Male gamete heterogeneity among females

TwoGener, a novel two-generation (parent-offspring) approach to the study of genetic structure is presented. It makes possible quantification of heterogeneity among the male gamete pools sampled by scattered maternal trees and to estimate mean pollination distance and effective neighborhood size. TwoGener is illustrated where the male gamete can be categorically or ambiguously determined and applied to an empirical study of Quercus alba in Misouri.