Search

A. Key to the extant genera of the subfamily Exothecinae Haliday (modified from Belokobylskij, 1984, 1994b, 2000; van Achterberg, 1983) 1. Prepectal carina present in front of precoxal sulcus; occipital carina complete dorsally and area in front of it rugulose; frons with crenulate median groove; precoxal sulcus very wide............................ Orientocolastes Belokobylskij, 1999 - Prepectal carina completely absent (Fig. 2F); occipital carina usually absent dorsally, if present then area in front of it smooth (Figs 3A, 4C); frons without crenulate median groove; precoxal sulcus absent or narrow (Fig. 2F); but distinctly developed in Colastes (Discolastes Belokobylskij, 2000)................................................................ 2 2. Pronope absent or small, shallow and transverse, rarely medium-sized; clypeus convex............................. 3 - Pronope present, large and deep (Fig. 3F); sometimes obsolescent in Xenarcha; clypeus flattened..................... 4 3. Metasomal tergite V markedly enlarged, bent posteriorly and concealing subsequent tergites; tergites I–V entirely sculptured; male without occipital carina, but large temporal tubercles developed; malar suture distinct.................................................................................................... Vietcolastes Belokobylskij, 1994 - Metasomal tergite V normal; tergites II–V either smooth or sculptured; in both sexes occipital carina developed and temporal tubercles absent; malar suture indistinct or obsolescent..................................... Colastes Haliday, 1833 4. Tergites I–III forming a carapace-like metasoma, coarsely longitudinally striate, and their epipleura less sclerotised than their nota; remaining tergites covered by tergites I-III or largely so; tergite III on posterior margin with more or less distinct transparent border........................................................... Colastinus Belokobylskij, 1984 - Tergites I–III not forming a carapace-like metasoma, with their epipleura and nota equally sclerotised; remaining tergites distinctly exposed, tergite III along posterior margin without transparent border................................... 5 5. Occipital carina complete, ventrally reaching up to base of mandible (Figs 3A, 4C, 6A–E); first subdiscal cell of fore wing elongate (Figs 5B, 5D); fore wing vein 1-M of male widened, as wide as or wider than the length of vein cu-a (Fig. 5D)...................................................................................... Occipitotus gen. nov. - Occipital carina incomplete, dorsally interrupted; first subdiscal cell of fore wing broad; fore wing vein slender.......... 6 6. Metasomal tergites I–V entirely sculptured; apical tergites not protruding (or only slightly) beyond tergite V; tergites IV and V with a lateral crease.................................................... Pseudophanomeris Belokobylskij, 1984 - Metasomal tergites less sculptured, at most tergite III completely and base of tergite IV sculptured; apical tergites distinctly protruding beyond tergite V; tergites IV and V without a lateral crease........................................... 7 7. Posterior half of notauli absent or obsolescent; vein r of fore wing usually departing between basal third and base of pterostigma; if departing near basal 0.4 of pterostigma, then vein m-cu of fore wing distinctly converging to vein 1-M posteriorly................................................................................. Shawiana van Achterberg, 1983 - Posterior half of notauli distinctly impressed, (nearly) complete; vein r of fore wing departing from basal 0.4–0.6 of pterostigma and vein m-cu of fore wing less converging posteriorly.................................... Xenarcha Foerster, 1863, Published as part of Singh, Longjam Roni Kumar, Achterberg, C. Van & Sheela, S., 2022, Studies on the subfamily Exothecinae (Hymenoptera: Braconidae) with the description of a new genus and a new species from India, pp. 40-52 in Zootaxa 5133 (1) on pages 41-42, DOI: 10.11646/zootaxa.5133.1.2, http://zenodo.org/record/6521264, {"references":["Belokobylskij, S. A. (1984) On the division of the tribe Exothecini s. l. (Hymenoptera, Braconidae) in two with the description of a new genus and subgenus. Zoologicheskii Zhurnal, 63, 1019 - 1026. [in Russian with English summary]","Belokobylskij, S. A. (1994 b) To the fauna of Indo - Malayan braconid wasps of the tribes Exothecini, Pambolini and Pentatermini (Hymenoptera, Braconidae). Proceedings of the Zoological Institute of the USSR Academy of Sciences, 245, 125 - 173. [In Russian]","Belokobylskij, S. A. (2000) New taxa of the subfamily Exothecinae (Hymenoptera, Braconidae) from tropical and subtropical regions of the Old World. II. Entomologicheskoe Obozrenie, 79 (1), 180 - 203. [in Russian]","van Achterberg, C. (1983) Revisionary notes on the Palaearctic genera and species of the tribe Exothecini Foerster (Hymenoptera, Braconidae). Zoologische Mededelingen, Leiden, 57, 339 - 355.","Belokobylskij, S. A. (1999) New genera of the subfamilies Rhyssalinae, Exothecinae and Gnamptodontinae from the Old World (Hymenoptera: Braconidae). Zoosystematica Rossica, 8 (1), 155 - 169.","Haliday, A. H. (1833) An essay on the classification of the parasitic Hymenoptera of Britain, which correspond with the Ichneumones minuti of Linnaeus. Entomological Magazine, 1 (iii), 259 - 276, 333 - 350.","Foerster, A. (1863) Synopsis der Familien und Gattungen der Braconen. Verhandlungen des Naturhistorischen Vereines der Preussischen Rheinlande und Westphalens, 19, 225 - 288. [1862]"]}

Occipitotus langpramensis Singh & van Achterberg sp. nov. (Figures 2–6) Holotype, female, length of body 3.1 mm and of fore wing 2.9 mm and hind wing 2.5 mm. Head.Antennae with 27–28 flagellomeres (left 28 and right 27), first flagellomere 1.1 times second flagellomere, length of first, second, third and penultimate flagellomeres 5.0, 4.4, 4.0 and 3.5 times their width, respectively (Figs 2C, 2D); terminal flagellomere acuminate; head in dorsal view slightly transverse, 1.4 times as broad as long (Fig. 3A) and 1.15 times wider than the width of mesoscutum; length of eye 1.9 times length of temple in dorsal aspect; OOL (ocular ocellar line): diameter of ocellus: POL (posterior ocellar line) = 9.5:3:4.5; vertex, temple smooth and conspicuously setose; frons smooth; face finely granulate, medially obscurely scabriculous (Fig. 2E); width of face: width of head: height of eye = 3.9: 7.3: 3.7; occipital carina complete (Fig. 3A) and reaching base of mandible, occipital flange rather wide (Fig. 6A); hypostomal carina raised into an elevated flange (Fig. 6A), its height almost equal to length of third segment of labial palp; clypeus slightly convex, dorsally smooth and ventral rim of clypeus finely foveate; hypoclypeal depression small, less than half as wide as face (Fig. 2E); malar space as wide as basal width of mandible and 0.4 times as long as the maximum width across mandibles; malar space smooth, anteriorly granulate; first tooth of mandible distinctly longer than second, both teeth of mandible acute, striate on outer side; length of maxillary palp 1.3 times height of head. Mesosoma. Length of mesosoma 2.0 times its height; pronope deep and large; a narrow anterior marginal pronotal sulcus crenulate; side of pronotum smooth to finely rugulose, with crenulate transverse depression (Fig. 2F); propleuron smooth, shiny and striate-rugose anteriorly; mesoscutum glabrous, shiny and anterior margin with a well-developed sulcus; notauli finely crenulate, posteriorly not strongly converging and ending in broad rugose, slightly depressed setose area (Fig. 3B); scutellar sulcus wide, deep, 0.3 times length of scutellum and rugose with a mid-longitudinally carina; scutellum weakly convex, setose and rugulose but medio-dorsally smooth; subalar depression smooth; mesopleuron largely smooth but rugose antero-dorsally and conspicuously setose ventrally (Fig. 2F); mesosternum with a foveate mid-longitudinal sulcus (mesosternal sulcus) and setose; mesopleural (epicoxal) sulcus extends dorsally and crenulate; metanotum largely smooth with a short fine streak antero-medially and posteriorly with a strongly raised median area and rugulose; propodeum and metapleuron coarsely rugose-reticulate; propodeum with a weak irregular median carina anteriorly. Wings. Fore wing (Fig. 5B): Length 3.65 times its maximum width; angle between veins 1-SR and C+SC+ R ≈ 35°; marginal cell long, vein SR1 almost reaching the wing margin, 0.95 times of distance between apex of pterostigma and wing tip; length of veins r: 3-SR: SR1 = 1.3: 4.0: 6.9; 1-SR+M evenly curved posteriorly, 0.8 times as long as 1-M; second submarginal cell narrow, length of veins 2-SR: 3-SR: r-m = 2.5: 4.0: 1.5; vein cu-a postfurcal and first subdiscal cell elongate; length of veins 1-CU1: 2-CU1: cu-a = 2.2: 7.0: 0.7; 3-CU1: CU1b = 2.2: 1.5. Hind wing (Fig. 5C): Length 5.7 times its maximum width; length of veins SC+ R 1: 2-SC+ R: 1r-m = 4.0: 0.9: 1.1; m-cu present, antefurcal; M+CU: 1-M: 1r-m = 1.2: 1.4: 0.35; 1r-m straight. Legs. Legs slender and relatively long; length of femur, tibia and tarsus of foreleg = 5.4: 7.0: 7.6; length of femur, tibia and basitarsus of hind leg = 6.5: 10.6: 4.4, 4.3, 9.6 and 6.1 times longer than their maximum width, respectively; tibia as long as tarsus; hind coxa medium-sized, without a basoventral tooth, 1.8 times longer than wide, 0.55 times as long as hind femur; length of inner and outer spurs of hind tibia 0.22 and 0.18 times as long as hind basitarsus, respectively; tarsal claws simple. Metasoma. Length of tergite I 1.2 times its apical width, its dorsal surface coarsely rugose-reticulate; its dorsal carinae not joined with weak mid-longitudinal carina (Fig. 3C); dorsope distinct, with continuous crenulate depression posteriorly; laterope absent; tergite II longitudinally striate (nearly complete); second metasomal suture weak, smooth; tergite III basally with weak oblique short striate; remaining tergites entirely smooth, sparsely setose; length of setose part of ovipositor sheath 0.7 times as long as tergite I; ovipositor with dorsal nodus, ventral valves with 3 ventral serrations (Figs 3D, 3E). Colour. Black; antenna dark brown except scapus (ventrally) and annellus yellowish-brown; mandible pale brownish, apical teeth darker; palpi, tegulae, wing veins, metasomal tergites VII and VIII yellowish-brown; pterostigma, parastigma and vein 1-M slightly darker; wing membrane subhyaline; legs yellowish, but apical third of hind tibia and tarsus infuscate; telotarsus darker and hind coxa basally dark brown to yellowish; ovipositor pale yellowish, but apically yellowish-brown. Variations. Length of body 2.6–3.2 mm (female) or 2.0– 2.8 mm (male), and of fore wing 2.2–2.9 mm (female) or 1.8–2.5 mm (male); antenna of female with 25 (1), 27 (3), 28 (2), 29 (2) or 30 (1) flagellomeres, and of male with 23 (1), 24 (3), 25 (4), 26 (6), 27 (6) or 28 (5) flagellomeres; occipital carina complete and continues up to base of the mandible (8 females; 24 males) (Figs 6B–E) or interrupted ventrally above base of mandible (1 female, holotype; 2 males) (Fig. 6A); area between occipital and hypostomal carinae smooth (Figs 6A, 6B, 6E); a short carina running close to occipital carina (Fig. 6C), or with oblique or transverse carinae and weakly crenulate (Fig. 6D); face largely smooth or finely granulate; side of pronotum largely smooth in males, with crenulate transverse depression; propodeum coarsely rugose-reticulate, with weak median carina anteriorly (Fig. 4D) or entire; vein 3- SR of fore wing 1.6–1.9 times as long as vein 2-SR in females and that of males 1.6–1.7 times; fore wing vein 1-M of male widened, as wide as or wider than the length of vein cu-a (Fig. 5D); metasomal tergite I with (feeble) or without median carina; tergite II longitudinally striate (nearly) completely (in female) (Figs 3C, 4A) or anterior half striate (mostly in males) (Fig. 5A); second metasomal suture weak (Figs 3C, 4A) or obsolescent (Fig. 5A); length of ovipositor sheath 0.5–0.7 times as long as tergite I; metasomal tergites largely black and sclerotized, but in some males, tergites III and remaining comparatively less sclerotized and dark brown in colour. Distribution. India. Type material. Holotype female: INDIA: Manipur, Tamenglong Dist., Langpram Village, Ariangiuky stream, 25°09’27.0”N 93°33’35.0”E; appx. 363 mt. a.s.l., 3.iv.2019, sweep net on Equisetum sp. (Equisetaceae), col. L. R. K. Singh (NZC, ZSI, Kolkata; Registration No. 26520/H3); paratypes: 9 females (NZC, ZSI, Kolkata; Registration Nos. 26521-26529/H3), 26 males (NZC, ZSI, Kolkata; Registration Nos. 26530-26555/H3), data same as the holotype. Etymology. The species is name after its type locality, Langpram Village, Tamenglong Dist., Manipur., Published as part of Singh, Longjam Roni Kumar, Achterberg, C. Van & Sheela, S., 2022, Studies on the subfamily Exothecinae (Hymenoptera: Braconidae) with the description of a new genus and a new species from India, pp. 40-52 in Zootaxa 5133 (1) on pages 43-49, DOI: 10.11646/zootaxa.5133.1.2, http://zenodo.org/record/6521264

Occipitotus Singh & van Achterberg gen. nov. (Figures 2–6) Type species: Occipitotus langpramensis Singh & van Achterberg sp. nov. Etymology. The generic name Occipitotus is masculine and refers to the complete occipital carina on the head; “occipit” stands for occipital carina and “totus” (Latin for “complete”). Diagnosis. Antenna with 22–30 flagellomeres (25–30 in female; 22–28 in male), longer than fore wing (Figs 2B, 4B), scapus small, subglobose, shorter ventrally than dorsally; occipital carina complete (Figs 3A, 4C), ventrally straight or slightly curved reaching base of mandible (rarely reduced near base of mandible) and raised into a flange; remaining widely separated from hypostomal carina (Figs 6A–E); hypoclypeal depression deep and small (Fig. 2E); clypeus slightly convex or flat, not protruding beyond the level of the face (Fig. 2A); malar suture obsolescent; pronope large and deep (Figs 3F, 4C); mesoscutum glabrous and shiny; notauli finely crenulate, terminating into rugose area posteriorly (Figs 3B, 4D); vein r of fore wing distinctly oblique and arising from basal 0.4–0.5 of pterostigma; first subdiscal cell of fore wing elongate (Figs 5B, 5D); fore wing vein 1-M of male widened, as wide as or slightly wider than length of cu-a (Fig. 5D); propodeum coarsely rugose-reticulate, with weak median carina anteriorly (Fig. 4D) or entire; dorsal carinae of metasomal tergite I not united medially and tergite coarsely rugosereticulate with (feeble) (Fig. 3C) or without (Fig. 3G) median carina; metasomal tergite II longitudinally striate (nearly) complete (Figs 3C, 4A) or anterior half striate (Fig. 5A); tergite II slightly longer than tergite III, feebly striate antero-laterally (Fig. 4A) or smooth and shiny (Fig. 5A); tergites IV–VI smooth and sparsely setose (Figs 3C, 4A); length of ovipositor sheath 0.5–0.7 times as long as first tergite (Figs 2B, 4A). Biology. Host unknown, but collected from horsetails, Equisetum sp. (Equisetaceae; Fig. 7B) along with an unidentified species of Coleoptera (Chrysomelidae; Fig. 7A); considering the biology of other species of the members of this subfamily Exothecinae, it is likely to be an ectoparasitoid of coleopterous larvae on Equisetum sp. Distribution. Oriental.

Tribe Adeliini Viereck, 1918 Diagnosis. Head transverse, sculptured (Figs 1B, 3D, 5B, 7B, 9C, 11K, 12I, 13B, 15C, 17B); ocelli small; frons flat or concave medially, with or without midlongitudinal groove (Figs 1C, 4C, 5C, 7C, 9D, 11C, 12C, 13C, 15D, 17C); eyes setose (Figs 1B, 3D, 5B, 7B, 12I, 13B, 15C, 17B) or glabrous (Figs 9C, 11K); malar suture complete and deep (Figs 1E, 3D, 5D & E, 7D, 10B & E, 11I & K, 15E); occipital carina often complete, joining hypostomal carina ventrally (Figs 4A, 6A, 8A, 10A, 14A, 16A), rarely absent ventrally (Figs 1A & E) (in Carinadelius gen. nov.) without joining hypostomal carina; palps short and thick, maxillary palp often 5-segmented (Figs 1E, 4A, 13E) rarely 3-segmented (Fig. 12F) (in Paradelius), labial palp 3-segmented; antenna more or less thickened, 20–22 antennomeres; scape rather long, wide; pedicel short; first flagellomere longer than second; subapical flagellomeres of female quadrate in several species (Figs 2A, 5A, 7C); tentorial pits deep (Figs 1B, 3D, 5B, 7B, 9C, 11K, 12I, 13B, 15C, 17B); clypeus smooth or sculptured (Figs 1B, 3D, 5B, 7B, 9C, 11K, 12I, 13B, 15C, 17B); labrum concealed by clypeus (Figs 1B, 3D, 5B, 7B, 9C, 11K, 12I, 13B, 15C, 17B); mesosoma often dorso-ventrally flattened (Figs 1E, 4B, 6B, 7F, 10B, 11I, 13E, 15E, 17E); propleuron short, crenulate posteriorly (Figs 1E, 4B, 6B, 7F, 10B, 11I, 13E, 15E, 17E); propleural lobe distinct or indistinct (Figs 1E, 4B, 6B, 7F, 10B, 11I, 13E, 15E, 17E); mesoscutum smooth to sculptured (Figs 1D, 4C, 5F, 7E, 9F, 11L, 12B, 13D, 15D, 17C); notauli absent (Figs 1D, 4C, 5F, 7E, 9F, 11L, 12B, 13D, 15D, 17C); prepectal carina often absent (Figs 1E, 4B, 6B, 7F, 10B, 11I, 13E, 15E), rarely present (Fig. 12H, 17E) (in Sinadelius and Paradelius); postpectal carina absent (Fig. 9E); scutellar sulcus narrow, crenulated (Figs 1D, 4C, 5F, 7E, 9F, 11L, 12B, 13D, 15D, 17C); mesopleuron convex, smooth or sculptured (Figs 1E, 4B, 6B, 7F, 10B, 11I, 12H, 13E, 15E, 17E); precoxal sulcus smooth or crenulate entirely (Figs 1E, 4B, 6B, 7F, 10B, 11I, 12H, 13E, 15E, 17E); metapleuron smooth (Figs 1E, 7F, 10B, 13E, 17E) or rugose (Figs 4B, 6B, 15E); propodeum smooth or sculptured, often areolate, either divided into distinct anterior and posterior part or not (Figs 1F, 4D & E, 6C, 8B, 10C & D, 11D, 12B, 13F, 14B, 16B, 17F, 18A), rarely with distinct midlongitudinal carina and vertical lateral carina posteriorly (Fig. 1F) (in Carinadelius gen. nov.); propodeal spiracles round; fore wing hyaline (Figs 7A, 10E, 11A, 12A, 14C, 18C) or infuscated (Figs 2C, 4F, 6D, 16C); pterostigma wide; vein 1-R1 distinct, long (Figs 2C, 4F, 6D, 10E, 12A, 14C, 18C) to nearly absent (Figs 7A, 9A, 11A); fore wing vein r present (Figs 7A, 9A, 11B & F) or absent (Figs 2C, 4F, 6D, 12A, 14C, 18C); vein r-m absent (Figs 2C, 4F, 6D, 7A, 10E, 11A, 12A, 14C, 16C, 18C); vein SR1 not reaching wing margin; vein 1SR+M either connected with parastigma or 1-M; vein m-cu often postfurcal or interstitial (Figs 2C, 4F, 6D, 7A, 10E, 11A, 12A, 14C, 16C, 18C); vein M+CU straight or distinctly curved (Figs 2C, 4F, 6D, 7A, 10E, 11A, 12A, 14C, 16C, 18C); vein cu-a antefurcal, interstitial or postfurcal (Figs 2C, 4F, 6D, 7A, 10E, 11A, 12A, 14C, 16C, 18C); subdiscal cell open; hind wing with 3 hamuli; basal cell of hind wing narrow; mid coxa with transverse groove dorsally (Figs 4B, 6B, 7F, 16D); hind coxa smooth or rugose; hind femur and tibia flattened; hind tibial spurs long, inner spur approx. 0.5 × as long as basitarsus; hind basitarsus long, 0.8–0.9 × as long as combined length of 2 nd– 5 th tarsomeres; tarsal claws short, simple (Fig. 12E); metasoma smooth or sculptured (Figs 2B, 4E, 6C, 8B, 10D, 11D, 12G, 14B, 16B, 18A), often compressed dorso-ventrally; metasomal tergites 1–3 formed into carapace, smooth or rugose; first and second metasomal suture present (Figs 12G, 14B, 16B, 18A) or absent (Figs 2B, 4E, 6C, 8B, 10D, 11D, 16B); spiracle of metasomal tergites 6–7 absent; ovipositor short, always less than 0.25 × as long as hind tibia. Genera included. Adelius Haliday, Carinadelius Ranjith & van Achterberg, gen. nov., Myriola Shestakov reinstated, Paradelius de Saeger, Sinadelius He & Chen, and Sculptomyriola Belokobylskij. Distribution. Cosmopolitan. Biology. Solitary endoparasitoids of leaf-mining Nepticulidae. Comments. Shimbori et al. (2019) discussed possible apomorphic and synapomorphic character states between Cheloninae and Adeliini. Apart from the synapomorphic characters, the number of antennomeres is considered as plesiomorphic character to some extent (E.M. Shimbori, personal communication). Within the tribe it has been also found (Kittel & Austin 2014; Kittel et al. 2015, 2016) that the division of the propodeum into distinct anterior and posterior parts (in Adelius, Sinadelius and Sculptomyriola) is a plesiomorphic character as it is absent in other genera (in Carinadelius gen. nov., Myriola and Paradelius). It is suggested that the new genus Carinadelius is probably derived from the genus Adelius., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on pages 2-3, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470, {"references":["Shimbori, E. M., Bortoni, M. A., Shaw, S. R., Souza-Gessner, C. D. S., Cerantola, P. D. C. M. & Penteado-Dias, A. M. (2019) Revision of the New World genera Adelius Haliday and Paradelius de Saeger (Hymenoptera: Braconidae: Cheloninae: Adeliini). Zootaxa, 4571 (2), 151 - 200. https: // doi. org / 10.11646 / zootaxa. 4571.2.1","Kittel, R. N. & Austin, A. D. (2014) Synopsis of Australian chelonine wasps (Hymenoptera: Braconidae: Cheloninae) with description of two new genera. Austral Entomology, 53, 183 - 202. https: // doi. org / 10.1111 / aen. 12070","Kittel, R. N., Jennings, J. T. & Austin, A. D. (2015) Systematics of the parasitic wasp genus Phanerotomella Szepligeti (Hymenoptera: Braconidae: Cheloninae) for Australia, with descriptions of 18 new species. Insect Systematics & Evolution, 46, 301 - 357. https: // doi. org / 10.1163 / 1876312 X- 45032120","Kittel, R. N., Austin, A. D. & Klopfstein, S. (2016) Molecular and morphological phylogenetics of chelonine parasitoid wasps (Hymenoptera: Braconidae), with a critical assessment of divergence time estimations. Molecular Phylogenetics and Evolution, 101, 224 - 241. https: // doi. org / 10.1016 / j. ympev. 2016.05.016"]}

Carinadelius Ranjith & van Achterberg gen. nov. Figs 1–2 urn:lsid:zoobank.org:act: 6A7F6D98-D1D5-40D6-AFBE-0E3D788FD5CD Type species: Carinadelius medicus Ranjith & van Achterberg sp. nov. Description. Female. Head. Antenna with 21 antennomeres (Fig. 2A). Scape without modifications, longer than wide (Fig. 2A). Pedicel short (Fig. 2A). Terminal antennomere blunt (Fig. 2A). Antennomeres 3–8 longer than wide, rest as long as wide (Fig. 2A). Head slightly transverse in anterior view (Fig. 1B). Eyes long setose (Figs 1B & C). Labrum concealed by clypeus (Fig. 1B). Malar suture present (Fig. 1B). Head wider than long in dorsal view (Fig. 1C). Ocelli small, arranged in equilateral triangle (Fig. 1C). Frons slightly concave medially (Fig. 1C). Occipital carina complete dorsally (Fig. 1C), absent ventrally, not connected with hypostomal carina (Figs 1A & E). Vertex faintly crenulate medially near occipital carina (Fig. 1C). Mandible curved (Fig. 1B). Face weakly convex in lateral view (Fig. 1E). Tentorial pit rather small (Fig. 1B). Palps short. Maxillary palp 5-segmented. Labial palp 3-segmented (Fig. 1E). Mesosoma. Mesosoma dorso-ventrally flattened (Fig. 1E). Propleuron crenulate posteriorly and laterally (Fig. 1E). Propleural lobe present (Fig. 1E). Mesoscutum smooth, setose, with indistinct lateral carina and distinct posterior transverse carina (Fig. 1D). Notauli absent (Fig. 1D). Scutellar sulcus arch-shaped, crenulated (Fig. 1D). Scutellum smooth, sparsely setose (Fig. 1D). Lateral sides of scutellum distinctly crenulate (Fig. 1D). Metanotum smooth (Fig. 1D). Mesopleuron smooth (Fig. 1E). Precoxal sulcus long, smooth sometimes weakly crenulate (Fig. 1E). Metapleuron smooth with distinct medial pit (Fig. 1E). Propodeum with distinctly marginated smooth areas (Fig. 1F). Propodeum slightly curved in lateral view, not differentiated into anterior and posterior part (Figs 1E & F). Posterior side of propodeum with distinct midlongitudinal carina and vertical lateral carina (Fig. 1F). Medial transverse carina interrupted by areola (Fig. 1F). Propodeal areola pentagonal, slightly sculptured anteriorly (Fig. 1F). Propodeal spiracles round (Fig. 1F). Wings. Fore wing: fore wing infuscated below pterostigma and near vein cu-a (Fig. 2C). Pterostigma wide (Fig. 2C). Vein 1SR+M connected with vein 1-M (Fig. 2C). Vein M+CU distinctly curved (Fig. 2C). Vein cu-a postfurcal (Fig. 2C). Vein r absent (Fig. 2C). Vein r-m absent (Fig. 2C). Vein m-cu postfurcal (Fig. 2C). Veins 2-SR and SR1 connected with pterostigma (Fig. 2C). Vein 1-R1 long (Fig. 2C). Veins 2-1A and CU1b absent (Fig. 2C). Subdiscal cell open (Fig. 2C). Hind wing: hind wing with 3 hamuli. Basal cell narrow. Vein M+CU longer than 1-M. Vein 1-1A absent. Legs. Middle coxa with dorsal transverse groove (Fig. 1E). Hind coxa smooth (Fig. 1E). Hind femur and tibia distinctly flattened (Fig. 1A). Hind tibial spurs long. Hind basitarsus long. Tarsal claw simple. Metasoma. Metasoma smooth, 6-segmented (Fig. 2B). Tergites 1–3 immovably joined (Fig. 2B). First tergite distinctly notched antero-medially (Fig. 2B). First and second metasomal suture absent (Fig. 2B). Ovipositor sheath exerted, setose apically. Ovipositor without dorsal nodus and ventral serrations. Etymology. Name formed by combining the generic name Adelius and ‘carina’, because of the midlongitudinal carina on the propodeum and the incomplete occipital carina. Gender: masculine. Distribution. Oriental region (India). Comparative diagnosis. The ventrally reduced occipital carina and the midlongitudinal carina and vertical lateral carina posteriorly of the propodeum are considered to be apomorphic characters of the new genus. In addition, the new genus exhibits another peculiar character, its propodeum is not divided into anterior and posterior parts by means of a complete transverse carina which separates it from other genera. This carina may be reduced in Adelius. These characters combined separate Carinadelius from other genera within the tribe. In the keys provided by Belokobylskij (1988, 1998), He & Chen (2000), Shimbori et al. (2019) Carinadelius comes close to Adelius because of the smooth metasoma and evenly fused metasomal tergites 1–3. The combination of the smooth metasoma and absence of first and second metasomal sutures places this genus within the group containing Adelius and Myriola. The smooth marginated areas on the propodeum found in Carinadelius are different from those in Adelius + Myriola. Apart from the apomorphic characters Carinadelius exhibits some other interesting characters like absence of fore wing vein r and position of vein cu-a. Carinadelius differs from Adelius in having the unique carination of the propodeum (propodeum without midlongitudinal carina posteriorly in Adelius), ventrally absent occipital carina which is not connected with hypostomal carina (occipital carina complete, connected with hypostomal carina in Adelius), fore wing vein M+CU distinctly curved (more or less straight in Adelius), vein r absent (present or absent in Adelius). Carinadelius is also similar to Myriola in having an undivided propodeum. However, it differs from Myriola in having the following characters: fore wing vein 1-R1 present and long (vein 1-R1 absent in Myriola), malar space distinctly longer than basal width of mandible (distinctly shorter than basal width of mandible in Myriola), fore wing vein r absent (present in Myriola), fore wing infuscated (hyaline in Myriola). The smooth metasoma in Carinadelius separates it from the genera Paradelius, Sculptomyriola and Sinadelius in which metasomal tergites 1–3 are sculptured. Even though other adeliine genera exhibit a fixed number of antennomeres (22 in Sinadelius and 20–21 in other genera), it is found that number of antennomeres is highly variable at least in the genus Adelius (E.M. Shimbori, personal communication).

Ranjith, A.P., Achterberg, C. Van, Samartsev, K.G., Nasser, M. (2021): Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India. Zootaxa 4926 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.4926.1.1

Carinadelius medicus Ranjith & van Achterberg sp. nov. urn:lsid:zoobank.org:act: 14CAFC1F-18D8-4E5C-A0E8-F4B957764516 Material examined. Holotype, female, “ INDIA: Kerala, Thiruvananthapuram, JNTBGRI, 06.i.2020, sweep net, coll. Ranjith, A.P. ”. Paratypes: 2 females with same data as holotype, except 07.i.2020, 2 males with same collection data except 08.i.2020 (DZUC). Description, holotype, female. Body length 4.0 mm, fore wing 3.2 mm Head. Head 1.2 × as wide as long in anterior view and dorsal view. Face smooth setose (Fig. 1B), 1.5 × as wide as long. Ratio of height of clypeus: intertentorial distance: tentorio-ocular distance = 1: 2.1: 1.3. Length of malar space 1.6 × basal width of mandible. Length of eye 1.4 × as long as temple in dorsal view. Frons, vertex and occiput smooth, setose except frons with a shallow depression medially (Fig. 1C). Ratio of POL: transverse diameter of ocellus: OOL = 1.1: 1: 2.6. Antenna as long as body. Scape 1.5 × as long as maximum width. Pedicel as long as wide. Third antennomere 1.1 × as long as fourth antennomere. Third, fourth and terminal antennomeres 1.9, 1.7 and 2.1 × as long as maximum width respectively. Mesosoma. Mesosoma 1.9 × as long as high. Scutellar sulcus divided by 11 crenulae (Fig. 1D). Propodeal areola 1.9 × as long as maximum width. Wings. Fore wing 1.1 × as long as maximum width. Pterostigma 2.7 × as long as wide. Vein 1-R1 as long as pterostigma. Vein m-cu 0.27 × as long as vein 1-M. Vein 2-SR 1.3 × as long as vein r. Vein r 1.2 × as wide as pterostigma. Legs. Hind femur, tibia and basitarsus 4.0, 4.3 and 5.5 × as long as maximum width, respectively. Metasoma. Metasoma 0.6 × as long as head and mesosoma combined. Metasomal tergites smooth, sparsely setose (Fig. 2B). Hypopygium acute apically. Ovipositor sheath 0.2 × as long as hind tibia. Colour. Body yellow to brownish yellow, except scape, pedicel, antennomeres 1–4, maxillary and labial palps, fore and mid legs, hind coxa, trochanter, femur and metasomal tergite 1–3 yellow; eyes, ocelli, antennomeres 5–19, sides and posterior end of scutellum, pterostigma, wing membrane below pterostigma, junction of fore wing veins 1-M, Cu1 and cu-a, hind femur apically, hind tibia in apical 2/3 rd, basitarsus, claw and metasomal tergites 4–6 brown. Etymology. Named after Friedrich Kasimir Medikus (1738–1808), a German physician and botanist. We dedicate this species with gratitude to all doctors and nurses for their timeless and uncompromising efforts to control COVID-19. Male. Same as female, except for slenderer antenna. Distribution. India (Kerala). Biology. Unknown., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on pages 6-7, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470

Sinadelius He & Chen, 2000 Figs 17–18 Type species: S inadelius guangxiensis He & Chen, 2000. Diagnosis. Antenna of female slender submedially, with 22 antennomeres; prepectal carina present (Fig. 17E); vein 1-SR of fore wing absent (Fig. 18C); maxillary palp with 3 segments; dorsal areola of propodeum medium-sized (Figs 17F, 18A); second metasomal tergite smooth (Figs 17F, 18A) (except basally); first metasomal suture deep, distinctly sculptured (Figs 17F, 18A); second metasomal suture indistinct and smooth (Figs 17F, 18A). Distribution. Oriental, East Palaearctic. Included species. Sinadelius guangxiensis He & Chen, 2000, S. nigricans He & Chen, 2000., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on pages 22-24, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470, {"references":["He, J. H., Chen, X. X. & Ma, Y. (2000) Hymenoptera Braconidae. Fauna Sinica. Insecta, 18, 1 - 757."]}

Paradelius de Saeger, 1942 Figs 12–14 Type species: Paradelius ghesquierei de Saeger, 1942. Diagnosis. Antenna of female slender submedially, with 20 antennomeres; maxillary palp with 3–5 segments (Figs 12F, 13B & E) (3 in type species); clypeus almost as wide as face (Figs 12I, 13B); eyes short setose (Figs 12I, 13B); prepectal carina present (Fig. 12H); vein 1-SR of fore wing absent (Figs 12A, 14C); dorsal areola of propodeum wide (Figs 12B, 13F); first and second metasomal tergites largely longitudinally sculptured (Figs 12G, 13F, 14B); first and second metasomal sutures distinct and crenulate (Fig. 12G). Distribution. Afrotropical, East Palaearctic, Oriental. Included species. Paradelius chinensis He & Chen, 2000, P. ghesquierei de Saeger, 1942, P. ussuriensis Belokobylskij, 1988., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on pages 17-18, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470, {"references":["He, J. H., Chen, X. X. & Ma, Y. (2000) Hymenoptera Braconidae. Fauna Sinica. Insecta, 18, 1 - 757.","Belokobylskij, S. A. (1988) The subfamily Adeliinae (Hymenoptera, Braconidae) in the Far East of the USSR. Trudy Vsesoyuznogo Entomologicheskogo Obshchestva, 70, 144 - 152. [in Russian]"]}

Adelius Haliday, 1833 Figs 3–6 Type species: Adelius subfasciatus Haliday, 1833. Synonyms: Acaelius Haliday, 1834 (invalid correction); Acoelius Haliday, 1835 (lapsis calami); Anomopterus Rohwer, 1914; Pleiomerus Wesmael, 1835; Pseudoscapiola Perepetchayenko, 1994 (syn. nov.). Diagnosis. Antenna of female more or less widened submedially, with 20 or 21 antennomeres; seventh antennomere distinctly (in Pseudoscapiola) or not protruding ventrally; eyes short or long setose (Figs 3D, 5B); maxillary palp with 3–5 segments (5 in type species); clypeus distinctly narrower than face (Figs 3D, 5B); prepectal carina absent (Figs 3C, 4B); vein 1- R1 of fore wing at least shortly developed (Figs 3A, 4F, 6D); vein 1-SR of fore wing present (Figs 3A, 4F, 6D); areola of dorsal face of propodeum usually absent or narrow, rarely wide (Figs 4D & E, 6C); first and second metasomal tergites smooth or nearly so; first and second metasomal sutures variable, usually absent or shallowly impressed and smooth (Figs 4D & E, 6C). Distribution. Cosmopolitan. Included species. Adelius adeleae Shimbori & Shaw, 2019, A. amplus Belokobylskij, 1998, A. angustus (Papp, 1997), A. australiensis (Ashmead, 1900), A. bolivariensis Bortoni & Penteado-Dias, 2019, A. boliviensis Bortoni & Penteado-Dias, 2019, A. caatinga Bortoni, Shimbori & Penteado-Dias, 2019, A. cadmium (Papp, 2003), A. canadensis Shimbori & Shaw 2019, A. clandestinus (Foerster, 1851), A. coloradensis Muesebeck, 1922, A. determinatus (Foerster, 1851), A. dubius (Foerster, 1851), A. ecuadoriensis Bortoni & Penteado-Dias, 2019, A. erythronotus (Foerster, 1851), Adelius excelsus Bortoni & Shimbori, 2019, Adelius fasciipennis (Rohwer, 1914), A. floridiensis Shimbori & Shaw, 2019, A. gauldi Shimbori & Shaw, 2019, A. germanus (Haliday, 1834), A. hyalinipennis (Foerster, 1851), A. janzeni Shimbori & Shaw, 2019, A. monteiroi Souza-Gessner, Cerântola & Penteado-Dias, 2019, A. morretesiensis Bortoni, Shimbori & Penteado-Dias, 2019, A. nigripectus Muesebeck, 1922, A. panamensis Shimbori & Shaw, 2019, A. pentagonalis Shimbori & Shaw, 2019, A. quiteriae Souza-Gessner, Cerântola & Penteado-Dias, 2019, A. rudnikovi (Perepechayenko, 1994) comb. nov., A. sancticaroli Bortoni, Penteado-Dias & Shimbori, 2019, A. stenolcos de Saeger, 1944; A. subfasciatus Haliday, 1833, A. viator (Foerster, 1851). Notes. We hereby synonymize Pseudoscapiola Perepetchayenko, 1994 (type species and only known species: Myriola rudnikovi Perepetchayenko, 1994) with Adelius as the second author examined the type of Myriola rudnikovi Perepetchayenko. We place this species in Adelius, even though it has a single aberrant character, the 7 th antennomere distinctly protruding ventrally. Adelius rudnikovi (Perepetchayenko, 1994) is a new combination., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on pages 7-9, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470, {"references":["Shimbori, E. M., Bortoni, M. A., Shaw, S. R., Souza-Gessner, C. D. S., Cerantola, P. D. C. M. & Penteado-Dias, A. M. (2019) Revision of the New World genera Adelius Haliday and Paradelius de Saeger (Hymenoptera: Braconidae: Cheloninae: Adeliini). Zootaxa, 4571 (2), 151 - 200. https: // doi. org / 10.11646 / zootaxa. 4571.2.1","Belokobylskij, S. A. (1998) 29. Subfam. Adeliinae (Acaeliinae). In: Lehr P. A. (Ed.), Key to the Insects of the Russian Far East. Neuropteroidea, Mecoptera, Hymenoptera. Vol. IV, pt 3. Dal'nauka, Vladivostok, pp. 553 - 558. [in Russian]"]}

Myriola Shestakov, 1932 re-instated Figs 7–11 Type species: Myriola gussakovskii Shestakov, 1932. Diagnosis. Antenna of female slender submedially, with 20 antennomeres; eyes often glabrous or short setose (Figs 9C, 11C & K); maxillary palp with 5 segments; clypeus almost as wide as face (Figs 7B, 9B, 11C & K); prepectal carina absent (Figs 7F, 10A & B, 11I); medial areola of dorsal face of propodeum narrow (Figs 8B, 10C & D, 11D); vein 1-R1 of fore wing completely absent (Figs 7A, 9A, 10E, 11A); vein 1-SR of fore wing distinct and slender; second metasomal tergite usually smooth (Figs 8B, 10D, 11D); metasomal sutures absent or shallowly impressed and smooth (Figs 8B, 10D, 11D). Distribution. East Palaearctic. Included species. Myriola aridus Tobias, 1967, M. ferulae Tobias, 1964, M. gussakovskii Shestakov, 1932, M. magna Tobias, 1967. Notes. Myriola Shestakov, 1932, is traditionally included in Adelius Haliday, 1833 (Yu et al. 2016), but there are several differences between both taxa as listed above in the key to genera. Therefore, we re-instate Myriola as a valid genus., Published as part of Ranjith, A. P., Achterberg, C. Van, Samartsev, K. G. & Nasser, M., 2021, Discovery of the chelonine tribeAdeliini Viereck, 1918 (Hymenoptera: Braconidae) from the Indian subcontinent with the description of a new genus from south India, pp. 1-25 in Zootaxa 4926 (1) on page 11, DOI: 10.11646/zootaxa.4926.1.1, http://zenodo.org/record/4500470, {"references":["Yu, D. S. K., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016. Ichneumonoidea 2015. Taxapad, Nepean, Ontario. Database on flash-drive. Available from: http: // www. taxapad. com (accessed 7 August 2020)"]}

Megischus alveolifer van Achterberg, 2004 (Fig. 56) Megischus ptosimae sensu van Achterberg, 2002, Zool. Verh. Leiden, 339, 141���144. Megischus alveolifer van Achterberg, 2004, Zool. Meded. Leiden, 78 (3), 115. Type material. Holotype, ♀, India: Tamil Nadu, ĽTiruvannamalai district, Annamalai Hills, (10��18���N & 77��0���E, 1067m), v.1969, Coll. P.S. Nathan, (CNC) CNC Type No. 23675 (OSUC 0021726) Digital loan examined. Diagnosis. Body length 23.5 mm; ovipositor sheath length 25.85 mm; fore wing length 13.7 mm. Antenna with 43 antennomeres; frons dorsally with strong up curved rugae and coarsely transversely rugose frontally; vertex with three strong regular carinae followed by a coarse densely reticulate flattened area; temples smooth, rounded and narrowed behind eyes; occipital carina strongly developed, reaching lower level of eyes; neck anteriorly deeply emarginate; pronotum with three robust curved carinae anteriorly, followed by four weak carinae posteriorly; pronotum posteriorly convex; convex part of metapleuron coarsely reticulate; propodeum coarsely and densely foveolate with wide smooth interstices; vein 1-M 7.4�� vein 1-SR and 1.2�� vein m-cu; vein 2-SR 1.1�� vein r; hind coxa rather slender, spindle-shaped; hind femur swollen and robust, bidentate ventrally; hind tibia steeply depressed at base on the outer side of widened part; first metasomal tergite 12.4�� as long as its maximum width; ovipositor sheath 1.1�� length of body, with a long subapical ivory part, ivory part 1.3�� as long as dark apical part. Male. Unknown. Host. Unknown. Distribution. Oriental (India: Tamil Nadu)., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 30-31, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. & Yang, Z-Q. (2004) New species of the genera Megischus Brulle and Stephanus Jurine from China (Hymenoptera: Stephanoidea: Stephanidae), with a key to world species of the genus Stephanus. Zoologische Mededelingen Leiden, 78 (3), 101 - 117.","van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206."]}

Parastephanellus mufeedae Narendran & Sureshan, 2001 (Fig. 72) Parastephanellus mufeedae Narendran & Sureshan, 2001, Zoos Print J., 16(10), 597���599. Type material. Holotype ♀, mounted on rectangular card, India: Kerala, Malappuram district, Tirur (11��00���N & 76��00���E, 40m), 08.ix.1999, Coll. C.A. Mufeeda, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.1890. Examined. Diagnosis. Body length 8.2 mm; ovipositor sheath length 13.0 mm. Frons with transverse irregular carinae with rugose and reticulate interstices; vertex with several cross carinae followed by a longitudinal medial grove; temple with distinct pale streak; pronotal fold indistinct; mesoscutum with a median pitted grove, notauli distinct, pitted on the posterior half; scutellum shiny with very fine reticulation, 1.5�� as long as mesoscutum; propodeum with scattered shallow pits and fine reticulate interstices; metapleuron foveate; hind femora bidentate ventrally with 3���4 denticles in between the teeth and a few denticles basally; first metasomal tergite strongly annulate, basally rugate reticulate, distally smooth, shorter than remaining segments; ovipositor sheath 2.5�� as long as metasoma, without a subapical pale band. Male. Unknown. Distribution. Oriental: India (Kerala). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 36-37, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sureshan, P. M. (2001) A new species of Parastephanellus Enderlein (Hymenoptera: Stephanidae) from India. Zoos' Print Journal, 16 (10), 597 - 599. https: // doi. org / 10.11609 / JoTT. ZPJ. 16.10.597 - 9"]}

Foenatopus priyae (Narendran & Sureshan, 2003) Diastephanus priyae Narendran & Sureshan, 2003, Entomon, 28(2), 116���118. Type material. Holotype ♀, India: Kerala Thrissur district, Thrissur (10�� 31��� N & 76�� 12��� E, 2.8 m), 12.vi.1999, Coll. Priya Menon. Paratype 4♂, same data as holotype, 19.iv.1999, (ZSIK) (Type specimen missing). Diagnosis. Body length 12.4 mm, ovipositor sheath length 8.8 mm. Frons trans-striate with longitudinal and oblique striae; vertex with four striae followed by fine striae, not forming any longitudinal medial line or grove; neck trans-striate dorsally; mesoscutum smooth anteriorly, faintly reticulate with shallow pits and reticulate interstices at sides; scutellum and axillae almost smooth and shiny with a few scattered pits on sides; metapleuron rugose-reticulate; propodeum with broad deep pits and shagreened interstices; hind coxa and femora reticulate and transverse striate proximally turning transversely striate distally; hind femora ventrally tridentate with two denticles basally; first metasomal tergite trans-carinate as long as remaining segments combined; ovipositor sheath distinctly longer than metasoma, with a subapical pale ivory patch. Male. Body length 7���10 mm. Similar to females except lower frons and face yellow, vertex with three and rarely four carinae; first metasomal tergite 1.1���1.3�� length of remaining segments. Distribution. Oriental: India (Kerala). Host. Unknown. Note. The types are missing, so the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 19-20, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138."]}

Parastephanellus narendrani Binoy, van Achterberg & Santhosh sp. nov. (Figs 73���83) urn:lsid:zoobank.org:act: DA69B4AC-DFB9-438B-A29E-18DBA6DEBAC3 Type material. Holotype ♀, India: Kerala, Idukki district, Elappara (9��38���18.1���N & 76��58���56.8���E, 1030 m), 23.v.2019, Coll. C. Binoy, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13597. Paratype 1 ♀, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13598, same collection data as that of holotype. Diagnosis. Head in dorsal view almost oblique behind eyes; pronotum posteriorly well emarginate, entire surface finely transverse rugose joining onto the mesoscutum and with scattered setae; sdc1 of fore wing robust and 2.3�� longer than wide; hind coxa with strong transverse rugae and without a dorsal tooth, long scattered setae arising from the surface; first three metasomal tergites dark brown to black, first tergite apically and second tergite basally brownish orange; first metasomal segment narrow in lateral view; first tergite 9.1�� as long as its maximum width with strong irregular coarsely transverse rugae. The new species runs close to P. malabaricus in having mesoscutum with median longitudinal row of pits and no median carina and metapleuron with a smooth area separating it from propodeum. But it differs from P. malabaricus as follows: ovipositor sheath 0.6�� as long as body (in P. malabaricus ovipositor sheath subequal to length of body); vertex rugate, rugae almost reaching the occiput (medially strong) (in P. malabaricus vertex smooth posteriorly, median area weakly rugate); pronotum finely transversely striate (in P. malabaricus pronotum smooth). Description. Holotype, ♀. Body length 13 mm, ovipositor sheath length 6.7 mm, fore wing length 3.9 mm. Colour. Brownish black; tegula and palpi dark brown; scape brown with apical tip honey brown, pedicel, F1���F4, F5 and F6 basally honey brown, rest of flagellomeres deep brown; malar space with a pale yellow patch below eyes (Fig. 53); mandibles brown with apices black; all coxae, and femora liver brown; trochanters paler, all tibia liver brown in basal half, followed by paler are apically; all tarsi brown; wing membrane slightly brownish at cubital cells, pterostigma and veins deep brown, base of pterostigma pale; ovipositor sheath brownish black with a small pale spot apically. Head. Antenna with 28 antennomeres; F2 onwards surface distinctly pitted; frons uniformly rugose near the inner eye margin, coarsely rugose medially, with scattered setae; five anterior coronal teeth, antero-medial one largest, followed by two pairs of posterior arcuate ones, with few carinae radiating from middle teeth to the last pair through the median ocellus (Fig. 76); a curving carinae present behind the posterior teeth forming a subtriangular to pentagonal space with a small median carinae and irregular rugae; vertex rugose beyond the lateral ocellus, turning into irregular fine striae reaching the occipital carina, widely smooth laterally; temples non-angulate, largely smooth and shiny (Fig. 75); occipital carina complete, reaching the mandibles anteriorly. Mesosoma. Neck short with strong transverse and emarginate rugae, in level with pronotum without forming a cavity medially; pronotum strongly rugose without any distinct carinae, sparsely setose, posterior margin emarginated; propleuron smooth and shiny, sparsely setose; mesonotum irregularly foveolate with smooth interspaces, truncating on a straight scutellar line posteriorly; median grove distinct, notauli distinctly foveolate delimiting large coarsely rugose axillae (Fig. 77); mesopleuron dorsally with coarse rugae extending as alutaceous sculpturing ventro-laterally with scattered white setae; scutellum small, coriaceous, medially smooth with scattered large setigerous pits; metapleuron finely rugose without setae (Fig. 78); propodeum variable, anteriorly with fine alutaceous rugae turning into large irregular areola posteriorly, a smooth area flanging apical fovea delimited by a transverse carina, a median protuberance apically joining first metasomal tergite; fore wing semi-hyaline with scattered minute setae, 1-M 2.9�� 1-SR, almost straight; vein r reaching before the apex of pterostigma; sdc1 robust, 2.3�� as long as its maximum width; cu-a obsolete or absent (Fig. 80); hind coxa robust, without a dorsal teeth and with strong rugae throughout; hind femur widened, smooth with alutaceous sculpturing and scattered sparse white setae, ventrally with 2 large teeth and several denticles in between; hind tibia alutaceous with moderate pubescence, 1.4�� as long as hind femur, apical inflated part (i.e., apical part with sculpture different from basal petiolate part) 1.9�� as wide as narrow basal part, narrow part 1.4�� as long as wider part, widened part with distinct oblique inverted obsolete V shaped carinae; hind tarsus with three tarsomeres, sub-parallel sided 4.3�� as long as wide medially and 2.6�� as long as second tarsal segment (Fig. 79). Metasoma. First tergite 9.15�� as long as its maximum width, 2.39�� as long as second tergite, cylindrical with irregular strong rugae throughout the length and white setae along the lateral margins, apically smooth (Fig. 81); second tergite smooth and shiny medially, fine alutaceous laterally and apically; third to seventh tergites finely reticulate to alutaceous with scattered small white setae; eighth tergite finely pitted, setose and deeply furcate apically, no pygidial protuberance present apically (Fig. 82); ovipositor sheath 0.5�� as long as body and 1.69�� as long as fore wing (Fig. 83). Male. Unknown. Distribution. Oriental: India (Kerala). Biology. Unknown, presumably attacking wood boring beetles infesting fallen woods. Etymology. The species is named after the late Prof. Dr. T.C. Narendran for his large contributions in the field of hymenopteran taxonomy in India., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 37-40, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367

Foenatopus chareshi Binoy, van Achterberg & Girish Kumar sp. nov. (Figs 4���20) urn:lsid:zoobank.org:act: 5B58D9E8-F54F-40DB-9393-816A512A96A1 Type material. Holotype ♀, mounted on triangular card, India: Tamil Nadu, Thirunelveli district, Kalakkad Mundanthurai Tiger Reserve (8��41���04.7���N & 77��18���30.5���E, 1800 m), 28.ix.2017, Coll. C. Charesh, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13591. Paratypes: 4 ♂ (ZSIK) Regd. Nos. ZSI/ WGRC /IR/INV.13592 13595, same collection data as that of holotype. Diagnosis. Frons finely and transversely rugose; vertex with fine transverse striae; pronotum slender, anteriorly transversely rugulose to weakly carinate and posteriorly coriaceous; scutellum microsculptured; propleuron and mesopleuron coriaceous; propodeum areolate with medium-sized, circular areola and with imbricate to coriaceous interspaces, inside of areola imbricate; 2-CU 1 of fore wing absent; subapical part of ovipositor sheath ivory. This species comes close to Foenatopus punctatus Elliott, 1919 in having length of ovipositor subequal to body, subapical ivory patch on the ovipositor sheath and first metasomal tergite shorter than combined lengths of post-petiolar segments. But can be distinguished by the combination of the following characteristics: vertex with two carinae (in F. punctatus vertex with 3 carinae); pronotum anteriorly weakly carinate proceeding as coriaceous posteriorly (pronotum strongly trans-rugose in F. punctatus); and hind femora bidentate ventrally (hind femora tridentate in F. punctatus). The species comes close to F. achterbergi Gupta & Gawas 2020 in having the pterostigma more than 10.0�� longer than its maximum width, ovipositor sheath shorter than body and vertex with two carinae between the ocelli, but varies from the same in having the subapical pale band of the ovipositor sheath 1.6�� apical dark part (subapical pale band in F. achterbergi 0.9�� apical dark part); ventral teeth of hind femora more or less equilateral (in F. achterbergi femoral teeth of varying length, basal one nearly equilateral and apical one narrow, much longer than wide). The species runs to the Oriental species F. flavidentatus (Enderlein) (Hong et al. 2011) in having hind femur bidentate ventrally, pygidial impression of female deep and reverse V-shaped, ovipositor sheath 0.9�� length of body and with a subapical ivory patch. It differs from F. flavidentatus as follows: vertex with two carinae (in F. flavidentatus vertex with three distinct carinae); pronotum granulate to reticulate sculpturing (in F. flavidentatus pronotum weakly carinate); scutellum finely alutaceous (in F. flavidentatus scutellum smooth); 2-CU 1 absent (in F. flavidentatus 2-CU 1 present basally); first metasomal tergite 0.8�� as long as length of remaining metasomal tergites combined (in F. flavidentatus first metasomal tergite 0.9�� as long as length of remaining metasomal tergites combined). Description. Holotype. ♀ (Figs 4 & 5���14). Body length 10.1 mm; ovipositor sheath length 9.5 mm; fore wing length 5.2 mm. Colour. Reddish brown with the following parts as follows: frons with short ivory streaks through the inner eye margin and medially, not reaching level of anterior coronal tooth; temple brown with ivory streak along lower half of the eye, streak becoming narrow apically to brown occiput (Fig. 6); lower face with base of mandible yellow (Fig. 8); subapical ivory-white part of ovipositor sheath about as long as apical black part (Fig. 14). Head. Antenna with 29 distinct antennomeres; first flagellomere 5.0�� as long as wide, and 0.68�� as long as second flagellomere; three anterior coronal teeth large and acute, both posterior ones short and wider; frons finely and transversely rugate; coronal area with weak longitudinal carinae (Fig. 8), with two transverse carinae between ocelli; vertex finely and transversely striate (but medially rugose) and with a shallow medio-longitudinal furrow (Fig. 7); temple non-angulate, smooth and shiny. Mesosoma. Pronotum slender and mostly coriaceous turning microreticulate or granulate posteriorly, without transverse rugulae anteriorly; weakly carinate dorso-medially and laterally; neck anteriorly deeply emarginated, neck at slightly lower level than middle part of pronotum; anterior half of mesoscutum finely rugose to coriaceous with irregular areola, posterior half with a median carina (Fig. 10); scutellum, propleuron and mesopleuron coriaceous to micropitted (Fig. 9); scutellum separated from axilla by means of a large furrow, not differentiated in the form of fovea with scattered small setae; axillae separated from each other by a large alveola; propodeum with medium-sized, circular areola with wide coriaceous interspaces, inside of foveae imbricate to alutaceous; fore wing with 2-CU1 absent; pterostigma elongate and subparallel-sided, turning acute apically; vein r and vein SR1 obtusely-angled, vein r ends 0.2�� length of pterostigma behind level of apex of pterostigma; vein SR1 subparallel to costal margin (Fig. 11); hind coxa weakly striate, ventrally with moderate striae; hind femur microreticulate, ventrally with two large acute teeth and with two smaller tubercle basally, three denticles in between the large teeth and a stub like lobe apically; hind tibia faintly coriaceous, as long as hind femur, basal narrow part of hind tibia 1.1�� as long as widened part, inner side of widened part basally distinctly depressed, followed by convex and setose area, apically densely setose; hind tarsi densely setose (Fig. 12). Metasoma. First tergite rugose with faint transverse striae, weakly annulate, sparsely setose, 13�� as long as its maximum width, 2.6�� as long as second tergite and 0.9�� as long as remainder of metasoma (Fig. 13); post-petiolar tergites largely smooth to weakly coriaceous; pygidial area distinctly differentiated, pygidial impression moderate, reverse V-shaped; length of ovipositor sheath 0.94�� as long as body length, length of subapical whitish band 1.6�� length of its dark apex (Fig. 14). Male. Description. Paratypes. ♂ (Figs 15���20). Body length 7.3���8.5 mm; fore wing length 3.0��� 4.7 mm; Colour. Black with following parts different: lower face with an ivory patch below eyes; posterior rim of pronotum yellowish golden; post-petiolar sternites reddish brown; fore and mid tibia and all tarsi brown; mid tibia brown with basal part paler (Fig. 15). Frons finely striate with shallow median grove; vertex with 2 carinae, scattered setae (Fig. 17); pronotum alutaceous with scattered setae, without strong carinae, dorso-medially four weak ridges; scutellum smooth to finely alutaceous with scattered setae, separated from axillae by wide ridges; axillae separated from one another by a pair of irregular alveola; propodeum with large imbricate irregular to circular areola with alutaceous wider interspaces than in females, inside of pits imbricate to alutaceous (Fig. 18); wing venation similar to that of female (Fig. 19); mesopleuron antero-dorsally with moderately dense silvery setae; metapleura rugose carinate; hind femora with 2 tooth with basal one white (variable within specimens) (Fig. 20). Distribution. Oriental: India (Tamil Nadu). Host. Unknown. Etymology. The species is named after C. Charesh (Laboratory Assistant at Western Ghat Regional Centre, Zoological Survey of India, Kozhikode), who collected the type specimens., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 8-11, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Gupta, A & Gawas, S. M. (2020) A new species of the genus Foenatopus Smith (Hymenoptera: Stephanoidea: Stephanidae) from India. Zootaxa, 4801 (2), 389 - 394. https: // doi. org / 10.11646 / zootaxa. 4801.2.13","Hong C. - D., van Achterberg & Xu, Z. - F. (2011) A newly recorded genus and species of Family Stephanidae (Hymenoptera, Stephanoidea) from China. Entomotaxonomia, 33 (1), 71 - 73."]}

Foenatopus sureshani Binoy, van Achterberg & Girish Kumar sp. nov. (Figs 32–43) urn:lsid:zoobank.org:act: 2077C090-758E-49B6-B7B4-E22914C0283C Type material. Holotype ♀, mounted on triangular card, India: Uttar Pradesh, Lalitpur district, Babina (25°16’957”N & 78°30’379”E, 303 m), Coll. P. Girish Kumar, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13596. Diagnosis. Frons with fine and transverse rugae; vertex with irregular rugulae; pronotum slender, anteriorly with weak transverse striae, posteriorly coriaceous to finely reticulate; scutellum microreticulate; propleuron and mesopleuron coriaceous; propodeum anteriorly microreticulate to alutaceous, postero-laterally longitudinal areola and coriaceous interspaces, inside of areola imbricate; 2-CU 1 of fore wing absent. This species runs close to F. sudhae in having the hind femora ventrally with two large teeth. However, the new species differs from F. sudhae as follows: vertex with two carinae (in F. sudhae vertex with four carinae); mesoscutum with a median longitudinal grove (in F. sudhae mesoscutum without such grove). The species also comes close to F. idukkiensis, F. chinnarensis and F. frontilinea in having vertex with two carinae, but differs as follows: stemmaticum without a distinct medial grove (a distinct medial grove present in F. idukkiensis); ovipositor sheath longer than metasoma (ovipositor sheath shorter than metasoma in F. chinnarensis); first metasomal tergite shorter than combined remaining segments (in F. frontilinea first tergite longer than combined remaining segments). The new species runs to the Oriental species F. maculiferus Hong et al. (Hong et al. 2011) in having third metasomal tergite with two large ivory patches, propodeum almost matt on anterior 2/3 rd and foveate on the rest. However, the new species differs from F. maculiferus in having: face with yellow patch reaching second set of tubercles (in F. maculiferus the patch not extending beyond anterior coronal teeth); vertex with two carinae (in F. maculiferus vertex with three carinae); scutellum black, flat medially and almost in same line with mesoscutum and propodeum (in F. maculiferus scutellum reddish brown, medially distinctly convex); hind femora, reddish brown and densely reticulate throughout the lamina (in F. maculiferus hind femora orange red, reticulate dorsally with punctate lamina ventrally); ovipositor sheath completely brownish black (in F. maculiferus ovipositor sheath with a brownish subapical band). Description. Holotype. ♀. Body length 6.1 mm; ovipositor length 4.43 mm; fore wing length 3.54 mm. Colour. Brownish black with the following parts as follows: frons till the anterior coronal teeth and lateral halfway along the outer margin of eyes yellow; temples brown; mandibles yellow with apex black; ovipositor sheath black. Head. Antenna with 22 distinct antennomeres; first flagellomere 3.9× as long as wide, and 0.7× as long as second flagellomere; three anterior coronal teeth large and acute, both posterior ones short and wider; frons finely and transversely rugate; coronal area with weak longitudinal carinae; vertex finely and transversely striate, two carinae and without a shallow median longitudinal furrow (Fig. 35); temple non-angulate, smooth and shiny (Fig. 34). Mesosoma. Pronotum slender and mostly coriaceous to faintly pitted, without transverse rugulae anteriorly; weakly carinate dorso-medially and laterally; neck anteriorly deeply emarginated, neck at slightly lower level than middle part of pronotum; anterior half of mesoscutum finely rugose to coriaceous with irregular areola, posterior half with a median carina (Fig. 36); scutellum, propleuron and mesopleuron coriaceous to micropitted (Fig. 38); scutellum separated from axilla by means of a large furrow, not differentiated in the form of fovea with scattered small setae; axillae separated from each other by a large alveola; propodeum with medium-sized, circular areola with wide coriaceous interspaces, inside of foveae imbricate to alutaceous; fore wing with 2-CU1 absent; pterostigma elongate and subparallel-sided, turning acute apically; vein r and vein SR1 obtusely-angled, vein r ends 0.2× length of pterostigma behind level of apex of pterostigma; vein SR1 subparallel to costal margin (Fig. 37); hind coxa weakly striate, ventrally with moderate striae (Fig. 39); hind femur microreticulate, ventrally with two large acute teeth and with two smaller tubercle basally, three denticles in between the large teeth and a stub like lobe apically; hind tibia faintly coriaceous, as long as hind femur, basal narrow part of hind tibia 0.96× as long as widened part, inner side of widened part basally distinctly depressed, followed by convex and setose area, apically densely setose; hind tarsi densely setose (Fig. 40). Metasoma. First tergite rugose with faint transverse striae, weakly annulate, sparsely setose, 6.86× as long as its maximum width (Fig. 39), 1.91× as long as second tergite and 0.7× as long as remainder of metasoma; post-petiolar tergites largely smooth to weakly coriaceous (Fig. 42); pygidial area distinctly differentiated, pygidial impression moderate, reverse V-shaped (Fig. 41); length of ovipositor sheath 0.72× as long as body length, without a subapical whitish band (Fig. 43). Male. Unknown. Distribution. Oriental: India (Uttar Pradesh). Host. Unknown. Etymology. The species is named after Dr. P.M. Sureshan (Scientist-E & Officer-in-Charge, Western Ghat Regional Centre, Zoological Survey of India, Kozhikode) for his valuable contributions towards the study of Indian Stephanidae.

Foenatopus keralensis (Narendran & Sureshan, 2003) (Fig. 25) Diastephanus keralensis Narendran and Sureshan, 2003, Entomon, 28(2), 121���124. Type material. Holotype ♀, India: Kerala Malappuram district, Edappal (10�� 43��� N & 76�� 1��� E, 10 m), 10.x.1999, Coll. P. Sudha. (NHM) Diagnosis. Body length 11.1 mm, ovipositor sheath length 8.7 mm. Frons trans-striate without any oblique or longitudinal striations; vertex with four carinae followed by fine striae, not forming any medial grove; pronotum weakly reticulate dorsally; mesoscutum faintly shiny, reticulate, anterior half separated by a fovea, posteriorly with a median ridge; scutellum shiny with faint reticulations; metapleuron microsculptured; propodeum with large pits, interstices transversely reticulate; wings hyaline with pterostigma translucent yellow medially; hind coxa carinate punctate basally turning circularly annulate elsewhere; hind femora tridentate ventrally with a basal tubercle; first metasomal tergite uniformly annulated, longer than remaining segments; ovipositor sheath longer than metasoma, with a subapical yellow band. Male. Unknown. Distribution. Oriental: India (Kerala). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 17-18, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138."]}

Foenatopus aequalis Elliott, 1928 Foenatopus aequalis Elliott, 1928, Entomologist, 61 (778), 63. Type material. Holotype ♀. India: Karnataka, Mysore, Chikkaballapura, Coll. Campbell, vii.1918 / (Type specimen missing). Diagnosis. Body length 13.5 mm; ovipositor sheath length 12 mm. Frons moderately trans-striate, impressed in upper half; vertex with three strong carinae; scape long, as long as cheeks; pronotum smooth and shining, semi annular with apical transverse line of punctures; mesoscutum irregularly punctate, usual rows of punctures obsolete; scutellum large, central lobe smooth, lateral lobes punctate; metapleura medially reticulate punctate; hind coxa trans-striate, basally more coarsely; hind femora finely trans-aciculate, bidentate; first metasomal segment trans-striate, white banded, shorter than rest of remaining metasomal segments; ovipositor sheath 1.3�� length of metasoma, with a subapical pale band. Male. Unknown. Distribution. Oriental: India (Karnataka). Host. Unknown. Note. The type of this species is missing, therefore, the above mentioned diagnosis is obtained from the original description. We have been unable to associate any extant specimens with this species., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 6-7, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1928) A hymenopteron (Stephanidae) new to science. The Entomologist, 61 (778), 63."]}

Key to Indian species of Parastephanellus 1. Vein 2-SR+M of fore wing absent (Fig. 91)....................... P. reductus Binoy, Girish Kumar & Santhosh sp. nov. - Vein 2-SR+M of fore wing largely present (Figs 72, 80)....................................................... 2 2. Pale streak of temple scarcely noticeable........................................................ P. scitus Elliott - Pale streak of temple distinctly developed.................................................................. 3 3. Ovipositor sheath distinctly longer than body (Fig. 72); mesoscutum without median longitudinal row of pits but with a median carina; metapleuron without a smooth area separating from propodeum; [vertex with several cross carinae and with a mediolongitudinal grove]........................................................ P. mufeedae Narendran & Sureshan - Ovipositor sheath approx. as long as or distinctly shorter than body; mesoscutum with median longitudinal row of pits and no median carina; metapleuron with a smooth area separating it from propodeum..................................... 4 4. Ovipositor sheath subequal to length of body; vertex mostly smooth, two strong carinae followed by weak striae (medially indistinct); pronotum smooth................................................ P. malabaricus Narendran & Sheela - Ovipositor sheath 0.5× as long as body; vertex rugate, almost reaching the occiput (medially strong); pronotum finely transverse striate............................................. P. narendrani Binoy, van Achterberg & Santhosh sp. nov.

Foenatopus bilineatus (Elliott, 1919) Diastephanus bilineatus Elliott, 1919b, Entomologist, 52, 162. Type material. Type ♀, India: Bihar, Pusa, Coll. G. R. Dutt, 12.v1909; India: West Bengal, Chapra, Coll. Mackensie, Date:?. (Type specimen missing). Diagnosis. Body length 8.5 mm; ovipositor sheath length 7 mm. Head finely trans-striate, temples smooth; vertex with two carinae; neck elongate, finely trans-striate, remainder smooth; mesonotum sub-glabrous, very diffusely punctate; metapleuron cribrate punctate; hind coxa trans-striate; hind femora tridentate ventrally, central tooth white; first metasomal tergite trans-striate, as long as remaining segments; ovipositor sheath longer than metasoma, with a subapical pale whitish band. Male. Unknown. Distribution. Oriental: India (Bihar, West Bengal). Host. Unknown. Note. The type of this species is missing, therefore, the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 8, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1919 b) Two new Stephanidae. The Entomologist, 52, 162 - 163."]}

Foenatopus achterbergi Gupta & Gawas, 2020 (Fig. 2) Foenatopus achterbergi Gupta & Gawas, 2020, Zootaxa, 4801 (2): 390���392. Type material. Holotype ♀ India: Karnataka, Sangam, Coll. Sandesh M. Gawas, 18.vii. 2018 (ICAR-NBAIR). Diagnosis. Body length 22.5 mm; ovipositor sheath length 21.0 mm; fore wing length 5.6 mm. Frons coarsely transversely striate; vertex with two strong carinae followed by fine transverse striae, forming a medial line or grove; occipital carina strong, complete; neck in level of pronotum, deeply emarginated; pronotum anteriorly with weak transverse rugulae, medially with transverse reticulate striae and posteriorly smooth; mesoscutum anterior half finely reticulate striate, posteriorly coarsely foveolate; scutellum and axillae shiny with reticulate striations and scattered punctures; mesopleuron anterior third densely setose, remainder largely aciculate; metapleuron largely strongly reticulate-foveolate; propodeum foveolate with microsculptured interspaces; fore wing hyaline with vein 2-CU1 weakly developed, 0.2�� vein cu-a; pterostigma elongate, subparallel-sided and acute apically; hind coxa mostly transversely striate, anterior third rugose; hind femur densely transversely strigate with two ivory coloured teeth ventrally, apical one longer; first metasomal tergite finely transversely striate; 1.6�� remainder of metasoma; ovipositor sheath 0.96�� length of body, with a subapical pale band, band 0.9�� as long as apical dark part. Male. Unknown. Distribution. Oriental: India (Karnataka). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 5-6, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Gupta, A & Gawas, S. M. (2020) A new species of the genus Foenatopus Smith (Hymenoptera: Stephanoidea: Stephanidae) from India. Zootaxa, 4801 (2), 389 - 394. https: // doi. org / 10.11646 / zootaxa. 4801.2.13"]}

Foenatopus sudhae (Narendran & Sureshan, 2003) (Fig. 31 ) Diastephanus sudhae Narendran & Sureshan, 2003, Entomon, 28(2), 119���121. Type material. Holotype ♀, India: Kerala Palakkad district, Kumaranellur (10��47���N & 76��02���E, 26 m), 08.viii.1999, Coll. P. Sudha, (NHM) B. M. TYPE HYM. 3a. 391 (NHMUK013458832). Additional material examined: 1♀, India: Arunachal Pradesh, Tirap district, Deomali (27��11���36.62���N & 95��28���11.70���E, Alt 161 m), 1971, Coll. G.S. Arora, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13603; 1♀, India: Goa, South Goa district, Bhagwan Mahaveer Wildlife Sanctuary (15��20���33.26���N & 74��15���8.62���E, Alt 85 m), 17.v.2018, Coll. P. Girish Kumar & party, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13604; 1♀, India: Uttar Pradesh, Lucknow, National Botanical Research Institute Campus (26��51���17.56���N & 80��57���11.89���E, Alt 117 m), 7.iii.2007, Coll. P. Girish Kumar, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13605. Diagnosis. Body length 12.0 mm, ovipositor sheath length 8.8 mm. Frons with weak transverse striae and reticulations; vertex with four strong carinae followed by reticulate area posteriorly, not forming any medial grove; pronotum with 12���15 trans-carinae anteriorly, posterior half weakly reticulate-striate; mesoscutum with deep irregular punctures and reticulations; scutellum smooth to faintly reticulated with scattered pits on sides; axillae finely reticulate; metapleuron foveate; propodeum with scattered large deep pits with a broad anterior smooth area with transverse reticulations; first metasomal tergite uniformly annulated, a little shorter than combined remaining segments; ovipositor sheath 1.4�� as long as metasoma, without an ivory band sub-apically. Male. Unknown. Distribution. Oriental: India (Arunachal Pradesh (present record), Goa (present record), Kerala, Uttar Pradesh (present record)). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 22-23, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138."]}

Foenatopus frontilinea (Morley, 1917) (Fig. 21 & 22) Diastephanus frontilinea Morley, 1917, Entomologist, 50, 109. Type material. Holotype ♀, India: Assam-Eastern Bengal border, 20.vi.1905, Coll.? from PUSA collection, (NHM). Diagnosis. Body length 10.5 mm, ovipositor sheath length 9 mm. Frons faintly reticulate, a longitudinal line on frons and head; vertex longitudinally aciculate, two carinae between the lateral ocelli; occiput finely and closely trans-aciculate; pronotum multicarinate, semi-annular, smooth and shiny; mesoscutum smooth and shiny, sparsely punctate; metapleuron smooth and subaciculate; scutellum smooth and shiny with three rows of punctures on either side; propodeum coriaceous between very largely and partly confluent pits; hind coxa trans-striate; hind-femora alutaceous with two teeth on ventral margin with few denticles; first metasomal tergite trans-aciculate, longer than combined remaining segments; ovipositor sheath longer than metasoma, with a subapical pale ivory band. Male. Unknown. Distribution. Oriental: India-Bangladesh border. Host. Unknown. Note. The types are in very bad condition, therefore, the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 13, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Morley, C. (1917) Some Stephanidae: with descriptions of new species. The Entomologist, 50, 105 - 113. https: // doi. org / 10.5962 / bhl. part. 3484"]}

Megischus ranjithi Binoy, Santhosh & Girish Kumar sp. nov. (Figs 57���71) urn:lsid:zoobank.org:act: 729938C3-25E2-4364-8C95-672EFF775570 Type material. Holotype ♀, India: Tamil Nadu, Coimbatore district, TNAU Campus (11��0���28���N & 76��56���14���E, 47 m), Coll. M. Ranjith, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.14031. Diagnosis. Head in dorsal view with smooth and shiny area past eyes, pronotum anteriorly with three strong carinae, posteriorly smooth with scattered setigerous pits; mesoscutum densely and obliquely punctate; scutellum and axillae smooth with scattered pits; ovipositor sheath with a subapical ivory band 0.4�� as long as the apical dark part. The new species falls out from the first couplet of key to old world Megischus (van Achterberg 2002) in having narrowed part of hind tibia 0.4�� as long as hind tibia; ivory part of ovipositor sheath 0.52�� as long as its dark apical part; area behind eyes in dorsal view smooth; mesopleuron smooth with setigerous pits. The new species runs close to M. ptosimae in key to Chinese Megischus (Hong et al. 2010) in having temple slightly convex behind eye; head largely blackish or dark brown; widened part of hind tibia of female distinctly concave ventrally. But it differs from M. ptosimae in having vein 1-M of fore wing 6.8�� as long as 1-SR and 0.8�� as long as vein m-cu (in M. ptosimae 1-M 4.2���5.5�� as long as vein 1-SR and 1.1���1.3�� vein m-cu); ovipositor sheath with subapical ivory band 0.52�� dark apical band (in M. ptosimae subapical ivory band 0.7���2�� dark apical band). The new species varies from the previously reported Indian specimen M. alveolifer van Achterberg, 2004 in having head black (in M. alveolifer, head orange red); head in frontal view with striae subparallel, converging medioventrally from sides); vertex with two strong and two weak more or less straight carinae (in M. alveolifer vertex with three deeply emarginated carinae); hind coxa robust (in M. alveolifer hind coxa rather slender, spindleshaped); subapical ivory part of ovipositor sheath 0.52�� as long as dark apical part (in M. alveolifer ivory part 1.3�� as long as dark apical part); and petiole reddish brown (in M. alveolifer petiole brownish black). Description. Holotype, ♀. Body length 17.06 mm; ovipositor length 3.93 mm; fore wing length 9.64 mm. Colour. Black with the following parts as follows: tegulae apically with a reddish brown patch; fore and mid coxae black with a reddish tint apically; fore and mid trochanters, femur, tibia and tarsi deep reddish; hind trochanters, tibia and tarsi red; petiole basally and apically brownish red with medial portion bright reddish; apex of second metasomal tergite reddish brown; ovipositor sheath black with subapical ivory band. Head. Antenna with 41 distinct antennomeres; first antennal segment 1.6�� as long as wide and 2�� as long as second segment; clypeus rugose with moderate pubescence; mandibles strong with thick setae arising from above and below teeth; ventrally more setose; three anterior coronal teeth large and acute, both posterior ones short and wider; frons finely and transversely rugate; rugae sub parallel sided almost forming a medial carina on lower face; coronal area with smooth areolate rugose channels enclosing the anterior ocellus (Fig. 60); vertex rugose-carinate with three carinae beyond the posterior ocelli; area behind the eye in dorsal view, smooth and shiny; temples non angulate (Fig. 61); face with scattered golden yellow setae along vertex and frons; clypeus rugate with moderate pubescence; thick yellowish brown setae arising from above and below mandibles; gena smooth with dense setigerous micropunctures halfway up the temple (Fig. 59). Mesosoma. Pronotum stout with neck less emarginate, weakly convex in level with middle part of pronotum (Fig. 62); anteriorly with three strong carinae (prominent laterally), posteriorly smooth with few setigerous punctures, laterally produced as shoulders, ending near propleura as buttressing ridges (Fig. 63); propleura with a row of setae along the margin; mesoscutum with oblique rows of irregular punctae forming with a median bare foveate line; scutellum and axillae smooth, shiny with few punctae; axillae separated by a large fovea; mesopleura smooth with scattered setigerous pits, more concentrated ventrally, dorso posteriorly smooth and shiny; metapleura dorsally and ventrally smooth and shiny, medially with irregular setigerous areola (Fig. 63); propodeum two thirds foveolate with inside of fovea and interstices smooth, a median rugose stripe present, posteriorly fovea turning into irregular rugae (Fig. 62); fore wing with vein 2-1 A and 3-CU 1 present; pterostigma elongate and acutely ending posteriorly; 1-M straight, 6.8�� as long as vein 1-SR, 0.8�� vein m-cu (Fig. 64); hind coxa smooth, strong broken striae to tuberculate sculpture anteriorly, posteriorly trans striate with thick setae arising from strong pits along the length (Fig. 65); hind femur smooth, with many setigerous pits on outer and inner disc, setae arising from the pits long and variable in colour (brown and white), a pair of strong tooth ventrally, basal one wider, a denticle near the ventral apex; hind tibia robust, narrowed part 0.45�� as long as hind tibia; expanded part micropitted with long setae similar to that on hind femur, below widened part distinctly concave (Fig. 66); apex of hind tibia and hind tarsi ventrally densely setose with thick yellow setae extending till the second tarsal segment; hind telotarsi almost bare (Fig. 67). Metasoma. First metasomal tergite anteriorly rugose carinate, regularly annulated along the length with a medio-lateral groove, sparsely setose ventrally (Fig. 68); 9.8�� as long as its maximum width, 2.98�� as long as second tergite and 1.3�� as long as remainder of metasoma; post-petiolar tergites largely smooth to weakly reticulate, scattered setae ventro-laterally; seventh and eighth tergites microreticulate (Fig. 69); pygidial area distinctly differentiated, pygidial impression moderate, reverse V-shaped with a pair of nodes with three pairs of setae arising from it (Fig. 70); length of ovipositor sheath 0.54�� as long as body length, with a subapical ivory band, 0.52�� apical dark part (Fig. 71). Male. Unknown. Distribution. Oriental: India (Tamil Nadu). Host. Unknown. Etymology. The species is named after M. Ranjith (Research Scholar, Department of Entomology, University of Agricultural and Horticultural Sciences, Shivamogga), who collected the type specimen., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 31-35, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206.","Hong C. - D., van Achterberg & Xu, Z. - F. (2010) A new species of Megischus Brulle (Hymenoptera, Stephanidae) from China, with a key to the Chinese species. ZooKeys, 69, 59 - 64. https: // doi. org / 10.3897 / zookeys. 69.738","van Achterberg, C. & Yang, Z-Q. (2004) New species of the genera Megischus Brulle and Stephanus Jurine from China (Hymenoptera: Stephanoidea: Stephanidae), with a key to world species of the genus Stephanus. Zoologische Mededelingen Leiden, 78 (3), 101 - 117."]}

Parastephanellus scitus Elliott, 1919 Parastephanellus scitus Elliott, 1919b, Entomologist, 52, 163. Type material. Holotype ♂, India: West Bengal, Pusha (22��03���32.6���N & 87��30���16.8���E, 61m), 21.vi.1911, Coll. T.B.F, Repository? (Type specimen missing). Diagnosis. Body length 8.0 mm. Frons strongly arcuate striate; vertex and occiput trans-striate; vertex with two carinae between posterior ocelli; short neck; pronotum trans-striate, extreme base smooth; mesoscutum finely punctate; metapleura coarsely punctate; wings slightly iridescent; hind coxa coarsely striate; hind femora finely trans-striate, ventrally bidentate; first metasomal tergite finely tans-striate, as long as the remaining metasomal segments combined. Female. Unknown. Distribution. Oriental: India (West Bengal). Host. Unknown. Note. The types of this species seems to be lost, therefore, the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 43, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1919 b) Two new Stephanidae. The Entomologist, 52, 162 - 163."]}

Key to genera of Stephanidae occurring in India (Modified from van Achterberg 2002) 1. First subdiscal cell of fore wing comparatively wide basally, wider than first discal cell or vein 1-SR of fore wing not differentiated and first discal cell absent because of absence of vein 1-SR+M (Figs 11, 37, 51); hind tibia usually hardly longer than hind femur (Figs 12, 40, 52); inner side of hind tibia with a long oblique depression..................................... 2 - First subdiscal cell of fore wing comparatively narrow basally, about as wide as first discal cell or narrower and vein 1-SR of fore wing differentiated with first discal cell present because of presence of vein 1-SR+M (Figs 1, 80, 91, 102); hind tibia usually distinctly longer than hind femur (Figs 66, 79, 92, 103); inner side of hind tibia variable, frequently without long oblique depression........................................................................................... 3 2. Veins 2-SR and 2-SR+M of fore wing absent; vein 1-SR of fore wing not or hardly differentiated because of absence of vein 1-SR+M; sclerite present between hind coxae; neck moderately to very slender and finely striate and no distinct pronotal fold, rarely with weakly developed pronotal fold and specialised (Figs 10, 18, 36, 49); outer side of hind tibia posteriorly usually without fine oblique striae and/or ventrally with fine carina; vein 2-CU1 of fore wing nearly always reduced, but sometimes complete; metapleuron slender; ovipositor sheath with or without a subapical ivory or whitish band (completely black: Figs 3, 24, 30, 43; with subapical ivory band: Figs 2, 4, 14, 21, 23, 25, 54)................................ Foenatopus Smith - Veins 2-SR and 2-SR+M of fore wing present (Figs 72, 80)[vein 2-SR+M absent in P. reductus sp. nov. (Fig. 91)], sometimes only pigmented; vein 1-SR of fore wing distinctly differentiated because of presence of vein 1-SR+M; sclerite absent between hind coxae; neck short and comparatively robust, without pronotal fold or transverse carinae; outer side of hind tibia with distinct oblique striae ventrally (but often fine or only ventrally distinctly developed and frequently with some rugulosity and/or apical half of tibia with ventral carina more or less developed), rarely without striae or ventral carina; vein 2-CU1 of fore wing completely developed; metapleuron robust; ovipositor sheath without ivory or whitish subapical band (Figs 83, 94)..................................................................................... Parastephanellus Enderlein 3. Hind tarsus of female with five tarsomeres; hind femur distinctly slender and elongate, coarsely striate, ventrally with 2���3 large teeth, rarely with 4 teeth (Fig. 103); inner side of hind tibia only with a short narrow oblique groove below a small convexity..................................................................................... Stephanus Jurine - Hind tarsus of female with three tarsomeres; hind femur comparatively robust and less elongate, largely smooth and with some punctures, ventrally with 2 large teeth; inner side of hind tibia usually with wide submedial ventral carina and or with oblique striae ventro-posteriorly (Fig. 66); [ovipositor sheath with ivory subapical band; pronotum without transverse protuberance (Figs 56, 71); temple without pale yellowish streak behind eye, at most with a ventral patch; outer side of hind tibia without oblique striae or rugulosity; vein 1-M of fore wing 3.5���8.0 �� as long as vein 1-SR (Fig. 1)]............. Megischus Brull��, Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 3-4, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206."]}

Genus Stephanus Jurine, 1800 Stephanus Jurine (in Panzer) 1800: 76, Fig. 13. Type species (by monotypy): Stephanus coronatus Jurine (in Panzer), 1800 (= Ichneumon serrator Fabricius, 1798). Diagnosis. Temple at most with a yellow patch, without any streak along the eye; sdc1 of fore wing basally narrow; 1-SR differentiated with dc1 present; 1-SR+M of fore wing present; hind wing with M+CU, no cu-a vein; hind femur distinctly slender and elongate with coarse striations, ventrally with 2���3 (rarely 4) large teeth; inner side of hind tibia only with a short narrow oblique groove below a small convexity; hind tarsus of female with five tarsomeres; ovipositor sheath completely blackish or brown without a subapical yellow patch. Distribution. Palaearctic and Oriental (van Achterberg 2002; van Achterberg & Yang 2004; Hong et al. 2011). Notes. Five species in the genus Stephanus are known worldwide. In this paper, we describe the male specimen of species S. bidentatus from India, previously described from China. Since part of Andaman and Nicobar islands (India: Nicobar islands) lies in the distinct Sundaland biogeographical region of the Oriental region, there is a high possibility of occurrence of species reported from the Sunda area (S. borneensis and S. soror) from the Indian mainland. A key to world species of Stephanus is also provided for identification, if encountered., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 43-44, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206.","van Achterberg, C. & Yang, Z-Q. (2004) New species of the genera Megischus Brulle and Stephanus Jurine from China (Hymenoptera: Stephanoidea: Stephanidae), with a key to world species of the genus Stephanus. Zoologische Mededelingen Leiden, 78 (3), 101 - 117.","Hong C. - D., van Achterberg & Xu, Z. - F. (2011) A newly recorded genus and species of Family Stephanidae (Hymenoptera, Stephanoidea) from China. Entomotaxonomia, 33 (1), 71 - 73."]}

Foenatopus longicaudalis (Elliott, 1919) (Fig. 26) Diastephanus longicauda Elliott, 1919a, Entomologist, 52, 30. Type material. Holotype ♀, India: Nilgiri hills (= Kerala / Tamil Nadu?) (3000 m), 14.iv1888, Coll. Sir George Hampson. (NHM). Diagnosis. Body length 15 mm, ovipositor sheath length 22 mm. Frons trans-rugose; vertex finely trans-striate with three stout carinae; pronotum finely trans-striate; metapleuron coarsely punctate; propodeum punctate; first metasomal tergite a little longer than remaining segments combined; hind coxae trans-striate; hind femur tridentate ventrally; wings infuscate; ovipositor sheath longer than metasoma, with a subapical ivory ring. Male. Unknown. Distribution. Oriental: India (Nilgiri hills = Kerala / Tamil Nadu). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 18, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1919 a) Some Stephanidae in the British Museum. The Entomologist, 52, 28 - 32."]}

Genus Foenatopus Smith, 1861 Foenatopus Smith 1861: 58. Type species (by monotypy): Stephanus indicus Westwood, 1841. Diastephanus Enderlein 1905: 473. Type species: Stephanus flavomaculatus Enderlein, 1901. Synonymized by Benoit, 1956. Neostephanus Kieffer 1904: 1���4. Type species (by monotypy): Neostephanus alluaudi Kieffer, 1904. Synonymized by Narendran et al. 2001. Diagnosis. Members of the genus are characterized by the presence of pale ivory to yellowish streak along both the inner and outer margins of eye; reduced venation with the absence of veins 2-SR and 2-SR+M of the fore wing and the absence of or hardly seen 1-SR vein; Vein 2 Cu1 of fore wing nearly always reduced, but sometimes complete; 2-SR and 2-SR+M of fore wing absent; anteriorly emarginate neck with fine striae, no distinct pronotal fold basally; the presence of a large sclerite in between the hind coxae; outer side of hind tibia usually without fine oblique striae, with or without ventral fine carinae; three tarsomeres in females, five tarsomeres in males; eighth metasomal tergite without apical protuberance; ovipositor sheath usually with ivory or yellowish subapical band (Hong et al. 2011; Gadallah & Edmardash 2015). Distribution. Afrotropical, Neotropical, Palaearctic and Oriental. Notes. Foenatopus Smith is represented by 159 species worldwide of which 20 were reported from India (Narendran & Sureshan 2003; Sureshan et al. 2013; Gupta & Gawas 2020). Two new species viz., F. chareshi Binoy, van Achterberg & Girish Kumar sp. nov. and F. sureshani Binoy, van Achterberg & Girish Kumar sp. nov. are described here with illustrations and is incorporated into a key to Indian species modified from Narendran et al. (2001) and Narendran & Sureshan (2003)., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 4, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Westwood, J. O. (1841) On Evania and some allied genera of Hymenopterous insects. Annals and Magazine of Natural History, Series 1, 7, 535 - 539.","Enderlein, G. (1905) Uber die Klassifikation der Stephaniden. Zoologischer Anzeiger, 28 (13), 473 - 477.","Benoit, P. L. G. (1956) Nouvelles especes africaines du genre Foenatopus Smith (Hym. - Stephanidae). Bulletin et Annales de la Societe Royale d'Entomologie de Belgique, 92 (7 - 8), 205 - 212.","Kieffer, J. J. (1904) Description de Stephanides et d'Evaniides nouveaux. Bulletin de la Societe d'Histoire Naturelle de Metz, Serie 11, 2, 23, 1 - 30.","Hong C. - D., van Achterberg & Xu, Z. - F. (2011) A newly recorded genus and species of Family Stephanidae (Hymenoptera, Stephanoidea) from China. Entomotaxonomia, 33 (1), 71 - 73.","Gadallah, N. S. & Edmardash, Y. A. 2015. First record of Stephanidae (Hymenoptera, Stephanoidea) for the fauna of Egypt. Zootaxa, 3980 (1), 147 - 150. https: // doi. org / 10.11646 / zootaxa. 3980.1.10","Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138.","Sureshan, P. M., Narendran, T. C. & Nikhil, K. (2013) Parasitoids (Hymenoptera) of xylophagous beetles (Coleoptera) attacking dead wood in southern Western Ghats, Kerala, India, with descriptions of two new species. Journal of Threatened Taxa, 5 (9), 4385 - 4391. https: // doi. org / 10.11609 / JoTT. o 3385.4385 - 91","Gupta, A & Gawas, S. M. (2020) A new species of the genus Foenatopus Smith (Hymenoptera: Stephanoidea: Stephanidae) from India. Zootaxa, 4801 (2), 389 - 394. https: // doi. org / 10.11646 / zootaxa. 4801.2.13"]}

Genus Megischus Brull��, 1846 Megischus Brull��, 1846: 537. Type species (designated by Viereck, 1914): M. annulator Brull��, 1846. Megiseleus Cameron, 1902: 32. Lapsus calami. Bothriocerus Sichel, 1861: 759. Type species (by monotypy): Bothriocerus europaeus Sichel, 1861 (= Stephanus anomalipes Foerster, 1855, synonymy according to Madl, 1991). Diagnosis. Largest known stephanids ranging up to 35 mm (excluding ovipositor); length of fore wing 5���20 (but usually 10���18) mm; rounded temple without conspicuous ivory-white streak behind eye; neck mostly elongate; pronotal folds distinct; fore wing with 2-SR and 2-SR+M present, vein r of fore wing straight, in most cases distinctly removed from apex of pterostigma; vein 2-SR of fore wing medium-sized and straight; vein 1-SR of fore wing differentiated because of presence of vein 1-SR+M; bidentate hind femora; hind tibia distinctly narrowed basally and at most basal half ventrally carinate; outer side of hind tibia without oblique striae or rugulosity; ovipositor sheath with ivory subapical band. Distribution. Mainly Indo-Australian and Neotropical, with scattered reports from subtropical Oriental, Southern Nearctic and Central and Southern Europe (van Achterberg 2002). Host. Probably attacking beetle larvae infesting dead trees (van Achterberg 2002). Notes. The genus comprises more than 50 species worldwide (New World: 10 spp.; West Palaearctic: 1 species; Australian continent: 2 species (Aguiar 2001), Oriental, Wallacea and Papua: 39 species (van Achterberg 2002)). The genus is represented by a single species viz., M. alveolifer van Achterberg from India. A new species viz., M. ranjithi Binoy, Santhosh & Girish Kumar sp. nov. is described here with illustrations and is incorporated into a key to Indian species., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 30, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Brulle, A. (1846) Histoire naturelle des Insectes. Hymenopteres, Paris, 4, i-viii + 1 - 680.","Viereck HL (1914) Type species of the genera of ichneumon flies. Bulletin of the United States National Museum, 83, I-V + 1 - 186. https: // doi. org / 10.5479 / si. 03629236.83.1","Cameron, P. (1902) On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society, 37, 29 - 131.","Sichel, J. (1861) Liste des Hymenopteres recueillis en Sicile par M. E. Bellier de la Chavignerie. Annales de la Societe entomologique de France, Series 3, 7, 749 - 764. [1860]","Madl, M. (1991) Zur Kenntnis der palaarktischen Stephanidae (Hymenoptera, Stephanoidea). Entomofauna, 12, 117 - 126.","van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206.","Aguiar, A. P. (2001) Revision of the Australian Stephanidae (Hymenoptera). Invertebrate Taxonomy, 15 (6), 763 - 822. https: // doi. org / 10.1071 / IT 00006"]}

Parastephanellus reductus Binoy, Girish Kumar & Santhosh sp. nov. (Figs 84���94) urn:lsid:zoobank.org:act: 1EBB1316-6C62-499F-96E0-B9CCC2A19C5D Type material. India: Tamil Nadu, Kanyakumari district, Kanyakumari Wildlife Sanctuary, Alagiapandiapuram (8��18���27.7���N & 77��26���19.1���E, 30m), 16.ii.2020, Coll. Souvik Sen & party, (ZSIK). Reg. No. ZSI/ WGRC /IR/ INV.14261. Diagnosis. The new species possesses an important and unique character in having the fore wing with vein 2- SR+M absent. Head in dorsal view rounded behind eyes; neck moderately emarginated; pronotum posteriorly well emarginate, forming a gap between a lower mesoscutum; hind coxa with strong transverse rugae and without a dorsal tooth; first metasomal tergite almost equal to rest of metasoma (excluding ovipositor), 6.7�� as long as its maximum width with strong irregular coarsely transverse rugae anteriorly, proceeding as annulate sculpturing beyond. Description. Holotype, ♀. Body length 6.01 mm, ovipositor sheath length 3.82 mm, fore wing length 3.26 mm. Colour. Black; tegula and palpi dark brown; scape brown with apical tip honey brown, pedicel, F1���F4, F5 and F6 basally honey brown, rest of flagellomeres deep brown; frons brown with yellowish patches near inner orbits and on lower face along the toruli; malar space with a pale yellow patch below eyes (Fig. 86) extending to the occiput posteriorly; mandibles yellowish brown with apices black (Fig. 88); all tibia liver brown, paler apically; all tarsi yellowish brown to testaceous, hind basitarsus pale yellowish; wings hyaline, pterostigma translucent, margins brown, veins mostly translucent brown; first metasomal tergite apically and second metasomal tergite basally brownish; second tergite onwards pale orange brown apical margins; apical tip of hypopygium orange brown; ovipositor sheath brownish black with a small pale spot on expanded ventral and apical area. Head. Antenna with 29 antennomeres; F2 onwards surface distinctly pitted; frons with subparallel uniform rugae, medially coarse, few scattered setae; coronal area smooth with few strong rugae; clypeus projecting, rugose, with apical brush of setae; five anterior coronal teeth, antero-medial one largest, reddish brown, followed by a pair of arcuate ones and blunt ones posteriorly; behind the posterior teeth a couple of straight carinae present; vertex faintly rugose beyond the lateral ocellus along the eye margin, medially bare, almost smooth (Fig. 87); temples non-angulate, largely smooth and shiny (Fig. 86); occipital carina strong, complete, almost straight, reaching the mandibles anteriorly. Mesosoma. Neck short, emarginate anteriorly, in level with pronotum without forming a cavity medially; pronotum medially moderately rugose without any distinct carinae, sparsely setose, posteriorly smooth, shiny with margin emarginated; pronotal margin forming a large gap at pronotal-mesoscutal margin; propleuron smooth and shiny forming buttressing ridges ventrally, sparsely setose; mesoscutum with medial foveolate furrow; anteriorly imbricate to alutaceous, medially with irregular fovea and a pair of converging accessorial carina beyond middle, 0.7�� as wide as long, sparsely setose; scutellum large, as long as wide, alutaceous with scattered punctures and sparsely setose; axillae separated from each other by a deep and an indistinct fovea (Fig. 89); mesopleuron alutaceous with moderately dense white setae, smooth posteriorly (Fig. 90); propodeum with large irregular areola and smooth to imbricate interstices, inside of areola smooth, a transverse curving carina present sub apically (Fig. 89); fore wing hyaline with scattered minute setae concentrated more on apically, 2-SR+M absent, 1-M 1.43�� 1-SR, almost straight; vein r reaching before the apex of pterostigma; 3-CU 1 obsolete, 2-1A absent (Fig. 91); hind coxa robust, without a dorsal teeth and with strong rugae anteriorly, turning annular posteriorly; hind femur moderately widened, smooth, alutaceous sculpturing and scattered sparse long white setae, ventrally with 2 large teeth and several denticles in between and beyond apical teeth, basal tooth wide, faintly rugose; hind tibia with uniform rugae ventrally, moderately pubescent, 1.27�� as long as hind femur, apical inflated part (i.e., apical part with sculpture different from basal petiolate part) 1.06�� as wide as narrow basal part, widened part with distinct oblique inverted obsolete V shaped carinae; hind tarsus with three tarsomeres, sub-parallel sided 4�� as long as second tarsal segment (Fig. 92). Metasoma. First tergite 8.6�� as long as its maximum width, 0.73�� as long as remaining metasomal tergites, cylindrical with irregular strong rugae throughout the length, apically smooth (Fig. 93); second tergite smooth and shiny medially, fine alutaceous laterally and apically; fifth to seventh tergites finely reticulate to alutaceous with scattered small white setae; eighth tergite finely reticulate, setose and furcate apically, no pygidial protuberance present apically; ovipositor sheath 0.63�� length of the body and 1.17�� as long as fore wing (Fig. 94). Male. Unknown. Distribution. Oriental: India (Tamil Nadu). Biology. Unknown. Etymology. The species is named after the peculiar feature in having vein 2-SR+M of fore wing reduced/absent., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 40-43, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367

Stephanus bidentatus van Achterberg, 2002 (Figs 95���105) Stephanus bidentatus van Achterberg, 2002, Zool. Meded. Leiden, 78 (3), 104. Type material. Holotype ♀, China: Henan, Longyuwan, Lianchuan, (29��55���46.1���N & 118��29���35.5���E, 700 m), on trunk of Quercus tree with Cerambycidae larvae, 13.vii.1996, Coll. Zhong-qi Yang, (CAFB). Material Examined: 1 ♂, mounted on triangular card, India: Andaman and Nicobar, Nicobar Islands, Great Nicobar, Chingen Basti (6��49���14.1���N & 93��53���05.1���E, 76 m), 08.xi.2010, Coll. E. E. Jehamalar, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13599. Description. ♂ (hitherto unknown). Body length 14.1 mm. Fore wing length 7.5 mm. Colour. Reddish brown with the following parts different: face reddish orange; eyes golden to yellowish orange; mandibles apically black; ocelli reflective golden yellow; antennae with scape and pedicel reddish orange; rest of the antennomeres brown or reddish brown; fore wing membrane mainly with brownish infumation; pterostigma dark brown; hind trochantellus apically, and fore and middle legs reddish brown; tarsi and palpi brown. Head. Third antennal segment moderately slender, 2.3�� as long as wide, and 0.8�� as long as fourth segment (antenna broken after 10 antennomeres); three anterior coronal teeth of head large, hardly larger than posterior ones (Fig. 97 & 98); with four curved carinae behind level of both posterior coronal lobes, fourth one shallow; remaining part of vertex medio- dorsally with finely curved rugae, becoming finer posteriorly and narrowly reaching occipital carina (Fig. 99); temples smooth and with sparse setae ventrally (Fig. 97); dorsally occipital carina distinct, carina ventrally weaker than laterally and reaching hypostomal carina; postgenal bridge steep. Mesosoma. Neck robust and rather short, anteriorly distinctly concave, lateral length of neck 0.83�� its maximum width, neck postero-dorsally lower than medial part of pronotum, and with five coarse carinae laterally, posterior two presented medio-anteriorly, large cavity in middle part of pronotum and pronotal fold absent medially; middle part of pronotum robust, coarsely rugose, without median carina, with some irregular and rather strong transverse rugae, middle part not differentiated from posterior part of pronotum, without steps between parts; pronotum setose dorso-laterally with dorso-median bare area; distinctly rugose; mesoscutum densely punctate with a distinct median groove formed of deep pits; scutellum smooth medially, scattered pits laterally; axillae smooth with few deep pits, separated from scutellum by a deep large fovea (Fig. 100); propleuron faintly convex, rugose and highly setose; mesopleuron punctate with rugose interspaces and densely setose, convex part rugose-reticulate with lesser setosity near the mesopleura-metapleura junction; metapleura shiny, with large rugae (Fig. 101); propodeum completely and coarsely punctate with pits and interspaces smooth and shiny; fore wing: vein 1-M almost straight, 4.4�� as long as vein 1-SR; wing basally and area below pv and pterostigma largely glabrous; vein M+CU of hind wing only partly pigmented, after middle of wing (Fig. 102); hind coxa slender, largely rugose and shiny, anteriorly and ventrally less sculptured, densely setose with long dirty white setae; hind femur with two large teeth, 3���4 denticles in between and few denticles apically, femur 3.2�� as long as its maximum width, basal half smooth, rest with micro pitted, with long dirty white setae arising from scattered pits; setosity concentrated more on ventral margin; basal narrow part of hind tibia parallel-sided and 0.55�� as long as widened part, and with ventral carina, outer side of widened part of hind tibia coriaceous, convex, antero-dorsally with weak tubercle, ventrally with distinct oblique striae and no median carina, and rather narrowed apically, inner side flattened and widened part largely with densely bristly setosity, no distinct punctures, and with indistinct depression below tubercle (Fig. 103); hind basitarsus parallel-sided, basally hardly curved, its ventral length 5�� its width. Metasoma. First tergite slender, 13.5�� as long as its apical width width, and largely irregularly and coarsely transversely rugose on basal third, remainder finely and more densely rugose, parallel-sided (Fig. 104); second tergite mainly smooth basally, remainder part and rest of tergites microsculptured with sparse setosity latero-ventrally (Fig. 105). Host. Probably a parasitoid of Cerambycidae larvae in Quercus species and other deciduous trees (van Achterberg &Yang 2004); unknown in the present study. Distribution. Palaearctic (China) (van Achterberg & Yang 2004; Hong et al. 2011), Oriental (India (present report))., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 44-47, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206.","van Achterberg, C. & Yang, Z-Q. (2004) New species of the genera Megischus Brulle and Stephanus Jurine from China (Hymenoptera: Stephanoidea: Stephanidae), with a key to world species of the genus Stephanus. Zoologische Mededelingen Leiden, 78 (3), 101 - 117.","Hong C. - D., van Achterberg & Xu, Z. - F. (2011) A newly recorded genus and species of Family Stephanidae (Hymenoptera, Stephanoidea) from China. Entomotaxonomia, 33 (1), 71 - 73."]}

Parastephanellus malabaricus Narendran & Sheela, 1995 Parastephanellus malabaricus Narendran & Sheela, 1995, Bull. Pure and Appl. Sci., 14A, 45���47. Type material. Holotype ♀, India: Kerala, Malappuram district, Nedumkayam forest (11��17���28.7���N & 76��20���24.3���E, 400 m), 12.i.1995, Coll. P. Aboobacker, (ZSIK) (Type specimen missing). Diagnosis. Body length 5 mm; ovipositor sheath length 4.8 mm. Frons transversely striate with oblique striation between coronal teeth; vertex mostly smooth with a pair of strong carinae followed by weak striae and a posterior longitudinal medial grove; pronotum smooth, pronotal fold concave; mesoscutum with deep scattered pits; scutellum polished, twice as long as mesoscutum; metapleuron rugose reticulate; propodeum rugose, sparsely punctured; hind coxa transversely striate; hind femora bidentate ventrally with few tubercles; first metasomal tergite finely trans-carinate, a little shorter than remaining segments except ovipositor; ovipositor sheath a trifle longer than body, without a subapical pale band. Male. Unknown. Distribution. Oriental: India (Kerala). Host. Unknown. Note. The holotype was not traceable, therefore, the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 36, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sheela, S. (1995) On a new species of Parastephanellus Enderlein (Hymenoptera: Stephanidae) from India. Bulletin of Pure and Applied Sciences, 14 A (1), 45 - 47."]}

Foenatopus iridipennis Elliott, 1919 Foenatopus iridipennis Elliott, 1919a, Entomologist, 52, 30. Type material. Syntype ♀, India: Uttarakhand, Dehradun (30��19���4.8���N & 78��1���44.4���E, 447m), ix.1907, Coll. Lt. Col. F.W. Thomson, I.M.S. Syntype ♂, India: Himachal Pradesh, Kangra Valley (32��10��� N & 76��30��� E, 610m), vii.1889, Coll. Dudgeon. Repository? (Type specimens missing). Diagnosis. Body length 15 mm, ovipositor sheath length 22 mm. Frons arcuate rugose; vertex distinctly transstriate with two short carinae; neck smooth, pronotum trans-striate with narrow smooth posterior margin; mesoscutum with large well separated punctures; metapleura cribrate punctate, separated by a carina; wings hyaline, strongly iridescent; hind coxa finely trans-striate; hind femora tridentate; first metasomal tergite trans-striate, as long as rest of metasomal tergites combined; ovipositor sheath half as long as body with a subapical pale ivory band. Male. Placed with the female; slender, head finely striate; metapleura striate; colour of head testaceous while red in female. Distribution. Oriental: India (Himachal Pradesh, Uttarakhand). Host. Unknown. Note. The types are not available, therefore, the above mentioned diagnosis is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 16, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1919 a) Some Stephanidae in the British Museum. The Entomologist, 52, 28 - 32."]}

Foenatopus anupam (Narendran and Sureshan, 2003) (Fig. 3) Diastephanus anupam Narendran and Sureshan, 2003, Entomon, 28(2), 124���127. Type material. Holotype ♀, mounted on triangular card, India: Arunachal Pradesh, West Kameng district, Bhalukpung (27��01���12.5���N & 92��38���25.9���E, 213 m), 25.x.1997, Coll. T. C. Narendran, (ZSIK) Regd. No. ZSI/ WGRC / IR/INV.1891. Examined. Additional material examined: 1♀, India: Tamil Nadu, Thirunelveli district, Kalakkad Mundanthurai Tiger Reserve, Karaiyar dam site (8��39���23.7609���N & 77��18���7.8669���E), 29.ix.2017, Coll. P. Girish Kumar, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13600. Diagnosis. Body length 10.4 mm; ovipositor sheath length 8.8 mm; fore wing length 5.2 mm. Frons trans-striate with interstices between striae microsculptured; vertex with four strong carinae followed by transverse striae not forming a medial line or grove; lower head white; pronotum strongly trans-carinate anteriorly becoming trans-reticulate posteriorly; scutellum finely reticulate; metapleuron rugose-reticulate; hind coxa transversely carinate; hind femora tridentate ventrally without any distinct tubercles in between; first metasomal tergite uniformly annulate, 1.2�� as long as remaining segments; ovipositor sheath 1.4�� length of metasoma, without a subapical pale band. Male. Unknown. Distribution. Oriental: India (Arunachal Pradesh, Tamil Nadu (present record)). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 7-8, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138."]}

Genus Parastephanellus Enderlein, 1906 Stephanus (part) Westwood, 1874: 126. Parastephanus Enderlein, 1905: 474. [Preoccupied by H��ckel for Protista]. Parastephanellus Enderlein, 1906: 301. Hemistephanus Moore, 1962: 93. [Misidentification] Parastephanellus (Parastephanellus) Narendran & Sheela, 1995: 45. Type species by original designation: Stephanus pygmaeus Enderlein, 1901. Diagnosis. Members of Parastephanellus Enderlein are characterised by a pale yellowish streak along the eye as in Foenatopus; short and robust neck, without a pronotal fold medially or carinae; 2-CU1 of fore wing complete; 2-SR and 2-SR+M of fore wing present (vein 2S-SR+M absent in P. reductus sp. nov.), sometimes only pigmented; 1-SR of fore wing straight; hind tibia with distinct oblique striae or carinae ventrally, rarely without; ovipositor sheath without ivory subapical band. Distribution. Australian, Palaearctic and Oriental. Notes. 58 species in the genus Parastephanellus are known worldwide, of which only three species are reported from India (Elliot 1919b; Narendran & Sheela 1995; Narendran & Sureshan 2001). Two new species viz., P. narendrani Binoy, van Achterberg & Santhosh sp. nov. and P. reductus Binoy, Girish Kumar & Santhosh sp. nov. are described here with illustrations and are incorporated into a key to Indian species., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 35, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Enderlein, G. (1906) Neue Beitrage zur Kenntnis und Klassifikation der Stephaniden. Stettiner Entomologische Zeitung, 47, 289 - 306.","Westwood, J. O. (1874) Thesaurus Entomologicus Oxoniensis; or illustrations of new, rare, and interesting insects, for the most part contained on the collections presented to the University of Oxford by the Rev. F. W. Hope ... with forty plates from drawings by the author. Oxford. At the Clarendon press, Oxford, 205 pp., 40 pls. https: // doi. org / 10.5962 / bhl. title. 14077","Enderlein, G. (1905) Uber die Klassifikation der Stephaniden. Zoologischer Anzeiger, 28 (13), 473 - 477.","Moore, K. M. (1962) Observations on some Australian forest insects. Proceedings of the Royal Zoological Society of New South Wales, 59, 87 - 95.","Narendran, T. C. & Sheela, S. (1995) On a new species of Parastephanellus Enderlein (Hymenoptera: Stephanidae) from India. Bulletin of Pure and Applied Sciences, 14 A (1), 45 - 47.","Narendran, T. C. & Sureshan, P. M. (2001) A new species of Parastephanellus Enderlein (Hymenoptera: Stephanidae) from India. Zoos' Print Journal, 16 (10), 597 - 599. https: // doi. org / 10.11609 / JoTT. ZPJ. 16.10.597 - 9"]}

Foenatopus indicus (Westwood, 1841) Stephanus indicus Westwood, 1841, Ann. Mag. Nat. Hist., VII, 298. Magischus indicus Westwood, 1851, Trans. Ent. Soc. Lond., 2, 1. Foenatopus ruficeps Smith, 1857, J. Proc. Linn. Soc. Zool., V, 58. Foenatopus indicus Enderlein, 1913, Zoologische, XIII, 290., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 15, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Westwood, J. O. (1841) On Evania and some allied genera of Hymenopterous insects. Annals and Magazine of Natural History, Series 1, 7, 535 - 539."]}

Foenatopus wynadensis (Sureshan & Narendran, 1997) (Fig. 55) Diastephanus wynadensis Sureshan & Narendran, 1997, Gebios, 16, 25���27. Diastephanus wayanadensis Narendran & Sureshan, 2003, Entomon, 28(2), 136 (invalid emendation). Type material. Holotype ♀, India: Kerala Wayanad district, Muthanga forest (10��47���N & 76��02���E, 26 m), 08.viii.1999, Coll. P. Sudha, (ZSIK). Reg. No. ZSIK / WGRS /IR-INV. 2340. Examined. Diagnosis. Body length 10.7 mm, ovipositor sheath length 10.8 mm. Frons trans-striate except above coronal teeth obliquely striate; vertex trans-carinate anteriorly and trans-striate posteriorly; pronotum with neck trans-striate dorsally and anteriorly semi annular trans-striate proceeding as polished posteriorly; mesoscutum faintly reticulate; scutellum and axillae smooth and polished; metapleuron rugose-reticulate; propodeum with broad and deep punctures, sparse anteriorly with shagreened interstices; first metasomal tergite uniformly trans-carinate, a little shorter than combined length of remaining segments; ovipositor very long, 3�� as long as metasoma, with a subapical ivory band. Male. Unknown. Distribution. Oriental: India (Kerala). Host. Unknown, collected from moist deciduous forest., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 29-30, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Sureshan, P. M. & Narendran, T. C. (1997) A new species of Diastephanus Enderlein (Hymenoptera: Stephanidae) from India. Geobios new Reports, 16, 25 - 27.","Narendran, T. C. & Sureshan, P. M. (2003) Systematic studies on Diastephanus Enderlein (Hymenoptera: Stephanoidea: Stephanidae) of Indian subcontinent. Entomon, 28 (2), 115 - 138."]}

Foenatopus rugiceps (Elliott, 1919) (Fig. 28) Diastephanus rugiceps Elliott, 1919a, Entomologist, 52, 29. Type material. Holotype ♀, India: South India Collection details incomplete. Coll. F. Smith. (NHM) B. M. TYPE HYM. 3a. 41 (NHMUK013458835). Diagnosis. Body length 12.0 mm, ovipositor sheath length 10.0 mm. Frons granulate rugose; vertex with three carinae; pronotum strongly trans-striate; mesoscutum coarsely punctate, apically smooth; first metasomal tergite trans-striate, as long as combined remaining segments; latter segments alutaceous; ovipositor sheath longer than metasoma, with an ill-defined subapical pale yellowish ring. Male. Unknown. Distribution. Oriental: India (South India). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 20-21, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Elliott, E. A. (1919 a) Some Stephanidae in the British Museum. The Entomologist, 52, 28 - 32."]}

Foenatopus similicus Narendran, 2001 (Fig. 29) Foenatopus similicus Narendran, 2001, Entomon, J. Adv. Zool., 22(2), 85���86. Type material. Holotype ♀, India: Himachal Pradesh, Shimla district, Shimla (31�� 6��� N & 77�� 10��� E, 2276 m), 01.vii.1918, Coll. Brunetti. (NHM) B. M. TYPE HYM. 3a. 375 (NHMUK013458833) Diagnosis. Body length 20.1 mm, ovipositor length 27.0 mm. Frons with irregular carinae and pits with microsculptured interstices; vertex with 4 or 5 carinae followed by closely packed cross striae and a median shallow grove; neck deeply emarginate, pronotal fold indistinct, trans-striate anteriorly weakening as proceeding posteriorly; mesoscutum with a broad reticulate area anteriorly, medial grove absent; scutellum smooth and shiny with faint aciculae and sparse pits laterally; metapleuron closely punctate; propodeum with close deep pits, interstices carinate with inside of pits and interstices reticulate; hind coxa finely and strongly annulate, irregular striae at base; hind femur tridentate and few denticle ventrally, basal tooth shorter; first metasomal tergite annulate, shorter than remaining segments; ovipositor distinctly longer than metasoma (ovipositor sheath broken). Male. Unknown. Distribution. Oriental: India (Himachal Pradesh). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 21, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367

Foenatopus maheshi Narendran, 2001 (Fig. 27) Foenatopus maheshi Narendran, 2001, J. Adv. Zool., 22(2), 81���83. Type material. Holotype ♀, India: Kerala Palakkad district, Nelliampathy forests (10�� 31��� N & 76�� 41��� E, 1500 m), 18.vi.1995, Coll. A. Mahesh Kumar, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.1301. Examined. Paratype 1♀, India: Kerala Palakkad district, Silent Valley National Park (11�� 8��� N & 76�� 28��� E, 2000 m), 18.v.1995, Coll. C. F. Binoy. Diagnosis. Body length 19.0 mm, ovipositor sheath length 28.0 mm. Frons with longitudinal irregular carinae and pits; vertex with two distinct cross carinae followed by distinct close and a distinct medial pitted longitudinal fovea; neck anteriorly emarginate; pronotum trans-carinate anteriorly, turning finely reticulate posteriorly; mesoscutum anteriorly cross reticulate, without a medial line of pits; scutellum smooth, shiny and faintly reticulate with shallow pits on sides; metapleuron irregularly rugose punctate; hind coxa irregularly rugose basally turning circular annulated elsewhere; hind femora tridentate ventrally with a couple of denticles in between; first metasomal tergite annulate, a little shorter than combined length of remaining segments; ovipositor sheath longer than metasoma, with a subapical ivory band. Male. Unknown. Distribution. Oriental: India (Kerala). Host. Unknown., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 19, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367

Key to species of the genus Stephanus Jurine (Modified from van Achterberg, 2002) 1. Temples smooth and shiny, at most with few punctures; hind femur shiny, largely smooth, superficially coriaceous or finely striate; outer side of hind tibia of female with fine striae ventrally and ventrally with carina; base of pterostigma with indistinct pale brownish spot; femora blackish or dark brown dorso-apically; head dark orange- or reddish-brown and with yellowish spot between base of mandible and eye, hardly contrasting with temple or blackish and with distinct ivory spot; hind tibia of female blackish or brown [East Palaearctic, Oriental (Sunda area)].................................................... 2 - Temples coriaceous, with several coarse punctures and rather matt; hind femur matt, distinctly finely granulate; outer side of hind tibia of female without striae ventrally and ventrally without carina, only more or less angulate; base of pterostigma with distinct white spot; femora narrowly white-tipped dorso-apically (female, more conspicuously so in male); head largely blackish or dark brown and with ivory spot between base of mandible and eye, distinctly contrasting with temple; hind tibia of female yellowish-brown (of male more or less dark brown) [West Palaearctic].......................... S. serrator (Fabricius) 2. First metasomal tergite about 9�� as long as its apical width (female; male: 13.5��) (Fig. 104); hind femur with 2 large ventral teeth (Fig. 103); pronotum with deep cavity and no distinct pronotal fold medially; small part of vein M+CU of hind wing pigmented (Fig. 102); scutellum densely rugose [East Palaearctic (China); Oriental (India (present report))]........................................................................................... S. bidentatus van Achterberg - First tergite 3���5�� as long as its apical width; hind femur with 3 large ventral teeth, rarely with 4 teeth; pronotum with indistinct to medium sized cavity and with a coarse to fine pronotal fold medially; large part of vein M+CU of hind wing pigmented; scutellum largely smooth................................................................................ 3 3. Head blackish and with a distinct ivory spot, contrasting with temple; temples rather angulate in dorsal view; pronotum without a distinct cavity below pronotal fold; medially vertex regularly and rather finely transversely striate; hind femur partly finely (rugose-)striate [East Palaearctic (China)]........................................... S. tridentatus van Achterberg - Head dark orange or reddish brown and with yellowish spot between base of mandible and eye, hardly contrasting with temple; temples roundly narrowed in dorsal view; pronotum with a cavity below pronotal fold; medially vertex irregularly and densely rugose; hind femur mainly smooth, at most superficially coriaceous [Oriental (Sunda area)].......................... 4 4. Hind basitarsus dark brown, and not contrasting with hind tibia; at least basal half of outer side of hind coxa finely irregularly rugose; hind leg more robust; pronotum and first metasomal tergite rather coarsely sculptured [Oriental (Borneo, Java)]................................................................................... S. borneensis (de Saussure) - Hind basitarsus rather pale brown, and contrasting with dark hind tibia; outer side of hind coxa largely smooth; hind leg slender; pronotum and first tergite finely sculptured [Oriental (West Malaysia)]......................... S. soror van Achterberg, Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 44, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["van Achterberg, C. (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen Leiden, 339, 1 - 206."]}

Foenatopus chinnarensis (Sureshan, 1997) Diastephanus chinnarensis Sureshan, 1997, Bioved, 3, 89���92. Type material. Holotype ♂, India: Kerala, Idukki district, Chinnar (10��30��� N & 77��20��� E, 450 m), 19.xi.1996, Coll. P.M. Sureshan, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.3598. (Type specimen missing). Additional material examined: ♀, India: Arunachal Pradesh, West Kameng district, Bhalukpung (27��01���12.5���N & 92��38���25.9���E, 213 m), 28.iv.2001, Coll. S.N. Ghosh. (NZC) 10072/H3 (Head missing). Diagnosis. Male. Body length 7 mm. Frons completely yellow with coronal teeth reddish trans-striate, tip of mandibles black; gena yellow; vertex with two trans-carinae turning rugose posteriorly and irregularly punctate; occipital carina indistinct; pronotum with pale brownish yellow near posterior margin, deeply reticulate; axillae and scutellum strongly reticulate; metapleuron deeply reticulate; hind femora ventrally with two tooth; propodeum anteriorly half reticulate, posteriorly with few scattered pits and carinae; first metasomal tergite weakly and longitudinally reticulate, longer than remaining segments, the latter with yellow patches. Female. Body length 13.65 mm, ovipositor sheath length 11.5 mm. Frons transversely striate with oblique striae in between coronal tooth; vertex with three carinae followed by fine transverse striae; neck transversely striae; pronotum rugose-reticulate; mesoscutum finely shagreened posteriorly with scattered pits and shagreened interstices; scutellum with smooth reticulations; propodeum with large close pits and shagreened interstices; hind coxa strongly striate; femora with two tooth and few denticles ventrally; first metasomal tergite shorter than remaining segments; ovipositor sheath with a subapical band. Distribution. Oriental: India (Kerala, Arunachal Pradesh). Host. Unknown. Note. The female specimen of F. chinnarensis designated from Arunachal Pradesh, India pose some confusion in the identity of the specimen. The female species designated shows quite distinct variations in the number of carinae present on the vertex, colouration on the pronotum, metasoma and the sculpturing on the metasoma with the designated holotype (male specimen) from Kerala, India. This is attributed to the possible sexual dimorphism within the specimens (Sheela & Ghosh, 2009). However, authors here feel the species extend beyond the limits of variation and the species to be a novel one. But, with the designated female specimen broken at the crucial part (head missing), it is quite impossible now to designate the same within the limits of variation or as a novel species. Hopefully future sampling of the species in India will allow a broader assessment of the consistency of these variations and novelty of the reported species. The holotype was untraceable, therefore, the above mentioned diagnosis of male is obtained from the original description., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 12-13, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Sheela, S. & Ghosh, S. N. (2009) First record and description of the female of Diastephanus chinnarensis Sureshan from India. Journal of Threatened Taxa, 1 (3), 182 - 183. https: // doi. org / 10.11609 / JoTT. o 1784.182 - 3"]}

Foenatopus weii Chen, van Achterberg & Xu, 2016 (Figs 44���54) Foenatopus weii Chen, van Achterberg & Xu, 2016a, Zootaxa, 612, 114���118. Material Examined: India: Tamil Nadu, Kanyakumari district, Kanyakumari Wildlife Sanctuary, Alagiapandiapuram (8��18���27.7���N & 77��26���19.1���E, 30m), 16.ii.2020, Coll. Souvik Sen & party, (ZSIK). Reg. No. ZSI/ WGRC / IR/INV.14032. Diagnosis. Body length 10.52 mm, ovipositor sheath length 8.41 mm. Frons finely and transversely carinaterugose, medially black, laterally with small yellow patches along the inner eye margin; coronal area with some longitudinal carinae (Fig. 48); vertex finely and transversely striate with two carinae (Fig. 47); temple non-angulate, shiny and smooth (Fig. 46). pronotum mostly coriaceous, anteriorly with transverse rugulae; mesoscutum anteriorly transverse coriaceous, posterior half with large foveae; median groove distinct; scutellum and axillae coriaceous; propodeum with medium-sized, circular foveae and with wide coriaceous interspaces, inside of fovea smooth (Fig. 49); propleuron and mesopleuron coriaceous (Fig. 50); vein 2-CU1 of fore wing absent; pterostigma parallel sided and elongate, apically acute (Fig. 51); hind coxa transversely striate with basal third rugose; hind femur microreticulate, ventrally with two large black teeth and a tubercle basally (Fig. 52); hind tibia coriaceous; first metasomal tergite transversely striate (Fig. 53), 1.1�� as long as remainder of metasoma; ovipositor sheath long, 1.3�� as long as metasoma, with a subapical ivory band (Fig. 54). Male. Unknown. Distribution. Oriental: China (Yunnan) (Chen et al. 2016); India (Tamil Nadu) (present record). Host. Unknown, collected from tropical semi evergreen forest. Note. The species is previously reported from Yunnan, China (Chen et al. 2016). This forms the second report of the species and a new record for India., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on page 27, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Chen, H. - Y., van Achterberg & Xu, Z. - F. (2016) Description of a new species of Foenatopus Smith from China and the male of Parastephanus brevicoxalis (Hymenoptera, Stephanidae). ZooKeys, 612, 113 - 123. https: // doi. org / 10.3897 / zookeys. 612.9781"]}

Foenatopus jodhpurensis Narendran, 2001 (Fig. 24) Foenatopus jodhpurensis Narendran, 2001, J. Adv. Zool., 22(2), 83���85. Type material. Holotype ♀, India: Rajasthan, Jodhpur (26��24��� N & 73�� 1��� E, 231 m), 10.ix.1987, Coll.?, (NHM) B. M. TYPE HYM. 3a. 374 (NHMUK013458830). Diagnosis. Body length 13.1 mm, ovipositor sheath length 9.6 mm. Frons with irregular carinae and pits; vertex with two strong carinae followed by close pits, median longitudinal grove absent; neck anteriorly deeply emarginate; pronotum trans carinate anteriorly turning finely reticulate posteriorly with a medial grove; mesoscutum faintly reticulate anteriorly, followed by deeply pitted area with carinate interstices and a longitudinal grove; scutellum finely alutaceous with sparse and scattered deep pits; metapleuron carinate with deep pits; propodeum with close pits and reticulate area anteriorly, remaining portion with carinate interstices; hind coxa annulate turning to weakly carinate medially, interstices microsculptured; hind femora bidentate with few denticles on the ventral side; first metasomal tergite uniformly annulate carinate, basally irregularly carinate, distal end weakly granulate, shorter than combined length of remaining segments; ovipositor sheath longer than metasoma, without a subapical pale ivory patch. Male. Unknown. Distribution. Oriental: India (Rajasthan). Plant Associate. Prosopsis cineraria (Fabales: Fabaceae)., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 16-17, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367

Foenatopus idukkiensis Sureshan and Narendran, 2013 (Fig. 23) Foenatopus idukkiensis Sureshan and Narendran, 2013, J. Threat. Taxa., 5(9), 4387���4389. Type material. Holotype ♀, mounted on triangular card, India: Kerala, Idukki district, Chinnar Wildlife Sanctuary, Kuttar (10��30��� N & 77��20��� E, 450 m), 04.iv.2012, Coll. P.M. Sureshan, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.2346. Ex. Dead wood infested with beetles. Paratypes 2♂ ZSI/ WGRC /IR/INV/2348(i���ii). Examined. Additional material examined: 1♀, India: West Bengal, N-24 Parganas district, Salt Lake (22��35���04.59���N & 88��25���02.08���E, Alt 8 m), 3.xii.2011, Coll. O. Biswas, (ZSIK) Regd. No. ZSI/ WGRC /IR/INV.13602. Diagnosis. Body length 11.0 mm, ovipositor sheath length 10.0 mm. Frons transversely striate to carinate; vertex with four complete and strong carinae followed by distinct transverse striae; ocellar area with longitudinal carinae; neck anteriorly deeply emarginate; pronotum trans carinate anteriorly turning finely reticulate posteriorly; mesoscutum anteriorly finely and transversely reticulate and with broad transverse band of pits posteriorly; scutellum finely alutaceous with sparse and scattered pits; metapleura carinate with deep pits; propodeum with broad shallow pits, interstices reticulate; hind coxa strongly annulate; hind femora bidentate with few denticles on the ventral side; first metasomal tergite uniformly annulate carinate, longer than combined length of remaining segments; ovipositor sheath longer than metasoma, with a subapical pale ivory ring. Male. Body length 9.0 mm. Antenna with fewer antennomeres than in females (30:26); otherwise resembles female. Distribution. Oriental: India (Kerala, West Bengal (present record)). Host. Unknown, presumably on wood boring beetles (reared from dead wood with unknown beetle infestation)., Published as part of Binoy, C., Achterberg, C. Van, Kumar, P. Girish, Santhosh, S. & Sheela, S., 2020, A review of Stephanidae (Hymenoptera: Stephanoidea) from India, with the description of five new species, pp. 1-51 in Zootaxa 4838 (1) on pages 14-15, DOI: 10.11646/zootaxa.4838.1.1, http://zenodo.org/record/4403367, {"references":["Sureshan, P. M., Narendran, T. C. & Nikhil, K. (2013) Parasitoids (Hymenoptera) of xylophagous beetles (Coleoptera) attacking dead wood in southern Western Ghats, Kerala, India, with descriptions of two new species. Journal of Threatened Taxa, 5 (9), 4385 - 4391. https: // doi. org / 10.11609 / JoTT. o 3385.4385 - 91"]}

Chelonus (Microchelonus) ensifer GRANGER 1949 Chelonus ensifer n.sp.: GRANGER 1949: 193 (key, descr. ♀, Madagascar), 196 (figs 197, 198). Microchelonus ensifer (GRANGER 1949): SHENEFELT 1973: 884 (tax., cat.). D i s t r i b u t i o n: Madagascar: Prov. Toliara: Bekily. Endemic., Published as part of Madl, M. & Achterberg, C. Van, 2014, A catalogue of the Braconidae (Hymenoptera: Ichneumonoidea) of the Malagasy subregion, pp. 5-220 in Linzer biologische Beiträge 46 (1) on page 73, DOI: 10.5281/zenodo.4507364, {"references":["GRANGER C. (1949): Braconides de Madagascar. Memoires de l'Institute Scientifique de Madagascar, Serie A, 2: 428 pp.","SHENEFELT R. D. (1973): Braconidae 5. Microgasterinae & Ichneutinae. In: VECHT J. VAN DER & R. D. SHENEFELT (eds), Hymenopterorum Catalogus (nova editio) Pars 9: 669 - 812."]}

Bracon (Bracon) atrolugens GRANGER 1949 Bracon atrolugens n.sp.: GRANGER 1949: 58 (key), 80 (fig. 105), 82 (fig. 111), 83 (descr. ♂ ♀, Madagascar). Bracon atrolugens GRANGER 1949: SHENEFELT 1978: 1467 (cat.). D i s t r i b u t i o n Madagascar: Prov. Antananarivo: near Antananarivo – Prov. Toamasina: Amparafaravola. Endemic., Published as part of Madl, M. & Achterberg, C. Van, 2014, A catalogue of the Braconidae (Hymenoptera: Ichneumonoidea) of the Malagasy subregion, pp. 5-220 in Linzer biologische Beiträge 46 (1) on page 40, DOI: 10.5281/zenodo.4507364, {"references":["GRANGER C. (1949): Braconides de Madagascar. Memoires de l'Institute Scientifique de Madagascar, Serie A, 2: 428 pp."]}

Lytopilus lucidus (GRANGER 1949) new combination Aerophilus lucidus n.sp.: GRANGER 1949: 307 (key, descr. ♀, Madagascar). Aerophilus lucidus GRANGER 1949: SHENEFELT 1970: 309 (cat.). Bassus lucidus (GRANGER 1949): MADL 2007: 995 (tax., cat.), 1006 (tab. 1: distr.). D i s t r i b u t i o n: Madagascar: Prov. Toamasina: Andekaleka (Rogez). Endemic., Published as part of Madl, M. & Achterberg, C. Van, 2014, A catalogue of the Braconidae (Hymenoptera: Ichneumonoidea) of the Malagasy subregion, pp. 5-220 in Linzer biologische Beiträge 46 (1) on page 15, DOI: 10.5281/zenodo.4507364, {"references":["GRANGER C. (1949): Braconides de Madagascar. Memoires de l'Institute Scientifique de Madagascar, Serie A, 2: 428 pp."]}