Octoglena Wood 1864

Genus Octoglena Wood, 1864 Octoglena Wood, 1864: 186; 1865: 229. Bollman, 1893: 117, 137, 187. Jeekel, 1971: 41. Hoffman, 1980: 73; 1999: 29. Shelley, 1996: 108 ���112; 2002 b: 92. Hypozonium Cook, 1904: 62. Cook and Loomis, 1928: 17. Chamberlin and Hoffman, 1958: 187. Buckett, 1964: 29. Jeekel, 1971: 39. Hoffman, 1980: 73. Kevan, 1983: 2962. Euzonium Chamberlin, 1950: 1. Chamberlin and Hoffman, 1958: 187. Buckett, 1964: 29. Jeekel, 1971: 38. Hoffman, 1980: 73. Type species. Of Octoglena, O. bivirgata Wood, 1864, by monotypy; of Hypozonium, H. anurum Cook, 1904, by monotypy; of Euzonium, E. crucis Chamberlin, 1950, by original designation. Description (adapted from the diagnosis in Shelley (1996)). Dorsum variably colored, either subuniformly pale yellow to white, with three dark longitudinal stripes on both pro- and metazona, or with transverse bands on prozona (Fig. 1). Dorsum convex and glabrous, smooth or lightly granular. Prozona relatively long, elevated above and extending entire widths of metaterga, strictures distinct, ridged; paranota absent; caudal metatergal margins detached from succeeding tergites, elevated and upturned to varying degrees; body broad (W/L ratio varying from 27���38 %), flattened "bell-shaped" in profile, sides extending strongly laterad. Head strongly pyriform (Fig. 4) with fine, light setation; three ocelli on each side arranged linearly in pigmented, diverging rows arising at level of antennal sockets and extending onto epicranium; antennae and antennomeres relatively short, becoming progressively more hirsute distad. Collum broad, overhanging epicranium and at least one pair of ocelli (Fig. 4); telson either broad and comprising nearly entire caudal width, or relatively narrow and comprising around half of breadth. Sterna moderately broad, opposing coxae separated in midline. Legs lightly hirsute, extending to lateral margins of body, podomeres small, claws indistinct. Gonopods in situ either upright and directed subventrad or recumbent and directed anteriad (Figs. 2���3). Anterior gonopods either curving submediad and overhanging sternal lobes or upright and lying parallel to latter; sterna with strong, rounded, apically hirsute lobes segregated to varying degrees; ultimate podomeres either in form of calyx (Fig. 6) or divided, with broad, hirsute, ventral lobes of varying sizes usually overhanging glabrous, dorsal counterparts, latter either short, broad, and apically blunt, curved slightly ventrad, and directed anteriomediad, or long, narrow, and acuminate, slightly sinuate and directed sublaterad; corners of 4 th and 5 th podomeres either extended on caudal sides or latter with variably broad, curvilinear or sinuate projection (Fig. 7); coxae with or without hirsute lobes, lengths variable. Posterior gonopods with ultimate podomeres simple and acicular, apically acuminate, fimbriate, angled, or lightly hirsute, projecting anteriad between dorsal and ventral lobes of anterior gonopod ultimate podomeres. Species. Six. Distribution. The Pacific Coast of North America from the vicinity of Vancouver (city), BC, to Santa Cruz Co., California, extending inland to the western slope of the Cascade Mountains from BC to central Oregon and the eastern slope of the Coast Range in southern Oregon and California, with localized, allopatric species some 74 and 320 mi (120 and 512 km) to the east in California and the Idaho Panhandle, and a widespread species in the southeastern US extending from westcentral South Carolina to southcentral Tennessee and northwestern Alabama (Fig. 9). Occurring along the Pacific Coast and in the southeastern states, the distribution of Hirudisomatidae is congruent with that of Andrognathidae (Platydesmida) (Shelley et al. 2005), which also occurs in northern Idaho (Gardner 1975, Hoffman 1999). Discovery of Hirudisomatidae in the latter is thus not surprising because congruence in the first two areas is matched in the third.
Published as part of Shelley, Rowland M., Richart, Casey H. & Bogan, Arthur E., 2010, Octoglena claraqua, n. sp. (Polyzoniida: Hirudisomatidae), a new milliped from Idaho, USA; first record of the order from the western interior of North America, pp. 55-64 in Zootaxa 2446 on pages 56-57, DOI: 10.5281/zenodo.195063
{"references":["Wood, H. C. (1864) Description of new genera and species of North American MYRIAPODA. Proceedings of the Academy of Natural Sciences, Philadelphia, 16, 186 - 187.","Bollman, C. H. (1893) The Myriapoda of North America. United States National Museum Bulletin, 46, 1 - 210.","Jeekel, C. A. W. (1971) Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging, 5, 1 - 412 + i - xii.","Hoffman, R. L. (1980 [1979]) Classification of the Diplopoda. Museum d'Histoire Naturelle, Geneve, Suisse, 237 pp.","Shelley, R. M. (1996 [1995]) The milliped family Hirudisomatidae in the New World (Polyzoniida). Brimleyana, 23, 103 - 143.","Cook, O. F. (1904) Myriapoda of northwestern North America. In: Harriman Alaska Expedition, 8 (Insects, pt. 1), 47 - 82.","Cook, O. F., and Loomis, H. F. (1928) Millipeds of the order Colobognatha, with descriptions of six new genera and type species, from Arizona and California. Proceedings of the United States National Museum, 72, 1 - 26.","Chamberlin, R. V., and Hoffman, R. L. (1958) Checklist of the millipeds of North America. United States National Museum Bulletin, 212, 1 - 236.","Buckett, J. S. (1964) Annotated list of the Diplopoda of California. Simmons Publishing Co., Davis, California. 34 pp.","Kevan, D. K. McE. (1983) A preliminary survey of known and potentially Canadian millipedes (Diplopoda). Canadian Journal of Zoology, 61, 2956 - 2975.","Chamberlin, R. V. (1950) Three new genera and eight new species of western millipeds. Chicago Academy of Sciences Natural History Miscellanea, 68, 1 - 6.","Shelley, R. M., McAllister, C. T., and Tanabe, T. (2005) A synopsis of the milliped genus Brachycybe Wood, 1864 (Platydesmida: Andrognathidae). Fragmenta Faunistica, 48 (2), 137 - 166.","Gardner, M. R. (1975 [1974]) Revision of the millipede family Andrognathidae in the Nearctic region. Memoirs of the Pacific Coast Entomological Society, 5, 1 - 61.","Hoffman, R. L. (1999) Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication, 8, 1 - 584."]}