Sisyphus fasciculatus Boheman 1857

Sisyphus fasciculatus Boheman 1857 (Figs. 2, 3C) Boheman 1857: 192; Harold 1869: 984; P��ringuey 1902: 101���106, Arrow 1927: 465; Haaf 1955: 376; Ferreira 1972: 799; Davis et al, 2008: 154; Montreuil 2015c: 3, 11. Type locality: Caffraria Size: Male: length: 9.4��� 7 mm, width: 5.7��� 3.5 mm. Female: length: 9.0���7.0 mm, width 5.6��� 3.5 mm. Diagnosis: S. fasciculatus is similar to S. muricatus and S. neobornemisszanus, but differs from these two species by bearing tufts of dense setae on the pronotum, which alternate with bare patches of short setae. Examined type material Holotype (photograph): (♀ NHRS): / Caffraria /, / J. Wahlb /, / type /, brown label, handwritten / fasciculatus Bohem /; red label / Typus: Sisyphus fasciculatus, det. Dr. E. Haaf, 1954/, /311:72/, /247:73/; /9415 E92 +/, brown label, handwritten / fasciculatus Boh /, /NHRS-SRAH 000000228/. Examined non-type material: See Supplementary information. Distribution: S. fasciculatus occurs in South Africa and Swaziland where it is a shade specialist (Davis et al. 1999). It is associated with riverine and lower hillside forest to dense partially shaded woodland thickets, particularly, on the eastern escarpment of Limpopo, Mpumalanga, west KwaZulu Natal and Swaziland. However, it also occurs in the warmer inland parts of the Maputaland Coastal Forest Mosaic ecoregion of Olson et al. (2001) southwards to the northernmost extent of the KwaZulu-Pondoland Coastal Forest Mosaic (Fig. 4). Conservation status: Although currently assessed as Least Concern (LC), S. fasciculatus is a potentially threatened species, due to the loss of forest and shaded woodland across its fairly restricted range (Davis 2013a). However, the species has been recorded in reserves where shaded vegetation is protected, such as in Hluhluwe, Umfolozi and Ithala Game Reserves (KwaZulu Natal) as well as Mlawula Nature Reserve (Swaziland). Because of this protection, it can be locally abundant in both coastal reserves and along the lower part of the northern escarpment of the Drakensberg.
Published as part of Daniel, Gimo M., Davis, Adrian Lv., Sole, Catherine L. & Scholtz, Clarke H., 2020, Taxonomic review of the tribe Sisyphini sensu stricto Mulsant, 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) in southern Africa, including new species descriptions, pp. 1-61 in Insect Systematics & Evolution 51 (1) on pages 13-15, DOI: 10.1163/1876312X-00002195, http://zenodo.org/record/3786629
{"references":["Boheman, C. H. (1857) Insecta Caffrariae annis 1838 - 1845 a JA Wahlberg collecta. Pars. II. Coleoptera (Scarabaeides). Holmiae: Offieina Nordstedtiana. - 395 pp, 1.","Harold, E. von (1869). Scarabaeidae. Gemminger, M. and Harold, E. Catalogus Coleopterorum. bucusque descriptorum, synonymicus et systematics, autotribus, 4: 976 - 1346. http: // www. iucnredlist. org / static / categories _ criteria _ 3 _ 1 access on 2017.03. 23.","Arrow, G. J. (1927) Notes on the Coleopterous genus Sisyphus. Annals and Magazine of natural History, 9 (19): 456 - 465.","Haaf, E. (1955) Uber die Gattung Sisyphus Latr. (Col. Scarab.). Entomologische Arbeitenaus dem Museum G. Frey, 6 (1): 341 - 381.","Ferreira, M. C. (1972) Os Escarabideos de Africa (sul do Saara). Revista de Entomologia de Mocambique, 11 (1968 - 1969): 5 - 1088.","Davis, A. L. V., Frolov, A. V., & Scholtz, C. H. (2008) The African dung beetle genera. Protea Book House. 148 - 155 pp.","Montreuil, O. (2015 c) Premiers cas de brachypterisme dans le genre Sisyphus Latreille (Coleoptera, Scarabaeidae, Sisyphini). Annales de la Societe entomologique de France, 51 (4): 281 - 293.","Davis, A. L. V., Scholtz, C. H., & Chown, S. L. (1999) Species turnover, community boundaries and biogeographical composition of dung beetle assemblages across an altitudinal gradient in South Africa. Journal of Biogeography, 26 (5): 1039 - 1055.","Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V., Underwood, E. C., & Loucks, C. J. (2001) Terrestrial Ecoregions of the World: A New Map of Life on Earth: A new global map of terrestrial ecoregions provides an innovative tool for conserving biodiversity. BioScience, 51 (11): 933 - 938.","Davis, A. L. V. (2013 a) Sisyphus fasciculatus. The IUCN Red List of Threatened Species 2013: e. T 137250 A 522146. http: // dx. doi. org / 10.2305 / IUCN. UK. 20132. RLTS. T 137250 A 522146. en. (accessed 07 September 2016)."]}