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Parachernes (Parachernes) nitidimanus (Ellingsen, 1905) Material examined. 1 female, Pernambuco, Brazil: Furna do Morcego cave, 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR010). Diagnosis. According to Ellingsen (1905) and Beier (1932b) can be easily distinguished from the other species of the genus Parachernes Chamberlin (1931) by the carapace as long as broad 1.14×, gradually narrowing forwards, the short frontal margin straightened. Metazone distinctly granulated, yet a little glossy before the first furrow. Two distinct transverse furrows, the anterior one about in the middle, the posterior one somewhat nearer to the hinder margin than to the anterior furrow. Tergite divided (except XI), slightly granulated, al setae rather long, slightly clavate. Chaetotaxy tergal: 10: 10: 10: 8(1): 8(2): 6(2): 8(2): 8(2): 8(2): 10(2): 4(2ST):2. Genital region opercula with a central group of 18 short, three setae on each side, and ten marginal setae. Chelicerae, female with galea very slender and short, with some fine teeth in and near the extremity on the underside, male with about 6 terminal branches; paired spermatheca, typical of the genus. Pedipalps moderately strong, all segments smooth and glossy. The setae on the inner side of the trochanter and femur are slightly clavate, the other setae of the palps are slightly dentate in the extremity, on the fingers simple with some longer ones intermixed; trochanter and the proximal part of femur slightly shagreened; femur 2.35×, strongly granulated; patella 2.02×, more slightly, rather shagreened. chela with pedicel 3.18×, chela without pedicel 2.47×, hand round about smooth and very glossy, fingers as general smooth and glossy. Measurements (mm). Body length: 2.359. Carapace: 0.771 / 0.672. Pedipalps: trochanter: 0.347 / 0.197, femur: 0.544 / 0.235, patella: 0.589 / 0.266, chela with pedicel: 1.064 / 0.384, chela without pedicel: 0.950 / 0.384, movable finger: 0.538. Chelicera: 0.277 / 0.141, movable finger length:0.202. Leg I: femur: 0.105 / 0.127, patella: 0.317 / 0.137, tibia: 0.272 / 0.091, tarsus: 0.308 / 0.071. Leg IV: femur+patella: 0.569 / 0.158, tibia: 0.318 / 0.103, tarsus: 0.255 / 0.052. Distribution. Brazil, Saint Vincent, and the Grenadines (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 320, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Ellingsen, E. (1905) Pseudoscorpions from South America collected by Dr. A. Borelli, A. Bertoni de Winkelried, and Prof. Goeldi. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Universita di Torino, 20 (500), 1 - 17. https: // doi. org / 10.5962 / bhl. part. 9304","Beier, M. (1932 b) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i - xxi + 1 - 294.","Chamberlin, J. C. (1931) The arachnid order Chelonethida. Stanford University Publications, Biological Science, 7 (1), 1 - 284.","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

Ceriochernes foliaceosetosus Beier, 1974 Material examined. 1 female, Pernambuco, Brazil: Meu Rei cave, 08°29’14.1”S, 37° 16’48.8”W, 777 m a.s.l., 19- I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-012). Diagnosis. According to the description made by Beier (1974), the specimens of Ceriochernes foliaceosetosus Beier, 1974 can be easily distinguished by the carapace reddish-brown, slightly longer than wide, without eyes and eyespots, strongly granulated, the granules of the metazone covered with an epicuticle, reticulate in the form of a rough honeycomb, both transverse furrow narrow and not very deep, but distinct, the anterior curved laterally, the posterior closer to the rear margin, with six setae on the anterior margin, and eight on the posterior margin. All tergites divided, setae clavate and broadly foliated, rounded at the tip, curved and almost sessile, partially covered by epicuticle, with midrib, pleural membrane very densely granulated. Chaetotaxy tergal: 6: 8: 8: 6: 6: 8: 8: 8: 8: 8: 8: 2. Genital region operculum with a set of 12 setae, and ten setae on posterior margin. Chelicerae with six to seven setae on hand, es very short, three blades on the rallum, Serrula with 17 blades, galea with 5 very short branches distally. Pedipalps strongly reticulated, trochanter 1.0×, femur 2.58×, patella 2.15×, fingers not as long as the hand without pedicel, each with about 35–40 marginal teeth, the accessory teeth small and inconspicuous, present on the medial side only on the distal part of the finger, trichobothrium ist midway between isb and it, est proximal of ist; trichobothrium st closest to t. Measurements (mm). Body length: 2.021. Carapace: 0.638 / 0.538. Pedipalps: trochanter: 0.344 / 0.244, femur: 0.522 / 0.204, patella: 0.569 / 0.204, chela with pedicel: 0.864 / 0.346, chela without pedicel: 0.833 / 0.346, movable finger: 0.383. Leg I: trochanter: 0.124 / 0.105, femur: 0.127 / 0.093, patella: 0.255 / 0.093, tibia: 0.275 / 0.072, tarsus: 0.283 / 0.053. Leg IV: trochanter: 0.183 / 0.140, femur+patella: 0.488 / 0.152, tibia: 0.377 / 0.109, tarsus: 0.385 / 0.088. Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 321, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Beier, M. (1974) Brasilianische Pseudoscorpione aus dem Museum in Genf. Revue Suisse de Zoologie, 81, 899 - 909. https: // doi. org / 10.5962 / bhl. part. 76050","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

Petterchernes brasiliensis Heurtault, 1986 Material examined. 1 male, Pernambuco, Brazil: Furna do Morcego cave, 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR011). Diagnosis. According to Heurtault (1986) and Mahnert (1994) the males of Petterchernes brasiliensis Heurtault, 1986 can be easily distinguished by the carapace without eyes and without eyespots, dark brown, metazone finely granulated, setae of the carapace and tergites strongly clavate, with two deep transverse furrows, for setae on anterior margin; tergites divided, granulate. Chaetotaxy tergal: 9: 9: 9: 10: 10: 10: 12: 12: 10: 10: 10: 2. Genital region operculum with 32 setae and ten setae on posterior margin. Chelicerae, galea with multiple branches, and three blades in the rallum. Pedipalps coarsely granulate, setae stout and enlarged, trochanter 1.67×, with high dorsal hump, femur 2.22×, abruptly enlarged, patella 2.15×, chela with pedicel 2.28×, chela without pedicel 2.09×, trichobothrium isb, ib, and est at same level, hand with well pronounced internal hump bearing some longer and stouter setae. Legs, tarsus IV without tactile setae, claws smooth and simple, subterminal seta smooth, curve. Measurements (mm). Body length: 1.251. Carapace: 0.554 / 0.473. Pedipalps: trochanter: 0.269 / 0.161, femur: 0.477 / 0.214, patella: 0.415 / 0.193, chela with pedicel: 0.766 / 0.335, chela without pedicel: 0.7020 / 0.335, movable finger: 0.418. Leg I: trochanter: 0.137 / 0.085, femur: 0.131 / 0.115, patella: 0.106 / 0.076, tibia: 0.127 / 0.078, tarsus: 0.202 / 0.047. Leg IV: trochanter: 0.153 / 0.121, femur+patella: 0.380 / 0.121, tibia: 0.164 / 0.086, tarsus: 0.276 / 0.058. Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 321, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Heurtault, J. (1986) Petterchernes brasiliensis, genre et espece nouveaux de Pseudoscorpions du Bresil (Arachnides, Pseudoscorpionida, Chernetidae). Bulletin du Museum National d'Histoire Naturelle, Paris, 8, 351 - 355.","Mahnert, V. (1994) New chernetid pseuuoscorpions (Pseudoscorpionida: Chemetidae) from Venezuela and Brazil, with remarks on the genus Ancalochernes Beier. Revue suisse de Zoologie, 101, 829 - 838. https: // doi. org / 10.5962 / bhl. part. 79929","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

In a study of pseudoscorpions in caves from the northeastern Brazil, eight species of Pseudoscorpiones were identified. The known distribution of the species Pseudochthonius biseriatus Mahnert, 2001, Cryptoditha aff. elegans, Parachernes (P.) nitidimanus (Ellingsen, 1905), Petterchernes brasiliensis Heurtault, 1986, Ceriochernes foliaceosetosus Beier, 1974, and Progarypus setifer Mahnert, 2001 is extended to include the State of Pernambuco. Additionally, Geogarypus gollumi sp. nov. (Geogarypidae) and Progarypus smaugi sp. nov. (Hesperolpiidae) are described as new species.

During a survey of caves in a seasonally dry tropical forest area in northeastern Brazil, a new species of the genus Maxchernes was discovered and it is formally described here. The new species, na...

Rhopalochernes luiscarlosi sp. nov. Figs. 1A–F, 2A–G, 3A–G, 7A ZooBank Registration: urn:lsid:zoobank.org:pub: 82D786A8-5B1A-4CF5-8194-64EF3B303633 Type material. female holotype: COLOMBIA, Bolívar, Municipio de San Jacinto, Vereda La Flecha, Finca Amanecer Gaitero (9°51′09.74″N; 75°10′32.32″W), 250 masl. 10.XI.2019, K. Marimon & E. Villarreal leg. (ICN.APs841). Paratypes: 1 male and 2 females, with same data as for holotype. (IAvH-I-3866 [1♀]; (IAvH-I-3876 [1♀]; ICN-APs-842 [1♂]), Etymology. The specific epithet is a patronym in honor of the late professor Luis Carlos Gutierrez Moreno, who was a lead researcher in the Caribbean region of Colombia, as well as founder of the research group Biodiversidad del Caribe Colombiano at Universidad del Atlántico. Diagnosis. Rhopalochernes luiscarlosi sp. nov. is characterized by the following combination of characters: cheliceral hand with four setae (Figs. 2F, 3F); serrula exterior with 11 blades; eyeless (Figs. 1A, 1C, 1D, 1F); carapace with two deep transverse furrows (Figs. 1C, 1F); nodus ramosus proximal to st (Figs. 2E, 3E); and trichobothria st closer to t than sb (Figs. 2E, 3E). Differential diagnosis: Rhopalochernes luiscarlosi sp. nov. can be differentiated from R. chamberlini, R. panamensis and R. ohausi by the lack of eyespots. It also differs from R. chamberlini by the position of the nodus ramosus which is midway between sb and st (Figs. 2E, 3E), but at the same level as st in R. chamberlini. It can be separated from R. panamensis by the serrula exterior which has 11‒12 blades in R. luiscarlosi sp. nov. and 14‒15 blades in R. panamensis. From the type species, R. ohausi, R. luiscarlosi sp. nov. is distinguished by having gaping chelal fingers (not gaping chelal fingers in R. ohausi) and the position of nodus ramosus which is proximal to or at same level of st in R. ohausi (see Mahnert, 2001 Fig. 110). Rhopalochernes luiscarlosi sp. nov. is similar to R. insulanus and R. catalinae sp. nov. by the absence of eyes, but can be differentiated from R. insulanus by the presence of 4 setae on the cheliceral hand (6 in R. insulanus), a trait that is also present in R. chamberlini, R. panamensis and R. catalinae sp. nov. (see Heurtault, 1998, Figs. 4, 9). It can be distinguished from R. catalinae sp. nov. by the furrows on the surface of the carapace, which are deep and conspicuous in R. luiscarlosi sp. nov. (Figs. 1C, 1F), but shallow and inconspicuous in R. catalinae sp. nov. (Figs. 4C, 4F). Additionally, they differ by the shape of the spermathecae, which have elongated and narrow ducts in R. luiscarlosi sp. nov. (Fig. 7A) but have short, thick ducts in R. catalinae sp. nov. (Fig. 7B). Finally, R. luiscarlosi sp. nov. has a slender tarsus IV (4.00–4.25 × longer than deep) while that of R. catalinae sp. nov. is more robust (2.75–3.75 × longer than deep). Description of adult: Colour: Carapace and pedipalps reddish brown. Tergites and legs yellowish brown (Fig. 1A–F). Carapace: heavily granulated; 1.15‒1.23 (♂), 1.09‒1.27 (♀) × longer than broad, eyes or eye-spots absent; with 2 deep transverse furrows, posterior furrow closer to posterior margin than anterior furrow; posterior margin of carapace straight; anterior margin with 4 setae (♂, ♀), posterior margin with 8 (♂), 10 (♀) setae (Fig. 3C, F). Chelicera: 1.67‒1.80 (♂), 1.29‒1.83 (♀) × longer than broad; with 4 setae on hand (sbs absent) and 1 distal seta (gs) on movable finger, is, bs and es short and acuminate, ls longer and thicker (Figs. 2F, 3F); galea with 4 rami (♀) or simple (♂) (Figs. 2D, 3G); rallum with 3 blades (Figs. 2G, 3D), anterior blade with several ramifications while others smooth; serrula exterior with 11‒12 blades; lamina exterior present. Pedipalp: trochanter, femur, patella and chelal hand heavily granulated, with short, feathered and acuminate setae (Figs. 2C, 3C); chelal fingers smooth; trochanter 1.50‒1.62 (♂), 1.40‒1.71 (♀), femur 2.13‒2.19 (♂), 2.31‒2.67 (♀), patella 2.00‒2.20 (♂), 1.88‒2.20 (♀), chela (with pedicel) 2.60‒2.74 (♂), 2.80‒3.95 (♀), chela (without pedicel) 2.40‒2.53 (♂), 2.45‒2.63 (♀), hand 1.10‒1.21 (♂), 1.20‒1.26 (♀) × longer than broad, movable finger 1.09‒ 1.18 (♂), 1.04‒1.27 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, accessory teeth not visible. Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria: eb and esb placed basally, est medially at equal distance from et and eb, and et distally near the tip of the fixed finger; ib at the same level as esb and close to ist, which is placed midway in the first half of the fixed finger, isb slightly more distal than est and it closer to et than isb; b, sb, st and t about an equal distance from each other, sb slightly ahead of esb, st in the middle of movable finger slightly closer to t than sb, t placed distally and closer to st than tip of the finger. Venom apparatus present only in movable chelal finger with nodus ramosus proximal to st (Figs. 2E, 3E). Coxal region: maxilla smooth except for the antero-lateral region; coxae smooth; manducatory process triangular in shape with 2 apical setae, 1 prolateral suboral seta and 12 (♂), 9 (♀) additional setae on maxilla. Median maxillary lyrifissure rounded and situated submedially. Chaetotaxy of coxae I‒IV, ♂, 9: 9: 8: 15; ♀, 10: 13: 11: 20. Legs: granulate; feathered setae on retrolateral margin and dentate setae on prolateral margins. Femur + patella of leg IV 2.45 ‒ 2.60 (♂), 2.73‒2.80 (♀) × longer than deep; tibia 3.14‒3.50 (♂), 3.14‒3.83 (♀) × longer than deep; tarsus 4.00‒4.25 (♂), 4.00‒4.50 (♀) × longer than deep. Tarsi III and IV with short tactile seta, located subdistally. Arolium undivided, slightly shorter than claws; claws slender and simple, not modified (Figs. 2A‒B, 3A‒B). Abdomen: tergites I‒X and sternites IV ‒X with a clear medial suture line (Figs. 7C). Tergal chaetotaxy: ♂, 10: 12: 12: 14: 14: 14: 15: 14: 14: 12: 9 (2 T): 2; ♀, 14: 12: 11: 15: 17: 16: 15: 14: 12: 13: 10 (2 T): 2; Sternal chaetotaxy: ♂, 17:(1) 8 (1): (1) 10 (1): 13: 14: 14: 12: 11: 9: 9: 2; ♀, 16: (1) 10 (1): (1) 9 (1): 14: 14: 15: 12: 10: 9: 4: 2. Genitalia: female: spermathecae paired, with long ducts ending in oval sacs (Fig. 7A). Male: genital opercula containing 9 setae on anterior operculum (sternite II), 8 setae clustered together just anterior to the aperture, 4 in posterior margin of aperture, and 4 in distal margin of posterior operculum (sternite III). Dimensions: holotype female (ICN APs-841), with range of 4 female paratype in parentheses: Body length 1.36 (1.12‒1.36). Pedipalps: trochanter 0.22/0.15 (0.21‒0.24/0.14‒0.15), femur 0.35/0.15 (0.33‒0.38/0.14‒0.16), patella 0.31/0.15 (0.31‒0.34/0.15‒0.17), chela (with pedicel) 0.56/0.2 (0.53‒0.56/0.18‒0.20), chela (without pedicel) 0.49 (0.49‒0.50), hand (without pedicel) length 0.24 (0.22‒0.24), movable finger length 0.27 (0.26‒0.28). Carapace 0.5/0.46 (0.49‒0.52/0.41‒0.46). Leg IV: femur + patella 0.28/0.10 (0.28‒0.30/0.22‒0.23), tibia 0.22/0.07 (0.22‒0.23/0.06‒0.07), tarsus 0.17/0.04 (0.16‒0.18/0.04). Allotype male (ICN APs-842), with other male in parentheses: Body length 1.18 (1.03). Pedipalps: trochanter 0.21/0.13 (0.21/0.14), femur 0.34/0.16 (0.35/0.16), patella 0.32/0.16 (0.33/0.15), chela (with pedicel) 0.52/0.19 (0.52/0.20), chela (without pedicel) 0.48 (0.48), hand (without pedicel) length 0.23 (0.22), movable finger length 0.25 (0.26). Carapace 0.47/0.41 (0.48‒0.39). Leg IV: femur + patella 0.27/0.11 (0.26/0.10), tibia 0.22/0.07 (0.21/0.06), tarsus 0.17/0.04 (0.16/0.04).

Rhopalochernes catalinae sp. nov. Figs. 4A–F, 5A–G, 6A–G, 7B ZooBank Registration: urn:lsid:zoobank.org:pub: 33CE2DF6-A490-4524-89DF-D6C5D5A884A7 Type material. female holotype: COLOMBIA, Atlántico, Municipio de Usiacurí, Reserva Campesina La Montaña (10°46’02,06” N, 75°02’34” W), 150m, 08.VI.2019, K. Marimon and E. Villarreal leg. (ICN-APs-843). Paratypes: 1 female and 3 males with the same data as the holotype (IAvH-I-3856 [2♂]; ICN-Aps-844 [1♂, 1♀]); 2 females and 2 males, Colombia, Atlántico, Municipio de Juan de Acosta, Corregimiento de Chorrera (10°47′48.34″ N, 75°00′45.37″ W), 270 masl. 09. VI.2018, K. Marimon, D. Tilano and E. Villarreal leg (IAvH-I-3846 [1♀]; ICN-Aps- 845 [1♀, 2♂]); 1 female, Colombia, Bolívar, Municipio de San Jacinto, Vereda La Flecha, Finca Amanecer Gaitero (9°51′9.74″ N; 75°10′32.32″ W), 250 masl. 11-12.IX.2019, K. Marimon & E. Villarreal leg. (ICN-APs-846). Etymology. This species is named for our colleague Catalina Romero-Ortiz, in appreciation of her research on Colombian pseudoscorpions. Diagnosis: Rhopalochernes catalinae sp. nov. can be distinguished from other species of the genus by the following combination of characters: cheliceral hand with four setae (Figs. 5E, 6E); serrula exterior with 11-12 blades; eyeless (Figs. 4C, 4F); carapace with shallow transverse furrows (only visible in lateral view); nodus ramosus at the same level or slightly close to st (Figs. 5G, 6G); and trichobothria st midway between t and sb (Figs. 5G, 6G). Differential diagnosis: Rhopalochernes catalinae sp. nov. can be differentiated from R. chamberlini, R. panamensis and R. ohausi by the lack of eyespots (Figs. 4C, 4F). It also differs from R. chamberlini by the thicker femur + patella of leg IV, 2.60‒2.89 versus 3.73 × longer than deep in R. chamberlini. It can be distinguished from R. panamensis and R. ohausi by the serrula exterior which has 11‒12 blades, but 14‒15 blades in R. panamensis and 16 blades in R. ohausi. It can also be distinguished from R. ohausi by having gaping chelal fingers (versus not gaping in R. ohausi, see Mahnert, 2001, Fig. 110). Rhopalochernes catalinae sp. nov. most closely resembles R. insulanus and R. luiscarlosi sp. nov. in the absence of eyes (Figs. 4C, 4F), but can be differentiated from R. insulanus by the number of setae on the cheliceral hand, 6 in R. insulanus versus 4 in R. catalinae sp. nov. and R. luiscarlosi sp. nov., a trait also found in R. chamberlini and R. panamensis (see Heurtault, 1998, Figs. 4 and 9). Rhopalochernes catalinae sp. nov. can also be distinguished from R. luiscarlosi sp. nov. by the shape of the furrows on the surface of the carapace, which are shallow and inconspicuous in R. catalinae sp. nov. (Figs. 4C, 4F), but are deep and conspicuous in R. luiscarlosi sp. nov. (Figs. 1C, 1F). They also differ in other features such as: the shape of the spermathecae, with short and thicker ducts (Fig. 7B) versus elongated and narrow ducts in R. luiscarlosi sp. nov. (Fig. 7A); the shape of tarsus IV, which is robust (2.75–3.75 × longer than deep) while that of R. luiscarlosi sp. nov. is slender (4.00–4.25 × longer than deep). Description of adult: Colour: Carapace reddish brown towards the anterior margin and lighter towards posterior margin, pedipalps reddish brown. Tergites and legs yellowish brown (Fig. 4). Carapace: heavily granulated; 1.24‒1.41 (♂) 1.24‒1.37 (♀) × longer than broad, eyes or eye‒spot absent; with 2 inconspicuous transverse furrows slightly visible on lateral view, posterior furrow closer to posterior margin than anterior furrow; posterior margin of carapace slightly v shaped; anterior margin with 6 setae (♂, ♀), posterior margin with 8‒9 setae (♂, ♀). (Fig. 4C, F) Chelicera: 1.43‒2.00 (♂), 1.33‒1.80 (♀) × longer than broad; hand with 4 setae (sbs absent) and 1 distal seta (gs) on movable finger, is, bs and es short and acuminate, ls longer and broad (Figs. 5E, 6E), galea with 4 rami (♀) or simple (♂) (Figs. 5C, 6D); rallum with 3 blades (Figs. 5D, 6C), anterior blade with several ramifications while others smooth; serrula exterior with 11‒12 blades (♂, ♀), lamina exterior present. Pedipalp: trochanter, femur, patella and hand heavily granulated, with short, feathered and acuminate setae (Figs. 5F, 6F), chelal fingers smooth; trochanter 1.20‒1.57 (♂), 1.20‒1.64 (♀), femur 2.07‒2.77 (♂), 2.43‒3.08 (♀), patella 1.62‒2.36 (♂), 1.93‒2.43 (♀), chela (with pedicel) 2.94‒3.27 (♂), 2.89‒3.24 (♀), chela (without pedicel) 2.40‒2.53 (♂), 2.45‒2.63 (♀). Hand 1.24‒1.40 (♂), 1.25‒1.41 (♀) × longer than broad, movable finger 1.10‒1.24 (♂), 1.09‒1.27 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, accessory teeth not visible. Fixed chelal finger with 8 trichobothria. movable chelal finger with 4 trichobothria: eb, esb and est in the first half of fixed finger, esb closer to eb than est, which is closer to the midpoint of the finger, et distal and close to the fingertip; ib at the same level of eb, ist slightly aligned to esb, isb at the same level as st, close to est, it closer to et than isb; b and sb grouped basally, st and t in the middle of the movable finger, st slightly closer to t than sb, t much closer to st than tip of the finger. Venom apparatus present only in movable chelal finger with nodus ramosus at the same level or slightly close to st (Figs. 5G, 6G). Coxal region: maxillae smooth except for the antero-lateral region, coxae smooth, manducatory process triangular in shape with 2-3 apical setae, 1 prolateral suboral seta and 14-15 (♂), 11 (♀) additional setae on maxilla. Median maxillary lyrifissure rounded and situated submedially. Chaetotaxy of coxae I-IV, ♂, 7-8: 8-10: 8: 12; ♀, 9: 9: 12: 24. Legs: granulate, feathered setae on retrolateral margin and dentate setae on prolateral margins. Femur + patella of leg IV 2.25‒2.70 (♂), 2.60‒2.89 (♀) × longer than depth, as well as tibia and tarsus 2.71‒3.33 (♂), 3.17‒3.67 (♀) and 2.75‒3.75 (♂), 3.25‒3.75 (♀) × longer than broad, respectively. Tarsi III and IV with tactile seta, located subdistally. Arolium undivided, slightly shorter than claws, claws slender and simple, not modified (Figs. 5A‒B, 6A‒B). Abdomen: tergites I-X and sternites IV-X with a clear medial suture line (Fig. 7D). Tergal chaetotaxy: ♂, 12: 11-12: 11-13:15-16: 14-15: 15-16: 15-16: 14-16: 13-14: 12-13: 9-10: 2, ♀, 12: 14: 13: 16: 17: 18: 19: 16: 16: 13: 9: 2. Sternal chaetotaxy: ♂, (1) 4-6 (1): (1) 7-9 (1): 13-14: 12-13: 13: 13: 12: 10: 4-6: 2, ♀, 20: (1) 5 (1): (1) 15 (1): 15: 15: 14: 14: 14: 9: 16: 2. Tergites with setae acuminate, sternites with setae leaf-like shape. Pleural membrane striated and without setae. Genitalia: female: spermathecae paired, with short ducts ending in oval sacs (Fig. 7A). Male: genital opercula containing 18 setae on the anterior operculum and 4 in distal margin of posterior operculum. Dimensions: holotype female (ICN APs-843), with a range of 5 females paratype in parentheses: Body length 1.4 (1.04‒1.50). Pedipalps: trochanter 0.21/0.14 (0.21‒0.23/0.13‒0.15), femur 0.33/0.13 (0.33‒0.37/0.12‒0.14), patella 0.27/0.14 (0.27‒0.34/0.14‒0.15), chela (with pedicel) 0.51/0.16 (0.51‒0.55/0.16‒0.18), chela (without pedicel) 0.47 (0.46‒0.50), hand (without pedicel) length 0.2 (0.20‒0.24), movable finger length 0.25 (0.25‒0.28). Carapace 0.45/0.36 (0.45‒0.49/0.36‒0.39). Leg IV: femur + patella 0.25/0.09 (0.25‒0.27/0.09‒0.1), tibia 0.19/0.06 (0.19‒ 0.22/0.06), tarsus 0.14/0.04 (0.13‒0.15/0.04). Paratype male (ICN APs-844), with a range of 7 male in parentheses: Body length 1.34 (1.11‒1.35). Pedipalps: trochanter 0.21/0.14 (0.18‒0.22/0.14‒0.15), femur 0.36/0.13 (0.29‒0.38/0.13‒0.16), patella 0.33/0.15 (0.21‒ 0.34/0.13‒0.16), chela (with pedicel) 0.51/0.17 (0.48‒0.55/0.15‒0.18), chela (without pedicel) 0.46 (0.43‒0.49), hand (without pedicel) length 0.21 (0.21‒0.23), movable finger length 0.26 (0.23‒0.27). Carapace 0.47/0.35 (0.46‒ 0.48/0.34‒0.37). Leg IV: femur + patella 0.26/0.11 (0.25‒0.27/0.10‒0.12), tibia 0.19/0.07 (0.17‒0.22/0.06‒0.07), tarsus 0.14/0.04 (0.11‒0.15/0.04)., Published as part of Marimon, Karla, Blanco, Eduardo Villarreal & Harvey, Rk. S., 2022, Two new species of Rhopalochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) from Colombia, pp. 397-410 in Zootaxa 5150 (3) on pages 403-407, DOI: 10.11646/zootaxa.5150.3.5, http://zenodo.org/record/6623085, {"references":["Heurtault, J. (1998) Pseudoscorpions of the genus Rhopalochernes (Chernetidae) from Panama and Venezuela. Journal of Arachnology, 26, 442 - 446."]}

Marimon, Karla, Blanco, Eduardo Villarreal, Harvey, Rk. S. (2022): Two new species of Rhopalochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) from Colombia. Zootaxa 5150 (3): 397-410, DOI: https://doi.org/10.11646/zootaxa.5150.3.5

Waste heaps of Atta are host to an extraordinary diversity of myrmecophiles insects and other arthropods. In this study, the presence of four species of pseudoscorpions is recorded in two Atta colombica waste heaps in the years 2016 and 2017. Two of these species, Cordylochernes scorpioides and Lustrochernes carolinensis are new records in waste heaps and except for the deutonymphs of L.carolinensis, the others all stages of post-embryonic development were present in the studied heaps, which could indicate that these two species carry out their entire life cycle in these waste mounds and live there permanently.

The pseudoscorpion genus Nudochernes Beier, 1935 (Chernetidae) is reported from Iran for the first time, and a new species, Nudochernes limusensis sp. n. is described on the basis of two females co...

Genus Nudochernes Beier, 1935 Nudochernes Beier, 1935: 122; Beier, 1959: 48; Harvey, 1991: 607. Type species: Nudochernes montanus Beier, 1935, by original designation. Key to the Chinese species of Nudochernes 1. Pedipalpal femur 3.20 times longer than broad............................................ N. lipsae Mahnert, 2003 - Pedipalpal femur 2.40���2.90 times longer than broad.......................................................... 2 2. Trichobothria st closer to t than to sb; pedipalpal chela (with pedicel) 3.40���3.80 times longer than broad............................................................................................... N. troglobius Mahnert, 2009 - Trichobothria s t situated midway between sb and t; pedipalp chela (with pedicel) 3.10 (♂) 2.62���2.68 (♀) times longer than broad.......................................................................... N. pseudptroglobius sp. nov., Published as part of Xu, Hongru, Gao, Zhizhong & Zhang, Feng, 2022, Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 from Guizhou, China (Pseudoscorpiones: Chernetidae), pp. 581-592 in Zootaxa 5105 (4) on pages 582-587, DOI: 10.11646/zootaxa.5105.4.7, http://zenodo.org/record/6333898, {"references":["Mahnert, V. (2003) Four new species of pseudoscorpions (Arachnida, Pseudoscorpiones: Neobisiidae, Chernetidae) from caves in Yunnan Province, China. Revue suisse de Zoologie, 110 (4), 739 - 748. https: // doi. org / 10.5962 / bhl. part. 80209","Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492"]}

Nudochernes pseudotroglobius sp. nov. (Figs 1–4) http://zoobank.org/NomenclaturalActs/ 20C7BD03-E125-4379-8D91-A3111A273DDF Type material. Holotype ♀ (Ps.- MHBU-GZXS202401): China, Guizhou Province, Xishui County, Daozuo Cave [28.511389°N, 107.153333°E], elev. 1601m a.s.l., 31 August 2020, Zegang Feng & Yanmeng Hou leg. Paratypes: 1♀ (Ps.- SWUC-GZXS202402), 1♂ (Ps.- MHBU-GZXS202403), same data as holotype. Etymology. The name refers to its similar habitat with Nudochernes troglobius Mahnert, 2009. Diagnosis. This species can be distinguished from other Nudochernes species by the presence of dentate setae on carapace and pedipalp, spermathecae with a long unpaired tube and two short apical tubules, st situated midway between sb and t, the presence of elongate setae on the posterior corners of coxae IV, and the tarsus IV with a tactile seta in middle of tarsus (TS=0.55 (♂) 0.57 (♀)). Description. (Fig. 2). Males smaller than females with 2.77 (♂) 3.11–3.16 (♀), carapace and pedipalps reddish brown, chelicera, sternites, tergites and legs yellowish brown. Carapace (Fig. 3A): evenly granulate; 1.12 (♂) 0.97–0.98 (♀) times longer than broad; eyes or eye-spots absent; with two distinct deep transverse furrows; subbasal transverse furrow situated closer to the posterior margin than to the median furrow; posterior area with a small longitudinal furrow; including 16 (♂) 10 (♀) setae on anterior margin and 15 (♂) 16 (♀) setae on posterior margin. Abdomen: all tergites and sternites narrowly divided (except the last one). Tergal chaetotaxy (I–XI): ♂, 7–7: 10–8: 8–7: 9–9: 8–8: 9–8: 9–9: 9–7: 7–7: 6–9: 11(2T); ♀, 7–7: 9–10: 9–8: 11–10: 10–10: 9–10: 9–10: 8–9: 7–8: 7–7: 10(2T); all setae clavate. Sternal chaetotaxy (IV–XI): ♂, 7–9: 11–11: 11–10: 9–10: 9–8: 8–8: 8–7: 10; ♀, 4–4: 6–6: 12–13: 13–12: 12–13: 13–13: 9–8: 11; all setae acuminate. Sternite II of ♀ with 18 setae arranged as an inverted “U” pattern (Figs 3E, 4E). Chelicerae (Figs 3D, 4D): 5 setae on hand and 1 subdistal seta on movable finger; sbs, bs, es dentate apically, ls, is, gs acuminate; fixed finger with 6 proximad-directed teeth, movable finger with broad tooth-like subapical lobe; galea with 6 long branchlets in distal part; rallum with 3 dentate blades (Fig. 4C); serrula exterior with 20 (♂, ♀) blades. Pedipalp (Figs 3B, 4A): pedipalp robust; trochanter, femur, patella and chelal hand coarsely granulate, with dentate setae; chelal fingers with short setae; trochanter 2.24 (♂) 1.50–1.62 (♀), femur 2.24 (♂) 2.33–2.61 (♀), patella 2.30 (♂) 2.12–2.38 (♀), chela (without pedicel) 2.71 (♂) 2.33–2.47 (♀), hand (without pedicel) 1.29 (♂) 1.21–1.26 (♀) times longer than broad, movable finger 1.27 (♂) 1.12–1.28 (♀) times as long as hand (without pedicel). Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria (Figs 3C, 4F): eb situated basally; est situated midway between esb and et; ib and esb situated sub-basally, it situated closer to tip of finger than to ib; s t situated midway between sb and t, sb situated closer to b than to s t, and t situated midway between tip of finger and st; fixed finger with 49 teeth, plus 5 prolateral accessory teeth and 5 retrolateral accessory teeth; movable finger with 47 teeth, plus 6 prolateral accessory teeth and 5 retrolateral accessory teeth. Venom apparatus only present in movable chelal finger, venom duct slender, nodus ramosus closer to t than to st. Coxae: with numerous short setae over entire ventral surface, with longer setae on posterior margin of coxae IV. Genitalia (Fig. 4B): ♂: typical of Chernetidae; ♀: spermathecae with a long unpaired tube and two short apical tubules. Legs I (Figs 3F, 4G): trochanter, femur, patella, tibia with numerous clavate setae; tarsus with numerous acuminate setae; proportions: trochanter 1.09 (♂) 1.20–1.31 (♀), femur 1.46 (♂) 1.51–1.84 (♀), patella 2.07 (♂) 2.09– 2.41 (♀), tibia 3.86 (♂) 3.33–3.70 (♀), tarsus 3.98 (♂) 4.30–5.01 (♀) times longer than wide; claws simple, arolium shorter than claws. Legs IV (Figs 3H, 4H): femur, patella, tibia with numerous clavate setae; trochanter and tarsus with numerous acuminate setae; proportions: trochanter 1.69 (♂) 1.65–1.77 (♀), femur + patella 5.50 (♂) 5.46–5.50 (♀), tibia 4.24 (♂) 4.58–5.10 (♀), tarsus 4.32 (♂) 4.20–4.87 (♀) times longer than deep, a tactile seta near middle of tarsus (Figs 3I, 4I) (TS=0.55 (♂) 0.57 (♀)); claws simple, arolium shorter than claws. Dimensions (length/breadth or depth, in mm). ♀: Total length 3.11–3.16. Carapace 0.70–0.76/0.71–0.78. Pedipalps: trochanter 0.41–0.42/0.25–0.31, femur 0.64–0.68/0.25–0.27, patella 0.64–0.68/0.25–0.29, chela (with pedicel) 1.14–1.31/0.36–0.42, chela (without pedicel) length 1.02–1.18, hand (without pedicel) length 0.48–0.53, movable finger length 0.54–0.68. Leg I: trochanter 0.17–0.18/0.13–0.15, femur 0.21–0.24/0.13–0.14, patella 0.23– 0.29/0.11–0.12, tibia 0.30–0.37/0.09–0.10, tarsus 0.30–0.35/0.07. Leg IV: trochanter 0.28–0.32/0.17–0.18, femur + patella 0.71/0.13–0.15, tibia 0.51–0.55/0.10–0.12, tarsus 0.39–0.42/0.08–0.10, TS=0.57. ♂: Total length 2.77. Carapace 0.72/0.64. Pedipalps: trochanter 0.40/0.22, femur 0.63/0.28, patella 0.65/0.28, chela (with pedicel) 1.15/0.37, chela (without pedicel) length 1.01, hand (without pedicel) length 0.48, movable finger length 0.61. Leg I: trochanter 0.15/0.14, femur 0.21/0.14, patella 0.25/0.12, tibia 0.33/0.09, tarsus 0.30/0.08. Leg IV: trochanter 0.26/0.15, femur + patella 0.59/0.11, tibia 0.42/0.10, tarsus 0.38/0.09, TS=0.55. Remarks. This new species is close related to N. troglobius Mahnert, 2009 (Hubei, China), but can be distinguished by the stouter pedipalp (chela (with pedicel) 3.10 (♂) 2.62–2.68 (♀) times longer than board versus 3.40–3.80 (♂, ♀) in N. troglobius) and the position of trichobothrium (s t situated midway between sb and t in N. pseudotroglobius sp. nov.; st closer to t than to sb in N. troglobius) (Mahnert 2009). This new species can be distinguished from N. lipsae Mahnert, 2003 (Yunnan, China) by the stouter legs I (patella 2.07 (♂) 2.09–2.41 (♀) times longer than deep vs 3.50 (♀) in N. lipsae; tibia 3.86 (♂) 3.33–3.70 (♀) times longer than deep vs 5.00 (♀) in N. lipsae), and by the position of trichobothrium (s t situated midway between sb and t in N. pseudotroglobius sp. nov., whereas s t closer to t than to sb in N. lipsae) (Mahnert 2003). Geographically, only one species, N. spalacis Beier, 1955, is described in Asia (Israel). The new species can be distinguished from N. spalacis by having slender legs IV (femur + patella 5.50 (♂) 5.46–5.50 (♀) times longer than deep versus 3.70 (♂) in N. spalacis) and the different position of trichobothrium (est situated midway between et and esb in N. pseudotroglobius sp. nov., whereas est closer to et than to esb in N. spalacis) (Beier 1955). Distribution. Known only from the type locality in Guizhou, China. Habitat. All specimens were collected under a rock within the dark zone of the cave.

Megachernes biyunensis sp. nov. (Figs 5–7) http://zoobank.org/NomenclaturalActs/31be2f78-81ae-4bb7-aa18-076eb4722f89 Type material. Holotype ♀ (Ps.- MHBU-GZ19080501): China, Guizhou Province, Pan County, Biyun Park [25.775000°N, 104.638333°E], 5 August 2019, Zegang Feng & Zhaoyi Li leg. Paratypes: 1♀ (Ps.-SWUC-GZ19080502); 1♂ (Ps.- MHBU-GZ19080503), same data as holotype. Etymology. The specific name refers to the type locality, Biyun Park, Pan County, Guizhou Province, China. Diagnosis. This new species can be recognized by the following combined characters: subbasal transverse furrow on carapace located midway between posterior margin and central furrow, the trichobothria st situated closer to sb than to t, t situated midway between tip of finger and st. Description. (Fig. 5). Carapace, chelicera, tergites and sternites yellowish brown, pedipalps reddish brown, colour of chelal hand darker than that of trochanter, femur and patella; soft parts pale. Carapace (Fig. 6A): evenly granulate; 1.07 (♂) 1.02 (♀) times longer than broad; eyes or eye-spots absent; with two distinct deep transverse furrows, subbasal transverse furrow situated midway between posterior margin and central furrow; posterior area with a small longitudinal furrow; with numerous setae including 16 (♂) 20 (♀) acuminate setae on anterior margin and 10 (♂) 14 (♀) on posterior margin. Abdomen: all tergites and sternites narrowly divided. Tergal chaetotaxy (I–XI): ♂, 7–6: 10–8: 7–8: 7–8: 7–8: 9– 8: 9–9: 7–9: 8–6: 7–8: 6(2 T)–7(2T); ♀, 7–7: 9–8: 8–8: 9–9: 9–8: 8–9: 8–9: 8–9: 9–7: 7–8: 7(2 T)–7(2T); I–X tergites with paired dark spot. Sternal chaetotaxy (IV–XI): ♂, 4–5: 8–9: 13–11: 13–12: 10–11: 11–19: 11–9: 6(2 T)–7(2T); ♀, 4–3: 10–12: 14–12: 14–14: 13–14: 12–14: 12–12: 6(2 T)–6(2 T); all sternal setae acuminate. Sternite II of ♀ with 33 setae arranged in a cluster (Figs. 6E, 7F). Chelicerae (Figs. 6D, 7D): 7 setae on hand and 1 subdistal seta on movable finger; basal 5 setae finely dentate apically, ls, is and gs acuminate; fixed finger with 3 proximad-directed teeth; galea comprising 2 parts, first part with 5 branches and the second with 4 branches (Fig. 7B). Serrula exterior with 27 (♂, ♀) blades; rallum with 3 dentate blades, the apex of the first one slightly twisted and with anterior margin dentate, the other two blades apically dentate (Fig. 7C). Pedipalp (Figs 6B, 7A): trochanter, femur, patella and chelal hand coarsely granulate, with acuminate setae; chelal fingers with short setae; trochanter 1.39 (♂) 1.67 (♀), femur 2.51 (♂) 2.31 (♀), patella 2.26 (♂) 2.23 (♀), chela (without pedicel) 3.02 (♂) 2.81 (♀), hand (without pedicel) 1.58 (♂) 1.51 (♀) times longer than board, movable finger 0.91 (♂) 0.96 (♀) times as long as hand (without pedicel). Fixed finger with 8 trichobothria, movable finger with 4 trichobothria (Figs 6C, 7E): eb situated basally; est situated closer to esb than to et; ib and esb situated sub-basally; it situated closer to tip of finger than to ib; st situated closer to sb than to t, sb situated closer to s t than to b, and t situated midway between tip of finger and st, fixed finger with 68 teeth, plus 13 prolateral accessory teeth, and 8 retrolateral accessory teeth; movable finger with 59 teeth, plus 12 prolateral accessory teeth, and 7 retrolateral accessory teeth. Venom apparatus only present in movable chelal finger, venom duct slender, nodus ramosus closer to t than to st. Coxae: with numerous short setae over entire ventral surface and with long setae on posterior margin of coxae IV; posterior corner of coxae IV slightly protruding. Genitalia (Figs 6E, 7H): ♂: typical of Chernetidae; ♀: spermathecae T-shaped and with extremely elongated ends. Leg I (Figs 6F, 7G): with long acuminate setae; proportions: trochanter 1.11 (♂, ♀), femur 1.50 (♂) 1.51 (♀), patella 2.60 (♂) 2.17 (♀), tibia3.71 (♂) 3.50 (♀), tarsus 4.41 (♂) 4.10 (♀) times longer than deep; claws simple, arolium shorter than claws. Legs IV (Figs 6H, 7I): with long acuminate setae; proportions: trochanter 1.76 (♂) 1.44 (♀), femur + patella 3.96 (♂) 3.43 (♀), tibia 5.01 (♂) 4.71 (♀), tarsus 3.80 (♂) 3.83 (♀) times longer than deep; tactile seta in middle tarsus (TS=0.51 (♂) 0.54 (♀)), length of tactile seta 2/3 of tarsus (Fig. 7J); claws simple, arolium shorter than claws. Dimensions: (length/breadth or depth, in mm). ♀: Total length 4.22. Carapace 1.20/1.18. Pedipalps: trochanter 0.65/0.39, femur 0.99/0.43, patella 1.07/0.48, chela (with pedicel) 1.95, chela (without pedicel) 1.80, hand (without pedicel) length 0.97, movable finger length 0.94. Leg I: trochanter 0.21/0.18, femur 0.30/0.21, patella 0.39/0.18, tibia 0.49/0.14, tarsus 0.41/0.10. Leg IV: trochanter 0.36/0.25, femur + patella 0.79/0.23, tibia 0.80/0.17, tarsus 0.46/0.12, TS=0.54. ♂: Total length 4.10. Carapace 1.14/1.05. Pedipalps: trochanter 0.59/0.43, femur 1.05/0.41, patella 1.06/0.47, chela (with pedicel) 1.93/0.64, chela (without pedicel) length 1.87, hand (without pedicel) length 1.01, movable finger length 0.92. Leg I: trochanter 0.22/0.20, femur 0.36/0.24, patella 0.52/0.2, tibia 0.51/0.14, tarsus 0.48/0.11. Leg IV: trochanter 0.36/0.21, femur + patella 1.06/0.27, tibia 0.81/0.16, tarsus 0.57/0.15, TS=0.51. Remarks. Based on the key of Megachernes species (Mahrad & Reza 2017), this new species is similar to M. limatus Hoff & Parrack, 1958 from Oceania, but can be distinguished by the character of carapace (the subbasal transverse furrow situated closer to the posterior margin than to the median furrow in M. limatus, whereas the subbasal transverse furrow situated midway in M. biyunensis sp. nov.); and by the different trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. limatus) (Hoff & Parrack 1958). It can be distinguished from M. himalayensis (Ellingsen, 1914) (China (Fujian); India; Nepal) by the number of accessory teeth on the chelal fingers (movable finger with 12 accessory teeth and fixed finger with 13 accessory teeth in M. biyunensis sp. nov., whereas movable finger with 6 accessory teeth and fixed finger with 9 accessory teeth in M. himalayensis); and by the stouter legs IV (tibia 5.01 (♂) 4.71 (♀) times longer than deep versus 6.20 (♀) in M. himalayensis; tarsus 3.80 (♂) 3.83 (♀) times longer than deep vs 5.00 (♀) in M. himalayensis) (Ellingsen 1914). It can be distinguished from M. titanius Beier, 1951 (China (Sichuan, Guangxi); Southeast Asia) by the trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. titanius); and by the stouter legs IV (5.01 (♂) 4.71 (♀) times longer than wide versus 6.20 (♀) in M. titanius; tarsus 3.80 (♂) 3.83 (♀) times longer than deep versus 5.20 (♀) in M. titanius) (Beier 1951). It differs from M. tuberosus Mahnert, 2009 (Sichuan, China) by different the trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. tuberosus); and by the morphology of the rallum (all three blades denticulate in M. biyunensis sp. nov., blades terminally scarcely dentate and apex of first blade slightly twisted in M. tuberosus) (Mahnert 2009). It differs from M. vietnamensis Beier, 1967 (China (Sichuan, Hubei, Yunnan); Vietnam) by the number of accessory teeth on the chelal fingers (movable finger with 12 accessory teeth and fixed finger with 13 accessory teeth in M. biyunensis sp. nov., whereas fixed finger with 21 accessory teeth in M. vietnamensis); and by the cheliceral hand setae (basal 5 setae with terminally dentateand acuminate in M. biyunensis sp. nov., whereas all setae with acuminate in M. vietnamensis) (Beier 1967). Distribution. Known only from the type locality, Guizhou, China. Habitat. All specimens were collected in wet leaf litter under Populus (Salicaceae).

Genus Megachernes Beier, 1932 Megachernes Beier, 1932: 128; Beier, 1933: 518; Beier, 1948: 476; Morikawa, 1960: 144; Harvey, 1991: 599. Type species: Chernes grandis Beier, 1930, by original designation. Key to the Chinese species of Megachernes 1. Leg IV femur + patella with numerous glandular microsetae........................... M. glandulosus Mahnert, 2009 - Leg IV femur + patella without glandular microsetae......................................................... 2 2. Male pedipalpal patella with hump................................................. M. tuberosus Mahnert, 2009 - Male pedipalpal patella without hump..................................................................... 3 3. Pedipalpal femur less than 4.00 times longer than board....................................................... 4 - Pedipalpal femur more than 4.00 times longer than board..................................... M. titanius Beier, 1951 4. Leg IV femur + patella more than 4.00 times longer than deep.................................................. 5 - Leg IV femur + patella less than 4.00 times longer than deep................................................... 6 5. Movable chelal finger with 9 retrolateral accessory teeth; all blades of rallum smooth..... M. himalayensis (Ellingsen, 1914) - Movable chelal finger with 21 retrolateral accessory teeth; only distal blade of rallum denticulate.................................................................................................... M. vietnamensis Beier, 1967 6. Only distal blade of rallum denticulate; st situated closer to t than to sb........................ M. grandis (Beier, 1930) - All blades of rallum denticulate; st situated closer to sb than to t................................ M. biyunensis sp. nov., Published as part of Xu, Hongru, Gao, Zhizhong & Zhang, Feng, 2022, Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 from Guizhou, China (Pseudoscorpiones: Chernetidae), pp. 581-592 in Zootaxa 5105 (4) on pages 587-591, DOI: 10.11646/zootaxa.5105.4.7, http://zenodo.org/record/6333898, {"references":["Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i - xxi + 1 - 294. https: // doi. org / 10.1515 / 9783111385402","Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492","Beier, M. (1951) Die Pseudoscorpione Indochinas. Memoires du Museum National d'Histoire Naturelle, Paris, Nouvelle Serie, 147 - 123.","Beier, M. (1967) Pseudoscorpione vom kontinentalen Sudost-Asien. Pacific Insects, 9, 341 - 369."]}

Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 are described from Guizhou Province, China, Nudochernes pseudotroglobius sp. nov. and Megachernes biyunensis sp. nov., with detailed diagnoses, descriptions and illustrations.

We herein report for the first time the phoretic association between the pseudoscorpion Cordylochernes scorpioides (Linnaeus, 1758) (Chernetidae) and the longhorn beetle Hylettus coenobita (Erichson, 1847) (Cerambycidae) in the central Amazon. In addition, this is the first record of pseudoscorpion phoresy for the genus Hylettus Bates, 1864.

The pseudoscorpion Chernes similis (Beier, 1932) is reported from the Baltic region for the first time. The new recordfrom Lithuania is based on a single male specimen found in a mould growing in a hollow of a Tilia L. tree in theBotanical Garden of Vilnius University. This finding represents the northernmost record of C. similis. With the newrecord of C. similis, nine pseudoscorpion species, belonging to six genera and three families, are currently known fromLithuania.

Pachychernes zehorum Muchmore, 1997 Figs. 4A–D, 5A–G, 6A–G, 7B Pachychernes zehorum Muchmore, 1997: 19–21, figs 1–4; Ceballos, 2004: 429; Villegas-Guzman, 2006: 133; Harvey 2013. Diagnosis. This species differs from all other Pachychernes species by the first leg of the males, which not only has long and short modified setae on the tibia, but also a conspicuous depression on the dorsal side of the tarsus (Muchmore 1997) (Fig. 4). Examined material. COLOMBIA, Bolívar, Municipio de San Jacinto, La Flecha, Finca Amanecer Gaitero (9°51'9.74" N; 75°10'32.32" W), 250 m, 4 ♀ (ICN-Aps-835) and 2 ♂ (IAvH-I-4079), 11–12.IX.2019, K. Marimon and E. Villarreal leg. Description (adults): Colour: carapace red-brown with a lighter tone below the transversal furrow. Tergites light brown, pedipalps reddish-brown. Legs light yellow-brown toned. Carapace: heavily granulated, subrectangular with broad posterior margin; 0.78–0.93 (♂) 0.86–1.13 (♀) × longer than broad, with two eye spots; anterior margin with 8 setae (♂) 7 setae (♀), posterior margin with 6 setae (♂, ♀). Chelicera: 2.13–2.54 (♂), 1.44–2.40 (♀) × longer than broad; with 5 setae on hand and 1 distal seta on movable finger, es, is and ls acuminate, bs and sbs dentate; galea with 5 rami; rallum with 3 blades, posterior blade with several ramifications while others smooth; serrula exterior with 24 blades (♂, ♀); lamina exterior present (Figs. 5, 6). Pedipalp: with robust trochanter, femur, patella and granulated hand with several denticulated setae and chelal fingers smooth; trochanter 1.30–1.36 (♂), 1.27–1.39 (♀), femur 2.70–2.92 (♂), 2.10–2.42 (♀), patella 2.38–2.50 (♂), 2.12–2.37 (♀), chela (with pedicel) 2.81–3.05 (♂), 2.61–2.89 (♀), chela (without pedicel) 2.63–2.86 (♂), 2.61–2.89 (♀), hand 1.53–1.63 (♂), 1.54–1.68 (♀) × longer than broad, movable finger 0.76–0.77 (♂), 0.59–0.86 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, external accessory teeth present on both fingers, with 6 (♂), 7 (♀) on the movable finger and 9 (♂), 11(♀) on the fixed finger. Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria: eb, esb, est and et slightly L-shaped, eb and esb situated basally, est located medially and at an equal distance from esb and et, which is located terminally; ib and ist basally located, with ist, eb and esb aligned, isb located in a medial position and closer to it and ib; b, sb and st placed in the first half of the movable finger with b and sb much closer to each other than to st, t located subterminally, midway from st and tip of finger. Venom apparatus present only in movable chelal finger with nodus ramosus located next to t (Figs. 5, 6). Coxal region: maxillae smooth except for the antero-lateral region, coxae smooth, manducatory process triangular in shape; regarding the setae, these individuals have one apical, one subapical and one sub oral while there are 25 additional setae (♂). Maxillary lyrifissure rounded and placed submedially. Chaetotaxy of coxae I–IV in ♂: 14: 12: 16: 44; ♀: 13: 11: 15: 36. Legs: leg I sexually dimorphic, with males having a tibia with a large number of setae plus a distinct depression on the surface of the tarsus while that of the females are chernetid in structure. Femur + patella of leg IV 2.67–2.77 (♂), 3.56–4.17 (♀) × longer than broad, as well as tibia and tarsus 3.79–4.11 (♂), 3.56–4.17 (♀) and 4.27–4.64 (♂), 3.89–4.67 (♀) × longer than broad, respectively. Tarsi III and IV with tactile seta, located subbasally, TS= 0.18 in both males and females approximately. Claws and arolium are equal in length. Abdomen: tergites I–X and sternites I–X with a clear medial suture line. Tergal chaetotaxy: ♂, 8: 7: 9: 8: 11: 13: 12: 12: 13: 12: 11: 2; ♀, 10: 10: 10: 10: 12: 12: 12: 13: 14: 13: 11: 2, all setae clavate. Sternal chaetotaxy: ♂, 47, (3) 16 (3): (1) 8 (1): 17: 19: 19: 23: 20: 17: 15: 2; ♀, 38: (2) 8 (2): (1) 9 (1): 20: 19: 18: 18: 18: 17: 16: 4. Spiracle with helix. Pleural membrane striated and without setae. Genitalia: spermatheca somewhat H-shaped, with four lobes, the anterior ones, oval in shape while the posterior ones have a tubular form with several cribriform plates along the ducts (Fig. 7B). Male genitalia consisting of 47 setae on the anterior operculum and 16 on the posterior operculum. Dimensions (mm): males, followed by females in parentheses: Body length 3.05–3.10 (2.44–3.30). Carapace 0.86–1.09/1.10–1.17 (0.81–1.05/0.82–1.22) Chelicera 0.32–0.33/0.13–0.15 (0.23–0.31/0.10–0.16), movable finger length 0.23–0.28 (0.20–0.25. Pedipalps: trochanter 0.48–0.49/0.36–0.37 (0.33–0.46/0.26–0.36), femur 1.10–1.14/0.37–0.39 (0.61–0.87/0.29–0.39), patella 1.00–1.15/0.42–0.46 (0.72–0.97/0.34–0.45), chela (with pedicel) 1.74/0.57–0.62 (1.39–1.89/0.46–0.62), chela (without pedicel) 1.63 (1.31–1.79), hand (without pedicel) length 0.93–0.95 (0.71–1.00), movable finger length 0.72 (0.57–0.85). Leg IV: femur 0.35–0.37/0.27–0.30 (0.26– 0.32/0.19–0.25), patella 0.77–0.8/0.35–0.36 (0.51–0.73/0.20–0.33), tibia 0.72–0.78/0.19 (0.50–0.72/0.12–0.18) tarsus 0.47–0.51/0.11 (0.35–0.49/0.09–0.11). Distribution and habitat. This species has been previously recorded from Mexico and Panama (Muchmore 1997; Ceballos 2004; Villegas-Guzman 2006), and according to Muchmore (1997) it is probably widespread through Central America. Here we expand the distribution to South America, with a population found in the Montes de María in northern Colombia. The new records demonstrate that this species is not necessarily restricted to environments such as rain forest (type locality) but also occurs in tropical dry forest (Figs. 8, 9)., Published as part of Marimon, Karla, Villarreal-Blanco, Eduardo, Romero-Ortiz, Catalina & Gutierrez, Luis C., 2021, A new species and new records of Pachychernes Beier, 1932 from Colombia (Pseudoscorpiones, Chernetidae), pp. 363-376 in Zootaxa 4999 (4) on pages 369-375, DOI: 10.11646/zootaxa.4999.4.6, http://zenodo.org/record/5119308, {"references":["Muchmore, W. (1997) An unusual new Pachychernes from Panama and Mexico (Pseudoscorpionida: Chernetidae). Entomological News, 108, 19 - 23.","Harvey, M. (2013) Pseudoscorpions of the world. Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 25 February 2021)"]}

In Colombia, the status of the Neotropical genus Pachychernes Beier, 1932 is unknown. The only available data in literature to support the presence of this genus is an adult of P. aff. subrobustus, possibly female, from copal resin without specific locality data, and a record at generic level from Northern Colombia. In this study, we extend the distribution of Pachychernes by describing a new species, P. florezi sp. n. and several reporting individuals of P. zehorum Muchmore, 1997 from Montes de María and the Caribbean coast of the country. These species are clearly differentiated from one another by the sexual dimorphism in the latter species, with a depression on the tarsus I in males. We also present an updated key for all species of the genus.

Allochernes solarii (Simon, 1898) is recorded for the first time from Slovakia. Six adults and two tritonymphs were found in two nests of Formica gagates Latreille, 1798. A description of the species is provided based on Slovakian specimens and the holotype from Italy.

New cases of phoresy of pseudoscorpions (Pseudoscorpiones: Atemnidae, Chernetidae) are recorded from Slovakia. The phoresy of the species Atemnus politus (Simon, 1878) involving owlet moth (Noctuidae) as a host is documented for the first time not only in Slovakia, but also worldwide. One phoretic female of Lamprochernes chyzeri (Tomosvary, 1883) was attached to the leg of a lance fly (Lonchaeidae). It represents the second known phoresy of the species from Slovakia and the record of a new host.

Three pseudoscorpion species from the families Cheliferidae and Chernetidae, Hysterochelifer tuberculatus (Lucas, 1849), Allochernes bulgaricus Hadži, 1939 and Chernes hahnii (C. L. Koch, 1839), are new for the pseudoscorpion fauna of North Macedonia. North Macedonia is the second known country of the distribution of A. bulgaricus in Europe.