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Plants use iron as a cofactor of proteins used in photosynthetic systems, electron transport chain and many more. Iron bioavailability for plants in soil is low because it tends to oxidise and create insoluable compounds.For this reasonplants haveevolvedtwo distinct iron uptake mechanisms.Because of the iron toxicitycaused by production ofreactiveoxygen species via the Fenton reaction and the unspecific transport of metals other than iron, plants have to regulate cellular iron concentrationtightly.Theyhave evolved a complex system of signalling networks that has recently begun to uncover. In additionto the regulation ofiron uptake, the plant cell combats iron toxicity by sequestering iron into storage organelles and by chelating it. Iron is essential for seed sprouting but this work is focused on transport of iron into the plant from the soil,subcellulartransportandlongdistance transport ofironin the vasculature.