Homalium (sect. Nisa) Warb

5. Homalium sect. Nisa (Noronha ex Thouars) Baill. ex Warb. in Engler & Prantl, Nat. Pflanzenfam. III(6a): 36. 1893. �� Nisa Noronha ex Thouars, Gen. Nov. Madagasc.: 24. 1806. �� Homalium [unranked as ��] Nisa (Noronha ex Thouars) Baill. in Bull. Mens. Soc. Linn. Paris 1: 575. 1886. Typus: Homalium nudiflorum (DC.) Baill. (�� Nisa nudiflora DC.) (designated by Sleumer, 1973: 295). Stipules opposite the petiole base and fused (in H. intercedens Sleumer axillary, free). Inflorescences spicate; bract and bracteoles large, broad, thick-textured, persistent. Flowers sessile (rarely subsessile); perianth 5-6(-7)-merous. Sepals obovate to oblanceolate (to nearly ligulate), oblanceolateoblong or narrowly elliptical, or ovate to oblong-ovate), spreading, accrescent; calyx tube short, broadly funnelform to cup-shaped, or in fruit hemispherical; sepal glands large, elliptical to elongated oblong-elliptical or roughly trapezoid. Petals ovate to oblong, usually smaller than sepals and curving over fruit (in H. louvelianum similar in size and spreading), accrescent; sepals and petals not ciliate or ciliolate in conjunction with overall surface pubescence). Stamens 1 per petal, inserted between glands; anthers dorsifixed, broadly oblongelliptical with oblong-elliptical locules and a large connective, the slits of dehiscence nearly parallel. Upper surface of ovary often nearly flat in flower, in fruit becoming hemispherical or broadly conical; styles fused basally into a short but distinct column (rarely fused for most of length), the branches 3-4. Locule of fruit subglobose or vertically compressed, glabrous (short-pubescent); seeds 1 per fruit, subglobose, largely filling the locule (or several small immature seeds). Distribution. ��� Madagascar. Species included. ��� Homalium intercedens, H. louvelianum, H. nudiflorum, H. stelliferum H. Perrier.
Published as part of Applequist, Wendy L., 2016, A reconsideration of the infrageneric classification of Homalium Jacq. (Salicaceae), pp. 231-256 in Candollea 71 (2) on page 253, DOI: 10.15553/c2016v712a9, http://zenodo.org/record/5721656
{"references":["Sleumer, H. (1973). Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes). Bull. Jard. Bot. Natl. Belg. 43: 239 - 328."]}