Riama Gray 2005

KEY TO THE SPECIES OF RIAMA GRAY 1a. Limbs overlapping when adpressed against body.....................................................2 1b. Limbs not overlapping when adpressed............................................................10 2a. Hexagonal dorsal scales.........................................................................3 2b. Quadrangular dorsal scales.......................................................................4 3a. Dorsal scales obtusely hexagonal, preanal pores in males 2................................... R. achlyens 3b. Dorsal scales acutely hexagonal, preanal pores in males 4���6................................... R. shrevei 4a Keeled, striated, or rugose dorsal scales............................................................5 4b Smooth dorsal scales..................................................................... R. laevis 5a. Lateral scale rows 0���6...........................................................................6 5b. Lateral scale rows 7���12..........................................................................9 6a. Transverse rows of dorsal scales 25���35.............................................................7 6b. Transverse rows of dorsal scales 39���43................................................... R. balneator 7a. Lateral scale rows 2���6...........................................................................8 7b. Lateral scale rows 0���1.................................................................... R. striata 8a. Rugose dorsal scales...................................................................... R. vieta 8b. Keeled or striated dorsal scales......................................................... R. hyposticta 9a. Femoral pores on one leg 0���6............................................................. R. oculata 9b. Femoral pores on one leg 9���14........................................................... R. luctuosa 10a. Venter immaculate except near lateral surfaces.....................................................11 10a. Ventral scales with dark pigmentation............................................................12 11a. Preanal pores present in both sexes........................................................ R. inanis 11b. Preanal pores absent.................................................................. R. vespertina 12a. Keeled, striated, or rugose dorsal scales...........................................................13 12b. Smooth dorsal scales.................................................................. R. meleagris 13a. Keeled or striated dorsal scales, dorsal stripes in some individuals, preanal pores present or absent, three or four supraoculars.......................................................14 13b. Keeled dorsal scales, no dorsal stripes, preanal pores present in males and females, four supraoculars............................................................ R. labionis 14a. Transverse rows of dorsal scales greater than 31...................................................15 14b. Transverse rows of dorsals scales 27���31................................................... R. petrorum 15a. Three or four supraoculars, lateral ocelli present or absent, lip barring variable..........................17 15b. Three supraoculars, no lateral ocelli, no barring on lip...............................................16 16a. Supralabial-subocular fusion absent........................................................ R. raneyi 16b. Supralabial-subocular fusion present................................................... R. laudahnae 17a. Superciliary series complete or incomplete, transverse rows of ventrals 19���27, anterodorsal thigh scales smooth, rugose, or striate.................................................18 17b. Superciliary series incomplete, transverse rows of ventral scales 18���20, anterodorsal thigh scales smooth.................................................... R. colomaromani 18a. Keeled or striated dorsal scales, longitudinal rows of dorsal scales 20���34, preanal pores present or absent, femoral pores on one leg of males 0���11................................19 18b. Keeled dorsal scales, longitudinal rows of dorsal scales 21���22, preanal pores absent, femoral pores of males on one leg 3���5........................................... R. columbiana 19a. Keeled or striate dorsal scales, subdigital lamellae on fourth toe 10���18, superciliary series complete or incomplete.........................................................20 19b. Keeled dorsal scales, subdigital lamellae on fourth toe 15���18, superciliary series complete........................................................................... R. orcesi 20a. Transverse rows of ventral scales 19���27, hemipenis with or without small or large spines.................21 20b. Transverse rows of ventral scales 19���20, hemipenis with small spines.......................... R. simotera 21a. Femoral pores of males 0���10, preanal pores of males 0���4, femoral pores of females 0���9, preanal pores of females 0���4....................................................................22 21b. Femoral pores of males 8���9, preanal pores of males 2���4, females lack femoral and preanal pores....................................................................... R. stigmatoral 22a. Hemipenis with small or absent spines, superciliary series usually incomplete...........................23 22b. Hemipenis with large spines, superciliary series complete................................... R. anatoloros 23a. Preanal pores present in both sexes...................................................... R. unicolor 23b. Preanal pores absent................................................................. R. cashcaensis KEY TO THE SPECIES OF PROCTOPORUS TSCHUDI 1a. Two to three supraoculars........................................................................ 2 1b. Four supraoculars...................................................................... P. pachyurus 2a. Venter uniformly dark or with dark stippling or mottling near lateral scale rows.......................... 3 2b. Venter clear yellow or orange without dark mottling......................................... P. guentheri 3a. No continuous series of lateral ocelli............................................................... 4 3b. Continuous series of lateral ocelli......................................................... P. unsaacae 4a. Frontonasal scale longer than frontal scale................................................. P. sucullucu 4b. Frontonasal scale equal in length to frontal scale........................................... P. bolivianus KEY TO THE SPECIES OF PROCTOPORUS TSCHUDI 1a. Two to three supraoculars........................................................................ 2 1b. Four supraoculars...................................................................... P. pachyurus 2a. Venter uniformly dark or with dark stippling or mottling near lateral scale rows.......................... 3 2b. Venter clear yellow or orange without dark mottling......................................... P. guentheri 3a. No continuous series of lateral ocelli............................................................... 4 3b. Continuous series of lateral ocelli......................................................... P. unsaacae 4a. Frontonasal scale longer than frontal scale................................................. P. sucullucu 4b. Frontonasal scale equal in length to frontal scale........................................... P. bolivianus
Published as part of Doan, Tiffany M. & Castoe, Todd A., 2005, Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus, pp. 405-416 in Zoological Journal of the Linnean Society 143 (3) on pages 413-414, DOI: 10.1111/j.1096-3642.2005.00145.x, http://zenodo.org/record/4634632