Phylogenetic placements of ustilaginomycetous anamorphs as deduced from nuclear LSU rDNA sequences<fnr rid="fn1"><fn id="fn1">Part 174 of the series ‘Studies in Heterobasidiomycetes’.</fn>

In order to integrate ustilaginomycetous anamorphs into the general phylogenetic system of Ustilaginomycetes, partial nuclear large subunit ribosomal DNA sequences of 56 teleomorphic and 19 anamorphic species of the Ustilaginomycetes were analysed. Maximum parsimony and neighbour joining confirm the new suprageneric system of Ustilaginomycetes and indicate that (i) the species of <e1>Pseudozyma</e1> represent anamorphs of Ustilaginales parasitizing grasses, (ii) <e1>Pseudozyma prolifica</e1>, the type of <e1>Pseudozyma</e1>, is very closely related to <e1>Ustilago maydis</e1>, (iii) <e1>Pseudozyma tsukubaënsis</e1> is probably synonymous with <e1>Ustilago spermophora</e1>, (iv) the species of <e1>Malassezia</e1> represent a group of its own within the Exobasidiomycetidae, (v) <e1>Tilletiopsis cremea</e1>, <e1>T. lilacina</e1> and <e1>T. washingtonensis</e1> belong to the Entylomatales and (vi) <e1>T. flava</e1>, <e1>T. fulvescens</e1> and <e1>T. minor</e1> are members of the Georgefischeriales. Like all <e1>Tilletiopsis</e1> species tested, <e1>T. albescens</e1> and <e1>T. pallescens</e1> are members of the Exobasidiomycetidae, but they cannot be ascribed to any of the known orders of this subclass. The description of the Malasseziales is emended.