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Some new findings of the family Pamphagidae obtained in the field trips carried out in Turkey are presented. New distributional, morphological, and zoogeographical information are given for the recorded 15 species and subspecies. A new species Nocaracris nebulosa Ünal, sp. nov. and the unknown male of Nocaracris tauricola Ramme, 1951 are described.

The genus Finotia was established by Bonnet in 1884 and included in the subfamily Pamphaginae; later, in 1916 Bolívar described the tribe Finotiini to include the genus and differentiate it from the other of the subfamily Pamphaginae, tribe Pamphagini. Differently from previous authors, who agreed with this placement, the present authors examined the phallic complex of Finotia maxima Jannone, 1938, finding out that it has the typical characters present in the subfamily Thrinchinae; thus they move the tribe Finotiini from the subfamily Pamphaginae to that of Thrinchinae. In addition, the genus Ocneridia Bolívar, 1912, differently from other genera within the tribe Pamphagini, lacks of the Krauss’ plate; coherently with the tribe placements within the subfamily Pamphaginae, the present authors establish the new tribe Ocneridiini to include the genus Ocneridia., This research received the support of the SYNTHESYS+ project http://www.synthesys.info/, financed by European Community Research Infrastructure Action under the FP7 “Capacities” Programme at the Museo Nacional de Ciencias Naturales, Madrid (CSIC) (B.M.: 2013: ES-TAF-2438; 2022: ES-TAF 3916; M.Ü.: ES-TAF-3897), the Museum für Naturkunde, Berlin (B.M.: 2014: DE-TAF-4109; M.Ü.: DE-TAF-558; DE-TAF-2761), the Naturhistorisches Museum, Vienna (B.M.: 2016: AT-TAF-5324; M.Ü.: AT-TAF-3896; AT-TAF-2757), the National Museum, Prague (B.M.: 2016: CZ-TAF-5559) and the Royal Belgian Institute of Natural Sciences, Bruxelles (B.M.: 2017: BE-TAF-6319).

The third part of a study series on the family Tettigoniidae from Turkey and the Middle East Region is presented, which includes only the subfamily Tettigoniinae of Turkey. A key to tribes, genera and subgenera found in Turkey is proposed. The synonymy of the tribe Platycleidini Brunner, 1893 (with Decticini Herman, 1874) is confirmed. Pezodrymadusa striolata ziyaretensis Koçak & Kemal, 2010 [with Pezodrymadusa affinis (I.Bolívar, 1899)], Anadolua rammei Karabağ, 1952 (with Anadolua burri Karabağ, 1952), Psorodonotus rize Kaya et Çıplak, 2014 (with Psorodonotus davisi Karabağ, 1956), Parapholidoptera intermixta Karabağ, 1961 [with Parapholidoptera indistincta (I.Bolívar, 1899)] are synonymized. Parapholidoptera indistincta (I.Bolívar, 1899) is reinstated as an independent species. Six new species, Pezodrymadusa dentata Ünal, sp. nov., Mixodusa retusa Ünal, sp. nov., Platycleis (Squamiana) supericola Ünal, sp. nov., Platycleis (Squamiana) goeksunica Ünal, sp. nov., Parapholidoptera yarpuzi Ünal, sp. nov. and Eupholidoptera singularis Ünal, sp. nov. are described. The invalidly described Schulmeisteri schulmeisteri Harz is discussed and its only specimen is included in the genus Pezodrymadusa as a new species. The genus Mixodusa Stolyarov and the subspecies, Pholidoptera aptera bulgarica Maran, 1953 are recorded from Turkey for the first time. An overlooked historical record of Psorodonotus venosus brunneri Stshelkanovtzev, 1914 from Turkey is determined. Discussions of many taxa and some records from new localities are presented. A full list of the Turkish Tettigoniinae is given. 357 figures are provided.

The genus Bradyporus Charpentier is taxonomically revised using information obtained from numerous recently collected specimens. Bradyporus is divided into two subgenera Bradyporus Charpentier s.str. and Callimenus Fischer von Waldheim. Five species groups are distinguished in the latter subgenus. Sixteen species and subspecies are listed. Bradyporus (Callimenus) multituberculatus multituberculatus (Fischer von Waldheim, 1833) and Bradyporus (Callimenus) multituberculatus montandoni (Burr, 1898) are designated subspecies for the first time. Previously synonymized Bradyporus (Callimenus) macrogaster skopjensis Karaman, 1961 is reinstated as a valid species. A new species, Bradyporus (Callimenus) gocmeni Ünal, sp. nov. from Eastern Turkey and the previously unknown female of Bradyporus (Callimenus) skopjensis Karaman, 1961 are described. A Neotype for Bradyporus (Callimenus) macrogaster longicollis (Fieber, 1853) is designated. Nomenclatural and taxonomical problems are discussed under the relevant taxa. Phylogenetic relationships based on morphology of the subgenera, species groups, and the species and subspecies of each species group are proposed. Morphological derived and ancestral characters are discussed. A key to the lower taxa of the genus is provided with 207 original figures including 3 distributional maps. A check-list of the lower taxa of the genus is given. Additionally the IUCN Red List statuses of the species and subspecies are included.

In the present paper, the male of Spinisternum castaneipictus Willemse, previously unknown is described. The similarity of Spinisternum with other genera causes confusion in identification. This problem is discussed and similar species in different genera are compared. A key to species of Spinisternum and illustrations of S. castaneipictus are provided.

The complicated genus Glyphotmethis Bey-Bienko, 1951 is revised using the previous materials including the type specimens preserved in Museums and an abundance of new material collected mainly in Turkey. Diagnoses, ecology, distribution and taxonomic characters for the genus are provided. The function of Krauss’ organ is discussed. The followingseven taxa are proposed as junior synonyms: Glyphotmethis holtzi spinosus Karabag, 1956 is synonym with G. holtzi pulchripes (Uvarov, 1943), Glyphotmethis holtzi extimus Bey-Bienko, 1951 is synonym with G. holtzi holtzi (Werner, 1901), Glyphotmethis escherichi coloripes Cejchan, 1965 is synonym with G. escherichi elatior (Ramme, 1951), Glyphotmethis guichardi Cejchn, 1964 is synonym with G. escherichi escherichi (Krauss, 1896), Glyphotmethis raggei Cejchan, 1965 and Glyphotmethis adaliae angorensis Cejchan, 1965 are synonyms with G. escherichi inermis (Uvarov, 1934) and Glyphotmethis heldreichi macedonicus, Bey-Bienko, 1951 is synonym with G. heldreichi heldreichi (Brunner, 1882). One species, Glyphotmethis arasi, is removed from the genus and three new taxa, Glyphotmethis holtzi brachypterus ssp.n., Glyphotmethis holtzi turcicus ssp.n. and Glyphotmethis efe sp.n. are described. The taxonomical status of some taxa are rearranged. A neotype is designated for Glyphotmethis eshcerichi escherichi (Krauss, 1896) from Ankara known as the type locality. Misidentified specimens and inaccurate records for taxa in previous papers are corrected and their geographical distributions adjusted. Many records are provided for new localities. Keys for the identification of species and subspecies are prepared using traditional and some new distinctive characters. A short description and comparison with close allies are given for each taxon. Distribution areas of the taxa and the contact zones between some subspecies are determined. In addition, 126 figures and 3 distributional maps are given.

The endemic Madagascan genera Oncodopus Brongniart and Colossopus Saussure are revised using museum specimens, including the types, and recently collected material. A new genus, Malagasopus gen. nov., and seven new species, Malagasopus desutterae sp. nov., Malagasopus meridianus sp. nov., Oncodopus janetae sp. nov., Oncodopus brongniarti sp. nov., Oncodopus saussurei sp. nov., Oncodopus soalalaensis sp. nov. and Colossopus parvicavus sp. nov. are described. Lectotypes are designated for Oncodopus zonatus Brongniart, 1897 and Colossopus redtenbacheri (Brongniart, 1897). A new term, mesothoracic auricle, is proposed for a structure on the episternum of the mesothorax. A tabulated key to the genera and keys to the species are presented. All species are described and diagnosed, and their phylogenetic relationships, geographical distributions, habitat preferences and phenologies are documented. The life history of Colossopus grandidieri is described, and the unusual possible mate-guarding behaviour of several species is discussed. Maps showing the distribution of the species are presented, as too are 57 photographs of museum specimens, 51 drawings of morphological characters, 17 photographs of living specimens and one habitat photograph.

The very rich material of Palaearctic Pamphagidae preserved in the collections including the type specimens of all taxa, the historical unidentified specimens and newly collected material have been studied. 58 genera and 295 species and subspecies are listed. The higher classification of the family is reviewed mainly based on the male phallic complex as well as the traditional and some new external characters. The tribe Haplotropiidini is transferred to the subfamily Thrinchinae. Previously synonymized subfamily Tropidaucheninae is validated as an independent tribe of Pamphaginae. The tribe Nocarodeini is taxonomically reviewed except the genus Bufonocarodes Mistshenko. The male phallic complex is also used as one of the main characters in decisions of the genus group taxa. The following 10 genera are synonymized: Pseudotmethis Bey-Bienko, 1948 and Paratmethis Zheng & He, 1996 (with Filchnerella Karny, 1908), Sinotmethis Bey-Bienko, 1959 and Kanotmethis Yin, 1994 (with Beybienkia Tsyplenkov, 1956), Paktia Pfadt, 1970 (with Mistshenkoella Cejchan, 1969), Pseudosavalania Demirsoy, 1973 (with Paranocarodes I. Bolívar, 1912), Nocaropsis Ramme, 1951 (with Paranothrotes Mistshenko, 1951), Paranocaracris Mistshenko, 1951 and Oronothrotes Mistshenko, 1951 (with Nocaracris Uvarov, 1928) and Savalania Mistshenko, 1951 (with Nocarodes Fischer von Waldheim, 1846). The genera Mistshenkoella Cejchan and Cryptonothrotes La Greca are proposed as subgenera of Saxetania Mistshenko. The following genera are divided: Eremocharis Saussure into 4 species groups, Paranocarodes I. Bolívar into 2 species groups, Paranothrotes Mistshenko into 4 species groups, Nocaracris Uvarov into 7 species groups and Nocarodes Fischer von Waldheim into 4 species groups. The following 24 species and subspecies are synonymized: Tmethis cinerascus cyanipes Werner, 1939 [with Eremopeza angusta (Uvarov, 1934)], Iranotmethis cyanipennis cyanipes Bey-Bienko, 1951 [with Iranotmethis cyanipennis iranicus (Werner, 1939)], Eremocharis granulosa roseipes Uvarov, 1943 [with Eremocharis granulosa granulosa (Walker, 1871)], Eremocharis subsulcata minor Cejchan, 1969 (with Eremocharis maior Ramme, 1952), Asiotmethis limbatus motasi Ramme, 1951 [with Asiotmethis limbatus (Charpentier, 1845)], Tropidauchen marginatum Bolívar, 1912 (with Tropidauchen escalerai Bolívar, 1912), Paranocarodes straubei serratus Uvarov, 1949, Paranocarodes straubei insularis Ramme, 1951 and Paranocarodes fieberi mytilenensis Ramme, 1951 [with Paranocarodes fieberi (Brunner von Wattenwyl, 1882)], Paranocarodes atympanicus Ramme, 1951 (with Paranocarodes tolunayi tolunayi Ramme, 1949), Paranocarodes cilicicus Ramme, 1951 (with Paranocarodes lubricus Mistshenko, 1951), Paranothrotes opacus hakkariana Demirsoy, 1973 [with Paranothrotes opacus opacus (Brunner von Wattenwyl, 1882)], Paranothrotes tenuicornis sordidus Mistshenko, 1951 and Paranothrotes iranicus (Ramme, 1951) (with Paranothrotes tenuicornis Mistshenko, 1951), Paranothrotes elbursianus (Ramme, 1951) [with Paranothrotes demawendi (Ramme, 1951)], Paranocarodes aserbeidshanicus Ramme, 1951 [with Paranothrotes margaritae (Miram, 1938)], Nocaracris burri obscurata Ramme, 1951 [with Nocaracris furvus furvus (Mistshenko, 1951)], Paranocaracris rimansonae ventosus Mistshenko, 1951 [with Nocaracris rimansonae (Uvarov, 1918)], Paranocaracris rigidus Mistshenko, 1951 (with Nocaracris tauricolus Ramme, 1951), Nocaracris cyanipes tristis Ramme, 1951 [with Nocaracris cyanipes (Motschulsky, 1846)], Nocarodes armenus Ramme, 1951 (with Nocarodes serricollis Fischer von Waldheim, 1846), Nocarodes specialis Mistshenko, 1951 [with Nocarodes iranicus (Werner, 1939)], Nocarodes gibbosus Mistshenko, 1951 (with Nocarodes humerosus Mistshenko, 1951), Nocarodes urmianus carinatus Mistshenko, 1951 (with Nocarodes urmianus Ramme, 1939). One new genus in the tribe Nocarodeini, Turkanocaracris Ünal gen. nov., and the following 29 new species and subspecies in four tribes are described: Eremopeza soltanii Ünal, sp. nov., Eremopeza kashmirensis Ünal, sp. nov., Eremocharis subsulcata beccalonii Ünal, ssp. nov., Orchamus massai Ünal, sp. nov., Paracinipe suezensis Ünal & Massa, sp. nov., Saxetania (Mistshenkoella) gorochovi Ünal, sp. nov., Paranocarodes anatoliensis anamas Ünal, ssp. nov., Paranothrotes buzuldagi Ünal, sp. nov., Paranothrotes dentatus Ünal, sp. nov., Paranothrotes eximius bitlis Ünal, ssp. nov., Paranothrotes siirt Ünal, sp. nov., Nocaracris furvus kazdagi Ünal, ssp. nov., Nocaracris istanbul Ünal, sp. nov., Nocaracris tunceli Ünal, sp. nov., Nocaracris tardus Ünal, Bugrov & Jetybayev, sp. nov., Nocaracris emirdagi Ünal, sp. nov., Nocaracris cejchani Ünal, sp. nov., Nocaracris judithae Ünal, sp. nov., Nocaracris minutus Ünal, sp. nov., Nocaracris karadagi Ünal, sp. nov., Nocaracris palandoken Ünal, sp. nov., Nocaracris monticolus Ünal, sp. nov., Nocaracris van Ünal, sp. nov., Nocaracris dilekensis Ünal, sp. nov., Nocaracris karshitoros Ünal, sp. nov., Nocaracris goektepe Ünal, sp. nov., Nocaracris crassipes Ünal, sp. nov., Turkanocaracris levigatus Ünal, sp. nov., Turkanocaracris davisi Ünal, sp. nov. Lectotypes are designated for the following 12 species: Strumiger desertorum desertorum Zubovski, 1896, Ocnerosthenus brunnerianus (Saussure, 1887), Orchamus gracilis (Brunner von Wattenwyl, 1882), Saxetania iranicum (Werner, 1939) [syn. of Saxetania (s.str.) cultricolle (Saussure, 1887)], Paranocarodes straubei (Fieber, 1853), Paranocarodes fieberi (Brunner von Wattenwyl, 1882), Paranothrotes shelkovnikovi (Uvarov, 1918), Paranothrotes nigripes (Stshelkanovtzev, 1916), Eunothrotes derjugini Adelung, 1907, Nocaracris rimansonae (Uvarov, 1918), Nocaracris tridentatus (Stshelkanovtzev, 1916) and Araxiana woronowi (Uvarov, 1918). Many species are given in new combinations and new statuses. Some taxa are proposed as reverted combinations, reverted statuses, reverted genera, species and subspecies. Nomenclatural problems are discussed under the relevant taxa. The authorship of 4 species are discussed and changed as follow: Eremopeza festiva (I. Bolívar, 1884), Paranocarodes tolunayi tolunayi Ramme, 1949, Nocaracris rubripes (Motschulsky, 1846), Nocaracris cyanipes (Motschulsky, 1846). Identification keys for subfamilies, tribes and genera (except the N. African and European genera of Pamphagini) and for each level of the tribe Nocarodeini are prepared. This study is also a review of the Pamphagidae species of Turkey. Therefore all Turkish species are studied in detail except the two genera recently revised by the author, Glyphotmethis Bey-Bienko and Asiotmethis Uvarov. Geographically this paper includes all Pamphagidae species of the following countries and regions: Bulgaria, Turkey, Cyprus, Syria, Israel, Palestine, Jordan, Iraq, Armenia, Georgia, Nakhichevan, Azerbaijan, Pakistan, Greece (except one species), Lebanon (except two species), Arabian Peninsula (except one species), Afghanistan (except some Saxetania) and Iran (except some Saxetania and Tropidauchen). In addition, 848 original figures including 10 distributional maps are provided.

Kaya et al. (2012) published a paper on the genus Poecilimon Fischer (Zootaxa, 3225). They synonymized eight species based on morphological characters by bioacoustic data (according to them). They frequently cited my paper (Ünal, 2010) during the text. But, I have seen that some cited informations from my paper and the reflecting my statements need correction, and some mistakes and wrong evaluations which are the bases of their taxonomic decisions should be re-evaluated. Furthermore many related results of Ünal (2010) were overlooked.

The genus Prionotropis Fieber, 1853 is revised. It is distributed in scattered areas of the Mediterranean region from Turkey in the East to Spain in the West. Overall, seven species are listed, namely P. maculinervis (Stal, 1878) (Turkey; P. urfensis Ramme, 1933 is here considered its synonym), P. willemsorum n. sp. (Greece, Epirus; previously considered P. appula), P. appula (O.G. Costa, 1836) (South Italy), P. hystrix (Germar, 1817) (Bosnia and Herzegovina, Slovenia, Croatia, North-East Italy; P. hystrix sontiaca is here synonymized), P. rhodanica Uvarov, 1923 resurrected status (France, Crau, Rhone delta; here considered a valid species), P. azami Uvarov, 1923 n. status (France, Var region; here considered a valid species), and P. flexuosa (Serville, 1838) (Spain; the ssp. pereezi Bolivar, 1921 and sulphurans Bolivar, 1921 are here considered its synonyms). A key to species is presented.

The species of Bradyporus Charpentier, 1825 found in Turkey has been revised by the second author (M. Unal, 2011). Bradyporus comprises thirteen species and subspecies at present. The species of the genus have a range of Central Palaearctic distribution; most of the species are Balkanian-Anatolian-Iranian, whereas some of them are restricted to Ukraine, the Caucasus, Transcaucasus, being inhabitants in the continental mountain steppe belt. It seems that Anatolia, with 8 species, and Balkans, with 6 species, are two diversity centres of this genus. Iran represented by only species, B. latipes (Stal, 1875). But, one more peculiar species is described from north west of Iran herein. The species of this genus are very similar to one another in their external features, however, detailed descriptions and characterizations are given for the authentic separation of the known species in the exhausting revision of the genus by the second author (Unal, 2011).

The members of the subfamily Bradyporinae including Bradyporini and Ephippigerini found in Turkey are studied. Two phylogenetic relationships based on morphology for the subgenera, species groups and for the all species of Bradyporus s.l. are proposed. A new diagram shows together the relationships and the schematized distributions of the taxa, Geographical Network Relationship, is proposed. A synonymic checklist of the known species of Bradyporus s.l. is given. Taxonomical changes, the doubtful and uncertain taxa names are discussed. Callimenus Fischer von Waldheim, 1830 is proposed as a subgenus of Bradyporus Charpentier, 1825. Four species groups, B. oniscus sp. group, B. macrogaster sp. group, B. dilatatus sp. group and B. latipes sp. group, are separated. Callimenus macrogaster pancici Brunner, 1882 is synonymized with Bradyporus (Callimenus) macrogaster longicollis (Fieber, 1853). Neotypes for B. (s.str.) dasypus (Illiger, 1800) and B. (Callimenus) macrogaster macrogaster (Lefebvre, 1831) are designated. Five new species, Bradyporus (Callimenus) avanos sp. n., B. (Callimenus) sureyai sp. n., B. (Callimenus) conophallus sp. n., B. (Callimenus) toros sp. n. and B. (Callimenus) karabagi sp. n. are described. Bradyporus (Callimenus) latipes (Stal, 1875) is recorded from Turkey for the first time. The species Schulmeisteri schulmeisteri Harz, 1992 is excluded from the subfamily Bradyporinae. All previous records of Bradyporinae species in Turkey with full synonymic list of each species are given. The previous misidentifications are determined and then the distribution of each species is adjusted. The characters used the separation of the taxa are briefly discussed. The titillator for Turkish species and the basal projecting of ventral valve of ovipositor are used for the first time. Descriptions and redescriptions of the studied Bradyporus species are given. Keys to tribes, subgenera, species groups and Turkish species of Bradyporus are provided. In addition 157 original figures including a distributional map are provided.