Bicellaria woodi Bart��k, Plant & Kub��k, 2013, sp. nov

Bicellaria woodi sp. nov. (Fig. 26) Type material. HOLOTYPE ♂, Japan, Hokkaido, Daisetsuzan Nt. Pk., Tenninkyo, 800 m, 10���14.vii. 1986, D.M. Wood (CNC). PARATYPES: Japan: 2 ♀, same data as holotype (CNC, CULSP). Diagnosis. All setae black; third antennal segment without elongate dorsal setae; 1���2 setae on each palpus; hind basitarsus very narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes long and broad, strip-like. Hind tibia with two long and strong preapical (ventral and anteroventral) setae. Etymology. The species is named in honour of collector of type series specimens, Monty D. Wood (Ottawa). Description. Male. Head blackish-brown, brownish-grey microtrichose, black setose (including lower occiput). Holoptic, facets in dorsal half much larger than in ventral half, about 15���18 facets in line of contiguity of eyes. One pair of ocellar setae (lost in all specimens). Occiput rather densely and not very long coarse setose. Face slightly narrowing ventrally, dorsal part about 0.07 mm broad and ventral 0.05 mm. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 1���2 setae. Labrum short, brown, polished. Antenna black, 2 nd segment short setose (longest setae almost 0.12 mm); 3 rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 15���16: 10���11: 19. Thorax blackish-brown, microtrichose, mesoscutum dark brownish black in dorsal view, prescutellar area evenly dark. All thoracic setae nearly completely rubbed out, apparently all black and coarse. Chaetotaxy: 8���9 biserial acrostichals; 8���10 uniserial dorsocentrals; some 10 setae in posthumeral and intrahumeral areas; 4���5 postpronotals; notopleuron with 7���8 setae arranged in irregular row (strong and long in front part of notopleuron); 1���2 supraalar- and 3���4 prealar; 1 postalar; at least 3 pairs of long scutellars (one preserved in one female is nearly 0.60 mm long). Legs including coxae black, microtrichose and black setose. Both forelegs in male missing (see female description). Hind coxa with long upright standing setae. Mid femur with sparse row of very short anteroventrals and with much stronger and longer posteroventrals subequally long as depth of femur. Mid tibia with 2 anterodorsal (one subbasal and one submedian) and 1 submedian posterodorsal setae 3 X as long as depth of tibia (longest about 0.30 mm) and 2 (anterior and anteroventral) conspicuously strong but rather short preapical setae, ventral setae slightly longer than depth of tibia. Hind femur with about 20 anterodorsal and 19 anteroventral setae subequally as long as depth of femur, posteroventrals slightly shorter than anteroventrals. Hind tibia slightly swollen in apical part (0.13 mm at broadest point in comparison with 0.08 mm in basal part) much narrower than hind femur (0.19 mm at broadest point), both dorsal and ventral setae slightly longer than depth of tibia, 4���5 pairs of dorsal setae, longest posterodorsal 0.32 mm long, two strong and long preapical setae in anterior and anteroventral positions (Fig. 26). Tarsi of all legs very thin and short setose. Wing brownish infuscated, fore margin dark brown, especially between veins C and R 1; stigma very long (1.00 mm), almost parallel-sided and narrow (0.05 mm), narrower than cell r 1 (about 0.08 mm broad at narrowest point). Distance between tips of veins R 1 and R 2 + 3 about 0.20 mm. Squama dark brown with black fringes, halter brown. Abdomen brown, microtrichose, entirely black setose, setae long, even those on 1 st sternite unusually long. Genitalia (undissected) with hypandrial processes long and broad, strip-like (as in B. shatalkini sp. nov.). Female. Fore coxa densely setose. Fore femur with row of fine anteroventrals half as long as depth of femur, with similar row of slightly longer posteroventrals. Fore tibia with posteroventrals distinctly longer than depth of tibia and without ventral pilosity (present as usual on fore basitarsus), with 1���3 setae dorsally about twice as long as depth of tibia. Tergites entirely pruinose including visible parts of tergites 6���7, sternite 8 polished. Length: body 3.6���3.9 mm, wing 3.4���3.6 mm. Remarks. All specimens of type series are rather damaged, devoid of most mesoscutal setae and most abdominal setae; however, their arrangement, number and length may be judged from their articulations and from several setae present on some places on some specimens. The new species somewhat resembles B. farkaci sp. nov. in general appearance but the palpus has only 1���2 setae, female sternite 8 is polished and genitalia quite different. Bicellaria woodi sp. nov. is easily recognizable in both sexes and unique in the World fauna due to two strong preapical anterior (and anteroventral) setae on the hind tibia. We decided not to damage the only male (holotype) by genital dissection, so are unable to present a full description of the genitalia.
Published as part of Bart��k, Miroslav, Plant, Adrian & Kub��k, ��t��p��n, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, pp. 233-256 in Zootaxa 3710 (3) on pages 252-253, DOI: 10.11646/zootaxa.3710.3.3, http://zenodo.org/record/223129