Chrysodema (Chrysodema) antennata subsp. obsoleta J. Thomson 1878

Chrysodema (Chrysodema) antennata obsoleta J. Thomson, 1878 (Figs 41–44, 46, 62, 67, 73) Chrysodema obsoleta J. Thomson (1878): 20 (original description); Kerremans (1892a): 39 (catalogue); Kerremans (1903): 76 (catalogue). Chalcophora obsoleta: Kerremans (1885): 126 (catalogue). Chrysodema (Chrysodema) obsoleta: Kerremans (1909): 520 (key), 582 (redescription); Obenberger (1926): 132 (catalogue). Chrysodema (Chrysodema) antennata obsoleta: Lander (2003): 48 (revision, downgraded to subsp. of C. (C.) antennata), 80 (colour Fig. 96, misidentification = C. (C.) sekerkai); Bellamy (2008): 527 (catalogue), 534 (listed as subspecies of C. (C.) antennata). Type locality. ‘ Celebes’ [Indonesia: Sulawesi Island]. Type material examined. LECTOTYPE (present designation): ♀ (22.00 × 7.00 mm; Figs 41–44), ‘Ex Musaeo | JAMES THOMSON [w(c), p (black border)] || TYPE [r, p] || Th. | TYPE [w, p (black border)] || Chrysodema | Obsoleta | (HDeyr) | Célèbes [y, h] || Obsoleta | (H. Deyr. M.s.s.) | Thoms. Type |T. B. 20. | Célèbes [w, h]’ (MNHN). The specimen was provided with an additional red printed label: ‘ LECTOTYPE ♀ | Chrysodema | obsoleta | THOMSON J., 1878 | David Frank des. IV. 2021 [date handwritten]’ and also white printed label ‘ Chrysodema (Chrysodema) | antennata | obsoleta | THOMSON J., 1878 | David Frank det. IV. 2021 [date handwritten]’. AddiTiOnal maTerial examined (3 ♂♂, 7 ♀♀). INDONESIA: Sulawesi Island (without accurate locality): Célèbes, 1 ♂ (MNHN; Figs 46, 67); Célebes, Carter, 1 ♀ (MNHN, ex coll. Théry); Celebes, 2 ♂♂, 1 ♀ (MNHN, ex coll. Lander); Indonesia, Celebes, 1 ♀ (DFPC). CENTRAL SULAWESI PROv.: Tonusu, b. Tentena, Poso-See, 21.- 26.4.1994, leg. Gerstmeier, Hiermeier, Romig, 1 ♀ (DFPC). NORTH SULAWESI PROv.: Sulawesi Utara, Dumoga Bone N. P., 0°33'N 123°40'E, 4.iii.1985, Base Camp Site, Malaise trap by fallen tree, C.H.C. Lyal, 1 ♀ (BMNH); Sulawesi Utara, Dumoga-Bone N. P., March 1985, Lowland forest edge ca 200m, Malaise trap, 1 ♀ (BMNH). SOUTH SULAWESI PROv.: Patunuang, Jan. 1896, H. Fruhstorfer, 1 ♀ (MNHN, ex coll. Van de Pool). Redescription of lectotype. Moderately well preserved ♀ specimen with missing right antenna from antennomere VI, and right middle and left hind legs; elytra slightly open. Length 22.00 mm, width 7.00 mm, length/ width ratio: 2.75. Body navicular, basic colour of dorsal side black-violet with metallic green macropunctation; ventral side green with coppery reflections, legs green. Head black with green area around eyes and punctures on dorsal side, green on ventral side. Eyes large, oval. Area around eyes green, finely macropunctate. Frons black with small green spot at center, with very sparse green macropunctures, 2× as wide as diameter of eye, impressed with deep medial sulcus, with very sparse short pubescence. Vertex more densely macropunctate than frons. Labrum pale brown, pubescent. Antennae serrate from antennomere IV. Antennomere I green, II green-brown, sparsely macropunctate and with short pubescence; radicula and antennomeres from III dark brown with sparse short pubescence. Antennomere II ca. 3× shorter than I and 2× than III. Antennomere IV long triangular, V–X more trapezoidal than triangular, XI oval. Maxillae, labium, maxillary palpi and labial palpi pale brown, densely pubescent. Pronotum trapezoidal, narrowing anteriad with rounded sides, widest at mid-length, black, with moderately dense irregular green macropunctures, micropunctate between macropunctures, 1.5× as wide as long.Anterior margin arcuate, lobe moderately protruding, densely pubescent. Medial line black, micropunctate, without macropunctures. Medial impressions weak and shallow only indicated by denser macropunctation. Principal impressions almost triangular, green, densely macropunctate. Lateral impressions green, moderately deep, macropunctate. Lateral margin green. Basal margin moderately bisinuate. Scutellum broadly trapezoidal, green and shiny. Elytra moderately wider than pronotum at base; subparallel at basal half, narrowing from beyond its midlength to apex; convex in lateral view. Lateral margins arcuate below humeral calli, serrate at apical third. Groups of macropunctures in basal half of almost the same size contain ca 5–10 macropunctures, arranged in almost regular rows. Black-violet areas between rows of groups of macropunctures look like weak costae which are interconnected at short intervals. Macropunctures at apical half in almost coherent rows between weak black-violet costae. Last two lateral intervals coherently very densely macropunctate therefore almost entire area is green. Humeral calli moderately furrowed. Epipleura horizontal, broad at basal quarter, green, moderately densely macropunctate and pubescent. Legs metallic green with bronze reflections; femora and tibiae macropunctate and pubescent more densely on inner side. Mesotibiae moderately curved at basal half and almost straight at apical half (Fig. 73). Tibiae with two apical ventral spurs. Tarsi metallic green with bronze reflections, tarsal claws divergent and simple. Ventral side green with bronze reflections. Hypomeron and prosternum macropunctate and with short pubescence. Anterior margin of prosternum with dense short pubescence. Prosternal process subparallel-sided, narrowed at apex, approximately 1.9× as long as wide, densely macropunctate at central part, macropunctures bigger than macropunctures on prosternum, apex and sides smooth, polished, nearly impunctate and sparsely pubescent. Metasternum densely macropunctate and pubescent. Abdominal ventrites I–IV centrally less, laterally very densely macropunctate, macropunctures at central part bigger than at lateral parts. Abdominal ventrite V very densely macropunctate at whole area. All abdominal ventrites with dense and short pubescence. Variation. Body ♂♂ (n = 3) length: 22.50–22.75 mm, width: 8.00– 8.50 mm, length/width ratio: 2.68–2.81; ♀♀ (n = 7): length: 22.00–27.25 (average 25.19) mm, width: 8.00–10.25 (average 9.25) mm, length/width ratio: 2.66– 2.75 (average 2.70). Groups of macropuctures can be variable but never large. Colour of macropunctures variable from green to red-bronze. Aedeagus (n = 2) length: 5.82 mm, width: 1.09 mm, length/width ratio: 5.34. Aedeagus narrowly rhomboidal, widest at mid-length. Parameres narrowly open at apex (Fig. 67). Differential diagnosis. Males of all subspecies of C. (C.) antennata have very dense and long pubescence on inner side of mesotibiae from mid-length to tarsi. Males of C. (C.) antennata obsoleta have antennae shorter than pronotum. From C. (C.) antennata antennata can be distinguished by shape of aedeagus which is narrowly rhomboidal. From C. (C.) antennata barriesi can be distinguished also by narrowly rhomboidal aedeagus, by parameres which are open only at apex (Fig. 67) and by V-shaped border of metallic green area of male abdominal sternite IX. Females can be distinguished from C. (C.) sekerkai by mesotibiae which are not bent at apical third (Fig. 73). Macropunctures on pronotum generally finer than in C. (C.) sekerkai and without golden-green margin. Chrysodema (C.) antennata obsoleta is generally the smallest subspecies of C. (C.) antennata. For additional characters see Key to species. Distribution. Indonesia: Sulawesi Island (Fig. 62). Remarks. Although J. Thomson (1878) wrote in his description of C. obsoleta ‘Le plus petit type à nous connu de la coupe actuelle;... [The smallest type known to me;...]’ it is not clear from this sentence if he had only one specimen or if one of specimens was the smallest. He also mentioned only a single length and width measurement ‘Long. 21 mill.; lat. 7 mill.’ but the specimen in MNHN labelled as type has 22 × 8 mm (Lander (2003) also mentioned 22 mm). On the other hand, the specimen has old type labels and it is almost certain that the specimen is the (holo)type or one of the syntypes. Therefore this specimen is designated as a lectotype in order to conserve the status of this taxon and avoid any further misinterpretations.
Published as part of Frank, David, 2022, Studies on the genus Chrysodema (Coleoptera: Buprestidae: Chrysochroinae) part II., pp. 507-543 in Zootaxa 5214 (4) on pages 530-531, DOI: 10.11646/zootaxa.5214.4.3, http://zenodo.org/record/7397816
{"references":["Thomson, J. (1878) Typi Buprestidarum Musaei Thomsoniani. E. Deyrolle, Paris, 103 pp.","Kerremans, C. (1892 a) Catalogue synonymique des Buprestides decrits de 1758 a 1890. Memoires de la Societe entomologique de Belgique, 1, 1 - 304.","Kerremans, C. (1903) Coleoptera Serricornia. Fam. Buprestidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 12 b. Verteneuil & Desmet, Bruxelles, pp. 49 - 112.","Kerremans, C. (1885) Enumeration des Buprestides decrits posterieurement au Catalogue de MM. Gemminger & de Harold. Annales de la Societe Entomologique de Belgique, 29, 119 - 157.","Kerremans, C. (1909) Monographie des Buprestides. Tome III. Chrysochroini - Chalcophorini: Chalcophorites (pars). Livraisons 13 - 19. J. Janssens, Bruxelles, pp. 385 - 604., pls. 17 - 22. https: // doi. org / 10.5962 / bhl. title. 9700","Obenberger, J. (1926) Buprestidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, Pars 84. W. Junk, Berlin, pp. 1 - 212.","Lander, T. (2003) Revision du genre Chrysodema. Collection Systematique. Vol. 8. Magellanes, Andresy, 94 pp.","Bellamy, C. L. (2008) A world catalogue and bibliography of the jewel beetles (Coleoptera: Buprestoidea). Vol. 1. introduction; fossil taxa; Schizopodidae; Buprestidae: Julodinae - Chrysochroinae: Poecilonotini. Pensoft Series Faunistica. No. 76. Pensoft, Sofia-Moscow, 625 pp."]}