A New Combination in Heterophyllium (Bryopsida, Sematophyllaceae), with a Key to the Himalayan Species

An evaluation of Asian Brotherella Loeske ex M. Fleischer reveals that a new combination in the genus Heterophyllium (Schimper) Kindberg is necessary to replace B. amblystega (Mitten) Brotherus for its noticeably dimorphic stem and branch leaves, strongly serrate leaf apices, differentiated alar cells in an excavate group, erect and symmetric capsules, and strongly papillose spores. Heterophyllium amblystegum (Mitten) Y. Jia, S. He & Crosby is here validated and lectotypified along with a full description and a key to the Himalayan species of Heterophyllium.