Expansion of grasslands across glacial Sundaland caused by enhanced precipitation seasonality.

The distribution of vegetation across glacial Sundaland and the underlying mechanisms have long been debated. We address this issue by comparing a compilation of vegetation reconstructions with climate simulations from the Community Earth System Model 2 (CESM2). A new n -alkane δ13C record derived from higher plants, covering the period from 21 to 5 cal ka BP in the southern South China Sea, is presented. This record, combined with previous pollen-based reconstructions, indicates that C 3 forests dominated the outer-to-mid shelf regions of the northern Sundaland between the Last Glacial Maximum (LGM) and Heinrich Stadial 1, while open grasslands occupied the inner shelf region. A compilation of vegetation reconstructions shows a glacial contraction of lowland rainforests in longitudinal extent compared to the Holocene, especially in southern Sundaland. Furthermore, the previously proposed "savannah corridor" or "dry tropical forests" existed in the interior of Sundaland during the LGM, extending from the Malay Peninsula to the Lesser Sunda Islands. Modelling studies suggest that changes in the seasonality of precipitation, rather than mean annual rainfall, are more relevant to vegetation changes between the LGM and the Holocene. The glacial emergence of Sundaland caused an eastward shift of the ascending branch of the Walker Circulation and a substantial reduction in atmospheric convection activity over Southeast Asia, leading to a decrease in water availability during the dry season in both hemispheres. The enhanced water stress thus favors the expansion of open environments. • Regional vegetation reconstructions are compared with hydroclimate simulations from the CESM2 model. • Glacial outer-middle continental shelf of northern Sundaland was covered by rainforests. • Savannah landscape or dry tropical forests occupied the glacial interior of Sundaland. • Enhanced dry-season water stress could account for glacial expansion of open environments. [ABSTRACT FROM AUTHOR]