Quadrupedal Dinosaurs did not evolve fully pronated Forearms: New Evidence from the Ulna

Therians (marsupials and placentals), archosaurs, and chameleons are remarkable in that they evolved postures and gaits with inturned forelimbs. However, recent studies have indirectly recognized that, unlike fully pronated therian and chameleon forearms, dinosaur forearms were mechanically constrained by semi-pronated (misaligned) joints. This has led to the hypothesis that quadrupedal dinosaurs mitigated this constraint via proximal migration of the radius, indirectly forming a more pronated, tubular manus distally. To test this hypothesis, a standardized pose was used to examine the forearm pronation of ornithischian dinosaurs that were obligatory quadrupeds and facultative bipeds. Results show that only restructuring of the distal, not the proximal radius, causes additional pronation of the pre-axial edge of the carpus, but also unexpectedly reveal that the ulna may help form a tubular manus by supinating the post-axial edge. Thus, relative to the plane of the elbow joint the wrist and finger joints r...