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1. Chrysodema (Chrysodema) sekerkai Frank 2022, sp. nov

Author: Frank, David
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Chrysodema sekerkai, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) sekerkai sp. nov. (Figs 38–40, 61, 62, 68, 75) urn:lsid:zoobank.org:act: 71D04526-4994-4F7D-AED4-5B104236BD15 Chrysodema (Chrysodema) antennata obsoleta [misidentification]: Lander (2003): 80 (colour Fig. 96) Type locality. Indonesia: Central Sulawesi Province, Banggai Island. Type material examined. HOLOTYPE: &male; (22.75 × 8.25 mm; Figs 38–40, 61, 68), ‘ Bangkei | H. Kühn | 1885 [w(c), p (black border)]’ (MNHN). PARATYPES (4 &male;&male;, 143 &female;&female;): INDONESIA: CENTRAL SULAWESI PROv.: Banggai Island: the same data as holotype (4 &female;&female; MNHN, 2 &female;&female; DFPC, 1 &female; LSPC); ‘Bangkei | H. Kühn | 1885 [w(c), p (black border)] || 13. [w, h]’, 1 &female; (MNHN); ‘Bangkei | H. Kühn | 1885 [w(c), p (black border)] || Betw. TALIABU IS | NE Sulawesi [w, h] || Chrysodema | Dalmani | Mann.? | Det. Dr. Obenberger [w, h/p] || Chrysodema | antennata | Saund. | Dét. T. LANDER 192001 [w, h/p]’, 1 &female; (NMPC); ‘Bangkei | H. Kühn | 1885 [w(c), p (black border)] || Collect. | Plason [w, p] || Chrysodema | antennata | Saund. | THERY det. [w, h/p]’, 1 &female; (NHMW); ‘Bangkei | H. Kühn | 1885. [w(c), p (black border)] || Coll. | Jul.Moser [w, p] || vagans Gory [w(c), h] || Chrysodema | obsoleta | Thoms., 1878 | Det. T. LANDER 2000 [w, p]’, 1 &female; (MFNB); ‘Bangkei | H. Kühn | 1885. [w(c), p (black border)] || Coll. | Jul.Moser [w, p] || Chrysodema | n.sp.? 50. [w(c), h] || Chrysodema | obsoleta | Thoms., 1878 | Det. T. LANDER 2000 [w, p]’, 1 &female; (MFNB); ‘Bangkei | H. Kühn | 1885. [w(c), p (black border)] || Coll. | Jul.Moser [w, p] || Chrysodema | obsoleta | Thoms., 1878 | Det. T. LANDER 2000 [w, p]’, 3 &female;&female; (MFNB); ‘Bangkei | Fruhstorfer. [w(c), h]’, 1 &female; (MNHN); ‘Bangkei | Fruhstorfer. [w(c), h] || VU PAR KERREMANS | antennata Snd. | POUR SA MONOGRAPHIE [w(c), p/h (black border)]’, 1 &female; (MNHN); ‘Bangkei | Staud. [w, h (Kerremans' handwritting)] || MUSEUM PARIS | COLL. CH. KERREMANS | 1923 [y, p] || hebes | Kerr. [w, h (Kerremans' handwritting)]’, 1 &female; (MNHN); ‘Bangkei | Staud. [w, h (Kerremans' handwritting)] || hebes | Kerr. [w, h (Kerremans' handwritting)]’, 1 &female; (MNHN); ‘Banggai Isl | Celebes | INDONESIE | Coll. T. LANDER [w, h/p]’, 2 &female;&female; (MHNG, ex coll. Lander). Potil-Kecil Island (ca 3 km NE of Banggai Island): ‘ &female; [w, p] || BANGGAI ARCH., | Potil Kecil, | 1°28'S – 123°34'E | 12–19.ii.1980 [w, p] || M.J.D.Brendell | B.M.1980–280 [w,p] || Chrysodema | antennata Saund.? | B. Levey det. 1982 [w, h/p]’, 1 &female; (BMNH). Peleng Island: ‘INDONESIA, prov. | Central Sulawesi, | PELENG isl., | native coll. [w, p] || ex coll. V. Kubá&ncaron; | National Museum | Prague, Czech Republic [w, p]’, 1 &female; (NMPC, ex coll. Kubá&ncaron;); ‘PELENG IS | INDONESIA [w, h (black border)] || Chrysodema | obsoleta | Thoms., 1878 | Dét. T. LANDER 192000 [w, h/p]’, 1 &female; (MHNG); ‘INDONESIA | CELEBES | Peleng | Coll. T. LANDER [w, h/p]’, 3 &female;&female; (MHNG, ex coll. Lander); ‘INDONESIA | CELEBES | Peleng Isl. | Coll. T. LANDER [w, h/p]’, 7 &female;&female; (MHNG, ex coll. Lander); ‘Peleng | March. [20]06 [w, p]’, 1 &female; (DFPC); ‘Peleng isl. | 10/2017 [w, p]’, 3 &female;&female; (DFPC); ‘Peleng | NOV. 2018 [w, p]’, 1 &female; (DFPC); ‘PELENG IS [w, p (green stamp)]’, 1 &female; (DFPC); ‘PELENG IS [w, p (green stamp)]’, 14 &female;&female; (SJPC); ‘PELENG IS | V.2019 [w, p/h (violet stamp)]’, 1 &female; (DFPC); ‘PELENG IS | V.2019 [w, p/h (green stamp)]’, 13 &female;&female; (DFPC); ‘INDONESIA: Sulawesi | Peleng Island | native leg. | Coll. B. & T. Liese [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p] || Chrysodema | spec. | det. S. Gottwald 2001 [w, p]’, 1 &male; (SGBG). SOUTH SULAWESI PROv.: ‘INDONESIA, | South Sulawesi, | Puncak, PALOPO, | Mar.2000, native coll. [w, p] || ex coll. V. Kubá&ncaron; | National Museum | Prague, Czech Republic [w, p]’, 1 &male;, 3 &female;&female; (NMPC, ex coll. Kubá&ncaron;); ‘Puncak | Palopo | SULAWESI | IX.1999 | Coll. T. LANDER [w, p]’, 5 &female;&female; (MHNG, ex coll. Lander); ‘Puncak | Palopo | CELEBES | IX.1999 | Coll. T. LANDER [w, p]’, 12 &female;&female; (11 &female;&female; MHNG, ex coll. Lander, 1 &female; DFPC); ‘Puncak | Palopo | CELEBES | XII.1999 | Coll. T. LANDER [w, p]’, 3 &female;&female; (MHNG, ex coll. Lander); ‘Palopo | Puncak | CELEBES | XII.1999 | Coll. T. LANDER [w, p]’, 1 &female; (MHNG, ex coll. Lander); ‘Puncak | Palopo | CELEBES | I.2000 | Coll. T. LANDER [w, p]’, 17 &female;&female; (16 &female;&female; MHNG, ex coll. Lander, 1 &female; DFPC); ‘ CELEBES | Puncak Palopo | 2000 | Coll. T. LANDER [w, h/p] || 5.2.2003 | Puncak Palopo | antenata | Coll. T. LANDER [w, h/p]’, 1 &male; (MHNG, ex coll. Lander); ‘Puncak Palopo | Maret—2000 [w, p]’, 1 &male; (DFPC); ‘ &female; [w, p] || INDONESIEN | Sulawesi, Puncac Palopo | 8 &9 2001 leg Johan [w, p] || ex coll. | Barries [y, h/p] || CHRYSODEMA | celebensis Kerr. 1909 | det Barries 2004 [w, p]’, 1 &female; (DFPC); ‘ &female; [w, p] || INDONESIEN | Sulawesi, Puncac Palopo | 8 &9 2001 leg Johan [w, p] || coll. | Barries [y, p] || CHRYSODEMA | celebensis Kerr. 1909 | det Barries 2004 [w, p]’, 3 &female;&female; (WBWA); ‘PUNCAK | PALOPO | 3.2000 [w, h] || Chrysodema | antennata | Saund., 1874 | Dét. T. LANDER 192007 [w, h/p]’, 1 &female; (EJBS); ‘PUNCAK | PALOPO | SULAWESI | APRIL 1996 [w, p] || Chrysodema | sp. | det. S. Gottwald 2001 [w, p]’, 1 &female; (SGBG); ‘Rautcpao [?Rantepao] | CELEBES | XII.1999 | Coll. T. LANDER [w, h/p]’, 6 &female;&female; (5 &female;&female; MHNG, ex coll. Lander, 1 &female; DFPC); ‘Rautcpao [?Rantepao] | CELEBES | XII.1999 | Coll. T. LANDER [w, p] || 10.4.2000 | CELEBES | &female; (1) 10'KOH | Dét. T. LANDER [w, h/p]’, 1 &female; (MHNG, ex coll. Lander); ‘INDONESIA, X.2017 | Central Sulawesi | PUNCAK PALOPO pass | local collector lgt. [w, p]’, 3 &female;&female; (SJPC). WEST SULAWESI PROv.: ‘Mamasa | Sulawesi | Jan. '03 [w, p] || ex Coll. | M. H. HEMPEL | (München) [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p]’, 2 &female;&female; (SGBG); ‘Mamasa | Sulawesi | Jan. '04 [w, p] || ex Coll. | M. H. HEMPEL | (München) [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p]’, 5 &female;&female; (SGBG). Sulawesi Island (without accurate locality): ‘ CELEBES [w, h]’, 6 &female;&female; (MHNG); ‘ SULAWESI | local collectors leg. | Coll. B. & T. Liese [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p] || C [w, h (blue letter)] || Chrysodema | sp. | det. S. Gottwald 2002 [w, p] || antennata | ssp. obsoleta [w, h]’, 1 &female; (SGBG); ‘INDONESIA: Sulawesi | native leg. 1999 | Coll. B. & T. Liese [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p] || 1 [w, h (blue letter)] || Chrysodema | sp. n. Lander in prep. | det. S. Gottwald 2000 [w, p]’, 1 &female; (SGBG); ‘ INDONESIA: Sulawesi | native leg. 1999 | Coll. B. & T. Liese [w, p] || Coll. | S. GOTTWALD | (Berlin) [w, p] || 7 [w, h (blue letter)] || Chrysodema | sp. n. Lander in prep. | det. S. Gottwald 2000 [w, p]’, 1 &female; (SGBG). Original specimens of unpublished manuscript names (see ‘Remarks’). Chrysodema grootaerti Ho&lstrok;y&nacute;ski (5 &female;&female;): ‘Bangkei | H. Kühn | 1885. [w(c), p; black border] || antennata | Saund [w, h] || 1993 | Chrysodema | grootaerti HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h/p] || Chrysodema | grootaerti | HO&Lstrok;Y&Nacute;SKI | HOLOTYPE [r, h]’, 1 &female; (IRSN); ‘[original label] Bangkei | H. Kühn | 1885. [w(c), p; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w, p; black border] [original label is glued on newer label] || 3458 [w, h] || Chrysodema | borneensis Kerr. | Det. Hoschek. 1941 [w, h/p; black border] || 1993 | Chrysodema | grootaerti HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h/p] || Chrysodema | grootaerti | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’, 1 &female; (IRSN); ‘[original label] Bangkei | H. Kühn | 1885. [w(c), p; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w, p; black border] [original label is glued on newer label] || 3884 [w, h] || Chrysodema | antennata | Saund | Théry det. [w, h/p; black border] || 1993 | Chrysodema | grootaerti HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h/p] || Chrysodema | grootaerti | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’, 1 &female; (IRSN); ‘Bangkei | H. Kühn | 1885. [w(c), p (black border)] || coll. R B HO&Lstrok;Y&Nacute;SKI | BPexf [w, h] || 1993 | Chrysodema | grootaerti HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h] || Chrysodema | grootaerti | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’, 1 &female; (RHMP); ‘Bangkei | H. Kühn | 1885. [w(c), p (black border)] || coll. R B HO&Lstrok;Y&Nacute;SKI | BPexg [w, h] || Chrysodema | grootaerti | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’, 1 &female; (RHMP). All specimens were provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] | Chrysodema (Chrysodema) | sekerkai sp. nov. | David Frank det. I. 2022 [date handwritten]’. Specimens ‘ Chrysodema grootaerti Ho&lstrok;y&nacute;ski’ were provided with an additional red printed label: ‘ Chrysodema | grootaerti | HO&lstrok;Y&nacute;SKI | unpublished manuscript name | David Frank I. 2022 [date handwritten]’. Description of holotype. Well preserved &male; specimen with all appendages, only one of claws on left fore leg and last antennomere of right antenna are missing. Length 22.75 mm, width 8.25 mm, length/width ratio: 2.76. Body navicular, basic colour of dorsal side black but it seems dark green because of dense metallic green punctures; ventral side green with golden reflections, legs green. Head black with green punctures, green on ventral side. Eyes large, oval, area around eyes with dense fine green macropunctures and sparsely pubescent. Frons black, with sparse green macropunctures, 2.1× as wide as diameter of eye, impressed with deep medial sulcus, with sparse short pubescence. Labrum dark brown, with long pubescence.Antennae longer than pronotum, serrate from antennomere IV. Antennomere I green, II bronze, sparsely macropunctate and pubescent; radicula and antennomeres from III dark brown with long pubescence. Antennomere II ca. 4× shorter than I and III. Antennomeres IV–X long triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi brown, densely pubescent. Pronotum trapezoidal, narrowing anteriad with rounded sides, black, moderately densely irregularly goldengreen macropunctate, 1.4× as wide as long, widest at mid-length. Anterior margin arcuate, lobe not protruding, densely pubescent. Medial line visible, interrupted at centre, without medial impressions. Principal impressions oval, golden-green, densely macropunctate. Lateral impressions moderately deep, densely macropunctate. Lateral margin green, shiny. Basal margin moderately bisinuate. Scutellum small, trapezoidal, green with golden reflections. Elytra moderately wider than pronotum at base; subparallel at basal half, narrowing from beyond its midlength to apex, strongly convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Groups of macropunctures of almost the same size contain ca 3–8 macropunctures, larger on lateral parts, arranged in almost regular rows. Black areas between rows of groups of macropunctures appear as weak costae which are interconnected at short intervals, along suture golden-green stripes with individual macropunctures. Last two lateral intervals coherently very densely macropunctate therefore almost entire area is green with golden reflections. Humeral calli golden-green, densely macropunctate. Epipleura horizontal, broad at basal third, green with golden reflections, moderately densely macropunctate and pubescent. Legs metallic green with golden reflections; femora and tibiae macropunctate and pubescent more densely on inner side. Mesotibiae curved at apical third and strongly bent at basal third, without long pubescence at basal third. Tibiae with two apical ventral spurs. Tarsi metallic green, tarsal claws divergent and simple. Ventral side metallic green with golden reflections. Hypomeron and prosternum macropunctate and with short pubescence. Anterior margin of prosternum with dense short pubescence. Prosternal process subparallel-sided, narrowed at apex, approximately 2× as long as wide, moderately densely and coarsely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, apex and sides smooth, polished, nearly impunctate and sparsely pubescent. Metasternum sparsely finely punctate and shiny at central part; more densely on sides, pubescent. Abdominal ventrites I–III moderately densely punctate at central part, laterally more densely punctate, pubescent. Abdominal ventrites IV–V more densely punctate than I–III at central part. Aedeagus. Broadly rhomboidal, widest at mid-length, length: 6.14 mm, width: 1.41 mm, length/width ratio: 4.35. Parameres narrowly open at apex, apices of parameres rounded (Fig. 68). Variation. Body &male;&male; (n = 5) length: 20.00–22.75 (average 21.50) mm, width: 7.50–8.25 (average 7.95) mm, length/width ratio: 2.67–2.76 (average 2.71); &female;&female; (n = 129): length: 22.00–30.50 (average 26.44) mm, width: 8.25– 11.75 (average 10.16) mm, length/width ratio: 2.51–2.74 (average 2.60). Groups of macropuctures can be variable but never large. Colour of macropunctures variable from golden-green to dark green-brown, very rare dark brown. Aedeagus (n = 4) length: 5.82–6.14 mm, width: 1.36–1.41 mm, length/width ratio: 4.28–4.48. Differential diagnosis. Males of C. (C.) sekerkai have antennae longer than pronotum, broadly rhomboidal aedeagus and strongly bent basal third of mesotibiae without long pubescence. Females can be distinguished by bent basal third of mesotibiae (Fig. 74) and from C. (C.) holynskii also by black basic colour of elytra between groups of macropunctures. Macropunctures on pronotum generally coarser than in C. (C.) antennata obsoleta and with golden-green margin. For additional characters see Key to species. Etymology. This species is dedicated to my good friend Lukáš Sekerka (Prague, Czech Republic), a wellknown specialist in world Cassidinae and Hispinae (Chrysomelidae)... and now also in Buprestidae. Distribution. Indonesia: Banggai, Potil-Kecil, Peleng and Sulawesi islands (Fig. 62). Remarks. I studied five specimens deposited in IRSN and RHMP labelled as holotype and paratypes of ‘ Chrysodema grootaerti Ho&lstrok;y&nacute;ski’. This name has never been published (R. Ho&lstrok;y&nacute;ski pers. comm. September 2021) and all specimens belong to the C. (C.) sekerkai sp. nov., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 534-537, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp."]}
Published: 2022
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2. Chrysodema (Chrysodema) eximia subsp. eximia Laporte de Castelnau & Gory 1835

Author: Frank, David
Subjects: Coleoptera, Chrysodema (chrysodema) eximia eximia laporte de castelnau & gory, 1835, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema eximia, Taxonomy
Abstract: Chrysodema (Chrysodema) eximia eximia Laporte de Castelnau & Gory, 1835 (Figs 1–10, 12, 13–17, 63, 64) Chrysodema eximia Laporte de Castelnau & Gory (1835): 8 (original description, incl. colour Fig. 9, Pl. 2); Deyrolle (1864): 14 (key), 20 (listed); Saunders (1871): 13 (catalogue); Kerremans (1892a): 38 (catalogue); Kerremans (1903): 75 (catalogue); Akiyama & Ohmomo (2000): Pl. 46, Figs 486-1, 486-2 (iconography). Chalcophora eximia: Lacordaire (1857): 23 (noted); Gemminger & Harold (1869): 1357 (catalogue). Chrysodema (Chrysodema) eximia: Kerremans (1909): 515 (key), 531 (redescription); Obenberger (1926): 131 (catalogue); Frank & Sekerka (2020): 8 (noted). Chrysodema (Chrysodema) eximia eximia: Lander (2003): 15 (key), 31 (redescription), 77 (colour Figs 48–51); Bellamy (2008): 530 (catalogue). Chrysodema intercostata Saunders (1874): 308 (original description); Kerremans (1892a): 38 (catalogue); Kerremans (1903): 76 (catalogue). Chalcophora intercostata: Kerremans (1885): 126 (catalogue). Chrysodema (Chrysodema) intercostata: Kerremans (1909): 515 (key), 529 (redescription); Obenberger (1926): 131 (catalogue); Lander (2003): 31 (as synonym of C. (C.) eximia), 77 (colour Fig. 51); Bellamy (2008): 530 (catalogue, in synonymy C. (C.) eximia), 532 (listed as synonym of C. (C.) eximia). Chrysodema eximia kawamurai Kurosawa (1989): 31 (original description, incl. Figs 3–4). Chrysodema (Chrysodema) eximia kawamurai: Lander (2003): 31 (as synonym of C. (C.) aurostriata); Bellamy (2008): 530 (catalogue, in synonymy C. (C.) eximia), 532 (listed as synonym of C. (C.) eximia). Type localities. Chrysodema eximia: ‘Indes-Orientales’ (see ‘Remarks’); C. intercostata: ‘N. Luzon’ [Philippines, Luzon Island (north)]; C. eximia kawamurai: ‘ Camiguin Island, Babuyan Islands, N. Philippines’ [Philippines, Cagayan Province, Camiguin Island]. Type material examined. Chrysodema eximia: SYNTYPES (3 &female;&female;): &female; (25.50 × 9.25 mm; Figs 1–3, 6), ‘ Chrysodema | Eximia Cast et Gory |Amboine [w, h; yellow border] || Coll. | Janson [w, p; black border] || MUSÉUM PARIS | 1935 | Coll. A. THÉRY [w, p]’ (MNHN); in front of row of specimens: ‘Eximia | Gory | Amboine Type [w(c), h; yellow border]’; under specimens: &female; (28.25 × 10.00 mm; Figs 4, 7), ‘TYPE [r, p] || MUSÉUM PARIS | 1952 | COLL. R. OBERTHUR [y, p] || SYNTYPE [r, p] || SYNTYPE | Chrysodema | eximia Laporte & Gory, 1835 [w, p] || MNHN | EC8630 [w, p]’ (MNHN); &female; (30.50 × 11.00 mm), ‘TYPE [r, p] || MUSÉUM PARIS | 1952 | COLL. R. OBERTHUR [y, p]’ (MNHN). All specimens were provided with an additional red printed label: ‘ SYNTYPE &female; | Chrysodema | eximia | LAPORTE DE CASTELNAU & GORY, 1835 | David Frank labelled IV. 2021 [date handwritten]’. Chrysodema intercostata: LECTOTYPE (present designation): &female; (31.75 × 11.50 mm; Figs 16, 25), ‘ N Luzon | 137 [w(c), h; round] || [2x] Type [w, p; round; red border] || Saunders | 74-18. [w(c), p] || intercostata | ES. Type [w(c), h]’ (BMNH). PARALECTOTYPE: &female; (31.25 × 11.50 mm), ‘ N Luzon | 137 [w(c), h; round] || COTYPE [w(c), p (red letters)] || Saunders | 74-18. [w(c), p] || MUSÉUM PARIS | 1935 | Coll. A. Théry [w(c), p; black border] || intercostata | Saund. | cotype [w(c), h]’ (MNHN). Both specimens were provided with an additional red printed label: ‘ LECTOTYPE [or PARALECTOTYPE respectively] &female; | Chrysodema | intercostata | SAUNDERS, 1874 | David Frank des. IV. 2021 [date handwritten]’ and also white printed label ‘ Chrysodema (Chrysodema) | eximia eximia | LAPORTE DE CASTELNAU & GORY, 1835 | David Frank det. IV. 2021 [date handwritten]’. Chrysodema eximia kawamurai: PARATYPE: &female; (27.00 × 9.50 mm; Figs 5, 8), ‘ Camiguin Is. | Babuyanes Iss. | Philippines | VI. 1988 [w, h] || PARATYPE [r, p] || ex coll. S. Bílý | National Museum | Prague, Czech Republic [w, p] || Chrysodema | eximia | kawakamii [sic!, see ‘Remarks’] | DET. K. Akiyama. 1993 [w, h/p]’ (NMPC). The specimen was provided with additional red printed labels ‘ PARATYPE &female; | Chrysodema | eximia | kawamurai | KUROSAWA, 1989 | David Frank labelled IV. 2021 [date handwritten] || ‘ kawakami ’ is only misinterpretation | by K. Akiyama. Acording to locality | data and measures this specimen | belongs to the type series of | Chrysodema eximia kawamurai | David Frank IV.2021 [r, p; date handwritten]’ (see ‘Remarks’) and white printed label ‘ Chrysodema (Chrysodema) | eximia eximia | LAPORTE DE CASTELNAU & GORY, 1835 | David Frank det. IV. 2021 [date handwritten]’. Original specimens of unpublished manuscript names (see ‘Remarks’). [dark specimens are marked ‘(d)’] Chrysodema eximia ab. atrocyanea: &female; (d) (33.75 × 11.50 mm; Figs 9, 13), ‘C.Engano, [Cape Engano, Palaui Is.] | N.Luzon | Whitehead leg. [w(c), p] || 2500 [gray, h] || TYPUS [w(c), p; black border] || Coll. G. Frey | NMB [blue, p] || Chrysodema | eximia C. G. | ab. | atrocyanea m. | Det. Hoscheck 19 34. [w(c), h/p; black border] || Chrysodema | eximia C.& G. | Dét. T. LANDER 1999 [w, h/p]’ (NHMB). Chrysodema eximia ab. purpureocuprea (2 &female;&female;): &female; (d) (30.25 × 10.25 mm), ‘C.Engano, | N.Luzon | Whitehead leg. [w(c), p] || 2499 [gray, h] || TYPUS [w(c), p; black border] || Chrysodema | eximia C. G. | purpureocuprea | m. n. ab. | Det. Hoscheck 19 41. [w(c), h/p; black border] || Chrysodema | eximia C.G. | det. R. Holynski 1999 [w, h/p]’ (IRSN); &female; (d) (34.00 × 11.75 mm; Figs 10, 14), ‘C.Engano, | N.Luzon | Whitehead leg. [w(c), p] || 2498 [gray, h] || TYPUS [w(c), p; black border] || Coll. G. Frey | NMB [blue, p] || Chrysodema | eximia C. G. | purpureocuprea | m. n. ab. | Det. Hoscheck 19 41. [w(c), h/p; black border] || Chrysodema | eximia C.& G. | Dét. T. LANDER 1999 [w, h/p]’ (NHMB). All specimens were provided with additional red printed labels ‘ Chrysodema eximia | ab. atrocyanea [or purpureocuprea respectively] | HOSCHECK | unpublished manuscript name | David Frank IV.2021 [r, p; date handwritten]’ and white printed label ‘ Chrysodema (Chrysodema) | eximia eximia | LAPORTE DE CASTELNAU & GORY, 1835 | David Frank det. IV. 2021 [date handwritten]’. AddiTiOnal maTerial examined (6 &male;&male;, 85 &female;&female;). [dark specimens are marked ‘(d)’] PHILIPPINES: ‘ Philipp [w, h] || TYPE [r, p] || MUSÉUM PARIS | 1952 | COLL. R. OBERTHUR [y, p]’ &female; (28.00 × 10.00 mm) (MNHN, ex coll. Oberthür) (labelled as type, see ‘ Remarks’); Philippines, Eydoux 118–38, 1 &female; (MNHN, ex coll. Oberthür); Philippines, Ch. Semper, 1 &female; (MNHN, ex coll. Oberthür); Philippines, Stauding [er], 1 &female; (MNHN, ex coll. Kerremans); Ins. Philipp., Coll. Janson, 1 &female; (MNHN, ex coll. Théry); Philip, 1 &female; (BMNH, ex coll. Saunders); Phillipine Islands, H. Cuming Esq. [determined as C. intercostata], 1 &female; (BMNH, ex coll. Saunders); Phillipine Islands, H. Cuming Esq., Bowring 63–47, 1 &female; (d) (BMNH); Philippinen, 2 &female;&female; (MFNB, ex coll. Moser); Philippinen, 1 &female; (MFNB); Philippines, 1 &female; (d) (MHNG, ex coll. Lander). LUZON ISLAND : Philippines, Luçon, Marche 1880, 1 &female; (MNHN); Phil, Luzon 56/93, 1 &female; (BMNH, ex coll. Saunders); Luzon /139, 1 &female; (RHMP); Luzon /139, 1 &female; (d) (BMNH, ex coll. Saunders); Philippines, S Luzon, vi.92, 1 &female; (NMPC, ex coll. Bílý); Ins. Luzon, 1 &female; (MFNB, ex coll. Moser); Philippines, Luzon [determined as C. intercostata], 1 &female; (RHMP); Insel Luzon, v.2007, 1 &female; (WBWA). Manila (see ‘ Remarks’): Manille, Ex Musaeo J. Thomson, 1 &female; (MNHN, ex coll. Théry); Manille, Ex Musaeo Mniszech [determined as subviolacea (Laferté)], 1 &female; (d) (MNHN, ex coll. Oberthür); Manilla, Parry, 1 &female; (BMNH, ex coll. Saunders); Manila, 1 &female; (MFNB, ex coll. Hauser). Abra Prov.: Tineg, May 2018, 1 &female; (d) (DFPC). Apayao Prov.: Conner, Sept [ember] 2014, 1 &female; (d) (WBWA). Aurora Prov.: Dingalan, vi.2012, 1 &male; (EJBS; Figs 15, 63); Sierra Madre, Dingalan, October 2018, 1 &female; (d) (DFPC); Sierra Madre, Casiguran, November 2018, 1 &female; (DFPC); Didin, June 2019, 1 &female; (DFPC); Dingalan, January 2020, 1 &female; (DFPC); Dingalan, June 2020, 1 &female; (DFPC); Dingalan, April 2021, 1 &female; (DFPC). Bulacan Prov.: Philippinen, Luzon, Sibul, 16- 19.6.1986, 1 &female; (MFNB, ex coll. Moser). Cagayan Prov.: C. Engano, [Cape Engano, Palaui Is.], N. Luzon, Whitehead leg., 1 &female; (red-bronze) (MFNB, ex coll. Moser); Sta. Ana, May 2018, 1 &male; (d) (DFPC); Sta. Ana, August 2018, 1 &female; (d) (DFPC). Ilocos Norte Prov.: Luzon P. I., Ilocos Norte, W. Jark leg., 1 &female; (IRSN); N. Luzon, Bangui, 1 &male; (NMPC). Isabela Prov.: San-Pablo, Sept [ember] 2014, 1 &female; (d) (WBWA); Sierra Madre, Cabagan, October 2018, 1 &female; (d) (DFPC). Laguna Prov.: P.I., Los Banos, Baker, 2 &female;&female; (MNHN (1 &female;, ex coll. Kerremans)); P.I., Laguna, Los Banos, 1 &female; (NMPC); P.I., Laguna, Mt. Makiling, 16.vii.[19]30, collected by F.C.Hadden, 1 &female; (NMPC); P.I., Luzon, Laguna, Mt. Makiling, 24.vi.[19]31, collected by F.C.Hadden, 1 &female; (NMPC); P.I., Los Baños, 15-vii-1923, E. Mangapit, 1 &female; (NHMB, ex coll. Frey). Laguna / Batangas Prov. (see ‘ Remarks’) : Philippines, Mt. Maquiling, Elev. 50 met, 24.vi.[19]46, Q. Tagorota, 1 &female; (NMPC); Philippines, Mt. Maquiling, Elev. 50 met, 10.xii.[19]46, S. Cabañgisan, 1 &female; (NMPC); Mt. Maquiling, Elev. 50 met, 22-ii-1947, M.Sandoval, 1 &female; (NMPC); Philippines, Mt. Maquiling, Elev. 600 met, 18.ix.[19]48, R.P. Rivera, 1 &female; (NMPC). Mountain Prov.: N Luzon, Mountain Prov., vi.[19]91, 2 &female;&female; (NMPC, ex coll. Bílý); Nord Luzon, Mt. Provinz, June 1991, 1 &male;, 1 &female; (ATMR, ex coll. Novak); Mountain Prov., Aug [ust] 1991, coll. Schulz, 2 &female;&female; (ATMR, ex coll. Novak); North Luzon, Mountain Province, June 1996, 1 &female; (SGBG). Nueva Ecija Prov.: Caranglan, July 2019, 1 &female; (DFPC). Nueva Vizcaya Prov.: Belance, Bambang, June 2012, (1 &female; TTOC; 1 &female;, 1 &female; (d) DFPC); Belance, Jul [y] 2013, 1 &female; (d) (TTOC); Belance, Aug [ust] 2013, 1 &female; (DFPC); Kayapa, Sept [ember] 2014, 1 &female; (WBWA). Pangasinan Prov.: Umigan, Elev. 250 m., 18-vi-1949, F. Badua, 1 &female; (NMPC, ex coll. Bílý). Quezon Prov.: Mt.Banabao, Luzon, 1 &male; (RHMP); Luzon, Novaliches, nördl. v. Manila. ii.[19]18, 1 &female; (RHMP). Quirino Prov.: Sierra Madre, Madela, Disimungal, October 2017, 1 &male; (Figs 17, 64), 1 &female; (DFPC); Sierra Madre, Madela, San Martin, April 2018, 1 &female; (d) (DFPC). Rizal Prov.: Montalban, Coll. Dr. A. Frh. v.Hoscheck, 1 &female; (IRSN); Luzon, Montalban, 1914, leg. G. Boettcher, 2 &female;&female; (MFNB, ex coll. Moser); Sierra Madre, Rodriguez, December 2017, 1 &female; (DFPC). BABUYAN ISLANDS: Babuyan Island: N-Luzon, Babuyan Isl., Sept [ember] [20]'04, (4 &female;&female; SGBG; 1 &female; DFPC); Babuyan Isl., 1 &female; (d) (MHNG, ex coll. Lander). Camiguin Island: Cagayan Prov., Camiguin Isl., 4.[20]'00, 1 &female; (SGBG); N-Luzon, Camiguin Isl., Sept [ember] [20]'04, (9 &female;&female; SGBG; 1 &female; DFPC). Redescription of syntype. Well preserved &female; specimen, right antenna missing from antennomere VII and slightly open elytra. Length 25.50 mm, width 9.25 mm, length/width ratio: 2.76. Body navicular, elytra and pronotum bright metallic green with golden and copper reflections; elytra towards apex gradually from more intensive copper to dark. Head, ventral side and legs brown with red-coppery reflections. Head brown, densely pubescent in lower half. Eyes large, oval, area around eyes finely macropunctate and pubescent in lower half. Frons with coppery-red reflections, 2× as wide as diameter of eye, impressed with medial sulcus, macropunctate, sparsely pubescent. Labrum brown, pubescent. Antennae brown, serrate from antennomere IV. Antennomeres I (scape) and II (pedicel) dark brown, sparsely macropunctate and pubescent; radicula (base of scape) and antennomeres from III pale brown, sparsely pubescent. Antennomere II ca. 3× shorter than I and III. Antennomere IV triangular, V–X more trapezoidal than triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi pale brown, densely pubescent. Pronotum trapezoidal, narrowing anteriad with almost equal sides, widest at base, green, 1.6× as wide as long. Anterior margin arcuate with moderately protruding lobe, densely pubescent. Pronotum densely coarsely macropunctate, without larger impunctate areas except for medial line. Pronotal surface except for medial line appears quite rugose due to deeply impressed macropunctures. Macropunctation gradually coarser and denser toward sides. Medial line well visible, flat, slightly elevated, sparsely micropunctate, with several macropunctures. Medial impressions weak, barely impressed. Principal impressions small, elongate, shallow. Lateral impressions moderately deep, with several macropunctures and numerous very distinct micropunctures. Lateral margin brownbronze. Basal margin bisinuate. Scutellum small, trapezoidal, green with gold-red reflections. Elytra slightly wider than pronotum at base; parallel at basal half, narrowing from mid-length to apex; moderately convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Serrations and apical quarter of suture coppery-red, apex of elytra black. Four distinctly elevated sharp costae on each elytron less distinct at basal third. 1 st costa reaches to apex of elytra where it is joined to suture, 2 nd joined to 1 st at apical fifth. 3 rd costa separated, shortened, reaching to 3/4 length of elytra. 4 th costa also separated but reaching to apex where it is almost joined to 1 st and suture. Basal third irregularly furrowed, more deeply toward base. Furrows and intercostal intervals very irregularly macropunctate and with short pubescence. Epipleura horizontal, broad at basal quarter, coppery-red, sparsely macropunctate and pubescent. Legs brown-violet; femora less, tibiae more macropunctate and pubescent. Tibiae with two apical ventral spurs. Tarsi metallic coppery-red, tarsal claws divergent and simple. Ventral side brown with red-coppery reflections. Hypomeron irregularly macropunctate more coarsely than prosternum.Anterior margin of prosternum densely pubescent.Prosternal process parallel-sided triangularly narrowed at apex, approximately 2.3× as long as wide, sparsely macropunctate and pubescent at central part, macropunctures bigger than macropunctures on prosternum, apex and sides smooth. Metasternum shiny, sparsely macropunctate at central part; not shiny, densely, coarsely macropunctate and pubescent on sides. Abdominal ventrites I–IV centrally sparsely macropunctate, laterally densely macropunctate in entire width, with short pubescence.Abdominal ventrites V densely macropunctate, with short pubescence, with shallow impression at apical part. Variation. Body &male;&male; (n = 6) length: 21.25–27.25 (average 25.21) mm, width: 7.25–9.25 (average 8.54) mm, length/width ratio: 2.89–2.95(3.15) (average 2.95); &female;&female; (n = 75): length: 23.50–34.50 (average 28.86) mm, width: 8.00–12.25 (average 10.12) mm, length/width ratio: 2.69–3.05 (average 2.85). Chrysodema (C.) eximia eximia is variable in colour from light green with golden reflections through dark green, green-red to bronze or brown. Some specimens can have well developed or slightly indicated intercostae. Well developed intercostae in all intercostal intervals are very rare and always thinner than main costae. Aedeagus (n = 4) length: 7.32–7.45 mm, width: 1.55– 1.59 mm, length/width ratio: 4.63–4.75. Parameres broadly regularly navicular widest at mid-length, narrowly open at apical third, apice of parameres rounded. Penis moderately impressed with densely arranged striate laminae at entire width, apex of penis narrowly triangular (Figs 63, 64). Differential diagnosis. Specimens of C. (C.) eximia eximia with well developed intercostae can be similar to C. (C.) eximia berliozi but they have entire head concolourous with ventral side. Dark specimens can be distinguished from C. (C.) fuscitarsis by the 1 st costa parallel with suture and narrower body. Green specimens can be distinguished from C. (C.) dohrnii by dark ventral side. For additional characters see Key to species. Distribution. Philippines: Luzon, Palaui, Babuyan and Camiguin islands (Fig. 12). Remarks. Laporte de Castelnau & Gory (1835) did not specify explicitly how many specimens they had at their disposal when describing Chrysodema eximia. They mentioned only depository in ‘Du cabinet des auteurs’, a single length and width measurement ‘Long. 11 lig. ½. Larg. 4 lig. (25.99 × 9.04 mm)’ and a locality ‘Indes Orientales’. I found three female specimens labelled as types in MNHN but none of them have the measurements indicated in the description. These specimens are from Oberthür’s collection where there is an old handwritten yellow bordered label ‘ Eximia, Gory, Amboine, Type’ in front of row of specimens and newer red printed labels ‘TYPE’ under each specimen. Two of them have no locality labels and the third is localised from Philippines (‘Philipp’). According to locality mentioned in the description Laporte de Castelnau & Gory probably did not know that the specimens are from Philippines because they mentioned ‘Philippines’ only for Chrysodema philippinensis Laporte de Castelnau & Gory, 1835 and C. rouxi Laporte de Castelnau & Gory, 1835. Therefore the specimen with locality label ‘Philipp’ is not included in the type series. I found also a specimen labelled ‘ Chrysodema | Eximia Cast et Gory | Amboine [w, h; yellow border; the label was originally in front of the specimen judging by holes after pins] || Coll. | Janson [w, p; black, Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 509-518, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Laporte Comte de Castelnau, F. L. N. & Gory, H. L. (1835) Chrysodema. In: Dumenil, P. (Ed.), Histoire naturelle de iconographie des insectes Coleopteres. Monographie des buprestides. Tome 1. J. B. Bailliere, Paris, pp. 1 - 27. https: // doi. org / 10.5962 / bhl. title. 110141","Deyrolle, H. (1864) Description des Buprestides de la Malaisie recueillis par M. Wallace. Annales de la Societe entomologique de Belgique, 8, 1 - 272., 4 pls. https: // doi. org / 10.5962 / bhl. title. 115963","Saunders, E. (1871) Catalogus Buprestidarum Synonymicus et Systematicus. J. Janson, London, 171 pp.","Kerremans, C. (1892 a) Catalogue synonymique des Buprestides decrits de 1758 a 1890. Memoires de la Societe entomologique de Belgique, 1, 1 - 304.","Kerremans, C. (1903) Coleoptera Serricornia. Fam. Buprestidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","Akiyama, K. & Ohmomo, S. (2000) The buprestid beetles of the world. In: Fujita, H. (Ed.), Mushi-Shaʼs Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, pp. 1 - 341., 30 pls.","Lacordaire, J. T. (1857) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique de critique de tous les genres proposes jusquʼici dans cet ordre dʼinsectes. Tome quartreme contenant les familles des buprestides, throscides, eucnemides, elaterides, cebrionides, cerophytides, rhipicerides, dascyllides, malacodermes, clerides, lymexylones, cupesides, ptiniores, bostrichides et cissides. Roret, Paris, 579 pp. https: // doi. org / 10.5962 / bhl. title. 8864","Gemminger, M. & Harold, E. von (1869) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus 5. Buprestidae, Trixagidae, Monommidae, Eucnemidae, Elateridae, Cebrionidae. E. H. Gummi, Monachii, 262 pp. [pp. 1347 - 1608] https: // doi. org / 10.5962 / bhl. title. 9089","Kerremans, C. (1909) Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars). Livraisons 13 - 19. J. Janssens, Bruxelles, pp. 385 - 604., pls. 17 - 22. https: // doi. org / 10.5962 / bhl. title. 9700","Obenberger, J. (1926) Buprestidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, pp. 1 - 212.","Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp.","Saunders, E. (1874) Notes on the Buprestidae collected by Professor Semper in the Philippines Islands; with descriptions of the new species. Transactions of the Royal Entomological Society of London, 1874 (2), 303 - 328.","Kerremans, C. (1885) Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique, 29, 119 - 157.","Kurosawa, Y. (1989) A list of the Buprestidae (Coleoptera) collected by Mr. S. Kawamura on the Island of Camiguin of the Babuyans, the Philippines, with descriptions of new taxa. Bulletin of the National Science Museum, Tokyo, Series A, Zoology, 15 (1), 29 - 32.","Descarpentries, A. (1948) Note sur deux Buprestides asiatiques. Bulletin de la Societe entomologique de France, 53 (4), 62 - 64. https: // doi. org / 10.3406 / bsef. 1948.16025","Horn, W. & Kahle, I. (1935) Uber entomologische Sammlungen, Entomologen & Entomo-Museikigie (Ein Beitrag zur Geschichte der Entomologie). Teil I. Entomologische Beihefte, Dahlem, 2, i - vi + 1 - 160., pls. 1 - 16.","Cambefort, Y. (2006) Des coleopteres, des collections et des hommes. Publications Scientifiques du Museum National dʼHistoire Naturelle, Paris, 375 pp. https: // doi. org / 10.4000 / books. mnhn. 2265","Kuban, V. (2016) Buprestidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Revised and updated edition. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden-Boston, pp. 19 - 27 + 456 - 461. https: // doi. org / 10.1163 / 9789004309142","Frank, D. & Sekerka, L. (2016) Revision of the subgenus Gelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae). Acta Entomologica Musei Nationalis Pragae, 56 (2), 671 - 719.","Gallagher, S. J., Kitamura, A., Iryu, Y., Itaki, T., Koizumi, I. & Hoiles P. W. (2015) The Pliocene to recent history of the Kuroshio and Tsushima Currents: a multi-proxy approach. Progress in Earth and Planetary Science, 2 (17), 1 - 23. https: // doi. org / 10.1186 / s 40645 - 015 - 0045 - 6"]}
Published: 2022
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3. Chrysodema (Chrysodema) antennata subsp. obsoleta J. Thomson 1878

Author: Frank, David
Subjects: Coleoptera, Insecta, Chrysodema antennata, Arthropoda, Chrysodema, Animalia, Chrysodema (chrysodema) antennata obsoleta j. thomson, 1878, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) antennata obsoleta J. Thomson, 1878 (Figs 41–44, 46, 62, 67, 73) Chrysodema obsoleta J. Thomson (1878): 20 (original description); Kerremans (1892a): 39 (catalogue); Kerremans (1903): 76 (catalogue). Chalcophora obsoleta: Kerremans (1885): 126 (catalogue). Chrysodema (Chrysodema) obsoleta: Kerremans (1909): 520 (key), 582 (redescription); Obenberger (1926): 132 (catalogue). Chrysodema (Chrysodema) antennata obsoleta: Lander (2003): 48 (revision, downgraded to subsp. of C. (C.) antennata), 80 (colour Fig. 96, misidentification = C. (C.) sekerkai); Bellamy (2008): 527 (catalogue), 534 (listed as subspecies of C. (C.) antennata). Type locality. ‘ Celebes’ [Indonesia: Sulawesi Island]. Type material examined. LECTOTYPE (present designation): &female; (22.00 × 7.00 mm; Figs 41–44), ‘Ex Musaeo | JAMES THOMSON [w(c), p (black border)] || TYPE [r, p] || Th. | TYPE [w, p (black border)] || Chrysodema | Obsoleta | (HDeyr) | Célèbes [y, h] || Obsoleta | (H. Deyr. M.s.s.) | Thoms. Type |T. B. 20. | Célèbes [w, h]’ (MNHN). The specimen was provided with an additional red printed label: ‘ LECTOTYPE &female; | Chrysodema | obsoleta | THOMSON J., 1878 | David Frank des. IV. 2021 [date handwritten]’ and also white printed label ‘ Chrysodema (Chrysodema) | antennata | obsoleta | THOMSON J., 1878 | David Frank det. IV. 2021 [date handwritten]’. AddiTiOnal maTerial examined (3 &male;&male;, 7 &female;&female;). INDONESIA: Sulawesi Island (without accurate locality): Célèbes, 1 &male; (MNHN; Figs 46, 67); Célebes, Carter, 1 &female; (MNHN, ex coll. Théry); Celebes, 2 &male;&male;, 1 &female; (MNHN, ex coll. Lander); Indonesia, Celebes, 1 &female; (DFPC). CENTRAL SULAWESI PROv.: Tonusu, b. Tentena, Poso-See, 21.- 26.4.1994, leg. Gerstmeier, Hiermeier, Romig, 1 &female; (DFPC). NORTH SULAWESI PROv.: Sulawesi Utara, Dumoga Bone N. P., 0°33'N 123°40'E, 4.iii.1985, Base Camp Site, Malaise trap by fallen tree, C.H.C. Lyal, 1 &female; (BMNH); Sulawesi Utara, Dumoga-Bone N. P., March 1985, Lowland forest edge ca 200m, Malaise trap, 1 &female; (BMNH). SOUTH SULAWESI PROv.: Patunuang, Jan. 1896, H. Fruhstorfer, 1 &female; (MNHN, ex coll. Van de Pool). Redescription of lectotype. Moderately well preserved &female; specimen with missing right antenna from antennomere VI, and right middle and left hind legs; elytra slightly open. Length 22.00 mm, width 7.00 mm, length/ width ratio: 2.75. Body navicular, basic colour of dorsal side black-violet with metallic green macropunctation; ventral side green with coppery reflections, legs green. Head black with green area around eyes and punctures on dorsal side, green on ventral side. Eyes large, oval. Area around eyes green, finely macropunctate. Frons black with small green spot at center, with very sparse green macropunctures, 2× as wide as diameter of eye, impressed with deep medial sulcus, with very sparse short pubescence. Vertex more densely macropunctate than frons. Labrum pale brown, pubescent. Antennae serrate from antennomere IV. Antennomere I green, II green-brown, sparsely macropunctate and with short pubescence; radicula and antennomeres from III dark brown with sparse short pubescence. Antennomere II ca. 3× shorter than I and 2× than III. Antennomere IV long triangular, V–X more trapezoidal than triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi pale brown, densely pubescent. Pronotum trapezoidal, narrowing anteriad with rounded sides, widest at mid-length, black, with moderately dense irregular green macropunctures, micropunctate between macropunctures, 1.5× as wide as long.Anterior margin arcuate, lobe moderately protruding, densely pubescent. Medial line black, micropunctate, without macropunctures. Medial impressions weak and shallow only indicated by denser macropunctation. Principal impressions almost triangular, green, densely macropunctate. Lateral impressions green, moderately deep, macropunctate. Lateral margin green. Basal margin moderately bisinuate. Scutellum broadly trapezoidal, green and shiny. Elytra moderately wider than pronotum at base; subparallel at basal half, narrowing from beyond its midlength to apex; convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Groups of macropunctures in basal half of almost the same size contain ca 5–10 macropunctures, arranged in almost regular rows. Black-violet areas between rows of groups of macropunctures look like weak costae which are interconnected at short intervals. Macropunctures at apical half in almost coherent rows between weak black-violet costae. Last two lateral intervals coherently very densely macropunctate therefore almost entire area is green. Humeral calli moderately furrowed. Epipleura horizontal, broad at basal quarter, green, moderately densely macropunctate and pubescent. Legs metallic green with bronze reflections; femora and tibiae macropunctate and pubescent more densely on inner side. Mesotibiae moderately curved at basal half and almost straight at apical half (Fig. 73). Tibiae with two apical ventral spurs. Tarsi metallic green with bronze reflections, tarsal claws divergent and simple. Ventral side green with bronze reflections. Hypomeron and prosternum macropunctate and with short pubescence. Anterior margin of prosternum with dense short pubescence. Prosternal process subparallel-sided, narrowed at apex, approximately 1.9× as long as wide, densely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, apex and sides smooth, polished, nearly impunctate and sparsely pubescent. Metasternum densely macropunctate and pubescent. Abdominal ventrites I–IV centrally less, laterally very densely macropunctate, macropunctures at central part bigger than at lateral parts. Abdominal ventrite V very densely macropunctate at whole area. All abdominal ventrites with dense and short pubescence. Variation. Body &male;&male; (n = 3) length: 22.50–22.75 mm, width: 8.00– 8.50 mm, length/width ratio: 2.68–2.81; &female;&female; (n = 7): length: 22.00–27.25 (average 25.19) mm, width: 8.00–10.25 (average 9.25) mm, length/width ratio: 2.66– 2.75 (average 2.70). Groups of macropuctures can be variable but never large. Colour of macropunctures variable from green to red-bronze. Aedeagus (n = 2) length: 5.82 mm, width: 1.09 mm, length/width ratio: 5.34. Aedeagus narrowly rhomboidal, widest at mid-length. Parameres narrowly open at apex (Fig. 67). Differential diagnosis. Males of all subspecies of C. (C.) antennata have very dense and long pubescence on inner side of mesotibiae from mid-length to tarsi. Males of C. (C.) antennata obsoleta have antennae shorter than pronotum. From C. (C.) antennata antennata can be distinguished by shape of aedeagus which is narrowly rhomboidal. From C. (C.) antennata barriesi can be distinguished also by narrowly rhomboidal aedeagus, by parameres which are open only at apex (Fig. 67) and by V-shaped border of metallic green area of male abdominal sternite IX. Females can be distinguished from C. (C.) sekerkai by mesotibiae which are not bent at apical third (Fig. 73). Macropunctures on pronotum generally finer than in C. (C.) sekerkai and without golden-green margin. Chrysodema (C.) antennata obsoleta is generally the smallest subspecies of C. (C.) antennata. For additional characters see Key to species. Distribution. Indonesia: Sulawesi Island (Fig. 62). Remarks. Although J. Thomson (1878) wrote in his description of C. obsoleta ‘Le plus petit type à nous connu de la coupe actuelle;... [The smallest type known to me;...]’ it is not clear from this sentence if he had only one specimen or if one of specimens was the smallest. He also mentioned only a single length and width measurement ‘Long. 21 mill.; lat. 7 mill.’ but the specimen in MNHN labelled as type has 22 × 8 mm (Lander (2003) also mentioned 22 mm). On the other hand, the specimen has old type labels and it is almost certain that the specimen is the (holo)type or one of the syntypes. Therefore this specimen is designated as a lectotype in order to conserve the status of this taxon and avoid any further misinterpretations., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 530-531, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Thomson, J. (1878) Typi Buprestidarum Musaei Thomsoniani. E. Deyrolle, Paris, 103 pp.","Kerremans, C. (1892 a) Catalogue synonymique des Buprestides decrits de 1758 a 1890. Memoires de la Societe entomologique de Belgique, 1, 1 - 304.","Kerremans, C. (1903) Coleoptera Serricornia. Fam. Buprestidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","Kerremans, C. (1885) Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique, 29, 119 - 157.","Kerremans, C. (1909) Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars). Livraisons 13 - 19. J. Janssens, Bruxelles, pp. 385 - 604., pls. 17 - 22. https: // doi. org / 10.5962 / bhl. title. 9700","Obenberger, J. (1926) Buprestidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, pp. 1 - 212.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp."]}
Published: 2022
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4. Chrysodema (Chrysodema) eximia subsp. berliozi Descarpentries 1948

Author: Frank, David
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Chrysodema (chrysodema) eximia berliozi descarpentries, 1948, Buprestidae, Chrysodema eximia, Taxonomy
Abstract: Chrysodema (Chrysodema) eximia berliozi Descarpentries, 1948 (Figs 12, 18–24, 65, 66) Chrysodema Berliozi Descarpentries (1948): 63 (original description); Akiyama & Ohmomo (2000): Pl. 46, Figs 487-1–487-4 (iconography). Chrysodema (Chrysodema) berliozi: Kurosawa (1954): 31 (redescription). Chrysodema (Chrysodema) eximia berliozi: Lander (2003): 31 (revision, downgraded to subsp. of C. eximia), 78 (colour Fig. 52); Mühle (2003): 45 (noted); Kubá&ncaron; (2006): 345 (catalogue); Bellamy (2008): 530 (catalogue); Qi, Ai & Song (2022): 361 (faunistics); Peng et al. (2022): 97 (redescription). Chrysodema (Cyalithoides) eximia berliozi: Kubá&ncaron; (2016): 460 (catalogue); Tamadera & Yoshitake (2017): 127 (faunistics). Chrysodema eximia berliozi: Ong & Hattori (2019): 20 (diagnosis, iconography). Type locality. ‘Kôtô-Sho (Botel Tobago) Formose’ [Taiwan, Orchid Island]. Type material examined. HOLOTYPE: &female; (34.00 × 12.00 mm), ‘KÔTÔ-&Scirc;HO | (Botel-Tobago Is.) | FORMOSA | IV. 1936 | COLL. Y. CHUJO [w(c), h; black border] || TYPE [r, p; black border] || 16 [w(c), h] || Chrysodema | Berliozi &female; |TYPE [red stamp] mihi | A. Descarpentries det [w, h/p]’ (MNHN). PARATYPE: &female; (30.25 × 11.00 mm; Figs 19–22), ‘ KÔTÔ-&Scirc;HO | (Botel-Tobago Is.) | FORMOSA | IV. 1936 | COLL. Y. CHUJO [w(c), h; black border; label identical to holotype] || Chrysodema | Berliozi &female; | PARATYPE [red stamp] mihi | A. Descarpentries. dét. [w, h/p]’ (NMPC). Both specimens were provided with an additional white printed label ‘ Chrysodema (Chrysodema) | eximia | berliozi | DESCARPENTRIES, 1948 | David Frank det. IV. 2021 [date handwritten]’. AddiTiOnal maTerial examined (4 &male;&male;, 30 &female;&female;). TAIWAN: Formosa, T. Kano, 1 &female; (MNCN, ex coll. Cobos); Yu Shan, vii.[20]06, J-M. Bousquet leg., 1 &female; (DFPC); Taiwan, Puli, Winkler, 1 &male;, 8 &female;&female; (Fig. 18) (ATMR, ex coll. Novak). Orchid Island: ‘KOTOSHO | (BOTEL-TOBAGO IS.) | FORMOSA | 20. VI-10. VII 1938 | COLL. M. CHUJO [w(c), p; black border] || Chrysodema | Berliozi | mihi | PARATYPE [red stamp] | A. Descarpentries. dét. [w, h]’ 1 &female; (MNHN) (labeled as paratype, see ‘ Remarks’); Lanyu Is., iv.1967, Coll. M. Chujo, 2 &female;&female; (MNHN); Tobago, 1 &female; (MNHN); Botel-Tobago I., 2.ix.1939, T. Kaneko, 1 &female; (NMPC); Botel-Tobago I., vi.1965, 1 &male; (NMPC, ex coll. Bílý); Botel-Tobago I., vii.1965, 2 &female;&female; (NMPC, ex coll. Bílý); Botel Tobako, 1 &male; (DFPC; Figs 23, 65); BotelTobago, 16.v.1957, 1 &female; (MNCN, ex coll. Cobos); Botel-Tobago, vii.1965, 2 &female;&female; (MNCN, ex coll. Cobos); Lanyu Is., v.1970, W. Chen leg., 1 &female; (MNCN, ex coll. Cobos); Lanyu Island, 25.5.1990, Coll. J. Dalihod, (3 &female;&female; NMPC, ex coll. Kubá&ncaron;; 1 &female; SGBG); Lan Yu Is., v.[19]59, 1 &female; (ATMR, ex coll. Novak); Lun-shu Is., 1 &male; (Figs 24, 66), 2 &female;&female; (ATMR, ex coll. Novak). WITHOUT LOCALITY DATA: R. I. Sc. N. B., I. G. 24.554, 2 &female;&female; (IRSN). Redescription of paratype. Well preserved &female; specimen, only left fore and middle tarsi partly missing. Length 30.25 mm, width 11.00 mm, length/width ratio: 2.75. Body navicular, dorsal side metallic blue-violet with greenish reflections at apical third and green at head; ventral side (including head) and legs brown-red. Head green on dorsal side, brown-violet on ventral side. Eyes large, oval, area around eyes finely macropunctate and sparsely pubescent. Frons green, with coppery-red reflections at lower part, 2× as wide as diameter of eye, impressed with medial sulcus, macropunctate, sparsely pubescent. Vertex green with coppery-red and bluish reflections, more coarsely macropunctate than frons. Labrum dark brown. Antennae serrate from antennomere IV. Antennomeres I and II dark green with violet reflections, sparsely macropunctate and pubescent; radicula and antennomere from III dark brown, sparsely pubescent. Antennomere II ca. 4× shorter than I and 3× than III. Antennomeres IV–VI triangular, VII–X more trapezoidal than triangular, XI oval. Pronotum trapezoidal, narrowing anteriad with almost equal sides, widest at base, blue-violet with greenish reflections, 1.8× as wide as long. Anterior margin arcuate, lobe not protruding, densely pubescent. Disc sparsely macropunctate with micropunctures between macropunctures, with sparse short pubescence. Medial line only slightly developed but visible, flat, almost unelevated, moderately impressed at base, only bordered by macropunctures, without medial impressions. Principal impressions elongate, deep. Lateral sides from principal impressions and lateral impressions coarsely macropunctate. Lateral margin green-violet. Basal margin bisinuate. Scutellum small, trapezoidal, shiny, violet with greenish reflections. Elytra slightly wider than pronotum at base; parallel at basal half, narrowing from beyond its mid-length to apex; convex in lateral view. Lateral margins arcuate below humeral calli, moderately serrate at apical third. Serrations and apex of elytra violet. Four distinct main costae along suture on each elytron. 1 st costa from base to apex, 2 nd from basal fifth and joined to 1 st at apical fifth. 3 rd costa separated, shortened, from basal fifth to 3/4 length of elytra. 4 th costa also separated, from basal fifth but reaching to apex where it is almost joined to 1 st and suture. Basal fifth in front of costae 2–4 irregularly furrowed, irregularly macropunctate. Humeral calli not furrowed, irregularly macropunctate. Intercostal intervals divided by less distinct but well visible intercostae. Main costae, intercostae and top of furrows blue-violet with a few macropunctures; lower areas of furrows and intercostae intervals green with golden reflections irregularly macropunctate and with very short pubescence. Epipleura horizontal, broad at basal quarter, green-brown at basal half, violet at apical half, sparsely macropunctate and pubescent. Legs brown-red with greenish reflections; femora less, tibiae more macropunctate and pubescent. Tibiae with two apical ventral spurs. Tarsi metallic red-brown, tarsal claws divergent and simple. Ventral side brown-red with greenish reflections. Hypomeron irregularly macropunctate more coarsely than prosternum. Anterior margin of prosternum densely pubescent. Prosternal process subparallel-sided, narrowed at apex, approximately 2.0× as long as wide, moderately densely and coarsely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, apex and sides smooth, polished and nearly impunctate. Metasternum sparsely macropunctate at central part; densely and coarsely macropunctate on sides; with short pubescence. Abdominal ventrites I–IV centrally sparsely macropunctate, laterally densely macropunctate in entire width, with short pubescence. Abdominal ventrites V densely macropunctate, with short pubescence. Variation. Body &male;&male; (n = 4) length: 25.25–25.75(27.50) (average 26.00) mm, width: 9.25–9.75 (average 9.44) mm, length/width ratio: (2.66)2.76–2.82 (average 2.76); &female;&female; (n = 27): length: 26.50–34.00 (average 29.89) mm, width: 9.50–12.00 (average 10.85) mm, length/width ratio: 2.61–2.90 (average 2.76). Colour of dorsal part from violet to blue-black, rarely bronze-brown or green. Intercostae can be thin or only slightly indicated but always visible. Aedeagus (n = 4) length: 6.91–7.32 mm, width: 1.64 mm, length/width ratio: 4.21–4.46. Parameres very broadly regularly navicular widest at mid-length, narrowly open at apical fifth, apice of parameres rounded. Penis moderately impressed with densely arranged striate laminae at center, apex of penis broadly triangular (Figs 65, 66). Differential diagnosis. Chrysodema (C.) eximia berliozi can be distinguished from the other species of this group by bicolourous head. Ventral side of head has brownish colour as well as entire ventral side but dorsal side has colour similar (or the same) to pronotum. Intercostae are always visible and most frequent colours of dorsal part are violet to blue-black. For additional characters see Key to species. Biology. Adults feed on leaves of Terminalia catappa, sometimes found on the leaves of Mallotus japonicus (Ong & Hattori 2019). Distribution. Taiwan: main, Orchid [Lanyu], and Green [Ludao] (Tamadera & Yoshitake 2017) islands (Fig. 12). Remarks. Descarpentries (1948) mentioned in his description that only two females (holotype deposited in Descarpentries’ collection and paratype from ‘même localité’ deposited in ‘Muséum de Paris’) belong to type series. The holotype is deposited in MNHN now but the paratype with the same locality label is deposited in NMPC. There is also the second female in MNHN from Botel-Tobago Is. with original Descarpentries’ type labels but with different dates of collecting. Also the Descarpentries’ (paratype) label is a little different. Therefore this specimen is not included in the type series and was provided with additional red printed labels ‘NO PARATYPE | This specimen cannot be paratype. | Descarpentries (1948: 64) mention- | ned only one paratype with the | same collecting data as holotype. | David Frank IV. 2021 [date handwritten] Lander (2003) downgraded Chrysodema (C.) berliozi to subspecies of C. (C.) eximia. However, majority of the specimens are very different from C. (C.) eximia but one of studied specimens was metallic green and it is similar to C. (C.) intercostata (which is in synonymy of C. (C.) eximia). Therefore it is kept as subspecies of C. (C.) eximia for the time being. It would be desirable to use molecular methods within C. (C.) eximia complex to definitively evaluate the status of this taxon (for more information see ‘Remarks’ under C. (C.) eximia)., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 518-520, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Descarpentries, A. (1948) Note sur deux Buprestides asiatiques. Bulletin de la Societe entomologique de France, 53 (4), 62 - 64. https: // doi. org / 10.3406 / bsef. 1948.16025","Akiyama, K. & Ohmomo, S. (2000) The buprestid beetles of the world. In: Fujita, H. (Ed.), Mushi-Shaʼs Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, pp. 1 - 341., 30 pls.","Kurosawa, Y. (1954) Buprestid-fauna of Eastern Asia, (Coleoptera) (I). Bulletin of the National Science Museum, Tokyo, 1 (1), 29 - 39.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Muhle, H. (2003) Taiwanese buprestids (Coleoptera, Buprestidae). Journal of the Zoological Society Wallacea, 1, 43 - 48.","Kuban, V. (2006) Buprestidae: Chrysochroinae: Chrysochroini. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 40 - 52 + 342 - 345.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp.","Qi, Z. - H., Ai, H. - M. & Song, H. - T. (2022) Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record. Zootaxa, 5099 (3), 335 - 368. https: // doi. org / 10.11646 / zootaxa. 5099.3.4","Kuban, V. (2016) Buprestidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Revised and updated edition. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden-Boston, pp. 19 - 27 + 456 - 461. https: // doi. org / 10.1163 / 9789004309142","Tamadera, Y. & Yoshitake, H. (2017) A New Record of Chrysodema (Cyalithoides) eximia berliozi (Coleoptera, Buprestidae) from Ludao Is., Taiwan. Elytra, Tokyo, New Series, 7 (1), 127 - 128.","Ong, U. & Hattori, T. (2019) Jewel beetles of Taiwan. Vol. 1. Kua Series, Donothing Image, Tainan, xv + 235 pp."]}
Published: 2022
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5. Chrysodema (Chrysodema) antennata Saunders 1874

Author: Frank, David
Subjects: Coleoptera, Insecta, Chrysodema antennata, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Key to the Chrysodema (Chrysodema) antennata species-group 1 Inner side of mesotibia without ridge...................................................................... 2 – Inner side of mesotibia with very distinct ridge (Figs 76, 77). Philippines: Mindanao, Dinagat and Samar islands. Figs 55–57, 59–60, 69, 76, 77.......................................................... C. (C.) variipennis Saunders, 1874 2 Lateral sides of abdominal ventrites II–V coarsely but sparsely macropunctate with smooth and shiny areas; inner side of male mesotibia of C. (C.) sekerkai without long setae, male of C. (C.) holynskii unknown................................. 3 – Lateral sides of abdominal ventrites II–V finely but densely macropunctate without smooth and shiny areas; inner side of male mesotibia densely pubescent with long setae................................................................ 4 3 Basic colour of elytra black; principal impressions of pronotum clearly visible; female mesotibia bent at apical third (Fig. 74). Generally more broad, length/width ratio: &female;&female; (n = 129) 2.51–2.74 (average 2.60). Indonesia: Banggai, Peleng and Sulawesi islands. Figs 38–40, 68, 74........................................................... C. (C.) sekerkai sp. nov . – Basic colour of elytra green-bronze; principal impressions inconspicuous; female mesotibia not bent at apical third (Fig. 75). Generally more slender, length/width ratio: &female;&female; (n = 37) 2.68–2.89 (average 2.80). Philippines: Masbate Is. Figs 48–50, 53, 75............................................................................. C. (C.) holynskii sp. nov . 4 Male antenna shorter than pronotum...................................................................... 5 – Male antenna longer than pronotum, metallic-coloured apical part of male sternite IX V-shape bordered at base (Fig. 78), penis narrowed at apical half (Figs 70, 71). Philippines: Mindanao Is., Camiguin (Mindanao) Is., Dinagat Is., Leyte Is., Samar Is. Figs 29–34, 70, 71, 78.................................................. C. (C.) antennata antennata Saunders, 1874 5 Metallic-coloured apical part of male sternite IX V-shape bordered at base, parameres shortly narrowly open at apex, (Fig. 67). Generally smaller: &female;&female; (n = 7): length: 22.00–27.25 (average 25.19) mm, width: 8.00–10.25 (average 9.25). Indonesia: Sulawesi Is. Figs 41–43, 46, 67, 73................................... C. (C.) antennata obsoleta J. Thomson, 1878 – Metallic-coloured apical part of male sternite IX arcuately bordered at base (Fig. 79), parameres open from mid-length, penis subparallel-sided (Fig. 72). Generally bigger: &female;&female; (n = 35): length: 24.25–30.00(31.50) (average 27.09) mm, width: 9.00–12.00 (average 10.12) mm. Philippines: Palawan Is. Figs 35–37, 72, 79................. C. (C.) antennata barriesi subsp. nov ., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on page 520, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Saunders, E. (1874) Notes on the Buprestidae collected by Professor Semper in the Philippines Islands; with descriptions of the new species. Transactions of the Royal Entomological Society of London, 1874 (2), 303 - 328.","Thomson, J. (1878) Typi Buprestidarum Musaei Thomsoniani. E. Deyrolle, Paris, 103 pp."]}
Published: 2022
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6. Chrysodema (Chrysodema) antennata subsp. antennata Saunders 1874

Author: Frank, David
Subjects: Coleoptera, Insecta, Chrysodema antennata, Arthropoda, Chrysodema, Animalia, Biodiversity, Chrysodema (chrysodema) antennata antennata saunders, 1874, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) antennata antennata Saunders, 1874 (Figs 26–34, 45, 47, 62, 70, 71, 78) Chrysodema antennatus Saunders (1874): 313 (original description). Chalcophora antennata: Kerremans (1885): 125 (catalogue). Chrysodema antennata: Kerremans (1892a): 37 (catalogue); Kerremans (1892b): 23 (comparison with C. hebes); Kerremans (1903): 74 (catalogue); Théry (1933): 3 (noted); Akiyama & Ohmomo (2000): Pl. 43, Figs 460-1, 460-2 (iconography). Chrysodema (Chrysodema) antennata: Kerremans (1909): 517 (key), 560 (redescription); Obenberger (1926): 130 (catalogue); Lander (2003): 14 (key), 48 (redescription), 80 (colour Fig. 94); Bellamy (2008): 527 (catalogue). Chrysodema hebes Kerremans (1892b): 23 (original description); Kerremans (1903): 74 (catalogue); Théry (1933): 4 (noted). Chrysodema (Chrysodema) hebes: Kerremans (1909): 518 (key), 562 (redescription); Obenberger (1926): 131 (catalogue); Lander (2003): 48 (as synonym of C. (C.) antennata), 80 (colour Fig. 95); Bellamy (2008): 527 (catalogue, in synonymy of C. (C.) antennata), 531 (listed as synonym of C. (C.) antennata). Chrysodema (Chrysodema) coriacea Kerremans (1919): 46 (original description); Obenberger (1926): 131 (catalogue); Lander (2003): 48 (as synonym of C. (C.) antennata), 80 (colour Fig. 96); Bellamy (2008): 527 (catalogue, in synonymy of C. (C.) antennata), 531 (listed as synonym of C. (C.) antennata). Chrysodema coriacea: Théry (1933): 3 (noted). Chrysodema samarensis Fisher (1926): 23 (original description). Chrysodema (Chrysodema) samarensis: Lander (2003): 48 (as synonym of C. (C.) antennata); Bellamy (2008): 527 (catalogue, in synonymy of C. (C.) antennata), 529 (listed as synonym of C. (C.) antennata). Type localities. Chrysodema antennata: ‘Mindanao and Camiguin de Mindanao’ [Philippines: Mindanao Island and Camiguin Island]; C. coriacea: ‘Ile Samar’ [Philippines: Samar Island]; C. hebes: ‘De Mindanao (Dr Staudinger, coll. G.-A. Baer)’ [Philippines: Mindanao Island]; C. samarensis: ‘Samar Island, Philippines’. Type material examined. Chrysodema antennata: LECTOTYPE (present designation): &male; (23.25 × 8.75 mm; Figs 27, 29–31, 70, 78), ‘ Mindanao | 143 [w(c), h; round] || Mindanao. [w(c), p] || Type [w, p; round, red border] || Saunders | 74-18. [w(c), p] || C. antennatus | Type &male; ES [w(c), h]’ (BMNH). PARALECTOTYPES (3 &female;&female;): &female; (27.50 × 10.50 mm), ‘ Camiguin | de | Mindanao | 143 [w(c), h; round] || Camiguin. | de Mindanao. [w(c), p] || Saunders | 74-18. [w(c), p]’ (BMNH); &female; (24.25 × 9.00 mm), ‘ Philippines | Semper | 143 [w(c), h; round] || Saunders | 74-18. [w(c), p]’ (BMNH); &female; (25.00 × 9.00 mm), ‘ Philippines | 143 [w(c), h; round] || Saunders | 74-18. [w(c), p] || antennata | K. [w(c), h]’ (BMNH). All specimens were provided with an additional red printed label: ‘ LECTOTYPE [or PARALECTOTYPE respectively] [sex] | Chrysodema | antennata | SAUNDERS, 1874 | David Frank des. IV. 2021 [date handwritten]’. Chrysodema hebes: LECTOTYPE (present designation): &female; (26.25 × 10.00 mm; Figs 28, 32), ‘Mindanao | Staudinger. [w(c), h (Kerremans' handwritting)] || Type [w, p; round; red border] || Kerremans. | 1903-59. [w(c), p] || hebes | Kerremans | Type. [w(c), h (Kerremans' handwritting)]’ (BMNH; figured in Lander (2003: Fig. 95) as holotype of C. antennata var. hebes). The specimen was provided with an additional red printed label: ‘ LECTOTYPE &female; | Chrysodema | hebes | KERREMANS, 1892 | David Frank des. IV. 2021 [date handwritten]’ and also white printed label ‘ Chrysodema (Chrysodema) | antennata antennata | SAUNDERS, 1874 | David Frank det. IV. 2021 [date handwritten]’. Chrysodema coriacea: SYNTYPES (3 &female;&female;): 2 &female;&female; (28.25 × 10.50; Figs 33, 47 and 25.50 × 9.50 mm), ‘ Samar | Meyer D. [w, h (Kerremans' handwritting)] || MUSEUM PARIS | COLL. CH. KERREMANS | 1923 [y, p] || TYPE [r, p] | coriacea | Kerr. Type [w, h (Kerremans' handwritting)]’ (MNHN); &female; (26.00 × 9.50 mm) ‘ Samar | Meyer D. [w, h (Kerremans' handwritting)] || &female; [w, h] || TYPE [r, p] | coriacea | Kerr. Type [w, h (Kerremans' handwritting)]’ (MNHN). All specimens were provided with an additional red printed label: ‘ SYNTYPE &female; | Chrysodema (Chrysodema) | coriacea | KERREMANS, 1919 | David Frank labelled IV. 2021 [date handwritten]’ and also white printed label ‘ Chrysodema (Chrysodema) | antennata antennata | SAUNDERS, 1874 | David Frank det. IV. 2021 [date handwritten]’. Chrysodema samarensis: HOLOTYPE: &female; (27.50 × 10.00 mm), ‘Samar | PI [w, p] || 2-vii-24. | coll. R.C.| McGregor [w, p] || Type No. | 28742 | U.S. N.M. [r, p/h] | HOLOTYPE [red letters] | CHRYSODEMA | samarensis | Fisher [w, h/p; red border]’ (USNM). ALLOTYPE: &male; (22.75 × 8.25 mm; Figs 34, 45, 71), ‘ Samar | PI [w, p] || 21-6-24. | coll. R.C. | McGregor [w, p] || AllotypeNo. | 28742 | U.S. N.M. [r, p/h]’ (USNM). AddiTiOnal maTerial examined (15 &male;&male;, 64 &female;&female;). PHILIPPINES: Philippines, 2 &female;&female; (MNHN); Philippines, 1 &female; (MNHN, ex coll. Janson); Philippin., v. Möllendorf, 1895 (1 &male;, 1 &female; IRSN; 1 &male;, 1 &female; SGBG; 1 &male; RHMP); Philippine Is., 1910–112, collected by R. McGregor, 2 &female;&female; (BMNH); Phil. Isla, 2 &female;&female; (BMNH); Philippines, 1 &female; (BMNH); Philippines, 1 &female; (NMPC, ex coll. Kubá&ncaron;); Philippines, 1 &female; (NMPC, ex coll. Lander). CAMIGUIN ISLAND: Phillippinen, Camiguin, 21.xii.1916, G. Boettcher, 1 &female; (MFNB, ex coll. Moser). DINAGAT ISLAND: Phillip. Islands, Dinagat, Semper, 1 &female; (BMNH, ex coll. Fry); Dinagat Is., Aug [ust]. 1991, 2 &male;&male; (NMPC, ex coll. Bílý); Dinagat Is., 22.11.1991, 1 &male;, 1 &female; (NMPC, ex coll. Bílý); Dinagat Is., 24.11.1991, 1 &male; (NMPC, ex coll. Bílý); Dinagat Is., xi.1991, 1 &female; (NMPC, ex coll. Bílý); Dinagat Is., 7.5.1995, 1 &male; (NMPC, ex coll. Bílý); Dinagat Is., 15.3.1992, 1 &male; (DFPC); Dinagat Isl., 15.iii.1992, 1 &female; (MHNG, ex coll. Lander); Dinagat Isl., 18.iii.1995, 1 &female; (MHNG, ex coll. Lander); Dinagat Isl., 1 &male;, 1 &female; (MHNG, ex coll. Lander); Dinagat Is., ix.1991, 1 &female; (RHMP). LEYTE ISLAND: Phillippinen, Leyte, St. Cruz, x.1915, G. Boettcher, 1 &female; (MFNB, ex coll. Moser). MINDANAO ISLAND: Mindanao, J. Waterstradt, 1903-1904, 2 &male;&male;, 14 &female;&female; (MNHN); Philippinés, Mindanao, 1 &female; (MNHN); Mindanao, 1 &female; (MNHN, ex coll. Oberthür); Mindanao, 1 &female; (NMPC); Mindanao, Zamboanga [without accurate specification], vi.[19]89, 2 &female;&female; (NMPC, ex coll. Bílý); Mindanao, R[Z?]amboanga [without accurate specification], vii.[19]89, 1 &female; (NMPC, ex coll. Bílý); Mindanao, 25.4.[19]95, Bourdant leg., 1 &female; (NMPC, ex coll. Bílý); Mindanao, 1 &male;, 1 &female; (RHMP). Lanao del Sur Prov.: Wao, November 2018, 1 &female; (DFPC). Misamis Oriental Prov.: P.I., Bugo, 1 &female; (NHMB, ex coll. Frey); Misamis, Balinagsag, Oct [ober]. 2014, 1 &female; (WBWA). South Cotabato Prov.: Koronadal, Aug. 30 1990, Gadayan, Rivera, Dajay, Frugalidad, 1 &female; (MHNG); Koronadal, Sept. 1991, Gadayan, Rivera, Dajay, Frugalidad, 1 &female; (MHNG). Zamboanga del Norte Prov.: Labuan, January 2018, &male; (DFPC). SAMAR ISLAND: Ile Samar, 3 &female;&female; (MNHN); Samar, vi.-vii.[18]96, J. Whithead, 2 &female;&female; (MFNB, ex coll. Moser, NHMB, ex coll. Frey); Samar, Meyer D., 2 &female;&female; (NMPC, ex coll. Kerremans); Samar, vi.1993, 1 &female; (NMPC, ex coll. Kubá&ncaron;); Samar, Eastern Visayan, July 1996, 1 &female; (NMPC, ex coll. Kubá&ncaron;); Eastern Visayae, Samar Isl. vi.1993, 1 &female; (MHNG, ex coll. Lander); Samar, vi.[19]93, 2 &female;&female; (NMPC, ex coll. Bílý); Island Samar, Baker, 1 &male;, 1 &female; (RHMP); Samar, Eastern Visayaz, July 1996, 1 &female; (SGBG); Samar, 5.6.1999, 1 &female; (DFPC). Northern Samar Prov.: Easter Visayas, Northern Samar, Lope de Vega, February 2019, 1 &female; (DFPC). Samar Prov.: Easter Visayas, Samar, Hinabangan, October 2017, 1 &female; (DFPC). Redescription of lectotype. Well preserved &male; specimen, only right antenna missing from antennomere III. Length 23.25 mm, width 8.75 mm, length/width ratio: 2.66. Body navicular, basic colour of dorsal side is black but it appears gold-green because of dense metallic green macropunctures; ventral side green with golden reflections, legs green. Head green with black frons.Eyes large, oval,area around eyes densely macropunctate with green macropunctures and densely pubescent. Frons black with green spot at center, sparsely macropunctate with green macro punctures, 1.6× as wide as diameter of eye, impressed with deep medial sulcus, with sparse short pubescence. Vertex green with golden reflections densely macropunctate. Labrum brown, pubescent. Antennae longer than pronotum, serrate from antennomere IV. Antennomeres I and II metallic green with coppery reflections, sparsely macropunctate and pubescent; radicula and antennomere from III brown. Antennomeres III–VI very long. Antennomere IV long triangular, V–X more trapezoidal than triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi brownblack, densely pubescent. Pronotum almost rectangular, black with irregular golden macropunctures or groups of macropunctures, moderately densely micropunctate, 1.5× as wide as long. Anterior margin arcuate, lobe not protruding, densely pubescent. Medial line black, micropunctate, without macropunctures. Medial impressions weak and shallow only indicated by denser macropunctation. Principal impressions oval, golden, densely macropunctate and with sparse short pubescence. Lateral impressions golden-green, moderately deep, macropunctate. Lateral margin green. Basal margin moderately bisinuate. Scutellum small, trapezoidal, green and shiny. Elytra wider than pronotum at base; widest at basal tenth, moderately narrowing to middle, strongly narrowing from beyond its mid-length to apex, moderately convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Groups of macropunctures of different sizes contain ca 5–35 macropunctures, arranged in almost regular rows. Black areas between rows of groups of macropunctures appear as weak costae which are interconnected at short intervals, sparsely macropunctate and micropunctate, along suture irregular row of individual macropunctures. The most lateral intervals coherently very densely macropunctate therefore almost entire area is golden-green. Humeral calli moderately furrowed. Epipleura horizontal, broad at basal quarter, green with golden reflections, moderately densely macropunctate and sparsely pubescent. Legs metallic green; femora and tibiae macropunctate and pubescent more densely on inner side. Inner side of mesotibiae from middle to tarsi very densely and longly pubescent. Tibiae with two apical ventral spurs. Tarsi metallic green, tarsal claws divergent and simple. Ventral side metallic green with golden reflections. Hypomeron irregularly punctate more coarsely than prosternum. Anterior margin of prosternum densely pubescent. Prosternal process subparallel-sided, narrowed at apex, approximately 2.0× as long as wide, densely and coarsely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, sides sparsely punctate and shiny. Metasternum sparsely punctate and shiny at central part; densely and finely punctate and pubescent on sides. Abdominal ventrites I–III moderately densely punctate at central part, laterally densely punctate and pubescent in entire width. Abdominal ventrites IV–V densely punctate and with short pubescence in entire area. Metallic green and pubescent apical part of sternite IX V-shape bordered at base (Fig. 78). Aedeagus. Narrowly regularly navicular, widest at mid-length, length: 6.18 mm, width: 1.00 mm, length/width ratio: 6.18. Parameres shortly narrowly open at apex, apices of parameres rounded. Penis distinctly narrowed from beyond its mid-length to apex, striate laminae only at the 2 nd quarter from apex. Apex of penis triangular (Figs 70). Variation (only specimens from selected localities, see ‘Remarks’). Body &male;&male; (n = 14) length: 20.50–23.75 mm, width: 7.50–8.75 mm, length/width ratio: 2.65–2.83; &female;&female; (n = 55): length: 22.75–29.25 (average 26.18) mm, width: 8.25–11.00 (average 9.66) mm, length/width ratio: 2.55–2.86 (average 2.71). Pronotum usually trapezoidal, rarely almost rectangular. Groups of macropuctures are variable in size as well as in count of macropunctures (2–>40). Colour of macropunctures variable from golden-green to dark brown. Aedeagus (n = 9) length: 5.55–6.18 mm, width: 0.91–1.18 mm, length/width ratio: 5.24–6.20. Differential diagnosis. Males of all subspecies of C. (C.) antennata have very dense and long pubescence on inner side of mesotibiae from middle to tarsi. Males of C. (C.) antennata antennata have antennae longer than pronotum. From C. (C.) antennata barriesi can be distinguished by shape of penis which is distinctly narrowed from beyond its middle to apex and by V-shaped border of metallic green apical part of male sternite IX (Fig. 78). From C. (C.) antennata obsoleta it can be distinguished by shape of aedeagus which is narrowly navicular. Groups of macropuctures are variable, the other subspecies have groups of macropunctures more or less uniform. For additional characters see Key to species. Distribution. Philippines: Mindanao, Camiguin (north of Mindanao), Dinagat, Leyte and Samar islands (Fig. 62). Specimens localised from Panay, Luzon and Mindoro islands are problematic (see ‘Remarks’). Remarks. Saunders (1874) did not specify explicitly how many specimens he had at his disposal when describing C. antennata nor mentioned any depository. Because he mentioned measurements ‘ &male;. Length 11 lines; breadth 4 lines. &female;. Length from 11 lines to 13 lines. . [&male; 23.32 × 8.48; &female;&female; 23.32–27.56 mm]’ and localities ‘ Hab.–Mindanao and Camiguin de Mindanao]’ it is clear that he had more specimens. I found one male specimen in BMNH (Figs 29–31) from Saunders’ collection, from ‘Mindanao’ which has almost identical length and width (23.25 × 8.75 mm) and type label ‘ C. antennatus | Type &male; ES [w(c), h]’ (Fig. 27). This specimen is designated as a lectotype in order to conserve the status of this taxon and avoid any further misinterpretations. There are also three females in Saunders’ collection labelled ‘ Camiguin de Mindanao’ (27.50 × 10.50 mm), ‘ Philippines, Semper’ (24.25 × 9.00 mm) and ‘Philippines’ (25.00 × 9.00 mm). According to measures in the description (23.32–27.56 mm) these female specimens are in this range therefore they are also included in the type series as paralecrotypes. Kerremans (1892b) did not specify explicitly how many specimens he had at his disposal when describing C. hebes. He mentioned only ‘De Mindanao (Dr Staudinger, coll. G.-A. Baer)’, a single length and width measurement ‘Long. 26; lat. 10 mill.’. I found one female specimen in BMNH (Fig. 32) with the label ‘Mindanao | Staudinger’ which has almost identical length and width (26.25 × 10.00 mm) and Kerremans’ handwriting type label ‘hebes | Kerremans | Type. [w, h]’ (Fig. 28). This specimen is designated as a lectotype in order to conserve the status of this taxon and avoid any further misinterpretations. Kerremans (1892b) described C. hebes based on the structure of elytra, matt appearance and non-metallic tarsi. Lander (2003) synonymized C. (C.) hebes with C. (C.) antennata antennata and this synonymy is accepted also by Bellamy (2008). I studied the type specimen of C. (C.) hebes and found all differences are within variability of C. (C.) antennata antennata. Therefore C. (C.) hebes is kept in synonymy of C. (C.) antennata antennata. Kerremans (1919) did not specify explicitly how many specimens he had at his disposal when describing C. coriacea from Samar Island but he had more specimens than one because he mentioned males as well as females and range of measures ‘Long. 25–28; larg. 8,5-10 10 mill.’. I found three females (25.50, 26.00 and 28.25 mm (Fig. 33)) in MNHN from Samar Is. with Kerremans’ handwriting type labels ‘coriacea | Kerr. Type [w, h]’ (Fig. 47). Since all three specimens morphologically belong to a single species, they are from type locality (Samar Is.) and have Kerremans’ handwritten type labels, there is no need of lectotype designation and I treat them as syntypes. Kerremans (1919) described C. coriacea based on stockier and more convex body, with much larger elytral punctuation and thicker reliefs, the ribs well marked. Lander (2003) synonymised C. (C.) coriacea with C. (C.) antennata antennata and this synonymy is accepted also by Bellamy (2008). I studied three syntypes of C. (C.) coriacea and twenty more specimens from Samar Is. and found all differences are within variability of C. (C.) antennata antennata. Therefore C. (C.) coriacea is kept in synonymy of C. (C.) antennata antennata. Fisher (1926) described Chrysodema samarensis from twenty two specimens from Samar Island. He wrote ‘This species is closely allied to Chrysodema antennata...’ and described it based on ‘absence of coppery spots on the elytra, and the reliefs of a violaceous black color...’. Lander (2003) synonymised C. (C.) samarensis with C. (C.) antennata antennata and this synonymy is accepted also by Bellamy (2008). I studied holotype (&female;) and allotype (&male;; Figs 34, 45) from USNM and found all differences are within variability of C.<, Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 520-524, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Saunders, E. (1874) Notes on the Buprestidae collected by Professor Semper in the Philippines Islands; with descriptions of the new species. Transactions of the Royal Entomological Society of London, 1874 (2), 303 - 328.","Kerremans, C. (1885) Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique, 29, 119 - 157.","Kerremans, C. (1892 a) Catalogue synonymique des Buprestides decrits de 1758 a 1890. Memoires de la Societe entomologique de Belgique, 1, 1 - 304.","Kerremans, C. (1892 b) Voyage de M. E. Simon aux iles Philippines (Mars et Avril 1890). 3 e Memoire (1), Buprestidae. Annales de la Societe entomologique de France, 61, 21 - 26.","Kerremans, C. (1903) Coleoptera Serricornia. Fam. Buprestidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","Thery, A. (1933) Buprestides recoltes par son Altesse Royale le prince Leopold de Belgique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 9 (31), 1 - 4.","Akiyama, K. & Ohmomo, S. (2000) The buprestid beetles of the world. In: Fujita, H. (Ed.), Mushi-Shaʼs Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, pp. 1 - 341., 30 pls.","Kerremans, C. (1909) Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars). Livraisons 13 - 19. J. Janssens, Bruxelles, pp. 385 - 604., pls. 17 - 22. https: // doi. org / 10.5962 / bhl. title. 9700","Obenberger, J. (1926) Buprestidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, pp. 1 - 212.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp.","Kerremans, C. (1919) Descriptions de Buprestides nouveaux. Annales de la Societe entomologique de Belgique, 59 (3), 41 - 62.","Fisher, W. S. (1926) Fauna Samarensis: Coleoptera, Buprestidae. Philippine Journal of Science, 31 (2), 235 - 244."]}
Published: 2022
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7. Chrysodema (Chrysodema) variipennis Saunders 1874

Author: Frank, David
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema variipennis, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) variipennis Saunders, 1874 (Figs 54–60, 62, 69, 76, 77) Chrysodema variipennis Saunders (1874): 310 (original description); Kerremans (1888 a): lxv (catalogue [sic!, locality ‘Iles Nicobar’]); Kerremans (1892a): 40 (catalogue); Kerremans (1903): 74 (catalogue [locality ‘Manille’]); Akiyama & Ohmomo (2000): Pl. 45, Figs 479-1, 479-2 (iconography, misidentification). Chalcophora variipennis: Kerremans (1885): 128 (catalogue). Chrysodema (Chrysodema) variipennis: Kerremans (1909): 516 (key), 539 (redescription); Obenberger (1926): 135 (catalogue); Lander (2003): 14 (key), 50 (redescription), 81 (colour Fig. 97); Bellamy (2008): 542 (catalogue). Chrysodema leopoldiana Théry (1933): 2 (original description). Chrysodema (Chrysodema) leopoldiana: Lander (2003): 50 (as synonym of C. (C.) variipennis); Bellamy (2008): 533 (listed as synonym of C. (C.) variipennis), 542 (catalogue, in synonymy of C. (C.) variipennis). Type localities. Chrysodema variipennis: ‘ Type from Mindanao’ [Philippines, Mindanao Island]; C. leopoldiana: ‘ Philippines, Mauo Samar’ [Philippines, Samar Island, Mauo]. Type material examined. Chrysodema variipennis: HOLOTYPE (by monotypy): &female; (20.75 × 8.50 mm; Figs 55– 58, 76), ‘Minda | nao [w(c), h; round] || Type [w, p; round, red border] || Saunders | 74-18. [w(c), p] || C. variipennis | ES. Type [w(c), h]’ (BMNH). The specimen was provided with an additional red printed label: ‘ HOLOTYPE (by monotypy) | Chrysodema | variipennis | SAUNDERS, 1874 &female; | David Frank labelled IV. 2021 [date handwritten]’. Chrysodema leopoldiana: HOLOTYPE: &female; (23.75 × 9.00 mm; Figs 54, 59), ‘ &female;? [w, p/h] || [original label] Philippines | Mauo–Samar | 8-IV-1932 | Prince Léopold [w(c), p]; [newer label] Coll. R. I. Sc. N. B. [y, p] [original label is glued on newer label] || Rog. Mus. Hist. Nat. | Belg. I.G. 9796 [w(c), p] || [original label] Chrysodema | Leopoldiana | Thery | TYPE [w(c), h/p (red stamp)]; [original label] Holotype [r, p; black border]; [newer label] d.: Bull. Mus. R. Hist. | nat. Belgique 1933, 9, 31:2,3 [w, h] [original labels are glued on newer label]’ (IRSN). The specimen was provided with an additional white printed label ‘ Chrysodema (Chrysodema) | variipennis | SAUNDERS, 1874 | David Frank det. IV. 2022 [date handwritten]’ AddiTiOnal maTerial examined (1 &male;, 1 &female;). PHILIPPINES: DINAGAT ISLAND: Philippines, Dinagat Is., 23.11.1991, 1 &male; (NMPC, ex coll. Bílý; Figs 60, 69, 77); Philippines, Dinagat Is., 15.3.1995, 1 &female; (NMPC, ex coll. Bílý). Redescription of holotype. Well preserved &female; specimen, only left antenna missing from antennomere IV. Length 20.75 mm, width 8.25 mm, length/width ratio: 2.52. Body navicular, basic colour of dorsal side black with dense metallic golden-green groups of macropunctures; ventral side metallic golden-green, legs green. Head black with green area around eyes and punctures on dorsal side, green on ventral side. Eyes large, oval. Area around eyes green, finely macropunctate. Frons black, 2.2× as wide as diameter of eye, impressed with deep medial sulcus. Vertex macropunctate and with short pubescence. Labrum very small, brown, with short pubescence. Antennae serrate from antennomere IV. Antennomeres I and II golden-green, sparsely macropunctate; radicula and antennomeres from III dark brown with sparse short pubescence. Antennomeres I, III and IV almost the same length, ca. 3× longer than II. Antennomere IV long triangular, V–X triangular, XI trapezoidal. Maxillae, labium, maxillary palpi and labial palpi pale brown, densely pubescent. Pronotum trapezoidal, black, moderately densely irregularly golden-green macropunctate, micropunctate on black areas, 1.7× as wide as long, widest at base. Anterior margin arcuate, lobe not protruding, densely pubescent. Medial line black, bordered rows of macropunctures create shallow medial impressions. Principal impressions small and shallow, golden, densely macropunctate and with sparse short pubescence. Lateral impressions shallow, densely macropunctate. Lateral margin black. Basal margin very slightly bisinuate. Scutellum small, trapezoidal, dark-green and shiny. Elytra wider than pronotum at base; parallel to middle, narrowing from mid-length to apex, strongly convex in lateral view. Lateral margins arcuate below humeral calli, serrate from beyond its mid-length to apex. Irregular groups of macropunctures have different sizes, but three fairly large at basal third and three large at apical third are situated in transverse rows on each elytron. Black areas between groups of macropunctures create visible costae which are interconnected between themselves, sparsely micropunctate, the first costa is visible for whole length, other costae from mid-length of elytra. The last lateral intervals coherently very densely macropunctate therefore almost entire area is golden-green. Humeral calli macropunctate. Epipleura horizontal, broad at basal quarter, green with golden reflections, macropunctate and with short pubescence. Legs metallic green; femora and tibiae macropunctate and pubescent more densely on inner side. Inner side of mesotibia with very distinct serrate and pubescent ridge (Fig. 76). Tibiae with two apical ventral spurs. Tarsi metallic green, tarsal claws divergent and simple. Ventral side metallic golden-green, shiny. Hypomeron and prosternum irregularly punctate. Anterior margin of prosternum densely pubescent. Prosternal process subparallel-sided narrowed at apex, approximately 1.7× as long as wide, densely and coarsely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, sides very sparsely punctate and shiny. Metasternum sparsely punctate and shiny at central part; densely punctate and pubescent on sides. Abdominal ventrites golden-green and densely pubescent and macropunctate at basal part, black with sparse macro- and micropunctures at apical part. Variation. Body &male; (n = 1) length: 20.00 mm, width: 7.50 mm, length/width ratio: 2.67; &female;&female; (n = 3): length: 20.75–23.75 mm, width: 8.25–9.00 mm, length/width ratio: (2.52)2.61–2.64. Groups of macropuctures variable in size as well as in count of macropunctures but three large groups at apical third always present. Colour of macropunctures moderately variable from golden-green to dark green. Serrate ridge on mesotibia more distinct in males where it is very obvious. Aedeagus (n = 1) length: 4.41 mm, width: 1.01 mm, length/width ratio: 4.36. Aedeagus small in contrast to other taxa. Parameres oval widest at mid-length, narrowly open from mid-length, apice of parameres obliquely truncated. Penis moderately impressed with densely arranged striate laminae at entire width, apex of penis narrowly triangular (Fig. 69). Differential diagnosis. Chrysodema (C.) variipennis can be easily distinguished from other taxa by serrate ridge on mesotibia (more obvious at males than females) (Figs 76, 77), small aedeagus and three large groups of macropunctures occure transverse at basal part of elytron. Distribution. Philippines: Mindanao, Dinagat and Samar islands (Fig. 62). Remarks. Saunders (1874) described C. variipennis as a typical form and five varieties (Var. 1–5). The typical form is described from one specimen from ‘Mindanao’ which has measures ‘Length 9 ½ lines; breadth 4 lines. [20.14 × 8.48 mm]’. I found one female specimen in BMNH (Figs 55–57) from Saunders’ collection, from ‘Mindanao’ which has the type label ‘ C. Variipennis | ES. Type [w(c), h]’ (Fig. 58). Because this specimen agrees with the primary description and has the type label it can be considered as holotype. All other varieties from Mindanao, Bohol and Luzon islands probably belong to different species (Lander (2003) mentioned C. jucunda, C. smaragdula and C. ruficornis) but I did not find (and study) these specimens. Théry (1933) described C. leopoldiana from one specimen from Mauo, Samar Is. Théry compared this species only with C. coriacea (= C. (C.) antennata antennata). Lander (2003) synonymized this species only based on the description without studying the type specimen. I studied the type and it certainly belongs to C. (C.) variipennis therefore it is kept in synonymy. Lander (2003) mentioned distribution for C. (C.) variipennis also from Luzon, Palawan and Borneo islands but I have never studied specimens from these islands. Occurrence in Borneo Is. is unlikely., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 537-541, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Saunders, E. (1874) Notes on the Buprestidae collected by Professor Semper in the Philippines Islands; with descriptions of the new species. Transactions of the Royal Entomological Society of London, 1874 (2), 303 - 328.","Kerremans, C. (1888) Buprestides du Musee de Calcutta. Bulletin ou Comptes-Rendus de la Seances de la Societe entomologique de Belgique, Serie III, 103, lxiii - lxix.","Kerremans, C. (1892 a) Catalogue synonymique des Buprestides decrits de 1758 a 1890. Memoires de la Societe entomologique de Belgique, 1, 1 - 304.","Kerremans, C. (1903) Coleoptera Serricornia. Fam. Buprestidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","Akiyama, K. & Ohmomo, S. (2000) The buprestid beetles of the world. In: Fujita, H. (Ed.), Mushi-Shaʼs Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, pp. 1 - 341., 30 pls.","Kerremans, C. (1885) Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique, 29, 119 - 157.","Kerremans, C. (1909) Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars). Livraisons 13 - 19. J. Janssens, Bruxelles, pp. 385 - 604., pls. 17 - 22. https: // doi. org / 10.5962 / bhl. title. 9700","Obenberger, J. (1926) Buprestidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, pp. 1 - 212.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp.","Thery, A. (1933) Buprestides recoltes par son Altesse Royale le prince Leopold de Belgique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 9 (31), 1 - 4.","Thomson, J. (1878) Typi Buprestidarum Musaei Thomsoniani. E. Deyrolle, Paris, 103 pp.","Laporte Comte de Castelnau, F. L. N. & Gory, H. L. (1835) Chrysodema. In: Dumenil, P. (Ed.), Histoire naturelle de iconographie des insectes Coleopteres. Monographie des buprestides. Tome 1. J. B. Bailliere, Paris, pp. 1 - 27. https: // doi. org / 10.5962 / bhl. title. 110141","Descarpentries, A. (1948) Note sur deux Buprestides asiatiques. Bulletin de la Societe entomologique de France, 53 (4), 62 - 64. https: // doi. org / 10.3406 / bsef. 1948.16025","Fisher, W. S. (1926) Fauna Samarensis: Coleoptera, Buprestidae. Philippine Journal of Science, 31 (2), 235 - 244."]}
Published: 2022
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8. Chrysodema (Chrysodema) holynskii Frank 2022, sp. nov

Author: Frank, David
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema holynskii, Taxonomy
Abstract: Chrysodema (Chrysodema) holynskii sp. nov. (Figs 48–53, 62, 75) urn:lsid:zoobank.org:act: 53F0BEEE-BCC0-4BB8-A632-422358F9DF61 Type locality. Philippines: Masbate Province, Masbate Island, Aroroy. Type material examined. HOLOTYPE: &female; (29.50 × 10.25 mm; Figs 48–51), ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || antennata E.S | Det. Dr. Obenberger [w, h/p] || Chrysodema | antennata Saund. | sensu Th. Lander 2000 [w, p]’ (NMPC). PARATYPES (44 &female;&female;): PHILIPPINES: MASBATE ISLAND: Masbate Prov.: ‘ Philippinen | Masbate |Aroroy [w(c), p/h; black border] || Chrysodema | antennata | Saund., 1874 | Dét. T. LANDER 192001 [w, h/p]’ 2 &female;&female; (NMPC); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || Chrysodema | antennata Saund. | sensu Th. Lander 2000 [w, p]’ 2 &female;&female; (NMPC); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || Slg.H.Pochon | Bern | 1968 [w, p] || 29. [w, h; blue letters] || Chrysodema | antennata Saund | Det. H. Pochon [w, h/p]’ 1 &female; (NMBE, ex coll. Pochon); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || Slg.H.Pochon | Bern | 1968 [w, p]’ 1 &female; (NMBE, ex coll. Pochon); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [underside]: G Boettcher | VIII.1917 [w(c), h] || Coll. | Jul.Moser [w, p] || antennata Snd | Det. Hoscheck. 1926 [w(c), h/p; black border]’ 1 &female; (MFNB, ex coll. Moser); ‘[upper side]: Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [underside]: G Boettcher | VIII.1917 [w(c), h] || Coll. | Jul.Moser [w, p]’ (5 &female;&female; MFNB, ex coll. Moser, 1 &female; DFPC); ‘[upper side]: Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [underside]: G Boettcher | VIII.1917 [w(c), h] || Coll. | Jul.Moser [w, p] || Chrysodema | antennata | Thoms., 1874 | Dét. T. LANDER 192001 [w, h/p]’ (13 &female;&female; MFNB, ex coll. Moser, 2 &female;&female; DFPC); ‘ Philippinen | Masbate |Aroroy [w(c), p/h; black border] || Collectio | H. MARCHAND | 1956 Basel [w, p] || Coll.NMB [r, p] || Chrysodema | antennata | E. Sd.| Det. Dr. Obenberger [w, h/p]’ 1 &female; (NHMB, coll. general, ex coll. Marchand); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || Coll.G. Frey | NMB [blue, p] || Chrysodema | antennata | E. Sd.| Det. Dr. Obenberger [w, h/p]’ 1 &female; (NHMB, ex coll. Frey); ‘Aroroy | Masbate [w, p] || Coll.G. Frey | NMB [blue, p] || Chrysod. | antennata. Snd. | Det. Hoscheck 19 26. [w(c), h/p; black border]’ 1 &female; (NHMB, ex coll. Frey); ‘Aroroy | Masbate [w, p] || Coll.G. Frey | NMB [blue, p]’ 1 &female; (NHMB, ex coll. Frey); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border]’ 1 &female; (DFPC); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || 915. [w, p] || Coll. S. Gottwald (Berlin) | ex. Coll. | A. DAHLHEUSER | (Marburg) [w, p] || Chrysodema | antennata [w, h]’ 1 &female; (SGBG); ‘1929 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘1931 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘1932 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘1933 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll. Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘1934 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘4267 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label]’ 1 &female; (IRSN); ‘4270 [w(c), h] || [original label] Philippinen | Masbate | Aroroy [w(c), p/h; black border]; [newer label] Koll.Dr.A.Frh.v.Hoschek [w(c), p; black border] [original label is glued on newer label] || antennata Sd. | Det. Hoschek. [w(c), p; black border]’ 1 &female; (IRSN); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || coll. R B HO&Lstrok;Y&Nacute;SKI | BPexd [w, h] || 1993 | Chrysodema | coolsi HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h] || Chrysodema | coolsi | HO&Lstrok;Y&Nacute;SKI | HOLOTYPE [r, h]’ 1 &female; (RHMP; Figs 52, 53); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || coll. R B HO&Lstrok;Y&Nacute;SKI | BPexe [w, h] || 1993 | Chrysodema | coolsi HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h] || Chrysodema | coolsi | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’ 1 &female; (RHMP); ‘ Philippinen | Masbate | Aroroy [w(c), p/h; black border] || coll. R B HO&Lstrok;Y&Nacute;SKI | BPdi- [w, h] || 1993 | Chrysodema | coolsi HO&Lstrok;. | det. R. Ho&lstrok;y&nacute;ski [w, h] || Chrysodema | coolsi | HO&Lstrok;Y&Nacute;SKI | PARATYPE [r, h]’ 1 &female; (DFPC). The last three specimens are labelled as holotype and paratypes of C. coolsi, however erroneously, for details see ‘Remarks’. All specimens were provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] &female; | Chrysodema (Chrysodema) | holynskii sp. nov. | David Frank det. I. 2022 [date handwritten]’. Description of holotype. Well preserved &female; specimen with all appendages intact. Length 22.75 mm, width 8.25 mm, length/width ratio: 2.76. Body navicular, head and elytra green-bronze, pronotum bronze; ventral side golden-green, legs green. Head bronze-violet with green areas around eyes and at central part on dorsal side, green on ventral side. Eyes large, oval, area around eyes densely finely macropunctate. Frons bronze-violet with green central part, sparsely macropunctate and with short pubescence, 2× as wide as diameter of eye, impressed with deep medial sulcus. Vertex more densely macropunctate than frons. Micropunctures sparse on frons. Labrum brown, sparsely pubescent. Antennae serrate from antennomere IV. Antennomeres I and II bronze-violet with green reflections, sparsely macropunctate and pubescent; radicula and antennomeres from III dark brown with sparse long pubescence. Antennomere II ca. 3× shorter than I and III. Antennomeres IV–V long triangular, VI–X more trapezoidal than triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi dark brown, densely pubescent. Pronotum trapezoidal, narrowing anteriad with rounded sides, black, moderately densely irregularly goldengreen macropunctate, sparsely micropunctate between macropunctures, 1.4× as wide as long, widest at half. Anterior margin arcuate, lobe not protruding, densely pubescent. Medial line without macropunctures, densely micropunctate. Medial and principal impressions only slightly indicated. Lateral impressions only at basal half, shallow, with several macropunctures. Lateral margin golden shiny. Basal margin moderately bisinuate. Scutellum small, trapezoidal, bronze with green margins. Elytra moderately wider than pronotum at base; parallel at basal half, narrowing from mid-length to apex, strongly convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Groups of macropunctures of almost the same size contain ca 5–10 macropunctures, arranged in almost regular rows. Areas between rows of groups of macropunctures sparsely micropunctate, at apical half appear as weak costae which are interconnected at short intervals, along suture a row of individual macropunctures. Most lateral interval coherently very densely macropunctate therefore almost entire area is golden-green and very shiny. Humeral calli moderately furrowed. Epipleura horizontal, broad at basal quarter, green with golden reflections, macropunctate and with short pubescence. Legs metallic green with golden reflections; femora and tibiae macropunctate and pubescent more densely on inner side. Mesotibiae moderately curved at basal third and almost straight at two apical thirds (Fig. 75). Tibiae with two apical ventral spurs. Tarsi metallic green, tarsal claws divergent and simple. Ventral side metallic green with golden reflections. Hypomeron irregularly macropunctate more densely than prosternum. Anterior margin of prosternum with very dense short pubescence. Prosternal process subparallel-sided, narrowed at apex, approximately 1.8× as long as wide, moderately densely and coarsely macropunctate at central part, apex and sides smooth,polished,nearly impunctate and sparsely pubescent.Metasternum macropunctate and pubescent. Abdominal ventrites densely macropunctate and pubescent at basal part,shiny with sparse macropunctures at apical part. Variation. Body &female;&female; (n = 38): length: 21.50–29.50 (average 26.37) mm, width: 7.50–10.50 (average 9.42) mm, length/width ratio: 2.68–2.89 (average 2.80); male unknown. Colour of dorsal part from green-bronze to bronze, rarely green. Groups of macropuctures can be moderately variable but never large and always arranged in almost regular rows. Differential diagnosis. Male of Chrysodema (C.) holynskii is unknown. Females of this species are the slenderest from whole group. Average length/width ratio is 2.80 in contrast to 2.60–2.72 of other species. Principal impressions of pronotum are only slightly indicated. Basic colour of elytra is green-bronze and not black. From subspecies of C. (C.) antennata can be distinguished by shiny and sparsely macropunctate apical part of abdominal ventrites, and from C. (C.) sekerkai by mesotibiae which are moderately curved at basal third and straight at two apical thirds. For additional characters see Key to species. Etymology. This species is dedicated to my friend Roman B. Ho&lstrok;y&nacute;ski (Milanówek, Poland), a specialist in Buprestidae, mainly in subfamily Chrysochroinae. Distribution. Philippines: Masbate Island (Fig. 62). Remarks. I studied three female specimens of C. (C.) holynskii labelled as types of Chrysodema coolsi. According to the description of Chrysodema (Thymedes) coolsi Ho&lstrok;y&nacute;ski, 1994 the type series includes one male (HT) and four female specimens. Two paratypes are deposited in IRSN, the holotype and two paratypes were in RHMP with Ho&lstrok;y&nacute;ski’s collection labels ‘BPeqp’ (HT), ‘BPeqo’ and ‘BPfke’. The paratype ‘BPfke’ has been sent to K. Akiyama (R. Ho&lstrok;y&nacute;ski pers. comm. September 2021). I studied two specimens from RHMP (BPeqp, BPeqo) and they certainly belong to the subgenus Thymedes. However, there are also three other female specimens in RHMP labelled as a holotype and paratypes of Chrysodema coolsi, det. R. Ho&lstrok;y&nacute;ski 1993 (Fig. 53), with Ho&lstrok;y&nacute;ski’s collection labels ‘BPexd’ (HT) (Fig. 52), ‘BPexe’ and ‘BPdi-’. These specimens are labelled erroneously and do not belong to the type series of Chrysodema (Thymedes) coolsi Ho&lstrok;y&nacute;ski, 1994 (R. Ho&lstrok;y&nacute;ski pers. comm. September 2021) or even to the subgenus Thymedes at all, evidently representing the new species of Chrysodema s.str. described hereby as Chrysodema (Chrysodema) holynskii sp. nov., Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 531-534, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816, {"references":["Thery, A. (1933) Buprestides recoltes par son Altesse Royale le prince Leopold de Belgique. Bulletin du Musee royal d'Histoire naturelle de Belgique, 9 (31), 1 - 4.","Saunders, E. (1874) Notes on the Buprestidae collected by Professor Semper in the Philippines Islands; with descriptions of the new species. Transactions of the Royal Entomological Society of London, 1874 (2), 303 - 328.","Holynski, R. B. (1994) A review of Chrysodema C. G. (Coleoptera: Buprestidae) I. The subgenera Tamamushia M. C. and Thymedes Wath. Annals of the Upper Silesian Museum, Entomology, 5, 69 - 96."]}
Published: 2022
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9. Chrysodema (Chrysodema) lewisii subsp. nakatai Frank & Sekerka 2020

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema (chrysodema) lewisii nakatai frank & sekerka, 2020, Chrysodema, Animalia, Chrysodema lewisii, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) lewisii nakatai Frank & Sekerka, 2020 (Figs 1D��F, 5I��K, 8E) Chrysodema (Chrysodema) lewisii nakatai Frank & Sekerka, 2020: 16. Material examined. 1 &female; (Figs 1D��F, 5I��K) (CHTS): CHINA, Wucheng Mountain, Tucheng District, New Taipei City, Taiwan; alt. 300 m; 6.XI.2004; Chi-Liang Cheng leg.; length 2.51 cm, width 0.88 cm, length/width ratio: 2.85. �� 1 &female; (CWIC): Wushibi, Yilan County, Taiwan; alt. 450 m; 18.VII.2004; Wen-I Chou leg. �� 1 &female; (CWIC); same collection data as for the preceding; alt. 300 m; 14.IX.2001. Distribution. China (north Taiwan) and Japan (Yaeyama Islands). Adult host plant. Machilus thunbergii and Terminalia catappa (need verification, see remarks) (Ong & Hattori, 2019); sometimes found on Castanopsis carlesii and Alnus formosana (Wen-I Chou, pers. comm. 2021). Remarks. Frank & Sekerka (2020) described this subspecies as elytra semi-matt and about half of specimens with impression at basal third of each elytron. However, all the three specimens we examined were without the impression mentioned above. It is the same in habitus of this species in Ong & Hattori (2019) and by personal communication with Wen-I Chou. Whether all specimens from Taiwan are without impression at basal third of each elytron needs to be verified. Ong & Hattori (2019) cited the former two host plants from Ohmomo & Fukutomi (2013) but we found it was misquoted. Four host plants listed by Ohmomo & Fukutomi (2013) are cited as the host plants of subspecies C. (C.) lewisii lewisii. It seems Terminalia catappa is not the real host and sometimes the species can be found on Castanopsis carlesii and Alnus formosana (Wen-I Chou, pers. comm. 2021)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 360, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1","Ong, U. & Hattori, T. (2019) Jewel beetles of Taiwan. Vol. 1. Kua Series, Donothing Image, Tainan, xv + 235 pp.","Ohmomo, S. & Fukutomi H. (2013) The buprestid beetles of Japan. Iconographic Series of Insect 7. Mushi-sha, Tokyo, 208 pp."]}
Published: 2022
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10. Chrysodema (Chrysodema) aurostriata Saunders 1866

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema aurostriata, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) aurostriata Saunders, 1866 Chrysodema aurostriata Saunders, 1866: 302. Distribution. China (Yunnan, need verification) and Laos. Host plant. Unknown. Remarks. Frank & Sekerka (2020) examined many specimens and noted this species was known only from the female holotype labelled just as Laos. However, they did not mention the specimen from Yunnan. Kub��&ncaron; (2006, 2016) mentioned occurrence of C. (C.) aurostriata in Yunnan and the record was followed also by Bellamy (2008). Unfortunately, no formal faunistic record was published. According to V. Kub��&ncaron; (pers. comm. 2021), it was based on specimens collected by Jing-Ke Li (often as Li Jingke, but Li is the family name), which he had seen personally during his expedition to Laos, where he met Jing-Ke Li. The material from Yunnan looked trustworthy to him as it contained morphotypes of common species typical for that region and hence he included it in the Palaearctic Catalogue. According to him the specimen(s) were certainly smooth and golden-green and thus probably belonged to C. (C.) vrabeci. He did not purchase any voucher specimen from Jing-Ke Li and thus the depository of the material is unknown., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 361, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1","Kuban, V. (2006) Buprestidae: Chrysochroinae: Chrysochroini. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 40 - 52 + 342 - 345.","Kuban, V. (2016) Buprestidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Revised and Updated Edition. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden-Boston, pp. 19 - 27 + 456 - 461.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76. Pensoft, Sofia-Moscow, 625 pp."]}
Published: 2022
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11. Chrysodema (Pseudochrysodema) dalmanni subsp. dalmanni dalmanni (Mannerheim 1837

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema (pseudochrysodema) dalmanni dalmanni (mannerheim, 1837), Chrysodema dalmanni, Taxonomy
Abstract: Chrysodema (Pseudochrysodema) dalmanni dalmanni (Mannerheim, 1837) (Figs 4A��F, 5D, 7C, 7D) Evides dalmanni Mannerheim, 1837: 40. = Chrysodema manillarum Thomson, 1879: 9. = Chrysodema granulosa Kerremans, 1895: 197. = Chrysodema mira Obenberger, 1928: 147. Material examined. 11 &male;&male; (Fig. 4A��C, 5D), 12 &female;&female; (Fig. 4D��F) (CHTS): CHINA, Ludao Island, Taitung City, Taiwan; 22��67��N, 121��49��E, alt. 49 m; Hai-Tian Song & Wen-I Chou leg.; body &male;&male; length: 2.35��2.81 cm, width: 0.80��0.98 cm, length/width ratio: 2.87��2.94; body &female;&female; length: 2.54��3.18 cm, width: 0.88��1.14 cm, length/width ratio: 2.79��2.89; aedeagus (n=1) length: 8.30 mm, width: 1.60 mm, length/width ratio: 5.19. Distribution. China (south to southeast of the main island and Ludao Island, Taiwan); Japan (Ryukyu Islands); Philippines and Indonesia. Adult host plant. Terminalia catappa (Ong & Hattori, 2019). Larva host plant. Casuarina equisetifolia (Ong & Hattori, 2019)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 365, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Ong, U. & Hattori, T. (2019) Jewel beetles of Taiwan. Vol. 1. Kua Series, Donothing Image, Tainan, xv + 235 pp."]}
Published: 2022
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12. Chrysodema (Chrysodema) lewisii subsp. nakatai Frank & Sekerka 2020

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema (chrysodema) lewisii nakatai frank & sekerka, 2020, Chrysodema, Animalia, Chrysodema lewisii, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) lewisii nakatai Frank & Sekerka, 2020 (Figs 1D–F, 5I–K, 8E) Chrysodema (Chrysodema) lewisii nakatai Frank & Sekerka, 2020: 16. Material examined. 1 &female; (Figs 1D–F, 5I–K) (CHTS): CHINA, Wucheng Mountain, Tucheng District, New Taipei City, Taiwan; alt. 300 m; 6.XI.2004; Chi-Liang Cheng leg.; length 2.51 cm, width 0.88 cm, length/width ratio: 2.85. • 1 &female; (CWIC): Wushibi, Yilan County, Taiwan; alt. 450 m; 18.VII.2004; Wen-I Chou leg. • 1 &female; (CWIC); same collection data as for the preceding; alt. 300 m; 14.IX.2001. Distribution. China (north Taiwan) and Japan (Yaeyama Islands). Adult host plant. Machilus thunbergii and Terminalia catappa (need verification, see remarks) (Ong & Hattori, 2019); sometimes found on Castanopsis carlesii and Alnus formosana (Wen-I Chou, pers. comm. 2021). Remarks. Frank & Sekerka (2020) described this subspecies as elytra semi-matt and about half of specimens with impression at basal third of each elytron. However, all the three specimens we examined were without the impression mentioned above. It is the same in habitus of this species in Ong & Hattori (2019) and by personal communication with Wen-I Chou. Whether all specimens from Taiwan are without impression at basal third of each elytron needs to be verified. Ong & Hattori (2019) cited the former two host plants from Ohmomo & Fukutomi (2013) but we found it was misquoted. Four host plants listed by Ohmomo & Fukutomi (2013) are cited as the host plants of subspecies C. (C.) lewisii lewisii. It seems Terminalia catappa is not the real host and sometimes the species can be found on Castanopsis carlesii and Alnus formosana (Wen-I Chou, pers. comm. 2021).
Published: 2022
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13. Chrysodema sonnerati Lap. & Gory 1835

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema sonnerati, Taxonomy
Abstract: Key to species of the Chrysodema sonnerati species-group 1. Lateral impression on elytron does not exceed half the length of elytron.............. C. (C.) sonnerati Lap. & Gory, 1835 - Lateral impression on elytron obviously present on more than half the length of elytron.............................. 2 2. Punctation on each elytron always regular, forms 4 wide longitudinal intercostal intervals; a large elliptical principal impression protrudes from lateral impression to 3rd intercostal interval at apical third of each elytron; penis oval.................................................................................................. C. (C.) hainanensis sp. nov. - Punctation on each elytron generally irregular or weakly regular; elytral lateral impression without any transverse extended impression; penis broadly-oval........................................................................... 3 3. Dorsum strongly shiny, green or green-purple; apices of parameres rounded.......... C. (C.) lewisii lewisii Saunders, 1873 - Dorsum semi-matt, usually dark blue; apices of parameres subtruncate, inner tip angular.............................................................................................. C. (C.) lewisii nakatai Frank & Sekerka, 2020
Published: 2022
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14. Chrysodema (Chrysodema) hainanensis Qi & Ai & Song 2022, sp. nov

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema hainanensis, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) hainanensis sp. nov. (Chinese common name: 海南fl彩吉丁) (Figs 1A–C, 2A–J, 5A, 5B, 5F–H, 6D, 6E) Type localiy. China, Hainan Island, Sanya City, Tianya District, Lizhiling Mountain. Type specimens. Holotype &male; (FAFS): CHINA • Lizhiling Mountain, Tianya District, Sanya City, Hainan Province; 18°21'N, 109°27'E, alt. 390 m; 19.VI.2021; Hai-Tian Song leg. Paratypes 4 &male;&male;, 5 &female;&female; (CHTS); same collection data as for holotype; Hai-Tian Song & Zhi-Hao Qi leg. • 2 &female; (CHTS); same collection data as for the preceding; 18.VI.2021 • 4 &female; (CHTS); same collection data as for the preceding; alt. 350 m; 17.VI.2021 • 2 &male;&male;, 2 &female;&female; (CHTS); same collection data as for the preceding; 17.VI.2021; Shi-Jie Lu & Jia-Sheng Lu leg. • 6 &female;&female; (CHTS); same collection data as for the preceding; 29.VI.–14.VII.2020; Jia-Sheng Lu leg. • 1 &female; (CHTS); same collection data as for the preceding; VII.2021; Jia-Sheng Lu leg. • 1 &female; (CHTS); Sanya City, Hainan Province; V–VI.2019; local collector leg. • 1 &female; (CHTS); Liuluo Valley, Tianya District, Sanya City, Hainan Province; VIII.2020; Rong-Jia Chen leg. • 1 &female; (CHTS); same collection data as for the preceding; VI.2020 • 1 &male; (CHTS); Jianfengling National Forest Park, Ledong Li Autonomous County, Hainan Province; 24.IV.2019; Shi-Liang Mo leg. Description of holotype. Well preserved &male; specimen (Fig. 1A–C, 2C, 5A). Length 2.03 cm, width 0.66 cm, length/width ratio: 3.08. Body metallic green, shiny. Impunctate (or micropunctate) areas on dorsal side dark blue to purple. Pronotum with bright golden green mixed macropunctation, principal impressions with golden green mixed punctation. Punctation on elytra variegate golden green mixed, and elytra with metallic green regular stripes; principal and lateral impressions bright golden-green; lateral margin metallic green. Ventral side bright metallic green. Legs including tarsi metallic green, ventral pads black, basally brown. Labrum, maxillae and labium yellowish-brown; maxillar and labial palpi brown to black. Scape and pedicel metallic green, remaining antennomeres brownish-black. Pronotum moderately densely macropunctate with narrow and sparsely micropunctate interspaces. Medial line well visible with longitudinal dense macropunctation from both sides of medial line to fifth of lateral margin, followed by relatively sparse punctation with many punctures connected and forming obviously narrow longitudinal impressed lines (principal impressions) at three fifths of medial line to lateral margin, next to lateral margin are moderately densely punctate. Medial impressions shallow but distinct. Principal impressions distinct and weakly delimited from disc. Lateral impressions not obvious. Lateral margin relatively straight. Each elytron regularly with four slightly but distinctly convex costae, 1st and 2nd costa united at both base and apex of elytron, gradually narrowed. 1st slightly wider than 2nd. 3rd costa starts from lateral base and ends at apical third of elytron, 4th costa starts at basal fourth of elytron near lateral edge and ends at apex. Punctation always regular. Subscutellar interval (row of puncture distributed along basal part of elytral suture) denser at basal fifth. Other punctures form 4 wide longitudinal intercostal intervals on each elytron (1st–4th intercostal intervals). Small oval impression occurrs at base of 2nd intercostal interval and obvious oval impression at about basal third. Lateral impression (4th intercostal interval) with condensed wax layer thus appears like elongate opaque yellow stripe. Large elliptical principal impression protrudes from lateral impression to 3rd intercostal interval at apical third of elytron, and interrupts 3rd intercostal interval. Lateral impression narrowed after large elliptical principal impression, and then rewidening, before narrowingly evenly to apex. Epipleura gradually narrowed at apical third of elytron, with continuous teeth. All tibiae and femora are metallic green, with dense punctation and short setae. Outer margin of apex of protibia has nearly triangular protrusion. Ventral side of body (Fig. 1B) and lateral sides (Fig. 1C) of all abdominal ventrites very densely punctate. Ventrite V triangularly excavated at tip (Fig. 2C). Aedeagus (Fig. 5A) length: 5.22 mm, width: 1.02 mm, length/width ratio: 5.12. Apices of parameres subtruncate, inner tip relatively rounded, lateral margin of parameres relatively straight, only slightly curved. Penis oval, in ventral view with shallow and linear impression along midline, lateral sides with deep canaliculus, which has dorsal side much broader than ventral. Variation. Body &male;&male; (n=7) length: 1.66–2.34 cm, width: 0.52–0.75 cm, length/width ratio: 3.08–3.19; &female;&female; (n=21): length: 1.83–2.76 cm, width: 0.59–0.92 cm, length/width ratio: 3.00–3.10. (Main specimens see Fig. 2.) Dorsal colouration usually with distinct dark blue purple tint. Oval impressions on 2nd intercostal interval vary from moderately large to small or absent, and the number from 3, 2, 1 to 0 on one side. These features are usually positively correlated with body size. The elliptical elytral principal impression also varies in size and shape. One specimen (Fig. 2J) with elliptical principal impression and oval impression at apical third obviously connected. Structure of elytra and pronotum nearly constant. Female Ventrite V slightly notched at the tip (Fig. 2D). Aedeagus (n=5) length: 4.18–5.35 mm, width: 0.88–1.08 mm, length/width ratio: 4.75–5.12. Small-sized specimens may have some variation in the parameres, penis nearly constant (Fig. 5B). Biology and adult host plant. Adults eat leaves of Toona ciliata and can be found on the tree frequently. Etymology. The specific name is after the type locality. Distribution. China (Hainan Island). Remarks. The characters of this new species seem to be between Chrysodema sonnerati species-group and Chrysodema aurostriata species-group (sensu Frank & Sekerka, 2020). The following two characters of this new species are similar to the former species-group: 1) The principal impressions on pronotum distinct, longitudinal, narrowed; 2) elytra without obvious transverse impressions but with large elliptical principal impressions. However, with slightly but distinctly convex costae and wide longitudinal intercostal intervals on elytra are more similar to the latter species-group. Since the new species is most similar to C. (C.) lewisii nakatai at first glance, we assign it to the Chrysodema sonnerati species-group. Although many species of genus Chrysodema have been sold as ornamental beetles in large quantities for years, only a limited number of host plants have been discovered, such as Terminalia catappa (Family: Combretaceae), Machilus thunbergii (Family: Lauraceae), Castanopsis sieboldii sieboldii, Lithocarpus edulis, Quercus phillyreoides (Family: Fagaceae), Mallotus japonicus (Family: Euphorbiaceae) and Casuarina equisetifolia (larva, Family: Casuarinaceae)(Ohmomo & Fukutomi, 2013; Ong & Hattori, 2019). The adult host plant of this new species is a newly recorded family and genus for Chrysodema. Differential diagnosis. This new species presents several diagnostic characters which allow it to be easily separated from other related species: 1) Punctation on each elytron always regular, forms 4 wide longitudinal intercostal intervals, 2) most specimens with large elliptical principal impression protruding from lateral impression to 3rd intercostal interval at apical third of each elytron, 3) most specimens with oval impression in 2nd intercostal interval of each elytron at basal third, 4) apices of parameres subtruncate, inner tip relatively rounded, lateral margin of parameres relatively straight, only a slight curvature; Penis oval, no obvious widening.
Published: 2022
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15. Chrysodema Qi & Ai & Song 2022

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Key to Chrysodema species from China 1. Pronotal principal impressions small to moderately large, not reaching lateral margin of pronotum, sometimes only linear or nearly absent......................................................................................... 2 - Pronotal principal impressions large and rounded, reaching to lateral margin of pronotum, which forms a sharp carina......6 2. Elytra with strongly convex narrow and sharp costae....................... C. (C.) eximia berliozi Descarpentries, 1948 - Elytra nearly flat or slightly convex and without sharply delimited costae......................................... 3 3. Each elytron without transverse impressions............................................................... 4 - Each elytron with 1 or 2 transverse large but shallow impressions................................................ 5 4. Punctation on each elytron always regular, forms 4 wide longitudinal intercostal intervals; a large elliptical principal impression protrudes from lateral impression to 3 rd intercostal interval at apical third of each elytron..... C. (C.) hainanensis sp. nov. - Punctation on each elytron generally irregular or weakly regular; elytral lateral impression without any transverse extended impression........................................................ C. (C.) lewisii nakatai Frank & Sekerka, 2020 5. Labrum, maxillae and labium including palpi pale brown........................... C. (C.) aurostriata Saunders, 1866 - Labrum, maxillae and labium including palpi yellow............................... C. (C.) tonkinea Kerremans, 1909 6. Legs with a strong reddish tinge, dorsal surface dark blue with golden green punctures.... C. (P.) yasumatsui Kurosawa, 1954 - Legs golden green with a slight reddish tinge or simply greenish tinges on green of basal coloration, dorsal surface nearly black with golden green punctures............................................................................. 7 7. Legs golden green with a slight reddish tinge.......................... C. (P.) dalmanni dalmanni (Mannerheim, 1837) - Legs simply greenish tinges on green of basal coloration................ C. (P.) dalmanni penghuensis Hattori & Ong, 202
Published: 2022
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16. Chrysodema (Chrysodema) lewisii subsp. lewisii lewisii Saunders 1873

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema lewisii, Biodiversity, Buprestidae, Chrysodema (chrysodema) lewisii lewisii saunders, 1873, Taxonomy
Abstract: Chrysodema (Chrysodema) lewisii lewisii Saunders, 1873 (Figs 1G��I, 5C) Chrysodema lewisii Saunders, 1873: 510. = Chrysodema oschimana Nonfried, 1895: 297. Material examined. 1 &male; (Fig. 1G��I) (CHTS): JAPAN, Yonaha Mountain, Kunigami, Okinawa; 8��9.VIII.2016; Hao Xu & Jian-Yue Qiu leg.; body length 1.93 cm, width 0.67 cm, length/width ratio: 2.88; aedeagus (Fig. 5C) length: 4.95 mm, width: 1.26 mm, length/width ratio: 3.93. Distribution. Japan (Hachij�� and Tsushima islands south to Miyako Island) (Frank & Sekerka, 2020). Adult host plant. Castanopsis sieboldii sieboldii, Lithocarpus edulis, Quercus phillyreoides, Quercus spp. (Ohmomo & Fukutomi, 2013). Remarks. According to Frank & Sekerka (2020) the specimen above can certainly be identified as C. (C.) lewisii lewisii by the habitus characters and distribution, but the characters of aedeagus is obviously similar to C. (C.) lewisii nakatai. Since the apices of parameres also exhibit variation in C. (C.) hainanensis sp. nov., we consider the elytral characters sometimes more significant than the apices of parameres. Hence, we finally identified the specimen as C. (C.) lewisii lewisii. Whether the two subspecies of C. (C.) lewisii should be divided into more subspecies or synonymised may need further research with molecular methods., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 360, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1","Ohmomo, S. & Fukutomi H. (2013) The buprestid beetles of Japan. Iconographic Series of Insect 7. Mushi-sha, Tokyo, 208 pp."]}
Published: 2022
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17. Chrysodema (Pseudochrysodema) yasumatsui Kurosawa 1954

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema yasumatsui, Taxonomy
Abstract: Chrysodema (Pseudochrysodema) yasumatsui Kurosawa, 1954 (Figs 4G��I, 5E, 8A, 8F) Chrysodema yasumatsui Kurosawa, 1954: 29. Material examined. 1 &male; (Figs 4G��I, 5E) (CHTS): CHINA, Lanyu Island, Taitung City, Taiwan; 20��29.VI.2020; Hsiang-Yu Chuang & Chia-Lung Huang leg.; body length: 2.73 cm, width: 0.98 cm, length/width ratio: 2.79; aedeagus length: 8.47 mm, width: 1.64 mm, length/width ratio: 5.16. Distribution. China (Lanyu Island, Taiwan). Adult host plant. Terminalia catappa (Ong & Hattori, 2019)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 366, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Ong, U. & Hattori, T. (2019) Jewel beetles of Taiwan. Vol. 1. Kua Series, Donothing Image, Tainan, xv + 235 pp."]}
Published: 2022
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18. Chrysodema (Chrysodema) tonkinea Kerremans 1909

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema tonkinea, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) tonkinea Kerremans, 1909 (Figs 3F��I, 5L��T, 8D) Chrysodema (Chrysodema) tonkinea Kerremans, 1909: 540. Material examined. 1 &female; (Fig. 3F, 5R��T) (CHTS): CHINA, Damingshan Nature Reserve, Nanning City, Guangxi; 11.VIII.2011; Hai-Tian Song leg. �� 1 &female; (CHTS); Guangxi; 2009 �� 2 &female;&female; (Fig. 3G��I, 5L��Q) (CHTS), 2 &female;&female; (CCHZ); Damingshan Nature Reserve, Nanning City, Guangxi; 2006; Ying-Bing Li leg.; body &female;&female; length: 2.39��2.64 cm, width: 0.77��0.89 cm, length/width ratio: 2.97��3.10. Distribution. China (Guangxi, new country record); Cambodia; Laos; Vietnam. Host plant. Unknown. Remarks. This is a variable species, which is mainly reflected in the number of transverse impressions on each elytron. Three of our four specimens only with an obvious transverse principal impression on the lateral impression towards elytral suture at apical third of each elytron. Specimens in Figs 3F and 8D have an extra transverse impression at basal third of each elytron, which angles slightly to the elytral suture. It is quite similar to Figs 138 and 139 in Lander (2003), only transverse impressions on apical third are larger. Apex of ovipositor varies, slightly protruding (Fig. 5L), flat (Fig. 5O) or slightly concave (Fig. 5R). Female tergite IX also changed (Fig. 5M, P, S) but female tergite VIII nearly constant (Fig. 5N, Q, T)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 361, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp."]}
Published: 2022
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19. Chrysodema sonnerati Lap. & Gory 1835

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema sonnerati, Taxonomy
Abstract: Key to species of the Chrysodema sonnerati species-group 1. Lateral impression on elytron does not exceed half the length of elytron.............. C. (C.) sonnerati Lap. & Gory, 1835 - Lateral impression on elytron obviously present on more than half the length of elytron.............................. 2 2. Punctation on each elytron always regular, forms 4 wide longitudinal intercostal intervals; a large elliptical principal impression protrudes from lateral impression to 3rd intercostal interval at apical third of each elytron; penis oval.................................................................................................. C. (C.) hainanensis sp. nov. - Punctation on each elytron generally irregular or weakly regular; elytral lateral impression without any transverse extended impression; penis broadly-oval........................................................................... 3 3. Dorsum strongly shiny, green or green-purple; apices of parameres rounded.......... C. (C.) lewisii lewisii Saunders, 1873 - Dorsum semi-matt, usually dark blue; apices of parameres subtruncate, inner tip angular.............................................................................................. C. (C.) lewisii nakatai Frank & Sekerka, 2020, Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on pages 360-361, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1"]}
Published: 2022
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20. Chrysodema (Pseudochrysodema) dalmanni subsp. penghuensis Hattori & Ong 2020

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Chrysodema (pseudochrysodema) dalmanni penghuensis hattori & ong, 2020, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema dalmanni, Taxonomy
Abstract: Chrysodema (Pseudochrysodema) dalmanni penghuensis Hattori & Ong, 2020 Chrysodema (Pseudochrysodema) dalmanni penghuensis Hattori & Ong, 2020: 300. Material examined. No specimens were observed, please refer to Hattori & Ong, 2020. Distribution. China (Penghu Islands, Taiwan). Adult host plant. Terminalia catappa (Hattori & Ong, 2020)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on page 365, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Hattori, T. & Ong, U. (2020) Studies on the Buprestidae (Coleoptera) of Asia 10) One New Subspecies in Genus Chrysodema and Four New Species of the Genera Nipponobuprestis and Coraebus from Taiwan. Elytra, 10 (2), 299 - 318."]}
Published: 2022
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21. Chrysodema Qi & Ai & Song 2022

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Key to Chrysodema species from China 1. Pronotal principal impressions small to moderately large, not reaching lateral margin of pronotum, sometimes only linear or nearly absent......................................................................................... 2 - Pronotal principal impressions large and rounded, reaching to lateral margin of pronotum, which forms a sharp carina......6 2. Elytra with strongly convex narrow and sharp costae....................... C. (C.) eximia berliozi Descarpentries, 1948 - Elytra nearly flat or slightly convex and without sharply delimited costae......................................... 3 3. Each elytron without transverse impressions............................................................... 4 - Each elytron with 1 or 2 transverse large but shallow impressions................................................ 5 4. Punctation on each elytron always regular, forms 4 wide longitudinal intercostal intervals; a large elliptical principal impression protrudes from lateral impression to 3 rd intercostal interval at apical third of each elytron..... C. (C.) hainanensis sp. nov. - Punctation on each elytron generally irregular or weakly regular; elytral lateral impression without any transverse extended impression........................................................ C. (C.) lewisii nakatai Frank & Sekerka, 2020 5. Labrum, maxillae and labium including palpi pale brown........................... C. (C.) aurostriata Saunders, 1866 - Labrum, maxillae and labium including palpi yellow............................... C. (C.) tonkinea Kerremans, 1909 6. Legs with a strong reddish tinge, dorsal surface dark blue with golden green punctures.... C. (P.) yasumatsui Kurosawa, 1954 - Legs golden green with a slight reddish tinge or simply greenish tinges on green of basal coloration, dorsal surface nearly black with golden green punctures............................................................................. 7 7. Legs golden green with a slight reddish tinge.......................... C. (P.) dalmanni dalmanni (Mannerheim, 1837) - Legs simply greenish tinges on green of basal coloration................ C. (P.) dalmanni penghuensis Hattori & Ong, 202, Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on pages 366-367, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Frank, D. & Sekerka, L. (2020) Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I. Zootaxa, 4720 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4720.1.1"]}
Published: 2022
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22. Chrysodema (Pseudochrysodema) dalmanni subsp. penghuensis Hattori & Ong 2020

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema (pseudochrysodema) dalmanni penghuensis hattori & ong, 2020, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema dalmanni, Taxonomy
Abstract: Chrysodema (Pseudochrysodema) dalmanni penghuensis Hattori & Ong, 2020 Chrysodema (Pseudochrysodema) dalmanni penghuensis Hattori & Ong, 2020: 300. Material examined. No specimens were observed, please refer to Hattori & Ong, 2020. Distribution. China (Penghu Islands, Taiwan). Adult host plant. Terminalia catappa (Hattori & Ong, 2020).
Published: 2022
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23. Chrysodema (Chrysodema) eximia subsp. berliozi Descarpentries 1948

Author: Qi, Zhi-Hao, Ai, Hong-Mu, and Song, Hai-Tian
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Chrysodema (chrysodema) eximia berliozi descarpentries, 1948, Buprestidae, Chrysodema eximia, Taxonomy
Abstract: Chrysodema (Chrysodema) eximia berliozi Descarpentries, 1948 (Figs 3A��E, 8B, 8C) Chrysodema berliozi Descarpentries, 1948: 63. Material examined. 1 &male; (Fig. 3A��C), 2 &female;&female; (Fig. 3D, E) (CWIC): CHINA, Lanyu Island, Taitung City, Taiwan; 22.VII.1999; Wen-I Chou leg. Distribution. China (Lanyu Island and Ludao Island, Taiwan) (Tamadera & Yoshitake, 2017). Adult host plant. Terminalia catappa, sometimes found on the leaves of Mallotus japonicus (Ong & Hattori, 2019)., Published as part of Qi, Zhi-Hao, Ai, Hong-Mu & Song, Hai-Tian, 2022, Notes on the genus Chrysodema from China (Coleoptera: Buprestidae: Chrysochroinae), with one new species and one new record, pp. 355-368 in Zootaxa 5099 (3) on pages 361-362, DOI: 10.11646/zootaxa.5099.3.4, http://zenodo.org/record/6078948, {"references":["Tamadera, Y. & Yoshitake, H. (2017) A New Record of Chrysodema (Cyalithoides) eximia berliozi (Coleoptera, Buprestidae) from Ludao Is., Taiwan. Elytra, 7, 127 - 128.","Ong, U. & Hattori, T. (2019) Jewel beetles of Taiwan. Vol. 1. Kua Series, Donothing Image, Tainan, xv + 235 pp."]}
Published: 2022
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24. Chrysodema (Chrysodema) dany Frank & Sekerka 2020, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema dany, Taxonomy
Abstract: Chrysodema (Chrysodema) dany sp. nov. urn:lsid:zoobank.org:act: AD74608E-0568-437F-9A89-89B797 D88961 (Figs 31–33, 79) Type locality. Indonesia, Maluku Province, Haruku Island. Type material. HOLOTYPE: &male;, ‘ INDONESIA, / Maluku Isls, / HARUKU Isl., / xii. 1993 [w, h]’ (NMPC, ex coll. VKSC). PARATYPES (5 &male;&male;, 3 &female;&female;): 4 &male;&male;, 2 &female;&female;, same data as holotype (3 &male;&male;, 1 &female; VKSC, 1 &male;, 1 &female; DFPC); 1 &female;, ‘ INDONESIA / HARUKU isl. / xii. 1993 [w, h] // Chrysodema / mniszechi / v. aeneoviolacea / Dét. T. LANDER 19 2007 [w, h/p]’ (EJBS); 1 &male;, ‘ Indonesia / Ceram is. 12/1993 [w, p] // Chrysodema / mniszechi / aeneoviolacea Deyr. / det. S. Gottwald 2002 [w, h/p]’ (SGBG). All specimens were provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Chrysodema) / dany sp. nov. / David Frank & / Lukáš Sekerka det. I. 2019 [date handwritten]’. Description of holotype. Well preserved &male; specimen, left middle tarsus missing as well as left antennomeres IX–XI and right antennomeres VIII–XI. Length 28.75 mm, width 9.50 mm, length/width ratio: 3.03. Body very dark metallic black-blue with golden-bronze punctate areas, elytral costae gradually more bluish towards apex of elytra. Ventral pads on all tarsomeres dorsally black, only their extreme base dark brown. Head sparsely punctate, punctures foveolate, each bearing short seta, thus frons sparsely pubescent. Vertex more densely punctate, punctures bearing adherent setae. Pronotum moderately densely macropunctate, shiny parts moderately densely micropunctate and weakly micro-reticulate. Central part of disc with shiny areas. Macropunctation laterally gradually coarser and denser. Each macropuncture bearing moderately long white adherent seta. Medial line well visible, sparsely micropunctate, more or less elevated along entire length. Medial impressions distinct but shallow. Principal impressions small, elongate, and moderately deeply impressed; their punctation internally forming line of small and very dense punctures. Lateral impressions shallow, barely marked, with several shallowly impressed punctures. Lateral margin carinate in basal 2/3, then widening and flattened reaching to approx. 3/4 length of pronotum. Elytra regularly convex with distinctly elevated rounded and broad costae along suture and on intervals 2, 4, 6 and 8. Interval 6 shortened, reaching almost to 4/5 length and almost connected to interval 4. Costae moderately densely micropunctate (at magnification 50 ×) and each with a few additional confusedly dispersed macropunctures. Intercostal depressions deep and very densely macropunctate. Punctation completely irregular, not forming rows or groups of punctures. Each macropuncture bearing short adherent white seta, but impressions do not appear densely pilose. Interspaces at maximum 2 × as wide as puncture diameter but mostly as wide as puncture diameter. Epipleura in basal 1/4 broad, densely punctate, then continuously gradually narrowing apically; punctures becoming finer and vanishing towards apex. Each puncture bearing long erect white seta. Fore and mid femora in central part strongly shiny, sparsely punctate; hind femora distinctly more densely punctate. Each puncture bearing seta, setation particularly conspicuous on lower side of hind femora. Ventral side of body (Fig. 32) overall densely and moderately coarsely punctate, only central part sparsely punctate with large shiny impunctate areas. Entire lateral sides of all abdominal ventrites densely and almost uniformly punctate, only towards external margins punctation finer but even denser (Fig. 33). Aedeagus length: 8.00 mm, width: 1.56 mm, length/width ratio: 5.13. Parameres cymbiform, apices narrow and rounded. Penis narrow, elongate, subparallel-sided, ventral side slightly impressed, lateral sides with deep canaliculus, which has both margins equal (Fig. 79). Variation. Body &male;&male; (n=6) length: 25.00– 28.75 mm, width: 8.75–9.50 mm, length/width ratio: 2.86–3.03; &female;&female; (n=3): length: 30.00– 31.50 mm, width: 10.25–11.00 mm, length/width ratio: 2.86–2.98. Whole series quite constant with similar structure of pronotum and elytra. Medial and principal impressions in some specimens less impressed. Colouration slightly variable; two specimens have more distinct blue tinge on elytral costae in combination with golden punctures in intercostal depressions; two specimens blue-black with blue tinge also in intercostal depressions; remaining four specimens with intercostal depressions more or less bronze and costae distinctly bluish. Aedeagus (n=3) length: 7.44–8.00 mm, width 1.50–1.56 mm, length/width ratio: 4.96–5.13. Differential diagnosis. Chrysodema (C.) dany sp. nov. is similar to C. (C.) aeneoviolacea and C. (C.) sibuyanica as all three have dark dorsum. It can be easily distinguished by distinctly and densely micropunctate costae on elytra, thus surface appears less regular and whole dorsum appears less shiny while C. (C.) aeneoviolacea has nearly polished costae without obvious micropunctures and thus dorsum is strongly shiny. Chrysodema (C.) dany sp. nov. has elytra usually distinctly bicoloured, costae in dark tones of blue or turquoise and intercostal depressions golden while C. (C.) aeneoviolacea has dorsum bronze-brown, pronotum often with a slight violet tinge. Chrysodema (C.) sibuyanica differs in densely pubescent frons with long setae while C. (C.) dany has it sparsely and pubescent with short setae. Etymology. This species is named after Daniel ‘Dany’ Frank, son of the senior author of this paper. The species epithet is a noun in apposition. Distribution. Indonesia: Maluku Province, Haruku Island and? Seram Island (see remarks). Remarks. One paratype was supposedly collected on Seram but since it was collected by native collectors and with the same date as the rest of the type series from Haruku it might have been mislabelled. On the other hand, it is possible that C. dany sp. nov. can be found in southern Seram as it is only 6 km air distance from Haruku. The occurrence on Seram should be verified with new material and until then we consider the occurrence there as doubtful., Published as part of Frank, David & Sekerka, Lukáš, 2020, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I., pp. 1-62 in Zootaxa 4720 (1) on pages 30-31, DOI: 10.11646/zootaxa.4720.1.1, http://zenodo.org/record/3602864
Published: 2020
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25. Chrysodema (Chrysodema) aeneoviolacea Deyrolle 1864

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema aeneoviolacea, Biodiversity, Buprestidae, Taxonomy
Abstract: Key to the Chrysodema (C.) aeneoviolacea species-group 1 Dorsum dark bronze brown.............................................................................. 2 - Dorsum metallic green................................................................................. 4 2 Frons densely pubescent with long setae. Pronotum coarsely and densely punctate, principal impressions hardly visible, narrow and very shallow. Philippines. Figs 34–36, 80...................................... C. (C.) sibuyanica Fisher, 1924 - Frons sparsely pubescent with short semi-adherent setae. Pronotum coarsely and moderately sparsely punctate, principal impressions obvious, moderately impressed. Species from Moluccas............................................... 3 3 Dorsum strongly shiny, costae on elytra polished, not obviously micropunctate. Dorsum usually bronze-brown, pronotum often with slight violet tinge. Indonesia, Kai Isls., Larat Is. Figs 25–30, 78................ C. (C.) aeneoviloacea Deyrolle, 1864 - Dorsum moderately shiny, costae very distinctly and densely micropunctate. Elytra distinctly bicoloured, costae in dark tones of turquioise or blue with golden intercostal depressions. Indonesia, Haruku Is. Figs 31–33, 79........ C. (C.) dany sp. nov. 4 Third costa shortened, reaching approximately to midlength of elytron, sometimes it continues as low fold but always densely punctate, not smooth. Indonesia, Ambon Is. Figs 37–39................................ C. (C.) excellens Théry, 1923 - Third costa long, reaching distinctly at least to apical third, smooth and strongly shiny along entire length............... 5 5 Principal impressions on pronotum regular, large and cup shaped, entire concavity densely covered with small punctures. Indonesia, Ambon and Seram islands. Figs 43–48, 81................................... C. (C.) wallacei Deyrolle, 1864 - Principal impressions on pronotum never cup shaped, moderately deep and irregularly circular to oval with stripe of dense and small punctures internally. Indonesia, Gorong and Seram islands. Figs 40–42........... C. (C.) mniszechii Deyrolle, 1864, Published as part of Frank, David & Sekerka, Lukáš, 2020, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I., pp. 1-62 in Zootaxa 4720 (1) on page 25, DOI: 10.11646/zootaxa.4720.1.1, http://zenodo.org/record/3602864, {"references":["Fisher, W. S. (1924) New buprestid beetles from the Philippine Islands I. Philippine Journal of Science, 24 (1), 45 - 52.","Deyrolle, H. (1864) Description des Buprestides de la Malaisie recueillis par M. Wallace. Annales de la Societe entomologique de Belgique, 8, 1 - 272, 4 pls. https: // doi. org / 10.5962 / bhl. title. 115963","Thery, A. (1923) Etudes sur les Buprestides. (Troisieme partie). Annales de la Societe Entomologique de Belgique, 62 (1922), 193 - 270."]}
Published: 2020
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26. Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I

Author: David Frank and Lukáš Sekerka
Subjects: Male, Synonym, Chrysodema, Zoology, Biology, Subspecies, biology.organism_classification, Coleoptera, Aedeagus, Animals, Key (lock), Animal Science and Zoology, Taxonomy (biology), Subgenus, Animal Distribution, Ecology, Evolution, Behavior and Systematics, Buprestidae
Abstract: Subgeneric classification of Chrysodema Laporte de Castelnau & Gory, 1835 is revised and following synonymy is established: Chrysodema=Cyalithoides Fisher, 1922 syn. nov.; Pseudochrysodema Saunders, 1874=Leganya Hołyński, 1994 syn. nov.,=Marcsikiella Hołyński, 2014 syn. nov. Four subgenera are recognized as valid: Chrysodema s. str., Gelaeus Waterhouse, 1905, Pseudochrysodema Saunders, 1874, and Thymedes Waterhouse, 1905, and a key to the subgenera is given. Four species-groups are established in the subgenus Chrysodema s. str.: C. aeneoviolacea-group, C. aurostriata-group, C. eximia-group, and C. sonnerati-group. All known species, except the currently accepted synonyms of Chrysodema (Chrysodema) eximia Laporte de Castelnau & Gory, 1835, included in these groups are revised based on comparative study of extensive material including types of all described taxa. Three new species and one subspecies are described: Chrysodema (Chrysodema) dany sp. nov. from Haruku Island, Indonesia; C. (C.) gottwaldi sp. nov. from Palawan Island, Philippines; C. (C.) lewisii nakatai subsp. nov. from Yaeyama Islands, Japan; and C. (C.) vrabeci sp. nov. from Thailand and Laos. Chrysodema (C.) aeneoviolacea Deyrolle, 1864 stat. rev., C. (C.) dohrnii Saunders, 1874 stat. rev., and C. (C.) fuscitarsis Kerremans, 1895 stat. rev. are removed from synonymy with C. (C.) mniszechii Deyrolle, 1864; C. (C.) tonkinea Kerremans, 1909 stat. rev. is removed from synonymy with C. (C.) aurostriata Saunders, 1866. Three new synonymies are established: C. (C.) aeneoviolacea=C. (C.) elongata Kerremans, 1900 syn. nov.=C. (C.) keyensis Théry, 1923 syn. nov. (the latter was formerly synonym of C. (C.) elongata); C. (C.) dohrnii=C. (C.) fairmairei Kerremans, 1895 syn. nov. (formerly synonym of C. (C.) mniszechii). Neotype is designated for C. (C.) sonnerati Laporte de Castelnau & Gory, 1835 and lectotypes are designated for following taxa to ensure their correct application and recognition in future: C. (C.) aurostriata, C. (C.) fairmairei, C. (C.) fuscitarsis, C. (C.) sumatrensis Kerremans, 1895, C. (C.) tonkinea, C. (C.) yerburyi Waterhouse, 1905, C. (P.) coelestina Obenberger, 1922, and C. (P.) inslabilis Deyrolle, 1864. Additionally, dark species of the subgenus Pseudochrysodema are revised and as a result C. (P.) coelestina stat. rev. and C. (P.) instabilis stat. rev. are removed from synonymy with C. (P.) radians (Guérin-Méneville, 1830) and a new species C. (Pseudochrysodema) jakli sp. nov. is described from Timor Island, Indonesia. All herein included taxa are illustrated with colour photographs of habitus and the male aedeagus when available. All new taxa are diagnosed and a key to species of the C. aeneoviolacea-group is given.
Published: 2020
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27. Chrysodema (Chrysodema) gottwaldi Frank & Sekerka 2020, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema gottwaldi, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Chrysodema) gottwaldi sp. nov. urn:lsid:zoobank.org:act: 352DDA6B-0083-4DFE-8D32-3A6664613262 (Figs 58–60, 84, 89) Chrysodema fairmairei [misidentification]: Akiyama & Ohmomo (2000): Pl. 46, Figs 485-1, 485-2 (colour photographs). Chrysodema mniszechii [misidentification]: Lander (2003): 78 (colour Fig 66). Type locality. Philippines, Palawan Province, Palawan Island, Brooke’s Point. Type material. HOLOTYPE: &male;, ‘Brookes Pt. / Palawan Isl. / Jul. ‘01 [w, p] // ex Coll. / M. H. HEMPEL / (München) [w, p] // Chrysodema / fairmairei / Kerremans [w, p]’ (NMPC). PARATYPES (6 &female;&female;): 1 &female;, ‘Brookes Pt. / PALAWAN / Mai. ‘01 [w, p] // Chrysodema / fairmairei Kerr. / det. S. Gottwald 2003 [w, p]’ (SGBG); 1 &female;, ‘ Palawan Is / PHILIPPINES / 12 / Coll. T. LANDER [w, h/p]’ (MHNG); 2 &female;&female;, ‘Brook’s Point July 1992 / South Palawan PHILIPPINES / Coll. VAN DE MERGHEL [w, p]’ (1 &female; MHNG figured in LANDER (2003: 78, Fig. 66) as C. miszechii), 1 &female; DFPC); 2 &female;&female;, ‘ Philippinen / South-Palawan / 12. 1979 [w, h] // ‘ Chrysodema 1864 / mniszechii Deyrolle, / det. G. Novak 2005 [w, p]’ (ATMR, coll. R. Novak. All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Chrysodema) / gottwaldi sp. nov. / David Frank & / Lukáš Sekerka det. I. 2019 [date handwritten]’. Description of holotype. Well preserved &male; specimen, missing only last left hind tarsomere. Length 24.75 mm, width 9.00 mm, length/width ratio: 2.75. Body generally bright metallic green with golden reflections. Ventral pads on all tarsomeres dorsally brownishblack. Antennae black, basal two antennomeres metallic green. Labrum, labium and maxillae pale brown. Ventral side strongly shiny, bright metallic green, centrally golden-copper. Head moderately densely punctate, punctures foveolate, each bearing short seta, thus frons sparsely pubescent. Vertex more densely punctate and punctures bearing adherent setae. Pronotum densely macropunctate, shiny parts moderately densely micropunctate and distinctly micro-reticulate, thus not strongly shiny. Macropunctation laterally gradually coarser and denser, thus the structure is strongly rugose. Each macropuncture bearing moderately long white adherent seta. Medial line more or less visible, sparsely micropunctate, without macropunctures, flat. Medial impressions indistinct. Principal impressions moderately large, oval, relatively deep; internally with more or less visible stripe of small and dense punctures. Lateral impressions shallow, with several more or less visible macropunctures and indistinct micropunctures. Lateral margin carinate in basal 4/5, then widening and flattened and reaching almost to anterior margin of pronotum. Elytra regularly convex with slight postscutellar hump, bearing distinctly elevated sharp and moderately broad costae along suture and on intervals 2, 4, 6 and 8. Interval 2 distinctly curved on apical third. Interval 6 shortened, reaching to 1/2 length of elytra, then interrupted, before continuing for short distance as isolated branch laterally connected to interval 4. Costae moderately densely micropunctate (at magnification 50 ×) and each with a few additional confusedly dispersed macropunctures. Intercostal depressions deep and moderately densely macropunctate. Punctation completely irregular, but partly with tendency to form small groups of punctures, which appear like larger puncture and particularly visible on basal half of elytra. Each macropuncture bearing short adherent white seta, but impressions do not appear pilose. Interspaces at maximum 2 × as wide as puncture diameter but mostly as wide as puncture diameter. Epipleura in basal 1/4 broad, sparsely and moderately punctate, then abruptly constricted with obtusely angled corner and then gradually narrowing apically; punctures becoming finer and vanishing towards apex. Each puncture bearing long erect white seta. Fore and mid femora in central part strongly shiny, sparsely punctate; hind femora somewhat more densely punctate. Each puncture bearing seta, setation particularly conspicuous on lower side of hind femora. Ventral side of body (Fig. 59) overall densely and moderately coarsely punctate only central part sparsely punctate with large shiny impunctate areas. Entire lateral sides of all abdominal ventrites densely and almost uniformly punctate, only towards external margins punctation finer but even denser (Fig. 60). Aedeagus length: 8.56 mm, width: 1.61 mm, length/width ratio: 5.32. Parameres elongate-rhomboidal in outline, apices moderately broad and subtruncate. Apex of penis strongly narrowed and projecting beyond parameres. Penis ventrally deeply concave without striate laminae (Fig. 84). Ventrite VIII impressed and very densely pubescent (Fig. 89). Variation. Body &female;&female; (n=7): length: 27.00– 31.25 mm, width: 10.25–11.50 mm, length/width ratio: (2.63)2.70– 2.72. Slightly variable species. Colouration as in holotype, just one specimen with predominantly copper dorsum. Structure of elytra constant only third costa interrupted to complete. Pronotum of variable shape and structure; most specimens with trapezoidal pronotum, but one with subquadrate. Punctation of pronotum can be either slightly sparser and finer than in holotype or somewhat denser and coarser. Principal impressions shallow to moderately deep. Differential diagnosis. Chrysodema (C.) gottwaldi sp. nov. is externally quite similar to C. (C.) dohrnii because of green dorsum and to C. (C.) fuscitarsis due to dense pubescence of elytra. However, it has entirely different morphology of the male genitalia with parameres elongate (vs. broadly rhomboidal), ventral side of penis with smooth and deep concavity (vs. slightly impressed with numerous laminae), and apex of penis narrowly pointed and projecting beyond parameres (vs. broadly triangular and barely projecting beyond parameres. Chrysodema (C.) gottwaldi sp. nov. is sympatric with C. (C.) fuscitarsis but can be distinguished by green colour (vs. brown-bronze) and sparsely pubescent lateral sides of pronotum (vs. densely pubescent). See also Table 3 on page 49. Etymology. This species epithet is dedicated to Stephan Gottwald (Berlin, Germany), specialist in the Buprestidae, who kindly provided the holotype specimen. Distribution. Philippines, Palawan Province, Palawan Island. Remarks. Colour photographs of this species were already published by Akiyama & Ohmomo (2000: Pl. 46, Figs 485-1, 485-2) under the name C. (C.) fairmairei and by Lander (2003: 78, Fig. 66) as C. (C.) mniszechii. We have studied the specimen illustrated by Lander, which is here designated as paratype of C. (C.) gottwaldi sp. nov. Based on external characters shown in the photograph we are convinced that also the specimen illustrated by Akiyama & Ohmomo (2000) belongs to C. (C.) gottwaldi sp. nov. but we did not study the specimen., Published as part of Frank, David & Sekerka, Lukáš, 2020, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I., pp. 1-62 in Zootaxa 4720 (1) on pages 47-48, DOI: 10.11646/zootaxa.4720.1.1, http://zenodo.org/record/3602864, {"references":["Akiyama, K. & Ohmomo, S. (2000) The buprestid beetles of the world. In: Fujita, H. (Ed.), Mushi-Sha's Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, pp. 1 - 341, 30 pls.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp."]}
Published: 2020
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28. Chrysodema (Chrysodema) dany Frank & Sekerka 2020, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema dany, Taxonomy
Abstract: Chrysodema (Chrysodema) dany sp. nov. urn:lsid:zoobank.org:act: AD74608E-0568-437F-9A89-89B797 D88961 (Figs 31–33, 79) Type locality. Indonesia, Maluku Province, Haruku Island. Type material. HOLOTYPE: &male;, ‘ INDONESIA, / Maluku Isls, / HARUKU Isl., / xii. 1993 [w, h]’ (NMPC, ex coll. VKSC). PARATYPES (5 &male;&male;, 3 &female;&female;): 4 &male;&male;, 2 &female;&female;, same data as holotype (3 &male;&male;, 1 &female; VKSC, 1 &male;, 1 &female; DFPC); 1 &female;, ‘ INDONESIA / HARUKU isl. / xii. 1993 [w, h] // Chrysodema / mniszechi / v. aeneoviolacea / Dét. T. LANDER 19 2007 [w, h/p]’ (EJBS); 1 &male;, ‘ Indonesia / Ceram is. 12/1993 [w, p] // Chrysodema / mniszechi / aeneoviolacea Deyr. / det. S. Gottwald 2002 [w, h/p]’ (SGBG). All specimens were provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Chrysodema) / dany sp. nov. / David Frank & / Lukáš Sekerka det. I. 2019 [date handwritten]’. Description of holotype. Well preserved &male; specimen, left middle tarsus missing as well as left antennomeres IX–XI and right antennomeres VIII–XI. Length 28.75 mm, width 9.50 mm, length/width ratio: 3.03. Body very dark metallic black-blue with golden-bronze punctate areas, elytral costae gradually more bluish towards apex of elytra. Ventral pads on all tarsomeres dorsally black, only their extreme base dark brown. Head sparsely punctate, punctures foveolate, each bearing short seta, thus frons sparsely pubescent. Vertex more densely punctate, punctures bearing adherent setae. Pronotum moderately densely macropunctate, shiny parts moderately densely micropunctate and weakly micro-reticulate. Central part of disc with shiny areas. Macropunctation laterally gradually coarser and denser. Each macropuncture bearing moderately long white adherent seta. Medial line well visible, sparsely micropunctate, more or less elevated along entire length. Medial impressions distinct but shallow. Principal impressions small, elongate, and moderately deeply impressed; their punctation internally forming line of small and very dense punctures. Lateral impressions shallow, barely marked, with several shallowly impressed punctures. Lateral margin carinate in basal 2/3, then widening and flattened reaching to approx. 3/4 length of pronotum. Elytra regularly convex with distinctly elevated rounded and broad costae along suture and on intervals 2, 4, 6 and 8. Interval 6 shortened, reaching almost to 4/5 length and almost connected to interval 4. Costae moderately densely micropunctate (at magnification 50 ×) and each with a few additional confusedly dispersed macropunctures. Intercostal depressions deep and very densely macropunctate. Punctation completely irregular, not forming rows or groups of punctures. Each macropuncture bearing short adherent white seta, but impressions do not appear densely pilose. Interspaces at maximum 2 × as wide as puncture diameter but mostly as wide as puncture diameter. Epipleura in basal 1/4 broad, densely punctate, then continuously gradually narrowing apically; punctures becoming finer and vanishing towards apex. Each puncture bearing long erect white seta. Fore and mid femora in central part strongly shiny, sparsely punctate; hind femora distinctly more densely punctate. Each puncture bearing seta, setation particularly conspicuous on lower side of hind femora. Ventral side of body (Fig. 32) overall densely and moderately coarsely punctate, only central part sparsely punctate with large shiny impunctate areas. Entire lateral sides of all abdominal ventrites densely and almost uniformly punctate, only towards external margins punctation finer but even denser (Fig. 33). Aedeagus length: 8.00 mm, width: 1.56 mm, length/width ratio: 5.13. Parameres cymbiform, apices narrow and rounded. Penis narrow, elongate, subparallel-sided, ventral side slightly impressed, lateral sides with deep canaliculus, which has both margins equal (Fig. 79). Variation. Body &male;&male; (n=6) length: 25.00– 28.75 mm, width: 8.75–9.50 mm, length/width ratio: 2.86–3.03; &female;&female; (n=3): length: 30.00– 31.50 mm, width: 10.25–11.00 mm, length/width ratio: 2.86–2.98. Whole series quite constant with similar structure of pronotum and elytra. Medial and principal impressions in some specimens less impressed. Colouration slightly variable; two specimens have more distinct blue tinge on elytral costae in combination with golden punctures in intercostal depressions; two specimens blue-black with blue tinge also in intercostal depressions; remaining four specimens with intercostal depressions more or less bronze and costae distinctly bluish. Aedeagus (n=3) length: 7.44–8.00 mm, width 1.50–1.56 mm, length/width ratio: 4.96–5.13. Differential diagnosis. Chrysodema (C.) dany sp. nov. is similar to C. (C.) aeneoviolacea and C. (C.) sibuyanica as all three have dark dorsum. It can be easily distinguished by distinctly and densely micropunctate costae on elytra, thus surface appears less regular and whole dorsum appears less shiny while C. (C.) aeneoviolacea has nearly polished costae without obvious micropunctures and thus dorsum is strongly shiny. Chrysodema (C.) dany sp. nov. has elytra usually distinctly bicoloured, costae in dark tones of blue or turquoise and intercostal depressions golden while C. (C.) aeneoviolacea has dorsum bronze-brown, pronotum often with a slight violet tinge. Chrysodema (C.) sibuyanica differs in densely pubescent frons with long setae while C. (C.) dany has it sparsely and pubescent with short setae. Etymology. This species is named after Daniel ‘Dany’ Frank, son of the senior author of this paper. The species epithet is a noun in apposition. Distribution. Indonesia: Maluku Province, Haruku Island and? Seram Island (see remarks). Remarks. One paratype was supposedly collected on Seram but since it was collected by native collectors and with the same date as the rest of the type series from Haruku it might have been mislabelled. On the other hand, it is possible that C. dany sp. nov. can be found in southern Seram as it is only 6 km air distance from Haruku. The occurrence on Seram should be verified with new material and until then we consider the occurrence there as doubtful.
Published: 2020
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29. Chrysodema (Pseudochrysodema) jakli Frank & Sekerka 2020, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Chrysodema jakli, Taxonomy
Abstract: Chrysodema (Pseudochrysodema) jakli sp. nov. urn:lsid:zoobank.org:act: 7845C32C-20AC-40D2-A069-15DE60A5BED1 (Figs 70–72) Type locality. Indonesia, East Nusa Tenggara Province, Timor Island, Kolon env. Type material examined. HOLOTYPE: &female;, ‘ INDONESIA, Lesser Sundas / WEST TIMOR I. / Kolon vill. env. / IV.2016 local collector leg. [w, p]’ (MHNG). PARATYPES (17 &female;&female;): 3 &female;&female;, same data as holotype (NMPC, DFPC, SJPC); 1 &female;, ‘ INDONESIA TIMOR isl. / Mutis, Gunuag / 9°34’00’’S, 124°14’00’’E / local collector 1998 [w, p] // Chrysodema / radians / (Guerin, 1831) / Dét. T. LANDER 19 2007 [w, h/p]’ (EJBS); 2 &female;&female;, ‘ INDONESIA, Lesser Sundas / WEST TIMOR, Mutis Mts. / MOLO MT., Soe region / 500 m alt., 10 - 20.I.2016 / local collector leg. [w, p]’ (SJPC); 1 &female;, ‘ Indonesia, Lesser Sundas / TIMOR Isl., Oemelu vill env / 200 m alt S slopes of Mt. FATULEU / 1.2009, local collectors leg [w, p]’ (DFPC); 1 &female;, ‘Timor / JAN.2017 [w, p]’ (SJPC); 1 &female;, ‘Timor / APR.2017 [w, p]’ (SJPC); 3 &female;&female;, ‘TIMOR IS. [w, p]’ (SJPC); 1 &female;, ‘RI-KI. Sundainseln / Timor Isl. Juli 2007 / coll. Jakl [w, p] // Timor / July 2007 [w, p] // Chrysodema 1831 / radians (Guerin), / Timor Isl. / det. G. Novak 2010 [w, p]’ (ATMR, coll. R. Novak; 2 &female;&female;, ‘RI-KI. Sundainseln / Timor Isl. Mt. Mutis 1100m / coll. Jakl Mai 2008 [w, p] // Timor / May. 2008 [w, p] // Chrysodema 1831 / radians (Guerin), / Timor Isl. / det. G. Novak 2010 [w, p]’ (ATMR, coll. R. Novak; 1 &female;, ‘C. Rousse / Moluques [w, h]’ (MHNG). WITHOUT LOCALITY DATA: 1 &female; (DFPC). All specimens were provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Pseudochrysodema) / jakli sp. nov. / David Frank & / Lukáš Sekerka det. I. 2019 [date handwritten]’. Redescription of holotype. Well preserved &female; with all appendages. Length 32.00 mm, width 11.25 mm, length/ width ratio: 2.84. Body generally obscure metallic bronze-brown, without additional reflections; pronotum distinctly darker coloured than elytra, principal impressions somewhat metallic iridescent. Apical half of abdominal ventrites strongly metallic bronze. Antennae dark brown, only base of antennomere I rust-coloured. Labrum, labium and maxillae pale brown. Ventral pads on all tarsomeres yellowish-brown. Head coarsely but sparsely punctate, punctures foveolate, each bearing short seta, thus frons sparsely pubescent. Vertex sparsely and moderately punctate; punctures bearing adherent setae. Pronotum moderately densely macropunctate; interspaces distinctly micro-reticulate and finely and sparsely micropunctate, not shiny. Macropunctation laterally gradually coarser and denser. Each macropuncture bearing moderately long white adherent seta. Medial line well visible, moderately elevated, anterior quarter very conspicuously micropunctate, then punctures gradually weaker, and apical 1/4 without micropunctures. Medial impressions distinct, moderately deep. Principal impressions large formed by one deep concavity; its punctation much finer and shallower, sparse centrally and gradually denser marginally. Lateral impressions moderately deep, with several macropunctures. Border between principal and lateral impression forming high and relatively sharp carina. Lateral margin reaching to approx. 5/6 length of pronotum, carinate in whole length. Base of pronotum with broad, impunctate and slightly elevated transverse ridge on each side. Elytra regularly convex; basal 1/4 smooth without costae, apical 3/4 with more or less distinct costae along suture and on intervals 2, 4, 6 and 8. Interval 6 not shortened and connected to interval 4 on apical slope. Costae low and irregular, partly connected by low transverse rugosities. Elytra completely irregularly punctate including intervals; without distinct micropunctation (at magnification 50 ×). Punctures deeply impressed and foveolate, moderately densely arranged; interspaces 1–8 × as wide as puncture diameter. Punctures centrally (between sutural and interval 8) forming irregular groups of 2–4 punctures; each group more or less impressed. Each macropuncture bearing short adherent white seta, but impressions do not appear pilose. Epipleura in basal 1/4 broad, densely punctate, then continuously constricted and gradually narrowing apically; punctures becoming finer and vanishing towards apex. Each puncture bearing long erect white seta. Fore and mid femora in central part strongly shiny, sparsely punctate; hind femora somewhat more densely punctate. Each puncture bearing seta, setation particularly conspicuous on lower side of hind femora. Ventral side of body (Fig. 71) overall densely and moderately coarsely punctate, only central part sparsely punctate with large shiny impunctate areas. Lateral sides of abdominal ventrite I very densely punctate with small punctures, only narrow area along apical margin shiny with several larger and sparsely arranged punctures. In remaining abdominal ventrites basal half of lateral sides densely punctate, and apical ones sparsely punctate (Fig. 72). Variation. Body &female;&female; (n=17): length: (29.50)31.00–36.00(38.00) mm, width: 10.25–13.50 mm, length/width ratio: (2.60)2.66–2.88. Quite constant species showing little variability in punctation, which can be weaker and thus appear slightly sparser. In some specimens micropunctation on pronotum very fine and practically invisible including medial line. Basal ridge of pronotum variable, in most specimens missing or more or less defined as impunctate area. Central part of each elytron at least between intervals 2 and 6 with impressed groups of punctures but in some specimens these groups small and thus not as obvious. Aedeagus unknown, all studied specimens are females. Differential diagnosis. For differential diagnosis see Table 4. Etymology. This species is named after our friend Stanislav Jákl (Prague, Czech Republic), specialist in the Cetoniinae and donor of the holotype. Distribution. Indonesia, East Nusa Tenggara Province, Timor Island. The species possibly also occurs in East Timor. Remarks. The old specimen from ‘Moluques’ is morphologically absolutely identical to recent material from Timor, therefore we designate it as paratype and assume it was perhaps mislabelled., Published as part of Frank, David & Sekerka, Lukáš, 2020, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part I., pp. 1-62 in Zootaxa 4720 (1) on pages 55-56, DOI: 10.11646/zootaxa.4720.1.1, http://zenodo.org/record/3602864
Published: 2020
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30. Jewel beetles (Coleoptera, Buprestidae) collected during the 2018 field survey on Iejima Island, the Okinawa Islands, Ryukyus, south-western Japan

Author: Hiraku Yoshitake and Yutaka Tamadera
Subjects: 0106 biological sciences, Agrilus, Insecta, Asia, Arthropoda, Fauna, Biodiversity & Conservation, 010607 zoology, Zoology, 010603 evolutionary biology, 01 natural sciences, Coraebus, distribution, Animalia, new record, Buprestoidea, Buprestidae, Chalcophora japonica, fauna, Ecology, Evolution, Behavior and Systematics, Ecology, biology, Chrysodema, Field survey, biology.organism_classification, Coleoptera, Geography, Buprestid beetle, Paratrachys, Taxonomic Paper
Abstract: Only one jewel beetle, Chrysodema (Marcsikiella) dalmanni (Eschscholtz) (Coleoptera, Buprestidae) has hitherto been recorded from Iejima Is. in the Okinawa Isls., Ryukyus, south-western Japan.A total of seven jewel beetles were collected on Iejima Is. The following six species are newly recorded from the island: 1)Paratrachys princeps chujoiKurosawa, 2) Chrysodema (Chrysodema) lewisii Saunders, 3)Chalcophora japonica oshimanaSchönfeldt, 4)Coraebus hastanusGory and Laporte, 5)Sambus quadricolor quadricolorSaunders and 6)Agrilus okinawensis shiozakiiTôyama.
Published: 2019

31. Chrysodema (Gelaeus) cyanicollis Kerremans 1900

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema cyanicollis, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) cyanicollis Kerremans, 1900, assign. nov. (Figs 42–44) Chrysodema cyanicollis Kerremans, 1900: 62 (original description). KERREMANS (1903):75 (catalogue); OBENBERGER (1926): 131 (catalogue). Chrysodema (Chrysodema) cyanicollis: KERREMANS (1909):518 (key), 567 (redescription); LANDER (2003): 13 (key), 17 (redescription), Fig. 7 (colour photograph); BELLAMY (2008): 529 (catalogue). Type locality. Indonesia, South Sulawesi Province, Selayar Islands, Tanahjampea (or Jampea or Djampea) Island. Type material examined. HOLOTYPE (by monotypy): &female;, ‘DJAMPEA, / Dec 95. / A. Everett [w, p] // Chrysodema / cyanicollis / Kerr. Type [w, h] // cyanicollis Kerr. [w, h] // Coll. / Jul. Moser // Chrysodema / cyanicollis / Kerr. / Det. T. LANDER 2000’ (MFNB). Specimen provided with an additional red printed label: ‘HOLOTYPE (by monopyty) / Chrysodema (Gelaeus) / cyanicollis / KERREMANS, 1900 &female; / David Frank & / Lukáš Sekerka labeled 2016 [year handwritten]’. Additional material examined. INDONESIA: SOUTH SULAWESI PROV.: ‘ Celebes, Somarisi’ [probably Somarisi on Selayar Island 16 km S offshore of Sulawesi], 1 &female; (MHNG), 1 &female; (DFPC). EAST NUSA TENGGARA PROV.: Lembata [or Lomblen] Island, 1 &female; (MHNG). Redescription of holotype. Preserved &female; specimen with all tarsi partly broken. Length 27.00 mm, width 9.50 mm, length/width ratio: 2.84. Head green with slight golden and bluish reflections. Pronotum brightly bluish-green, gradually more bluish towards base, punctation in impressions green. Scutellum bright bluishgreen. Elytra bluish-black, only lateral impressions and two outer rows of punctures dark copper-green (Fig. 42). Ventral side bright green gold, legs green, femora with bluish tint, tarsi metallic green with bluish tint (Fig. 43). Pronotum in general coarsely and moderately densely punctate. Medial line well marked, impunctate. Medial impressions barely impressed, moderately densely punctate, punctures slightly smaller than those lateral to them. Principal impressions moderately deep, elongate, not connected to basal impressions, apparently impunctate only with row of very fine punctures. Lateral impressions shallow and punctate. Basal impressions moderately deep, subtriangular. Elytra moderately and quite densely punctate, with shallow lateral impressions present as round groups of coarser and paler coloured punctures. Posterior impression larger than anterior and somewhat more impressed (Fig. 44). Puncture rows visible from base to apex, distinctly impressed on apical 2/3, more or less regularly arranged, and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra more or less regularly arranged, not grouped, inner rows somewhat less regular; spaces 1–2× as broad as diameter of punctures, flat not reticulate. Intervals basally flat, not elevated, broad, approximately 3–4× as wide as rows of punctures, impunctate; apically and laterally slightly elevated and punctate. Mid femora in central part strongly shiny, sparsely punctate with fine punctures, on sides punctures smaller but coarser and densely arranged. Ventral side of body (Fig. 43) overall coarsely and quite densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Lateral side of abdominal ventrite I moderately dense and coarsely punctate; ventrites II to V gradually more sparsely punctate in apical half. Variation. Body &female;&female; (n = 4) length: 25.50–27.50 mm, width: 8.75–9.75 mm, length/width ratio: 2.82–3.00. Known only from female specimens. Pronotum metallic green, green with purple tint or bluish-green. Elytra from bluish-black to dark blue or green. Ventral side metallic green to golden-green. Structure of pronotum and elytra quite uniform in examined specimens, only one specimen with denser and coarser punctation and moderately deep medial impression. Differential diagnosis. Chrysodema (G.) cyanicollis belongs among species with distinctly dichromatic dorsal side having pronotum brighter coloured than elytra, latter in tones of green or blue. All similar taxa (C. (G.) walkeri, C. (G.) sara sp. nov., C. (G.) katka sp. nov. and C. (G.) cupriventris) differ in pale coloured tarsi, at most with terminal tarsomere metallic while C. (G.) cyanicollis has tarsi uniformly metallic green. The uniformly green specimen (from Lembata Is.) of C. (G.) cyanicollis is also very similar to C. (G.) florensis as both have metallic tarsi and distinct lateral impressions on elytra. It can be distinguished by lateral sides of abdominal ventrites II–III variously punctate with large impunctate areas (Fig. 4) while C. (G.) florensis has them densely and uniformly punctate without impunctate areas (Fig. 5). Distribution. Indonesia: South Sulawesi Prov.: Tanahjampea (or Jampea) Is., Selayar Is. and East Nusa Tenggara Prov.: Lembata Is. Remarks. KERREMANS (1900) described Chrysodema cyanicollis from the collection of Meyer Darcis and explicitly mentioned only a single female specimen with size ‘Long., 26; larg., 8,5 mill.’. The latter collection was purchased by Julius Moser and is nowadays deposited in MFNB. The holotype is however somewhat larger (27.0 × 9.5 mm) than indicated in the original description. Nevertheless, we consider it as a mistake as the specimens otherwise perfectly fits the original description. Chrysodema (G.) cyanicollis was until now placed in the nominotypical subgenus, however, it posses deep triangular basal impression of the pronotum typical for Gelaeus and therefore we assign it to the latter subgenus. The uniformly green specimen from Lembata resembles C. (G.) florensis, however, in our opinion it is morphologically more similar to C. (G.) cyanicollis despite considerable geographic separation. More specimens from Selayar Islands and Lembata Island are necessary to verify its identity., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 681-682, DOI: 10.5281/zenodo.5310800, {"references":["KERREMANS C. 1900: Buprestides Indo-Malais, troisieme partie. Memoires de la Societe Entomologique de Belgique 7: 61 - 93.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","OBENBERGER J. 1926: Buprestidae I. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, 212 pp.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","LANDER T. 2003: Revision du genre Chrysodema. Collection Systematique, Vol. 8, Magellanes, Andresy, 94 pp.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp."]}
Published: 2016
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32. Chrysodema (Gelaeus) walkeri subsp. walkeri walkeri (Waterhouse 1892

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema (gelaeus) walkeri walkeri (waterhouse, 1892), Chrysodema, Chrysodema walkeri, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) walkeri walkeri (Waterhouse, 1892) (Figs 24–26, 62) Pseudochrysodema (?) Walkeri Waterhouse, 1892: 411 (original description). Chrysodema Walkeri: KERREMANS (1903): 76 (catalogue); KUROSAWA (1982): 190 (noted as type species of Gelaeus). Gelaeus Walkeri: WATERHOUSE (1905): 584 (noted); OBENBERGER (1926): 129 (catalogue); THÉRY (1935): 248 (key, catalogue); VOLKOVITSH (2001): 46 (noted); BELLAMY (2003): 35 (catalogue). Chrysodema (Gelaeus) Walkeri: KERREMANS (1909): 509 (key), 510 (redescription, incl. colour Fig. 5, Pl. 22); HO&Lstrok;Y&Nacute;SKI (1994): 71 (noted as type species of Gelaeus in key to subgenera); LANDER (2003): Fig. 152 (colour photograph, not typical C. (G.) walkeri), BELLAMY (2008): 542 (catalogue). Type locality. Indonesia, Maluku Province, Damar Island. Type material examined. LECTOTYPE (present designation): &female;, ‘Damma I. [6.–11. xi.1891, J. J. Walker lgt.] / 92—20. [w, p] / Pseudochrysodema? / Walkeri / (Type) Waterh. [w, h] / Type [r, p] / 5766 [w, p]’ (BMNH). The specimen provided with an additional red printed label: ‘ LECTOTYPE &female; / Chrysodema (Gelaeus) / walkeri walkeri / (WATERHOUSE, 1892) / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Additional material examined. INDONESIA: MALUKU PROV.: Damar Island, 1 &female; (NMPC), 1 &female; (MNHN), 9 &female;&female; (MFNB), 2 &female;&female; (DFPC), 1 &female; (RMNH), 1 &male; 2 &female;&female; (RMNH, ex coll ZMAN), 6.–11. xi.1891, 1 &female;, J. J. Walker lgt. (BMNH). Redescription of lectotype. Well preserved &female; specimen missing only left fore ultimate tarsomere with claws. Length 31.75 mm, width 11.75 mm, length/width ratio: 2.70. Body strongly shiny. Body generally bright green with golden and copper reflections, only tibiae with bluishgreen reflections (Figs 24–26). Elytra deep blue-violet, apex with purple reflections (Figs 24, 26). Tarsi brown with metallic green distal half of last tarsomere and claws. Pronotum in general coarsely but sparsely punctate, only laterally punctures dense. Medial line well marked and impunctate. Medial impressions very weak, impressed only along medial line, basically present as rows of dense punctures, punctures slightly smaller than remaining ones on disc. Principal impressions present as narrow, moderately deep and sharply bordered line almost connected to basal impressions, with small dense punctures. Lateral impression shallow and punctate. Basal impressions large and triangular. Elytra without lateral impressions, moderately punctate. Rows visible from base to apex, only apically weakly impressed, generally regularly arranged, and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra quite regular and sparse, forming distinct rows, not isolated groups; spaces much broader than puncture diameter, flat, not reticulate (Fig. 26). Intervals basally flat, not elevated, broad, approximately 4–6× as wide as rows of punctures, impunctate; apically and laterally somewhat elevated and punctate. Mid femora in central part strongly shiny, sparsely punctate, punctures large, on sides punctures smaller and densely arranged. Ventral side of body (Fig. 25) overall coarsely and quite densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Almost whole lateral sides of abdominal ventrite I densely and coarsely punctate, punctures small. Lateral sides of ventrite II mostly similarly punctate to ventrite I only basal line impunctate. Ventrites III–V largely smooth and sparsely punctate, dense punctation present as patch in lateroapical corners. Variation. Body &male; (n = 1) length: 31.25 mm, width: 11.00 mm, length/width ratio: 2.84; &female;&female; (n = 18) length: (26.50)29.25–35.00 mm, width: (9.50) 10.50–12.75 mm, length/width ratio: (2.66)2.71–2.85. Medial impressions in some specimens more impressed thus medial line looks elevated. Pronotum a variable tone of green but mostly with more or less distinct bluish tint. Principal impressions variable but always with more or less deeply impressed furrow. Two specimens have tarsi brown except for metallic claws. Remaining 17 studied specimens have last tarsomere metallic green. Aedeagus (Fig. 62) (n = 1) length: 8.11 mm, width 1.63 mm, length/width ratio: 4.97. Apex of parameres rounded. Penis elongate, subparallel-sided. Differential diagnosis. The nominotypical subspecies can be easily distinguished from others by usually metallic coloured ultimate tarsomere (brown in other subspecies) and has smoothest elytra with sparsest punctation of all the subspecies. For additional characters see Table 1. Chrysodema (G.) walkeri can be distinguished from other species of Gelaeus by dichromatic dorsal side of body with green pronotum and blue elytra. Similar colour combination is also present in C. (G.) cyanicollis (differing in uniformly metallic green tarsi) and in C. (G.) sara sp. nov. (differing in smooth surface of elytra without impressed rows or elevated intervals). Distribution. Indonesia: Maluku Prov.: Damar Is. Remarks. WATERHOUSE (1892) did not state how many specimens he had at his disposal when describing C. (G.) walkeri. We have located two specimens in BMNH with identical locality label ‘Damma I. / 92—20’, however, only one is labelled as type and has the original label handwritten by C. O. Waterhouse. The latter specimen perfectly matches the original description and has identical size to that of mentioned in the original description hence we designate this specimen a lectotype in order to fix identity of this taxon. We do not consider the other specimen as part of the type series as it has larger size than mentioned in the original description and the specimen does not bear any proof that it was studied by Waterhouse when describing C. (G.) walkeri. It is listed among Additional material examined with supplemented data according to the BMNH collection register. The number ‘[18]92—20’ refers to the following entry in the register: ‘458 coleoptera, various localities presented by the Lords of the Admiralty: collected by Mr J. J. Walker during the cruise of HMS ‘Penguin’; [Nos.] 5762–5988 (266 coleoptera) Damma Island Nov[ember] 6–11 [18]91’ (M. Barclay, pers. comm.). AKIYAMA & OHMOMO (2000: plate 42) illustrated three forms of C. (G.) walkeri, however, they belong to C. (G.) walkeri nigriventris subsp. nov. (Figs 445-1, 445-2) and C. (G.) sara sp. nov. (Fig. 445-3)., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 705-708, DOI: 10.5281/zenodo.5310800, {"references":["WATERHOUSE C. O. 1892: Two new Buprestidae from Damma Island. Annals and Magazine of Natural History, Series 6 10: 410 - 412.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","KUROSAWA Y. 1982: A remarkable convergence found in Malayan buprestid beetles, with description of two new species from Thailand and Hainan. Bulletin of the National Science Museum, Series A, Zoology (Tokyo) 8: 173 - 204.","WATERHOUSE C. O. 1905: Observations on Coleoptera of the family Buprestidae, with descriptions of new Species. Annals and Magazine of Natural History, Series 7 15: 577 - 584.","OBENBERGER J. 1926: Buprestidae I. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, 212 pp.","THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","VOLKOVITSH M. G. 2001: The comparative morphology of antennal structures in Buprestidae (Coleoptera): evolutionary trends, taxonomic and phylogenetic implications. Part 1. Acta Musei Moraviae, Scientiae Biologicae 86: 43 - 169.","BELLAMY C. L. 2003: An illustrated summary of the higher classification of the superfamily Buprestoidea (Coleoptera). Folia Heyrovskyana, Supplementum 10: 1 - 197 pp. + 44 pls.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","HOLYNSKI R. B. 1994: A review of Chrysodema C. G. (Coleoptera: Buprestidae) I. The subgenera Tamamushia M. C. and Thymedes Wath. Annals of the Upper Silesian Museum, Entomology 5: 69 - 96.","LANDER T. 2003: Revision du genre Chrysodema. Collection Systematique, Vol. 8, Magellanes, Andresy, 94 pp.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","AKIYAMA K. & OHMOMO S. 2000: The buprestid beetles of the world. In: FUJITA H. (ed.): Mushi-Sha's Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, 341 pp. + 30 pls."]}
Published: 2016
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33. Chrysodema (Gelaeus) florensis Lansberge 1880

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema florensis, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) florensis Lansberge, 1880, assign. nov. (Figs 5–8, 56) Chrysodema Florensis Lansberge, 1880: cxxxii (original description). KERREMANS (1903): 75 (catalogue); OBENBERGER (1926): 131 (catalogue). Chalcophora florensis: KERREMANS (1885): 126 (catalogue). Chrysodema (Chrysodema) florensis: KERREMANS (1909): 514 (key), 527 (redescription); LANDER (2003): 13 (key), 18 (redescription), Figs 8–9 (colour photograph); BELLAMY (2008): 531 (catalogue). Gelaeus foraminifera ? var. florensis: THÉRY (1935): 248 (catalogue). Type locality. Indonesia, East Nusa Tenggara Province, Flores Island. Type material examined. LECTOTYPE (present designation):&male;, ‘Florensis / Lansbge / Flores / Type [w, h] // Flores.[w, p] // TYPE [r, p] // MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [y, p] // SYNTYPE [r, p] // MNHN / EC4802 // Chrysodema / florensis / Lansb / Dét. T. LANDER 2000 [w, h/p]’ (MNHN). PARALECTOTYPE: &female;, ‘ Flores. [w, p] // TYPE [r, p] // MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [y, p] // SYNTYPE [r, p] // MNHN / EC4803 // SYNTYPE / Chrysodema / (Chrysodema) / florensis Lansberge, 1880 ’ (MNHN). Both specimens provided with an additional red printed label: ‘ LECTOTYPE [or PARALECTOTYPE respectively] [sex] / Chrysodema (Gelaeus) / florensis / LANSBERGE, 1880 / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Additional material examined. INDONESIA: EAST NUSA TENGGARA PROV.: Flores Island, 1 &female; (MHNG; the locality label is identical to the type specimens), x.1896, 1 &female;, Everett lgt. (MFNB). WEST NUSA TENGGARA PROV.: Sangeang Island, iii. 2008, 1 &female; (DFCP); Lombok Island, xii. 2000, 1 &male; (MHNG). Redescription of lectotype. Well preserved &male; specimen with all appendages intact. Length 25.75 mm, width 9.00 mm, length/width ratio: 2.86. Body generally metallic green. Pronotum green with slight bluish reflections (in lateral or apical view), basal impressions golden. Elytra green with centre and first two elevated ribs golden-copper, anterior lateral impressions golden and posterior ones copper (Figs 6, 8). Ventral parts with slight golden reflections laterally (Fig. 7). Legs including tarsi and base of claws metallic green. Pronotum densely (laterally) to moderately dense (centrally) and coarsely punctate. Punctures on lateral sides foveolate, more deeply impressed. Medial line well visible, impunctate but not elevated. Medial impressions hardly impressed basically on same level as medial line and visible as stripe of moderately coarse punctures. Principal impressions weak and hardly separated from disc, sloping and present as elongate groups of small coarse densely arranged punctures with long setae. Surface between medial and principal impressions with sparsely dispersed punctures without larger impunctate areas. Lateral impressions moderately deep with several large punctures. Basal impressions large, oval. Elytra with broad (ca. 5× wider than puncture rows), nearly impunctate and strongly elevated sutural, 2nd and 4th intervals. Puncture rows impressed from base to apex, in apical half irregularly double and in apical third gradually coalesced, particularly between suture and 2nd interval. Punctation of elytra overall moderately coarse and very dense (except on intervals), distance between punctures mostly smaller than puncture diameter. Puncture rows V–VIII in mid third of elytra arranged in small groups of 2–4 punctures separated by more or less elevated impunctate folds forming weak reticulation. Each elytron with three impressions: 1) small, elongate situated between weakly visible 6th and 8th interval in basal third length, 2) very small, round, situated centrally between 2nd and 4th interval in basal third length, and 3) large, round, occupying whole width between 4th and 8th interval situated in apical third length (Figs 6, 8). Femora in central part not strongly shiny, moderately densely punctate, punctures large; on sides with very densely arranged punctures. Ventral side of body (Fig. 7) overall coarsely and very densely punctate, only central parts of thoracic ventrites sparsely punctate. All of lateral sides of all abdominal ventrites densely and coarsely punctate (Fig. 5), only last two with faint impunctate line apically. Aedeagus (Fig. 56) length: 6.32 mm, width 1.33 mm, length/width ratio: 4.75. Apices of parameres obliquely truncate. Penis stout-oval. Variation. Body &male;&male; (n = 2) length: 25.50–25.75 mm, width: 9.00 mm, length/width ratio: 2.83–2.86; &female;&female; (n = 4): length: 26.00– 30.50 mm, width: 9.25–11.00 mm, length/width ratio: 2.63–2.81. Body colouration always green in background in some specimens with strong copper sheen on most of body. Lateral impressions on elytra present in all examined specimens and of constant size, anterior one always elongate and posterior always round. Central impression variable in size or absent. Structure of pronotum in all specimens quite similar or punctation only slightly sparser. Aedeagus (n = 2) length: 6.32–6.73 mm, width 1.33–1.43 mm, length/width ratio: 4.71–4.75. Differential diagnosis. Chrysodema (G.) florensis can be easily separated from the other taxa by elytra with large lateral impressions in combination with dense punctation of elytra. The only similar species is C. (G.) foraminifera, which however differs in the mostly yellow tarsi with only last tarsomere metallic (completely metallic in C. (G.) florensis) and posterior lateral impression elongate or if round then divided into two. Other taxa of Gelaeus have either the lateral impressions of elytra barely indicated or they are absent entirely. However we have examined a single uniformly green specimen of C. (G.) cyanicollis, which has small but distinct lateral impressions on elytra and green tarsi and thus is superficially very similar to C. (G.) florensis. However, it can be separated at once by lateral portion of abdominal ventrites II–III variously punctate with large impunctate areas while C. (G.) florensis has lateral portions of respective ventrites densely and uniformly punctate without impunctate areas. Distribution. Indonesia: East Nusa Tenggara Prov.: Flores Is., Lombok Is. and West Nusa Tenggara Prov.: Sangeang Is. Remarks. LANSBERGE (1880) described Chrysodema florensis but did not mention the number of specimens he had at his disposal or where they were deposited. However, he gave range of lengths and mentioned male and female, therefore he must have had at least two specimens. The collection of Lansberge was purchased by R. Oberthür (nowadays in MNHN) and some duplicate specimens are also deposited in IRSN, collection of J. Neervoort van de Poll (purchased by A. Théry and nowadays in MNHN), and Museo Civico di Storia Naturale Giacomo Doria, Genova (HORN & KAHLE 1935, 1936). We consulted the respective museums and have located two specimens in MNHN (ex coll. R. Oberthür), one having original identification label by Lansberge. LANSBERGE (1880) stated: ‘Long. 12–15, lat. 5–6 mill.’, however, the actual size of the specimens is 25 mm in male and 30 in female. As both fit well the original description except of the size of body we consider them syntypes and suppose that the small size printed in description is the result of an error. We designate the male specimen a lectotype to conserve the status of this species for the purpose of this revision and avoid any further misinterpretation as additional specimens might exist., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 691-694, DOI: 10.5281/zenodo.5310800, {"references":["LANSBERGE J. W. VAN 1880:. Description de quelques Coleopteres de la Malaisie et de la Papouasie. Comptes-Rendus des Seances de la Societe Entomologique Belgique 23: cxviii - cxxxix.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","KERREMANS C. 1885: Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique 29: 119 - 157.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","LANDER T. 2003: Revision du genre Chrysodema. Collection Systematique, Vol. 8, Magellanes, Andresy, 94 pp.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","LANSBERGE J. W. VAN 1879: Diagnoses de quelques especes nouvelles de Buprestides et de Scarabaeides de la Malaisie. Comptes-Rendus des Seances de la Societe Entomologique de Belgique 22: cxlvii - clv.","KERREMANS C. 1898: Buprestides nouveaux de l'Australie et des regions voisines. Annales de la Societe Entomologique de Belgique 42: 113 - 182.","NOVAK G. 2010: Chrysodema moluensis n. sp. von den Inseln Molu und Larat, Indonesien (Coleoptera: Buprestidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 62: 107 - 109.","WATERHOUSE C. O. 1892: Two new Buprestidae from Damma Island. Annals and Magazine of Natural History, Series 6 10: 410 - 412.","HORN W. & KAHLE I. 1935: Uber entomologische Sammlungen, Entomologen & Entomo-Museologie (Ein Beitrag zur Geschichte der Entomologie). Teil I. Entomologische Beihefte (Dahlem) 2: i - vi + 1 - 160 + 1 - 16 pls.","HORN W. & KAHLE I. 1936: Uber entomologische Sammlungen, Entomologen & Entomo-Museologie (Ein Beitrag zur Geschichte der Entomologie). Teil II. Entomologische Beihefte (Dahlem) 3: 160 - 296 + pls. 17 - 26."]}
Published: 2016
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34. Chrysodema (Gelaeus) walkeri subsp. nigriventris Frank & Sekerka 2016, subsp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Chrysodema walkeri, Animalia, Chrysodema (gelaeus) walkeri nigriventris, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) walkeri nigriventris subsp. nov. (Figs 4, 33–35, 65) Gelaeus Walkeri var. nigriventris Théry, 1935: 248 (unavailable infrasubspecific name). BELLAMY (2008): 542 (catalogue, listed as synonym of C. (G.) walkeri). Gelaeus walkeri walkeri [misidentification]: AKIYAMA & OHMOMO (2000): Pl. 42, Figs 445-1, 445-2 (iconography). Chrysodema walkeri: LANDER (2003): Fig. 152 (colour photograph). Type locality. Indonesia, Maluku Province, Moa Island. Type material. HOLOTYPE: &male;, ‘Ins. Moa [w, p] // Coll. / Jul. Moser [w, p]’ (MFNB). PARATYPES (5 &male;&male; 77 &female;&female;): 1 &female;, ‘Ins. Moa [w, p] // COLLECTION / de BONNEUIL [w, p] // Pseudochrysodema Saund / Walkeri / I. Moa? [w, h] // Pseudochrysodema / Walkeri / v. nigriventris / Thery / PARATYPE [w, h]’ (NHMB, coll. G. Frey); 2 &female;&female;, ‘Ins. Moa [w, p]’ (NHMB, coll. G. Frey); 1 &female;, ‘Ins. Moa [w, p] // Chrysodema / walkeri Waterh. [w, h]’ (NHMB, coll. G. Frey); 1 &female;, ‘Moa / b. Timor [w, p] // Chrysodema / walkeri Wat. / Moa (Timor) [w, h] // walkeri [w, h]’ (NHMB, coll. G. Frey); 1 &female;, ‘Ins. Moa [w, p] // Chrysodema / walkeri / Wat / A. Descarpentries det. [w, h/p]’ (MHNH); 1 &female;, ‘Ile Moa [w, h] // Molukken / Ins. Moa [w, h] // cupriventris / Kerrem. [w, h] // Walkeri / Waterh. [w, h] // Chrysodema / Walkeri v. / nigriventris Théry / A. Descarpentries det. / ne doit pas éte décrit [w, h/p]’ (MHNH); 1 &male; 45 &female;&female;, same data as holotype (1 &male; 43 &female;&female; MFNB, 2 &female;&female; DFPC); 1 &female;, same data as holotype plus additional label ‘ Gelaeus // Walkeri Wat. [w, h]’ (MFNB); 1 &female;, ‘ Ins. Moa [w, p] // Museum Leiden / Ex coll. / G. van Roon [w, p] // Chrysodema / walkeri (WATH.) / nigriventris (THY.) / det. R. HOLY&Ncaron;SKI 2004 [w, p/h]’ (RMNH); 1 &female;, ‘ Ins. Moa [w, h] // Gelaeus Walkeri / Wat. / Det. Dr. Obenberger [w, h/p]’ (NMPC); 4 &female;&female;, ‘Ile Moa / INDONESIE / v.1992 / Coll. T. LANDER [w, h/p] // Galaeus / walkeri / (Waterh. 1892) / Dét. T. LANDER 1992 [w, h/p]’ (MHNG); 1 &female;, ‘ Ile de Moa / v.1992 / Coll. T. LANDER [w, h/p] // Galeus / walkeri / (Waterh. 1892) / Dét. T. LANDER 1993 [w, h/p]’ (MHNG); 1 &female;, ‘INDONESIE / Moa Isl. / vii.1998 / Coll. T. LANDER [w, p/h] // Gelaeus / walkeri / (Wat. 1892) / Dét. T. LANDER 2000 [w, h/p]’ (MHNG); 1 &female;, ‘ GELAEUS WALKERI WALKERI / (bon A2, lot de 5 pcs) / Moa / INDONESIE, iii.2001 / 8462 L [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p]’ (NMPC); 3 &female;&female;, ‘ INDONESIA, iii.2001 / Moa Is. / local collector [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p]’ (2 &female;&female; NMPC, 1 &female; IRSN); 1 &female;, ‘ INDONESIA, iii.2001 / Moa Is. / local collector [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p] / Gelaeus / walkeri Wat. / Sv. Bílý det.’ (NMPC); 4 &female;&female;, ‘ INDONESIA, / Maluku, March 2001, / MOA isl., / native coll. [w, p]’ (3 &female;&female; VKSC, 1 &female; DFPC); 1 &male;, ‘ Moa / Feb. 2002 [w, p]’ (VKSC); 1 &male;, ‘ Moa / Juni. 2003 [w, p]’ (DFPC); 4 &female;&female;, ‘MOA ISL. / 2/2008 [w, h]’ (3 &female;&female; SJPC, 1 &female; DFPC); 2 &male;&male; 2 &female;&female;, ‘Indonesia, SW Moluccas / MOA ISLAND., 0 - 200 m alt / E of TIMOR., 1.2009 / KAIWATU vill / local collectors lgt [w, p]’ (1&male; 2&female;&female; SJPC, 1&male; DFPC); 1&female;,‘SUMATRA / Pulau [= Island] Moa [w, p] / GALEUS / walkeri Waterhouse / det. Barries 1995 [w, p]’ (WBWA). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / walkeri / nigriventris subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved &male; specimen with all appendages, missing only left antenna from antennomere VIII on. Length 29.50 mm, width 11.00 mm, length/width ratio: 2.68. Body weakly shiny. Head, pronotum and scutellum dark green, impunctate areas on pronotum with dark purple tint. Elytra dark viridian, apical half with purple-violet metallic reflections (Fig. 33). Ventral side dark green with purple and golden reflections (Fig. 34). Legs green with strong golden reflections, whole tarsi brown, only claws metallic. Pronotum in general moderately coarsely and densely punctate. Medial line well marked, impunctate, not elevated. Medial impressions shallow, more impressed internally than externally, coarsely and moderately densely punctate, punctures subequal to those on disc, basally denser. Principal impressions elongate and shallow, not sharply bordered, well separated from basal impressions, impunctate but rugose. Lateral impressions shallow, oval, punctate. Basal impressions large and triangular. Elytra without lateral impressions, moderately strongly punctate. Rows visible from base to apex, weakly impressed and irregularly double in apical half, internal rows more or less regularly arranged, external gradually irregular. Punctures in rows V–VIII in middle third of elytra irregular but not forming isolated groups; spaces ca. 3× as broad as puncture diameter, not elevated (Fig. 35). Intervals basally flat, not elevated, broad, approximately 4–5× as wide as rows of punctures, impunctate; apically and laterally vanishing, strongly narrowed, slightly elevated and punctate. Ventral side (Fig. 34) including legs as in nominotypical subspecies. Only lateral sides of abdominal ventrites II–V densely punctate in basal half and very sparsely punctate in apical half; punctures moderately large, subequal in size (Fig. 4). Aedeagus (Fig. 65) length 7.50 mm, width 1.63 mm, length/width ratio: 4.60. Apex of parameres rounded. Penis elongate, subparallel-sided. Variation. Body &male;&male; (n = 6) length: (21.00) 24.25–29.50 mm, width: (7.50)8.75–11.00 mm, length/width ratio: 2.63–2.80; &female;&female; (n = 74) length: (22.75)24.00–35.75(38.00) mm, width: (8.25)9.25–13.00(14.00) mm, length/width ratio: (2.43)2.54–2.85. Head, pronotum and scutellum green with blue, turquoise, purple, golden or copper sheen. Impressions on pronotum usually golden. Elytra predominantly viridian with purple-violet reflection on apical half. Purple colour sometimes extends also to basal half of elytra. Rarely elytra dark blue-green. Underside nearly always dull, only exceptionally shiny, varying from almost uniformly dark green to mostly golden-green or copper-green. Punctation of pronotum variable in density and coarseness, medial impressions variously impressed, principal impressions oval to round and variously impressed, lateral impressions variously impressed, and basal impressions oval to triangular, always deeply impressed. Elytra exceptionally with lateral impressions; larger specimens smoother while smaller specimens rougher, particularly on apical third. Aedeagus (n = 5) length: 6.17–8.06 mm, width 1.17–1.63 mm, length/width ratio: 4.60–5.04(5.45). Differential diagnosis. Among the subspecies of C. (G.) walkeri this one is easily distinguished by viridian colouration of elytra (various shades of blue in other subspecies) and rounded apices of parameres (truncate in other subspecies). For additional characters see Table 1. Etymology. The epithet is combination of Latin words niger = black and venter = underside reflecting darkly coloured ventral side of body. Distribution. Indonesia, Maluku Prov.: Moa Is. Remarks. THÉRY (1935: 248) described Gelaeus walkeri var. nigriventris, however, in the same work he described also a subspecies Gelaeus walkeri wetteriana therefore the variety nigriventris is infrasubspecific name according to the Article 45.6.4 (ICZN 1999). Nor it was validated before 1985 (Article 45.6.4.1) and hence it is an unavailable name according to ICZN (1999). We have located one female specimen in NHMB from the locality Moa mentioned by THÉRY (1935) and provided with his original paratype label as the variety nigriventris. Additionally we have found a series of specimens in MFNB with identical locality label (specimens originated in the collection of Julius Moser) agreeing also morphologically with the NHMB specimen. In our opinion this taxon is a distinct subspecies of C. (G.) walkeri and we decided to use the same name as Théry. LANDER (2003) provided a colour photograph of C. (G.) walkeri, however, the specimen does not have the last tarsomere metallic green thus it does not belong to the nominotypical subspecies. In our opinion the depicted specimen belongs to C. (G.) walkeri nigriventris subsp. nov. based on general shape.
Published: 2016
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35. Chrysodema (Gelaeus) walkeri subsp. rejzeki Frank & Sekerka 2016, subsp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Chrysodema (gelaeus) walkeri rejzeki, Arthropoda, Chrysodema, Chrysodema walkeri, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) walkeri rejzeki subsp. nov. (Figs 30–32, 64) Type locality. Indonesia, East Nusa Tenggara Province, Alor Island. Type material. HOLOTYPE: &female;, ‘Alor [w, p] // Galaeus / walkeri / (Waterh., 1892) / Dét. T. LANDER 1990 [w, h/p]’ (MHNG). PARATYPES (1 &male; 2 &female;&female;): &female;, ‘Alor / Staud. [w, h] // SYN- / TYPE [b, p] // cupriventris / Kerr. / Type [w, h] // Kerremans. / 1903—59. [w, p] // Chrysodema / erronea / HO&Lstrok;Y&Nacute;SKI / HOLOTYPE [r, h] // 1978 Chrysodema / erronea Hol. / det. R. Ho&lstrok;y&nacute;ski [w, h; unpublished manuscript name]’ (BMNH); &male;, ‘Alor [w, p] // COLLECTION / de BONNEUIL [w, p] // MUSÉUM PARIS / 1935 / Coll. A. THÉRY [w, p]’ (MHNH); &female;, ‘I. Alor / H. F. Holz. [w, p]’ (MHNH). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / walkeri / rejzeki subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved &female; specimen, missing right antenna and last tarsomeres on left fore-leg and both hind-legs. Length 30.25 mm, width 10.75 mm, length/width ratio: 2.81. Body strongly shiny. Body colouration as in the nominotypical subspecies but elytra dark blue including apex without purple reflections and with greenish apical half of suture, lateral sides, lateral impressions, and partly also punctures (Figs 30, 32). Whole tarsi brown, only claws metallic. Pronotum moderately coarsely and moderately densely punctate. Medial line well marked, impunctate. Medial impressions distinctly impressed on both sides, coarsely and densely punctate, punctures smaller than those on disc. Principal impressions oval and shallow, not sharply bordered, well separated from basal impressions, weakly separated from disc, with small dense punctures. Lateral impressions oval, shallow, punctate. Basal impressions large and oval. Elytra with conspicuous anterior and posterior lateral impressions, moderately punctate. Both impressions shallow but distinct, anterior round, larger, posterior oval and smaller (Fig. 32). Rows visible from base to apex, weakly impressed and irregularly double in apical half, internal rows more or less regularly arranged, external gradually irregular. Punctures in rows V–VIII in middle third of elytra quite densely and more or less regularly arranged, not forming isolated groups; spaces approximately 2× as broad as puncture diameter, flat not reticulate (Fig. 32). Intervals basally flat, not elevated, broad, approximately 4× as wide as rows of punctures, impunctate; apically and laterally narrowed, somewhat elevated and punctate. Ventral side (Fig. 31) including legs as in nominotypical subspecies. Only lateral sides of abdominal ventrites III–V densely punctate in basal half and very sparsely punctate in apical; punctures moderately large, subequal in size. Variation. Body &male; (n = 1) length: 26.00 mm, width: 9.5 mm, length/width ratio: 2.74, &female;&female; (n = 3) length: 30.25–31.00 mm, width: 10.75–11.25 mm, length/width ratio: 2.73–2.81. Head, pronotum, underside of body including legs from bright green to golden-green with more or less expanded copper reflections. Elytra from turquoise-green to blue or violet-blue and with only anterior lateral impression. In one &female; specimen elytra mostly blue, lateral margin green only in basal third of elytra. Aedeagus (Fig. 64) length 6.53 mm, width 1.43 mm, length/width ratio: 4.57. Apex of parameres horizonatlly truncate. Penis elongate-oval, distinctly rounded on sides. Differential diagnosis. Among the subspecies of C. (G.) walkeri this one is easily distinguished by elytra with distinct large lateral impressions and green lateral sides and suture, while the other subspecies do not have differently coloured margins of elytra or suture and the lateral impressions are rarely present and if so they are very small and shallow (basically present only as a group of several punctures). For additional characters see Table 1. Etymology. This species is dedicated to Roman Rejzek (Prague, Czech Republic), excellent collector of Buprestidae. Distribution. Indonesia, East Nusa Tenggara Prov.: Alor Is. Remarks. The paratype of C. (G.) walkeri rejzeki subsp. nov. deposited in BMNH is labelled as syntype of C. (G.) cupriventris. The latter species was described from ‘Macassar’ and KERREMANS (1898) must have had at least two specimens as he gave length span ‘Long., 21- 28; larg., 7-9 mill.’. However, this ‘syntype’ has locality Alor and measures 30.75 × 11.25 mm and is very different from the other syntype of C. (G.) cupriventris deposited in BMNH. Therefore we do not consider it as a part of the type series of the latter species. We assume that Kerremans acquired this specimen after the description of C. (G.) cupriventris and erroneously labelled it as its type. Most likely it represents a voucher specimen for the record from Alor mentioned by him (KERREMANS 1903). See additional remarks under C. (G.) cupriventris., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 713-714, DOI: 10.5281/zenodo.5310800, {"references":["KERREMANS C. 1898: Buprestides nouveaux de l'Australie et des regions voisines. Annales de la Societe Entomologique de Belgique 42: 113 - 182.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112."]}
Published: 2016
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36. Chrysodema (Gelaeus) walkeri subsp. bilyi Frank & Sekerka 2016, subsp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Chrysodema walkeri, Animalia, Chrysodema (gelaeus) walkeri bilyi, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) walkeri bilyi subsp. nov. (Figs 1–2, 36–38, 66) Type locality. Indonesia, Maluku Province, Tanimbar Islands, Selaru Island. Type material. HOLOTYPE: &male;, ‘ Selaru / Jan. 2010 [w, p]’ (NMPC). PARATYPE: 1 &female;, same data as holotype (DFPC). Both specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / walkeri / bilyi subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved&male; specimen, left antenna missing from antennomere VII, right from antennomere IV. Length 28.50 mm, width 9.75 mm, length/width ratio: 2.92. Body weakly shiny. Head, pronotum and scutellum bluish-green, elytra dark blue, apex with purple reflections (Fig. 36). Ventral side dark green with golden reflections (Fig. 37). Legs green with strong bluish luster, whole tarsi brown, only claws metallic. Pronotum in general coarsely and densely punctate. Medial line well marked, impunctate and somewhat elevated. Medial impressions distinctly impressed on both sides, coarsely and densely punctate, punctures smaller than those on disc. Principal impressions elongate and shallow, not sharply bordered, well separated from basal impressions, with small dense punctures. Lateral impressions oval, shallow, punctate. Basal impressions large and oval. Elytra without lateral impressions, moderately strongly punctate. Rows visible from base to apex, weakly impressed and irregularly double in apical half, internal rows more or less regularly arranged, external irregular. Punctures in rows V–VIII in middle third of elytra more or less regular, not forming isolated groups; spaces 2–3× as broad as puncture diameter, slightly elevated, not reticulate (Fig. 38). Intervals basally flat, not elevated, broad, approximately 5× as wide as rows of punctures, impunctate; apically and laterally strongly narrowed, slightly elevated and at most with a few additional punctures. Ventral side (Fig. 37) including legs as in nominotypical subspecies. Only lateral sides of abdominal ventrites II–V densely punctate in basal half and very sparsely punctate in apical half; punctures moderately large, subequal in size. Aedeagus (Fig. 66) length 7.60 mm, width 1.43 mm, length/width ratio: 5.32. Apex of parameres obliquely truncate. Penis elongate, parallel-sided. Variation. Body &female; (n = 1) length: 30.25 mm, width: 11.00 mm, length/width ratio: 2.75. Head, pronotum and scutellum with golden reflections in general and copper reflections in impressions. Elytra blue with slightly greenish shade. Underside with copper reflections, and abdomen with purple reflections. Pronotum with denser and more numerous punctures, particularly between medial and principal impressions. Internal rows of elytra slightly more impressed in basal 1/6 length. Differential diagnosis. Chrysodema (G.) walkeri bilyi subsp. nov. do not posseses any striking character like other subspecies but has the slimmest penis. It has dull underside of body like C. (G.) walkeri kubani subsp. nov. and C. (G.) walkeri nigriventris subsp. nov. and looks morphologically intermediate between them as it has less sculptured elytra with sparser punctation than C. (G.) walkeri kubani but more than C. (G.) walkeri nigriventris. However, C. (G.) walkeri bilyi is widely separated geographically, the only taxon of the C. (G.) walkeri species-group occurring on Tanimbar Islands while the two latter occur on Moa and nearby Romang islands. For additional characters see Table 1. Etymology. This species is dedicated to Svatopluk Bílý (Prague, Czech Republic), a wellknown specialist in Buprestidae, particularly Anthaxiini. Distribution. Indonesia: Maluku Prov.: Tanimbar Isls.: Selaru Is., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 708-709, DOI: 10.5281/zenodo.5310800
Published: 2016
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37. Chrysodema (Gelaeus) iris subsp. moluensis Novak 2010, stat. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Chrysodema iris, Arthropoda, Chrysodema, Chrysodema (gelaeus) iris moluensis novak, 2010, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) iris moluensis Novak, 2010, stat. nov. (Figs 15–17, 59) Chrysodema moluensis Novak, 2010: 107 (original description; partim). Type locality. Indonesia, Maluku Province, Tanimbar Islands, Molu Island. Type material. Not examined. Additional material examined. INDONESIA: MALUKU PROV.: Molu Island, ii.2008, 2 &female;&female;, local collectors lgt. (NMPC), iv.2009, 3 &female;&female; (DFPC); Molu Island, Tutunametal village env., xi.2008, 1 &male;, local collectors lgt. (DFPC), 1 &female; (SJPC), 0–50 m a.s.l., xi.2012, 1&male; (DFPC); Tanimbar Islands, 11.i.2008, 1&male; 2&female;&female;, R. Geweiler [lgt.?] (SGBG). Redescription. Well preserved &male; specimen with all appendages, missing only right antennomere XI. Length 25.00 mm, width 8.50 mm, length/width ratio: 2.94. Colouration as in nominotypical subspecies but whole tarsi metallic green and elytra iridescent only in apical third, basal two thirds purple and fern green (Fig. 15). Pronotum in general as in nominotypical subspecies, medial impression densely punctate along whole length, basal impression triangular and not open to principal impression, latter elongate. Elytra without clear lateral impressions, coarsely punctate. Punctures large, deeply impressed, with large fovea. Rows visible from base to apex, moderately impressed, generally regularly arranged, and barely double in apical half. Punctures in rows V–VIII in middle third of elytra quite regularly and densely arranged forming distinct rows not isolated groups, laterally somewhat less regular; spaces slightly broader than puncture diameter, flat not reticulate (Fig. 17). Intervals more or less convex from base to apex, moderately broad, approximately 3× as wide as rows of punctures, nearly impunctate, only with a few coarse punctures apically. Mid femora and ventral side as in nominotypical subspecies (Fig. 16). Aedeagus (Fig. 59) length 6.63 mm, width 1.38 mm, length/width ratio: 4.80. Apices of parameres truncate. Penis oval, distinctly rounded on sides. Variation. Body &male;&male; (n = 3) length: 25.00–29.00 mm, width: 8.50–10.00 mm, length/width ratio: 2.90–3.03; &female;&female; (n = 7) length: 27.00–33.00 mm, width: 9.25–11.75 mm, length/width ratio: 2.64–2.92. In contrast to nominotypical subspecies elytral pattern more variable, basal 2/3 from completely purple or violet, sometimes partly with golden reflections, to mostly fern green with slight purple reflections. Colouration of underside nearly constant. Medial impressions on pronotum shallow to moderately deep, variously punctate, sometimes punctures grouped together and more numerous basally. Elytra in larger specimens appear slightly less sculptured. Four specimens have a more or less indicated posterior lateral impressions on elytra as a group of small dense punctures of olive-green colour, but distinctly impressed only in one specimen. Differential diagnosis. This subspecies can be easily recognized from the nominotypical (characters of latter in brackets) by uniformly metallic tarsi (yellow with last tarsomere metallic), iridescent pattern limited to apical third of elytra (present on lateral slope from base to apex), and coarser and distinctly impressed punctation of elytra with larger punctures. Distribution. Indonesia: Maluku Prov.: Tanimbar Isls.: Molu Is. Remarks. NOVAK (2010) described Chrysodema moluensis from series of 24 specimens. Holotype, allotype and 16 paratypes were collected on Molu Island and six additional paratypes from Larat Island. Unfortunately, G. Novak (Vienna, Austria) has passed away and his collection is currently inaccessible. Nevertheless, he provided the description with photographs of both populations allowing us to recognize them with certainty. However, in our opinion the population from the Larat Island is so strikingly different that it deserves its own taxon and is here described as C. (G.) oborili laratensis subsp. nov. See further comments under the latter taxon. The redescription of C. (G.) iris moluensis presented here is based on a topotypic male specimen collected on the same date and obtained from the same batch. NOVAK (2010) did not assign his species to any of the existing subgenera of Chrysodema. In our opinion this taxon represents a subspecies of C. (G.) iris as it has deep basal impression on pronotum (absent in other subgenera of Chrysodema) and has similar but different elytral pattern to C. (G.) iris., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 698-699, DOI: 10.5281/zenodo.5310800, {"references":["NOVAK G. 2010: Chrysodema moluensis n. sp. von den Inseln Molu und Larat, Indonesien (Coleoptera: Buprestidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 62: 107 - 109."]}
Published: 2016
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38. Chrysodema (Gelaeus) oborili subsp. laratensis Frank & Sekerka 2016, subsp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema oborili, Chrysodema (gelaeus) oborili laratensis, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) oborili laratensis subsp. nov. (Figs 21–23, 61) Gelaeus Iris var. cupripennis Théry, 1935: 248 (unavailable infrasubspecific name). BELLAMY (2008):532 (catalogue, listed as synonym of C. iris). Chrysodema moluensis Novak, 2010: 107 (partim, incl. colour Figs 2–3). Type locality. Indonesia, Maluku Province, Tanimbar Islands, Larat Island. Type material. HOLOTYPE: &male;, ‘ Larat - Ins. / (Tenimber I.) [w, p]’ (NMPC). PARATYPES (7 &male;&male; 19 &female;&female;): 5 &male;&male; 15 &female;&female;, ‘Larat - Ins. / (Tenimber I.) [w, p] // Coll. / Jul. Moser [w, p]’ (5 &male;&male; 14 &female;&female; MFNB, 1 &female; DFPC); 2 &male;&male; 2 &female;&female;, ‘Indonesia, S MOLLUCAS / Tanimbar Isls, 0-50 M / LARAT ISLAND, 2.2008 / local collectors lgt. [w, p]’ (2 &male;&male; 1 &female; DFPC, 1 &female; SJPC); 1 &female;, Tanimbar, feb. 2003 [w, p]’ (DFPC); 1 &female;, Tanimbar, 03/ 2003 [w, h]’ (DFPC). All specimens provided with an additional red printed label:‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / oborili / laratensis subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved &male; specimen missing left two distal antennomeres and left hind leg. Length 27.00 mm, width 9.50 mm, length/width ratio: 2.84. Head and pronotum olive green, latter with barely visible violet reflections along anterior margin and anterior corners. Scutellum olive green. Elytra bronze centrally, gradually changing laterally and apically to copper and dark violet (Fig. 21), lateral serrations and apex dark teal (Fig. 23). Ventral side and legs as in nominotypical subspecies (Fig. 22). Pronotum as in nominotypical subspecies. Elytra as in nominotypical subspecies, only intervals distinctly more elevated (Figs 21, 23). Legs and ventral side as in nominotypical subspecies (Fig. 22). Aedeagus (Fig. 61) length 7.55 mm, width 1.53 mm, length/width ratio: 4.93. Apices of parameres obliquely truncate. Penis elongate, subparallel-sided. Variation. Body &male;&male; (n = 8) length: 26.25–28.25 mm, width: 9.00–10.00 mm, length/ width ratio: 2.79–2.86(3.03); &female;&female; (n = 19) length: (21.00)25.25–34.25(36.50) mm, width: (7.50)9.00–13.00 mm, length/width ratio: 2.67–2.92. Structure of pronotum quite variable, principal and medial impressions from shallow to moderately deep, latter sometimes slightly coarser punctate, principal impressions always appears impunctate except of inner row. Basal impressions always deep, variously open and connected with principal impression. Pronotum from uniformly olive green to partly violet; most specimens with green pronotum at most with slight violet reflections. Elytra from bronze to nearly copper, always with gradually darker lateroapical margins. Ventral side often with violet, purple or copper reflections. Aeadeagus (n = 5) length: 7.34–7.75 mm, width 1.38–1.53 mm, length/width ratio: 4.93–5.42. Differential diagnosis. Chrysodema (G.) oborili laratensis subsp nov. is on average narrower than the nominotypical subspecies (see body length/width ratios in Variation) and has much more convex elytra in lateral view (see Figs 20 vs 23). Additionally it has brighter colouration of elytra with whole central portion bronze to copper while the nominotypical subspecies is less shiny and has dark-violet body with only more or less bronze intervals. Etymology. This subspecies is named after its type locality Larat Island, the subspecies epithet being an adjective. Distribution. Indonesia, Maluku Prov.: Tanimbar Isls.: Larat Is. Remarks. THÉRY (1935: 248) described Gelaeus iris var. cupripennis, however, in the same work he described also a subspecies Gelaeus walkeri wetteriana therefore the variety cupripennis is infrasubspecific name according to the Article 45.6.4 (ICZN 1999). Nor was it validated before 1985 (Article 45.6.4.1) and hence it is an unavailable name according to ICZN (1999). THÉRY (1935: 248) separated his variety cupripennis based on uniformly copper coloured elytra while C. (G.) iris has iridescent pattern on elytra. He mentioned only ‘Tenimber’ in catalogue as the distribution of this variety. We were unable to locate any original ‘type’ specimens for this variety. However, the historical series from Larat (ex collection of J. Moser; MFNB) was loaned to us under the latter name. We presume that the var. cupripennis is the same as C. (G.) oborili laratensis subsp. nov. because most specimens have copper or bronze elytra, which are distinctly darker in C. (G.) oborili oborili and iridescent in C. (G.) iris., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 703-704, DOI: 10.5281/zenodo.5310800, {"references":["THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","NOVAK G. 2010: Chrysodema moluensis n. sp. von den Inseln Molu und Larat, Indonesien (Coleoptera: Buprestidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 62: 107 - 109."]}
Published: 2016
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39. Chrysodema (Gelaeus) sara Frank & Sekerka 2016, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema sara, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) sara sp. nov. (Figs 45–47, 68) Gelaeus walkeri walkeri [misidentification]: AKIYAMA & OHMOMO (2000): Pl. 42, Figs 445-3. Type locality. Indonesia, Maluku Province, Babar Island, Tepa. Type material. HOLOTYPE: &male;, ‘ Indonesia, S. Moluccas / BABAR ISL., 0 – 250 m alt / Tepa env., 7.2010 / local collection lgt’ (NMPC). PARATYPES (2 &female;&female;): 1 &female;, same data as holotype (SGCB); 1 &female;, ‘ Indonesia, S. Moluccas / WETANG ISL, 4.2009 / Rumalewan Besar vill env / Babar archipelago / local collectors lgt’ (DFPC). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / oborili sp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved&male; specimen with all appendages, only left two distal antennomeres missing. Length 24.00 mm, width 8.00 mm, length/width ratio: 3.00. Head purple-green. Pronotum and scutellum copper-green. Elytra bluish-black with apicolateral margins bright bluish-green and violet (Figs 45, 47). Ventral side dark violet laterally and purple-red centrally (Fig. 46). Legs copper-red with golden reflections on femora, violetgreen tibiae, tarsi brown with metallic claws. Pronotum in general coarsely but sparsely punctate, laterally punctures moderately dense. Medial line well marked impunctate. Medial impressions shallow, more impressed along medial line, moderately densely punctate, punctures slightly smaller than remaining ones on disc. Principal impression, oval, moderately deep, not connected to basal impressions, variously punctate with several moderately large punctures and line of fine punctures. Lateral impressions shallow and rugose. Basal impressions oval, moderately deep. Elytra without lateral impressions, finely punctate. Rows visible from base to apex, not impressed, basally regularly arranged, apically gradually irregular and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra more or less regular and sparse forming distinct rows, not isolated groups; spaces much broader than puncture diameter, flat not reticulate (Fig. 47). Intervals flat, not elevated, broad, approximately 4–6× as wide as rows of punctures, basally impunctate, apically gradually narrowed and partly vanishing. Mid femora in central part strongly shiny, very sparsely and finely punctate, on sides punctures slightly larger but still sparse. Thoracic ventrites (Fig. 46) centrally nearly impunctate, laterally moderately densely punctate, punctures large. Abdominal ventrites sparsely punctate, only ventrite I laterally moderately densely punctate. Aedeagus (Fig. 68) length 6.12 mm, width 1.53 mm, length/width ratio: 4.00. Apices of parameres obliquely truncate. Penis stout-oval and short in contrast to other taxa. Variation. Body &female;&female; (n = 2): length: 26.75–34.00 mm, width: 9.50–12.75 mm, length/width ratio: 2.67–2.82.All three specimens quite similar, not showing much variation. Female from Babar Island appears more sparsely punctate due to larger size of its body. Differential diagnosis. Readily recognizable species due to its smooth elytra with fine punctation. Chrysodema (G.) sara sp. nov. has the finest punctation of the elytra of all Gelaeus taxa. In generall it is most similar to C. (G.) walkeri walkeri, which has quite smooth elytra in comparison to remaining taxa but still has distinctly impressed rows of punctures and slightly elevated intervals posteriorly while C. (G.) sara does not have impressed rows or elevated intervals. Additionally, C. (G.) sara has the shortest and stoutest penis of all Gelaeus species and is practically the only with a distinctly broadly oval penis. Etymology. This species is named after Sára Franková, daughter of the first author of this paper. The species epithet is a noun in apposition. Distribution. Indonesia: Maluku Prov.: Babar Is. and Wetang Is. Remarks. Figure 445-3 published by AKIYAMA & OHMOMO (2000) as C. (G.) walkeri walkeri is in fact C. (G.) sara sp. nov.
Published: 2016
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40. Chrysodema (Gelaeus) iris subsp. iris iris (Kerremans 1898

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Chrysodema (gelaeus) iris iris (kerremans, 1898), Insecta, Chrysodema iris, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) iris iris (Kerremans, 1898) (Figs 12–14, 58) Pseudochrysodema Iris Kerremans, 1898: 114 (original description). Chrysodema Iris: KERREMANS (1903): 76 (catalogue). Chrysodema (Gelaeus) Iris: KERREMANS (1909): 509 (key, redescription, incl. colour Fig. 4, Pl. 22); BELLAMY (2008): 532 (catalogue). Gelaeus Iris: OBENBERGER (1926): 129 (catalogue); THÉRY (1935): 248 (key, catalogue). Type locality. Original type locality ‘Banda’ here changed to: Indonesia, Maluku Province, Tanimbar Islands. Type material examined. LECTOTYPE (present designation): &male;, ‘Banda Ins. [w, h] // Iris / Kerr / Type [w, h] // iris Kerr. [w, h] // Coll. / Jul. Moser // Chrysodema (Gelaeus) / iris / Kerr. / Det. T. LANDER 2000’ (MFNB). Specimen provided with an additional red printed label: ‘ LECTOTYPE &male; / Chrysodema (Gelaeus) / iris / (KERREMANS, 1898) / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Additional material examined. INDONESIA: MALUKU PROV.: Banda Island [probably erroneous locality data], 1 &female; (NMPC); Tanimber [= Tanimbar Islands], 1 &male; 2 &female;&female; (RMNH, ex coll. ZMAN); Tenimber [= Tanimbar Islands], 1 &female; (NHMB, coll. G. Frey), 1 &female; (DFPC), 1 &female; (RMNH), 1 &female; (MHNG); Key [= Kai Islands; probably erroneous locality data], 1 &female;, J. M. Bedoc coll. (MHNG). Redescription of lectotype. Quite well preserved &male; specimen missing both antennae, left fore and hind, and right middle tarsi. Length 28.00 mm, width 9.50 mm, length/width ratio: 2.95. Head, pronotum, ventral side and legs except tarsi fern green with purple reflections, the latter particularly obvious on impunctate parts of pronotum and abdomen (Fig. 13). Tarsi yellow, last tarsomere metallic green. Elytra iridescent, purple on top of disc and basally, changing laterally and apically to fern green, then golden, red, violet, blue and finally turquoise (Fig. 12). Pronotum in general moderately strongly and sparsely punctate, only laterally punctures more densely arranged. Medial line well marked, impunctate, appearing slightly elevated. Medial impressions distinctly impressed, moderately deep, sparsely punctate, punctures in basal 2/3 smaller and arranged in an irregular row and partly grouped together, in anterior 1/3 impressions shallower with punctures almost equal in size to remaining punctures along anterior margin. Principal impressions large, round, bordered internally by row of very fine punctures, centrally and externally smooth with only a few larger punctures. Basal impressions large, oval, broadly open and connected with principal impression. Lateral impressions deep and impunctate. Elytra without lateral impressions, moderately strongly punctate. Punctures small, shallowly impressed basally, and moderately impressed laterally. Rows visible from base to apex, weakly impressed, generally regularly arranged, and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra quite regularly and sparsely arranged, forming distinct rows not isolated groups, laterally somewhat less regular; spaces much broader than puncture diameter, flat not reticulate (Fig. 14). Intervals basally flat, not elevated, broad, approximately 5–6× as wide as rows of punctures, impunctate; apically and laterally barely elevated and nearly impunctate. Mid femora in central part strongly shiny, sparsely punctate, punctures large, on sides punctures smaller and dense. Ventral side of body (Fig. 13) overall coarsely and quite densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Almost whole lateral side of abdominal ventrite I densely and coarsely punctate, punctures small. Lateral sides of ventrites II–V densely and coarsely punctate basally and sparsely punctate with large punctures apically. Aedeagus (Fig. 58) length 7.55 mm, width 1.53 mm, length/width ratio: 4.93. Apices of parameres truncate. Penis elongate, subparallel-sided. Variation. Body &female;&female; (n = 6): length: 26.00–31.00 mm, width: 9.00–11.00 mm, length/width ratio: 2.82–2.95. In contrast to the other taxa this one seems to be rather constant in colouration. Only one specimen has more extensive purple colour on the disc of the elytra and the fern green colour is limited to a thin line. Differential diagnosis. This species can be easily recognized from other species of Gelaeus by iridescent colouration of elytra. The subspecies C. (G.) iris moluensis differs in whole tarsi metallic coloured, much coarser punctation of elytra and iridescent pattern restricted to apical half of elytra while the nominotypical subspecies also has iridescent lateral sides. Distribution. Indonesia: Maluku Prov.: Tanimbar Isls (see Remarks). Remarks. KERREMANS (1898) did not mention explicitely how many specimens he had at his disposal just gave a single length and width measurement: ‘Long., 28; larg. 9 mill.’ and stated that the material is from collection of Meyer Darcis. The latter collection was purchased by Julius Moser and is nowadays deposited in MFNB. We have found a single specimen in the latter collection, however since Kerremans did not mention the number of specimens it is not certain that this is the only existing specimen used in making the description. Therefore we designate the MFNB specimen a lectotype in order to conserve status of this species and avoid any further misinterpretations. KERREMANS (1898) gave ‘Banda’ as the type locality. There is only one other known specimen with the same locality data on handwritten label by J. Obenberger (deposited in NMPC). There is also one specimen with the label Kai Islands. The remaining seven studied specimens were collected on Tanimbar Islands, however, without more precise locality data. Nevertheless we assume that the type locality Banda Island as well as the record from Kai Island are erroneous and the species is restricted to Tanimbar Islands because all specimens are quite similar each to other, not displaying much variation and on the Tanimbar Islands also occurs C. (G.) iris moluensis the other subspecies of this taxon. Therefore we change the type locality to Tanimbar Islands. More accurately localized material is needed to confirm and clarify the distribution of this species., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 697-698, DOI: 10.5281/zenodo.5310800, {"references":["KERREMANS C. 1898: Buprestides nouveaux de l'Australie et des regions voisines. Annales de la Societe Entomologique de Belgique 42: 113 - 182.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","OBENBERGER J. 1926: Buprestidae I. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, 212 pp.","THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256."]}
Published: 2016
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41. Chrysodema (Gelaeus) oborili subsp. oborili oborili 2016, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema oborili, Biodiversity, Chrysodema (gelaeus) oborili oborili, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) oborili oborili sp. nov. (Figs 18–20, 60) Type locality. Indonesia, Maluku Province, Tanimbar Islands, Yamdena Island, 20 km NE of Saumlaki. Type material. HOLOTYPE: &male;, ‘ INDONESIA. Tanimbar is. / YAMDENA ISL. 20km NE / of Saumlaki. 150m / 1.- 30.i.2007 M. Obo&rcaron;il lgt. [w, p]’ (NMPC). PARATYPES (29 &male;&male; 17 &female;&female;): 29 &male;&male; 16 &female;&female;, same data as holotype (19 &male;&male; 10 &female;&female; MOOC, 7 &male;&male; 4 &female;&female; DFPC, 1 &male; HMMG, 1 &male; 1 &female; MFNB, 1 &male; 1 &female; MHNG); 1 &female;, ‘Tanimbar Is., Yamdena / Mams village / 21km N of Saumlaki / 27.11.- 11.12.2005 / J. Horák leg. [w, p]’ (NMPC). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / oborili / oborili subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved&male; specimen with all appendages. Length 28.25 mm, width 9.00 mm, length/width ratio: 3.14. Head and pronotum olive green with violet reflections, particularly on impunctate parts. Scutellum olive green. Elytra dark metallic brown-violet, intervals more or less bronze (Fig. 18), latero-apical slope dark violet, lateral serrations and apex dark teal (Fig. 20). Thoracic ventrites olive green with violet reflections, abdominal ventrites largely dark violet, only basally more or less olive green with some golden reflections (Fig. 19). Legs olive green with violet reflections, tarsi brown, last tarsomere metallic olive green. Pronotum in general moderately strongly and sparsely punctate, only laterally punctures denser. Medial line well marked impunctate, appears slightly elevated. Medial impression sdistinctly impressed, moderately deep, irregularly and moderately densely punctate, punctures somewhat smaller than external ones. Principal impressions moderately deep, elongate, bordered internally by row of very fine punctures, centrally and externally smooth with only a few larger punctures. Basal impressions large, oval, broadly open and connected with principal impressions. Lateral impressions deep and impunctate. Elytra without lateral impressions, moderately strongly punctate. Punctures moderately strong and moderately impressed. Rows visible from base to apex, weakly impressed, generally regularly arranged, internal ones irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra quite regular and moderately dense, forming distinct rows not isolated groups, laterally slightly less regular; spaces approximately as wide as puncture diameter, flat not reticulate (Fig. 20). Intervals basally flat, barely elevated, narrow, approximately 1–2× as wide as rows of punctures, with several punctures apically. Mid femora in central part strongly shiny, very sparsely punctate, punctures large, on sides punctures smaller and dense. Ventral side of body (Fig. 19) overall coarsely and quite densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Lateral sides of abdominal ventrites laterobasally with patch of dense small punctures, rest sparsely punctate with larger punctures. Aedeagus (Fig. 60) length 7.40 mm, width 1.43 mm, length/width ratio: 5.17. Apices of parameres obliquely truncate. Penis elongate, subparallel-sided. Variation. Body&male;&male; (n = 30) length: 24.75–29.75(32.00) mm, width: 7.75–10.25 mm, length/ width ratio: (2.94)2.97–3.22; &female;&female; (n = 16) length: (24.00)27.00–34.00(36.00) mm, width: (8.00)9.25–11.75(12.25) mm, length/width ratio: (2.73)2.80–3.00. Structure of pronotum quite variable, principal and medial impressions from shallow to moderately deep, latter sometimes slightly more coarsely punctate, principal impressions always appear impunctate except for inner row. Basal impressions always deep variously open and connected with principal impressions. Pronotum from nearly uniformly olive green to mostly violet but always impressions and punctures olive green; most specimens with extensive violet reflections. Intervals on elytra sometimes not differently coloured, mostly dark bronze or olive green-bronze. Elytra sometimes with very small apparent lateral impressions due to grouped small punctures. Sometimes also thoracic ventrites predominantly violet. Aedeagus (n = 5) length: 7.04–8.36 mm, width 1.38–1.53 mm, length/width ratio: 5.10–5.46. Differential diagnosis. This species can be easily separated from other taxa of Gelaeus by brown or bronze violet colouration of elytra while remaining taxa are green, blue or black. It is similar only to C. (G.) iris, which however differs in the distinct iridescent pattern of the elytra. For distinguishing characters from C. (G.) oborili laratensis subsp. nov. see Differential diagnosis under the latter subspecies. Etymology. This species is named after Martin Obo&rcaron;il (Olbramovice, Czech Republic), a specialist in Buprestidae and collector of most of the type series. Distribution. Indonesia: Maluku Prov.: Tanimbar Isls.: Yamdena Is., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 702-703, DOI: 10.5281/zenodo.5310800
Published: 2016
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42. Chrysodema (Gelaeus) foraminifera Lansberge 1879

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Chrysodema foraminifera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) foraminifera Lansberge, 1879 (Figs 9–11, 57) Chrysodema foraminifera Lansberge, 1879: cxlviii (original description). LANSBERGE (1880): cxxxii (comparative note); KERREMANS (1903): 76 (catalogue); OBENBERGER (1926): 131 (catalogue). Chalcophora foraminifera: KERREMANS (1885): 126 (catalogue). Chrysodema (Chrysodema) foraminifera: KERREMANS (1909): 514 (key), 527 (redescription). Gelaeus foraminifera: THÉRY (1935): 247 (key), 248 (catalogue). Chrysodema (Gelaeus) foraminifera: LANDER (2003): 70 (noted), Figs 150–151 (colour photo); BELLAMY (2008): 531 (catalogue). Type locality. Indonesia, East Nusa Tenggara Province, Flores Island (see Remarks). Type material examined. LECTOTYPE (present designation): &female;, ‘v Lansb. / Flores [w, h] // Chrysodema / Foraminifera / Flores Lansb [w, h] // Type [r, p]’ (RMNH). Specimen provided with an additional red printed label: ‘ LECTOTYPE &female; / Chrysodema (Gelaeus) / foraminifera / LANSBERGE, 1879 / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Additional material examined. INDONESIA: EAST NUSA TENGGARA PROV.: Timor Island, 1 &male; 1 &female;, Macklot lgt. (RMNH), 1&male; 2&female;&female; (MNHN), 3&female;&female; (MHNG), 1&female; (DFPC), viii. 1935, 1 &female; (MHNG); Timor Island, Buraen env., 60 km SE of Kupang, 350 m, 10.–21.ii.2006, 2 &female;&female;, St. Jákl lgt. (DFPC); Semau Island? (see Remarks), 1 &female; (RMNH). Redescription of lectotype. Well preserved &female; specimen missing only mid last tarsomeres with claws and slightly open elytra with exposed wings. Length 25.25 mm, width 9.50 mm, length/width ratio: 2.66. Body generally metallic green. Pronotum slightly lighter green and somewhat shinier than elytra, basal and principal impressions golden with slight traces of copper. Elytra including elevated ribs uniformly dark green, lateral impressions slightly paler and with slight copper reflections (Fig. 9). Ventral parts bright metallic green with golden reflections (Fig. 10). Legs bright metallic green, tarsi brown, last tarsomere including claws metallic green. Pronotum laterally densely and coarsely punctate, centrally sparsely punctate. Punctures on lateral sides foveolate, more deeply impressed. Medial line well defined, impunctate but not elevated. Medial impressions hardly impressed, basically on same level as medial line and visible as stripes of moderately coarse punctures, more densely arranged in basal half. Principal impressions round, moderately deep, more or less distinctly separated from disc, densely punctate. Surface between medial and principal impressions with sparsely dispersed punctures basally with larger impunctate areas. Lateral impressions moderate with several large punctures. Basal impressions moderate, oval. Elytra distinctly punctate-striate, puncture rows posteriorly irregularly double or triple. Intervals basally slightly elevated, ca. 4× wider than puncture rows, impunctate; apically partly obscured due to punctation. Punctation of elytra overall moderate and dense (except on intervals), distance between punctures mostly larger than diameter of the punctures. Puncture rows V–VIII in mid third of elytra arranged in small groups of 2–4 separated by impunctate spaces not forming elevated reticulations. Each elytron with two oval lateral impressions situated between weakly visible 6th and 8th intervals in basal third and two thirds respectively (Fig. 11). Mid femora in central part strongly shiny, sparsely punctate, punctures large; on sides with densely arranged small punctures. Ventral side of body (Fig. 10) overall coarsely and very densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Whole lateral side of abdominal ventrite I densely and coarsely punctate. Ventrites II–V gradually less punctate apically with small and dense punctation restricted to basal half. Variation. Body &male; (n = 2) length: 25.00– 26.25 mm, width: 9.00– 9.50 mm, length/width ratio: 2.76–2.78; &female;&female; (n = 12): length: (23.00)25.00– 31.75 mm, width: (8.50) 9.25–12.25 mm, length/width ratio: 2.55–2.77. Body colouration always basically green, in some specimens with strong copper or turquoise sheen on most of the body. Medial pronotal impressions in some specimens shallowly impressed, principal impressions from round to oval and variably deeply impressed, lateral impressions punctate to impunctate, basal impression oval to triangular. Lateral impressions on elytra present in all examined specimens and of rather constant size. Two specimens have posterior lateral impressions enlarged, occupying width between 4th and 8th interval, round and more or less divided into two impressions. One specimen has similarly enlarged posterior lateral impressions and additional oval impressions internally next to it and two small round impressions in centre of elytron in basal third length. Tarsi laterally with metallic spots in four specimens. Aedeagus (Fig. 57) (n = 2) length: 6.68–6.94 mm, width 1.43 mm, length/width ratio: 4.67–4.85. Apices of parameres rounded. Penis elongate, subparallel-sided. Differential diagnosis. This species can be separated from similar C. (G.) florensis by pale colouration of tarsi (all tarsomeres green in the latter species) and by lateral portion of abdominal ventrites II–III variously punctate with large impunctate areas (Fig. 4) while C. (G.) foraminifera has lateral portions of respective ventrites densely and uniformly punctate without impunctate areas (Fig. 5). The curious uniformly green specimen of C. (G.) cyanicollis differs by the green tarsi and coarser sculpture of elytra. Some specimens of C. (G.) foraminifera with apparently differently coloured pronotum and elytra are quite similar to C. (G.) cupriventris, which differs in the elytra always lacking lateral impressions (with two impressions in C. (G.) foraminifera), red underside (predominantly green in C. (G.) foraminifera) and in lateral portions of abdominal ventrites II–IV with band of denser but yet sparse punctures along anterior margin while the rest is nearly impunctate (very densely punctate basally and sparsely apically in C. (G.) foraminifera). Distribution. Indonesia: East Nusa Tenggara Prov.: Flores Is., Timor Is. and Semau Is. The species possibly also occurs in East Timor. Remarks. LANSBERGE (1879) did not state how many specimens he had at his disposal when describing Chrysodema foraminifera nor mentioned any depository. However, he gave a range of lengths: ‘Long. 25–30 millim.’ thus he must have had at least two specimens. The collection of Lansberge was purchased by René Oberthür (nowadays in MNHN) and some duplicate specimens are also deposited in IRSN, collection of J. Neervoort van de Poll (purchased by A. Théry and nowadays in MNHN), and Museo Civico di Storia Naturale Giacomo Doria, Genova (HORN & KAHLE 1935, 1936). We have searched above mentioned museums as well as other collections and found only a single specimen labelled as type in RMNH. The specimen came from Lansbergeʼs collection, bears handwritten label, but not by Lansberge (H. Huijbregts, pers. comm.), with name and additional small red label ‘Type’. The specimen agrees with the primary description and thus we assume it is a part of the type series. Since there is possibility of existence of additional type specimens we designate the RMNH specimen the lectotype in order to fix the identity of this species. The lectotype is the only specimen of this species known to us from Flores. We have examined additional eleven specimens morphologically identical to the lectotype but these were collected on Timor. This is quite strange as collectors collected extensively in the past on both, Flores and Timor, but there is no other specimen of C. (G.) foraminifera from Flores. Therefore we assume that the type locality might be erroneous and the species is in fact restricted to Timor but this need to be verified by recent material from Flores. One specimen bears locality data as ‘Macklot / Poeloe / Samoe’, however, we think it is very unlikely that the specimen was collected in Samoa as H. C. Macklot never visited Samoa but collected in New Guinea, Timor and surrounding islands (EFFERT 2008: 132). Poeloe is an old transcription of pulau which means island(s). Samoe in our opinion is most likely misspelled Semau Island situated at southwest end of Timor Island. Since we do not have actual proof whether the specimen was collected on the latter island or not we list Semau Island with a question mark and the record has to be verified by additional specimens. LANDER (2003: 70) briefly mentioned C. (G.) foraminifera and noted that it appears to be a variety of C. (G.) walkeri with white spots but did not propose any synonymy., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 694-696, DOI: 10.5281/zenodo.5310800, {"references":["LANSBERGE J. W. VAN 1879: Diagnoses de quelques especes nouvelles de Buprestides et de Scarabaeides de la Malaisie. Comptes-Rendus des Seances de la Societe Entomologique de Belgique 22: cxlvii - clv.","LANSBERGE J. W. VAN 1880:. Description de quelques Coleopteres de la Malaisie et de la Papouasie. Comptes-Rendus des Seances de la Societe Entomologique Belgique 23: cxviii - cxxxix.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","OBENBERGER J. 1926: Buprestidae I. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, 212 pp.","KERREMANS C. 1885: Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique 29: 119 - 157.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","LANDER T. 2003: Revision du genre Chrysodema. Collection Systematique, Vol. 8, Magellanes, Andresy, 94 pp.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","HORN W. & KAHLE I. 1935: Uber entomologische Sammlungen, Entomologen & Entomo-Museologie (Ein Beitrag zur Geschichte der Entomologie). Teil I. Entomologische Beihefte (Dahlem) 2: i - vi + 1 - 160 + 1 - 16 pls.","HORN W. & KAHLE I. 1936: Uber entomologische Sammlungen, Entomologen & Entomo-Museologie (Ein Beitrag zur Geschichte der Entomologie). Teil II. Entomologische Beihefte (Dahlem) 3: 160 - 296 + pls. 17 - 26.","EFFERT R. 2008: Royal cabinets and auxiliary branches. Origins of the National Museum of Ethnology 1816 - 1883. Mededelingen van het Rijksmuseum voor Volkenkunde 37: i - viii + 1 - 345."]}
Published: 2016
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43. Chrysodema (Gelaeus) cupriventris Wetar Is., DFPC

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Chrysodema cupriventris, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) cupriventris (Kerremans, 1898), stat. restit. (Figs 3, 48–50, 69) Pseudochrysodema cupriventris Kerremans, 1898: 114 (original description). Chrysodema cupriventris: KERREMANS (1903): 74 (catalogue); WATERHOUSE (1905): 584 (as syn. of Gelaeus walkeri); KERREMANS (1909): 510 (catalogue, in synonymy of C. (G.) walkeri); OBENBERGER (1926): 29 (catalogue, in synonymy of G. walkeri); BELLAMY (2008): 542 (listed as synonym of C. (G.) walkeri wetteriana). Gelaeus walkeri var. cupriventris: THÉRY (1935): 248 (key, catalogue). Gelaeus walkeri ssp. cupriventris: AKIYAMA & OHMOMO (2000): Pl. 42, Figs 445-4 [sic!] (iconography). Type locality. ‘Macassar’ (see Remarks). Type material examined. SYNTYPES (2 &female;&female;): &female;, ‘ Macassar / Meyer D. [w, h] // cupriventris / Kerr. / Type [w, h] // SYN- / TYPE [b, p] // Kerremans / 1903—59. [w, p]’ (BMNH); &female;, ‘Macassar / Meyer D. [w, h] // cupriventris / Kerr. / Type [w, h] // MUSEUM PARIS / COLL CH. KERREMANS / 1923 [y, p] // TYPE [r, p] // C. cuprive- / ntris Kerr. / Macassar Type [w, h]’ (MNHN). Both specimens were provided with an additional red printed label: ‘ SYNTYPE &female; / Chrysodema (Gelaeus) / cupriventris / (KERREMANS, 1898) / David Frank & / Lukáš Sekerka labeled 2016’. Additional material examined. INDONESIA: MALUKU PROV.: Wetter Ins. [= Wetar Island], 5 &female;&female; (MFNB), 1 &female; (DFPC), 3 &female;&female;, C. Schädler [lgt.?] (2 RMNH, 1 MNHN; two specimens labelled as types of unpublished manuscript name ‘ C. wetteriana var. igniventris Théry’); Wetar Island, iii.2002, 1 &female; (SGBG), v.2004, 1 &male; (DFPC), iii.2006, 1 &female; (DFPC). Redescription of syntype (BMNH). Unusually small &female; specimen, missing left antenna and with partly cracked elytra and somewhat distorted left elytron in apical third. Length 21.50 mm, width 7.75 mm, length/width ratio: 2.77. Head, pronotum and scutellum bright green with golden reflections. Elytra fern green with blue lateroapical margins without any reflections (Fig. 48). Ventral side bright green with strong copper and golden reflections. Abdominal ventrite I mostly green while II–V mostly copper (Fig 49). Legs bright green partly with golden reflections, tarsi yellow, last tarsomere and claws metallic green. Pronotum in general coarsely and moderately densely punctate. Medial line well marked, impunctate. Medial impressions shallow, distinctly impressed along medial line, densely punctate, punctures anteriorly coarse and large, basally smaller than those lateral to them. Principal impressions moderately deep, elongate, densely punctate with small punctures, not connected with basal impression. Lateral impressions deep and rugose. Basal impressions deep, oval. Elytra moderately punctate, without lateral impressions. Puncture rows visible from base to apex, barely impressed on apical 1/4 length, more or less regularly arranged, and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra quite irregularly and sparsely arranged, not grouped, internally forming more or less distinct rows with intervals 1–3× as broad as diameter of punctures, flat not reticulate (Fig. 50). Intervals basally flat, not elevated, broad, approximately 3–4× as wide as rows of punctures, impunctate; apically barely elevated and distinctly punctate. Mid femora in central part strongly shiny, sparsely punctate, punctures large, on sides punctures smaller but coarser and moderately densely arranged. Ventral side of body (Fig. 49) moderately punctate, abdominal ventrites and central parts of thoracic ventrites sparsely and finely punctate. Lateral side of abdominal ventrite I moderately densely punctate; ventrites II–V with narrow irregular line of dense punctures along basal margin otherwise very sparsely punctate (Fig. 3). Variation. Body &male; (n = 1) length: 22.50 mm, width: 8.00 mm, length/width ratio: 2.81; &female;&female; (n = 13) length: (21.50)24.25–30.00 mm, width: (7.75)9.00–11.00 mm, length/width ratio: 2.62–2.77(2.97). Punctation of pronotum variable. Medial impression shallow to moderately impressed. Principal impression variously impressed but always elongate. Pronotum from mostly green to mostly golden or copper. Colouration of elytra quite constant only one specimen with golden tint. Ventral side always extensively copper or copper-red on abdomen, thoracic ventrites centrally green and laterally variously copper; some specimens with nearly whole underside copper-red. All specimens except syntypes with uniformly brown tarsi. Aedeagus (Fig. 69) (n = 1) length: 6.12 mm, width 1.28 mm, length/width ratio: 4.78. Apices of parameres obliquely truncate. Penis wedge-shaped, narrowed basally. Differential diagnosis. This species can be easily distinguished by very sparsely and finely punctate abdominal ventrites II–V (Fig. 3). All other taxa have coarser and denser punctation (Figs 4–5). Chrysodema (G.) katka sp. nov. is dorsally most similar, see its diagnosis for additional characters. Distribution. Indonesia: Maluku Prov.: Wetar Is. Remarks. KERREMANS (1898) gave ‘Macassar’ as the type locality of C. (G.) cupriventris and did not mention how many specimens he had at his disposal, however, he gave a range of lengths: ‘Long., 21-28; larg., 7-9 mill.’ thus he must have had at least two specimens. Later on, KERREMANS (1903) mentioned ‘Iles Alor et Dammer’ as distribution of C. (G.) cupriventris. We have located two specimens labelled as types with Kerremans original label in BMNH. One has the locality Alor, measures 30.75 × 11.25 mm and does not agree at all with the primary description therefore we exclude it from the type series of C. (G.) cupriventris. The specimen is here included as paratype of C. (G.) walkeri rejzeki subsp. nov., see remarks under that taxon. We believe that it is a voucher specimen for the record published by KERREMANS (1903). The other specimen is curiously small female with somewhat distorted elytra but agrees well with the primary description as well as with the type locality. We found one more specimen from Macassar labelled as type of C. (G.) cupriventris in the collection of Kerremans (MNHN). The specimen perfectly agrees with the primary description therefore we consider the two specimens from Macassar syntypes. The type locality given by Kerremans is peculiar because Macassar is usually understood to be the city in southern Sulawesi. However, in our opinion it is very unlikely that the species lives on Sulwesi as no other species of Gelaeus is know from that island. There is also Pante Macassar in the Oecusse exclave of East Timor in the Indonesian (West) Timor which is a more likely explanation of the type locality. However, as we have not seen any other specimen of C. (G.) cupriventris from Timor mainland we think that the type locality is erroneous. We have examined a series of specimens from the Wetar Island, which perfectly matches syntypes, with exception of uniformly brown tarsi (last tarsomere metallic in syntypes). On the other hand, C. (G.) cupriventris is morphologically most similar to C. (G.) katka sp. nov., which has about half of specimens with metallic tarsi and half with brown tarsi. Thererefore we assume that this character is subject to intraspecific variation in these two taxa. Chrysodema (G.) katka is endemic to Timor and all specimens have densely punctate lateral sides of abdominal ventrites II–V while all specimens of C. (G.) cupriventris have them very sparsely and finely punctate. This character is constant in the series of specimens from Wetar and thus we think this species is maybe restricted to the Wetar Island and was given the locality Macassar because the original type specimens were received through the relatively nearby Pante Macassar, which served as an important port., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 679-680, DOI: 10.5281/zenodo.5310800, {"references":["KERREMANS C. 1898: Buprestides nouveaux de l'Australie et des regions voisines. Annales de la Societe Entomologique de Belgique 42: 113 - 182.","KERREMANS C. 1903: Coleoptera Serricornia. Fam. Buprestidae. In: WYTSMAN P. (ed.). Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","WATERHOUSE C. O. 1905: Observations on Coleoptera of the family Buprestidae, with descriptions of new Species. Annals and Magazine of Natural History, Series 7 15: 577 - 584.","KERREMANS C. 1909: Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars) [livraisons 13 - 19]. J. Janssens, Bruxelles, pp. 385 - 604 + pls. 17 - 22.","OBENBERGER J. 1926: Buprestidae I. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, 212 pp.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","AKIYAMA K. & OHMOMO S. 2000: The buprestid beetles of the world. In: FUJITA H. (ed.): Mushi-Sha's Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, 341 pp. + 30 pls."]}
Published: 2016
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44. Chrysodema (Gelaeus) katka Frank & Sekerka 2016, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Chrysodema katka, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) katka sp. nov. (Figs 51–53, 70) Type locality. Indonesia, East Nusa Tenggara Province, Timor Island, Soe. Type material. HOLOTYPE: &male;, ‘ Soë (alt. 880m) / Timor central [w, p] // COLL ON LE MOULT / Naturaliste, Paris [w, p]’ (IRSN). PARATYPES (72 &male;&male; 47&female;&female;): 1&male; 6&female;&female;, same data as holotype (1&male; 3&female;&female; IRSN, 2&female;&female; DFPC, 1&female; NMPC); 1 &male; 1 &female;, same data as holotype plus additional labels ‘R. Mus. Hist. Nat. / Belg. I. G. 12.595 [w, p] // J. De Walsche det.,1943: / Gelaeus / Walkeri Wat. [w, p/h]’ (IRSN); 18 &male;&male; 2 &female;&female;, same data as holotype plus additional labels ‘R. Mus. Hist. Nat. / Belg. I. G. 12.595 [w, p] // J. De Walsche det.,1943: / GELAEUS / Walkeri(Wath.) [w, p]’ (16 &male;&male; 2 &female;&female; IRSN, 2 &male;&male; DFPC); 9 &male;&male; 6 &female;&female;, ‘Soë-(880m.) / Timor central [w, p] // COLL ON LE MOULT / Naturaliste, Paris [w, p]’ (7 &male;&male; 2 &female;&female; IRSN, 1 &female; MHNG, 1 &female; LSPC, 1 &male; NMPC, 1 &male; 2 &female;&female; DFPC); 2 &male;&male; 1 &female;, ‘Soë-(880m.) / Timor central [w, p] // COLL ON LE MOULT / Naturaliste, Paris [w, p] // R. Mus. Hist. Nat. / Belg. I. G. 12.595 [w, p] // J. De Walsche det.,1943: / Gelaeus / Walkeri Wat. [w, p/h]’ (IRSN); 38 &male;&male; 14 &female;&female;, ‘Soë-(880m.) / Timor central [w, p] // COLL ON LE MOULT / Naturaliste, Paris [w, p] // R. Mus. Hist. Nat. / Belg. I. G. 12.595 [w, p] // J. De Walsche det.,1943: / GELAEUS / Walkeri(Wath.) [w, p]’ (32 &male;&male; 11 &female;&female; IRSN, 1 &male; MFNB, 1 &male; LSPC, 4 &male;&male; 3 &female;&female; DFPC); 1 &male; 1 &female;, ‘Soë (alt. 880m) / Timor central [w, p] // COLL ON LE MOULT / Naturaliste, Paris [w, p] // MUSÉUM PARIS / 1935 / Coll.A. THÉRY’ (MNHN); 2 &male;&male; 10 &female;&female;, ‘ INDONESIEN / Timor / Camplong NTT / XII. 1996 [w, h]’ (1 &male; 7 &female;&female; HMCM, 1 &male; 1 &female; MHNG, 2 &female;&female; DFPC); 1 &female;, ‘ INDONESIA, / TIMOR Isl., / Feb. 2005 [w, h]’ (VKSC); 1 &female;, ‘ Indonesia, Lesser Sundas / TIMOR isl. 5.2005 / local collectors lgt [w, p] // CHRYSODEMA (Gelaeus) / walkeri Waterhouse 1892 / det Barries 2008 [w, p]’ (WBWA); 1 &female;, ‘ INDONESIEN / WESTTIMOR / Mt. Mutis / 5/2005 [w, p/h] // Chrysodema / (Gelaeus) / walkeri (Waterh.) / det. S. Gottwald 2007 [w, p]’ (SGBG); 3 &female;&female;, ‘Timor / Feb. 05 [w, p]’ (2 &female;&female; DFPC, 1 &female; MFNB). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / katka sp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Quite well preserved &male; specimen missing mouthparts and left antenna from antennomere V on. Length 23.00 mm, width 8.50 mm, length/width ratio: 2.71. Head, pronotum and scutellum bright green with strong golden and copper reflections, impunctate areas of pronotum with violet reflections. Elytra pine green with slight purple reflection (Fig. 51), blue lateroapical margins and suture, and somewhat paler green lateral punctation with golden reflections (Fig. 53). Ventral side bright green with strong copper and golden reflections (Fig. 52). Legs bright green partly with golden and bluish reflections, tarsi yellow, only apical half of last tarsomere and claws metallic green. Pronotum in general coarsely and moderately densely punctate. Medial line well marked, with a few moderately large punctures. Medial impressions very weak, more or less impressed only along medial line, densely punctate, punctures smaller than those lateral to them. Principal impressions deep, round, not connected to basal impressions, and with small dense punctures. Lateral impressions shallow and punctate. Basal impressions moderately deep, subtriangular. Elytra moderately strongly and quite densely punctate, with slightly marked lateral impressions as round groups of coarser and paler coloured punctures (Fig. 53). Rows visible from base to apex, distinctly impressed on apical 2/3, more or less regularly arranged, but irregularly double or triple in apical 2/3 length. Punctures in rows V–VIII in middle third of elytra quite irregular and dense, and partly grouped, only internally forming more or less distinct rows; spaces 1–2× as broad as diameter of puncture, flat, not reticulate (Fig. 53). Intervals basally flat, not elevated, broad, approximately 3–4× as wide as rows of punctures, impunctate; apically and laterally somewhat elevated and punctate, first interval somewhat vanishing apically. Mid femora in central part strongly shiny, sparsely punctate, punctures large, on sides punctures smaller but coarser and dense. Ventral side of body (Fig. 52) overall coarsely and quite densely punctate, only central parts of thoracic and abdominal ventrites sparsely punctate. Lateral sides of abdominal ventrites densely and coarsely punctate with small punctures, latter on ventrites II to V gradually sparser in apical half and replaced with sparse larger punctures. Lateral side of ventrite V densely punctate only in basal third. Aedeagus (Fig. 70) length 5.95 mm, width 1.22 mm, length/width ratio: 4.88. Apices of parameres obliquely truncate. Penis elongate, subparallel-sided. Variation. Body&male;&male; (n = 49) length: 18.00–24.25(25.75) mm, width: 6.25–9.20 mm, length/ width ratio: 2.60–2.85(2.96); &female;&female; (n = 42) length: 19.25–28.50(30.75) mm, width: 7.25–11.50 mm, length/width ratio: 2.51–2.78. Very variable taxon. Impressions on pronotum shallow to moderately deep and of variable size; punctation of pronotum, particularly in smaller specimens, coarser, punctures with larger fovea. Principal impressions round to elongate oval. Colouration of pronotum from mostly vivid green with bluish tint to mostly copper or violet, however majority of specimens with green background colour and golden/copper impressions and foveae of punctures; largely copper to violet pronotum is present only in specimens from Camplong. Colouration of elytra in series from Soë quite uniform, varying only in intensity of purple tint; only six specimens with bluish tint to mostly blue elytra and two with nearly black (? artificially) elytra. In series from Camplong five specimens with blue elytra and green pronotum and seven with pine green elytra and purple pronotum. Lateral impressions on elytra absent to variously present, if present posterior impression deeper and better indicated than anterior, which is only exceptionally truly impressed. Ventral side in most specimens green with variable extent of golden and copper reflections, sometimes nearly whole underside copper; exceptionally dark violet (five specimens with purple pronotum from Camplong). Specimens from Soë were badly damaged and about one quarter of them have missing tarsi. Of the remaining specimens 49 have last tarsomere metallic and 40 uniformly brown. Aedeagus (n = 7) length: 5.36–6.83 mm, width 1.12–1.43 mm, length/width ratio: (4.29)4.78–4.94(5.31). Differential diagnosis. Chrysodema (G.) katka sp. nov. is most similar to C. (G.) cupriventris as both have similarly green elytra and paler pronotum with golden and copper reflections. They can be easily separated by punctation of abdomen, which is moderately strong and very sparse in C. (G.) cupriventris while dense and coarse in C. (G.) katka. Additionally, C. (G.) cupriventris has lighter tone of elytra (i.e. fern green) and brightly copper-red ventral side. At first glance C. (G.) katka is also similar to C. (G.) foraminifera (also known from Timor) but the latter differ in brightly green ventral side of body and uniformly equally coloured pronotum and elytra while C. (G.) katka has dark ventral side and elytra distinctly darker coloured than pronotum. Chrysodema (G.) foraminifera also differs in narrower and more elongate penis than C. (G.) katka (Figs 56 vs 70). Etymology. This species is dedicated to Katka Štajerová for her support and love to the second author of this paper. The epithet is noun in apposition. Distribution. Indonesia / East Timor: Timor Is. We have examined several specimens labelled just Timor and therefore we cannot be sure whether they were collected in the Indonesian part of the island or in the East Timor., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 700-701, DOI: 10.5281/zenodo.5310800, {"references":["WATERHOUSE C. O. 1892: Two new Buprestidae from Damma Island. Annals and Magazine of Natural History, Series 6 10: 410 - 412."]}
Published: 2016
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45. Chrysodema (Gelaeus) wetteriana Frank & Sekerka 2016, stat. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema wetteriana, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) wetteriana (Théry, 1935), stat. nov. (Figs 54–56, 71) Gelaeus Walkeri subsp. wetteriana Théry, 1935: 248 (original description). Chrysodema (Gelaeus) walkeri subsp. wetteriana: BELLAMY (2008): 542 (catalogue). Type locality. Indonesia, Maluku Province, Wetar Island. Type material examined. LECTOTYPE (present designation): &male;, ‘K. Schädler / Wetter / acq. 1898. [w, h] // Gelaeus / Walkeri / ssp. Wetteriana / Thery / &male; TYPE [w, h] // C. wetteriana / THY. / det. HO&Lstrok;Y&Nacute;SKI / PARATYPE [g, h]’ (RMNH). PARALECTOTYPES (1 &male; 2 &female;&female;): 1 &female;, ‘K. Schädler / Wetter / acq. 1898. [w, h] // Gelaeus / Walkeri / ssp. Wetteriana / Thery / &female; TYPE [w, h] // wetteriana THY. / HOLOTYPE! / (cf. size) / det. R. HO&Lstrok;Y&Nacute;SKI [r, h]’ (RMNH); 1&male;, ‘K. Schädler / Wetter / acq. 1898. [h] // Paratype [y, p] // 1934–692 [p] // Gelaeus / Wetteriana / Thery / PARATYPE [w, h]’ (BMNH); 1 &female;, ‘K. Schädler / Wetter / acq. 1898. [w, h] // Ceram / Jilo / C.Ribbe 1884 [w, p, sic!] // Gelaeus / Wetteriana / Thery / PARATYPE [w, h]’ (MHNH). All specimens provided with an additional red printed label: ‘ LECTOTYPE [or PARALECTOTYPE respectively] [sex] / Chrysodema (Gelaeus) / wetteriana / (THÉRY, 1935) / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Redescription of lectotype. Well preserved &male; specimen with all appendages intact, elytra with transverse crack at base. Length 27.00 mm, width 10.00 mm, length/width ratio: 2.70. Body generally black (Figs 54–56) partly with dark metallic reflections: purple on principal impressions of pronotum, marginal punctures on elytra, and most of ventral side; basal impressions of pronotum, and legs and mouthparts at joints with greenish tint. Apical half of last tarsomere and claws darkened but not metallic. Pronotum in general coarsely and densely punctate particularly anteriorly and laterally, punctures foveolate. Medial line well marked and impunctate. Medial impressions distinct, moderately deep, densely punctate, towards centre somewhat sparser; punctures slightly smaller than those along anterior margin. Principal impressions shallow and weakly bordered from disc, densely punctate. Lateral impressions shallow and punctate. Basal impressions small with anterior part less impressed than basal part. Elytra without lateral impressions, with strongly elevated intervals and distinctly impressed rows of punctures from base to apex. Punctation of elytra overall very coarse and dense. Rows visible from base to apex, irregularly double (internal) or triple (external) from basal third of elytra to apex. Punctures in rows V–VIII in middle third of elytra forming groups of 2–4 separated by elevated impunctate folds forming irregular elevated reticulate sculpture (Fig. 56). Intervals approximately 4× as wide as rows of punctures, with shallow transverse wrinkles, and apically with several additional punctures. Mid femora in central part moderately densely punctate, punctures large, on sides punctures smaller and denser. Ventral side of body (Fig. 55) overall coarsely and quite densely punctate. Abdominal ventrite I laterally densely and coarsely punctate, punctures large. Ventrite II laterobasally densely punctate, punctures distinctly smaller than on ventrite I, lateroapical area sparsely punctate. In each of subsequent ventrites densely punctate area proportionally smaller to the sparsely punctate area. Aedeagus (Fig. 71) not fully sclerotised and broken between parameres, therefore its width was not measured, length: 7.29 mm. Apices of parameres rounded. Penis elongate, subparallel-sided. Variation. Body &male;&male; (n = 2) length: 27.00– 27.25 mm, width: 10.00 mm, length/width ratio of body: 2.70–2.73; &female;&female; (n = 2): length: 29.50–30.50 mm, width: 11.00– 12.50 mm, length/ width ratio of body: 2.44–2.68. One female has pronotum slightly smaller and with sparser punctation, mainly in central part. Subhumeral denticle in paralectotypes distinctly larger, particularly in males. Differential diagnosis. Chrysodema (G.) wetteriana is readily distinguished from other taxa of the subgenus by nearly black body with slight metallic reflections on pronotum and strongly elevated intervals on elytra. It is the most sulcate species of all Gelaeus. Distribution. Indonesia, Maluku Prov.: Wetar Is. Remarks. THÉRY (1935) described this taxon as subspecies of C. (G.) walkeri based on the material from RMNH but did not mention the number of specimens he had at disposal. At the beginning of the description he stated: ‘Taille du type (long 28 mm; 5.25 mm).’ but did not mentioned the sex of the ‘type’ nor is it indirectly indicated in the description. The width of ‘type’ must have been given erroneously as the width of studied specimens is 10–11 mm; the length is also not in agreement with presently studied specimens as they are either smaller or larger. The description itself is rather short followed by longer paragraph comparing C. (G.) wetteriana to C. (G.) walkeri where Théry mentioned a female of C. (G.) wetteriana. In the last sentence he compared the apical notch on abdominal ventrite V in both sexes making obvious he must have had more than one specimen. There are two specimens (male and female) of C. (G.) wetteriana in RMNH, however, both were labelled by Théry as ‘type’. Additionally, there are two specimens labelled as ‘paratype’ deposited in BMNH and MNHN respectively. All four beetles bear identical locality label and Théryʼs original identification labels. As two of the four specimens are labelled ‘holotype’ and the description is very short it is not possible to identify any of the specimens as the type and we consider all four to be syntypes. According to the reference number 1934–692 the BMNH specimen was presented to the museum by Théry himself. In order to conserve the concept of this taxon and associate it with the morphology of male genitalia we designate the male specimen deposited at RMNH as the lectotype. The MNHN syntype has two locality lables, one identical to other syntypes, however, on the other is printed Ceram, Jilo. We assume the second locality label was accidentally pinned under the specimen as it was remounted. As mentioned above Théry described this taxon as subspecies of C. (G.) walkeri. In general C. (G.) wetteriana is most similar to the latter species-group, particularly to C. (G.) walkeri kubani subsp. nov. but it is very distinct (see differential diagnosis). Therefore we rise its rank to species., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 715-716, DOI: 10.5281/zenodo.5310800, {"references":["THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp."]}
Published: 2016
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46. Chrysodema (Gelaeus) sara Frank & Sekerka 2016, sp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema sara, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) sara sp. nov. (Figs 45–47, 68) Gelaeus walkeri walkeri [misidentification]: AKIYAMA & OHMOMO (2000): Pl. 42, Figs 445-3. Type locality. Indonesia, Maluku Province, Babar Island, Tepa. Type material. HOLOTYPE: &male;, ‘ Indonesia, S. Moluccas / BABAR ISL., 0 – 250 m alt / Tepa env., 7.2010 / local collection lgt’ (NMPC). PARATYPES (2 &female;&female;): 1 &female;, same data as holotype (SGCB); 1 &female;, ‘ Indonesia, S. Moluccas / WETANG ISL, 4.2009 / Rumalewan Besar vill env / Babar archipelago / local collectors lgt’ (DFPC). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / oborili sp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved&male; specimen with all appendages, only left two distal antennomeres missing. Length 24.00 mm, width 8.00 mm, length/width ratio: 3.00. Head purple-green. Pronotum and scutellum copper-green. Elytra bluish-black with apicolateral margins bright bluish-green and violet (Figs 45, 47). Ventral side dark violet laterally and purple-red centrally (Fig. 46). Legs copper-red with golden reflections on femora, violetgreen tibiae, tarsi brown with metallic claws. Pronotum in general coarsely but sparsely punctate, laterally punctures moderately dense. Medial line well marked impunctate. Medial impressions shallow, more impressed along medial line, moderately densely punctate, punctures slightly smaller than remaining ones on disc. Principal impression, oval, moderately deep, not connected to basal impressions, variously punctate with several moderately large punctures and line of fine punctures. Lateral impressions shallow and rugose. Basal impressions oval, moderately deep. Elytra without lateral impressions, finely punctate. Rows visible from base to apex, not impressed, basally regularly arranged, apically gradually irregular and irregularly double in apical half. Punctures in rows V–VIII in middle third of elytra more or less regular and sparse forming distinct rows, not isolated groups; spaces much broader than puncture diameter, flat not reticulate (Fig. 47). Intervals flat, not elevated, broad, approximately 4–6× as wide as rows of punctures, basally impunctate, apically gradually narrowed and partly vanishing. Mid femora in central part strongly shiny, very sparsely and finely punctate, on sides punctures slightly larger but still sparse. Thoracic ventrites (Fig. 46) centrally nearly impunctate, laterally moderately densely punctate, punctures large. Abdominal ventrites sparsely punctate, only ventrite I laterally moderately densely punctate. Aedeagus (Fig. 68) length 6.12 mm, width 1.53 mm, length/width ratio: 4.00. Apices of parameres obliquely truncate. Penis stout-oval and short in contrast to other taxa. Variation. Body &female;&female; (n = 2): length: 26.75–34.00 mm, width: 9.50–12.75 mm, length/width ratio: 2.67–2.82.All three specimens quite similar, not showing much variation. Female from Babar Island appears more sparsely punctate due to larger size of its body. Differential diagnosis. Readily recognizable species due to its smooth elytra with fine punctation. Chrysodema (G.) sara sp. nov. has the finest punctation of the elytra of all Gelaeus taxa. In generall it is most similar to C. (G.) walkeri walkeri, which has quite smooth elytra in comparison to remaining taxa but still has distinctly impressed rows of punctures and slightly elevated intervals posteriorly while C. (G.) sara does not have impressed rows or elevated intervals. Additionally, C. (G.) sara has the shortest and stoutest penis of all Gelaeus species and is practically the only with a distinctly broadly oval penis. Etymology. This species is named after Sára Franková, daughter of the first author of this paper. The species epithet is a noun in apposition. Distribution. Indonesia: Maluku Prov.: Babar Is. and Wetang Is. Remarks. Figure 445-3 published by AKIYAMA & OHMOMO (2000) as C. (G.) walkeri walkeri is in fact C. (G.) sara sp. nov., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 704-705, DOI: 10.5281/zenodo.5310800, {"references":["AKIYAMA K. & OHMOMO S. 2000: The buprestid beetles of the world. In: FUJITA H. (ed.): Mushi-Sha's Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, 341 pp. + 30 pls."]}
Published: 2016
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47. Chrysodema (Gelaeus) walkeri subsp. nigriventris Frank & Sekerka 2016, subsp. nov

Author: Frank, David and Sekerka, Lukáš
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Chrysodema walkeri, Animalia, Chrysodema (gelaeus) walkeri nigriventris, Biodiversity, Buprestidae, Taxonomy
Abstract: Chrysodema (Gelaeus) walkeri nigriventris subsp. nov. (Figs 4, 33–35, 65) Gelaeus Walkeri var. nigriventris Théry, 1935: 248 (unavailable infrasubspecific name). BELLAMY (2008): 542 (catalogue, listed as synonym of C. (G.) walkeri). Gelaeus walkeri walkeri [misidentification]: AKIYAMA & OHMOMO (2000): Pl. 42, Figs 445-1, 445-2 (iconography). Chrysodema walkeri: LANDER (2003): Fig. 152 (colour photograph). Type locality. Indonesia, Maluku Province, Moa Island. Type material. HOLOTYPE: &male;, ‘Ins. Moa [w, p] // Coll. / Jul. Moser [w, p]’ (MFNB). PARATYPES (5 &male;&male; 77 &female;&female;): 1 &female;, ‘Ins. Moa [w, p] // COLLECTION / de BONNEUIL [w, p] // Pseudochrysodema Saund / Walkeri / I. Moa? [w, h] // Pseudochrysodema / Walkeri / v. nigriventris / Thery / PARATYPE [w, h]’ (NHMB, coll. G. Frey); 2 &female;&female;, ‘Ins. Moa [w, p]’ (NHMB, coll. G. Frey); 1 &female;, ‘Ins. Moa [w, p] // Chrysodema / walkeri Waterh. [w, h]’ (NHMB, coll. G. Frey); 1 &female;, ‘Moa / b. Timor [w, p] // Chrysodema / walkeri Wat. / Moa (Timor) [w, h] // walkeri [w, h]’ (NHMB, coll. G. Frey); 1 &female;, ‘Ins. Moa [w, p] // Chrysodema / walkeri / Wat / A. Descarpentries det. [w, h/p]’ (MHNH); 1 &female;, ‘Ile Moa [w, h] // Molukken / Ins. Moa [w, h] // cupriventris / Kerrem. [w, h] // Walkeri / Waterh. [w, h] // Chrysodema / Walkeri v. / nigriventris Théry / A. Descarpentries det. / ne doit pas éte décrit [w, h/p]’ (MHNH); 1 &male; 45 &female;&female;, same data as holotype (1 &male; 43 &female;&female; MFNB, 2 &female;&female; DFPC); 1 &female;, same data as holotype plus additional label ‘ Gelaeus // Walkeri Wat. [w, h]’ (MFNB); 1 &female;, ‘ Ins. Moa [w, p] // Museum Leiden / Ex coll. / G. van Roon [w, p] // Chrysodema / walkeri (WATH.) / nigriventris (THY.) / det. R. HOLY&Ncaron;SKI 2004 [w, p/h]’ (RMNH); 1 &female;, ‘ Ins. Moa [w, h] // Gelaeus Walkeri / Wat. / Det. Dr. Obenberger [w, h/p]’ (NMPC); 4 &female;&female;, ‘Ile Moa / INDONESIE / v.1992 / Coll. T. LANDER [w, h/p] // Galaeus / walkeri / (Waterh. 1892) / Dét. T. LANDER 1992 [w, h/p]’ (MHNG); 1 &female;, ‘ Ile de Moa / v.1992 / Coll. T. LANDER [w, h/p] // Galeus / walkeri / (Waterh. 1892) / Dét. T. LANDER 1993 [w, h/p]’ (MHNG); 1 &female;, ‘INDONESIE / Moa Isl. / vii.1998 / Coll. T. LANDER [w, p/h] // Gelaeus / walkeri / (Wat. 1892) / Dét. T. LANDER 2000 [w, h/p]’ (MHNG); 1 &female;, ‘ GELAEUS WALKERI WALKERI / (bon A2, lot de 5 pcs) / Moa / INDONESIE, iii.2001 / 8462 L [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p]’ (NMPC); 3 &female;&female;, ‘ INDONESIA, iii.2001 / Moa Is. / local collector [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p]’ (2 &female;&female; NMPC, 1 &female; IRSN); 1 &female;, ‘ INDONESIA, iii.2001 / Moa Is. / local collector [w, p] // ex coll. S. Bílý / National Museum / Prague, Czech Republic [w, p] / Gelaeus / walkeri Wat. / Sv. Bílý det.’ (NMPC); 4 &female;&female;, ‘ INDONESIA, / Maluku, March 2001, / MOA isl., / native coll. [w, p]’ (3 &female;&female; VKSC, 1 &female; DFPC); 1 &male;, ‘ Moa / Feb. 2002 [w, p]’ (VKSC); 1 &male;, ‘ Moa / Juni. 2003 [w, p]’ (DFPC); 4 &female;&female;, ‘MOA ISL. / 2/2008 [w, h]’ (3 &female;&female; SJPC, 1 &female; DFPC); 2 &male;&male; 2 &female;&female;, ‘Indonesia, SW Moluccas / MOA ISLAND., 0 - 200 m alt / E of TIMOR., 1.2009 / KAIWATU vill / local collectors lgt [w, p]’ (1&male; 2&female;&female; SJPC, 1&male; DFPC); 1&female;,‘SUMATRA / Pulau [= Island] Moa [w, p] / GALEUS / walkeri Waterhouse / det. Barries 1995 [w, p]’ (WBWA). All specimens provided with an additional red printed label: ‘ HOLOTYPE [or PARATYPE respectively] [sex] / Chrysodema (Gelaeus) / walkeri / nigriventris subsp. nov. / David Frank & / Lukáš Sekerka des. 2016 [year handwritten]’. Description of holotype. Well preserved &male; specimen with all appendages, missing only left antenna from antennomere VIII on. Length 29.50 mm, width 11.00 mm, length/width ratio: 2.68. Body weakly shiny. Head, pronotum and scutellum dark green, impunctate areas on pronotum with dark purple tint. Elytra dark viridian, apical half with purple-violet metallic reflections (Fig. 33). Ventral side dark green with purple and golden reflections (Fig. 34). Legs green with strong golden reflections, whole tarsi brown, only claws metallic. Pronotum in general moderately coarsely and densely punctate. Medial line well marked, impunctate, not elevated. Medial impressions shallow, more impressed internally than externally, coarsely and moderately densely punctate, punctures subequal to those on disc, basally denser. Principal impressions elongate and shallow, not sharply bordered, well separated from basal impressions, impunctate but rugose. Lateral impressions shallow, oval, punctate. Basal impressions large and triangular. Elytra without lateral impressions, moderately strongly punctate. Rows visible from base to apex, weakly impressed and irregularly double in apical half, internal rows more or less regularly arranged, external gradually irregular. Punctures in rows V–VIII in middle third of elytra irregular but not forming isolated groups; spaces ca. 3× as broad as puncture diameter, not elevated (Fig. 35). Intervals basally flat, not elevated, broad, approximately 4–5× as wide as rows of punctures, impunctate; apically and laterally vanishing, strongly narrowed, slightly elevated and punctate. Ventral side (Fig. 34) including legs as in nominotypical subspecies. Only lateral sides of abdominal ventrites II–V densely punctate in basal half and very sparsely punctate in apical half; punctures moderately large, subequal in size (Fig. 4). Aedeagus (Fig. 65) length 7.50 mm, width 1.63 mm, length/width ratio: 4.60. Apex of parameres rounded. Penis elongate, subparallel-sided. Variation. Body &male;&male; (n = 6) length: (21.00) 24.25–29.50 mm, width: (7.50)8.75–11.00 mm, length/width ratio: 2.63–2.80; &female;&female; (n = 74) length: (22.75)24.00–35.75(38.00) mm, width: (8.25)9.25–13.00(14.00) mm, length/width ratio: (2.43)2.54–2.85. Head, pronotum and scutellum green with blue, turquoise, purple, golden or copper sheen. Impressions on pronotum usually golden. Elytra predominantly viridian with purple-violet reflection on apical half. Purple colour sometimes extends also to basal half of elytra. Rarely elytra dark blue-green. Underside nearly always dull, only exceptionally shiny, varying from almost uniformly dark green to mostly golden-green or copper-green. Punctation of pronotum variable in density and coarseness, medial impressions variously impressed, principal impressions oval to round and variously impressed, lateral impressions variously impressed, and basal impressions oval to triangular, always deeply impressed. Elytra exceptionally with lateral impressions; larger specimens smoother while smaller specimens rougher, particularly on apical third. Aedeagus (n = 5) length: 6.17–8.06 mm, width 1.17–1.63 mm, length/width ratio: 4.60–5.04(5.45). Differential diagnosis. Among the subspecies of C. (G.) walkeri this one is easily distinguished by viridian colouration of elytra (various shades of blue in other subspecies) and rounded apices of parameres (truncate in other subspecies). For additional characters see Table 1. Etymology. The epithet is combination of Latin words niger = black and venter = underside reflecting darkly coloured ventral side of body. Distribution. Indonesia, Maluku Prov.: Moa Is. Remarks. THÉRY (1935: 248) described Gelaeus walkeri var. nigriventris, however, in the same work he described also a subspecies Gelaeus walkeri wetteriana therefore the variety nigriventris is infrasubspecific name according to the Article 45.6.4 (ICZN 1999). Nor it was validated before 1985 (Article 45.6.4.1) and hence it is an unavailable name according to ICZN (1999). We have located one female specimen in NHMB from the locality Moa mentioned by THÉRY (1935) and provided with his original paratype label as the variety nigriventris. Additionally we have found a series of specimens in MFNB with identical locality label (specimens originated in the collection of Julius Moser) agreeing also morphologically with the NHMB specimen. In our opinion this taxon is a distinct subspecies of C. (G.) walkeri and we decided to use the same name as Théry. LANDER (2003) provided a colour photograph of C. (G.) walkeri, however, the specimen does not have the last tarsomere metallic green thus it does not belong to the nominotypical subspecies. In our opinion the depicted specimen belongs to C. (G.) walkeri nigriventris subsp. nov. based on general shape., Published as part of Frank, David & Sekerka, Lukáš, 2016, Revision of the subgenusGelaeus of Chrysodema (Coleoptera: Buprestidae: Chrysochroinae), pp. 671-719 in Acta Entomologica Musei Nationalis Pragae 56 (2) on pages 711-713, DOI: 10.5281/zenodo.5310800, {"references":["THERY A. 1935: Note sur les Buprestides du Museum de Leiden. Zoologische Mededeelingen (Leiden) 18: 241 - 256.","BELLAMY C. L. 2008: A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Volume 1: introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica No. 76, Pensoft, Sofia-Moscow, 625 pp.","AKIYAMA K. & OHMOMO S. 2000: The buprestid beetles of the world. In: FUJITA H. (ed.): Mushi-Sha's Iconographic Series of Insects 4. Gekkan-Mushi Co., Tokyo, 341 pp. + 30 pls.","LANDER T. 2003: Revision du genre Chrysodema. Collection Systematique, Vol. 8, Magellanes, Andresy, 94 pp."]}
Published: 2016
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48. Chrysodema Laporte & Gory 1835

Author: Bellamy, C. L. and Lander, T.
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: Genus Chrysodema Laporte & Gory, 1835 Chrysodema Laporte & Gory 1835: 1; Obenberger 1926: 129; Bellamy 2003: 35. Type species: Chrysodema sonnerati Laporte & Gory 1835 (fixed by subsequent designation: Opinion 2076, ICZN 2004: 128). Cyalithoides Fisher 1922: 4; Obenberger 1926: 135; Levey 1978: 154; Hoyski 1980: 274; Bellamy 2003: 31; 2006: 151 (gender). syn. nov. Type species: Cyalithoides fulgida Fisher 1922 (fixed by original designation)., Published as part of Bellamy, C. L. & Lander, T., 2008, The synonymy of Cyalithoides fulgida Fisher, 1922 with Chrysodema robusta Deyrolle, 1864 (Coleoptera: Buprestidae: Chrysochroinae), pp. 34-36 in Zootaxa 1811 (1) on page 34, DOI: 10.11646/zootaxa.1811.1.2, http://zenodo.org/record/5125098, {"references":["Obenberger, J. (1926) Buprestidae 1. In: W. Junk & S. Schenkling. (Eds.). Coleopterorum Catalogus, W. Junk, Berlin, Volume 12, Pars 84, 1 - 212.","Bellamy, C. L. (2003) An illustrated summary of the higher classification of the superfamily Buprestoidea (Coleoptera). Folia Heyrovskyana, Supplementum 10, 197 pp, 44 plates.","ICZN. (2004) Opinion 2076 (Case 3193). Chrysodema Laporte & Gory, 1835 and Iridotaenia Deyrolle, 1864 (Insecta, Coleoptera): usage conserved by the designation of C. sonnerati Laporte & Gory, 1835 as the type species of Chrysodema. Bulletin of Zoological Nomenclature 61 (2), 128 - 129.","Fisher, W. S. (1922) New buprestid beetles from Borneo and the Philippines Islands. Proceedings of the United States National Museum (Number 2428), 61 (7), 1 - 18.","Bellamy, C. L. (2006) Nomenclatural notes and corrections in Buprestidae (Coleoptera). The Pan-Pacific Entomologist, 81 (3 / 4), 145 - 158."]}
Published: 2008
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49. Chrysodema ventralis Waterhouse 1885

Author: Swezey, O. H.
Subjects: Coleoptera, Insecta, Arthropoda, Chrysodema ventralis, Chrysodema, Animalia, Biodiversity, Buprestidae, Taxonomy
Abstract: 2.Chrysodema ventralis Waterhouse, Ann. Mag. Nat. Hist., V, 15: 381, 1885. Obenberger, Coleopt. Catalog. (84): 135, 1926. Dededo, Aug. 11, on corn leaf, Swezey; Piti, Aug. 13, 24, Sept. 21, Oct. 12, 25, 27, Nov. 6, Swezey. Described from Guam, Ladrone Islands and not recorded elsewhere. This sdinewhat smaller bright green species was more abundant. One or two specimens were swept from Glochidion 111,arianum trees on several occasions at Piti., Published as part of Swezey, O. H., 1942, Miscellaneous Families of Guam Coleoptera, pp. 150-171 in Insects of Guam I, Honolulu, Hawaii :Bernice P. Bishop Museum on page 164, DOI: 10.5281/zenodo.5167701
Published: 1942
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