Rocky reef fish assemblages at six Mediterranean marine protected areas: broad scale patterns in assemblages structure, species richness and composition

Fish assemblages inhabiting shallow rocky reefs at six Italian marine protected areas (MPAs) were investigated to assess broad-scale patterns in assemblage structure, species richness and composition, and to evaluate the presence of a latitudinal gradient in the distribution pattern of a thermophilic fish, the ornate wrasse Thalassoma pavo. Fish abundance was estimated by visual censuses carried out within the integral reserve zones, within the 1--7 m depth range and during four sampling periods. Multivariate analyses revealed significant differences in the assemblage structure among MPAs, although patterns of variation changed considerably from time to time and they were not consistent with latitudinal or longitudinal gradient. Fish assemblages of Miramare (northern Adriatic) and those of the other MPAs clearly separated into two groups. Multivariate differences among MPAs mainly relied upon some relatively common species (e.g. Diplodus puntazzo and Muraena helena). Substantial contribution to t...