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Bicellaria ingrata Collin, 1960 Material examined. Muğla prov.: 40♂, 18♀, Muğla University campus, edge of pine wood + Quercus shrubs, 710 m, MT, 37��09���39������N, 28��22���20������E, Bart��k, Kub��k, xi.2012 ��� iii.2013; 2♂, 1♀, same locality, 730 m, MT, 37��09���38������N, 28��22���11������E, Bart��k, Kub��k, xi.2015 ��� iv.2016; 3♀, same locality, MT, 720 m, 37��09���42������N, 28��22���13������ E, H. Kavak, i.2015; 1♀, same locality, PT, 645 m, 37��09���42������N, 28��22���21������ E, O. Dursun, 8.vi.2012 ���all CULSP. Remarks. The species has been known up to now only from Israel and Lebanon. Very few specimens were found with four setae on the palpus, which may cause difficulties using the key by Bart��k et al. (2013). These are the first records for Turkey., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2018, Hybotidae (Diptera) from Turkey, with descriptions of seven new species, pp. 453-482 in Zootaxa 4410 (3) on page 477, DOI: 10.11646/zootaxa.4410.3.2, http://zenodo.org/record/1221772, {"references":["Collin, J. E. (1960) Some Empididae from Palestine. Annals and Magazine of Natural History, 13, 385 - 420.","Bartak, M., Plant, A. & Kubik, S. (2013) Species of Bicellaria (Diptera: Hybotidae) from Asia. Zootaxa, 3710, 233 - 256."]}

Bicellaria halterata Collin, 1961 (Figs 11���12) Bicellaria halterata Collin, 1961: 256. Type material examined. SYNTYPES: Great Britain: 23, Tomintoul, Banff, 8���11, August 1937 (OUM); 53, 1��, Banffshire, Tomintoul, 8���11.viii. 1937, 1100 ft, R.L. & B.M. Coe; 13, Aberdeenshire, Braemar, 1.viii. 1937, R.L. Coe���(BMNH). Additional material examined (133, 15��). Czech Republic: 13, ��umava Mts., Star�� Brunst, along brook, 49 �� 10 'N, 13 �� 16 'E, 960 m, 21.viii. 1989; 1 ��, ��umava Mts, S slopes of Panc��ř, damp meadow, 49 ��09'N, 13 �� 16 'E, 1000 m, 24.viii. 1989 ���all M. Bart��k; 13, 1��, Krkono��e Mts., Slunečn�� str��ň, nr. pond, 50 �� 38 ' 12.5 ''N, 14 �� 49 ' 23.6 ''E, 645 m, Malaise trap, 1.ix.- 13.x. 2009; 73, 6��, same locality, 4.viii.��� 1.x. 2009; 23, same locality, 30.vi.��� 4.viii. 2009; 13, Krkono��e Mts, Zrcadlov�� potok, nr. brook, 50 �� 38 ' 1.9 ''N, 14 �� 43 ' 54.6 ''E, Malaise trap, 18.viii.��� 1.ix. 2009; 1 ��, Krkono��e Mts, Pekelsk�� potok, nr. brook, 50 �� 38 ' 13.3 ''N, 14 �� 40 ' 29.3 ''E, Malaise trap, 30.vii.- 18.viii. 2009 ���all J. Van��k; 13, Orlick�� hory Mts, Popelov, valley of Zdobnice river, 14.ix. 1995, B. Mocek; 1 ��, Podyj�� NP, Braitava, 48 �� 52 ' 35 ''N, 15 �� 50 '03''E, 26.vi.��� 30.vii. 2002, Malaise trap; 1 ��, same locality, 30.vii.��� 31.viii. 2002 ���all M. Bart��k et ��. Kub��k���(CULSP). Russia: 2 ��, Moscow, Losiny ostrov, park vegetation, 55 �� 19 'N, 37 �� 45 'E, 4.ix. 1988, M. Bart��k (CULSP); 1 ��, Permskaja oblasť Kamenka, env. of Lysva, 31.vii. 1963, V. Zherikhin (ZMMU). Slovak Republic: 1 ��, Ko��eck�� Podhradie, mixed wood, 48 �� 59 'N, 18 �� 17 'E, 270 m, 29.vii. 1989, M. Bart��k (CULSP). Diagnosis. The species is a member of B. nigra complex of species as proposed by Chv��la (1991) characterized by swollen hind basitarsus, short posteroventral setae on hind tibia, and narrow face and comprising three species in Europe (B. nigra, B. nigrita, B. halterata). Males may be recognised easily according to the structure of the genitalia, possessing both phallic hooks, the left one with lateral triangular-shaped process best visible from below [not apparent in Collin (1961, fig. 89, p. 256)] and relatively short postgonites (Fig. 11). It is possible to distinguish B. nigra from B. halterata without dissection of the genitalia according to the shape of the hypandrial processes which are parallel in B. halterata but divergent towards apex in B. nigra (but this character should always be verified by dissection). Females sometimes have only slightly swollen hind basitarsi, so they may erroneously lead in the key to the R. sulcata complex, but it may be distinguished by yellow halteres, pale setose base of abdomen and swollen hind tibiae (opposite of B. simplicipes which is not a member of this complex). Distribution. Great Britain, Czech and Slovak Republics, European Russia. Probably more broadly distributed in temperate Europe but mixed in collections with B. nigra or B. intermedia. Remarks. The species was described (including illustrations of genitalia) from an unknown number of males and a single female taken at Tomintoul and Braemar (G. Britain) early in August 1937 by R.L. Coe. Since the original description no other record of this species has been published., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 258-259, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria austriaca Tuomikoski (Figs 7���8) Bicellaria austriaca Tuomikoski, 1955: 70. Type material examined. HOLOTYPE 3: Austria, Obertillach, R. Frey (labelled: ���Mus. Zool. H:fors, Spec. typ. No. 8202 Bicellaria austriaca Tuomikoski ��� and ��� Holotypus Bicellaria austriaca Tumikoski, 1955, Chv��la 1980 ���). PARATYPES: 13, 2��, Obertillach, R. Frey (male labelled ���Mus. Zool. H:fors, Spec. typ. No. 8203 Bicellaria austriaca Tuomikoski ���)���(NHMH). Additional material examined. (453, 7��) Austria: 13, Niedere Tauern, S��lker Pass, 47 �� 16 'N, 14 ��04'E, 1900 m, 7���8.vii. 1995, M. Bart��k; 1 3, Tauernalm, pasture, 15.vii. 1998, M. Bart��k; 23, 1��, Gerlos, spruce wood, 1600 m, 47 �� 16 'N, 12 ��03'E, 9.viii. 1988, M. Bart��k (CULSP); 13, Tirol, Stafflach, 19���20.vi. 1933, A.C.W. Wagner (NHMH). Bulgaria: 23, Pirin Mts, Begovica, 17.1 km NW from Sandanski, 41 �� 40 '09''N, 23 �� 25 ' 33 ''E, 1760 m, 10.viii. 2005, J. Fechtner; 13, Pirin Mts, 5 km SE Vikhren, peak, 2100���2300 m, 1.vii. 1988, B. Mocek; 13, Pirin Mts, Banderica, 7.vii. 1982, 1800 m, F. Kocourek (CULSP). Czech Republic: 13, ��umava Mts, Ant��gl, 2.viii. 1995, ��umava Mts, Popeln��, along brook, 49 ��06'N, 13 �� 38 'E, 880 m, 7.vii. 1988, M. Bart��k; 13, ��umava Mts, S. slope of Panc��ř, meadow, 49 ��09'N, 13 �� 16 'E, 1000 m, 24.viii. 1989, M. Bart��k; 13, ��umava Mts, Nov�� hůrka, peat-bog, 49 ��09'N, 13 �� 20 'E, 850 m, 1.vii. 1995, M. Bart��k; 13, same locality, 15.vi. 1999; 23, 1��, ��umava Mts, Tetřev�� slať, peat-bog, 49 ��01'N, 13 �� 20 'E, 23.vii. 1989, M. Bart��k; 13, ��umava Mts, Rakousk�� louka, peat-bog, 48 �� 46 'N, 13 �� 50 'E, 1350 m, 16.viii. 1994, M. Bart��k; 13, ��umava Mts, Pr����ily nr. Křemeln�� brook, 49 ��08'N, 13 �� 23 'E, 810 m, 8.vii. 1994, M. Bart��k; 13, ��umava Mts, Zhůřsk�� slat��, peat-bog, 1130 m, 49 ��04'N, 13 �� 34 'E, 17.vi. 1999, M. Bart��k et ��. Kub��k; 13, ��umava Mts, Boub��n, mixed wood, 48 �� 58 ' 40 ''N, 13 �� 49 '00''E, 990 m, 2��� 17.vii. 2002, K. Spitzer; 13, B��lina, ��t��p��nov, nr. brook, 50 �� 33 'N, 13 �� 50 'E, 370 m, 18.vi. 1994; 13, B��lina, Chloumek, damp meadow nr. woods, 50 �� 33 'N, 13 �� 50 'E, 410 m, 18.vi. 1994; 13, Janov nad Nisou, 50 �� 46 'N, 15 �� 12 'E, 700���800 m, 19.vi. 1982; 13, ��tiř��n, near pond, 49 �� 55 'N, 14 �� 37 'E, 380 m, 11.vii. 1991 ���all M. Bart��k; 13, 2��, Krkono��e Mts, Lučn�� bouda, 50 �� 44 ' 19 ''N, 15 �� 40 ' 38 ''E, 1250 m, 9���16.viii. 2007; 13, same locality, 16��� 26.vii. 2007; 13, same locality, 16���25.v. 2007; 13, Krkono��e Mts, ��pa, peat-bog, 50 �� 44 ' 14 ''N, 15 �� 42 ' 55 ''E, 1407 m, Malaise trap, 28.vi.- 12.vii. 2007; 13, Krkono��e Mts, Labsk�� rokle, nr. brook, 50 �� 46 ' 19 ''N, 15 �� 32 ' 43 ''E, 1300 m, 25.vii.- 15.viii. 2007; 13, 1��, Krkono��e Mts, Li��č�� hora, upper forest line, 50 �� 42 '04''N, 15 �� 40 ' 35 ''E, 1320 m, 29.vi.- 26.vii. 2005; 13, Krkono��e Mts, Dvorsk�� potok, nr. brook, 50 �� 45 ' 54 ''N, 15 �� 34 ' 41 ''E, 18���29.viii. 2006, Malaise trap��� all J. Van��k; 13, Podyj�� NP, Braitava, mixed wood, 48 �� 52 'N, 15 �� 49 'E, 27.vi. 2002; 13, 1��, Beskydy Mts, Muřinkov�� vrch, peat-bog, 49 �� 31 'N, 18 �� 39 'E, 950 m, 23.viii. 1987; 13, Beskydy Mts, Lys�� hora, hilltop, 49 �� 32 ' 46 ''N, 18 �� 26 ' 53 ''E, 1323 m, 7���29.vii. 2004, Malaise trap; 13, 2��, Jesen��k Mts, Velk�� Kotlina, 25���26.vi. 1982; Horn�� Bečva, 26.vi. 1975 ���all M. Bart��k���(CULSP). Germany: 13, Bayerischer Wald, Spiegelau, 48 �� 57 'N, 13 �� 22 'E, 760 m; 13, Bayerischer Wald, Hochgericht, Stangenfiltz, 1185 m, 6.vii. 1992, Weiss���(CULSP). Italy: 43, S. Stefano di Cadore, fiuma Piave, viii. 1979, G. Raffone; 13, Lago Calaita, viii. 1970; 13, Passo Monte, Croce Carnico, 23.viii. 1964 (MSNV). Switzerland: 13, GR, Lenzerheide, pasture, 2000 m, Malaise trap, 14��� 21.vii. 2000, B. Merz (CULSP). Diagnosis. Bicellaria austriaca is a member of B. austriaca complex (incl. B. italica sp. nov., B. collini Tuomikoski, and B. intermedia Lundbeck) characterized by 1���2 long dorsal setae on the third antennal segment, 2��� 3 setae on palpus and both phallic hooks present and long. All four species of this complex may be easily identified in the male sex according to the key and genitalia. Bicellaria austriaca has two rather strong unequally long and unequally bent phallic hooks and relatively broad hypandrial processes with long setae on the apical part. We believe that females may be also distinguished from other species of the B. austriaca complex according to characters given in the key despite finding no specimens in copula. To be certain faunistic records based solely on females are omitted here. Distribution. Europe, from Italy and Bulgaria to about 60 �� N latitude in Scandinavia. Remarks. The species was described (and hypandrium illustrated) on the basis of many specimens from the mountains of Central and Southern Europe. Chv��la (1983) redescribed this species and provided illustrations of the face, antenna, hind leg, genitalia, and identification of the holotype., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 255-256, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria globulicauda sp. nov. (Figs 9, 10, 11) Type material. HOLOTYPE ♂, Uzbekistan, Amankutan, pasture, 1600 m, 39.17 N, 66.54 E, 23.v. 1989, M. Bart��k (CULSP). PARATYPES: 4 ♂, 3 ♀, same data as holotype (CULSP). Diagnosis. Thoracic setae and most abdominal setae black, base of abdomen on sides with pale setae; third antennal segment without elongate dorsal setae; 2 setae on palpus; front tibia with posteroventral setae longer than ciliation (which is very short or absent); hind basitarsus not swollen and hind tibia very slightly swollen in both sexes. Male hypandrium long with apically bifurcate processes, postgonites with ventral projections; left phallic hook long, right one slightly smaller but equally shaped. Female visible parts of tergites 5���7 and apical half of 8 th tergite microtrichose, 8 th sternite polished. Etymology. The species is named after peculiarly enlarged epandrium. Description. Male. Head black, light grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 12���13 facets in line of contiguity of eyes. One pair of rather long ocellar setae. Occiput grey, sparsely setose. Face slightly narrowed ventrally, ventral part about 0.03 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest setae about 0.07 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 13���14: 4���7: 18���21. Thorax brownish-black, grey microtrichose, mesoscutum brownish grey microtrichose in dorsal view, prescutellar area differently microtrichose than other parts of mesoscutum, appearing light grey. All thoracic setae black. Chaetotaxy: 6���8 rather narrowly biserial long acrostichals (about 0.13 mm long in front of suture); 6���9 uniserial dorsocentrals as long as or slightly longer than acrostichals; some 5���10 setae in intrahumeral and posthumeral areas; 2���4 postpronotals; notopleuron with 3���5 setae arranged in irregular row; 1 supraalar- and 1 prealar; 1 long postalar; 2 pairs of scutellars. Legs including coxae brownish-black, microtrichose and black setose, hind coxa in some specimens with several pale setae. Fore femur with sparse rows of anteroventrals and posteroventrals half as long as depth of femur. Fore tibia with posterodorsals as long as depth of tibia, longer setae scarcely discernable, posteroventrals as long as depth of tibia or longer in basal third but shorter in apical part, pilosity absent or very short. Mid femur with sparse row of anteroventrals more than one third as long as depth of femur (even in middle of row) and with slightly longer posteroventrals slightly shorter than depth of femur. Mid tibia with 1���4 setae dorsally in basal half up to 2 X as long as depth of tibia, otherwise short setose. Hind femur with 13���17 anterodorsal setae on basal third slightly longer than depth of femur, shorter apically, with row of 13���15 subequally long anteroventrals; posteroventrals short, scarcely one-third as long as depth of femur. Hind tibia very slightly swollen in apical part (0.09���0.10 mm at broadest point in comparison with 0.06���0.07 mm in basal part), several anterodorsals and posterodorsals up to 0.20 mm long, ventral setae short. Tarsi of all legs short, thin, short setose, basitarsus of hind leg not swollen (0.05 mm). Wing slightly yellowish infuscated; stigma light brownish yellow, 0.55���0.60 mm long, slightly narrowing apically and narrow (0.04 mm in middle), distinctly narrower than cell r 1 (about 0.09 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.30���0.35 mm. Squama yellowish with dark fringes, halter clear yellow. Abdomen brownish-black, almost velvety brown in dorsal view and pale grey in lateral view, microtrichose, lateral parts of basal sternites and tergites apparently polished (however, difficult to observe because all male specimens have abdomen distorted). Setae on abdomen pale, dorsally and laterally also with black setae, posteromarginals on sides of tergites 2���4 slightly longer than corresponding segments, slightly shorter on subsequent tergites. Genitalia (Figs 9, 10, 11): hypandrium long with apically bifurcate processes, inner arm with short setae; postgonites short, with ventral projections; left phallic hook long, right one slightly smaller but similarly shaped (appearing as single structure in lateral view). Female. Body with setae somewhat less numerous than in male. Visible parts of tergites 5���7 and apical half of 8 th tergite microtrichose, 8 th sternite polished. Length: body 2.8���3.6 mm, wing 2.4 ���3.0 mm. Remarks. Bicellaria globulicauda sp. nov. is somewhat similar to B. amankutanensis sp. nov. and B. chimganensis sp. nov. according to combination of the following characters: palpus with 2 setae, 3 rd antennal segment without dorsal seta, hind basitarsus not swollen, fore tibia with rather long posteroventrals and without elongate ventral pilosity, and abdomen at least partly pale setose, but it may be easily differentiated on account of the very peculiar form of its male genitalia, as described in the Diagnosis., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 240-241, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria chimganensis sp. nov. (Figs 3, 4) Type material. HOLOTYPE ♂, Uzbekistan, Chimgan, damp valley, 1200 m, 41.38 N, 70.02 E, 18.v. 1989, M. Bart��k (CULSP). PARATYPES: Uzbekistan: 12 ♂, 42 ♀, same data as holotype; 1 ♂, 1 ♀, same locality as holotype, except 1800 m, 41.38 N, 70.06 E; 8 ♂, 4 ♀, Aktash, deciduous wood, 500 m, 41.38 N, 69.46 E, 17.v. 1989; 1 ♀, Karamazar, damp meadow, 500 m, 41.32 N, 69.48 E, 18.v. 1989; 1 ♀, same locality, 800 m, 41.30 N, 69.49 E��� M. Bart��k���(CULSP). Diagnosis. Nearly all setae including acrostichals, dorsocentrals and those on legs pale (white to brownish yellow); halter yellow in both sexes; third antennal segment without elongate dorsal setae; hind basitarsus narrow and hind tibia swollen in both sexes. Male hypandrial processes narrow; only one (left) long phallic hook present; female tergites 5���8 and sternites 6���8 polished. Etymology. The species is named after type locality (Chimgan Mountain in Uzbekistan). Description. Male. Head brownish-black, rather light grey microtrichose, pale setose. Holoptic, facets in dorsal half much larger than in ventral half, about 11���13 facets in line of contiguity of eyes. One pair of rather long ocellar setae. Face slightly narrowed ventrally, ventral part about 0.03���0.04 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 3 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest setae about 0.08 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 12���16: 5���7: 16���20. Thorax brownish-black, rather sparsely light grey microtrichose, mesoscutum almost subshining in dorsal view, prescutellar area differently microtrichose than other parts of mesoscutum, appearing light grey. All thoracic setae pale. Chaetotaxy: 6���9 broadly biserial acrostichals; 7���13 uniserial dorsocentrals (both acrostichals and dorsocentrals about 0.10 mm long before suture); about 6���9 setae in intrahumeral and posthumeral areas; 1���2 long and 1���2 shorter postpronotals; notopleuron with 3���6 setae arranged in one irregular row; 1 supraalar and 1 prealar; 1 long postalar; 2 pairs of scutellars. Legs including coxae blackish brown, microtrichose and pale setose, tarsi with darker setae. Fore femur short setose, both anteroventrals and posteroventrals much shorter than depth of femur (except preapicals). Fore tibia with 4���5 posterodorsals up to 2 X longer than depth of tibia, otherwise short setose, posteroventrals in apical third about as long as depth of tibia, pilosity apparently absent (similar to B. spuria). Mid femur with sparse row of anteroventrals one-third as long as depth of femur and with somewhat longer posteroventrals slightly shorter than depth of femur. Mid tibia with 1���2 pairs of setae dorsally in basal half up to twice as long as depth of tibia, otherwise short setose. Hind femur with 15���20 anterodorsal setae on basal third up to 2 X as long as depth of femur, much shorter apically and with row of 16���22 anteroventrals slightly longer than depth of femur at about middle and shorter both basally and apically, posteroventrals half as long as depth of femur. Hind tibia swollen in apical part (0.14���0.16 mm at broadest point in comparison with 0.06 mm in basal part), several anterodorsals and posterodorsals up to 0.20 mm long, ventral setae short. Tarsi of all legs thin, short setose, basitarsus of hind leg very slightly swollen (0.09 mm). Wing slightly yellow-brownish infuscated; stigma brown, long (0.70 mm), almost parallel-sided and narrow (0.05 mm), narrower than cell r 1 (about 0.09 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.30 mm. Squama yellowish with pale fringes, halter clear yellow. Abdomen blackish-brown in ground-colour, brown in dorsal view and pale grey in lateral view, microtrichose, pale setose, dorsally with short setae, posteromarginals on sides of tergites as long as or slightly longer than corresponding segments. Genitalia (Figs 3, 4) with hypandrial processes medium long and narrow, with conspicuous but not very long spine at tip; postgonites elongate-trapezoid, slightly asymmetrical (right one slightly longer and broader at apex); left phallic hook long, right one strongly reduced. Female mesoscutal setae less numerous and hind tibia narrower (0.11 mm) than in male. Tergites 5���8 polished, sternites 5���8 polished (5 th sternite with narrow sparsely microtrichose central stripe), other parts of abdomen microtrichose. Length: body 2.2 ���3.0 mm, wing 2.3 ���3.0 mm. Remarks. Bicellaria chimganensis sp. nov. can be easily recognized from all other Palaearctic species of Bicellaria by its mostly pale setose body (including all mesoscutal setae) and narrow hind basitarsus. Moreover, females have the last four sternites polished. Bicellaria amankutanensis sp. nov. is undoubtedly closely allied with this species. Beside the main diagnostic character given in the key (different colouring of mesoscutal setae), the species described above has slightly smaller broad part of third antennal segment, less setae on the intrahumeral and posthumeral areas and more sparsely microtrichose mesoscutum giving it a subshining appearance. The genitalia of both species are very similar, B. chimganensis sp. nov. has slightly narrower hypandrial processes. Female 5 th sternite is polished in B. chimganensis sp. nov. with the exception of sparsely microtrichose central stripe but is densely microtrichose (with the exception of narrow polished lateral parts) in B. amankutanensis sp. nov., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 236-237, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria femorata Collin (Figs 7, 8) Bicellaria femorata Collin, 1960: 392. Identification of syntypes. Bicellaria femorata was described from five specimens: ���a male labelled ���Jerusalem [=Yerushalayim], O. Theodor���, and a female ���Ginegar, Gallilee, 10.xi. [sic!] 34. O. Theodor���. Also two females from ���B. Alta [= Bet Alfa], 16.i. 40, Dr. A. Shulov���, and one from ���Judean Hills, 1.ii. 31. O. Theodore��� [sic!]���. All the above specimens are present in TAU: 1 male labelled ���Jerusalem Palestine [=Yerushalayim] O. Theodor / COM. INST. ENT. COLL. No. 12611 / ♂ TYPE / B. femorata det. Coll. 58 ���, 1 female labelled ���G 10 1 / Ginegar Galilee Palestine O. Theodor 10.ii. 34 / COM. INST. ENT. COLL. No. 12611 / B. femorata det. Coll. 58 ���, 1 female labelled ���B. Alfa 16.i. 40 / 135 / Palestine Dr. A. Shulov / B. femorata det. Coll. 56 / Bicellaria femorata n. sp. ���, 1 female labelled ���B. Alfa [=Bet Alfa] 16.i. 40 / 133 / Palestine Dr. A. Shulov / B. femorata det. Coll. 56 ���, and one female labelled ���BK VIII 34 / Kir. Anavim [=Qiryat 'Anavim] Judean Hills Palestine O. Theodor 1.2. 31 / COM. INST. ENT. COLL. No. 12611 / ♀ TYPE / B. femorata det. Coll. 58 ���. We consider all above mentioned specimens syntypes in spite of small difference in date of the female from Ginegar and they were provided by red labels ��� Syntype Bicellaria femorata rev. Bart��k 2013 ���. Moreover, in TAU there are another two specimens collected before 1960: 1 female without locality labelled ��� 11.i. 22 / 5.i. 22 / 212 ��� and another female labelled ���C J VIII 30 / Jerusalem Palestine [=Yerushalayim] O. Theodor 5.i. 36 ��� and another 5 females labelled ���Jerusalem Palestine [=Yerushalayim] O. Theodor��� which were probably also collected before 1960. These are neither provided with circular handwritten Collin��s identification labels nor listed in the original description. For this reason we identified and labelled as syntypes of both Collin��s species (B. femorata and B. spuria ingrata) only specimens provided by the above mentioned circular label (see also under B. ingrata). Diagnosis. The species has unique combination of several features: palpus with many setae, halter yellow, mesoscutum with stripes, hind femur spinose, male phallic hooks curiously shaped (Fig. 8). It may be easily identified according to the key. Additional material examined (13 ♂, 34 ♀). Israel: 2 ♂, 9 ♀, 2 exx, Horvat Nappah, 10.xii. 1973, A. Freidberg���(TAU); 3 ♀, same data���(CULSP); 5 ♂, 8 ♀, same locality, 20.xii. 1973, A. Freidberg���(TAU); 2 ♂, same data���(CULSP); 1 ♂, 1 ♀, same locality, 1.i. 1974, A. Freidberg; 1 ♂, Upper part of Nahal Tirza, 20.ii. 1974, A. Freidberg; 1 ♂, Kesalon, 10.ii. 1973, M. Kaplan; 1 ♀, Shelomi, 8.i. 1975, F. Kaplan; 1 ♀, Rosh ha'Ayin, 31.xii. 1986, Y. Nussbaum���(TAU); 1 ♂, same data���(CULSP); 1 ♀, Nahal 'Ammud, 23.ii. 1983, I. Yarom; 1 ♀, 'Ofra, 29.i. 1987; 1 ♀, Nahal Rabba, 20.xii. 1981; 1 ♀, sine loc. (��� 212 ��� and ��� 11.i. 22 ��� and ��� 5.i. 22 ���); 6 ♀, Jerusalem, Palestine���(TAU); 1 ♀, Carmel, 14.ii. 1976, A. Freidberg���(CULSP). Distribution. Israel. Remarks. This species was described in full detail by Collin (1960). Bicellaria femorata is included in the B. austriaca -complex of species as defined by Chv��la (1991), but differs in having a pale halter, long setose hypandrium and swollen and spinose hind femur., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 238-239, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria vana Collin Bicellaria vana Collin, 1926: 190. Distribution and diagnosis. See Bart��k and Kub��k (2013), published record from Asia: Yakutsk (Tuomikoski 1955)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 252, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria longisetosa Chv��la (Figs 19���20) B. longisetosa Chv��la, 1991: 9. Material examined (73, 2��). Austria: 13, Radstadter Tauern, Obertauern, puddle Gruenwaldkopf, 2000 m, 26.viii. 1999, Chv��la (CULSP). Russia: 53, 2��, Amurskaja oblast, g. (= town) Zeja, 7.ix. 1981 (ZMMU and CULSP). Switzerland: 13, GR, Lenzerheide, pasture, 2000 m, Malaise trap, viii. 2000, Merz (CULSP). Diagnosis. The species may be easily identified according to the key. It differs from any other European species not only in external features (e.g., possessing curiously long lateral setae on the third antennal segment), but also in the structure of male genitalia (Figs 19���20): hypandrium with rather short and broad processes ending in a single long seta (not visible on Chv��la���s original illustration), two subequally long but differently shaped phallic hooks (right one branched at apex), median extension of postgonites not connecting cranially (ventral apodeme absent), and presence of a sclerotized phallus forming valve-like flat structure. Distribution. Mountains of Central and Southern Europe, Russia (Amur region). Remarks. This species was described from several specimens taken in high European mountains (2000���2380 m), viz Rila (Bulgaria), Belansk�� Tatry (Slovak Republic), and W��ltzer Tauern (Austria). The illustrations in the original description are not clear, including the hypandrium and fore tibia. Details of the phallus are illustrated here for the first time (Figs 19���20)., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 262-263, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Key to species of Bicellaria of Europe Remarks. Absolute measures given in the text (performed on dry mounted material) are only benchmarks: they may slightly differ in unusually small or large individuals. Only those setae on the palpus are counted which are longer than 0.05 mm (those occurring on the palpifer are not counted). Long setae on dorsal side of the third antennal segment may be broken or their presence/absence may be variable (we found a single specimen of B. subpilosa with long fine seta on the third antennal segment). Those setae on intrahumeral and posthumeral areas are counted which occur outside the single row of dorsocentrals in the presutural area. Prescutellar depression in some species contains differently long and more erect microtrichia giving this area lighter appearance in dorsal view, this area is mostly sharply delimited and this character state is referred in the text as follows: ���prescutellar depression lighter than other parts of mesoscutum in dorsal view���. Nearly all characters sometimes vary (including precise shape of male hypandrial processes) which makes identification extremely difficult. Females are sometimes indistinguishable (most notably: B. italica - B. intermedia), thus faunistic records should be based only on males and always be supported by dissection of the genitalia. Both B. albopilosa and B. spuria probably represent complexes of sibling species. For this reason, some recently published records should be considered carefully due to possible misidentifications (e.g. Raffone 2006, 2008; Jonassen 1984; Parvu 2003, 2005). 1 Antennal stylus together with slender portion of third segment half as long as broad basal portion of third segment (Fig. 47); acrostichals and dorsocentrals (except last two pairs of dorsocentrals) very short (less than 0.05 mm long); all tibiae without any setae obviously longer than setulae; male hypandrial processes long setose along the whole inner margin; very small species (wing length 1.9 mm).................................................... B. dispar Oldenberg - Antennal stylus together with slender portion of third segment at least as long as broad basal portion of third segment (as in Fig. 48); acrostichals and dorsocentrals longer than 0.05 mm; remaining characters disagree....................... 2 2 (1) Palpus with 4 or more black setae; third antennal segment with dorsal seta(e); all body setae black.................. 3 - Palpus with 1���3 black setae (rarely with an additional pale seta), if palpus with more than 4 setae (B. andorra sp. nov.), then sides of abdomen with yellowish-white setae; third antennal segment with or without dorsal seta(e)............. 6 3 (2) Mesoscutum with two pale (silvery) stripes between rows of acrostichals and dorsocentrals apparent from both dorsal and caudal views (in female covering most of mesoscutum); usually less than 9 setae outside rows of dorsocentrals at presutural area; abdomen light bluish grey in lateral view; male genitalia as in Figures 3���4 (B. pilosa complex-part).............................................................................................. B. alpina Bezzi - Mesoscutum uniformly dark velvety black without any stripes; usually 10 or more setae outside rows of dorsocentrals at presutural area; abdomen dark greyish to blackish brown in lateral view....................................... 4 4 (3) Third antennal segment usually with 4 or more dorsal setae, second segment with curiously long lateral setae pointing forwards; fore tibia with posteroventral setae shorter than diameter of tibia and with a row of 4���6 erect posterodorsal setae about 4 X longer than diameter of tibia (0.25 mm long); hind tibia broadly thickened on apical third in male and slender in female; male hypandrium with short and broad processes bearing long apical seta, two phallic hooks present, phallus strongly sclerotized, forming flat and long valve-like structure (Figs 19���20)................... B. longisetosa Chv��la - Third antennal segment usually with 1���3 dorsal setae, second segment with short lateral setae (but with long ventral setae); fore tibia differently setose; hind tibia more evenly dilated in both sexes; male hypandrial processes without long apical seta, three phallic hooks (left hook bifurcate), phallus membranous, not forming valve-like structure. (B. pilosa complexpart)............................................................................................ 5 5 (4) Face narrow (about 5���6 facets combined), almost parallel throughout (0.06���0.07 mm); about 10 setae on posthumeral and intahumeral areas; male hypandrial processes broadened near apex, left phallic hook bifurcate, its anterior arm bent, postgonites strongly asymmetrical, right one with ventral extension (Figs 29���30).................. B. setipalpus sp. nov. - Face broad (about 8���9 facets combined), in lower half even broader (0.11 mm) than in upper half (0.10 mm); usually more than 10 setae on posthumeral and intrahumeral areas; male hypandrial processes apically narrow; left phallic hook bifurcate, its anterior arm straight (Figs 27���28).............................................. B. pilosa Lundbeck 6 (2) Hind basitarsus (and usually also the hind tarsomeres 2) distinctly swollen (usually more than 2.3 X broader than hind tarsomere 3 and usually broader than 0.10 mm); additional characters: fore tibia with ventral pilosity on apical third at most slightly shorter than on fore basitarsus and posteroventral setae short, at most as long as pilosity and usually shorter than depth of tibia (as in Fig. 46), face very narrow (less than 3 facets combined, less than 0.03 mm at narrowest point), abdomen at least partly pale setose (exception: male B. nigrita), third antennal segment without dorsal seta(e), hypandrial processes never strikingly shortened (B. nigra complex)...................................................... 7 - Hind basitarsus slender (usually less than 2.0X broader than hind tarsomere 3 and usually narrower than 0.10 mm); additional characters in different combinations.............................................................. 9 7 (6) Third antennal segment arched dorsally; mid tibia usually without dorsal setae differentiated from short hairs; hind femur shorter than 1.2 mm; hind tibia dilated in basal third; male: postgonites shortened (about 0.10 mm long), both phallic hooks present, long and strong, longer than postgonites, right one slightly shorter and narrower than left one (Fig. 25), squamae mostly with dark fringes................................................................ B. nigrita Collin -. Third antennal segment nearly straight dorsally; mid tibia with 1���3 outstanding dorsal setae, about twice as long as short hairs on limb; hind femur longer than 1.3 mm; hind tibia equally slender in basal third; male: postgonites not reduced (slightly shortened in B. halterata), phallic hooks different, squamae mostly with pale fringes...................... 8 8 (7) All large setae on mesoscutum black; male: halter usually dark, postgonites long (more than twice as long as broad), left phallic hook long, without lateral process, right one strongly reduced (Fig. 23); female: broad portion of third antennal segment usually longer than 0.15 mm, lustrous portions of abdominal tergites 6���7 comprise about basal third of these tergites................................................................................... B. nigra (Meigen) - Large setae on mesoscutum mostly white to brownish-yellow (at least scutellars, last pair of dorsocentrals and notopleurals); male: halter yellow, postgonites shorter (about 1.5 X as long as broad), both phallic hooks long, left one with lateral process (best visible from below, Fig. 11); female: broad portion of third antennal segment usually shorter than 0.14 mm, lustrous portions of abdominal tergites 6���7 comprise more than basal half of these tergites......... B. halterata Collin 9 (6) Third antennal segment with one to several long dorsal setae; male with two phallic hooks (both exceeding postgonites) (B. austriaca complex)................................................................................ 10 - Third antennal segment without long dorsal setae; male with 1���2 phallic hooks................................ 16 10 (9) Male........................................................................................... 11 - Female......................................................................................... 14 11 (10) Prescutellar depression lighter than other parts of mesoscutum in dorsal view (lighter area reaches hind acrostichals); halter yellow; hypandrial processes swollen apically, bearing several rather long setae on apex, both phallic hooks almost equally long, right one more sharply bent (Figs 15���16).............................................. B. italica sp. nov. - Prescutellar depression usually evenly dark in dorsal view (rarely lighter in a narrow area not reaching halfway between scutellum and hind acrostichals); halter dark brown to yellowish brown; remaining characters disagree............. 12 12 (11) Face parallel-sided or slightly broadened ventrally (about 5 facets combined, 0.06 mm broad at narrowest point); hypan- drial processes short (0.09���0.10 mm) and broadly U-shaped (distance between their inner margins of processes about 0.04��� 0.05 mm), both phallic hooks equally long and equally bent (Figs 9���10)...................... B. collini Tuomikoski - Face narrowing ventrally (about 3 facets combined, 0.04 mm broad at narrowest point); hypandrial processes longer and narrowly U-shaped (distance between inner margins of processes about 0.02���0.03 mm), phallic hooks different...... 13 13 (12) Usually more than 9 setae in posthumeral and intrahumeral areas; hypandrial processes broader, with several rather long setae near apex, phallic hooks unequal, shorter and broader (Figs 7���8).................... B. austriaca Tuomikoski - Usually less than 7 setae in posthumeral and intrahumeral areas; hypandrial processes narrower, with at most a single longer seta on apex, other setae much shorter, long and narrow phallic hooks nearly equal in length (Figs 13���14)....................................................................................... B. intermedia Lundbeck 14 (10) Face parallel-sided or slightly broadening ventrally (about 5 facets combined, 0.06 mm broad at narrowest point); abdominal tergites 6���7 broadly lustrous on basal half or more.................................... B. collini Tuomikoski - Face narrowed ventrally (about 3 facets combined, 0.04 mm broad at narrowest point); abdominal tergites 6���7 nearly completely microtrichose.............................................................................. 15 15 (14) Halter brown; usually more than 9 setae in posthumeral and intrahumeral areas............. B. austriaca Tuomikoski - Halter yellow; usually less than 9 setae in posthumeral and intrahumeral areas................................................................................................. B. intermedia Lundbeck, B. italica sp. nov. 16 (9) Fore tibia with ventral pilosity on apical third at most slightly shorter than pilosity on fore basitarsus and/or posteroventral setae on apical third of tibia shorter than depth of tibia (as in Fig. 46, observe in exactly posterior view)............ 17 - Fore tibia with ventral pilosity on apical third at most one-third as long as on fore basitarsus and posteroventral setae on apical third of tibia at least twice as long as pilosity, and usually longer than depth of tibia (as in Fig. 45)............ 20 17 (16) Hind tibia slender throughout; lower occipital setae mostly pale; male: hypandrial processes short and broad, right phallic hook long, simply curved and exceeding postgonites, left one slightly shorter, triangular in shape (Figs 31���32).................................................................................... B. simplicipes (Zetterstedt) - Hind tibia distinctly swollen apically (less so in female); lower occipital setae black; male: hypandrial processes not conspicuously short and broad, right phallic hook strongly reduced or absent, left one long and simply curved (B. sulcata complex)........................................................................................... 18 18 (17) Abdomen with long white setae on sides; the anteromost acrostichals and dorsocentrals subequally long as those just before suture (first three about 0.15 mm long or even longer in males); hind femur without posteroventral setae................................................................................................ B. andorra sp. nov. - Abdomen entirely black setose; the anteromost acrostichals and dorsocentrals distinctly shorter than those just before suture (first three about 0.09 mm long); hind femur with posteroventral setae (if abdomen partly pale setose, compare B. halterata)....................................................................................... 19 19 (18) Face very narrow (less than 2 facets combined, 0.02 mm), narrowed ventrally; male presutural area evenly dark in dorsal view (male) or with pale spot sharply triangular-shaped (female); hind tibia more equally swollen towards tip, at its broadest point subequally broad as hind femur even in male (both 0.12���0.13 mm); male hypandrial processes slender; apical corner of postgonites broadly rounded (Figs 43���44).............................................. B. vana Collin - Face broader (about 3 facets combined, 0.03 mm), parallel-sided; presutural area lighter in dorsal view than remaining parts of mesoscutum in both sexes, of rectangular to semicircular appearance; hind tibia strongly swollen towards tip (usually more than 0.15 mm at broadest point in male and some 0.14 mm in female), at its broadest point broader than hind femur (0.12 mm); male hypandrial processes broader; apical corner of postgonites narrower and more pronounced (Figs 39���40)........................................................................ B. sulcata (Zetterstedt) 20 (16) Abdomen partly pale setose (at least on sides of tergite 2)................................................. 21 - Abdomen entirely black setose....................................................................... 22 21 (20) Legs microtrichose, with longer and slender tarsi (combined length of fore+mid+hind basitarsus usually more than 1.3 mm); usually less than 7 setae in posthumeral and intrahumeral areas; male hind tibia gradually slightly swollen, lateral parts of abdomen brown; hypandrial processes shorter (0.11 mm), both phallic hooks narrow, connected ventrally (Figs 1��� 2); female tergites 6���8 at least partly microtrichose...................................... B. albopilosa Chv��la - Legs lustrous, with shorter tarsi (combined length of fore+mid+hind basitarsus usually less than 1.2 mm); usually more than 8 setae in posthumeral and intrahumeral areas; male hind tibia strongly and abruptly swollen, lateral parts of abdomen pale and almost silvery grey; hypandrial processes long (0.15 mm), both phallic hooks present, left one hoe-like broadened laterally (best visible from behind) (Figs 35���36); female tergites 6���8 almost entirely lustrous.................................................................................................. B. stackelbergi Tuomikoski 22 (20) Halter yellow; male hypandrium large with contrastingly short, narrow, broadly U-shaped and long setose processes (Fig. 18); female tergites 5���8 almost entirely lustrous......................................... B. kocoureki sp. nov. - Halter brownish yellow to black; male hypandrium different; female tergites 5���8 at least partly microtrichose........ 23 23 (22) Male........................................................................................... 24 - Female......................................................................................... 27 24 (23) Postgonites slightly broadened laterally and narrow in lateral view, hypandrium with short (0.06���0.09 mm) and broad processes (3��� 4 X as long as broad), two equally shaped and long phallic hooks (Figs 41���42)........... B. uvens Melander - Postgonites not broadened laterally and trapezoid in lateral view, hypandrium with narrower and longer processes, a single long phallic hook present (the other less than one-quarter as long)........................................... 25 25 (24) Face broad in ventral portion (usually more than 5 facets combined, 0.06 mm); mid femur with complete row of anteroventrals at least half as long as depth of femur at mid-length; hind tibia with ventral setae about twice as long as depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view; phallic hook bent in S-shaped (Fig. 37). .................................................................................. B. subpilosa Collin - Face narrow in ventral portion (less than 4 facets combined, 0.04 mm); mid femur with anteroventrals shorter than half of femur depths at mid-length; hind tibia with ventral setae at most slightly longer than tibia depth of tibia; prescutellar depression paler or evenly dark as remaining parts of mesoscutum in dorsal view; phallic hook simply bowed (Figs 33, 21)................................................................................................ 26 26 (25) Hypandrium with rather long (0.13 mm), slender and straight processes (Fig. 34).................. B. spuria (Fall��n) - Hypandrium with rather short (0.09���0.11 mm), apically foot-like bent and broadened processes (Fig. 22).. B. mera Collin 27 (23) Face broad in ventral part (usually broader than 0.05 mm, more than 4 facets combined); mid femur with complete row of anteroventrals at least half as long as depth of femur at mid-length; hind tibia with ventral setae about twice as long as depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view..... B. subpilosa Collin - Face narrower in ventral part (usually less than 0.04 mm, about 3 facets combined); mid femur with anteroventrals shorter than one-third of femur depth at mid-length; hind tibia with ventral setae at most slightly longer than depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view or evenly dark...................... 28 28 (27) Usually two dorsal setae on fore trochanter; usually more than 9 setae in posthumeral and intrahumeral areas; hind tibia swollen (usually about 0.13 mm at broadest point)......................................... B. uvens Melander - Usually a single seta on fore trochanter; usually less than 9 setae in p, Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 273-276, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria Macquart Bicellaria Macquart, 1823: 155. Type species: Bicellaria nigra Macquart, 1823: 156 [= spuria (Fall��n, 1816)] (monotypy). Cyrtoma Meigen, 1824: 1. Type species: Cyrtoma atra Meigen, 1824: 2, des. Westwood, 1840: 133 [= spuria (Fall��n, 1816)]. Enicopteryx Stephens, 1829: 264 (catalogue name). Calo Gistel, 1848: VIII, unjustified new name for Cyrtoma. Type species: Cyrtoma atra Meigen, 1824 [= spuria (Fall��n, 1816)]. Diagnosis. Species of the genus are small flies (wing length 1.9 ~ 4 mm), brown to dull black with black or pale setae (almost exclusively pale in B. chimganensis sp. nov.). Head hemispherical with prominent ocellar triangle bearing one to two pairs of ocellar setae and short proboscis, palpus with one to ~ 10 setae. Antenna black, pedicel with circlet of setae, ventral ones often elongated, lateral ones elongated in B. longisetosa Chv��la, 1991, third segment with one to several elongate dorsal setae situated on proximal part of the segment (B. austriaca complex) or on distal part of the segment (B. dispar Oldenberg, 1920) or without seta(e), apical stylus two-segmented. Face sometimes very narrow (about 0.02 mm), even narrower below (B. vana, B. koreana sp. nov., B. nigra complex), in most species slightly narrowed ventrally, but also broadened ventrally (B. pilosa Lundbeck, 1910). Thorax dull black, more or less arched, conspicuously setose. Acrostichals biserial and distinctly separated from dorsocentrals, setae sometimes long (more than 0.15 mm) but shortened in some species (B. koreana sp. nov., extremely so in B. dispar), taxonomically important but overlooked by previous authors is number of setae in intrahumeral and posthumeral areas (i.e., in presutural area laterad from dorsocentrals). Legs slender, at most hind tibia at least slightly swollen apically, usually less distinctly so in females. Fore femur usually with sparse rows of short antero- and posteroventral setae; fore tibia with species-specific ciliation on ventral part in distal third: some species bearing pilosity similar to that of fore basitarsus but absent or very short in other species and posteroventral setae very short, absent or distinctly longer than diameter of tibia in other species (usually short posteroventral setae found in species with long pilosity���typical feature of B. sulcata and B. nigra complexes). Mid femur usually with very short anteroventrals (somewhat longer in B. setitibia sp. nov.) and longer posteroventrals. Mid tibia usually with one to several dorsal pairs of setae situated on basal two-thirds, in B. setitibia sp. nov. with long setae also on apical part. Hind femur usually with two rows of elongate setae anterodorsally and anteroventrally, posteroventrals usually much shorter or (rarely) absent. Hind tibia mostly swollen in males and less so in females, but sometimes almost clavate [B. stackelbergi, B. sulcata (Zetterstedt, 1842)] or narrow and not at all swollen [B. simplicipes (Zetterstedt, 1842)], hind tibia with long and strong apical setae in B. woodi sp. nov. Wing usually at least slightly darkened, cell dm absent, base of vein M often depigmented, vein A 1 reaching (or nearly so) hind wing margin, area between tip of veins Sc and R 1 stigma-like darkened. Abdomen elongate and narrow, in males usually darker in dorsal view and lighter in lateral view. A detailed description of the male genitalia was given by Bart��k and Kub��k (2013) and the following presents a more detailed comparison with respect to Asian fauna. Male genitalia usually small (somewhat larger and almost globular in B. globulicauda sp. nov.). Epandrium, cerci and hypoproct usually very similar between species, apparently without useful diagnostic features (except B. globulicauda sp. nov. with swollen epandrium). Hypandrium usually with two processes (absent in B. farkaci sp. nov. or with additional rounded process between processes in one unidentified specimen from Georgia), in most species bearing several moderately long spines near apex, but also long setose along its length (B. femorata Collin, 1960, B. japonica Kato, 1971). Shape of hypandrial processes very important: in several species processes long and narrow (e.g., B. koreana sp. nov.), or contrastingly short and broad (B. uvens), or long and broad (B. shatalkini sp. nov.). Hypandrium articulates dorsally with symmetrical postgonites (slightly asymmetrical in B. chimganensis sp. nov.), mostly roughly rectangular or trapezoid in lateral view, sometimes short (B. shatalkini sp. nov.) or elongate (B. kovalevi sp. nov.), triangular or broadened laterally (narrow in lateral view in B. uvens), precise shape of postgonites species-specific. Left and right postgonites connected dorsally; middle connected part articulates closely with phallic hooks. Phallus membranous in most species, in B. longisetosa dorsal side sclerotized forming roof-like flat sclerite. The structure of the phallic hooks provides many important differentiating characters. Usually two phallic hooks are developed and sometimes the right one is reduced, although in B. setitibia the left one is reduced (see remarks under the species). Species of B. pilosa -complex have three phallic hooks (left hook bifurcate, extremely so in B. femorata, Fig. 7). Female terminal segments with at least 8 th sternite nearly always shining (except B. farkaci sp. nov.), and basal part of tergite 8 shining in most species (often also basal parts of tergites 6���7, but dry specimens often hidden under preceding tergites); rarely (B. amankutanensis sp. nov.) tergites 5���8 and sternites 6���8 also shining., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 234, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

B. longisetosa Chv��la B. longisetosa Chv��la, 1991: 9. Distribution and diagnosis. See Bart��k and Kub��k (2013)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 246, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria nigra (Meigen) (Figs 23���24) Cyrtoma nigra Meigen, 1824: 3. Cyrtoma rufa Meigen, 1824: 3. Material examined (273, 44��). Czech Republic: 1 ��, ��umava Mts, Nov�� Hůrka, spruce wood, 49 ��09' 17 ''N, 13 �� 19 ' 46 ''E, 880 m, 24.vi.- 28.vii. 2000; 1 ��, ��umava Mts, Horsk�� Kvilda, 49 ��05'N, 13 �� 33 'E, 1130 m, 21.vii.��� 21.viii. 1999 ���all M. Bart��k & ��. Kub��k; 13, 1��, Vracov, 48 �� 58 'N, 17 �� 16 'E, 200 m, 16.v. 1990; 33, 6��, ��umava Mts, ��mauzy, pine wood, 49 �� 10 'N, 13 �� 14 'E, 960 m, 20.viii. 1989; 1 ��, ��umava Mts, Mal�� Niva, peat-bog, 48 �� 55 'N, 13 �� 49 'E, 780 m, 5.vii. 1988; 1 ��, same locality, 21.v. 1992; 3 ��, ��umava Mts, S slope of Panc��ř, meadow, 49 ��09'N, 13 �� 16 'N, 1000 m; 1 ��, ��umava Mts, Sp��lenec, damp meadow, 48 �� 56 'N, 13 �� 57 'E, 800 m, 10.vi. 1989; 3 ��, Jizersk�� hory Mts, Janov nad Nisou, 50 �� 46 'N, 15 �� 12 'E, 700���800 m, 19.vi. 1982; 23, Kokoř��nsk�� důl, mixed forest, 50 �� 26 'N, 14 �� 35 'E, 260 m, 8.vii. 1985; 13, 1��, Kunice, garden vegetation, 49 �� 56 'N, 14 �� 40 'E, 13.vii. 1985; 13, Břežansk�� rokle, mixed wood, 49 �� 58 'N, 14 �� 25 'E, 280 m, 20.vi. 1990; 1 ��, Kosteln�� Lhota, pine wood, 50 ��08'N, 15 ��02'E, 190 m, 12.v. 1988; 13, 2��, Sadsk��, pine forest, 50 �� 10 'N, 14 �� 59 'E, 180 m; 13, Velk�� Popovice, 49 �� 54 'N, 14 �� 39 'E, 470 m, 10.vii. 1988; 23, Vr��ž nr. P��sek, mixed wood, 49 �� 23 'N, 14 ��08'E, 400 m, 22.v. 1995; 1 ��, same locality, 1.vi. 1994; 1 ��, B��lina, Chloumek, hilltop steppe, 50 �� 32 ' 38 ''N, 13 �� 51 ' 32 ''E, 480 m, 15���25.vi. 1998; 13, same locality, 1.vi. 1996; 2 ��, same locality, 13���28.v. 1998; 13, 1��, B��lina, ��trbick�� vrch, edge of wood, 50 �� 33 ' 10 ''N, 13 �� 50 ' 40 ''E, 400 m, 31.v.- 1.vi. 1997; 13, same locality, 8.v. 1994; 13, Duchcov envir., oak wood, 50 �� 37 'N, 13 �� 41 'E, 29.v. 1993; 13, same locality, 29.iv.- 7.v. 1996; 2 ��, same locality, 16.vii. 1992; 13, MS Beskydy, Horn�� Lomn��, 49 �� 32 'N, 18 �� 37 'E, 650 m, 20.vi. 1986; 1 ��, same locality, 740 m, vi.-vii. 1999; 13, MS Beskydy, Ostr��, meadow, 49 �� 37 'N, 18 �� 41 'E, 19.v. 1990 ���all M. Bart��k; 13, Podyj��, Braitava, mixed wood, 48 �� 52 ' 32 ''N, 15 �� 50 '05''E, 530 m, 2.vii.��� 3.viii. 2004, M. Bart��k & ��. Kub��k���(CULSP); 1 ��, Moravsk�� Krumlov, steppe + forest, 31.v. 1995; 2 ��, Vysok�� Chvojno, deciduous forest, 15 �� 59 'N, 50 ��08'E, 296 m, 4.vi. 1998; 1 ��, same locality, 12.v. 1989; 1 ��, same locality, 28.v. 1998 ���all B. Mocek���(MHK). France: 1 ��, Grand Bois (30 km S of Lyon), 45 �� 30 ' 55 ''N, 4 �� 46 ' 20 ''E, 440 m, 21.v. 2006, M. Bart��k (CULSP). Georgia: 2 ��, West Georgia, Borzhomskiy Zapovednik [Borzhomi Nature Reserve], 10���13.viii. 1969; 1 ��, West Georgia, Bakuriani, 18.vii. 1969 ���all V. Kovalev���(ZMMU). Germany: 1 ��, Bayerischer Wald, Spiegelau, 48 �� 57 'N, 13 �� 22 'E, 890 m, 8.vi. 1995, M. Bart��k (CULSP). Italy: 13, Cansiglio, Ponte Val Cappella, 20.vi. 1981, L. Munari; 1 ��, S. Stefano di Cadore, fiuma Piave, viii. 1979, G. Raffone��� (MSNV); 33, 1��, Passo Nigra, meadow + wood, 46 �� 26 ' 39 ''N, 11 �� 35 ' 18 ''E, 1700 m, 5.vii. 2011, M. Bart��k (CULSP). Russia: 33, Salmi, R. Tuomikoski; 13, Salla [probably collected in Russian part of county], R. Frey��� (NHMH); 2 ��, Moskovskaya oblast: Skorotovo, bog in Picetum, 55 �� 41 'N, 36 �� 53 'E, 30.v. 1989, 180 m; 1 ��, Serpuchov, mixed wood, 54 �� 52 'N, 37 �� 36 'E, 150 m, 27.v. 1989; 13, 1��, Abramcevo, along brook, 56 �� 14 'N, 37 �� 59 'E, 200 m, 28.v. 1989 ���all M. Bart��k���(CULSP). Diagnosis. Bicellaria nigra is a member of the B. nigra complex of species as proposed by Chv��la (1991), for characters see under B. halterata. Males may be recognised easily according to the structure of the genitalia, possessing long postgonites (more than twice as long as broad), left phallic hook long, without lateral process, right one strongly reduced (Fig. 23). Identification of females is difficult (see characters given in the key) and notes under B. halterata. Distribution. West Palaearctic species, in Europe ranging from Spain (Chv��la 1981) to the extreme north of Scandinavia (Chv��la 1983). Remarks. Tuomikoski (1955) provided a short redescription and illustration of the hypandrium. Collin (1961) provided a short redescription, illustrated the male hind leg and genitalia, and provided notes on the synonymy. Chv��la (1983) redescribed the species and provided illustrations of the face, antenna, hind leg, mid tibia and genitalia, and provided a synonymy., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 263-265, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria vana Collin (Figs 43 ���44, 46) Bicellaria vana Collin, 1926: 190. Material examined (643, 58��). Andorra: 23, 1��, Pto. de Envalira, pine wood, 42 �� 33 'N, 14.42 'E, 1800 m, 8.vii. 1990, M. Bart��k (CULSP). Austria: 13, 1��, Tauernalm, pasture, 15.vii. 1998; 13, Boden nr. Villach, edge of wood, 15.vii. 1998 ���all M. Bart��k���(CULSP); 13, Bludenz, R. Frey (NHMH). Czech Republic: 13, Kunice, 21.ix. 1974, M. Bart��k (���B. sulcata ��� det. Chv��la); 13, Podyj�� NP, Havran��ky, forest steppe, 48 �� 48 ' 52 ''N, 15 �� 59 ' 48 ''E, 330 m, 16.v.��� 11.vi. 2002; 1 ��, Podyj�� National Park, ��irok�� Pole, river bank, 48 �� 51 ' 30 ''N, 15 �� 51 '01''E, 230 m, 3.v.��� 2.vi. 2003; 1 ��, Podyj�� NP, Č��žovsk�� rybn��k, nr. pond, 48 �� 52 ' 55 ''N, 15 �� 53 ' 11 ''E, 380 m, 27.v.��� 7.vi. 2001; 1 ��, same locality, 28���30.viii. 2001; 1 ��, same locality, 9.ix.��� 28.x. 2004; 13, Podyj�� NP, Břečkov, nr. pond, 48 �� 53 ' 28 ''N, 15 �� 52 ' 32 ''E, 400 m, 26.vi.��� 24.vii. 2001; 13, Podyj�� NP, Pod Ledov��mi slujemi, wetland nr. river, 48 �� 53 '09''N, 15 �� 50 ' 29 ''E, 400 m, 26.vi.��� 24.vii. 2001; 23, 1��, same locality, 30.vii.��� 31.viii. 2002 ���all M. Bart��k & ��. Kub��k; 33, 3��, ��umava Mts, ��pič��ck�� sedlo, meadow, 49 �� 10 'N, 13 �� 13 'E, 900 m, 21.viii. 1989; 1 ��, ��umava Mts, Horsk�� Kvilda, damp meadow, 49 ��03'N, 13 �� 34 'E, 1000 m, 21.vii. 1992; 1 ��, ��umava Mts, Ant��gl, 49 ��02'N, 13 �� 30 'E, 23.viii. 1989; 13, ��umava Mts, S Slope of Panc��ř, meadow, 49 ��09'N, 13 �� 16 'E, 1000 m, 24.viii. 1989; 23, ��umava Mts, Sp��lenec, damp meadow, 48 �� 56 'N, 13 �� 57 'E, 800 m, 15.ix. 1990; 2 ��, ��umava Mts, Popeln��, 49 ��06'N, 13 �� 38 'E, 3���9.vii. 1988 ���all M. Bart��k���(CULSP); 1 ��, ��umava Mts, Čern�� Kř��ž, peat���bog, 48 �� 52 'N, 13 �� 50 'E, 760 m, 13.vii. 1986, K. ��patenka; 3 ��, ��umava Mts, Rokyteck�� slať, peat���bog, 49 ��00' 59 ''N, 13 �� 25 '05''E, 1100 m, 20.vii.��� 24.ix. 1999; 13, same locality, 21.vii.��� 20.viii. 1999 ���all M. Bart��k & ��. Kub��k; 1 ��, ��umava Mts, Jezern�� slať, peat���bog, 49 ��02'N, 13 �� 34 'E, 980 m, 5.viii.��� 6.ix. 1996; 13, same locality, 5���7.viii. 1996; 43, ��umava Mts, Chalupsk�� slať, 49 ��01'N, 13 �� 39 'E, 860 m, 5���7.viii. 1996; 1 ��, same locality, 4.vii. 1996 ���all M. Bart��k & J. Roh��ček; 1 ��, ��umava Mts, Mal�� Niva, peat���bog, 48 �� 54 'N, 13 �� 49 'E, 21.viii. 1997, ��. Kub��k (CULSP); 13, Kunice, 49 �� 56 'N, 14 �� 40 'E, 30.vii. 1985; 13, same locality, 17.viii. 1975; 1 ��, same locality, 21.ix. 1974; 1 ��, same locality, 11.vi. 1983; 13, same locality, 17.vi. 1984; 13, same locality, 14.ix. 1984; 1 ��, same locality, 19.ix. 1982; 1 ��, Buchlov, edge of mixed wood, 49 ��07'N, 17 �� 18 'E, 400 m, 16.x. 1987; 13, Velk�� Popovice, 49 �� 54 'N, 14 �� 39 'E, 470 m, 16.vii. 1989; 1 ��, same locality, 26.ix. 1982; 13, Lukavec, damp meadow, 49 �� 34 'N, 15 ��00'E, 14.v. 1990; 23, 3��, S��zava, damp meadow, 49 �� 55 'N, 14 �� 55 'E, 26.ix. 1991; 13, same locality, 10.vi. 1990; 13, 1��, Davle, edge of mixed wood, 49 �� 54 ���N, 14 �� 23 'E, 300 m, 22.v. 1988; 33, Duchcov, 50 �� 37 'N, 13 �� 41 'E, 29.v. 1993; 1 ��, same locality, 21.ix. 1992; 1 ��, Čel��kovice, lowland wood, 50 �� 10 'N, 14 �� 49 'E, 180 m, 10.ix. 1991; 13, Skuteč, edge of damp wood, 49 �� 49 'N, 16 ��00'E, 440 m, 1.vii. 1994; 13, Kozlov, meadow, 49 �� 24 'N, 15 �� 41 'E, 340 m, 27.v. 1986; 13, Mnichovice, meadow nr. pond, 49 �� 56 'N, 14 �� 41 'E, 370 m, 3.vi. 1984; 13, Vr��ž nr. P��sek, 49 �� 24 'N, 14 �� 7 'E, 400 m, 20���26.vi. 2009; 13, same locality, 18���22.vi. 2007; 1 ��, B��lina, ��t��p��nov, mixed wood, 50 �� 31 ' 59 ''N, 13 �� 51 ' 39 ''E, 380 m, 13���28.v. 1998; 1 ��, B��lina, Chloumek, 50 �� 33 'N, 13 �� 51 'E, 25.vi.��� 24.vii. 1998, 480 m; 1 ��, same locality, 18.vi. 1994 ���all M. Bart��k��� (CULSP). France: 13, Col du Cabaretous, edge of wood, 43 �� 32 '02''N, 2 �� 45 ' 24 ''E, 940 m, 25.v. 2006; 13, 1��, Grand Bois, edge of wood, 45 �� 30 ' 55 ''N, 4 �� 46 ' 20 ''E, 21.v. 2006; 43, Sainte Baume, 43 �� 20 'N, 5 �� 44 'E, 400���600 m, 15���16.v. 2012 ���all M. Bart��k���(CULSP). Georgia: 1 ��, West Georgia, env. Poti, 22.v. 1969; 1 ��, West Georgia, Borzhomskij Nature Reserve, 7.vii. 1969 ���all V. Kovalev���(ZMMU). Germany: 13, Hamburg, Eppendorfer moor, 10.ix. 1933, O. Kr��ber; 13, Allg��u, Burgegg, 14���28.vii. 1933, O. Kr��ber; 13, Allg��u, Ein��dsbach, 1933, O. Kr��ber���(NHMH). Great Britain: 1 ��, Coppersole, 27.viii. 1950; 1 ��, Pennington Hants, 24.v. 1948 ���all C.N. Colyer���(BMNH); 1 ��, Surrey, Ranmore, 30.v. 1948; 1 ��, Surrey, Bookham, 13.vii. 1947; 1 ��, same locality, 12.vi. 1948 ���all L. Parmenter���(BMNH). Italy: 11 ��, Torbiera di Lipoi, 7.vi. 1981, L. Munari; 13, Lases, 4.v. 1990, Brigolli; 13, 1��, Veneto, Onara Tombolo, 20.iv. 1985; 1 ��, Sila, 18.v. 1985; 13, Torino, 26.v. 1918, T. Sangone; 1 ��, S. Stefano di Cadore, fiuma Piave, viii. 1979, G. Raffone���(MSNV); 13, Lago di S. Croce, 46 ��07'N, 12 �� 20 'E, nr. lake, 1.viii. 1988; 13, Passo Nigra, 46 �� 26 ' 39 ''N, 11 �� 35 ' 18 ''E, 1700 m, meadow + wood, 5.vii. 2011; 1 ��, Weisslahnbad, edge of wood, 46 �� 28 ' 40 ''N, 11 �� 34 ' 11 ''E, 1400 m, 4.vii. 2011 ���all M. Bart��k���(CULSP). Romania: 13, Orsova, 3.vi. 1912 (��� B. sulcata ��� det.?) (SDEI). Russia: 13, Moskovskaja oblast, Abramcevo, 3.viii. 1965, Usačev; 13, Moskovskaja oblast, Golicyno, 7.vii. 1983, A. Shatakin; 13, Karachay���Cherkesskaya Republic, Caucasus, Teberda, 1200 m, 14.vi. 1965, N. Lap��ina���(ZMMU); 13, Moscow Prov., Choťkovo, 56 �� 14 'N, 37 �� 53 'E, mixed wood, 31.viii. 1988; 13, Moscow Prov., Abramcevo, along brook, 56 �� 14 'N, 37 �� 59 'E, 200 m, 28.v. 1989 ���all M. Bart��k���(CULSP). Slovak Republic: 13, N��zk�� Tatry, Demanovsk�� dol., 2.vii. 1975, M. Bart��k (��� R. sulcata ��� det. Chv��la); 13, Vysok�� Tatry, Doln�� Smokovec, 28.v. 1968, M. Chv��la (��� R. sulcata ��� det. Chv��la)���(OUMNH). Spain: 13, Picos Europa, R. Frey (NHMH). Ukraine: 1 ��, Carpathians, Jasinja, Stanislav-Gat, 26.vii. 1972, V. Kovalev (ZMMU). Diagnosis. The species may be easily identified and distinguished from allied B. sulcata according to the genitalia: B. sulcata has distinctly pronounced (elongated) upper dorsal angle of the postgonites, which is more rounded and less pronounced in B. vana [this character was properly illustrated by both Chv��la (1983, fig. 356: B. vana and fig. 352: B. sulcata) and Collin (1961, fig. 91: B. sulcata and fig. 92: B. vana)]. Left phallic hook long, right one much shortened; ventral apodeme broadened in lateral view, but very narrow in dorsal view; hypandrium of both species is also different with much narrower processes in B. vana (Figs 43���44) than in B. sulcata (Figs 39��� 40). We believe that also females of both species may be distinguished according to characters given in the key despite we had no pair in copula. We have specimens similar to B. vana from southern Spain, which may represent different species, see section ���Unidentified specimens���. Distribution. Broadly distributed West Palaearctic species. Published record [omitted in catalogs by Chv��la (2012) and Yang (2007)]: Romania (Parvu 2003). Remarks. Collin (1961) provided a short redescription, and illustrations of the hind leg and genitalia. Chv��la (1983) also redescribed this species and provided illustrations of the face, antenna, hind leg, and genitalia., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 272-273, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria collini Tuomikoski (Figs 9���10) Bicellaria collini Tuomikoski, 1955: 70. Material examined (533, 9��). Austria: 13, Boden nr. Villach, edge of wood, 15.vii. 1998; 53, Sulzgau, along river, 47 �� 33 'N, 13 �� 11 'E, 30.vii. 1988; 83, 1��, Brennersee, nr. lake, 47 ��00' 58 ''N, 11 �� 30 ' 18 ''E, 1320 m���all M. Bart��k���(CULSP); 33, Tirol, Stafflach, 19���20.vi. 1933, A.C.W. Wagner (NHMH). Italy: 53, 4��, Passo Rolle, alpine meadow, 46 �� 13 'N, 11 �� 42 'E, 1900 m, 8.viii. 1988; 193, 1��, Passo Nigra, meadow + wood, 46 �� 26 ' 39 ''N, 11 �� 35 ' 18 ''E, 1700 m, 5.vii. 2011; 63, 1��, Weisslahnbad, edge of forest, 46 �� 28 ' 40 ''N, 11 �� 34 ' 11 ''E, 1400 m, 4.vii. 2011 ���all M. Bart��k���(CULSP). Romania: 53, 2��, Mt. Bucegi, 16.vii. 1970, M. Chv��la (��� B. collini ��� det. Chv��la) (OUMNH). Diagnosis. Bicellaria collini is a member of the B. austriaca complex of species as defined by Chv��la (1991), for characters see under B. austriaca. All species of this complex may be easily identified according to the key using the male sex. Males have two rather long and almost equally shaped phallic hooks and broadly U-shaped hypandrial processes armed with rather long setae (Figs 9���10). Distribution. Mountains of Central and Southern Europe. Remarks. This species was described (and hypandrium illustrated) on the basis of four males and four females from ���Bayern, Allg��u, Burgegg��� (Germany)., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 256-257, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria stackelbergi Tumikoski (Figs 35���36) Bicellaria stackelbergi Tumikoski, 1955: 74. Type material examined. HOLOTYPE 3: Russia: Kamtschatka, Ozernaja, 30.vii. 1917, Y. Wuorentaus (labelled: ���Mus. Zool. H:fors, Spec. typ. No. 8200 Bicellaria Stackelbergi Tuomikoski ���). PARATYPE: 13, Kamtschatka, Bolscherjetsk [now Ust���Bolsheretsk], 20.vii. 1917, Y. Wuorentaus���(NHMH). Additional material examined (33, 2��). Russia: 23, 2��, Archangel'skaya oblast, Kanin, B. Poppius; 13, Republic Khakassia, r. Abakan, s. Nemir (r. = river, s. = village), 11.v. 197?, Wagner���(NHMH). Diagnosis. The species may be easily recognized according to the key. Almost silvery dusted abdomen is similar to B. alpina, but this latter species has black setose body and a long seta on the third antennal segment (Fig. 48). Lustrous legs is a unique feature of this species among European fauna (remaining known species have microtrichose legs except sometimes posterior part of hind femur). Male postgonites are narrowed apically, both phallic hooks present, nearly equally long, left one strongly (right one slightly) broadened laterally (best visible from behind). Interestingly, one female paratype represents in fact B. uvens. Distribution. Poorly known, probably Palaearctic species, in Europe known only from extreme North (Kola, Kanin). Remarks. The species was described (and hypandrium illustrated) on the basis of specimens from Kola, Kamtschatka (Bolscherjetsk [now Ust���Bolsheretsk]), Ozernaja river), Jakutsk and Altai., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 269, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria intermedia Lundbeck Bicellaria intermedia Lundbeck, 1910: 25. Distribution and diagnosis. See Bart��k and Kub��k (2013)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 244, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria mera Collin (Figs 21���22) Bicellaria mera Collin, 1961: 258. Type material examined. SYNTYPES: Great Britain: 1 ��, Upware Cambs, 12.vii. 1875; 2 ��, Wicken, Cambs, 21.viii. 1905; 13, same locality, 21.vi. 1905 ���all G.H. Verrall���(BMNH). Additional material examined (103, 2��). Czech Republic: 63, 2��, Čel��kovice, lowland wood, 50 �� 10 'N, 14 �� 49 'E, 180 m, 10.ix. 1991 (CULSP). Great Britain: 13, Wood Walton, Fen, Hunts, 16���18.vi. 1923, F.W. Edwards (BMNH). Slovak Republic: 13, ��amor��n, near river, 6���10.vi. 1977 (��� B. spuria ��� det. Chv��la); 13, ��tůrovo, 2 km N, damp meadow, 47 �� 49 'N, 18 �� 56 'E, 110 m, 24.vii. 1989; 13, Jursk�� ��ur, deciduous wood, 48 �� 15 'N, 17 �� 16 'E, 130 m, 24.viii. 1989 ���all Bart��k���(CULSP). Diagnosis. Bicellaria mera is very similar to B. spuria. Both species may be identified in the male sex according to the hypandrial processes which are foot-like bent near apex in B. mera but straight and narrow in B. spuria (Fig. 22). This character is sometimes poorly visible and requires dissection. Identification of females is difficult because two candidate characters from Collin���s description we tried to specify based on limited materials at hand (hind tibia at its broadest point usually narrower than 0.09 mm in B. mera but usually broader than 0.09 mm in B. spuria; narrow apical part of third antennal segment mostly about as long as the first segment of stylus in B. mera, but twice as long in B. spuria) proved to be variable. The only female character which appears constant is given in the key. Distribution. Temperate Central and Western Europe. Published record [omitted by Chv��la (2012) and Yang (2007)]: Romania (Parvu 2005), record from Italy (Raffone 2006) is doubtful. Remarks. Described (alongside illustration of genitalia and hind leg) on the basis of unknown number of specimens taken in Wicken, Chippenham, Fens (Cambs) (England). Chv��la (1983) redescribed this species and provided illustrations of the face, antenna, hind leg, and genitalia., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 263, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria italica sp. nov. (Figs 15���16) Type material. HOLOTYPE 3, Italy, Monti del Matese, pasture, 41 �� 25 ' 27 ''N, 14 �� 27 ' 20 ''E, 1550 m, 18.viii. 2004, M. Bart��k (CULSP). PARATYPES: Andorra, 53, Pyrenees, Pto de Envalira, meadow nr. wood, 42 �� 35 'N, 1 �� 40 'E, 1400m, M. Bart��k (CULSP). Austria, 13, Lienz, 5 km E, edge of wood, 46 �� 50 'N, 12 �� 45 'E, 31.vii. 1988, M. Bart��k (CULSP). France, 143, Fabian, pasture nr. brook, 42 �� 48 'N, 0�� 11 'E, 9.vii. 1990, 1100 m, M. Bart��k (CULSP); 13, Ax-les-Thermes, along river, 42 �� 44 'N, 1 �� 50 'E, 8.vii. 1990, 750 m, M. Bart��k (CULSP). Italy, 63, same data as holotype (CULSP); 83, Mte. Terminillo, meadow nr. wood, 42 �� 26 'N, 13 ��07'E, 2000 m, 4.viii. 1988, M. Bart��k (CULSP); 183, S. Martino, spruce wood, 46 �� 11 'N, 11 �� 41 'E, 1300���1500 m, 8.viii. 1988, M. Bart��k (CULSP); 13, Lago di S. Croce, deciduous wood nr. lake, 46 ��07'N, 12 �� 20 'E, 1.viii. 1988, M. Bart��k (CULSP); 93, Weisslahnbad, edge of forest, 46 �� 28 ' 40 ''N, 11 �� 34 ' 11 ''E, 1400 m, 4.vii. 2011, M. Bart��k (CULSP); 23, S. Stefano di Cadore, fiuma Piave, viii. 1979, G. Raffone (MSNV). Diagnosis. Mesoscutum black setose; third antennal segment with dorsal seta; abdomen pale setose on sides of basal segments; halter yellow; male hypandrial processes swollen apically, with several rather long setae on apical part; female abdominal segments 6���7 nearly completely microtrichose. Description. Male. Head black, brownish-grey microtrichose, black setose (lower occiput with pale setae). Eyes meeting on frons, facets on dorsal half much larger than on ventral half, about 11���14 facets in line of contiguity of eyes. One pair of long black ocellar setae. Face slightly narrowing ventrally, dorsal part just below antennae about 0.05 mm broad and ventral part about 0.04 mm broad at narrowest point. Palpus short, brown, with 2���3 setae. Labrum brown, lustrous. Antenna black, second segment short setose (longest ventral setae about 0.10 mm); third segment with single elongate dorsal seta; ratio of broad part of segment 3: narrow part of segment 3: stylus (in 0.01 mm scale) = 13���15: 4���5: 19���22. Thorax black to brownish black, microtrichose, prescutellar area and scutellum lighter than disc of mesoscutum, light prescutellar area triangular-shaped reaching hind acrostichals. All thoracic setae black, only fringes on squama mostly pale. Chaetotaxy: acrostichals broadly biserial (7���8 setae in each row) and long even in presutural area (about 0.10���0.13 mm in front but longer posteriorly); dorsocentrals nearly evenly long, about 5���7 setae in posthumeral and intrahumeral areas; 1 long and several slightly shorter postpronotals; notopleura with 3���5 unequally long setae in one irregular row; 1 supraalar and 0���1 seta in prealar area; 1 long postalar; 2 pairs of scutellars. Legs including coxae black, microtrichose and black setose (only hind coxa with several pale setae posteriorly). Fore femur with row of short anteroventrals in basal half and row of longer posteroventrals along whole length (half as long as depth of femur). Fore tibia with longest posterodorsals 1.5 X as long as depth of tibia, posteroventrals about as long as depth of tibia (much longer than very short pilosity). Mid femur with row of short anteroventrals (one third as long as depth of femur) and with much longer posteroventrals about as long as depth of femur. Mid tibia with ventral setae equally long (about as long as depth of tibia) and with 1���2 antero- and 0���1 posterodorsal setae up to 3 X as long as depth of tibia (0.20 mm). Hind femur slender (0.10 mm), with 14���16 dorsal and 16���19 anteroventral setae up to 2 X as long as depth of femur (on basal part, apically shorter) and with much smaller posteroventrals. Hind tibia very slightly and evenly swollen in apical half (0.11 mm at broadest point), dorsal and ventral setae slightly longer than depth of tibia. Tarsi of all legs thin and short setose. Wing slightly brownish infuscated, stigma dark brown (0.06 mm broad), about 3.6 X as long as distance between tips of veins R 1 and R 2 + 3. Squama brown with pale fringes, halter yellow. Abdomen black, brownish-grey microtrichose, setae on basal segments and ventrally pale, on apical segments and dorsally dark. Genitalia (Figs 15���16) with hypandrial processes swollen and with several rather long setae apically; both phallic hooks almost equally long, right one more sharply bent. Female. Similar to male. Tergites 6���7 nearly entirely microtrichose, sternite 8 lustrous. Length: body 2.8���3.9 mm, wing 2.7���3.4 mm. Etymology. The species is named after the country of origin (Italy). Distribution. Andorra, Austria, France, Italy. Remarks. The species described above is a member of the B. austriaca complex of species as defined by Chv��la (1991), for characters see under B. austriaca. Males of all four species of this complex (B. italica, B. austriaca, B. intermedia, B. collini) may be easily identified according to the key due to different genitalia. However, identification of females is difficult. In fact we found no characters to reliably differentiate B. italica sp. nov. from B. intermedia (originally described from Denmark). From this reason females were excluded from the type series of the above described species as well as from the faunistic records of B. intermedia., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 260-261, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria nigrita Collin (Figs 25���26) Bicellaria nigrita Collin, 1926: 190. Material examined (73, 9��). Czech Republic: 1 ��, Dobř����, 3 km W, edge of wood, 49 �� 47 'N, 14 ��08'E, 400 m, 19.vi. 1988; 13, Bojovsk�� ��dol��, damp valley, 49 �� 54 'N, 14 �� 22 'E, 230 m, 22.v. 1988 ���all M. Bart��k; 13, Podyj�� NP, Hnanice, damp forest, 48 �� 48 ' 12 ''N, 15 �� 58 ' 22 ''E, 300 m, 24���27.v. 2001; 1 ��, Podyj�� NP, Terasy, mixed wood, 48 �� 53 ' 22 ''N, 15 �� 50 ' 18 ''E, 460 m, 3.v.��� 2.vi. 2003; 1 ��, Podyj�� NP, Hardegg, vyhl��dka, mixed wood, 48 �� 51 ' 30 ''N, 15 �� 51 ' 35 ''E, 420 m, 21.v.��� 12.vi. 2004 ���all M. Bart��k & ��. Kub��k���(CULSP); 13, Pojihlav��, Mohelno reserve, 30.v. 1995, B. Mocek (MHK). France: 13, 3��, Col du Cabaretous, edge of wood, 43 �� 32 '02''N, 2 �� 45 ' 24 ''E, 940 m, 25.v. 2006; 33, 2��, Saint Etienne, forest edge, 43 �� 28 ' 15 ''N, 2 �� 51 ' 33 ''E, 540 m, 25.v. 2006 ���all M. Bart��k��� (CULSP). Greece: 1 ��, 7 km E of Mt. Olympos, 40 ��06' 13 ''N, 22 �� 25 ' 36 ''E, 1100 m, 23.v. 2007, M. Bart��k (CULSP). Diagnosis. Bicellaria nigrita may be, as are all three European species of the B. nigra complex, easily identified in males according to the genitalia. Postgonites are shortened, both phallic hooks present and longer than the postgonites. Moreover, males have a black setose body which differentiate it from both other species of this complex. However, identification of females may be difficult, differing by characters given in the key; however, the variation range of these characters remains unknown. Distribution. Temperate and southern Europe from Great Britain to central European Russia, south to Bulgaria and Greece. Remarks. Tuomikoski (1955) provided a short redescription and illustration of the hypandrium. Collin (1961) briefly redescribed this species and provided an illustration of the genitalia. Chv��la (1983) redescribed this species and provided illustrations of the face, antenna, mid tibia, hind leg, and genitalia., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 265, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria kocoureki sp. nov. (Figs 17���18) Type material. HOLOTYPE 3: Bulgaria, Pirin, Banderica, 1800 m, 7.vii. 1982, F. Kocourek (CULSP). PARATYPES: Bulgaria, 53, 7��, same data as holotype (CULSP); 13, 2��, Pirin, Begovica, 1760 m, 17.1 km NW from Sandanski, 41 �� 40 '09''N, 23 �� 25 ' 33 ''E, 10.viii. 2005, J. Fechtner (CULSP). Diagnosis. Entirely black setose species; third antennal segment without dorsal setae, broadened part of third segment very short in comparison to its narrow part and stylus; halter yellow in both sexes; male hypandrium with short and long setose processes; female tergites 5���8 almost entirely lustrous. Description. Male. Head black, brownish grey microtrichose, black setose (including lower occiput). Eyes meeting on frons, facets on dorsal half much larger than on ventral half, about 10���12 facets in line of contiguity of eyes. One pair of long black ocellar setae. Occiput rather sparsely setose. Face nearly parallel-sided and about 0.05 mm broad, clypeus slightly paler grey microtrichose. Palpus short, brown, with 2 setae. Labrum brown, lustrous. Antenna black, second segment short setose (longest ventral setae about 0.08 mm); third segment without dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: stylus (in 0.01 mm scale) = 9���12: 5���8: 17���25. Thorax black to brownish black, slightly subshining, prescutellar area (broad and reaching hind acrostichals) and scutellum lighter than disc of mesoscutum. All thoracic setae black including fringes on squama. Chaetotaxy: acrostichals biserial and long even in presutural area (about 0.15 mm in front but longer posteriorly); dorsocentrals (with exception of first 1���2 pairs) slightly longer than corresponding acrostichals; about 3���8 setae in posthumeral and intrahumeral areas; 1 longer and 2���3 slightly shorter postpronotals; notopleura with 3���6 unequally long setae in one irregular row; 1 supraalar and 1���2 setae in prealar area; 1 long postalar; 2���3 pairs of scutellars. Legs including coxae black, microtrichose (only trochanters partly lustrous) and black setose. Fore femur with row of short anteroventrals and row of slightly longer posteroventrals (half as long as depth of femur). Fore tibia with 1���3 longest posterodorsals (in basal half of tibia) 1.5 X as long as depth of tibia, posteroventrals slightly longer than depth of tibia. Mid femur with sparse row of anteroventrals half as long as depth of femur, with posteroventrals subequally as long as depth of femur. Mid tibia with ventral setae about as long as depth of tibia, with 1���2 pairs of dorsal setae up to 0.20 mm long (longest about 4 X as long as diameter of tibia). Hind femur with 14���15 anterodorsal and 14���15 anteroventral setae slightly longer than depth of femur, posteroventrals shorter than corresponding anteroventrals. Hind tibia slightly swollen in apical half (about 0.11 mm at broadest point), subequally broad or slightly narrower than hind femur (about 0.12 mm at broadest point), ventral setae about as long as depth of tibia, 2���3 pairs of dorsal setae up to 3 X as long (0.20 mm). Tarsi of all legs thin and short setose. Wing slightly brownish infuscated, stigma light brown and narrow (0.04���0.05 mm), about 2.2 X as long as distance between tips of veins R 1 and R 2 + 3. Squama brown with black fringes, halter yellow. Abdomen black, dull brownish black microtrichose in dorsal view and grey in lateral view and posteriorly, rather short and entirely black setose. Genitalia (Figs 17���18) unusual in view of European species of Bicellaria: hypandrium long, processes narrow and short, broadly U-shaped, covered with several rather long setae; postgonites small and narrow, both phallic hooks subequal in length, phallus with two smaller additional hooks derived probably from ejaculatory apodeme (appearing as additional smaller hook in lateral view), ventral apodeme slightly broadened laterally. Female. Similar to male. Last three abdominal segments (6���8) almost entirely lustrous. Length: body 2.5���3.5 mm, wing 2.3���2.9 mm. Etymology. The species is named after the collector of the type series, Dr. Franti��ek Kocourek (Prague). Distribution. Bulgaria. Remark. The species described above can be easily identified according to the key. It is rather isolated in position among European Bicellaria with possible affinity to the B. uvens complex of species., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 261-262, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria albopilosa Chv��la (Figs 1 ��� 2) Bicellaria albopilosa Chv��la, 1991: 14. Material examined (773, 97��). Austria: 23, 1��, Niedere Tauern, S��lker Pass, 47 �� 16 'N, 14 ��04'E, 1900 m, 7��� 8.viii. 1995; 13, Schladminger Tauern, Donnersbachwald-Stallaalm, 1600 m, viii.-x. 1999, Malaise trap; 13, 1��, Grossglockner, spruce wood, 47 ��06'N, 12 �� 43 'E, 1400���1700 m, 31.vii. 1988 ���all M. Bart��k���(CULSP). Czech Republic: 363, 19��, Krkono��e Mts (Lučn�� hora, Lučn�� bouda, Labsk�� důl, Labsk�� bouda, Velk�� Koteln�� j��ma, Dvorsk�� potok, Labsk�� rokle, Pančava, Medv��d��n, U bufetu, Li��č�� hora), altitude range: 1040���1500 m, vii.-ix.���all J. Van��k et M. Bart��k���(CULSP). Germany: 13, 2��, Bayerische Wald, Spiegelau, 890 m, 48 �� 57 'N, 13 �� 22 'E, 7.ix. 1995, M. Bart��k (CULSP). Slovak Republic: 263, 73��, Vysok�� Tatry, several localities (Javorov�� dolina, Tich�� dolina, Mal�� Studen�� dolina, Hrebienok, Star�� Smokovec), spruce forest, meadow and clearing in forest, 22.vii.- 17.viii., altitude range: 1100���1700 m (CULSP). Italy: 103, Passo Rolle, larch wood and alpine meadow, 46 �� 13 'N, 11 �� 42 'E, 1700���1900 m, 8.viii. 1988, M. Bart��k (CULSP). Switzerland: 1 ��, GR, Lenzerheide, pasture, 2000 m, 14.��� 21.vii. 2000, B. Merz (CULSP). Diagnosis. The species has elongate-trapezoid postgonites, relatively narrow V-shaped divergent hypandrial processes, both phallic hooks present, connecting ventrally (forming closed ring, right hook slightly shorter than left hook), median extension of postgonites saw-toothed ventrally (Figs 1 ��� 2). Presutural area mostly similarly coloured as rest of mesoscutum in males but lighter in females. Distribution. Mountains of Central and Southern Europe. Remarks. Bicellaria albopilosa was described from numerous specimens originating from the Krkono��e Mts, Orlick�� hory, and Carpathian Mts (Slovak Republic and Romania), exclusively from high altitudes (from 800 to 2200 m) (Chv��la 1991). Chv��la (ibid.) illustrated the fore and hind legs, antenna and hypandrium. There may be at least one similar species (see Bicellaria sp. 5 in section ���Unidentified specimens���) and, therefore, careful examination of genitalia is necessary., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 253, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria simplicipes (Zetterstedt) (Figs 31���32) Cyrtoma simplicipes Zetterstedt, 1842: 331. Material examined (73, 6��). Czech Republic: 1 ��, P��sek-Čimelice (car net), 28.vi. 1992; 2 ��, Vr��ž nr. P��sek, 49 �� 23 'N, 14 ��08'E, 400 m, 30.v. 1994; 23, 1��, same locality, 2.vi. 1994; 1 ��, Čertova voda nr. D��č��n, 130 m, 50 �� 48 ' 47.3 ''N, 14 �� 13 ' 35.3 ''E, meat-baited Malaise trap, 21.v.- 20.vi. 2009; 13, Čel��kovice, 4 km E, lowland wood, 50 �� 10 'N, 14 �� 49 'E, 180 m, 6.vi. 1990; 1 ��, Přerov nad Labem, river bank, 50 �� 11 'N, 14 �� 48 'E, 180 m, 6.ix. 1983 all M. Bart��k���(CULSP). Finland: 13, Tv��rminne, R. Frey; 13, Liminka, R. Tuomikoski (NHMH). Russia: 13, Tyt��rsaari (= Bolshoy Tyuters), W. Hell��n; 13, Suistamo, R. Tuomikoski (NHMH). Diagnosis. An easily recognisable species due to narrow (apically almost not swollen) hind tibia and peculiar genitalia with shortened postgonites (bearing small triangular-shaped extension best visible in ventral view), nearly triangular-shaped left phallic hook (somewhat similar to the situation in the B. pilosa complex), not visible in Chv��la (1983, fig. 309, p. 145) or Collin (1961, fig. 92, p. 263) is probably due to the occasional alignment of the cranial part of the hook with the median extension of the postgonites, and short and broad hypandrial processes armed with apical setae. Distribution. Europe. Published record from Italy (Raffone 2006) was based on a misidentification [not included in recent catalogs by Chv��la (2012) and Yang (2007)]. Remarks. Tuomikoski (1955) provided a short redescription and illustration of the hypandrium. Collin (1961) also wrote a short redescription, illustrated the male genitalia and hind leg, and provided notes to the synonymy. Chv��la (1983) gave a redescription, synonymy, lectotype designation, and illustrations of the genitalia, face, antenna, and hind leg. The species is rather isolated in position without clear affinities., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 267, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria koreana sp. nov. (Figs 20, 21) Type material. HOLOTYPE ♂, North Korea, S slope of Paekdu Mt., 16.viii. 1989, M. Koz��nek (CULSP). PARATYPES: N. Korea: 3 ♂, 10 ♀, same data as holotype; 8 ♀, Paekdusan, Mupo, 18.vii. 1989, M. Koz��nek; 1 ♀, N. Korea, Paekdusan, Samjion, 40 km NW, 17.viii. 1989, M. Koz��nek; 1 ♀, N. Korea, Paekdusan, Onsupjong, 19.viii. 1989, M. Koz��nek; 1 ♀, N. Korea, Čchong-Rion-San, 1.viii. 1989, M. Koz��nek���(CULSP). Russia: [labels in Cyrillic] Chita: 1 ♀, Černy��evsk [= Chernyshevsk], 17.vii. 1977; 1 ♂, same locality, 19.vii. 1977; 1 ♂, same locality, 23.vii. 1977; 5 ♂, Kuenga River, above Chernyshevsk, 8.viii. 1977; 2 ♀, same locality, 10.viii. 1977; 1 ♂, Ukur'evskoe, 1.viii. 1977, V. Kovalev; 1 ♂, same locality, 6.viii. 1977, V. Kovalev; Amur: 1 ♂, 1 ♀, town Zeja, 2.ix. 1981, A. Shatalkin; Tyva Republic [= Tuva]: 1 ♀, Saryg-Sep, 28.viii. 1973, V. Kovalev; Kamchatka: 1 ♀, Kamenskoe, 29.vii. 1973, Negrobov; Magadan: 1 ♀, Palatka, Hasych, 28.viii. 1978, V. Kovalev���(ZMMU). Diagnosis. Halter brown in both sexes; third antennal segment without elongate dorsal setae; fore tibia with posteroventrals not longer than pilosity; body setae both black and pale; hind basitarsus narrow and hind tibia swollen in male and less so in female. Male hypandrial processes long and narrow; only one (left) long and simply curved phallic hook present. Etymology. The species is named after the country where holotype specimen was taken (Korea). Description. Male. Head brownish-black, brownish grey microtrichose, both black and pale setose. Holoptic, facets in dorsal half much larger than in ventral half, about 15���16 facets in line of contiguity of eyes. Two pairs of black ocellar setae, posterior pair much shorter. Occiput dark setose on upper part and pale setose below. Face strongly narrowed ventrally, dorsal part about 0.06 mm and ventral part about 0.02 mm broad at narrowest point (as in B. vana). Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2 pale setae. Labrum short, brown, polished. Antenna black, 2 nd segment very short setose (longest ventral setae about 0.06 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 12���15: 4���5: 14���16. Thorax blackish-brown, rather light greyish-brown microtrichose, prescutellar area evenly coloured as remaining mesoscutum, scutellum paler grey. Thoracic setae varying from pale to black, with marked tendency to be pale in some specimens (pale in holotype), most notably scutellars and notopleurals (also costal seta) in some specimens whitish-yellow. Chaetotaxy: 7���9 broadly biserial acrostichals; 9���12 uniserial dorsocentrals, both acrostichals and dorsocentrals very short (0.08���0.10 mm) except last 2���3 pairs of dorsocentrals; some 5���8 setae in intrahumeral and posthumeral areas; 3 subequally long postpronotals; notopleuron with 3���5 setae arranged in irregular row (1���2 setae of row often longer and stronger than others); 1 supraalar- and 1���2 prealar; 1 long postalar; 2 pairs of scutellars. Coxae blackish brown, microtrichose, pale (in holotype) but sometimes brown setose. Legs brown, microtrichose and mostly dark setose with at least some pale setae present. Fore femur with sparse rows of fine antero- and posteroventral setae at most half as long as depth of femur. Fore tibia with dorsal setae about as long as depth of tibia or slightly longer, posteroventrals almost absent in distal half of tibia (not longer than pilosity), ventral pilosity slightly shorter than on fore basitarsus. Mid femur with very short or almost absent anteroventrals, posteroventrals about as long as depth of femur. Mid tibia with 1���2 pair(s) of submedian setae dorsally 2 X as long as depth of tibia. Hind femur thin (0.09���0.10 mm), with 15���17 anterodorsal setae more than 2 X as long as depth of femur on basal part but much shorter apically and with row of 13���16 anteroventrals subequally as long as anterodorsals (except on basal part), posteroventrals scarcely half as long as depth of femur. Hind tibia strongly swollen in apical part (0.13���0.15 mm at broadest place in comparison with 0.05 mm in basal third), longest anterodorsals and posterodorsals up to 0.18 mm long, ventral setae slightly longer than depth of tibia in middle part, much shorter distally. Tarsi of both fore and mid legs very thin and short setose, basitarsus of hind leg thin (0.05���0.06 mm), but distinctly slightly broader than 2 nd tarsal segment. Wing light brownish infuscated; stigma brown, long (0.60���0.65 mm), almost parallel-sided, 0.05 mm broad, slightly narrower than cell r 1 (about 0.07 mm at broadest point). Distance between tips of veins R 1 and R 2 + 3 about 0.20���0.25 mm. Squama dark brown with pale (holotype) or dark fringes, halter brown. Abdomen brown, darker (brownish-black) in dorsal view and lighter (greyish-brown) in lateral view, microtrichose, pale setose on sides of basal 2���3 segments (holotype) or only on sides of 2 nd segment, subsequent segments dark setose; posteromarginals on sides of tergites 2���4 longer than subsequent segments, on tergites 5���7 slightly shorter. Genitalia (Figs 20, 21) with hypandrial processes long and narrow, with several short setae on apical part; postgonites elongate-ovate in shape, with concave lower margin; left phallic hook long and simply bent, right one almost absent. Female. Abdominal setae shorter and paler than in male. Tergite 8 polished on basal part, sternite 8 polished, otherwise abdomen microtrichose. Length: body 2.1���2.9 mm, wing 2.4���2.9 mm. Remarks. Bicellaria koreana sp. nov. belongs to the B. sulcata complex of species as proposed by Chv��la (1991). It differs from the only other Asian representative of the complex (B. vana) according to characters given in the key. However, the species described above is very similar to Nearctic B. furcifer Melander, 1928 (Figs 18, 19). In fact we found only one reliable character distinguishing both species and this is the longer phallic hook in B. koreana sp. nov. (compare Figs 18 and 20)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 244-245, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria pilosa Lundbeck (Figs 27���28) Bicellaria pilosa Lundbeck, 1910: 27. Material examined (463, 29��). Czech Republic: 323, 25��, Krkono��e Mts, many localities (Pančava, Lučn�� bouda, Labsk�� bouda, ��psk�� mire, Lučn�� hora, Kotel, Čertův důl, Labsk�� louka), 1250���1500 m, vi.���viii., M. Bart��k or M. Chv��la; 23, 2��, B��lina (Choumek, Holibka), 50 �� 31���32 'N, 13 �� 49���51 'E, v.-vi. 1998, M. Bart��k; 93, 9��, Kosteln�� Lhota, 50 ��08'N, 15 ��02'E, 190 m, 12.v. 1988, M. Bart��k���(CULSP). Norway: 13, Hardangervidda Nat. Park, 1300 m, vii. 2001, J. Farkač (CULSP). Romania: 1 ��, Fagaras Mts., Bilea env., 13.vii. 2001, P. Heřman (CULSP). Russia: 13, 1��, Pummanki [= Zemljanoe], W. Hell��n; 1 ��, Kutsajoki river near Kandalak��a), W. Hell��n; 13, Vuoremi [= Vorjema], W. Hell��n���(NHMH). Diagnosis. Bicellaria pilosa is very similar to B. alpina (see notes under latter). Chv��la (1991) considered both to be identical, listing B. alpina Bezzi as a junior synonym of B. pilosa (but he had no specimens of B. alpina on hand). However, we remove this species from synonymy despite not finding any differences in the male genitalia (see also remarks under B. alpina). Distribution. Europe. Remarks. Tuomikoski (1955) provided a short redescription, illustration of the hypandrium, and differentiation from B. alpina. Collin (1961) also briefly redescribed the species and illustrated the genitalia. Chv��la (1983) provided a redescription, synonymy, and illustrations of the genitalia, face, antenna, and hind leg. The species was described from several localities in Denmark; however, according to Chv��la (1983), all syntypes from the Lundbeck collection were destroyed., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 265-266, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria setitibia sp. nov. (Figs 12, 17) Type material. HOLOTYPE ♂, Georgia, [in Cyrillic] East Georgia, Lagodekhi zapovednik (Natural Reserve), 26���28.vii. 1969, V. Kovalev (ZMMU). PARATYPES: 2 ♀, same data; 1 ♂, 1 ♀, same locality, 27.vii. 1969 (CULSP: 1 ♂, 1 ♀, ZMMU: 2 ♀). Russia: Republic of North Ossetia-Alania: 1 ♂, Kakadur, Vladikavkaz env., 27.vii. 1925, Kiritschenko (NHMH). Diagnosis. Halter blackish brown in both sexes; third antennal segment without elongate dorsal setae; both fore and mid tibiae with longer setae also on distal part; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes narrow, with rather long apical seta; only right phallic hook present, long and sharply bent, left one reduced; female tergites 5���8 and sternites 6���7 partly shining, sternite 8 polished. Etymology. The species is named after the long setose tibiae. Description. Male. Head brownish-black, dark brownish grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 15���17 facets in line of contiguity of eyes. One pair of rather long ocellar setae and additional pair of shorter setae on hind part of ocellar triangle. Occiput dark brownish black. Face slightly narrowed ventrally, dorsal part about 0.07 mm and ventral part about 0.05 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2���3 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest ventral setae about 0.15 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 14���16: 4��� 8: 18���21. Thorax brownish-black, dark brownish grey microtrichose, prescutellar area with very small lighter patch in front of scutellum. All thoracic setae black. Chaetotaxy: 7���9 broadly biserial long acrostichals (up to 0.15 mm in front of suture); 9���12 uniserial dorsocentrals as long as or slightly longer than acrostichals; about 16���19 setae in intrahumeral and posthumeral areas; 3���5 subequally long postpronotals; notopleuron with 4���5 unequally long setae arranged in irregular row; 1 supraalar- and 3 prealar; 1 long postalar; 2 pairs of scutellars. Legs including coxae blackish brown to brownish black, microtrichose and black setose, parts of trochanters and hind femur posteriorly shining. Fore femur with anteroventral setae one-third as long as depth of femur and posteroventral setae slightly shorter than depth of femur. Fore tibia with posteroventrals as long as (in apical third) or slightly longer than depth of tibia, ventral pilosity very short, row of anterodorsals 0.10���0.15 mm long (3���4 are slightly stronger than others), developed also on hind part of tibia and similar row of slightly longer (about 0.20 mm) posterodorsals. Mid femur with sparse row of anteroventrals scarcely half as long as depth of femur (in middle part) but longer both proximally and distally, with posteroventrals slightly longer than depth of femur. Mid tibia with 4���5 pairs of setae dorsally (about 0.20 mm long) present also on apical third of tibia, other (short) setae including ventral ones slightly longer than depth of tibia. Hind femur with 15���17 anterodorsal setae slightly longer than depth of femur (longer proximally), with row of 16���18 subequally long anteroventrals, posteroventrals 2 / 3 as long as depth of femur. Hind tibia slightly swollen in apical part (0.11 mm at broadest point in comparison with 0.07 mm in basal third), longest anterodorsals and posterodorsals up to 0.15 mm long, ventral setae slightly longer than depth of tibia in middle part, longer proximally and much shorter distally. Basitarsus of all legs thin and short setose. Wing light brownish infuscated; stigma brown, long (0.60���0.70 mm), almost parallel-sided and narrower (0.06���0.07 mm) than cell r 1 (0.09���0.10 mm at broadest point). Distance between tips of veins R 1 and R 2 + 3 about 0.35���0.40 mm. Squama dark brown with dark fringes, halter blackish brown. Abdomen blackish-brown, darker (brownish-black) in dorsal view and lighter (blackish-brown) in lateral view, microtrichose, entirely black setose, posteromarginals on sides of tergites much longer than segment (longest lateral as well as ventral setae up to 0.35 mm long). Genitalia (Figs 12, 17) similar to B. spuria and B. ingrata: hypandrial processes narrow, with rather long apical seta; postgonites elongate trapezoid in shape, only right phallic hook present, long and sharply bent, left one reduced. Female. Thoracic setae shorter and less numerous than in male (about 10 in intra- and posthumeral areas), abdominal setae shorter. Tergite 5 polished on basal half, tergites 6���8 shining except dorsoapical corner, lateral parts of sternites 6���7 partly polished, sternite 8 polished, otherwise abdomen microtrichose. Length: body 2.7���3.1 mm, wing 2.4���3.2 mm. Remarks. Bicellaria setitibia sp. nov. belongs to the B. spuria complex of species (characterised by at most 3 setae on apical part of palpus, hind basitarsus slender, fore tibia on apical third without long ventral pilosity and with posteroventrals at least as long as depth of tibia, third antennal segment without long dorsal seta, body entirely black setose and male hypandrial processes long and narrow and only a single phallic hook present). The genitalia are very similar to B. spuria (compare Bart��k & Kub��k (2013, figs 33, 34) with Figs 12 and 17). However, both species may be differentiated according to characters given in the key. The small number of specimens did not enable us to decide if the long right hook of the phallus is ���typical��� or only an exception (all studied specimens of B. spuria and B. ingrata had left phallic hook long and right one reduced)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 246-247, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria setipalpus sp. nov. (Figs 29���30) Type material. HOLOTYPE 3: Italy, Passo Rolle, 46 �� 13 'N, 11 �� 42 'E, 1700 m, 8.viii. 1988, larch wood, M. Bart��k (CULSP). PARATYPES: France: 1 ��, Pic Long, along brook, 42 �� 49 'N, 0��09'E, 1800 m, 9.vii. 1990; 1 ��, same locality, alpine meadow, 2200 m ���all M. Bart��k���(CULSP). Italy: 13, same data as holotype (CULSP); 13, Mte. Terminillo, 42 �� 26 'N, 13 ��07'E, 2000 m, 4.viii. 1988, meadow nr. wood, M. Bart��k (CULSP); 2 ��, S. Stefano di Cadore, fiuma Piave, viii. 1979; 1 ��, Camposilvano, TN, 10.ix. 1974; 13, Passo Monte, Croce Carnico, 23.viii. 1964 ���all G. Raffone���(MSNV); 13, 1��, S. Martino, spruce wood, 46 �� 11 'N, 11 �� 41 'E, 1500 m, M. Bart��k (CULSP). Diagnosis. Entirely black setose species; third antennal segment with dorsal setae; palpus with about 10 strong setae; male hypandrial processes broadened apically; three phallic hooks present (left one bifurcate and its anterior arm bent); postgonites asymmetrical. Description. Male. Head black, dark brownish grey microtrichose, black setose (including lower occiput). Eyes meeting on frons, facets on dorsal half much larger than on ventral half, about 16���17 facets in line of contiguity of eyes. One to two pairs of ocellar setae, posterior pair shorter. Occiput in middle hind part densely setose. Face nearly parallel-sided, about 0.06 mm broad in lower half and 0.07 mm broad in dorsal part, clypeus slightly paler grey microtrichose. Gena broad and lustrous (not easily visible if head collapsed). Palpus short, brown, with about 10 setae, some rather strong and more than 0.15 mm long. Labrum brownish black, lustrous. Antenna black, second segment long setose ventrally (longest ventral setae about 0.15 mm); third segment with 1��� 2 dorsal setae (as in Fig. 48); ratio of broad part of segment 3: narrow part of segment 3: stylus (in 0.01 mm scale) = 16���18: 7���8: 19���20. Thorax brownish black, slightly subshining, prescutellar area similarly coloured as rest of mesoscutum, scutellum slightly paler. All thoracic setae black including fringes on squama. Chaetotaxy: acrostichals broadly biserial (8���9 in each row) and long even in presutural area (about 0.15 mm in front but longer posteriorly); dorsocentrals slightly longer than corresponding acrostichals in postusutural area; about 10���12 setae in posthumeral and intrahumeral areas; 1 longer and 2���3 slightly shorter postpronotals; notopleura with 5���6 unequally long setae; 1 supraalar and 1���2 setae in prealar area (and 1���2 additional setae close to dorsocentral row); 1 long postalar; 2 pairs of scutellars. Legs including coxae black, microtrichose (only trochanters partly lustrous) and black setose. Fore femur in basal two-thirds with row of strong anteroventrals, half as long as depth of femur and complete row of posteroventrals slightly longer than depth of femur. Fore tibia with 3 posterodorsals 3 X as long as depth of tibia (0.20 mm long), posteroventrals slightly longer than depth of tibia. Mid femur with sparse row of anteroventrals about as long as depth of femur in basal half but shorter apically and with posteroventrals slightly longer than depth of femur. Mid tibia with unequally long ventral setae, longest up to 2 X as long as depth of tibia, with 2���3 pairs of dorsal setae up to 0.25 mm long (longest about 3 X as long as diameter of tibia). Hind femur with 16���18 dorsal and 13���16 anteroventral setae 1.5 X as long as depth of femur, and with complete row of fine and much shorter posteroventrals. Hind tibia slightly swollen in apical half (0.16 mm at broadest point), subequally broad as hind femur, ventral setae about as long as depth of tibia (in basal half and shorter in apical third), 3 antero- and 5 posterodorsal setae up to 0.25 mm long. Tarsi of all legs thin and short setose. Wing brownish infuscated, stigma brown and 0.10 mm wide. Squama dark blackish-brown with black fringes, halter brown. Abdomen black, dull brownish-black in dorsal view and lighter brownish grey microtrichose in lateral view, very long black setose (marginals up to 0.50 mm long). Genitalia (Figs 29���30): hypandrial processes long, broadened apically and somewhat truncate; postgonites asymmetrical: right one with ventral extension arising from caudal part of median extension; three long phallic hooks present (left hook bifurcate with anterior arm sharply bent). Female. Similar to male. Abdominal tegites 6���8 lustrous on basal third, sternite 8 lustrous. Length: body 2.0���4.0 mm, wing 3.0��� 3.6 mm. Etymology. The species is named after the strongly setose palpus. Distribution. France, Italy. Remarks. The species described above is a member of the B. pilosa complex of species (characterized under B. alpina) comprising three European species (B. pilosa, B. alpina and B. setipalpus). Bicellaria setipalpus may be distinguished from both other species of this complex by narrower face, strongly bent anterior arm of the left phallic hook and asymmetrical postgonites. If numerous setae on the palpus are overlooked, the species is superficially very similar to B. austriaca; however, genitalia of both species are strikingly different., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 266-267, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria woodi sp. nov. (Fig. 26) Type material. HOLOTYPE ♂, Japan, Hokkaido, Daisetsuzan Nt. Pk., Tenninkyo, 800 m, 10���14.vii. 1986, D.M. Wood (CNC). PARATYPES: Japan: 2 ♀, same data as holotype (CNC, CULSP). Diagnosis. All setae black; third antennal segment without elongate dorsal setae; 1���2 setae on each palpus; hind basitarsus very narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes long and broad, strip-like. Hind tibia with two long and strong preapical (ventral and anteroventral) setae. Etymology. The species is named in honour of collector of type series specimens, Monty D. Wood (Ottawa). Description. Male. Head blackish-brown, brownish-grey microtrichose, black setose (including lower occiput). Holoptic, facets in dorsal half much larger than in ventral half, about 15���18 facets in line of contiguity of eyes. One pair of ocellar setae (lost in all specimens). Occiput rather densely and not very long coarse setose. Face slightly narrowing ventrally, dorsal part about 0.07 mm broad and ventral 0.05 mm. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 1���2 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest setae almost 0.12 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 15���16: 10���11: 19. Thorax blackish-brown, microtrichose, mesoscutum dark brownish black in dorsal view, prescutellar area evenly dark. All thoracic setae nearly completely rubbed out, apparently all black and coarse. Chaetotaxy: 8���9 biserial acrostichals; 8���10 uniserial dorsocentrals; some 10 setae in posthumeral and intrahumeral areas; 4���5 postpronotals; notopleuron with 7���8 setae arranged in irregular row (strong and long in front part of notopleuron); 1���2 supraalar- and 3���4 prealar; 1 postalar; at least 3 pairs of long scutellars (one preserved in one female is nearly 0.60 mm long). Legs including coxae black, microtrichose and black setose. Both forelegs in male missing (see female description). Hind coxa with long upright standing setae. Mid femur with sparse row of very short anteroventrals and with much stronger and longer posteroventrals subequally long as depth of femur. Mid tibia with 2 anterodorsal (one subbasal and one submedian) and 1 submedian posterodorsal setae 3 X as long as depth of tibia (longest about 0.30 mm) and 2 (anterior and anteroventral) conspicuously strong but rather short preapical setae, ventral setae slightly longer than depth of tibia. Hind femur with about 20 anterodorsal and 19 anteroventral setae subequally as long as depth of femur, posteroventrals slightly shorter than anteroventrals. Hind tibia slightly swollen in apical part (0.13 mm at broadest point in comparison with 0.08 mm in basal part) much narrower than hind femur (0.19 mm at broadest point), both dorsal and ventral setae slightly longer than depth of tibia, 4���5 pairs of dorsal setae, longest posterodorsal 0.32 mm long, two strong and long preapical setae in anterior and anteroventral positions (Fig. 26). Tarsi of all legs very thin and short setose. Wing brownish infuscated, fore margin dark brown, especially between veins C and R 1; stigma very long (1.00 mm), almost parallel-sided and narrow (0.05 mm), narrower than cell r 1 (about 0.08 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.20 mm. Squama dark brown with black fringes, halter brown. Abdomen brown, microtrichose, entirely black setose, setae long, even those on 1 st sternite unusually long. Genitalia (undissected) with hypandrial processes long and broad, strip-like (as in B. shatalkini sp. nov.). Female. Fore coxa densely setose. Fore femur with row of fine anteroventrals half as long as depth of femur, with similar row of slightly longer posteroventrals. Fore tibia with posteroventrals distinctly longer than depth of tibia and without ventral pilosity (present as usual on fore basitarsus), with 1���3 setae dorsally about twice as long as depth of tibia. Tergites entirely pruinose including visible parts of tergites 6���7, sternite 8 polished. Length: body 3.6���3.9 mm, wing 3.4���3.6 mm. Remarks. All specimens of type series are rather damaged, devoid of most mesoscutal setae and most abdominal setae; however, their arrangement, number and length may be judged from their articulations and from several setae present on some places on some specimens. The new species somewhat resembles B. farkaci sp. nov. in general appearance but the palpus has only 1���2 setae, female sternite 8 is polished and genitalia quite different. Bicellaria woodi sp. nov. is easily recognizable in both sexes and unique in the World fauna due to two strong preapical anterior (and anteroventral) setae on the hind tibia. We decided not to damage the only male (holotype) by genital dissection, so are unable to present a full description of the genitalia., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 252-253, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria intermedia Lundbeck (Figs 13���14) Bicellaria intermedia Lundbeck, 1910: 25. Material examined (223). Czech Republic: 33, ��umava Mts, Pr����ily, nr. Křemeln�� brook, 49 ��08'N, 13 �� 23 'E, 810 m, 8.vii. 1994; 23, ��umava Mts, Jezern�� slať, spruce wood, 49 ��02'N, 13 �� 34 'E, 980 m, 6���7.viii. 1996; 13, ��umava Mts, Nov�� Hůrka, spruce forest, 49 ��09' 21 ''N, 13 �� 20 '00''E, 880 m, 24.vi.- 28.vii. 2000, Malaise trap; 13, Ka��persk�� hory, nr. Losenice river, 49 ��08'N, 13 �� 33 'E, 600 m, 1.vii. 1995; 13, Kunice, Vrchy, edge of mixed wood, 49 �� 56 'N, 14 �� 42 'E, 450 m, 11.vi. 1983; 13, Kunice, 27.vii. 1975; 13, Vr��ž nr. P��sek, 49 �� 23 'N, 14 ��08'E, 400 m, 30.v. 1994; 23, N��dek, 49 �� 39 'N, 18 �� 46 'E, 2.viii. 1981, 550 m���all M. Bart��k���(CULSP); 13, ��umava Mts, Horsk�� Kvilda, spruce forest, 49 ��05'N, 13 �� 33 'E, 1130 m, 21.vii.- 21.viii. 1999, Malaise trap; 13, Podyj�� NP, Terasy, mixed wood, 48 �� 53 ' 22 ''N, 15 �� 50 ' 18 ''E, 460 m, 2 vi.- 2.vii. 2003, Malaise trap; 13, Podyj�� NP, Li��č�� sk��la, Quercetum, 48 �� 49 ' 52 ''N, 15 �� 56 ' 35 ''E, 410 m, 2.vii.- 4.viii. 2004, Malaise trap���all M. Bart��k et ��. Kub��k���CULSP); 13, Česk�� Meziř��č��, Zbytka reserve, 50 �� 17 'N, 16 ��04'E, 250 m, 28.vi. 1996, B. Mocek (MHK). Italy: 13, San Vigilio di Marebbe, 13.vii. 1989, G. Raffone (MSNV). Russia: 13, Salla (probably collected in Russian part of county), R. Tuomikoski (NHMH); 13, Irkutskaya oblast, Bajkal port, 24.vi. 1965, Negrobov (ZMMU). Slovak Republic: 23, Vysok�� Tatry, Star�� Smokovec, meadow, 49 ��09'N, 20 �� 14 'E, 1400 m, 22���23.vii. 1986; 13, Vysok�� Tatry, Hrebienok, clearing in forest, 49 �� 10 'N, 20 �� 14 'E, 1250 m, 24.vii. 1986 ���all M. Bart��k���(CULSP). Diagnosis. The species is a member of the B. austriaca complex (see under this species). Males of all four species of this complex (B. italica, B. austriaca, B. intermedia, B. collini) may be easily identified according to the key due to different genitalia. Males of B. intermedia may be easily recognized according to two long and narrow, irregularly bent phallic hooks [accurately illustrated by both Collin (1961) and Chv��la (1983)]. We found no reliable character enabling to differentiate females from B. italica. Distribution. Eurosiberian species. Remarks. The species was originally described from many localities in Denmark. Tuomikoski (1955) provided a short redescription and illustration of the hypandrium. Collin (1961) provided a short redescription and illustrations of the hind leg, head, antenna and genitalia, in addition to notes to its identity (comparison with type specimens). Chv��la (1983) also redescribed this species, provided illustrations of the face, antenna, hind leg, genitalia (incl. variations in the shape of hypandrium), and designated a lectotype., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 259-260, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria sulcata (Zetterstedt) (Figs 39���40) Cyrtoma sulcata Zetterstedt, 1842: 331. Bicellaria sulcata vanella Tuomikoski, 1936: 82. Material examined (273, 24��). Austria: 13, Fels��l��v�� (= Obersch��tzen), 24.v. 1909 (SDEI). Czech Republic: 13, Božejovice-Jezviny, 4.vi. 1983, M.Chv��la (OUMNH); 13, Žehuňsk�� obora, Kopič��ck�� rybn��k, 28.iv. 2004; 23, Žď��rsk�� vrchy Mts, Radost��n, peat-bog, 16.v. 1995; 1 ��, B��leč nad Orlic��, damp meadow, 25.v. 1995 ���all B. Mocek��� (MHK); 13, B��leč nad Orlic��, nr. river, 28.iv. 1995; 13, Vr��ž nr. P��sek, damp meadow, 49 �� 23 'N, 14 ��08'E, 12.v. 1991; 1 ��, B��lina, Holibka, lowland wood, 50 �� 31 ' 19 ''N, 13 �� 49 ' 47 ''E, 360 m, 9.viii.- 23.ix. 1998; 43, Beskydy Mts, Muřinkov�� vrch, peat-bog, 49 �� 31 'N, 18 �� 39 'E, 950 m, 13.vi. 1987; 13, ��umava Mts., Lib��nsk�� sedlo, 48 �� 59 'N, 13 �� 59 'E, 800 m, 9.v. 1987; 13, same locality, 17.v. 1997; 13, 2��, ��umava Mts, Sp��lenec, damp meadow, 48 �� 56 'N, 13 �� 57 'E, 800 m, 21.v. 1992; 5 ��, ��umava Mts, Nov�� Hůrka, peat-bog, 49 ��09'N, 13 �� 20 'E, 870 m, 15.vi. 1999; 13, ��umava Mts, Kyselovsk�� les, Alnetum, 48 �� 42 'N, 14 ��04'E, 725 m, 17.v. 1997; 1 ��, same locality, 16.vi. 1997; 1 ��, ��umava Mts, Chalupsk�� slať, damp meadow, 49 ��01'N, 13 �� 39 'E, 860 m, 4.vi. 1996; 1 ��, same locality, peat-bog, 10.vi. 1989 ���all M. Bart��k���CULSP). Croatia: 13, 1��, Jasenak, 30.v. 1905 (?), K. Kert��sz (CULSP). Finland: 13, Joutseno, 30.v. 1962, E. Thuneberg; 23, Utsjoki, R. Frey; 13, Malla, R. Frey���(NHMH); 1 ��, Esbo, Kolmpera, 6.vii. 1980, M. Chv��la (OUMNH). Germany: 1 ��, Bayerischer Wald, Spiegelau, 48 �� 57 'N, 13 �� 22 'E, 760 m, 8.vi. 1995, M. Bart��k (CULSP); 13, Berlin, Jungfhd., v. 97; 13, 1��, Berlin, Schildhorn; 1 ��, Mecklenburg, B��tzov, 23.v. (?), C. F. Kettel; 1 ��, same loc., 16.v. ���(SDEI). Russia: 4 ��, Moscow prov.: Serpuchov, mixed wood, 54 �� 52 'N, 37 �� 36 'E, 150 m, 27.v. 1989; 13, Moscow, Losiny ostrov, park vegetation, 55 �� 49 'N, 37 �� 45 'E, 180 m, 26.v. 1989; 13, Abramcevo, along brook, 56 �� 14 'N, 37 �� 59 'E, 200 m, 28.v. 1989 ���all M. Bart��k���(CULSP). Slovakia: 1 ��, V. Fatra, Kantorsk�� dol. (= valley), 29.vi. 1975, M. Bart��k (OUMNH); 13, Vysok�� Tatry, Doln�� Smokovec, 28.v. 1968, M.Chv��la���all ��� B. sulcata ��� det. Chv��la���(OUMNH). Sweden: 1 ��, Lk. Tarraure, 19.vii. 1962, A.C. Pont; 13, Sala Distr., 1km E Broddbo, 18.vi. 1986, A.C. Pont���(OUMNH). Diagnosis. Males may be easily identified according to the key (but see Bicellaria sp. 3 in ���Unrecognized specimens��� section); differences in genitalia of two allied species, B. sulcata and B. vana, are discussed under the latter species. The left phallic hook is long and broad, right one strongly reduced. Females may be identified according to characters given in the key. Distribution. A broadly distributed West Palaearctic species. Record from East Siberia (Tuomikoski 1955) needs verification. Remarks. Tuomikoski (1955) briefly redescribed this species, illustrated the hypandrium and synonymized it with his B. sulcata vanella. Collin (1961) provided a short redescription and illustrated the male genitalia. Chv��la (1983) also redescribed this species, illustrated the face, antenna, hind leg, and genitalia, discussed the synonymy, and designated the lectotype., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on page 270, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria uvens Melander Bicellaria uvens Melander, 1928: 74. Bicellaria bisetosa Tuomikoski, 1936: 84. Bicellaria montana Kato, 1971: 283, syn. nov. Distribution, diagnosis. See Bart��k and Kub��k (2013). Additional material examined. Japan: 1 ♂ (dissected, ��� Bicellaria montana / uvens, det. B.J. Sinclair 2012), Hokkaido, Daisetzusan Aizankei, 17.vii. 1986, D.M. Wood, Malaise trap (CNC); Russia: 1 ♂, Magadanskaya oblast (= region), Aborigen Field Stn., 500 m, 25.vii.��� 10.viii. 1990, D.M. Wood (CNC). Remarks. Kato (1971) described B. montana from Japan, illustrating the hypandrial processes and hind leg. We found on examination that his species is identical with B. uvens (= B. bisetosa Tuomikoski) and while we have not been able to examine type material we here propose formally that B. montana Kato, 1971 is a junior synonym of B. uvens Melander 1928. This proposal was supported by T. Saigusa (pers. comm. to B. Sinclair, 2013) who compared illustrations in Bart��k and Kub��k (2013) with one dissected paratype of B. montana and stated they are almost identical. He also provided photographs of genitalia which were identical to our specimens of B. uvens., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 251, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria kovalevi sp. nov. (Figs 15, 16) Type material. HOLOTYPE ♂, Georgia, [printed in Cyrillic] West Georgia, Borzhomskiy Zapovednik, [= Borzhomskiy Nature Reserve] 8.viii. 1969, V. Kovalev (ZMMU). PARATYPES: Georgia: 2 ♂, same data as the holotype; 1 ♂, ♀, same locality, 7.viii. 1969, V. Kovalev���(ZMMU); 3 ♂, 1 ♀, same data (CULSP); 1 ♂, West Georgia, Bakuriani, 18.vii. 1969, V. Kovalev; 2 ♀, East Georgia, Lagodekhi Zapovednik, [= Lagodekhskiy Nature Reserve] 25.vii. 1969, V. Kovalev���(ZMMU); 1 ♀, same locality, 27.��� 28.vii. 1969 (CULSP); 1 ♂, same locality, 26��� 28.vii. 1969; 1 ♂, Adzharia, Kintrishi Zapovednik, okr. [= Kintrishskiy Nature Reserve] env. Kobuleti, 13.v. 1971, V. Kovalev���(ZMMU); 1 ♂, same data (CULSP). Diagnosis. Halter blackish brown in both sexes; third antennal segment without elongate dorsal setae; all body setae black; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes long and narrow, abruptly axe-like broadened at apex with rather long seta in the middle of broadened part; both phallic hooks present. Etymology. The species is named in honour of the late Vladimir Kovalev, collector of the type series and well known Russian dipterologist. Description. Male. Head brownish-black, dark brownish grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 13���15 facets in line of contiguity of eyes. One pair of rather long ocellar setae. Occiput dark grey. Face slightly narrowed ventrally, dorsal part about 0.05 mm and ventral part about 0.04 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2���3 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest ventral setae about 0.10 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 10���13: 4���6: 17���19. Thorax brownish-black, dark brownish grey microtrichose, prescutellar area evenly dark or with only very small lighter patch in front of scutellum. All thoracic setae black. Chaetotaxy: 5���9 broadly biserial long acrostichals (up to 0.15 mm in front of suture); 8���9 uniserial dorsocentrals as long as or slightly longer than acrostichals; about 5���9 setae in intrahumeral and posthumeral areas; 3 almost subequally long postpronotals; notopleuron with 4���5 setae arranged in irregular row (1���2 setae often longer and stronger than others); 1 supraalar- and 2���3 prealar; 1 long postalar; 2 pairs of scutellars. Coxae blackish brown, microtrichose, black setose, fore and mid ones with rather strong upright anterior setae. Legs brown, microtrichose and black setose. Fore femur with both antero- and posteroventral setae slightly shorter than depth of femur. Fore tibia with setae about as long as depth of tibia or slightly longer including posteroventrals, 1���3 posterodorsals up to twice as long as depth of tibia, ventral pilosity not developed. Mid femur with sparse row of anteroventrals scarcely onethird as long as depth of femur (in middle part), with much longer posteroventrals as long as or slightly longer than depth of femur. Mid tibia with 2���3 antero- and posterodorsal setae in basal half (up to 0.20 mm long), other short setae slightly longer than depth of tibia. Hind femur with 14���16 anterodorsal setae slightly longer than depth of femur, with row of 14���18 anteroventrals slightly shorter than anterodorsals, posteroventrals half as long as depth of femur. Hind tibia slightly swollen in apical part (0.13 mm at broadest point in comparison with 0.07 mm in basal third), longest anterodorsals and posterodorsals up to 0.20 mm long, ventral setae slightly longer than depth of tibia in middle part, longer proximally and much shorter distally. Tarsi of both fore and mid legs thin and short setose, basitarsus of hind leg thin (0.06���0.07 mm), slightly broader than 2 nd segment. Wing light brownish infuscated; stigma brown, long (0.60���0.70 mm), almost parallel-sided and subequally broad as cell r 1 (about 0.08 mm at broadest point). Distance between tips of veins R 1 and R 2 + 3 about 0.20���0.25 mm. Squama dark brown with dark fringes, halter blackish brown. Abdomen blackish-brown, darker (brownish-black) in dorsal view and lighter (greyish-brown) in lateral view, microtrichose, entirely black setose, posteromarginals on sides of tergites much longer than segments (longest lateral and ventral setae up to 0.40 mm long). Genitalia (Figs 15, 16) with hypandrial processes very long and axe-like broadened apically, with single long seta in middle of broadened apex; postgonites elongate-ovate in shape, median extension of postgonites slightly saw-toothed ventrally; left phallic hook long, right one slightly shorter. Female prescutellar area often lighter than rest of mesoscutum; abdominal setae shorter than in male. Tergite 8 polished at least on basal half, sternite 8 polished, otherwise abdomen microtrichose. Length: body 2.2���2.7 mm, wing 2.3���2.7 mm. Remarks. Bicellaria kovalevi sp. nov. is very similar to B. spuria. Differential characters are given in the key. Most striking differences are in the shape of the male genitalia: hypandrial processes are apically broadened in B. kovalevi sp. nov. but narrow in B. spuria and median extension of postgonites is saw-toothed in B. kovalevi sp. nov. and smooth in B. spuria. Moreover, right phallic hook is much longer in above described species than in B. spuria. Females are difficult to differentiate. Based on materials at disposal it seems that B. kovalevi sp. nov. has smaller broad part of 3 rd antennal segment and hind basitarsus distinctly slightly broader than 2 nd hind tarsomere (in B. spuria broad part of 3 rd antennal segment usually longer than 0.14 mm and hind basitarsus equally thick as second hind tarsomere)., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 245-246, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria shatalkini sp. nov. (Figs 22, 23) Type material. HOLOTYPE ♂, Russia, Magadan: r. [=river] Indigirka, ust [= river mouth] Injali, 23.vi. 1976, listvennik s ernikom i mkhom na sklone sopki [= larch forest with black crowberry and mosses on slope of a hill], V. Kovalev (ZMMU). PARATYPES: Russia: 7 ♂, 3 ♀, same data as holotype; Chita: 2 ♂, 1 ♀, Undurga River, 9.vii. 1977, V. Kovalev; 1 ♂, 1 ♀, same locality, 13.vii. 1977, V. Kovalev (ZMMU: 9 ♂, 3 ♀, CULSP: 1 ♂, 2 ♀). Diagnosis. Halter blackish brown in both sexes; third antennal segment without elongate dorsal setae; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes long and broad, postgonites triangle-shaped and saw-toothed ventrally; both phallic hooks present, right one slightly shorter than left; female tergite 8 entirely microtrichose. Etymology. The species is named in honour of Prof. Anatoliy Shatalkin, well known Russian dipterologist (Moscow). Description. Male. Head brownish-black, dark brownish grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 14���16 facets in line of contiguity of eyes. One pair of ocellar setae (sometimes an additional shorter pair present). Occiput brownish black. Face slightly narrowed ventrally, dorsal part about 0.05 mm and ventral part about 0.04 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest ventral setae about 0.09 mm); 3 rd without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 12���14: 4���5: 17���19. Thorax brownish-black, brownish grey microtrichose, prescutellar area with broad lighter spot in front of scutellum reaching hind dorsocentrals and acrostichals (in some specimens divided by narrow dark midline). All thoracic setae black. Chaetotaxy: 8���11 broadly biserial acrostichals (about 0.12 mm long in front of suture); 9���11 irregularly uniserial dorsocentrals as long as or slightly longer than acrostichals; 10���15 setae in intrahumeral and posthumeral areas; 1 longer and 2���3 shorter postpronotals; notopleuron with 4���6 unequally long setae arranged in irregular row; 1���2 supraalar-, 0���2 prealar and 1���4 additional setae between supraalar- and dorsocentrals (situated laterad of dorsocentral row); 1 long postalar; 2 pairs of scutellars. Legs including coxae blackish brown, microtrichose, black setose. Fore femur with both antero- and posteroventral setae half as long as depth of femur and similar row of slightly longer posterodorsals. Fore tibia with irregular rows of unequally long antero- and posterodorsals, longest about twice as long as depth of tibia (up to 0.15 mm long), posteroventrals about as long as depth of tibia, ventral pilosity not developed or very short. Mid femur with sparse row of anteroventrals less than half as long as depth of femur (in middle part), with posteroventrals up to as long as depth of femur. Mid tibia usually with 2 anterodorsals in basal half and 3 posterodorsal setae in basal 2 /3, 2��� 3 X as long as depth of tibia, other short setae slightly longer than depth of tibia. Hind femur with 17���19 anterodorsal setae slightly longer than depth of femur (in basal part, shorter in apical part), with row of 12���18 anteroventrals slightly shorter than anterodorsals, posteroventrals about half as long as corresponding anteroventrals. Hind tibia swollen in apical part (0.13���0.14 mm at broadest point in comparison with 0.07���0.08 mm in basal third), longest anterodorsals and posterodorsals up to 0.15 mm long, ventral setae subequally as long as depth of tibia in middle part, longer proximally and much shorter distally. Tarsi of both fore and mid legs thin and short setose, basitarsus of hind leg thin (0.06 mm), at most slightly broader than 2 nd segment. Wing brownish infuscated; stigma brown, long (0.55���0.60 mm), almost parallel-sided (about 0.08 mm at broadest point) and slightly broader than narrowest part of cell r 1. Distance between tips of veins R 1 and R 2 + 3 about 0.20���0.25 mm. Squama dark brown with dark fringes, halter brownish black. Abdomen blackish-brown, slightly darker (brownish-black) in dorsal view than in lateral view, microtrichose, entirely black setose, posteromarginals on sides of tergites much longer than segment. Genitalia (Figs 22, 23) with hypandrial processes long and broad (longer than 0.15 mm), with very short setae on apical third; postgonites triangle-shaped, sawtoothed ventrally; both phallic hooks present, right one slightly shorter than left. Female. Setae shorter (especially on abdomen) than in male, and less numerous, narrower hind tibia (0.11 mm in broadest part). Tergites entirely microtrichose including 8 th one, sternite 8 polished. Length: body 2.7���3.1 mm, wing 2.5 ���3.0 mm. Remarks. Bicellaria shatalkini sp. nov. probably belongs to the B. uvens complex of species; and differs from other species of this complex by its long hypandrial processes., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 248-249, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria amankutanensis sp. nov. (Figs 1, 2) Type material. HOLOTYPE ♂, Uzbekistan, Amankutan, pasture, 1300 m, 39.19 N, 66.55 E, 23.v. 1989, M. Bart��k (CULSP). PARATYPES: 28 ♂, 33 ♀, same data as the holotype (CULSP). Diagnosis. Halter yellow in both sexes; third antennal segment without elongate dorsal setae; mesoscutal setae black and abdominal setae mostly pale; hind basitarsus narrow and hind tibia swollen in both sexes. Male hypandrial processes moderately broad; only one (left) long phallic hook present; female tergites 5���8 and sternites 6���8 polished. Etymology. The species is named after the type locality (Amankutan, also Aman-Kutan in Uzbekistan). Description. Male. Head brownish-black, grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 14���16 facets in line of contiguity of eyes. One pair of rather long black ocellar setae. Occiput dark grey, black setose. Face slightly narrowed ventrally, ventral part about 0.03���0.04 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2���3 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest setae about 0.10 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: stylus (at 0.01 mm resolution) = 16���19: 5���7: 16���20. Thorax brownish-black, grey microtrichose, mesoscutum brownish grey microtrichose in dorsal view, prescutellar area differently microtrichose than other parts of mesoscutum, appearing light grey. All thoracic setae black, except on hind coxa. Chaetotaxy: 6���8 broadly biserial long acrostichals (up to 0.15 mm in front of suture); 8���11 uniserial dorsocentrals as long as or slightly longer than acrostichals; about 11��� 16 setae evenly spread in intrahumeral and posthumeral areas; 1 long and 1���3 somewhat shorter postpronotals; notopleuron with 3���5 setae arranged in one irregular row (one of them often conspicuously longer and stronger than other); 1 supraalar and 1���2 prealar; 1 long postalar; 2 pairs of scutellars. Legs including coxae blackish brown, microtrichose and black setose, hind coxa and trochanter (and in some specimens also base of hind femur) with some pale setae. Fore femur with short and sparse anteroventrals one-third as long as depth of femur and with row of slightly longer posteroventrals. Fore tibia with row of posterodorsals up to 2 X longer than depth of tibia, otherwise short setose, posteroventrals in apical third slightly shorter than depth of tibia, ventral pilosity absent or very short. Mid femur with sparse row of anteroventrals scarcely one-third as long as depth of femur and with much longer posteroventrals as long as depth of femur. Mid tibia with irregularly arranged setae dorsally up to 2 X long as depth of tibia (0���3 in each anterodorsal and posterodorsal rows), otherwise short setose. Hind femur with 15���18 anterodorsal setae on basal third up to 2 X long as depth of femur but shorter apically and with row of 16���20 anteroventrals slightly longer than depth of femur at about middle and shorter both basally and apically; posteroventrals short, scarcely 0.5 X long as depth of femur. Hind tibia swollen in apical part (0.14���0.17 mm at broadest point in comparison with 0.06���0.07 mm in basal part), anterodorsals and posterodorsals up to 0.20 mm long, ventral setae short. Tarsi of all legs thin, short setose, basitarsus of hind leg very slightly swollen (0.08 mm wide). Wing slightly brownish infuscated; stigma brown, long (0.60 mm), almost parallel-sided and narrow (0.06��� 0.07 mm), slightly narrower than cell r 1 (about 0.08���0.09 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.25 mm. Squama light brownish with dark fringes, halter clear yellow. Abdomen blackish-brown in ground colour, dark blackish brown in dorsal view and pale grey in lateral view, microtrichose, mostly pale setose, dorsally with short and partly black setae; posteromarginals on sides of tergites 2���5 much longer than corresponding segments, on last tergites subequally as long as corresponding segments. Genitalia (Figs 1, 2) with hypandrial processes moderately long and broad, postgonites elongate-trapezoid, slightly asymmetrical (right one slightly longer and broader at apex); left phallic hook long, right one strongly reduced. Female. Mesoscutal setae less numerous and hind tibia slightly narrower than male. Tergites 5���8 polished, sternites 6���8 polished, sternite 5 densely microtrichose (with exception of narrowly polished lateral parts), otherwise abdomen faintly microtrichose. Length: body 2.5���3.1 mm, wing 2.5���2.9 mm. Remarks. Bicellaria amankutanensis sp. nov. is very similar to B. chimganensis sp. nov. Differential characters are given in the Remarks following the description of the latter species. Both species are rather similar to Nearctic B. halteralis (Loew, 1862), however, the latter species has much narrower face and very long terminal seta on the hypandrial processes., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 235-236, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria alpina Bezzi (Figs 3 ���4, 48) Bicellaria alpina Bezzi, 1918: 75. Type material examined. Val Bitto, 4.viii.1899, 13 (labelled: ��� B. alpina ��� det. Bezzi - probably a syntype, SDEI). Additional material examined (43, 6��). Italy: 13, Cusiano(?), viii. 1895; 2 ��, Val Genova, vii.(?) 1899; 1 ��, same loc. 4.viii. 1899; 1 ��, San Martino, 30.vii. 1914; 13, Venino, 11.vii. 1902 (��� B. alpina ��� det. Bezzi) (SDEI); 1 ��, Passo Sella, alpine meadow, 8.viii.1988, 46�� 26 'N, 11 �� 43 'E, 2100 m, M. Bart��k (CULSP). Switzerland: 13, Lenzerheide, pasture, 2000 m, 14���21.vii. 2000, Malaise trap, B. Merz; 13, same locality, 21���31.vii. 2000, B. Merz (CULSP); 1 ��, Maloja, 23.vii. 1906? (SDEI). Diagnosis. Bicellaria alpina is a member of the B. pilosa complex of species (characterized by 4 or more black setae on palpus, 1���3 dorsal setae on third antennal segment, all body setae black and bifurcate left phallic hook), the third species of this complex (B. setipalpus sp nov.) differs from both remaining species in the structure of the genitalia (sharply bent anterior arm of left phallic hook). Chv��la (1991) considered B. alpina and B. pilosa Lundbeck to be identical, listing B. alpina as a junior synonym of B. pilosa (although he had no specimens of B. alpina on hand). However, we remove this species from synonymy despite not finding any differences in the male genitalia (the presence or absence of a long seta on the base of the hypandrial processes is apparently a variable character in all three species of the B. pilosa complex). Bicellaria alpina may be easily recognised from B. pilosa according to the key. Distribution. Mountains of Central and Southern Europe. Remarks. Tuomikoski (1955) provided a short redescription and illustration of the hypandrium. For further remarks see under B. pilosa., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 253-254, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Key to Bicellaria of Asia Note. When determining the identity of Asian species of Bicellaria it is important to also consult the key of Bart��k and Kub��k (2013) to European species. The distribution of most species in the Asian part of the Palaearctic remains largely unknown and many ���European��� species in fact have broader distribution [for example, the surprising finding of B. longisetosa in the Amur region and B. intermedia in the Bajkal area as reported in Bart��k & Kub��k (2013)]. It is likely that other species presently known only from Europe will eventually be found in Asia. 1 Palpus with 4 or more black setae........................................................................ 2 - Palpus with 1���3 setae (those occurring on the palpifer are not counted)........................................... 5 2 (1) Halter yellow. Mesoscutum with stripes. Hind femur spinose. Male phallic hooks curiously shaped (Fig. 7) (Israel).............................................................................................. B. femorata Collin - Halter brown to blackish-brown. Mesoscutum without stripes. Hind femur with only ordinary setae. Male phallic hooks differ- ent................................................................................................ 3 3 (2) Third antennal segment without long dorsal setae. Male without hypandrial processes (Fig. 6). Female with pruinose visible part of 8 th sternite (China: Yunnan)......................................................... B. farkaci sp. nov. - Third antennal segment with long dorsal seta(e) (if without this seta(e), follow couplet 5). Male with hypandrial processes. Female with visible part of 8 th sternite polished............................................................. 4 4 (3) Second antennal segment without conspicuously long lateral seta. Altogether 6���7 setae in intra- and posthumeral areas. Male hypandrial processes very narrow at apex, long setose even on base (see Kato 1971, fig. 4 A) (Japan)...... B. japonica Kato - Second antennal segment with at least 1 very long lateral or lateroventral seta 0.2���0.3 mm long. More than 10 setae in intra- and posthumeral areas. Male hypandrial processes broad at apex, short setose on base (see Bart��k & Kub��k 2013, fig. 20) (Europe, East Palaearctic).............................................................. B. longisetosa Chv��la 5 (1) Hind basitarsus distinctly swollen, more than 2 X as broad as hind tarsomere 3. Face very narrow (0.02 mm, about as 1���2 fac- ets). Fore tibia without posteroventrals on apical third in exactly posterior view (Japan)................. B. flavipes Kato - Hind basitarsus not swollen, less than 2 X as broad as hind tarsomere 3. Other characters different (except B. vana and B. kore- ana sp. nov.)......................................................................................... 6 6 (5) Third antennal segment with 1 or more dorsal seta(e) (Europe, Siberia)....................... B. intermedia Lundbeck - Third antennal segment without long seta(e) dorsally........................................................ 7 7 (6) Fore tibia with ventral pilosity on apical third at most slightly shorter than on fore basitarsus, posteroventral setae at most as long as pilosity and usually shorter than depth of tibia (observe in exactly posterior view)............................ 8 - Fore tibia with ventral pilosity on apical third at most 1 / 3 as long as on fore basitarsus and/or posteroventral setae at least 2 X as long as pilosity and usually longer than depth of tibia......................................................... 9 8 (7) Body entirely black setose. Male phallic hook slightly bent (see Bart��k & Kub��k 2013, fig. 43)............ B. vana Collin - At least first two abdominal segments pale setose on sides. Male phallic hook strongly bent (Fig. 20) (East Palaearctic)......................................................................................... B. koreana sp. nov. 9 (7) Abdomen partly pale setose (at least on sides of tergite 2)..................................................... 10 - Abdomen entirely black setose......................................................................... 13 10 (9) Halter brown to yellowish-brown. Legs polished. Male: two phallic hooks present, hoe-like broadened (best visible from behind���see Bart��k & Kub��k 2013, fig. 35) (Palaearctic species?)......................... B. stackelbergi Tuomikoski - Halter yellow. Legs microtrichose. Male: 1���2 hook(s) of phallus present, if two, then not broadened................... 11 11 (10) Hind tibia very slightly swollen. Male left epandrial lamella large, almost globular (Fig. 11), hypandrial processes bifurcate apically (Fig. 10). Female 5 th tergite microtrichose (Uzbekistan)............................. B. globulicauda sp. nov. - Hind tibia distinctly swollen. Male left epandrial lamella triangle-shaped as usual, not conspicuously large, hypandrial pro- cesses simple. Female 5 th tergite polished................................................................. 12 12 (11) Setae on mesoscutum all pale (Uzbekistan)............................................. B. chimganensis sp. nov . - Setae on mesoscutum all black (Uzbekistan).......................................... B. amankutanensis sp. nov . 13 (9) Hind tibia with 2 long and strong anteroventral/ventral setae (Fig. 26) (Japan)........................ B. woodi sp. nov. - Hind tibia without conspicuously long and strong anteroventral/ventral setae..................................... 14 14 (13) Male.............................................................................................. 15 - Female........................................................................................... 21 15 (14) Two phallic hooks long, protruding beyond postgonites with tip visible below postgonites in lateral view, the shorter hook more than 2 / 3 as long as the longer hook. Usually more than 15 setae in intrahumeral and posthumeral areas. Hypandrial processes either broad or narrow and apically axe-like broadened................................................. 16 - Only a single long phallic hook present, the shorter hook less than 1 / 3 as the longer hook, not protruding beyond postgonites and not visible below postgonites inlateral view. Usually less than 14 setae in intrahumeral and posthumeral areas (except B. setitibia sp. nov.). Hypandrial processes narrow or broad (B. thailandica sp. nov.)................................ 18 16 (15) Postgonites broadened laterally, narrow in lateral view, hypandrial processes very short (usually less than 0.11 mm) (see Bart��k & Kub��k 2013, figs 41, 42) (Holarctic species)............................................ B. uvens Melander - Postgonites triangle-shaped in lateral view, hypandrial processes longer (usually more than 0.12 mm)................. 17 17 (16) Hypandrial processes narrow, axe-like broadened apically, with a single long seta in the middle of apex. Postgonites elongate triangular (Fig. 15) (Georgia)............................................................. B. kovalevi sp. nov. - Hypandrial processes strip-like, not broadened apically, without any long seta. Postgonites short triangular (Fig. 22) (East Palaearctic)......................................................................... B. shatalkini sp. nov. 18 (15) Mid tibia with 4���5 pairs of long setae dorsally, at least one pair on apical fourth of tibia. About 15 setae in posthumeral and intrahumeral areas. Hypopygium in Figs 12 and 17 (North Ossetia-Alania, Georgia)................. B. setitibia sp. nov. - Mid tibia with at most 3 pairs of long setae dorsally, lacking even one pair on apical fourth of tibia. Usually less than 14 setae in posthumeral and intrahumeral areas (except some B. ingrata).............................................. 19 19 (18) Legs with generally longer setae (even on hind basitarsi). Hypandrial processes broad, median extensions of postgonites setose and saw-toothed ventrally (Figs 24, 25) and ventral apodeme very narrow in dorsal view (about as broad as phallic hook) (Thailand)......................................................................... B. thailandica sp. nov. - Legs with generally shorter setae (including hind basitarsi). Male hypandrial processes long and narrow. Remaining characters in different combination.............................................................................. 20 20 (19) Face narrower (usually less than 0.04 mm, diameter of 2���3 facets). Prescutellar depression evenly dark. Usually less than 9 setae in intrahumeral and posthumeral areas. Median extensions of postgonites smooth ventrally (see Bart��k & Kub��k 2013, fig. 33) (Palaearctic?).................................................................... B. spuria (Fall��n) - Face broader (usually more than 0.05 mm, diameter of 4���5 facets). Prescutellar depression lighter than disc of scutum. Usually more than 10 setae in intrahumeral and posthumeral areas. Median extensions of postgonites saw-toothed ventrally (Fig. 13) (Israel)............................................................................... B. ingrata Collin 21 (14) Mid tibia with 4���5 pairs of long setae dorsally, at least one pair on apical fourth of tibia. About 10 setae in posthumeral and intrahumeral areas, abdominal tergites 6���8 shining except dorsoapical corner....................... B. setitibia sp. nov. - Mid tibia with at most 3 pairs of long setae dorsally, lacking one pair on apical fourth of tibia (females belonging to this section are difficult to differentiate without associated males)....................................................... 22 22 (21) Legs with generally longer setae (even on hind basitarsi), ground ciliation longer than diameter of limbs. Halter yellowishbrown............................................................................. B. thailandica sp. nov. - Legs with generally shorter setae (including hind basitarsi), ground ciliation at most as long as diameter of limbs. Halter brown to brownish-black................................................................................... 23 23 (22) Usually 2 dorsal setae on fore trochanter. Usually more than 9 setae in posthumeral and intrahumeral areas. Hind tibia swollen (usually about 0.13 mm at broadest point).................................................. B. uvens Melander - Not as above....................................................................................... 24 24 (23) Face broad (usually more than 0.05 mm, diameter of 4���5 facets). Usually more than 9 setae in posthumeral and intrahumeral areas. Legs longer setose (mid femur in middle with anteroventrals at least half as long as depth of femur). B. ingrata Collin - Face narrow (usually less than 0.04 mm, diameter of 2���3 facets). Usually less than 8 setae in posthumeral and intrahumeral areas. Legs shorter setose (mid femur in middle with anteroventrals at most �� as long as depth of femur)............. 25 25 (24) Abdominal tergite 8 entirely microtrichose................................................ B. shatalkini sp. nov. - A bdominal tergite 8 shining at least on basal third........................................................... 26 26 (25) Broad part of third antennal segment usually shorter than 0.11 mm. Hind basitarsus slightly broader than second hind tarsomere.................................................................................. B. kovalevi sp. nov. - Broad part of third antennal segment usually longer than 0.12 mm. Hind basitarsus equally thick as second hind tarsomere......................................................................................... B. spuria (Fall��n), Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 253-254, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria dispar Oldenberg (Fig. 47) Bicellaria dispar Oldenberg, 1920: 221. Taxonomic notes. The species was described on the basis of two males and one female from Tschamintal (=Val di Ciamin, Italy) and one female from Linthal (Switzerland). There are two specimens in Oldenberg���s collection (SDEI), one male from Tschamintal and one female from Linthal. The male labelled as ���Tschamintal 19.6.14 ��� ��� dispar m.��� ��� dispar m. Old.���, ���coll. Oldenberg��� ��� Type ��� ��� Syntypus ��� and an unreadable label, was labelled as LECTOTYPE of B. dispar and it is herein designated. The female belongs obviously to a different species (leading in the key near to B. spuria, but it has an unusually pale grey thorax, almost globular broad part of third antennal segment and pale wings with M veins depigmented. This female syntype cannot be identified more precisely because a great part of the abdomen and halteres are missing). Diagnosis. Small species (wing 1.9 mm long); thoracic setae very short (less than 0.05 mm); antennal stylus together with slender portion of third segment half as long as broadest portion of third segment (Fig. 47). Redescription. Male. Head black, brownish-grey microtrichose, black setose (occiput with somewhat paler setae). Eyes meeting on frons, facets on dorsal half much larger than on ventral half, about 18 facets in line of contiguity of eyes. One pair of short (0.11 mm) black ocellar setae. Occiput sparsely and short setose, setae approx. 0.06 mm long. Face slightly narrowing ventrally, dorsal part just below antennae about 0.04 mm broad and ventral part about 0.03 mm broad at narrowest point (but very slightly collapsed). Palpus short, brown, hardly visible (retracted within head), probably with 2 setae. Labrum short, brown, lustrous. Antenna black, peculiar and quite distinct among all known European species, with second segment short setose; third segment with single dorsal seta near apex (Fig. 47); ratio of broad part of segment 3: stylus (in 0.01 mm scale) = 15: 6. Thorax brownish black, microtrichose, prescutellar area not paler than remaining parts of mesoscutum. All thoracic setae black, some setae on squama appear paler. Chaetotaxy: acrostichals narrowly biserial (11���12 setae in each row) and very short (0.04 mm); dorsocentrals evenly short and irregularly uniserial, only 2 last pairs somewhat longer; 6���7 setae in posthumeral and intrahumeral areas; 1 longer and 3 slightly shorter postpronotals; notopleura with 2���3 setae; several very short setae in supraalar and prealar areas; 2 pairs of scutellars. Legs including coxae brown, microtrichose and dark setose (some setae appear paler). Fore and mid femora with very short setulae, less than �� as long as depth of femur. Fore tibia conspicuously spindle-shaped dilated, very short setose (posteroventrals �� as long as depth of tibia). Mid tibia narrow and very short setose, without prominent setae. Hind femur with 18 anteroventrals slightly shorter than depth of femur, arising from small warts, posteroventrals slightly shorter than anteroventrals, dorsal setae not differentiated. Hind tibia very slightly swollen in apical half (0.06 mm at broadest point and 0.04 at narrowest), short setose (all setae shorter than depth of tibia). Tarsi of all legs thin and short setose. Wing clear including apical section of cell sc; vein M depigmented almost throughout length; clear stigma about 0.40 mm long; distance between tips of veins R 1 and R 2 + 3 = 0.21 mm. Squama brownish yellow with dark or somewhat faded fringes, halter yellow with dark stem. Abdomen brown, microtrichose, very short setose. Genitalia: (undissected) epandrium and cercus of usual shape and setosity; hypandrium scarcely visible, processes apparently long with long setae along inner margins (about 0.07 mm long); postgonites probably of triangular shape, both phallic hooks equal, rather long and simply bent. Female. Unknown. Length: body 1.8 mm (but somewhat collapsed), wing 1.9 mm. Distribution. Italy. Remarks. The specimen is rather immature and long time stored in collection. So it is difficult to decide if some setae are pale or only faded. We decided not to damage this specimen by genital dissection so its description is based on characters which are rather poorly visible. However, we hope that according to the key and redescription it is possible to recognize it easily. The species was omitted in previous studies including those of Tuomikoski (1955) and Chv��la (1983) probably because syntypes were not found in the SDEI collection., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 257-258, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159

Bicellaria thailandica sp. nov. (Figs 24, 25) Type material. HOLOTYPE ♂, Thailand, Kamphaeng Phet, Mae Wong NP, Chong Yen, 1306 m, 16 �� 5.212��N, 99 �� 6.576��E, Malaise trap, 31.xii. 2007 ��� 7.i. 2008, C. Piluek leg (NMWC). PARATYPES: Thailand: 2 ♂, 5 ♀, same locality as holotype, 24���31.xii. 2007; 1 ♀, same locality, 17���24.xii. 2007 ���C. Piluek; 3 ♀, same locality, 10��� 17.xii. 2007; 1 ♀, same locality, 28.xi.��� 3.xii. 2007; 1 ♀, same locality, 12���19.xi. 2007 ���C. Piluek & A. Inpuang; 1 ♂, Chiang Mai, Doi Phahompok NP Kiewlom 2, montane forest, 2112 m, 20 �� 3.426��N, 99 �� 8.553��E, Malaise trap, 14��� 21.ix. 2007; 1 ♂, same locality, 28.ix.��� 4.x. 2007 ���P. Wongchai; 1 ♀, Chiang Mai, Doi Phahompok NP Kiewlom 1, montane forest, 2174 m, 20 �� 3.549��N, 99 �� 8.552��E, Malaise trap, 28.v.��� 7.vi. 2008, P. Wongchai ���(CULSP: 1 ♂, 1 ♀, NMWC: 3 ♂, 11 ♀). Diagnosis. Halter yellowish brown in both sexes; third antennal segment without elongate dorsal setae; all body setae black; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes broad, median extension of postgonites saw-toothed; only one (left) long phallic hook present; female tergite 8 polished in basal two -thirds, sternite 8 polished. Etymology. The species is named after the country of origin (Thailand). Description. Male. Head brownish-black, dark brownish grey microtrichose, black setose. Holoptic, facets in dorsal half much larger than in ventral half, about 10���16 facets in line of contiguity of eyes. One pair of rather short ocellar setae. Occiput dark grey. Face slightly narrowed ventrally, dorsal part about 0.05 mm and ventral part about 0.03 mm broad at narrowest point. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 2 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest ventral setae about 0.10 mm); 3 rd without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 12���16: 4���5: 15���19. Thorax brownish-black, brownish grey microtrichose and slightly subshining including prescutellar area. All thoracic setae black. Chaetotaxy: 5���7 broadly biserial long acrostichals (up to 0.10 mm in front of suture); 6���8 uniserial dorsocentrals as long as or slightly longer than acrostichals; some 3���5 setae in intrahumeral and posthumeral areas; 1 long and 1���2 somewhat shorter postpronotals; notopleuron with 3���4 setae arranged in irregular row (1���2 setae often longer and stronger than others); 1 supraalar- and 1 prealar; 1 long postalar; 2 pairs of scutellars. Legs including coxae blackish brown, microtrichose and black setose, hind femur with polished area in proximal posterior and ventral half. Fore femur with short and sparse anteroventrals half as long as depth of femur, with row of slightly longer posteroventrals. Fore tibia with setae about as long as depth of tibia including posteroventrals, 1���2 posterodorsal up to 2 X as long as depth of tibia, ventral pilosity very short. Mid femur with sparse row of anteroventrals scarcely one-third as long as depth of femur (in middle part), with posteroventrals as long as or slightly longer than depth of femur. Mid tibia with one anterodorsal seta in proximal third and another at about middle (up to 0.20 mm long) and 1 submedian posterodorsal, other short setae slightly longer than depth of tibia. Hind femur with 14 anterodorsal setae slightly longer than depth of femur, with row of 17 anteroventrals slightly longer than anterodorsals at about middle and shorter both basally and apically, posteroventrals short, scarcely half as long as depth of femur, slightly longer on basal part. Hind tibia slightly swollen in apical part (0.11 mm at broadest point in comparison with 0.06���0.07 mm in basal third), longest anterodorsals and posterodorsals up to 0.25 mm long, ventral setae slightly longer than depth of tibia. Tarsi of all legs thin and short setose, basitarsus of hind leg thin (0.06 mm), with rather long setae dorsally (1���2 setae up to twice as long as depth of basitarsus). Wing brownish infuscated; stigma brown, long (0.70 mm), almost parallelsided and narrow (0.06���0.07 mm), slightly narrower than cell r 1 (about 0.09���0.10 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.30 mm. Squama light brownish with dark fringes, halter yellowish brown. Abdomen blackish-brown, dark and almost velvety brown in dorsal view and dark brownish grey in lateral view, microtrichose, pale setose, dorsally with short and partly black setae, posteromarginals on sides of tergites 2���5 much longer than segment, on subsequent tergites subequally long as their corresponding segments. Genitalia (Figs 24, 25) with hypandrial processes broad; postgonites elongate-trapezoid in shape, median extension of postgonites saw-toothed and setose ventrally; left phallic hook long, right hook strongly reduced. Female. Abdominal setae much shorter than in male. Tergites 6���8 polished in basal 1 / 4 to 1 / 2, sternite 8 polished, lateral parts of tergites 2���5 slightly subshining, otherwise abdomen microtrichose. Length: body 2.7���3.3 mm, wing 2.4���3.1 mm. Remarks. All specimens of type series are rather damaged, devoid of most mesoscutal setae; however, their arrangement, number and length may be judged from their articulations and several setae present on some specimens. The species described above is very similar to B. spuria and differential characters are given in the key. Most striking differences are in the shape of male genitalia: the hypandrial processes are much narrower in B. spuria than in the above described species and the median extension of postgonites is saw-toothed in B. thailandica sp. nov. and smooth in B. spuria. Moreover, the ventral apodeme (not illustrated) is very narrow in dorsal view in B. thailandica sp. nov. (not broader than phallic hook), whereas it is much broader (about three times broader than phallic hook) in B. spuria. Another peculiar feature of the species is the setose median extensions of postgonites., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on page 250, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria farkaci sp. nov. (Figs 5, 6) Type material. HOLOTYPE ♂, China, N. Yunnan, Xue Shan nr. Zhongdian, 4200 m, 24.vi. 1996, 27.49 N, 99.34 E, J. Farkač, P. Kab��tek and A. Smetana (CULSP). PARATYPES: 3 ♂, 4 ♀, same data as holotype (CULSP). Diagnosis. All setae black; third antennal segment without elongate dorsal setae; 4���6 setae on each palpus; front tibia with posteroventral setae slightly shorter than depth of tibia, but longer than ciliation; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes absent, phallus with two nearly identical hooks; visible part (approximately posterior half) of female 8 th sternite microtrichose. Etymology. The species is named in honour of one of the collectors and donator of the specimens of the type series, Prof. Jan Farkač (Prague), a specialist on ground beetles (Coleoptera, Carabidae). Description. Male. Head blackish-brown, brownish-grey microtrichose, black setose (including lower occiput). Holoptic, facets in dorsal half much larger than in ventral half, about 10���13 facets in line of contiguity of eyes. One pair of rather short ocellar setae. Occiput rather densely and long setose. Face slightly broadened ventrally, ventral part about 0.07 mm broad. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 4���6 setae. Labrum short, brown, polished. Antenna black, 2 nd segment long setose ventrally (longest setae almost 0.20 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 15���20: 3���5: 20���23. Thorax blackish-brown, microtrichose, mesoscutum dark brownish black in dorsal view, prescutellar area differently microtrichose than other parts of mesoscutum, appearing pale brownish grey. All thoracic setae black including fringes on squama. Chaetotaxy: 7���8 narrowly biserial acrostichals; 8���9 uniserial dorsocentrals; 1 intrahumeral, 1 posthumeral and 0���2 additional setae in posthumeral and intrahumeral areas; 1 long and 1���2 shorter postpronotals; notopleuron with 3���4 unequally long setae arranged in one irregular row; 1 supraalar and 1 strong prealar; 1 long postalar; 3 pairs of long scutellars; pleura, antepronotum and sternites without any setae. Legs including coxae black, microtrichose and black setose. Fore femur with row of very short anteroventrals and row of longer posteroventrals (slightly shorter than depth of femur). Fore tibia with 0���2 dorsal and 0���1 submedian posterodorsal setae about 1.5 X as long as depth of tibia, otherwise short setose, posteroventrals in apical third about as long as depth of tibia, pilosity apparently absent (similar to B. spuria); conspicuous anteroventral sense organ ovoid, about 0.10 mm long, consisting of central area of very short hairs and surrounding ovoid circle devoid of microtrichiae. Mid femur with sparse row of anteroventrals one-third as long as depth of femur and with much stronger and longer posteroventrals slightly longer than depth of femur. Mid tibia with 2 anterodorsal (1 subbasal and 1 submedian) and 1 submedian posterodorsal setae 2 X as long as depth of tibia and 2 ventral preapical setae. Hind femur with 15���17 anterodorsal setae on basal third slightly longer than depth of femur, with row of 18���20 anteroventrals about as long as depth of femur, much shorter posteroventrals absent on apical half to third. Hind tibia very slightly swollen in apical part (0.12 mm at broadest point in comparison with 0.08 mm in basal part) being much narrower than hind femur (0.18 mm at broadest point), both dorsal and ventral setae slightly longer than depth of tibia, several anterodorsals up to 0.20 mm long. Tarsi of all legs slender, short setose. Wing slightly brownish infuscated; stigma dark brown, long (0.80 mm), almost parallel-sided and narrow (0.05 mm), much narrower than cell r 1 (about 0.13 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.40 mm. Squama dark brown, halter dark brown. Abdomen brown, microtrichose, entirely black setose, dorsally with short setae, marginals on sides of tergites 2���5 slightly longer than corresponding segments, on subsequent tergites slightly shorter. Genitalia (Figs 5, 6) with hypandrial processes absent, distal part of hypandrium with some 5 long setae on each half; postgonites triangle-shaped, two nearly identical phallic hooks present. Female. Tergites entirely pruinose, sternite 8 microtrichose on apical (visible) half, polished on basal half. Length: body 2.7���3.5 mm, wing 3.2 ���4.0 mm. Remarks. All specimens of the type series are rather damaged, devoid of most mesoscutal setae; however, their arrangement, number and length may be judged from their articulations and several setae present on some specimens. The new species is quite different from all known species of Bicellaria in having hypandrium rounded caudally, without processes. Also the combination of setose palpus with third antennal segment lacking long dorsal setae is not present in any known species and the microtrichose female 8 th abdominal sternite is also a very peculiar feature. According to all these characters this species is easily recognizable from all other species of Bicellaria worldwide., Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 237-238, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129

Bicellaria uvens Melander (Figs 41���42) Bicellaria uvens Melander, 1928: 74. Bicellaria bisetosa Tuomikoski, 1936: 84, syn. nov. Type material examined. B. bisetosa: PARALECTOTYPES: Russia: 13, Kuusamo [probably collected in area of today���s Russia], 21.vii. 1934, Krogerus (labelled: ���Mus. Zool. H:fors, Spec. typ. No. 8143 Bicellaria bisetosa Tuomikoski ���); 13, same locality, 19.vii. 1934, Krogerus (labelled: ���Mus. Zool. H:fors, Spec. typ. No. 8142 Bicellaria bisetosa Tuomikoski ���); 13, same locality, 13.vii. 1934, Krogerus (labelled: ���Spec. Typ.���)���(NHMH). Additional material examined (193, 9��). NEARCTIC: Canada: BC, 13, Summit L., Alaska Hwy mi. 392, 4500 ���, 17���19.vii. 1959, E.E. McDougall; 13, Atlin, 2200 ���, 23.vii. 1955, B.A. Gibbart; Labrador, 13, Hebron, 10.vii. 1954, J.F. McAlpine; Quebec, 13, Ft. Chimo [= Kuujuaq], 16.vii. 1948, H.N. Smith; 13, Indian House Lk., 24.vii. 1954, W.R. Richards; 2 ��, same locality, 5.vii. 1954, R. Coyles���all ��� B. uvens ��� B. Sinclair det.���(CNC). USA: Alaska, 13, King Salmon, Naknek, 13.vii. 1952, J.B. Hartley; 13, Unalakleet, 4.vii. 1961, B.S. Heming; 1 ��, same locality, 5.viii. 1961, B.S. Heming���all ��� B. uvens ��� B. Sinclair det.���(CNC). PALAEARCTIC: Finland: 13, Utsjoki, R. Tuomikoski; 13, same locality, R. Frey; 1 ��, Muonio, 2.vii. 1867, J. Sahlberg���(MHNH). Russia: 1 ��, Kuusamo, 13.vii. 1934, Krogerus; 13, same locality, 19.vii. 1934; 53, 2��, same locality, 19.vii. 1934; 1 ��, same locality, 21.vii. 1934; 13, same locality, 29.vii. 1934; 13, Suistamo, R. Tuomikoski; 13, Kamtschatka, Bolscherjetsk [now Ust���Bolsheretsk], 20.vii. 1917, Y.Wuorentaus; 1 ��, same locality, 3.vii. 1917 (���Mus. Zool. H:fors, Spec. typ. No. 8201 Bicellaria stackelbergi Tuomikoski ������ paratype of B. stackelbergi)���(MHNH); 13, Amurskaja oblast, Zeja, 8.vii. 1981, A. Shatalkin (ZMMU). Diagnosis. The species belongs to a distinct complex of species [along with at least B. pilipes (Loew) and one still undescribed speices from Siberia] characterized by broad hypandrial processes and usually more than 10 setae on intra- and posthumeral areas. Two dorsal setae usually occur on the fore trochanter (hence the name ��� bisetosa ���); however, we found also specimens with only a single seta here (two of them being paralectotypes of B. bisetosa), and, contrary, other species may have occasionally two such setae (e.g., B. sulcata). Males of B. uvens may be easily identified according to the genitalia with both phallic hooks present and postgonites slightly broadened laterally and narrow in lateral view (Figs 41���42). There is considerable variability between individual specimens of the species in colour of mesoscutum (being usually darker in Nearctic specimens), in length of ventral setae on hind tibia, in number of scutellar setae (4���6), and in the shape of hind basitarsus (which is sometimes slightly broadened). However, these characters are not consistently different between specimens from North Europe, Eastern Asia and North America; moreover, genitalia of all specimens studied are identical. There may be problems with identification of females of this complex of species. Distribution. Holarctic species. In Europe known only from Fennoscandia. Published records from Italy (Raffone 2008) and Romania (Parvu 2003) are both probably based on misidentifications and were not included in recent catalogs (Chv��la 2012; Yang 2007). Remarks. The species B. uvens was described from numerous specimens from Fort Chimo [= Kuujuaq], Ungava Bay, Hudson Straits, and northern Labrador (Canada). The species B. bisetosa was described from an unknown number of specimens from Finland and North European part of Russia. In a later work (Tuomikoski 1955), ���Ussuri��� was added. Chv��la (1983) redescribed this species (under B. bisetosa), provided illustrations of the face, antenna, fore and hind leg, and genitalia, and designated the lectotype. According to the study of Nearctic materials of B. uvens, we consider both species identical., Published as part of Bart��k, Miroslav & Kub��k, ��t��p��n, 2013, Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species, pp. 251-278 in Zootaxa 3647 (2) on pages 270-272, DOI: 10.11646/zootaxa.3647.2.2, http://zenodo.org/record/224159