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Basing on the external morphological characters, an isolated population of the Arctic Apollo, Parnassius arcticus (Eisner, 1968) (Lepidoptera, Papilionidae), from the Arga-Tas Range mountains (North-Eastern Yakutia, Russia) is described here as a new subspecies, Parnassius arcticus shavlovi Yakovlev & Bakhaev, subsp. nov.

Parnassius arcticus shavlovi Yakovlev & Bakhaev, subsp. n. http://zoobank.org/ urn:lsid:zoobank.org:act: 4268E6A7-6D90-4958-B165-A9D59DCEBB6B Figs 9−19 Material examined. Holotype, male, Russia, North-Eastern Yakutia, Verkhnekolymsky District, Arga-Tas Range, uppers of Tuora-Bygyttakh river, H− 1100−1300 m, 20−28.vi. 2022, leg. Y. Bakhaev (Zoological Institute, Saint-Petersburg, Russia). Paratypes. 76 males, 19 females, the same locality and data (Zoological Institute, Saint-Petersburg, Russia; private collection of Roman V. Yakovlev, Barnaul and Yuri I. Bakhaev, Lipetsk). Description. Male. Wingspan of holotype 43 mm, paratypes – 34−46 mm. Antennae coal-black, shorter than half of fore wing in length. Head, thorax and abdomen covered with coal-black scales. Fore wing white from above, with dense sputtering of black scales basally, black scales sputtering also along costal margin from root to apex of wing, clearly expressed black stroke in centre of discal cell, black cuneal or round spot at top of discal cell, series of two-three (partially fused) black strokes between radial veins postdiscally, series of two-four small black strokes between radial cells postdiscally (closer to outer margin of wing), outer margin of wing (from apex to Cu 1) semi-transparent, with sputtering of black scales, fringe white. Fore wing from underside with completely the same pattern as on wing from upside. Hind wing from upside white, with vast black portion, occupying area from anal margin and wing base to middle of discal area and portion along Cu 2 vein, postdiscal area in cell Sc+R− Rs and cell M 1 −M 2 with tiny dark spots translucent from underside; in some males, black spot on wing upside only in cell Sc+R− Rs, very thin rim along wing margin from Rs to М 2 (rarely, M 3) (in almost half of specimens, rim torn into separate strokes), fringe white. Hind wing from underside with more contract pattern than from upside: postdiscally in cell Sc+R− Rs and cell M 1 −M 2 well developed small black spots (often developed in cells M 3 −Cu 1, Cu 1 −Cu 2 and Cu 2 −2A as well); in most specimens, tiny black spots developed submarginally in cells between medial and cubital veins, small black spots between medial veins along wing margin. Male genitalia (Fig. 17) is similar to the nominotypical P. arcticus. Uncus forked, short, with uncinate apices diverged to sides; valve cup-like, with even edges, apically semicircular; conical harpe on inner surface of valve, slightly protruding beyond the apex of valve; juxta oval, with pair of flask-shaped processes directed dorsally; saccus robust, conical; phallus thin, slightly curved along all length, apically needle-like, 1/3 longer than valve. Female. Wingspan 35−46 mm. Antennae coal-black, shorter than half of fore wing in length. Head, thorax and abdomen covered with coal-black scales. Fore wing from upside with intensive black sputtering at root, costal margin with black sputtering; in vast majority of specimens, black sputtering distributed throughout wing field, except for some light spots and bands; black spot of medium size in middle of discal cell, large black spot at top of discal cell; postdiscally from costal margin to M2 – band of fused black spots; postdiscally closer to outer margin – relatively wide light band (or light portion), submarginally – expressed thin (often, fragmentary) light band, outer margin of wing semitransparent, with intensive grey sputtering, fringe white. Fore wing from underside with the same pattern as from upside, but significantly lighter. Hind wing from upside often with very wide dark-grey root portion (sometimes occupying all root, discal and partially postdiscal areas), large black spots (round of shaped as wide longitudinal strokes) postdiscally in cells Sc+R− Rs, M 1 −M 2, M 3 −Cu 1 and Cu 1 −Cu 2; postdiscally (closer to outer margin of wing) – wide light band with small black strokes in the range of cells from Rs− M 1 to Cu 1 −Cu 2 (steadily developed in cells Rs− M 1 and M 1 −M 2, rarely – in all cells), outer margin dark-grey or black, fringe white. Hind wing from underside in general with the same pattern as on upside but slightly lighter and more contrasting (in root area, discal cell and cell Sc+R− Rs with developed long bright black longitudinal strokes). Diagnosis. The new subspecies clearly differs from the known subspecies in a series of characters. From P. arcticus arcticus: • the new subspecies on an average is larger in size: male wingspan is 34−46 mm, females – 35−46 mm (in P. arcticus arcticus: 32−41 mm and 37−40 mm, respectively); • in the males of the new subspecies, the black spots on the fore wing postdiscally and especially, submarginally, are significantly less developed (in P. arcticus arcticus, the rows of black spots are developed steadily); • in the males of the new subspecies, the black spot on the hind wing in cell M 1 −M 2 is absent steadily; additionally, in most specimens there is no black spot in cell Sc+R− Rs (in P. arcticus arcticus, the black spots of the submarginal row on the hind wing are well expressed); • along the outer margin of the hind wing in most males there are small black spots (in P. arcticus arcticus, the outer margin of the wing is white); • in the females of the new subspecies, the black sputtering on the wings from upside is much more expressed; the orange nucleoli in the black spots of the postdiscal row on the hind wing are very rare (in P. arcticus arcticus, the black sputtering on the wings from above is significantly less expressed; in most specimens, there are orange nucleoli in the black spots of the postdiscal row on the hind wing). From P. arcticus arbugaevi: • in the males of the new subspecies, the black spot is steadily present in the discal cell on the fore wing (in P. arcticus arbugaevi, the black spot in the discal cell on the fore wing is steadily absent); • in the males of the new subspecies, the black spot on the hind wing in cell M 1 −M 2 is steadily absent; additionally, in most specimens there is no black spot in cell Sc+R− Rs (in P. arcticus arbugaevi, the black spots of the submarginal row on the hind wing are clearly expressed); • in most of the males, along the outer margin of the hind wing there are small black spots (in P. arcticus arbugaevi, the outer margin of the wing is white); • in the females of the new subspecies, the black sputtering on the wings from upside is less expressed, the pattern of fore and hind wings is more contrasting, on the fore wing there are well developed postdiscal and submarginal light bands (in P. arcticus arbugaevi, the black sputtering on the wings from upside is significantly more intense, the pattern (especially, on the fore wing) is more blurred, not distinctive; there are no postdiscal and submarginal light bands on the fore wing). Bionomy. The adults were indicated in the middle of June on gravel screes at altitudes of 1100−1300 m (Figs 20−21). The adults were observed feeding on the flower of Smelowskia jacutica (Botsch. & Karav.) Al-Shehbaz & S.I. Warwick (Brassicaceae) (Fig. 22). The leaves damaged by the caterpillars of P. arcticus shavlovi were found on Corydalis gorodkovii Karav. (Papaveraceae) (Figs 23−24). Etymology. The subspecies is named after Roman Shavlov (Yakutsk) − an amateur zoologist well known in Yakutia, a friend on one of the authors of this article. Distribution. Known only from Arga-Tas Range in North-Eastern Yakutia. Discussion. It seems surprising that the new subspecies P. arcticus shavlovi was formed in the same mountain range as P. arcticus arbugaevi (Fig. 25). But there is a large distance between their places (more than 230 km), and most importantly, a biotopic gap (red line) (Fig. 26): a low-mountain zone. As it is known, P. arcticus lives only in the highlands (a zone of stony tundra at a height of more than 1100 m). So, the highmountain areas of the Arga-Tas Range are an isolate, which contributed to the formation of quite significant differences at the subspecies level over time.

Parnassius arcticus (Eisner, 1968) Tadumia simo arctica Eisner, 1968: 15 Type locality: den Bergen östlich von Werchnosensk [Verkhoyansk?, Yakutia, Russia]., Published as part of Yakovlev, Roman V. & Bakhaev, Yuri I., 2023, A new subspecies of Arctic Apollo - Parnassius arcticus (Eisner, 1968) (Lepidoptera, Papilionidae) from the Arga-Tas Range (North-Eastern Yakutia), pp. 1-14 in Ecologica Montenegrina 63 on page 1, DOI: 10.37828/em.2023.63.1, http://zenodo.org/record/8029884, {"references":["Eisner, C. (1968) Parnassiana Nova. XLIII. Nachtragliche Betrachtungen zu der Revision der Subfamilie Parnassiinae (Fortsetzung 16). Zoologische Mededelingen, 43 (2), 9 - 17."]}

Parnassius arcticus arbugaevi Yakovlev & Shapoval, 2020 Figs 5−8 Yakovlev et al. 2020: 99−103. Type locality: Russia, North-Eastern Yakutia, Momsky District, 70 km E of Khonuu village. Distribution: Momsky Range (North-Eastern Yakutia). The conspecificity of P. arcticus and P. ammosovi does not look completely convincing due to the fact that there is still no good comparative material from the type locality of P. arcticus. Also, the subspecies belonging of the specimen collected on Yablonovyi Pass in Magadan Territory (Gorbunov 2008) is not known., Published as part of Yakovlev, Roman V. & Bakhaev, Yuri I., 2023, A new subspecies of Arctic Apollo - Parnassius arcticus (Eisner, 1968) (Lepidoptera, Papilionidae) from the Arga-Tas Range (North-Eastern Yakutia), pp. 1-14 in Ecologica Montenegrina 63 on page 4, DOI: 10.37828/em.2023.63.1, http://zenodo.org/record/8029884, {"references":["Yakovlev, R. V., Shapoval, N. A., Bakhaev, Y. I., Kuftina, G. N., Khramov, B. A. (2020) A new subspecies of Parnassius arcticus (Eisner, 1968) from the Momsky Range (Yakutia, Russia). Acta Biologica Sibirica, 6, 93 - 105. https: // doi. org / 10.3897 / abs. 6. e 55925","Gorbunov, P. Y. (2008) 3. Apollon arctichesky. In: Chereshnev I. A. (Ed.) Red Data Book of Magadan Region. Administration of Magadan Region, Department of natural resources; Institute of Biological problems of North, Magadan, 30 - 31. [in Russian]."]}

Parnassius arcticus arcticus (Eisner, 1968) Figs 1−4 =? Parnassius ammosovi Korshunov, 1988: 69. Type locality: Russia, Yakutia, Suntar-Khayata Range, 180 km ENE from the Khandyga settlement, river Vostochnaya Khandyga upper flow, 232nd km of the road from Khandyga to Magadan. Distribution: Verkhoyansky and Suntar-Khayata Range (North-Eastern Yakutia)., Published as part of Yakovlev, Roman V. & Bakhaev, Yuri I., 2023, A new subspecies of Arctic Apollo - Parnassius arcticus (Eisner, 1968) (Lepidoptera, Papilionidae) from the Arga-Tas Range (North-Eastern Yakutia), pp. 1-14 in Ecologica Montenegrina 63 on pages 1-4, DOI: 10.37828/em.2023.63.1, http://zenodo.org/record/8029884, {"references":["Eisner, C. (1968) Parnassiana Nova. XLIII. Nachtragliche Betrachtungen zu der Revision der Subfamilie Parnassiinae (Fortsetzung 16). Zoologische Mededelingen, 43 (2), 9 - 17.","Korshunov, Y. P. (1988) New butterflies (Lepidoptera, Rhopalocera) from Khakassia, Tuva and Yakutia. In: Zolotarenko G. S. (Ed.) Taxonomy of animals of Siberia, Nauka, Novosibirsk, 65 - 80. [in Russian]."]}

Vavizola hela sp. n. https://zoobank.org/ urn:lsid:zoobank.org:act: 2883141C-35E7-487C-8365-D3C74CB66789 (Figs 1–10, 21–22, 28, 34) Holotype: ♂, Tanzania, Arusha National Park, Miriakamba Hut, -3.2294225707906707, 36.79956440000001, 2500 m, 20.X.2004, slide 17.446, LBEOW1233-11 (MWM/ZSM). Paratypes (2♂, 3♀). Tanzania: ♂, like holotype, slide 1267 (CGM). Kenya: ♂, Taita- Taveta, Voi, 19.XII.1911, leg. Svatosh (ZISP); ♀, Taita- Taveta, SW of Voi, 8–12.XII.2009, leg. Snižek, slide 17.447, LBEOW975-11 (CGM); ♂, Lamu, E of Garsen, W of Witu, 28.IV.2011, leg. Snižek, slide 20.690 (CGM); ♀, Kitui, 202 km E Thika, Sosoma, 4.XII.2010, leg. Snižek, slide 0501 (CGM); ♀, Kitui, E of Mwingi, W of Enguni, 1.XII.2010, leg. Snižek (CGM). Description. Male (Figs 1, 5–6, 8–9). Flagellum covered with creamy and brown scales, rami brown. Head and thorax speckled creamy and brown; thorax dorsally has longitudinal dark stripes. Abdomen brown or reddish brown. Forewing. Forewing length: 19–23 mm; wingspan: 38–45 mm. Elongated, apex blunt, outer margin with indentation on veins and concavity between M 2 and CuA 1. Background color speckled creamy with brown scales. Pattern diagonal (Fig. 1), consist of more or less pronounced discal stroke, speckled medial field, postmedial fascia with dark edges vanishing towards apex, postmedial field and external fascia practically merged into speckled field with dark lines vanishing towards apex as well, and dark external field. Cilia creamy on vines and dark brown between them. Hindwing. Somewhat ovaloid with some indentation on veins. Background color brown or dark brown with dark external field. Cilia speckled creamy and brown, darker between veins. Genitalia (Figs 3, 21–22). Tegumen a narrow band bearing a pair of socia laterally. Socii papilla-shaped, covered with chaetae. Vinculum ventrally elongated, distally bears cubile. Cubile clearly divided medially into a pair of somewhat triangle processes with caudal dent. Cucullus tubercle-like with papilla-shaped medial extension. Sacculus about twice bigger than cucullus, tubercle-like with elongated finger-like apically pointed extension loosely covered with chaetae. Juxta a tiny medial extension of aedeagus. Aedeagus c-shaped with ventral apical spur. Vesica short, wrinkled, caudally narrows. Eight sternite a c-shaped band, posterior margin with medial concavity. Eight tergite somewhat oval widened posteriorly. Female (Figs 2, 7, 10). Reminds male in pattern but paler, larger in size, antenna pectination much shorter. Head and thorax speckled creamy with brown scales. Thorax medially paler with two longitudinal dark stripes. Abdomen brown or orangish with creamy speckles. Forewing. Forewing length: 33–35 mm; wingspan: 67–68 mm. Elongated, apex blunt, outer margin with indentation on veins. Background color speckled creamy with brown scales. Pattern diagonal (Fig. 2), consist of speckled medial field, wavy postmedial fascia with faded dark edges, pale postmedial field, wavy external fascia with faded dark edges, and dark external field. Cilia creamy on vines and dark brown between them. Hindwing. Somewhat ovaloid with some indentation on veins. Background color creamy with dark speckled external field. Cilia speckled creamy and brown, darker between veins. Genitalia (Figs 4, 28). Papillae anales oval, densely covered with chaetae. Posterior and anterior apophyses about the same length. Antevaginal plate somewhat pentagonal with round corners, bends outwards with help of male cubile; together with postvaginal plate forms cup-like antrum. Ostium wrinkled, amorphous. Ductus bursae short. Corpus bursae wrinkled. Variability. Male. One paratype male (Fig. 8) looks distinct compared to the others (Figs 5–6, 9). It was collected in the lowlands, in April. The other adults were collected from altitudes above 500 meters in October and December. It has no significant difference in genitalia and originates near the others, so we consider it within the same species. Overall, male may have more (Fig. 8) or less pronounced and contrasting pattern (Figs 5–6, 9). Dark stroke in forewing medial field may present (Fig. 8) or absent (Figs 5–6, 9). Hindwing may have dark blurred medial fascia (Fig. 8). Cubile may be more sclerotized basally (Fig. 21). Female. Females have stable wing patterns, only the abdomen may get more or less orangish. Antevaginal plate with more (Fig. 4) or less (Fig. 28) protruded medial extension. Etymology. The species is named in honor of Dr. Hela Gayatri. She has a motto: “To live in sync with nature with deep appreciation of even the smallest part of it.” In addition to being an ardent lover of nature, she is also a distinguished scientist and a successful inventor and entrepreneur. She has a master’s degree in organic chemistry, a doctorate in chemistry and has international research experience with the Medical Research Council (UK), École Normale Supérieure (France), Texas Medical Center (USA) and National Tsing Hua University (Taiwan). Dr. Hela Gayatri also has expertise in synthetic chemistry, nanotechnology and other leading technologies. She is a serial entrepreneur and graduated from Goldman Sachs 10K (Cohort 2) initiative at NSRCEL, IIM, Bangalore and is an alumni of Goldman Sachs 10K Women Entrepreneur program, reimagining business at ISB, India. Distribution. Southern Kenya and north-eastern Tanzania (for details visit https://bit.ly/ Vavizola _hela). Biology. Adults were collected in April, October, and December from lowlands up to 2,500 meters a.s.l. Preimaginal stages are unknown.

Vavizola gen. n. https://zoobank.org/ urn:lsid:zoobank.org:act: 06558427-344F-44B3-A355-3686BD8978B9 (Figs 1–10, 21–22, 28, 34) Diagnosis. The detailed description of morphology is provided for the new species below, since the genus is monotypic. Habitus of Vavizola gen. n. reminds us of some other members of “ Pachypasa sensu lato ” revised by Zolotuhin & Gurkovich (2009): 1) Seydelora Zolotuhin & Gurkovich, 2009; 2) Gufria Zolotuhin & Gurkovich, 2009; 3) Lasiocesa Koçak, 2013; 4) Braura Walker, 1865; and 5) Eutricha Hübner, 1814. Adults of all genera have a diagonal wing pattern, cubile in male genitalia and a corresponding antevaginal plate in female genitalia. 1) Seydelora includes only Seydelora semna (Hering, 1941) from DRC. It has very dark forewings with dark brown medial, postmedial and external fields and a complex speckled white or yellow and brown external fascia; hindwings have a well pronounced dark external field (Figs 13–14). Male adults of Vavizola gen. n. are overall lighter, their forewings are more elongated, medial field is more elongated towards the wing apex, postmedial fascia is developed, the medial white triangle on thorax is absent (compare Figs 5–6, 8–9 and 13); male genitalia of Vavizola gen. n. differ by shorter cucullus and sacculus, and smaller apodema of cubile (compare Figs 21–22 and 25). Female adults of Vavizola gen. n. are overall lighter, their forewings are more elongated, forewing pattern is shifted more towards the wing apex, postmedial and external lines are doubled, medial pale speckled thoracic spot is absent (compare Figs 7, 10 and 14); female genitalia of Vavizola gen. n. differ in the shape of sterigma and presence of cup-like antrum (compare Figs 28 and 29). 2) Zolotuhin & Gurkovich (2009) considered the genus Gufria to be monotypic with the only member Gufria limosa de Villiers, 1827 distributed from Southern Europe to North Africa. We suppose that European and African populations will show a genetic divergence, sufficient to be considered a separate species, similar to experiences with Lemonia philopalus (Donzel, 1842) (see Prozorov et al., 2022b). Here, for comparison, we take adults of G. limosa from Tunisia and Morocco and call them Gufria limosa powelli (Oberthür, 1916) – the earliest taxon from North Africa. Adults are colored in combinations of grey, creamy and brown; medial field on forewing stretches until the wing apex, postmedial field is paler than the others (Figs 17–18). Male adults of Vavizola gen. n. have more elongated forewings with medial field not reaching the wing apex, and postmedial lines (compare Figs 5–6, 8–9 and 17); male genitalia of Vavizola gen. n. differ with shorter cucullus and sacculus, shorter distal outgrowths and lack of lateral dents (compare Figs 21–22 and 23). Female adults of Vavizola gen. n. have more elongated forewings, their forewing pattern consists of more elements and both wings have dark external field (compare Figs 7, 10 and 18); antevaginal plate in female genitalia of Vavizola gen. n. is better developed and antrum is wider (compare Figs 28 and 31). 3) Lasiocesa includes 4 species. Here, for comparison, we take adults (Figs 15–16) and male genitalia (Fig. 26) of the type-species Lasiocesa fulgurata (Aurivillius, 1909) and female genitalia (Fig. 32) of Lasiocesa lanceolata (Hering, 1932) due to only one known bad quality slide of female genitalia of L. fulgurata, all from DRC. Adults are colored in combinations of brown and creamy, forewing has full set of fields and fasciae, while hindwing may be completely brown or creamy with brown external field. Wing pattern of both genera is very similar, however, Vavizola gen. n. is much paler and duller, have narrower postmedial field (compare Figs 5–10 and 15–16). Male genitalia of Vavizola gen. n. differ with shorter cucullus, larger sacculus, and lack medial ridges on processes of cubile (compare Figs 21–22 and 26). Female genitalia of Vavizola gen. n. differ with larger antevaginal plate and presence of cup-like antrum (compare Figs 28 and 32). 4) Braura includes 9 species. For comparison, we take the type-species Braura ligniclusa (Walker, 1865) from RSA. Adults have dark brown forewings with occasionally paler medial field, hind wings dark brown or creamy with darker external field. Male adults of Vavizola gen. n. are overall lighter, but head and thorax cranially are not contrasting (compare Figs 5–6, 8–9 and 11); male genitalia of Vavizola gen. n. differ with smaller cucullus and basally larger sacculus, smaller caudal processes of cubile (compare Figs 21–22 and 24). Female adults of Vavizola gen. n. are overall lighter, but head, thorax cranially, and forewing medial field are not contrasting (compare Figs 7, 10 and 12); female genitalia of Vavizola gen. n. differ in better developed antevaginal plate and presence of cup-like antrum (compare Figs 28 and 30). 5) Eutricha includes 5 species ranging in coloration from creamy to dark brown. Here, for comparison, we use Eutricha capensis (Linnaeus, 1767) from RSA, the type-species of the genus. Adults have well pronounced contrasting postmedial and external fasciae. Male adults of Vavizola gen. n. are overall lighter, their forewings are more elongated, medial field is more elongated towards the wing apex (compare Figs 5–6, 8–9 and 19); male genitalia of Vavizola gen. n. differ with smaller cucullus and sacculus, smaller apical dent of aedeagus (compare Figs 21–22 and 27). Female adults of Vavizola gen. n. are overall paler and duller with contrasting external field (compare Figs 7, 10 and 20); female genitalia of Vavizola gen. n. differ in shape of sterigma (compare Figs 28 and 33). DNA comparison (Fig. 34). Two specimens of Vavizola hela sp. n. were sequenced: the holotype male from Tanzania (Fig. 5) and the paratype female from Kenya (Fig. 10). The two have a 0.8% p -distance which is a little higher than we expected for specimens of a single species collected so closely together. It may be explained by the1200 meters difference in the altitude between collecting localities of the two. The new genus is compared with 18 sequences belonging to 12 biological index numbers (BINs) and 9 genera from the “ Pachypasa sensu lato ” group, missing Seydelora; Pachyna Weymer, 1892; Beriola Zolotuhin & Gurkovich, 2009; Euphorea Zolotuhin & Gurkovich, 2009; and Sophyrita Zolotuhin & Gurkovich, 2009 (see Table 1). We will only compare the new genus with the others without investigation of their internal concerns such as potential polyphyly of Pachytrina Zolotuhin & Gurkovich, 2009 or polytypy of Muzunguja which follow from the tree and p -distances. These differences require a detailed investigation. We can see that intergeneric p -distance lays between 4.7 and 12% (Fig. 34), where the lowest is between Pallastica Zolotuhin & Gurkovich, 2009 and Cleopatrina Zolotuhin & Gurkovich, 2009, and the highest is between Muzunguja and Lasiocesa. The nearest neighbor of Vavizola gen. n. (BOLD:AAV0301) found on BOLD is Eutricha morosa (Walker, 1865) from Malawi (BOLD:ABZ6351) at 5.92%. We selected two other Eutricha species which are not much farther: 6.4 and 7.4%. Other sequences, except Pachytrina sp. at 6.1% (Fig. 34, 10), are farther than any Eutricha. Etymology. Name of the new genus is devoted to Prof. Dr. Vadim Viktorovich Zolotuhin (1967–2021), Russian entomologist specialized on the Old World Lasiocampidae. It is formed by a combination of the first letters of his name by analogy with Tarsozeuzera vavizola Yakovlev, 2006 (Cossidae)., Published as part of Prozorov, Alexey M., Prozorova, Tatiana A., Nedoshivina, Svetlana V., Yakovlev, Roman V., Volkova, Julia S., Saldaitis, Aidas, Revay, Edita E. & Müller, Günter C., 2023, Vavizola hela - new species and genus of Afrotropic Lasiocampini (Lepidoptera, Lasiocampidae), pp. 55-66 in Ecologica Montenegrina 62 on pages 57-63, DOI: 10.37828/em.2023.62.8, http://zenodo.org/record/8044383, {"references":["Zolotuhin, V. V. & Gurkovich, A. V. (2009) A review of the genus Pachypasa Walker, 1855 sensu lato in Africa (Lepidoptera, Lasiocampidae). Neue Entomologische Nachrichten, 63, 1 - 75.","Kocak, A. O. (2013) Nomenclatural notes in the family Lasiocampidae (Lepidoptera). Miscellaneous Papers, 160, 1 - 5.","Walker, F. (1865) List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII. Supplement Part 2, 323 - 706.","Hering, E. M. (1941) Neue Heteroceren aus dem Congo-Gebiet. Revue de Zoologie et Botanique Africaines, 35 (1), 72 - 84.","de Villiers, A. - P. (1827) Note sur trois Lepidopteres inedits ou peu connus de la France meridionale. Memoires de la Societe Linneenne de Paris, 5, 471 - 485.","Donzel, M. H. (1842) Description de deux lepidopteres nouveaux recueillis en Barbarie par le capitaine Charlon. Annales de la Societe entomologique de France, 11, 197 - 199.","Prozorov, A. M., Prozorova, T. A., Volkova, Ju. S., Yakovlev, R. V., Nedoshivina, S. V., Pinzari, M., Pinzari, M., Scalercio, S., Bianco, G., Saldaitis, A., Hausmann, A., Revay, E. E. & Muller, G. C. (2022 b) Revision of the Lemonia taraxaci complex, with a description of a new species from Italy and clarification of the status of Lemonia strigata (Lepidoptera: Brahmaeidae: Lemoniinae). Zootaxa, 5195 (4), 337 - 360. https: // doi. org / 10.11646 / zootaxa. 5195.4.2","Oberthur, Ch. (1916) Etudes de lepidopterologie comparee, 12. Imprimerie Oberthur, Rennes, 527 pp. https: // doi. org / 10.5962 / bhl. title. 8792","Aurivillius, Ch. (1909) Diagnosen neuer Lepidopteren aus Afrika. 9. Arkiv for Zoologi, 5 (5), 1 - 29. https: // doi. org / 10.5962 / bhl. part. 3496","Hering, E. M. (1932) Neue Heteroceren aus Afrika. Revue de Zoologie et Botanique Africaines, 22 (1), 102 - 117.","Linnaeus, C. (1767) Systema Naturae, 1 (2). Editio duodeciima reformata. Impensis Direct. Laur. Salvii, Holmiae, 533 - 1327 pp.","Weymer, G. W. (1892) Exotische Lepidopteren VI. Aus dem Afrikanischen Faunagebiet. Stettiner Entomologische Zeitung, 53 (4 - 5), 79 - 125.","Yakovlev, R. V. (2006) New Cossidae (Lepidoptera) from Asia, Africa and Macronesia. Tinea, 19 (3), 188 - 213."]}

The article presents an illustrated catalog of the genus Inguromorpha Edwards, 1888 (Lepidoptera: Cossidae, Hypoptinae), which includes 22 species. The following new combinations are established: Inguromorpha centrosoma (Dyar, 1925) comb. nov., Inguromorpha clathrata (Dognin, 1910) comb. nov., Inguromorpha clymene (Schaus, 1921) comb. nov., Inguromorpha crassiplaga (Schaus, 1905) comb. nov., Inguromorpha racana (Dognin, 1920) comb. nov. Eight new species are described: Inguromorpha arawaka Naydenov, Yakovlev & Penco sp. nov., Inguromorpha bachmanni Naydenov, Yakovlev & Penco sp. nov., Inguromorpha catarinea Naydenov, Yakovlev & Penco sp. nov., Inguromorpha muisca Naydenov, Yakovlev & Penco sp. nov., Inguromorpha paraguaica Naydenov, Yakovlev & Penco sp. nov., Inguromorpha scutulata Naydenov, Yakovlev & Penco sp. nov., Inguromorpha texasensis Naydenov, Yakovlev & Penco sp. nov., Inguromorpha willinki Naydenov, Yakovlev & Penco sp. nov. The distributional maps for all the species of the genus are provided.