The first fossil record of Cretaceous Ocelloveliinae (Heteroptera: Gerromorpha: Veliidae) revealed its habitat and palaeoclimate.

A new genus and species, Longivelia circuliplsus gen. et sp. nov., is established and assigned to the subfamily Ocelloveliinae by the following characters: ocelli present, all tarsi 3-segmented, pretarsus inserted preapically, etc. This is the first record of the Ocelloveliinae from mid-Cretaceous Kachin amber which also enriches the geographical record. Based on the new species preserved with a true fly (Diptera) in the same piece of amber, we hypothesize that the palaeoclimate environment of the habitat of L. circuliplsus gen. et sp. nov. at that time might be similar with the modern South African Cape, which can be recognized by the freshwater environment. • First fossil record of Ocelloveliinae from mid-Cretaceous Kachin amber. • The habitat of the new species is determined by its morphological characteristics and co-occurring organisms. • The palaeoclimate of Kachin is inferred from the climate in which the extant taxa survived. [ABSTRACT FROM AUTHOR]