal half; hypopygium bilobate with median notch deep and narrow (Fig. 9i). Genitalia (Fig. 9j–k): ventral arm of paramere with long setae, apex of dorsal arm obtuse; margin of cuspis distinctly rounded in ventral view with long setae; digitus sickle-shaped and papillate dorsally; aedeagus bottle-shaped Female. Unknown. Type material. Holotype ♂ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′14″E, 35°30′15″N, 3– 14.vii.2009, Huayan Chen, No. 202016014. Distribution. China (Ningxia). Etymology. The specific name “ quadraticapitis ” compounded with two Latin words: “ quadrat ” (square) and “ capitis ” (head) refers to the sides of the head posterior to the eye parallel and the head is almost square. Diagnosis. This species can be distinguished from other species of this genus by having DPV 2.47 × DAO, notch of hypopygium deep and narrow (Fig. 9i), and cuspis rounded with long setae (Fig. 9k)., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on page 377, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728

Bethylus incurvus Wang, He & Chen sp. nov. Fig. 6 Description. Holotype (Fig. 6a). Female. Body length 3.52 mm. Forewing length 2.24 mm. Color. Body dark castaneous, metasoma castaneous. Mandible castaneous with teeth yellowish-brown. Antenna yellow to yellowish-brown. Legs castaneous, trochanters, tibiae and tarsi yellow. Forewing hyaline tinged with yellowish; veins and pterostigma light castaneous. Pubescence. Body with sparse setae, denser in frons, ventral surface of propleuron and mesopectus. Antenna with short appressed setae. Forewing with dense short setae. Legs with dense setae. Head (Fig. 6 b-e). Head longer than wide, LH 1.16 × WH. Mandible with four apical teeth; upper most one with apex rounded. Median clypeal lobe developed with apex rounded; median clypeal carina slightly extending posterad into frons. Antennomeres II–VI in ratio of 0.94:0.85:1.02:1.18:1.0 in length and respectively 1.85, 1.67, 1.63, 2.18, 1.96 × width; antennal scrobal carina absent. Frons and vertex coriaceous with punctures separate 1.0–2.0× their own diameter. WF 1.40 × LE; LE 1.0 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL 1.05 × AOL; OOL 2.33 × WOT; DPV 1.50 × DAO. Vertex crest almost straight; sides of head posterior to eyes converging posteriorly. Malar space narrow, malar line between mandible and eye absent. Gena coriaceous, ventral area coriaceous with shallow punctures. Mesosoma (Fig. 6 f-g). Pronotum coriaceous; dorsal pronotal area shorter than wide, coriaceous with shallow punctures. Mesoscutum coriaceous with shallow punctures; mesoscutum 0.80× mesoscutellum; parapsidal signum complete. Mesoscutellum coriaceous with shallow punctures; mesoscutellar pits elongated. Metapostnotum rugulose, median ridge weakly coriaceous; dorsal area of propodeum coriaceous; lateral marginal carina of metapectalpropodeal disc complete but weak basally; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with three pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Prosternum coriaceous with a distinct longitudinal sulcus. Mesopectus coriaceous with shallow punctures; mesopleural pit and fovea weak; ventral surface coriaceous with subpleural signum; mesodiscrimen absent. Forewing (Fig. 6h). Rs+M 2 v extremely short; 2r-rs&Rs 2 v slightly rounded with apex abruptly curved. Metasoma. Metasomal terga weakly coriaceous, shiny apically; median longitudinal sulcus of first metasomal tergum distinctly exceeding first metasomal spiracles. Metasomal sterna weakly coriaceous with tiny punctures; first metasomal sternum rugulose, median longitudinal carina present. Male. Unknown. Variation. Body length 3.26–3.73 mm. Length of forewing: 1.94–2.28 mm. LH 1.11–1.19 × WH; WF 1.13– 1.48 × LE; LE 0.95–1.02 × DEV; POL 1.05–1.27 × AOL; OOL 2.23–2.81 × WOT; DPV 1.12–1.53 × DAO. Type material. Holotype ♀ (ZJUH), China, Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3– 14.vii.2009, Huayan Chen, No. 202016002. Paratypes: ♀ (ZJUH), Ningxia, Jingyuan, Erlong River, 3.vii.2008, Jiemin Yao, No. 200808482; ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3–14.vii.2009, Huayan Chen, No. 202016003; ♀ (ZJUH), Ningxia, Liupan Mountain, 106°20′12″E, 35°39′31″N, 3–14.vii.2009, Huayan Chen, No. 202016005. Distribution. China (Ningxia). Etymology. The specific name “ incurvus ” refers to median clypeal carina incurved in lateral view. Diagnosis. This species can be distinguished from other species of this genus by having LH at least 1.15 × WH, apex of median clypeal lobe distinctly protuberant with apex rounded (Fig. 6b), and median clypeal carina incurved in lateral view (Fig. 6e)., Published as part of Wang, Chung-Hong, He, Jun-Hua & Chen, Xue-Xin, 2021, Taxonomy of Bethylus (Hymenoptera, Bethylidae) from China with description of nine new species, pp. 361-382 in Zootaxa 4974 (2) on pages 370-373, DOI: 10.11646/zootaxa.4974.2.6, http://zenodo.org/record/4775728

Bethylus sinensis Xu, He & Terayama, 2002 Fig. 10 Bethylus sinensis Xu, He & Terayama, 2002: 112–113. Description. Holotype (Fig. 10a). Female. Body length 2.92 mm. Forewing length 0.18 mm. Color. Body dark castaneous to black. Mandible dark castaneous, teeth castaneous. Antenna yellow-brown, castaneous distad; tegula castaneous. Legs dark castaneous, tibiae and tarsi yellow. Pubescence. Body with setae in different density, denser in legs. Antenna with appressed short setae. Head (Fig. 10 b-e). Head distinctly longer than wide, LH 1.26 × WH. Mandible with four apical teeth; upper most with apex rounded. Median clypeal lobe weak with apex rounded; median clypeal carina slightly extending posterad into frons. Antennomeres II–VI in ratio of 1.20: 0.90:0.95:1.12:1.0 in length and respectively 1.90, 1.55, 1.38, 1.54, 1.53 × width; antennal scrobal carina absent. Frons and vertex coriaceous with shallow sparse punctures. WF 1.19 × LE. LE 0.97 × DEV. Anterior ocellus far away from eyes; frontal angle of ocellar triangle acute; POL 1.49 × AOL; OOL 1.83 × WOT; DPV 0.93 × DAO. Vertex crest straight. Sides of head posterior to eyes almost parallel. Occipital carina absent. Malar space narrow. Gena coriaceous; ventral area coriaceous and flat. Mesosoma (Fig. 10 f-g). Pronotum coriaceous; dorsal pronotal area distinctly longer than wide with shallow punctures; posterior margin straight (Fig. 10f). Mesoscutum coriaceous with few shallow punctures; mesoscutum 0.77 × mesoscutellum; parapsidal signum absent. Mesoscutellum coriaceous with pair of oval pits (Fig. 10f). Dorsal surface of metapectal-propodeal disc coriaceous, medially elevated; lateral marginal carina of metapectal-propodeal disc complete, weak basally; propodeal declivity coriaceous; anterior metapleural area coriaceous, metapleural line with two pits; lateral surface of metapectal-propodeal complex coriaceous. Propleuron coriaceous. Mesopectus coriaceous with shallow punctures; mesopleural pit present; mesopleural fovea absent. Forewing. Extremely small, oval, extending to one fifth of metapectal-propodeal complex. Metasoma. Metasomal terga weakly coriaceous, apical margin shiny; median longitudinal sulcus of first metasomal tergum fovea-like nearly invisible. Metasomal sterna weakly coriaceous; first metasomal sternum coriaceous, median longitudinal carina present on basal half. Male. Unknown. Type material. Holotype ♀ (ZJUH), China, Liaoning Province, Shenyang, Dongling, vi-vii.1994, Juxian Lou, No. 947820. Specimen examined. Holotype ♀ (ZJUH), China, Liaoning Province, Shenyang, Dongling, vi-vii. 1994, Juxian Lou, No. 947820; ♀ (ZJUH), China, Hebei Province, Zhangjiakou, Xiaowutai Mountain, 20–23.viii.2005, Jingxian Liu, No. 200609404. Distribution. China (Hebei, Liaoning).

17. Megacampsomeris stoetzneri (Betrem, 1928) Campsomeris (Megacampsomeris) stoetzneri Betrem, 1928: 139, ♀, China (holotype in ETHZ, Zürich); Betrem, 1941: 63; Tsuneki, 1972a: 7; Kim, 2009: 101. Megacampsomeris stoezneri: Osten, 2005: 23. Bellimeris stoezneri: Schulten et al., 2011: 887. Distribution. China (Beijing, Sichuan, Taiwan); Japan and Korea., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 110, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Kim, J. K. (2009) Taxonomic review of the tribe Campsomerini (Scoliinae, Scoliidae, Hymenoptera) in Korea. Animal Systematics, Evolution and Diversity, 25 (1), 99 - 106. https: // doi. org / 10.5635 / KJSZ. 2009.25.1.099","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Schulten, G. G. M., Feijen, H. R. & Feijen, C. (2011) The genus Bellimeris Betrem (Hymenoptera: Scoliidae, Campsomerinae). Zoologische Mededelingen, 85, 887 - 903."]}

19. Micromeriella marginella (KIug, 1810) Scolia marginella KIug, 1810: 214, ♂, India (type in Berlin Museum); Dalla Torre, 1897: 169. Elis (Campsomeris) hirsuta de Saussure, 1858: 234. Elis hirsuta: de Saussure, 1892: 20; Bingham, 1897: 98. Elis (Dielis) hirsuta: de Saussure, 1864: 216. Scolia hirsuta: Dalla Torre, 1897: 163. Campsomeris (Campsomeris) marginella marginella: Betrem, 1928: 135; Bradley & Betrem, 1968: 329. Campsomeris (Campsomeris) marginella formosana var. flavocincta Betrem, 1928: 136. Campsomeris (Campsomeriella) marginella marginella: Betrem,1941: 90. Campsomeris marginella: Chhotani & Ray, 1975: 26. Campsomeris (Micromeris) marginella: Jonathan, 1976: 113. Micromeriella marginella: Betrem, 1972: 119. Micromeriella marginella marginella: Bradley, 1974: 443; Krombein, 1978: 19; Gupta & Jonathan, 2003: 37. Campsomeris marginella formosana Betrem, 1928: 136; Betrem, 1941: 91; Tsuneki, 1972a: 16. Distribution. China (Fujian, Guangdong, Guangxi, Taiwan, Zhejiang); India and Sri Lanka. Notes. Seven subspecies have been recognized in this species (Gupta & Jonathan 2003). We checked all specimens in RMNH, including the types of the nominate subspecies, and the related literature. It is easy to separate them into two groups by absence or presence of vein 2r-m of the fore wing: the vein is absent in bariensis (Tsuneki), billitonensis (Tullgren), modesta (Smith), terminata (Smith), and present in formosana (Betrem), marginella (Klug), and thaiana (Tsuneki). The other differences concern the extent of yellow maculation of males and females. DNA sequencing is needed to research the status of these subspecies., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 110, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Dalla Torre, C. G. (1897) s. n. In: Catalogus Hymenoptorum hucusque descriptorum systematicus et synonymicus. VIII. W. Engelmann, Lipsiae (Leipzig), pp. 144 - 188.","Saussure, H. de (1858) Description de diverses especes nouvelles ou peu connues du genre Scolia. Annales de la Societe entomologique de France, 3 (6), 193 - 249.","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Bradley, J. C. & Betrem, J. G. (1968) Friedrich Klugs material of Scoliidae with especial reference to his type specimens. Deutsche Entomologische Zeitschrift, 15, 321 - 334. https: // doi. org / 10.1002 / mmnd. 19680150406","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Chhotani, O. B. & Ray, K. K. (1975) Fauna of Rajasthan, India. Hymenoptera. Records of Zoological Survey of India, 71, 13 - 49.","Jonathan, J. K. (1976) Records of additional Campsomerinae from India (Hymenoptera: Aculeata). Newsletter of the Zoological Survey of India, Calcutta, 2 (3), 112 - 114.","Betrem, J. G. & Bradley, J. C. (1972) The African Campsomerinae (Hym., Scoliidae). Mono-grafieen van de Nederlandse Entomologische Vereniging, 6, 1 - 326.","Krombein, K. V. (1978) Biosystematic studies of Ceylonese Wasps, II: A monograph of the Scoliidae (Hymenoptera: Scolioidea). Smithsonian Contributions to Zoology, 283, 1 - 56. https: // doi. org / 10.5479 / si. 00810282.283","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41."]}

5. Colpa tartara (de Saussure, 1880) Trielis tartara de Saussure, 1880: 26, ♀, Russia (type in Zoological Institute, St. Petersburg). Campsoscolia (Crioscolia) tartara tartara Bradley, 1950: 432. Crioscolia (Crioscolia) tartara: Betrem, 1972: 65. Colpa (Crioscolia) tartara: Lelej & Mokrousov, 2017: 12; Osten, 2005: 23. Scolia (Trielis) mongolica Morawitz, 1889: 116. (= Trielis konowi Gribodo, 1895) ? Campsoscolia mongolica: Betrem, 1941: 50. Campsoscolia (Campsoscolia) mongolica: Bradley, 1950: 433. Colpa (Crioscolia) tartara mongolica: Osten, 2005: 23. Distribution. China (Neimenggu, Xinjiang); Afghanistan, Armenia, Japan, Kazakhstan, Mongolia, Russian, Tajikistan, Turkmenistan and Uzbekistan. Notes. The male from China in RMNH identified as C. mongolica by Betrem (Fig. 1) has the head missing and differs from the nominate C. tartara (Fig. 2) because of the robust marginal cell of the fore wing (elongate in C. tartara); the medially interrupted yellow bands on the first two metasomal tergites (broad and continuous in C. tartara); and strongly constricted first metasomal tergite (less constricted in C. tartara). Colpa mongolica was treated as a subspecies of C. tartara (Osten 2005) or as a full synonym (Lelej & Mokrousov 2017); the reported flat surface of the area between the antennal sockets (“spatium frontale”) for C. mongolica is apparently a character too variable to be of any use to differentiate the species., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on pages 105-107, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Bradley, J. C. (1950) The most primitive Scoliidae. Eos, Torno extraord., 427 - 437.","Betrem, J. G. & Bradley, J. C. (1972) The African Campsomerinae (Hym., Scoliidae). Mono-grafieen van de Nederlandse Entomologische Vereniging, 6, 1 - 326.","Lelej, A. S. & Mokrousov, M. V. (2017) The types of Scoliidae (Hymenoptera), described by Eduard Eversmann, with some taxonomic notes and checklist of Russian Scoliidae. Far Eastern Entomologist, 340, 1 - 17. https: // doi. org / 10.25221 / fee. 340.1","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Morawitz, F. (1889) Insecta, a Cl. G. N. Potanin in China et in Mongolia novissime lecta. IV. Hymenoptera Aculeata. Horae Societatis Entomologicae, 23, 112 - 168.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188."]}

36. Scolia (Discolia) inouyei Okamoto, 1924 re-instated Scolia inouyei Okamoto, 1924: 205, ♀, Korea (type in Agricultural Experimental Station, Suigen, S. Korea?); Yamane, 1995: 43. Scolia nobilis f. inouyei: Uchida, 1933: 250. Scolia nobilis inouyei: Betrem, 1941: 140; Osten, 2005: 39. Distribution. China (Taiwan, Yunnan); Korea. Notes. We re-instate Scolia (Discolia) inouyei Okamoto, 1924 as a valid species, after examining a specimen from Yunnan. This species is clearly different from S. nobilis; the legs are reddish, with wings pale as the spots of the tergites, the pronotum dorso-posteriorly (“scapula”) bright yellow laterally, the lateral carina extending over the propodeal spiracle (just reaching to the spiracle in S. nobilis); the median suture of the frons (“frontal fissure”) absent (distinct in S. nobilis); the antenna longer, with penultimate joints nearly as long as wide (antenna distinctly shorter, with penultimate joints wider than long in S. nobilis); and the upper part of the metapleuron with sparse punctures only medially (largely densely punctate in S. nobilis)., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 116, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Okamoto, H. (1924) The insect fauna of Quelpart Island. Bull. Bulletin of the Agricultural Experimental Station of the Government-General of Chosen, I, 1 - 233.","Yamane, S. (1995) On Some Taiwanese and Ryukyu Scoliinae (Hymenopt. era, Scoliidae). Journal of Taiwan Museum, 48 (1), 39 - 46.","Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

10. Megacampsomeris formosensis (Betrem, 1928) Campsomeris (Megacampsomeris) formosensis Betrem, 1928: 150, ♀, China (type in RMNH, Leiden [examined]); Betrem, 1941: 72; Tsuneki, 1972a: 14. Campsomeris formosensis: Uchida, 1933: 257; Uchida, 1936: 61. Megacampsomeris formosensis: Betrem (in Betrem & Bradley), 1972: 164 (referring to Megacampsomeris as a genus); Osten, 2005: 11. Campsomeris (Megacampsomeris) formosensis chinensis Betrem, 1941: 73. Distribution. China (Anhui, Fujian, Guangdong, Guizhou, Hunan, Jiangsu, Shanghai, Taiwan, Zhejiang). Notes. Specimens of the two subspecies M. f. formosensis and M. f. chinensis in RMNH show some interesting differences: basitarsus of hind leg as long as length of tarsomeres 2–4 combined in M. formosensis (hind basitarsus 0.8 times length of tarsomeres 2–4 combined in M. chinensis); setae on metasomal tergites entirely reddish brown (yellow, except blackish on epipygium); spiracular angles nearly glabrous (densely setose). More specimens are necessary to confirm the status of the subspecies M. f. chinensis., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 108, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Uchida, T. (1936) Family Scoliidae. In: Fauna Nipponica. 10 (9). Sanseido, Tokyo, 72 pp. [in Japanese]","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

7. Megacampsomeris binghami (Betrem, 1928) Campsomeris (Megacampsomeris) binghami Betrem, 1928: 143, ♀, China (type in Berlin Museum); Betrem, 1941: 63; Tsuneki, 1972a: 9. Campsomeris binghami: Uchida, 1933: 256; Uchida, 1936: 47; Yasumatsu, 1934: 64; Kontyu, 1937: 294; He et al., 2004: 944. Megacampsomeris binghami: Betrem (in Betrem & Bradley), 1972: 164 (referring Megacampsomeris as a genus); Osten, 2005: 6. Distribution. China (Guangdong, Taiwan, Zhejiang); India and Japan., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 108, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Uchida, T. (1936) Family Scoliidae. In: Fauna Nipponica. 10 (9). Sanseido, Tokyo, 72 pp. [in Japanese]","Yasumatsu, K. (1934) Les Hymenopteres de l'ile Yakushima. Mushi, 1934 (7), 61 - 67.","He, J. H., Chen, X. X., Fan, J. J., Li, Q., Liu, C. M., Lou, X. M. & Xu, Z. F. (2004) Hymenopteran Insect Fauna of Zhejiang. Science Press, Beijing, 1373 pp.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

24. Carinoscolia nipponensis (Uchida, 1933), new status Scolia (Carinoscolia) vittifrons f. nipponensis Uchida, 1933: 242, ♀, Japan (type in Entomological Museum, Sapporo). Scolia nipponensis fascinata var. nipponensis: Betrem, 1941: 114. Carinoscolia vittifrons nipponensis: Osten, 2005: 45. Distribution. China (Hubei, Sichuan); Japan and Korea. Notes. The status of C. nipponensis has been changed several times. Originally, it was described as a variety of C. vittifrons (Sichel) when found by Uchida in Japan for its black head, darker wings, and the very small spots on T 3 in both sexes (Uchida 1933). Betrem (1941) treated it as a variety of C. fascinata Smith by displaying the differences between C. fascinata and C. vittifrons. Osten (2005) treated it as a subspecies of C. vittifrons again. We found a male from Sichuan province of China (IOZ) and a female from Hubei (RMNH) of C. nipponensis, both are different from the nominate C. vittifrons. We consider C. nipponensis a separate species because of the distinct carina present on the area between the antennal sockets (“frontal spatium”) in both sexes and the nearly completely dark integument of both sexes., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 112, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

Subgenus Annulimeris Betrem, 1967 Campsomeriella - subgenus Annulimeris (section Annulimeris) Betrem, 1967: 29. Campsomeriella - subgenus Annulimeris Betrem & Bradley, 1972: 305. Type-species: Tiphia annulata Fabricius, 1793 (by original designation). Distribution. Indo-Australian and Palaearctic Regions., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 104, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. (1967) The natural groups of Campsomeriella Betr., 1941 (Hymenoptera: Scoliidae). Entomologische Berichten, 27, 25 - 29.","Betrem, J. G. & Bradley, J. C. (1972) The African Campsomerinae (Hym., Scoliidae). Mono-grafieen van de Nederlandse Entomologische Vereniging, 6, 1 - 326.","Fabricius, J. C. (1793) Entomologia systematica. Vol. 2. VIII. impensis Christ. Gottl. Proft, Hafniae, 519 pp."]}

42. Scolia (Discolia) rufispina Morawitz, 1889 Scolia rufispina Morawitz, 1889: 115, ♂, China (type in RMNH, Leiden?); Betrem, 1928: 261; Betrem,1935: 5; Betrem, 1941: 168; Osten, 2004: 42; Hua, 2006: 303. Distribution. China (Gansu, Shandong)., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 117, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Morawitz, F. (1889) Insecta, a Cl. G. N. Potanin in China et in Mongolia novissime lecta. IV. Hymenoptera Aculeata. Horae Societatis Entomologicae, 23, 112 - 168.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1935) Beitrage zur Kenntnis der Palaarktischen Arten des Genus Scolia. Tijdschrift voor Entomologie, 78, 1 - 78.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Osten, T. & Arens, W. (2004) Beitrag zur Kenntnis der Scoliiden-Fauna Griechenlands (ohne Zypern) (Hymenoptera, Scoliidae). Entomofauna, 25 (20), 305 - 320.","Hua, L. Z. (2006) List of Chinese Insects. Vol. IV. Sun Yat-sen University Press, Guangzhou, 540 pp."]}

47. Scolia (Discolia) tigrimaculosa Yamane, 1995 Scolia (Discolia) tigrimaculosa Yamane, 1995: 40, ♀, China (type in Kagoshima University, Kagoshima); Osten, 2005: 44. Distribution. China (Taiwan)., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 118, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Yamane, S. (1995) On Some Taiwanese and Ryukyu Scoliinae (Hymenopt. era, Scoliidae). Journal of Taiwan Museum, 48 (1), 39 - 46.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

28. Megascolia (Regiscolia) azurea (Christ, 1791) Scolia azurea Christ, 1791: 256. (Type lost). Scolia rubiginosa Fabricius, 1793: 230; de Saussure & Sichel, 1864: 45. Scolia ornata Lepeletier, 1845: 517. Scolia magnifica de Saussure, 1859: 175. Scolia (Triscolia) rubiginosa: Bingham, 1897: 76–77. Triscolia azurea azurea: Micha, 1927: 117. Scolia (Triscolia) azurea rubiginosa: Betrem, 1928: 231. Scolia (Triscolia) azurea christiana Betrem & Guiglia, 1958: 96. Megascolia (Regiscolia) azurea christiana: Betrem & Bradley, 1964: 444; Gupta & Jonathan, 2003: 45; Kumar & Pham, 2015: 326; Nidup et al., 2017: 13. Distribution. China (Anhui, Fujian, Guangdong, Hainan, Hong Kong, Hunan, Guangxi, Guizhou, Yunnan); Bangladesh, Bhutan, Burma, India, Indonesia, Laos, Malaysia, Myanmar, Nepal, Sri Lanka, Thailand and Vietnam., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 114, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Fabricius, J. C. (1793) Entomologia systematica. Vol. 2. VIII. impensis Christ. Gottl. Proft, Hafniae, 519 pp.","Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Lepeletier, A. (1845) Historie naturelle des insects. Vol. 3. Hymenopteres. Librairie encyclopedique de Roret, Paris, 646 pp.","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Micha, I. (1927) Beitrag zur Kenntnis der Scoliiden (Hym. Acul.) (Liacos Guer., Diliacos Sauss. Sich. und Triscolia Sauss. Sich). Mitteilungen aus dem Zoologischen Museum in Berlin, 13 (1), 1 - 156.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Guiglia, D. & Betrem, J. G. (1958) The identity of the Scoliidae described by J. L. Christ. Annali del Museo Civico di Storia Naturale di Genova, 70, 92 - 99.","Betrem, J. G. & Bradley, J. C. (1964) Annotations on the genera Triscolia, Megascolia and Scolia. Zoologische Mededelingen, 39, 433 - 444. [first part]","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Kumar, P. G. & Pham, P. H. (2015) New distributional records of scoliid wasps (Insecta: Hymenoptera: Scoliidae) from India. Records of the Zoological Survey of India, 115 (4), 325 - 334.","Nidup, T., Klein, W., Girish Kumar, P. & Dorji, P. (2017) New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres, 8, 9 - 19. https: // doi. org / 10.11609 / jott. 3204.9.7.10487 - 10489"]}

14. Megacampsomeris lindenii (Lepeletier, 1845) Campsomeris lindenii Lepeletier, 1845: 500, ♀, Indonesia (plesiotype in Halle Museum); Betrem, 1928: 151; Uchida, 1933: 259; Uchida, 1936: 60; Betrem, 1941: 75; He et al., 2004: 946. Megacampsomeris (Lindenimeris) lindenii: Osten, 2005: 15. Distribution. China (Fujian, Guangxi, Taiwan, Zhejiang); India, Indonesia, Japan, Myanmar., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 109, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Lepeletier, A. (1845) Historie naturelle des insects. Vol. 3. Hymenopteres. Librairie encyclopedique de Roret, Paris, 646 pp.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Uchida, T. (1936) Family Scoliidae. In: Fauna Nipponica. 10 (9). Sanseido, Tokyo, 72 pp. [in Japanese]","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","He, J. H., Chen, X. X., Fan, J. J., Li, Q., Liu, C. M., Lou, X. M. & Xu, Z. F. (2004) Hymenopteran Insect Fauna of Zhejiang. Science Press, Beijing, 1373 pp.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62."]}

50. Scolia (Scolia) concolor Eversmann, 1849 Scolia concolor Eversmann, 1849: 432, ♀, Kazakhstan (type in Zoological Institute, St. Petersburg); de Saussure, 1880: 22; Dusmet, 1930: 36; Betrem, 1935: 47; Lelej & Mokrousov, 2017: 3–4. Scolia (Scolia) concolor: Osten, 2005: 30; Samin et al., 2014: 717; Betrem, 1935: 47. Scolia (Scolia) incana Nagy: Osten, 2005: 1463; Samin et al., 2014: 717. Distribution. China; Iran, Kazakhstan, Kyrgyzstan, Mongolia, Tadzhikistan, Turkey, Turkmenistan and Uzbekistan., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 119, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Eversmann, E. (1849) Fauna hymenopterologica Volgo-Uralensis III. Idem, XXII (2), 430.","Dusmet, J. M. (1930) Himenopteros de la India inglesa cazados por el P. Ignacio Sala de Castellarnau. SJ Boletin de la Sociedad Entomologica de Espana Zaragoza, 1, 99 - 107.","Betrem, J. G. (1935) Beitrage zur Kenntnis der Palaarktischen Arten des Genus Scolia. Tijdschrift voor Entomologie, 78, 1 - 78.","Lelej, A. S. & Mokrousov, M. V. (2017) The types of Scoliidae (Hymenoptera), described by Eduard Eversmann, with some taxonomic notes and checklist of Russian Scoliidae. Far Eastern Entomologist, 340, 1 - 17. https: // doi. org / 10.25221 / fee. 340.1","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Samin, N., Badryacyk, N. & Gadallah, N. S. (2014) A checklist of Iranian Scoliidae (Hymenoptera: Vespoidea). Munis Entomology & Zoology, 9 (2), 713 - 723."]}

49. Scolia (Discolia) wusheensis Tsuneki, 1972 Scolia (Discolia) wusheensis Tsuneki, 1972a: 31, ♂, China (type in Fukui University, Fukui); Yamane, 1995: 45. Scolia (Discolia) wusheensis koreana Tsuneki, 1972a: 32. Distribution. China (Taiwan); Korea., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 118, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Yamane, S. (1995) On Some Taiwanese and Ryukyu Scoliinae (Hymenopt. era, Scoliidae). Journal of Taiwan Museum, 48 (1), 39 - 46."]}

34. Scolia (Discolia) desidiosa Bingham, 1896 Scolia desidiosa Bingham, 1896: 424, ♀, India (holotype in BMNH, London); Yamane, 1995: 43. Scolia (Scolia) decorata desidiosa: Betrem, 1928: 321; Uchida, 1934: 249. Scolia (Scolia) histrionica histrionica: Betrem, 1941: 63. Scolia (Discolia) desidiosa: Bingham, 1897: 86; Tsuneki, 1972a: 24; Gupta & Jonathan, 2003: 243; Kumar, 2017: 6; Nidup et al., 2017: 13. Distribution. China (Taiwan); Bhutan, India and Myanmar., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 116, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Bingham, C. T. (1896) On some Exotic Fossorial Hymenoptera in the Collection of the British Museum, with Descriptions of New Species and of a New Genus of the Pompilidae. Zoological Journal of the Linnean Society, 25 (164), 422 - 445. https: // doi. org / 10.1111 / 0044 - 0124.00164 - i 1","Yamane, S. (1995) On Some Taiwanese and Ryukyu Scoliinae (Hymenopt. era, Scoliidae). Journal of Taiwan Museum, 48 (1), 39 - 46.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Kumar, P. G. & Rajmohana, K. (2017) Fauna of India Checklist. A Checklist of Scoliidae (Insecta: Hymenoptera: Vespoidea) of India. Zoological Survey of India, Kolkata, 9 pp.","Nidup, T., Klein, W., Girish Kumar, P. & Dorji, P. (2017) New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres, 8, 9 - 19. https: // doi. org / 10.11609 / jott. 3204.9.7.10487 - 10489"]}

31. Scolia (Discolia) binotata Fabricius, 1804 Scolia binotata Fabricius, 1804: 244, ♂, Tranquebar (type in Copenhagen Museum); Bradley, 1964: 11; Bradley and Betrem, 1968: 325. Scolia (Discolia) cucullata Bingham, 1897: 82. Scolia quadripustulata: Paiva, 1907: 14. Scolia quadripustulata var. binotata: Bingham, 1908: 352. Scolia (Scolia) quadripustulata: Betrem, 1928: 314. Scolia (Scolia) 4-pustulata var. bipunctata KIug: Betrem, 1928: 316; Guiglia, 1965: 322. Scolia (Scolia) 4-pustulata kansarensis Betrem, 1928: 318. Scolia (Discolia) quadripustulata: Guiglia, 1965: 322. Scolia (Discolia) binotata binotata: Krombein, 1978: 41; Gupta & Jonathan, 2003: 223; Kumar & Pham, 2015: 326; Nidup et al., 2017: 13–14. Distribution. China (Anhui, Beijing, Fujian, Hong Kong, Jiangsu, Jilin, Shangdong, Shanghai, Sichuan, Taiwan, Zhejiang); Bhutan, India, Japan, Laos, Malaysia, Myanmar, Singapore, Sri Lanka and Vietnam., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 115, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Fabricius, J. C. (1804) Systema Piezatorium secundum ordines, genera, species, synonymis, locis, observationibus, descriptionibus. Reichard, Brunsvigae, 439 pp. https: // doi. org / 10.5962 / bhl. title. 10490","Bradley, J. C. & Betrem, J. G. (1968) Friedrich Klugs material of Scoliidae with especial reference to his type specimens. Deutsche Entomologische Zeitschrift, 15, 321 - 334. https: // doi. org / 10.1002 / mmnd. 19680150406","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Paiva, C. A. (1907) Records of Hemiptera and Hymenoptera from Himalayas. Records of the Indian Museum, 1, 13 - 20.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Guiglia, D. (1965) Hymenoptera Scolioidea, Risultati scientifici della spedizione del Dr. Fred Keiserall Isola di Ceylon. Verhandlungen der Naturforschenden Gesellschaft in Basel, 76, 315 - 324.","Krombein, K. V. (1978) Biosystematic studies of Ceylonese Wasps, II: A monograph of the Scoliidae (Hymenoptera: Scolioidea). Smithsonian Contributions to Zoology, 283, 1 - 56. https: // doi. org / 10.5479 / si. 00810282.283","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Kumar, P. G. & Pham, P. H. (2015) New distributional records of scoliid wasps (Insecta: Hymenoptera: Scoliidae) from India. Records of the Zoological Survey of India, 115 (4), 325 - 334.","Nidup, T., Klein, W., Girish Kumar, P. & Dorji, P. (2017) New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres, 8, 9 - 19. https: // doi. org / 10.11609 / jott. 3204.9.7.10487 - 10489"]}

Genus Megascolia Betrem, 1964 Scolia - subgenus Triscolia de Saussure and Sichel: Betrem, 1927: xcvii; Betrem, 1933: 254; Betrem, 1941: 117. Scolia - subgenus Triscolia (section Triscolia) de Saussure and Sichel: Betrem, 1928: 228. Scolia - subgenus Triscolia (section Megascolia Betrem), 1928: 239. Megascolia Betrem (in Betrem and Bradley), 1964: 437. Type-species: Scolia procer Illiger, 1802 (by original designation). Distribution. Oriental Region and Mediterranean Subregion of Palaearctic Region., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 114, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. & Bradley, J. C. (1964) Annotations on the genera Triscolia, Megascolia and Scolia. Zoologische Mededelingen, 39, 433 - 444. [first part]","Betrem, J. G. (1927) Een nieuwe indeeling der Scoliiden. Tijdschrift voor Entomologie, 70 (Verslag), XCIIl - XCVIll.","Betrem, J. G. (1933) Die Scoliiden der indoaustralischen and palaearktischen Region aus dem Staatlichen Museum fur Tierkunde zu Dresden (Hym.). Stettiner Entomologische Zeitung, 94, 236 - 263.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388."]}

2. Campsomeriella (Campsomeriella) collaris (Fabricius, 1798) Scolia collaris Fabricius, 1798: 255, ♂, India (type in Copenhagen Museum). Campsomeris thoracica var. Guérin-Méneville, 1838: 251. Scolia senilis Burmeister, 1853: 24. Elis phalerata de Saussure, 1858: 233. Elis fimbriata Bingham, 1906: 39. Dielis fimbriata Fabr.: Betrem, 1928: 127; Dover, 1926: 234. Campsomeris collaris: de Saussure, 1892: 225. Campsomeris quadrifasciata Betrem: 1928: 336; Betrem, 1932: 413; Betrem, 1933: 242. Campsomeris (Campsomeriella) quadrifasciata: Betrem, 1941: 89; Tsuneki, 1972b: 6. Campsomeris collaris: Betrem, 1947: 413. Campsomeris (Campsomeriella) collaris quadrifasciata: Bradley & Betrem, 1964: 19; Campsomeriella (Campsomeriella) collaris quadrifasciata: Betrem, 1966: 29; Gupta & Jonathan, 2003: 45; Osten, 2005: 8. Campsomeriella (Campsomeriella) collaris collaris: Nidup et al., 2017: 11. Distribution. China (Fujian, Guangdong, Guangxi, Hainan, Jiangxi, Yunnan); Bhutan, India, Indonesia, Malaysia, Myanmar and Philippines., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on pages 104-105, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Fabricius, J. C. (1798) Supplementum Entomologiae Systematicae. Hafniae, Copenhagen, 572 pp.","Guerin-Meneville, F. E. (1838) Insects: In: Duperrey, L. I. & Lesson, R. P. (Eds.), Voyage autour du Monde, execute par ordre du Roi, sur la corvette de S. M. la Coquille, pendant les annees 1822 - 1825. Section 1. Zoology. A. Bertrand, Paris, pp. 1 - 319.","Saussure, H. de (1858) Description de diverses especes nouvelles ou peu connues du genre Scolia. Annales de la Societe entomologique de France, 3 (6), 193 - 249.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Dover, C. (1926) A contribution to a list of the aculeate Hymenoptera (excepting ants) of HongKong. The China Journal of Science & Arts, 233 - 235.","Betrem, J. G. (1932) De Scoliiden-fauna van Banka. Entomologische Berichten, 9, 412 - 415.","Betrem, J. G. (1933) Die Scoliiden der indoaustralischen and palaearktischen Region aus dem Staatlichen Museum fur Tierkunde zu Dresden (Hym.). Stettiner Entomologische Zeitung, 94, 236 - 263.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Tsuneki, K. (1972 b) Some scoliid wasps from South East Asia (Hymenoptera). Etizenia, 63, 1 - 9.","Betrem, J. G. (1947) Analyse van enkele fauna-elementen van de Maleische Scoliiden. Tijdschrift voor Entomologie, 88 (1945), 409 - 416.","Betrem, J. G. & Bradley, J. C. (1964) Annotations on the genera Triscolia, Megascolia and Scolia. Zoologische Mededelingen, 39, 433 - 444. [first part]","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Nidup, T., Klein, W., Girish Kumar, P. & Dorji, P. (2017) New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres, 8, 9 - 19. https: // doi. org / 10.11609 / jott. 3204.9.7.10487 - 10489"]}

6. Megacampsomeris bella (Bingham, 1897) Elis bella Bingham, 1897: 93, ♀, Myanmar (type in BMNH, London). Campsomeris (Megacampsomeris) bella: Betrem, 1928: 140. Megacampsomeris (Bellimeris) bella: Bradley, 1972: 6. Sericocampsomeris bella: Gupta & Jonathan, 2003: 60. Megacampsomeris bella: Osten, 2005: 6. Bellimeris bella: Schulten et al., 2011: 889. Distribution. China (Fujian, Guangxi); India, Indonesia and Myanmar., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 108, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Bradley, J. C. (1972) Scoliid types in the Museum fur Naturkunde of the Humboldt Universitat zu Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 48 (1), 3 - 19. https: // doi. org / 10.1002 / mmnz. 19720480102","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Schulten, G. G. M., Feijen, H. R. & Feijen, C. (2011) The genus Bellimeris Betrem (Hymenoptera: Scoliidae, Campsomerinae). Zoologische Mededelingen, 85, 887 - 903."]}

29. Scolia (Discolia) affinis Guérin-Méneville, 1845 Scolia affinis Guerin, 1838: 254, ♀, Senegal (types in Paris Museum); Dalla Torre, 1897: 144. Scolia (Lacosi) jurinei de Saussure, 1855: 45; Bradley, 1974: 446. Scolia instabilis Smith, 1855: 88; Smith, 1871: 345; Betrem (in Bradley & Betrem), 1967: 30. Scolia (Discolia) aureipennis: Magretti, 1892: 239; Bingham, 1897: 79. Scolia aureipennis: Dalla Torre, 1897: 147; Paiva, 1907: 14. Scolia (Scolia) aureipennis: Betrem, 1928: 280. Scolia (Scolia) jurinei: Betrem, 1941: 136. Scotia (Discolia) jurinei: Betrem & Bradley, 1964: 90; Guiglia, 1965: 320. Scolia (Discolia) affinis: Bradley, 1973: 218; Krombein, 1978: 31; Gupta & Jonathan, 2003: 176. Distribution. China (Fujian, Guangdong, Zhejiang); Bangladesh, India, Myanmar, Nepal and Pakistan., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on pages 114-115, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Dalla Torre, C. G. (1897) s. n. In: Catalogus Hymenoptorum hucusque descriptorum systematicus et synonymicus. VIII. W. Engelmann, Lipsiae (Leipzig), pp. 144 - 188.","Saussure, H. de (1855) Melanges Hymenopterologiques. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 14, 1 - 67.","Smith, F. (1855) Mutillidae and Pompilidae. In: Catalogue of the Hymenopterous Insects in the collection of the British Museum. Taylor and Francis, London, pp. 1 - 206.","Magretti, P. (1892) Imenotteri; Viaggio de Leonardo Fea in Birmanicae Regioni vicini 43, parte prima Mutilledei Scollidei Tiphiidae. Annali del Museo Civico di Storia Naturale Giacomo Doria, 12, 97 - 266.","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Paiva, C. A. (1907) Records of Hemiptera and Hymenoptera from Himalayas. Records of the Indian Museum, 1, 13 - 20.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Betrem, J. G. & Bradley, J. C. (1964) Annotations on the genera Triscolia, Megascolia and Scolia. Zoologische Mededelingen, 39, 433 - 444. [first part]","Guiglia, D. (1965) Hymenoptera Scolioidea, Risultati scientifici della spedizione del Dr. Fred Keiserall Isola di Ceylon. Verhandlungen der Naturforschenden Gesellschaft in Basel, 76, 315 - 324.","Bradley, J. C. (1973) The scoliid types of Guerin-Meneville (Insecta: Hymenoptera: Scoliidae). Bulletin du Museum National D'Histoire Naturelle, Series 3, 112, Zoologie, 86, 217 - 221.","Krombein, K. V. (1978) Biosystematic studies of Ceylonese Wasps, II: A monograph of the Scoliidae (Hymenoptera: Scolioidea). Smithsonian Contributions to Zoology, 283, 1 - 56. https: // doi. org / 10.5479 / si. 00810282.283","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp."]}

Subgenus Campsomeriella Betrem, 1941 Campsomeris s. str. Betrem, 1928: 122. Campsomeris Betrem, 1933: 241. Campsomeriella - subgenus Campsomeriella (section Campsomeriella) Betrem, 1941: 86; Betrem, 1967: 29. Campsomeriella - subgenus Campsomeriella: Betrem & Bradley, 1972: 272. Type-species: Scolia thoracica Fabricius, 1787 (by original designation). Distribution. Afrotropical, Indo-Australian and Palaearctic Regions., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 104, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1933) Die Scoliiden der indoaustralischen and palaearktischen Region aus dem Staatlichen Museum fur Tierkunde zu Dresden (Hym.). Stettiner Entomologische Zeitung, 94, 236 - 263.","Betrem, J. G. (1967) The natural groups of Campsomeriella Betr., 1941 (Hymenoptera: Scoliidae). Entomologische Berichten, 27, 25 - 29.","Betrem, J. G. & Bradley, J. C. (1972) The African Campsomerinae (Hym., Scoliidae). Mono-grafieen van de Nederlandse Entomologische Vereniging, 6, 1 - 326."]}

13. Megacampsomeris limbata (de Saussure & Sichel, 1864) Elis (Dielis) limbata de Saussure, 1864: 206, ♀, Indonesia (plesiotype in RMNH, Leiden). Campsomeris limbata: Uchida, 1936: 56; He et al., 2004: 946. Distribution. China (Fujian, Taiwan, Zhejiang); Indonesia., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 109, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Uchida, T. (1936) Family Scoliidae. In: Fauna Nipponica. 10 (9). Sanseido, Tokyo, 72 pp. [in Japanese]","He, J. H., Chen, X. X., Fan, J. J., Li, Q., Liu, C. M., Lou, X. M. & Xu, Z. F. (2004) Hymenopteran Insect Fauna of Zhejiang. Science Press, Beijing, 1373 pp."]}

20. Phalerimeris phalerata (de Saussure, 1858) Elis (Campsomeris) phalerata de Saussure, 1858: 233, ♀, Indonesia (types in Copenhagen Museum); de Saussure, 1859: 268. Elis (Dielis) phalerata: de Saussure & Sichel, 1864: 201. Elis (Dielis) iris Magretti, 1892: 247; Bingham, 1897: 94. Campsomeris albopilosa Rohwer, 1911: 480. Campsonleris (Dielis) phalerata phalerata: Betrem, 1928: 103; Betrem, 1933: 238; Betrem, 1937: 95; Betrem, 1941: 101. Campsomeris lindenii: Yano, 1932: 317. Campsomeris phalerata: Uchida, 1934: 258. Campsomeris (Phalerimeris) phalerata: Betrem in Bradley & Betrem, 1967: 294; Tsuneki, 1972a: 15. Phalerimeris phalerata phalerata: Bradley, 1974: 460; Gupta & Jonathan, 2003: 68; Kumar & Pham, 2015: 326; Nidup et al., 2017: 11. Distribution. China (Fujian, Guangdong, Hong Kong, Hainan, Hunan, Jiangxi, Taiwan, Yunnan); Bhutan, India, Indonesia, Malaysia, Myanmar, Nepal and Thailand., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 111, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Saussure, H. de (1858) Description de diverses especes nouvelles ou peu connues du genre Scolia. Annales de la Societe entomologique de France, 3 (6), 193 - 249.","Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Magretti, P. (1892) Imenotteri; Viaggio de Leonardo Fea in Birmanicae Regioni vicini 43, parte prima Mutilledei Scollidei Tiphiidae. Annali del Museo Civico di Storia Naturale Giacomo Doria, 12, 97 - 266.","Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma: Hymenoptera, 1 (Wasps and Bees). Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 100738","Rohwer, S. A. (1911) On some hymenopterous insects from the Island of Formosa. Proceeding U. S. National Museum, 39, 477 - 485. https: // doi. org / 10.5479 / si. 00963801.1794.477","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. (1933) Die Scoliiden der indoaustralischen and palaearktischen Region aus dem Staatlichen Museum fur Tierkunde zu Dresden (Hym.). Stettiner Entomologische Zeitung, 94, 236 - 263.","Betrem, J. G. (1937) The males of the Indo-Australian Campsomeris species with yellow metanotum (Hym., Scoliidae). The Proceedings of the Royal Entomological Society of London (B), 6 (5), 91 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1937. tb 00305. x","Betrem, J. G. (1941) Etude systematique des Scoliidae de Chine et leurs relations avec les autres groups de Scoliidae. Notes d'Entomologie Chinoise, 8 (4), 47 - 188.","Bradley, J. C. & Betrem, J. G. (1967) The types of Scoliidae described by Frederick Smith (Hymenoptera). Bulletin of The British Museum (Natural History) Entomology, 1967, 287 - 327.","Tsuneki, K. (1972 a) Studies on the scoliid wasps of eastern Asia (Hymenoptera). Etizenia, 62, 1 - 41.","Gupta, S. K. & Jonathan, J. K. (2003) Fauna of India and the adjacent countries, Hymenoptera: Scoliidae. The Director, Zoological Survey of India, Kolkata, 277 pp.","Kumar, P. G. & Pham, P. H. (2015) New distributional records of scoliid wasps (Insecta: Hymenoptera: Scoliidae) from India. Records of the Zoological Survey of India, 115 (4), 325 - 334.","Nidup, T., Klein, W., Girish Kumar, P. & Dorji, P. (2017) New record of scoliid wasps (Hymenoptera: Scoliidae: Scoliinae) from Bhutan. Halteres, 8, 9 - 19. https: // doi. org / 10.11609 / jott. 3204.9.7.10487 - 10489"]}

38. Scolia (Discolia) minowai Uchida, 1933 Scolia minowai Uchida, 1933: 246, ♂, China (types in Entomological Museum, Sapporo). Scolia minowai: Yamane, 1995: 43. Distribution. China (Taiwan)., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 116, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Uchida, T. (1933) Revision der Japanischen Scoliiden mit Beschreibungen der neuen Arten und Formen. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 32, 229 - 262.","Yamane, S. (1995) On Some Taiwanese and Ryukyu Scoliinae (Hymenopt. era, Scoliidae). Journal of Taiwan Museum, 48 (1), 39 - 46."]}

Genus Colpa Dufour, 1841 Colpa Dufour, 1841: 378, 413, 486; Argaman, 1996: 184; Osten, 2005: 3. Trielis de Saussure, 1863: 18; de Saussure & Sichel, 1864: 140; Ashmead, 1903: 8; Schulz, 1912: 81; Betrem, 1962: 146; Betrem, 1972: 32. Campsoscolia Betrem, 1933: 240, 259 (part); Bradley, 1950: 429. Type species: Scolia interrupta Fabricius, 1781 [= C. sexmaculata (Fabricius, 1781)] (designated by Betrem, 1933). Distribution. Afrotropical, Palaearctic and Nearctic Regions., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 105, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Argaman, Q. (1996) Generic synopsis of Scoliidae (Hymenoptera, Scolioidea). Annls Historico-Naturales Musei Nationalis Hungarici, 88, 171 - 222.","Osten, T. (2005) Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Naturforschende Gesellschaft, 220, 1 - 62.","Saussure, H. de (1863) Sur quelques scolies de Basse-Californie. Annales de la Societe entomologique de France, Series 4, 3, 17 - 19. https: // doi. org / 10.1080 / 00379271.1863.11755423","Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Ashmead, W. H. (1903) Classification of the fossorial, predaceous and parasitic wasps, or the superfamily Vespoidea. The Canadian Entomologist, 35, 3 - 8. https: // doi. org / 10.4039 / Ent 353 - 1","Betrem, J. G. (1962) The taxon Trielis (Hymenoptera: Scoliidae) and its type. Entomological News, 73, 146.","Betrem, J. G. & Bradley, J. C. (1972) The African Campsomerinae (Hym., Scoliidae). Mono-grafieen van de Nederlandse Entomologische Vereniging, 6, 1 - 326.","Betrem, J. G. (1933) Die Scoliiden der indoaustralischen and palaearktischen Region aus dem Staatlichen Museum fur Tierkunde zu Dresden (Hym.). Stettiner Entomologische Zeitung, 94, 236 - 263.","Bradley, J. C. (1950) The most primitive Scoliidae. Eos, Torno extraord., 427 - 437.","Fabricius, J. C. (1781) Species insectorvm exhibentes eorvm differentias specificas, synonyma avctorvm, loca natalia, metamorphosin adiectis observationibvs, descriptionibvs. Vol. 1. Impensis Carol. Ernest. Bohnii, Hamburg, 552 pp. https: // doi. org / 10.5962 / bhl. title. 36509"]}

Genus Scolia Fabricius, 1775 Scolia Fabricius, 1775: 355. Scolia - subgenus Lacosi Guérin-Méneville, 1839: 243; Betrem, 1928: 56. Lisoca Costa, 1858: 8; Betrem and Bradley, 1964: 434. Scolia - subgenus Discolia de Saussure: de Saussure and Sichel, 1864: 55. Scolia - subgenus Scolia Fabricius: Betrem, 1927: xcviii. Scolia Fabricius: Betrem and Bradley, 1964: 89. Type-species: Scolia quadripustulata Fabricius, 1775 (by subsequent designation). Distribution. Cosmopolitan (but mainly subtropical and tropical areas)., Published as part of Liu, Zhen, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2021, A checklist of Scoliidae (Insecta: Hymenoptera) from China, pp. 101-126 in Zootaxa 4966 (2) on page 114, DOI: 10.11646/zootaxa.4966.2.1, http://zenodo.org/record/4736250, {"references":["Fabricius, J. C. (1775) Systema Entomologiae sistens insectorumn, classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensburgi et Lipsiae, 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510","Betrem, J. G. (1928) Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. Treubia, 9 (Supplement), 1 - 388.","Betrem, J. G. & Bradley, J. C. (1964) Annotations on the genera Triscolia, Megascolia and Scolia. Zoologische Mededelingen, 39, 433 - 444. [first part]","Saussure, H. de & Sichel, J. (1864) Catalogue des especes de l'ancien genre Scolia, contenant les diagnoses, les descriptions et la synonymie des especes, avec des remarques explicatives et critiques. V. Mason, Geneve et Paris, 350 pp. https: // doi. org / 10.5962 / bhl. title. 9323","Betrem, J. G. (1927) Een nieuwe indeeling der Scoliiden. Tijdschrift voor Entomologie, 70 (Verslag), XCIIl - XCVIll."]}