Generic recircumscription in the Loxocarpinae (Gesneriaceae), as inferred by phylogenetic and morphological data

12 The Loxocarpinae, also known as the “Boea group”, are the subtribe of Gesneriaceae 13 which includes Boea and a number of segregated genera and close relatives. This group 14 currently comprises over 200 species in 15 genera. Here we present the most up-to-date 15 phylogeny, covering all the genera known to belong to the group, based on Bayesian 16 Inference and Parsimony of the nuclear ITS and the plastid regions trnL-trnF (intron 17 and spacer) and ndhFtrnL (spacers). The results show discrepancies between the 18 current generic delimitation in the subtribe and the clades delineated by the phylogeny. 19 As a result Boea, Damrongia, Streptocarpus and Paraboea are recircumscribed in an 20 attempt to establish a more natural classification and a number of new combinations are 21 made. The new genus Middletonia is described. 22