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Hoplopeza colombiana sp. nov. (Figs 2A–G, 5A–G, 7A–B, 8) urn:lsid:zoobank.org:act: Diagnosis. Antenna dark brown (Fig. 2C); occiput without pruinosity, shiny black (Fig. 2D, E). Scutum shiny black, except posterior margin of postpronotal and postalar callus covered with dense yellow pruinosity (Fig. 2E); fore leg yellow, except tarsomeres 3–5 brown (Fig. 2A); fore tibia with several long, slender yellow hairs at mid apical half on ventral face (Fig. 2A); mid femur with 1 anterior apical black seta; hind femur slightly swollen with one preapical wide black ring and 3 strong black anterodorsally spaced setae (Fig. 2 F). Left epandrial lamella without spines along inner margin (Fig. 5A, B); hypoproct with irregular apex (Fig. 5G). basal medial cell; br = basal radial cell; CuA = anterior branch of cubital vein; cua = anterior cubital cell; CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; CuA = anterior cubital vein; dm = discal medial cell; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R 4+5 = third branch of radius; Rs = radial sector. Description. Holotype male. Body length: 4.5 mm (Fig. 2A). Wing length: 4.0 mm (Fig. 2G). Head. Frons shiny black, very narrow at lower margin, narrower than anterior ocellus, slightly widening towards anterior ocellus (Fig. 2D). Face bare, 1.5 times length of frons. Antenna (Fig. 2C), dark brown, postpedicel lanceolate, covered with dense brownish microtrichia; 1 pair of short proclinate ocellar setae. Proboscis short, yellow; palpus very small, covered with dense, dark yellow pruinosity with 1 long, slender seta near apex. Occiput smooth, shiny black; postocular setae dark, long and stout, and row of occipital yellow setae, longer than postocular setae. Thorax (Fig. 2E). Antepronotum black, covered with dense gray pruinosity. Scutum black, shiny, except posterior margin of postpronotal lobe brown, with anterior dark yellow spot; postalar callus dark yellow. Pleura brown, except yellow spot on upper margin of katepimeron, anterior margin of laterotergite, posterior margin of metanepisternum and metepimeron yellowish. Pleura with dense yellow pruinosity, except bare middle region of katepimeron. Scutellum black, except lateral and apical margins brown, with yellow pruinosity; 1 thin, short pair of lateral setae and 1 long, strong, dark yellow apical pair of scutellar bristles. Mediotergite black, with yellow pruinosity. Acrostichals uniserial; dorsocentrals biserial, short, slender and sparse on scutum; notopleuron with grey pruinosity, 2 bristles long, strong, upper longer and stronger, about half-length of scutum, lower seta yellow, shorter, about 1/3 length of upper bristle; several slender and short postpronotal setae; 1 anterior stronger, longer (about 1/3 length of scutum) bristle and 1 posterior short, slender postalar seta. Wing (Fig. 2G). Membrane light brown; pterostigma brown, elongate and narrow; R 1 long, ending at apical 1/2 of wing; vein bm-m slightly sinuous; vein CuA evanescent, not reaching CuP vein, arising near two-third length of bm cell; CuA+CuP long, ending just short of wing margin; veins M 1 and M 4 reaching wing margin. Halter yellow. Legs. Yellow (hind leg greenish yellow), except hind tibia with base and apex dark brown; hind femur slightly swollen, with dark brown preapical wide ring (Fig. 2F); tarsomere 1 of fore and mid legs brown at apex; fore, mid and hind tarsomeres 3–5 brown. Chaetotaxy: fore femur with row of slender yellow setae along anterodorsal surface and 1 long, yellow, slender anteroventral basal seta. Mid femur with 1 long anteroapical black seta. Hind femur with 3 long, strong setae on anterodorsal surface: 1 at basal 1/3, 1 near middle and 1 preapical. Fore tibia with several slender yellow ventral setae at mid apical half. Mid tibia with 1 short dorsal and 2 strong anterodorsal setae on apical half; 2 long, slender light brown setae: 1 anteroventral and 1 posteroventral. Hind tibia with 2 black apical spurs: 1 shorter anterodorsal and 1 long anteroventral. Abdomen (Fig. 2A). Shiny. Syntergites 1+2 brown, with greenish yellow spot on lateral margins; tergites 2–7 brown, with anterior margins yellow, weakly sclerotized; tergites 7 and 8 completely brown. All segments with short, long, slender yellow setae. Male terminalia (Fig. 5A–G). Right surstylus basally broad, with several long and strong setae on ventral margin, dorsal margin serrated, apex elongated, digitiform with rounded apex (Fig. 5A, B, E). Left surstylus shorter than right, almost subrectangular, with slightly elongated and rounded apex (Fig. 5A, B, D). Asymmetrical cerci, strongly sclerotized, digitiform, covered with brown, slender, long setae; left cercus slightly longer than right (Fig. 5A, B). Right epandrial lamella broader than left, rounded sides, with strong, dark rounded-tipped spines along inner margin in dorsal view (Fig. 5A, E). Left epandrial lamella as long as wide with strong setae along inner margin on dorsal view (Fig. 5A, D). Phallus elongate, phallic shaft gradually arched, cylindrical, without protuberances, distal articulation filiform, not bifid (Fig. 5F). Hypandrium subtriangular with several long, slender ventral setae at apex (Fig. 5C, F). Dorsal connection between lamellae narrow and short (Fig. 5A, B). Bacilliform sclerites divided, right basally fused with hypoproct, shorter and bare; left longer with apical half with scattered setae (Fig. 5G). Subepandrial sclerite with basal margin rounded, slightly widening toward at apex. Hypoproct sclerotized, subcylindrical with irregular apex (Fig. 5G). Female. Similar to male, except as noted: Body length: 5.0– 5.5 mm (Fig. 2B). Wing length: 4.5–5.0 mm. Abdomen. Tergites 2–4 brown with lateral yellow patches, segment 8 proximal half yellow. Terminalia (Fig. 7A, B). Tergite 8 and sternite 8 fused. Tergite 8 broad with posterolateral subtriangular extension. Sternite 8 subrectangular divided posteriorly into pair of extended sclerites with acute posterior apex. Tergite 10 undivided, separate from cercus. Membrane between sternite 8 and 10 hyaline with small spines arranged in longitudinal row. Sternite 10 undivided, slightly sclerotized, apex subtriangular. Cercus short, digitiform, length nearly twice width, bearing long, fine setae. Distribution. This species is known from Colombia (Antioquia) (Fig. 8). Type Material. HOLOTYPE m#: COLOMBIA: Antioquia department, Belmira Páramo de Santa Inés, El Morro location, 3005–3080 m, 6.634233; -75.658654, Forest. Malaise trap floor, 25-mar–5-abr-2017, Proy. Moscas de las flores, A.L. Montoya, C. Rodriguez & J.P. Carmona (CEUA 32646, dry pinned; genitalia preserved in glycerine in microvial). PARATYPES: COLOMBIA: Antioquia department, Belmira Páramo de Santa Inés, Río Arriba rural settlement, Laguna de Sabanas location, 2950–3050 m, 6.634233, -75.8654, Forest, Malaise trap floor, proy. Moscas de las Flores, 2–12.xii.2017, J. Torres-Toro & J.P. Carmona (1 m# CEUA 103671, 1 f# CEUA 102980, both body in 96% ethanol); same data except, Emergence trap (3 f#, CEUA 103672, body in 96% ethanol); same data except, pan trap, 16–27.ix.2017, A.L. Montoya & J.P. Carmona (2 m# CEUA 103106, wing mounted in Euparal on microscope slide; 1 f# CEUA 103107 body in 96% ethanol); same data except, sweep net (2 m# CEUA103104); same data except, 25.iii.–5.iv.2017, A.L. Montoya, C. Rodriguez & J.P. Carmona (1 m# CEUA 102474, 1 f# CEUA 102475, both with body in 96% ethanol); same data except, 25.iii.–5.iv.2017, Malaise trap floor (1 f#, CEUA 102828, body in 96% ethanol), Malaise trap canopy (2 f#, CEUA 102825, body in 96% ethanol), Malaise trap canopy (2 f# CEUA 102827, both body in 96% ethanol); Antioquia department, Belmira Páramo de Santa Inés, La Candelaria rural settlement, El Morro location, 3185–3200 m, 6.645901, -75.670441, Páramo, Malaise trap canopy, 21–30.vi.2017, C. Rodríguez & A.L. Montoya (1 f# CEUA 102956, body in 96% ethanol); same data except, Grassland, 21–30.vi.2017, C. Rodriguez & A.L. Montoya (1 f# CEUA 102824, body in 96% ethanol); same data except, 25.iii.–5.iv.2017, A.L. Montoya, C. Rodriguez & J.P. Carmona (1 f# CEUA 102823, body in 96% ethanol); Antioquia department, San José de La Montaña municipality, Páramo El Congo rural settlement, El Congo, 3165– 3190 m, 6.762229, -75.723144, Páramo, emergence trap, 20–30.vi.2017, A.L. Montoya & A. Sepúlveda (1 f# CEUA 102982, body in 96% ethanol) same data except, 25.iii–5.iv.2017 (1 f# CEUA 102822), 16–27.ix.2017, C. Henao-Sepúlveda (1 m# CEUA 103096); Antioquia department, Envigado municipality, Parcelación Nubarrones location, 2000 m, 6.139736, -75.5538545, forest, Malaise trap floor, 11.ix.2016 – 7.viii.2018, M. Wolff & C. Henao-Sepúlveda (1 f# CEUA 32633, 1 f# 32634, 1 f# 32635, 1 f# 32636, 1 f# 32637, 1 f# 32638, 1 f# 32639, 1 f# 32640, 1 f# 32641, 1 f# 32642, 1 f# 32643, all with body in 96% ethanol), same data except, Malaise trap canopy, 25.iii.–5.iv.2017 (1 f#, CEUA 102822, body in 96% ethanol); Antioquia, Yarumal municipality, El Respaldo rural settlement, La Maruja farm location, 2200 – 2600 m, 6.91046, -76.42792, forest, Malaise trap floor, 15.xii.2016, C. Henao-Sepúlveda (2 f# CEUA 32644, body in 96% ethanol); same data except, 1–22.iv.2017, C. Henao-Sepúlveda & D. Medina (1 f# CEUA 32645, body in 96% ethanol). Etymology. The specific epithet colombiana (nominative, adjective feminine) of this species refers to the country of Colombia, where the type specimens were collected. Remarks. Hoplopeza colombiana sp. nov. and H. chloropa Bezzi, 1909 share the shiny occiput, but the two species are easily separated by the absence of the row of long spiny setae on the hind basitarsomere in the new species, which is present in H. chloropa. Moreover, in terminalia structures H. colombiana can be differentiated from H. chloropa, by the absence of strong apical spine on the left cercus, which is present in H. chloropa (Rafael 1995, fig. 7). Hoplopeza colombiana sp. nov. can be differentiated from H. rafaeli sp. nov. by the shiny occiput, which is pruinose in H. rafaeli. In male terminalia, the strong curved spines along the inner margin of the left epandrial lamella are absent in H. colombiana, whereas they are present in H. rafaeli., Published as part of Henao-Sepúlveda, Carolina, Barros, Luana Machado, Ale-Rocha, Rosaly & Wolff, Marta, 2023, Two new species of the genus Hoplopeza Bezzi (Diptera, Hybotidae, Ocydromiinae) from the high Andean forests of Colombia, pp. 87-100 in Zootaxa 5297 (1) on pages 89-94, DOI: 10.11646/zootaxa.5297.1.4, http://zenodo.org/record/7989319, {"references":["Collin, J. E. (1933) Diptera of Patagonia and South Chile. Part IV. Empididae. British Museum (Natural History), London, 334 pp.","Bezzi, M. (1909) Beitrage zur Kenntnis der sudamerikanischen Dipterenfauna auf Grand der Sammelergebnisse einer Reise in Chile, Peru und Bolivia, ausgefuhrt in den Jahren 1902 - 1904 von W. Schnuse. Fam. Empididae. Nova acta Academiae Caesareae Leopoldino-Carolinae, 91, 297 - 406.","Rafael, J. A. (1995) Revis \" o das especies Neotropicais de Empididae (Diptera) descritas por Mario Bezzi. III. Ocydromiinae. Revista Brasileira de Entomolgia, 39 (4), 751 - 754."]}

Key for identification of Andean species of Hoplopeza Bezzi 1 Occiput bare, shiny black (Fig. 2D, E); fore tibia with several long, slender and wavy yellow setae on ventral surface (Fig. 2A)................................................................................................ 2 - Occiput with pruinosity, dull black (Fig. 3C–E); fore tibia without several long, slender and wavy yellow setae on ventral surface (Fig. 3A)...................................................................................... 3 2 Hind tarsomere 1 without short and strong spine-like setae on dorsal surface (Fig. 2F); left epandrial lamella without row of long spine-like setae on dorsal surface (Fig. 5A, B); left cercus without apical spine (Fig. 5A, B).................................................................................................. Hoplopeza colombiana sp. nov. - Hind tarsomere 1 with row of short and strong spine-like setae on dorsal margin (Fig. 4B); left epandrial lamella with row of long spine-like setae on dorsal surface (Rafael 1995, fig. 7); left cercus with long apical spine (Rafael 1995, fig. 7)........................................................................................ Hoplopeza chloropa Bezzi 3 Antenna light brown; pleura tawny-brown; postpronotal lobe yellow (Fig. 4A); fore femur without long and strong dorsal setae near apex; short and slender apical spur at apex of hind tibia.............................. Hoplopeza annulata Collin - Antenna black; pleura dark brown (Fig. 3A, B); postpronotal lobe black, except posterior margin paler; fore femur with one long and strong dorsal seta near apex; long and strong apical spur at apex of hind tibia (Fig. 3F)... Hoplopeza rafaeli sp. nov., Published as part of Henao-Sepúlveda, Carolina, Barros, Luana Machado, Ale-Rocha, Rosaly & Wolff, Marta, 2023, Two new species of the genus Hoplopeza Bezzi (Diptera, Hybotidae, Ocydromiinae) from the high Andean forests of Colombia, pp. 87-100 in Zootaxa 5297 (1) on page 89, DOI: 10.11646/zootaxa.5297.1.4, http://zenodo.org/record/7989319, {"references":["Rafael, J. A. (1995) Revis \" o das especies Neotropicais de Empididae (Diptera) descritas por Mario Bezzi. III. Ocydromiinae. Revista Brasileira de Entomolgia, 39 (4), 751 - 754."]}

Hoplopeza rafaeli sp. nov. (Figs 3A–G, 6A–G, 7C–D, 8) urn:lsid:zoobank.org:act: Diagnosis. Antenna black (Fig. 3C); occiput black, covered with dense yellow pruinosity (Fig. 3E). Scutum shiny black, except posterior margin of postpronotal lobe paler and anterior margin of postalar callus yellow (Fig. 3E). Mid basitarsomere with 1 strong posteroventral basal seta; hind tibia brown, paler on base (Fig. 3F). Tergites of abdomen brown, greenish anteriorly, syntergite 1+2 greenish on lateral margin (Fig. 3A, B). Left epandrial lamella with spines along the inner margin (Fig. 6A, B); hypoproct with rounded apex (Fig. 6G). Description. Holotype male. Body length: 4.5 mm (Fig. 3 A). Wing length: 4.7 mm (Fig. 3G). Head. Frons shiny brown, narrower than anterior ocellus, slightly widening towards anterior ocellus (Fig. 3D). Face with short whitish setulae, 1.6 times length of frons. Antenna (Fig. 3C) black, postpedicel lanceolate, covered with dense brownish microtrichia; 2 pairs of ocellar setae proclinate, posterior pair very short and slender, anterior pair long and stronger. Proboscis short, brown; palpus very small, yellow, covered with dense yellow pruinosity; 1 long, slender seta. Occiput black, covered with dense yellow pruinosity, patch without pruinosity next to ocellar triangle; postocular setae long, strong and scattered, upper setae longer, stronger; row of long, brown occipital setae. Thorax (Fig. 3E). Antepronotum black, covered with dense yellow pruinosity, with row of short, slender setae on anterior margin. Scutum black, shiny, except posterior margin of postpronotal lobe paler; anterior margin of postalar callus yellow. Pleura brown, except yellow spot on upper margin of katepisternum; upper margin of katepimeron, anterior margin of laterotergite, posterior margin of metanepisternum and metepimeron yellowish. Pleura with dense yellow pruinosity, except bare middle region of katepimeron. Scutellum black, except lateral and apical margins brown, with yellow pruinosity; 1 thin, short pair of lateral setae and 1 long, strong, black apical pair of scutellar bristles. Mediotergite black, with yellow pruinosity. Acrostichals uniserial, dorsocentrals biserial, short, slender and sparse on scutum; notopleuron with yellow pruinosity, 2 bristles long, strong, upper longer and stronger, about half length of scutum, lower seta yellow, shorter, about 1/3 length of upper bristle; several slender and short postpronotal setae; 1 stronger, longer (about 1/3 length of scutum) anterior and 1 shorter, slender posterior postalar seta. Wing (Fig. 3G). Membrane brownish; pterostigma brown, elongate and narrow; R 1 long, ending at apical 1/2 of wing; cell dm aligned with cell br; vein bm-m strongly sinuous; vein CuA incomplete, not reaching vein CuP, arising near mid-length of cell bm; CuA+CuP long, reaching wing margin; veins M 1 and M 4 reaching wing margin. Halter yellow, knob slightly darker. Legs. Yellow, except brown apical spot on dorsal surface of mid femur; hind femur with wide subapical ring and apex brown; hind tibia (Fig. 3F) brown, paler basally; fore and mid tarsomere 1 brown at apex; fore and mid tarsomere 2 pale brown; fore and mid tarsomeres 3–5 brown; hind tarsomeres brown, except yellow greenish on base of tarsomere 1. Hind femur slightly swollen. Chaetotaxy: fore femur with row of thin setae along anterodorsal surface, decreasing in size toward apex, one long and strong dorsal seta near apex. Mid femur with 2 long, strong anterodorsal setae: 1 at mid-length and 1 subapical. Hind femur with 3 long, strong anterior setae: 1 subbasal, 1 at mid-length and 1 subapical. Mid tibia with 2 long, strong anterodorsal setae: 1 subbasal and 1 at mid-length; 1 long, strong posterodorsal subbasal seta, 2 long, strong anteroventral and 2 posteroventral apical setae. Hind tibia (Fig. 3F) with 2 long, strong anterodorsal setae: 1 basal and 1 at mid-length; 1 long, stronger lateral apical spur: 2 short, strong apical setae. Mid basitarsomere with 1 strong posteroventral basal seta. Abdomen (Fig. 3A). Shiny. Syntergites 1+2 brown, except yellow greenish spot on lateral margin; tergites 3–6 brown, except anterior margin greenish yellow, weakly sclerotized; tergites 7 and 8 brown.All segments with short, long slender setae. Male terminalia (Fig. 6A–G). Right surstylus basally broad, apex with elongated projection, digitiform and truncated apex, with several long, strong setae on ventral margin; dorsal margin serrated (Fig. 6A, B, D). Left surstylus shorter than right, almost subrectangular, with apex slightly pointed; several short, slender setae near apex (Fig. 6A, B, D). Asymmetrical cerci, strongly sclerotized, digitiform, covered with brown, slender and long setae; left cercus slightly longer than right (Fig. 6A, B). Right epandrial lamella broader than left, rounded sides, with strong dark spines along inner margin in dorsal view (Fig. 6A, B, D). Left epandrial lamella as long as wide with strong dark curved spines along inner margin in dorsal view (Fig. 6A, B, E). Phallus elongate, shaft gradually arched, cylindrical, without protuberances; distal articulation filiform, not bifid (Fig. 6F). Hypandrium subtriangular in ventral view with several long and slender setae at apex (Fig. 6C, F). Dorsal connection between epandrial lamellae narrow and short (Fig. 6A, B). Bacilliform sclerites divided basally, thin, entire length covered with several short, slender setae (Fig. 6G). Subepandrial sclerite with basal margin truncate, slightly narrower toward apex. Hypoproct clavate, with rounded apex and bare, strongly sclerotized, tube-like (Fig. 6G). Female. Similar to male, except as noted: Body length: 5.0– 5.5 mm (Fig. 3 B). Wing length: 4.5–5.0 mm. Mid and hind tarsomeres 3–5 light brown. Abdomen. Tergites 2–4 brown, yellow laterally, tergite 7 completely yellow. Terminalia (Fig. 7C, D). Tergite 8 and sternite 8 fused. Tergite 8 broad subrectangular with lateral posterior subtriangular extensions. Tergite 10 undivided, separate from cerci. Sternite 10 slightly sclerotized, apex broadly oval. Sternite 8 subrectangular, divided posteriorly into pair of extended sclerites with acute apex. Membrane between sternite 8 and 10 hyaline with small spines scattered randomly. Cercus short, rounded, length less than twice width, bearing long, fine setae. Distribution. This species is known from Colombia (Antioquia) (Fig. 8). Type Material. HOLOTYPE m#: COLOMBIA: Antioquia department, Belmira Páramo de Santa Inés, El Morro location, 3100–3300 m, 6.632639; -75.645267 Páramo, Malaise trap canopy, 02–12-dec-2017, Proy. Moscas de las flores, C. Rodriguez & A. Echeverry (1m# CEUA 102957, dry pinned, wing mounted in Euparal on microscope slide; genitalia preserved in glycerine in microvial). PARATYPES: COLOMBIA: Antioquia, San José de La Montaña municipality, El Congo rural settlement, 3100–3150 m, 6.763791; -75.701738, Páramo, sweep net, 20–30.vii.2017, Proy. Moscas de las flores, A. Sepúlveda & A.L. Montoya (1 m# CEUA 103105, body in 96% ethanol; 1 m# CEUA 102960, dry pinned); same data except, 25.iii.–5.iv.2017, C. Henao-Sepúlveda & M. Wolff (1 m# CEUA 103108, body in 96% ethanol); same data except, 16–27.ix.2017, C. Henao-Sepúlveda & A. Sepúlveda (1f# CEUA 138330, dry pinned). Caldas, Pensilvania, Valle alto, 3378 m, 5.345971; -75.286728, forest, ground Malaise trap, 28–30.iv.2014, A. Cardona & J. Alvarez (1f# CEUA 138331, body in 96% ethanol with genitalia preserved in glycerine in microvial). Etymology. Named in honor of José Albertino Rafael, researcher from Instituto Nacional de Pesquisas da Amazônia, Brazil, for his contribution to the knowledge of the family Hybotidae. Remarks. Hoplopeza rafaeli sp. nov. differs from H. chloropa by the absence of several long, slender and waved yellow setae on the ventral face of the fore tibia. Hoplopeza rafaeli sp. nov. differs from H. annulata by the color of the pleura, which is all dark brown in H. rafaeli, whereas in H. annulata the side of the scutum and some areas of the pleura are tawny-brown according to Collin (1933) and are visible in photos of the type (Fig. 4A). Hoplopeza rafaeli sp. nov. differs from H. colombiana sp. nov. by the left surstylus with smooth apex, whereas the left surstylus of H. colombiana sp. nov. has a serrated apex. Also, H. rafaeli sp. nov. can be differentiated from the other Andean species by two obvious structures of the male terminalia, including the strong, curved spines along the inner margin of the left epandrial lamella, and the long and wide hypoproct., Published as part of Henao-Sepúlveda, Carolina, Barros, Luana Machado, Ale-Rocha, Rosaly & Wolff, Marta, 2023, Two new species of the genus Hoplopeza Bezzi (Diptera, Hybotidae, Ocydromiinae) from the high Andean forests of Colombia, pp. 87-100 in Zootaxa 5297 (1) on pages 95-98, DOI: 10.11646/zootaxa.5297.1.4, http://zenodo.org/record/7989319, {"references":["Collin, J. E. (1933) Diptera of Patagonia and South Chile. Part IV. Empididae. British Museum (Natural History), London, 334 pp."]}

Argentinomyia belmira Montoya & Wolff sp. nov. Proposed standard English name: Belmira’s long-antennae flower fly. (Figs 16, 17 and 90A) Type material. HOLOTYPE. Adult Male, pinned, deposited at Colección Entomológica Universidad de Antioquia. Original label: “ COLOMBIA, Antioquia, Belmira, Páramo Santa Inés, Vereda Río Arriba, Sector Laguna de Sabanas” / “ 6,640931, -75,665335, 2850-2950 m, Forest, Net” / “ 4-14.vii.2016, A. L. Montoya; J. Torres-Toro; J. D. Carmona ”. “ HOLOTYPE / Argentinomyia belmira / Montoya & Wolff 2023” (CEUA 95284, dissected). Identified as Argentinomyia sp. 13 by Montoya 2019. PARATYPE. COLOMBIA. Label with the same data as Holotype (3♁, CEUA 112574–76) except: Malaise, 2-12.ii.2017, A.L. Montoya; C. Rodríguez; J.D. Carmona (1♀, CEUA 95481). Length (n= 3): Body, 7.9–8.2 mm; Wings, 7.5–7.7 mm. Diagnosis. Face black and somewhat brassy; wing slightly brownish; 3 rd and 4 th terga almost entirely yellow, with a pair of wider yellow maculae from the base to the apical margin of the segment, rounded on the lateral edge and with a thin median black stripe and narrow posterior black margin; 5 th tergum almost entirely yellow with only a small posteromedian black triangle, black pilose in the black areas and on the medio-apical region of the 3 rd to the 5 th terga; surstylus with dorsal margin slightly concave and ventral margin slightly convex; hypandrium narrowed laterally towards the apex; aedeagal lobe circular, apex rounded. Description (Holotype). MALE. Head (Fig. 16A): Face black and somewhat brassy, whitish-grey pollinose and pilose. Facial tubercle at least with two slightly pronounced transversal grooves. Gena black, white-grey pollinose and yellow pilose. Frontal triangle more brownish pollinose, pile black. Ocellar triangle black with black pile. Occiput white pollinose, black pilose on dorsal 2/3 and white pilose on ventral 1/3. Antenna dark reddish-brown, short, ratio 1.0:1.5:2.3, pedicel and scape combined longer than basoflagellomere; basoflagellomere blackish dorsal and reddish ventral; arista brown, with a few short pile. Thorax (Figs 16B–C): Mesonotum, scutellum and pleura black-blue semi-opaque, golden yellow pilose. Scutum with two whitish vittae on anterior half, reaching beyond the notopleural suture, golden pilose, except black pile in the postpronotum, notopleuron and postalar callus. Pleura shining black, with white to brownish pollen, yellowish pilose, except anterior anepisternum and katerpimeron with black pile. Scutellum shining, slightly rugose on apical half, short white pile in addition to long marginal black pile. Wing (Fig. 16C): Slightly brownish, stigma yellowish, membrane microtrichose, except cell bm bare on basal 2/4, r only slightly bare on basal 1/4 or less, cell cua bare on anterior 1/3; calypter white, border and fringe brown; plumula white; halter white, capitulum white. Legs (Fig. 16C): Brown to black, black pilose, 1 st and 2 nd metatarsi black, not strongly contrasting with the dark color of other segments. Abdomen (Fig. 8E, 16B): Mostly yellow, 1 st- 2 nd terga black shining, with long lateral yellowish pile; 3 rd and 4 th terga almost entirely yellow, with a pair of wider yellow maculae from the base to the apical margin of the segment, rounded on lateral edge and with a thin median black stripe and narrow posterior black margin; 5 th tergum almost entirely yellow with only a small posteromedian black triangle, black pilose in the black areas and on medioapical region of 3 rd to 5 th terga, genitalia shining black, black pilose; male genitalia: surstylus in lateral view (Fig. 17A) with dorsal margin slightly concave and ventral margin slightly convex; hypandrium in ventral view (Fig. 17C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 17C) circular, apex rounded. FEMALE (Figs 16D–F). Similar to male except for usual sexual dimorphism and differing in the 2 nd tergum almost entirely yellow, with a pair of wider yellow maculae from the base to the apical margin of the segment, this tergum black shining in male, terga 3 rd to 4 th with wider yellow maculae from the base to the apical margin of the segment, rounded on lateral edge and with a thin median black stripe and narrow posterior black margin, 5 th tergum with a short yellow macula, slightly widen than in male. Taxonomic notes. Argentinomyia belmira sp. nov. is similar to A. agonis but differs by the following combination of characters: Antenna dark reddish-brown (Figs 16A, C, D-F); wing slightly brownish (Figs 16 B-C, E-F); 3 rd and 4 th terga almost entirely yellow, with a pair of wider yellow maculae from the base to the apical margin of the segment, rounded on lateral edge and with a thin median black stripe and narrow posterior black margin, 5 th tergum almost entirely yellow with only a small posteromedian black triangle, black pilose in the black areas and on medio-apical region of the 3 rd to the 5 th terga (Figs 16B, E). A. agonis is characterized by the antenna black (Figs 10A, C, D-F); wing hyaline, vein R4+5 not sinuate; cell r1 open; vein M1 not recurrent (Figs 10 B-C, E-F); abdomen elongate, 2 nd to 4 th terga with large orange maculae, black lateral margins, in addition to a thin median black vitta and narrow posterior black margin (Figs 10B, E). Based on males, A. belmira sp. nov. differs from A. agonis in having the surstylus (Fig. 17A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad [versus surstylus in lateral view with dorsal and ventral margins approximately of the same width in the whole length, elongated, three to four times longer than broad in A. agonis (Fig. 11A)] (see “differential diagnosis” under each species or key). Etymology. Argentinomyia belmira sp. nov. is a noun in apposition, toponymy that refers to the type locality “ Belmira ”, which corresponds to the “Páramo Complex of Santa Ines Belmira” in the northwestern Central Cordillera in Colombia where the type specimens were collected. Remarks. The type material was collected in two sampling events in February 2016 and April 2017. The species was found inside a well-preserved patch of Andean forest (Fig. 11D) covered by abundant leaf litter; tree trunks covered by mosses and abundant epiphytes such as bromeliads and orchids. The species occurs notably in forests dominated by the Colombian Oak, Quercus humboldtii Bonpl. (Fagaceae) forming a closed canopy. The series types were collected flying in leks formation over the native Bromeliad species, Guzmania coriostachya (Griseb.) Mez (Bromeliaceae). Argentinomyia belmira sp. nov. is a forest species that occur sympatrically throughout high elevations of northwestern Colombia with A. humboldti sp. nov., A. ivani sp. nov., A. serendipia sp. nov., A. teresae sp. nov. and A. andina. Geographical range. Argentinomyia belmira sp. nov. (n= 5) is known exclusively from the Páramo Complex of Santa Ines Belmira, northwestern in the Colombian Central Cordillera (Antioquia) (Fig. 90A). The species is endemic to the Northern Andes domain, distributed at middle altitudes (2950 m) in the Magdalena province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 35-38, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Argentinomyia teresae Montoya sp. nov. Proposed standard English name: Teresa’s long-antennae flower fly. Argentinomyia sp. 17a by Montoya, unpublished (Figs 79, 80 and 90B) Type material. HOLOTYPE. Adult male, pinned, deposited at Colección Entomológica Universidad de Antioquia. Original label: “ COLOMBIA, Antioquia, Belmira, Páramo Santa Inés, Cabaña Cabildo Verde, El Morro-Alto de La Gallina ” / “ 6,632639, -75,645267 ” / “ 3170–3200 m, Páramo, ex Espeletia occidentalis, Red entomológica” / “ 4-14.xii.2016, A. L. Montoya, J. Torres, J. D. Carmona ”. “ HOLOTYPE / Argentinomyia teresae / Montoya 2023” (CEUA 98384). Identified as Argentinomyia sp. 17b by Montoya 2019. PARATYPES. Label same data as Holotype, except, 2.xii.2017, C. Rodríguez & A. Echeverry Leg. (1♁, CEUA 138733); …, Malaise Canopy #2, 20-30.vi.2017, A. L. Montoya, C. Rodríguez, J. D. Carmona (1♁, CEUA 97991); …, Net, 16-27.ix.2017, C. Rodríguez, A. Rúa (1♁, CEUA 97996); …, Vereda Río Arriba, Sector Laguna de Sabanas, 6,645 901, -75,670441, 3200 m, Páramo, Net, 28-31.v.2016, A. L. Montoya (1♁, CEUA 93563); Sonsón, Vereda San Francisco, Cerro Las Palomas A, 5,725 972, -75,249253, 3364– 3310 m, Net, 1-6.iv.2018, A.L. Montoya; J. P. Carmona (1♁, CEUA 114186); …, Vereda Nori, Cerro Nori, 5,809 667, -75,269028, 2853 m, Forest, Malaise canopy, 31.iii-7.iv.2018, A.L. Montoya; J. P. Carmona (1♁, CEUA 114224). Length (n= 3): Body, 6.8 mm; Wings, 7.3 mm. Diagnosis. Body mainly black. Antennae brownish, orange ventrally. Frontal triangle golden pollinose; metanotum golden pile; legs black, except pro- and mesotibiae yellow on apical 1/5 and basal 1/3; terga 3 rd and 4 th with a pair of basolateral rectangular maculae extending 1/3 of tergum length; surstylus shorter than the epandrium, with dorsal margin slightly concave and ventral margin slightly convex; hypandrium expanded laterally towards the apex; dorsal area of the apex of hypandrium (superior lobes) in lateral view, no extending dorsally backward, without a small sclerotized inner spur; aedeagal lobe circular, apex rounded; the base of the aedeagus normal, only slightly globose. Description. MALE. Head (Fig. 79A): Face black, sides yellow golden pilose and pollinose. Frontal triangle with transversal golden maculae, black pilose. Gena black, white pollinose and pilose. Ocellar triangle opaque, with a velvety brown cross band black pilose. Occiput white pollinose, white pilose on dorsal 2/3 and black pilose on ventral 1/3.Antenna brownish, orange ventrally, short, ratio 1.1:1.3:2.6; basoflagellomere apically rounded and longer than the scape and pedicel; arista black. Thorax (Figs 79B–C): Bluish gray, yellow pilose and pollinose; with two grayish vitae in the anterior half, just before the notopleural suture. Pleura bright bluish, golden pollinose, scattered whitish-golden pilose. Metanotum golden pile. Wing (Fig. 79C): Smoky brown, stigma brown, microtrichose; calypter wholly brown; plumule white; halter white, capitulum white. Legs (Fig. 79C): Black, except pro- and mesotibiae yellow on apical 1/5 and basal 1/3; pile yellow basally and black apically, coxae brown. Abdomen (Fig. 79B): Parallel sides, 3 rd and 4 th terga with a pair of basolateral rectangular maculae extending 1/3 of tergum length, golden pilose, except black pilose on median apical region of each segment; male genitalia: surstylus in lateral view (Fig. 80A) shorter than the epandrium, with dorsal margin slightly concave and ventral margin slightly convex; hypandrium in ventral view (Fig. 80C) expanded laterally towards the apex; dorsal area of the apex of hypandrium (superior lobes) in lateral view, no extending dorsally backward, without a small sclerotized inner spur; aedeagal lobe in ventral view (Fig. 80C) circular, apex rounded; the base of the aedeagus normal, only slightly globose. FEMALE. Unknown. Taxonomic notes. Argentinomyia teresae sp. nov. is similar in appearance to A. serendipia sp. nov. from which it differs in the antennae brownish, orange ventrally; frontal triangle golden pollinose (Figs 79A, C); terga 3 rd and 4 th with a pair of small basolateral rectangular maculae extending 1/3 of tergum length (Fig 79B). In A. serendipia sp. nov. the antenna black; frontal triangle dark pollinose (Figs 74A, C); terga 3 rd and 4 th with a pair of small rectangular maculae extending 1/6 of tergum length and restricted to basolateral margins of tergites (Fig 74B). Based on males, A. teresae sp. nov. differs from A. serendipia sp. nov. in having the surstylus in lateral view (Fig. 80A) shorter than the epandrium [versus surstylus larger than the epandrium in A. serendipia sp. nov. (Fig. 75A)]; dorsal area of the apex of hypandrium (superior lobes) in lateral view (Fig. 80A) no extending dorsally backward, without a small sclerotized inner spur [versus extending dorsally backward with a small sclerotized inner spur in A. serendipia sp. nov. (Fig. 75A)] (see “diagnosis” under each species or key). Etymology. The name of this beautiful Colombian fly is a matronym in honor of the first author’s mother, Teresa Giraldo (Educator for more than 34 years), in gratitude for her love and dedication, teaching to enjoy and marvel at the little things in nature, transmitting values that left indelible fingerprints as well as material and emotional encouragement that made possible my professional development. Remarks. Field observations suggest that A. teresae sp. nov. seems to be associated with the pollination of the Colombian endemic “Frailejón”, Espeletia occidentalis. Geographical range. Argentinomyia teresae sp. nov. (n= 8) is exclusively known from the Páramo complexes Belmira, El Congo and Sonsón in the massif at northwestern Colombia (Antioquia) (Fig. 90B). The species is endemic to the North Andes domain at middle and high altitudes (2950–3200 m) in the Magdalena province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 129-131, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Argentinomyia rugosonasa (Williston, 1891) (Figs 2A, 4C, 9D, 69, 70 and 96A) Melanostoma rugosonasus Williston, 1891: 13. Type locality: Mexico. Guerrero, Omiltemi, 8000ft. HOLOTYPE Female BMNH (Examined). Refs.: Fluke, 1945: 7 (redesc.) 26 (Fig. 6, head, lateral view, male); 1957: 269 (Figs 6, 15, male genitalia) 279 (Fig. 130, male genitalia); Thompson et al. 2010: 767 (Fig. 23, head, lateral view, male) Rugonasus Aldrich, 1905: 361, Williston, 1907: 2, Coquillett, 1910 a: 601, misspellings. Rhysops rugonasus. Curran, 1937: 2 (key) misspellings Rhysops rugosonasus. Fluke, 1945: 7 Rhysops rugosonasus. Fluke, 1957: 266 Rhysops rugosonasus. Thompson et al. 1976: 44 Melanostoma rugosonasus. Papavero & Ibañez-Bernal, 2003: 166 Argentinomyia rugosonasus. Thompson et al. 2010: 767 Type material. HOLOTYPE. Adult female, MEXICO. Omiltemi, 8000ft, Guerrero, July, H.H. Smith (BMNH). PARATYPE. MEXICO. Mexico City, Omiltemi, 2600 m, vii-1936, H. G. Meyer (8♁, 1♀, BMNH); Guerrero, Xucumanatlan ( Note: The correct writing of Xucumanatlan is“ Xocomanatlan ”and refers to the locality Xocomanatlan / Chilpancingo de Los Bravo) 7000ft, July. H.H. Smith (1♀, BMNH). Length (n= 7): Body, 9.8–10.2 mm; Wings, 9.1–9.8 mm. Diagnosis. Face vertical, flat, not produced downward, with at least six or seven transversal grooves, a large oval median shining vitta, the rest of face whitish pollinose; antennae elongate, scape and pedicel a little longer than basoflagellomere; notopleural tubercle very prominent; wings hyaline; protibia reddish to brownish-yellow; metafemur black or dark brown with long pile on outer sides, metatibia with about one-half dozens of longer brownish pile on outer sides; metabasitarsomere elongate, black or dark brown, not contrasting with the tibia and/ or rest of tarsus; abdomen elongate, opaque black, with metallic maculae more shining alongside and broadly on anterior corners of 3 rd and 4 th terga; surstylus triangular-like, shorter than broad. Redescription. MALE. Head (Figs 2A, 4C, 69A, 70D): Face moderately shining, submetallic black, flat, with a large oval median shining vitta, but the rest of the face is whitish pollinose. Face shining black, partly concealed beneath whitish pollen, which, on sides, is faintly striate; in middle, with well-marked, slender, about six or seven transversal grooves. Frontal triangle and vertex with a black opaque cross-band below ocelli. Ocellar triangle black with thin yellowish pollen, pile black. Antennae brownish, orange ventrally, short, ratio 1.7:1.0:2.5, scape and pedicel a little longer than basoflagellomere, basoflagellomere blackish in dorsal area, arista black, pilose. Thorax (Figs 69B–C, 70D): Black, mesonotum moderately shining, brownish-steel-blue, yellow pilose; with two darker, not conspicuous, pollinose vittae on anterior half, pile yellow. Pleura thinly grayish-brown pollinose, yellow pilose. Scutellum steel-blue, pile short, with long yellow pile; fringe yellow; notopleural tubercle very prominent. Wing (Fig. 69C): Hyaline, stigma yellow, microtrichose, except costal cell only slightly bare on basal 1/4, cells bm and r bare on basal 2/3 and 1/3, respectively, cell cua bare on anterior 1/3; calypter wholly brown; plumula white, halter orange, capitulum orange-brown. Legs (Fig. 69C): Pro- and mesolegs reddish or brownish-yellow, femur and tarsus blackish basally; metafemur black, with long yellow pile on outer sides; metatibia and tarsus brown, except reddish-yellow basally, metatibia with about half a dozen of longer brownish pile on outer sides, metabasitarsomere elongate. Abdomen (Figs 9D, 69B, 70D): Elongate, shining metallic bluish; 2 nd to 4 th tegum with a posterior black band extending more or less forward in middle, more shining along sides and broadly on anterior corners of 3 rd and 4 th terga; sterna black shining, yellow pilose; male genitalia: surstylus in lateral view (Fig. 70A) triangular-like, shorter than broad; hypandrium in ventral view (Fig. 70C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 70C) circular, apex rounded. FEMALE (Figs 64D–F, 65E). Similar to male except for usual sexual dimorphism and differing in the pile of legs is much briefer, with very few long hairs on femur and none on tibia. Taxonomic notes. Argentinomyia rugosonasa is a dark species that can be easily differentiated from A. crenulata and A. sagoti sp. nov. by the presence of face vertical, flat, not produced downward, with at least six or seven transversal grooves, a large oval median shining vitta, the rest of face whitish pollinose; antennae elongate, scape and pedicel a little longer than basoflagellomere (Figs 68A, C, D, F); notopleural tubercle very prominent; wings hyaline (Figs 68 B-C, E-F); protibia reddish to brownish-yellow; metafemur black or dark brown with long pile on outer sides, metatibia with about one-half dozens of longer brownish pile on outer sides; metabasitarsomere elongate, black or dark brown, not contrasting with the tibia and/or rest of tarsus (Figs 68A, C, D, F); abdomen elongate, opaque black, with metallic maculae more shining alongside and broadly on anterior corners of 3 rd and 4 th terga (Figs 68B, E). In A. crenulata the face has seven or eight narrow, shallow, transverse grooves above the tubercle and sides of the face with a thin white line of pollen; antennae elongate, scape longer than pedicel and basoflagellomere, pedicel and basoflagellomere about equal in length (Figs 26A, C, D, F); wings with two short medial transverse brown fasciae on anterior cross-vein (Figs 26 B-C, E-F); metafemur and protibia black, black pilose (Figs 26A, C, D, F); abdominal maculae metallic (Figs 26B, E). In A. sagoti sp. nov. the face with four or more distinct transverse grooves above the tubercle; antenna short (Figs 26A, C, D, F); wing lightly brown pigmented anteriorly; protibia yellow on basal 1/3 (Figs 26 B-C, E-F); metafemur and tibia yellow on apically 1/3 (Figs 26A, C, D, F); 2 nd to 4 th terga without maculae (Figs 26B, E). Based on males, A. rugosonasa differs from A. crenulata and A. sagoti sp. nov. in having the surstylus in lateral view (Fig. 69A) triangular-like [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. crenulata (Fig. 27A); surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. sagoti sp. nov. (Fig. 72A)] (see “diagnosis” under each species or key). Biology. Label data suggest that A. rugosonasa visits flowers of Helianthus salicifolius (Asteraceae). Geographical range. Argentinomyia rugosonasa (n= 76) is distributed in Costa Rica * (Cartago, San José, Puntarenas) and Mexico (Chiapas, Desierto de los Leones, Guerrero, Toluca, Mexico City, Omiltemi) (Fig. 96A). The species is endemic to the Mesoamerican domain, mainly distributed at middle and high altitudes (1766–2990 m) in the provinces: Chiapas Highlands (2510 m), Pacific Lowlands (2438–2600 m) and Puntarenas-Chiriquí (1766–2990 m). Non-type material examined. COSTA RICA. Cartago, 9,6, -83,7, 2438 m, vi.1996 (InBio); Puntarenas, Cerro Biolley, Estación Altamira, ACIA PILA, 9,076 418, -82,980144, 1766 m, 2.viii.1995, L. Donzo (1♀, INBioCRI 002251037, Cat. N. 5379); R.I. Ujaras, Salitre, Cabagra, Buenos Aires, 9,072 968, -82,909872, 2990 m, 19ix.2003, M. Alfaro, Libre (3♁, INBioCRI 0003753632; 3753633; 37553634, Cat. N. 74941); San José, Estación Cuerici, R. Negro, Mora, 9,537 281, -83,708056, 1800 m, 1.iv.1992, H. Vargas, F.G. Zumbado; M.A. Zumbado (1♁, INBioCRI 000409368); …, camino al sendero El Carbón 4, 6 km al E de Villa Mills, 9,564 652, -83,707899, 2600 m, 30.v.1996, A. Picado (3♁, INBioCRI 002466351; 2466353; 2394364; 2394365); 31.iii.1996, A. Picado (1♀, INBioCRI 002394362, Cat. N. 7028); …, límite de la Finca Cuerici, 3km E. Villa Mills, 9,564 652, -83,707899, 2800 m, 23.vi.1996, A. Picado (6♁, INBioCRI 002452405; 2452407; Cat. N. 7698, 7699, 5902); …, Rt 95 btw km 42-43, 1 km N La Cima near train overpass, sweeping Helianthus salicifolius, 8.viii.1989, A.L. Norrbom (1♁, 1♀, USNM ENT 01443825; 01443649); …, alrededores de la Estación, 9,564 652, -83,707899, 2600 m, 20.viii.1995, A. Picado (6♁, INBioCRI 002338183; 2338185); MEXICO. Chiapas, San Cristóbal de las Casas, L. C. Huitepec, 16,736 812, -92,637332, 2510 m, 26.ii.2011, P. Sagot (13♁, 5♀, ECO-TAP-E); …, 11.ii.2009, P. Sagot (1♁, ECO-TAP-E 1073); 8.viii.2009, P. Sagot (1♁, ECO-TAP-E 24469); 20.iii.2010, P. Sagot (1♀, ECO-TAP-E 24468); 12.x.2009, P. Sagot (1♁, ECO-TAP-E 24424); Mexico city, 19,431 447, -99,131481, 2675 m, 1.vii.1936, Meyer (2♁, 1♀, AMNH); …, (2♁, 2♁, IRC); Desierto de los Leones, 19,305 81, -99,303274, 2800 m, 7.vi.1961, George W. Byers (7♁, Berkeley University Collection); …, E. Toluca, 19,308 081, -99,680517, 2850 m, 31.vii.1962, U. Kansas Mexico exploración (2♁, 4♀, Berkeley University Collection)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 115-117, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Williston, S. W. (1891 - 1892) Fam. Syrphidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Zoologia- Insecta-Diptera, 3, 57 - 72 (December 1891), 73 - 78 (May 1892).","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Thompson, F. C., Rotheray, G. E. & Zumbado, M. A. (2010) Family Syrphidae. In: Brown, B. E. (Ed.), Manual of Diptera of Central America. NRC Press, Ottawa, pp. 763 - 792.","Aldrich, J. M. (1905) A catalogue of North American Diptera. Smithsonian miscellaneous collections, 46 (2), 1 - 680.","Williston, S. W. (1907) Class I, Hexapoda. Order IV, Diptera. Dipterological notes Journal of the New York Entomological Society, 15, 1 - 2.","Coquillett, D. W. (1910) The Type species, of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499 - 647. https: // doi. org / 10.5479 / si. 00963801.37 - 1719.499","Curran, C. H. (1937) The Neotropical species of Melanostoma and allies (Syrphidae: Diptera). American Museum Novitates, 926, 1 - 4.","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Papavero, N. & Ibanez-Bernal, S. (2003) Contribution to a history of Mexican Dipterology. Part II. The Biologia CentraliAmericana. Acta Zoologica Mexicana, 88, 143 - 232. https: // doi. org / 10.21829 / azm. 2003.88881793"]}

Argentinomyia festiva (Fluke, 1945) (Figs 32, 33 and 94D) Rhysops festiva Fluke, 1945: 10. Type locality: Ecuador. Tungurahua, Baños, 1800m. HOLOTYPE Male AMNH (Examined). Refs.: Fluke, 1945: 26 (Fig. 13, head, lateral view, male) 28 (Fig. 42, male abdomen) Rhysops festivus. Thompson et al. 1976, 43 Argentinomyia festiva. Martín-Armijos et al. 2017: 168 Type material. HOLOTYPE. Adult Male, ECUADOR. Tungurahua, Baños, 1800 m, 4.vii.1938, W. C. MacIntyre (AMNH). Length (n= 3): Body, 8.5–9.1 mm; Wings, 8.3–8.9 mm. Diagnosis. Facial pubescence golden and very thick, face heavily golden pollinose, a shining area just dorsal the antennae, down the middle of the face with dorsal part forked by a broad dash of pollen, which reaches to the base of very low but elongate tubercle; mesonotum bronze reticulate, heavily coated with yellowish pollen and three dark brown vittae on anterior half, the median one slender; costal cell bare on basal 1/3; abdominal with three pairs of yellow vittae on 2 nd to 4 th terga of male, 2 nd to 5 th terga of female, maculae on 3rd and 4th terga reaching from the base to the apical 1/4 and 1/3, 5 th tergum with a pair of small rounded maculae; metafemur black, only the narrow yellow basally 3/4. Redescription. MALE. Head (Fig. 32A): Face heavily golden pollinose, a shining area just dorsal the antennae, down the middle of the face with dorsal part forked by a broad dash of pollen, which reaches to the base of very low but elongate tubercle, also a shining area from the eyes to the oral angles; pile everywhere yellow whitish except for a few black hairs near the frontal ocellus. Gena shining black with thin, more whitish pollen, pile white. Ocellar triangle black with thin yellowish pollen, yellowish-white pilose. Occiput with yellow hairs dorsal, the long cilia yellow-white, pollen and paler pile ventrally. Antennae orange-yellow, relatively short, ratio 1.2:1.9:1.3, scape and basoflagellomere subequal, the pedicel a little more than 1/2 as long as the basoflagellomere; arista yellow at the base and dark on apical 1/3, black pilose. Thorax (Figs 32B–C): Bronze reticulate, mesonotum rather heavily coated with yellowish pollen; with three dark brown vittae on anterior half, the median one slender. Pleura yellow whitish pollinose; pile everywhere yellow. Scutellum aeneous, the disc slightly rugose, the apex with two longer pale hairs. Wing (Fig. 32C): Hyaline, stigma brown; microtrichose, except costal cell bare on basal 1/3; calypter wholly whiteyellowish; plumula whitish, halter whitish, capitulum slightly yellowish. Legs (Fig. 32C): Yellowish red to brown; metafemur and tibia brown except the narrow base and tip yellow whitish; pro- and mesofemora as well as tibia orange, mesofemur slightly brown on anterior edge, pro and mesotibiae mostly yellow, tarsi orange, pro-metatarsus brown anteriorly; pile mostly yellowish, black on metatibia and dorsal sides of all the tarsi. Abdomen (Fig. 32B): Opaque black, the sides shining, three pairs of yellow vittae on 2 nd to 4 th terga of male, 2 nd to 5 th terga of female, lateral maculae on 2 nd tergum reaching from the basal corners along the sides to the apical 1/5, maculae on 3 rd and 4 th terga reaching from the base to the apical 1/4 and 1/3, 5 th tergum with pair of small rounded maculae; pile mostly yellow, only black on maculae; male genitalia: surstylus in lateral view (Fig. 33A) with dorsal and ventral margins approximately of the same width in the whole length, elongated, at least two times longer than broad; hypandrium in ventral view (Fig. 33C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 33C) with apex acute. FEMALE (Figs 32D–F). Similar to male except for usual sexual dimorphism and differing in Frontal triangle is wide with large transversal dark brown pollinose maculae, including a narrow short brown vitta in front of the lunule. Mesonotum more shining copper bronze, with golden vittae. The abdomen is more oval, semi-shining on disc, highly polished alongside; yellow spots on 2 nd tergum are comparatively wider than in male, pro and mesolegs yellow. The female of A. lanei differs from A. Festiva by the frontal triangle with two rounded maculae, mesonotum punctulate, shining bronze to aeneous, 3 rd to 4 th terga with a pair of oblique maculae. Taxonomic notes. Argentinomyia festiva is similar to A. fastigata and A. lanei, differing by the following combination of characters: Facial pubescence golden and very thick, face heavily golden pollinose, a shining area just dorsal the antennae, down the middle of the face with dorsal part forked by a broad dash of pollen, which reaches to the base of very low but elongate tubercle (Figs 32A, C, D-F); mesonotum bronze reticulate, heavily coated with yellowish pollen and three dark brown vittae on anterior half, the median one slender (Figs 32B, E); costal cell bare on basal 1/3; metafemur black, only the narrow yellow basally3/4 (Figs 32A, C, D-F); abdominal with three pairs of yellow vittae on 2 nd to 4 th terga of male, three pairs on 2 nd to 5 th terga of female, maculae on 3rd and 4th terga reaching from the base to the apical 1/4 and 1/3, 5 th tergum with a pair of small rounded maculae (Figs 32B, E). Based on males, A. festiva differs from A. fastigata and A. lanei in having the surstylus in lateral view (Fig. 33A) with dorsal and ventral margins approximately of the same width in the whole length, elongated, at least two times longer than broad [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad in A. lanei (Fig. 41A); surstylus with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad in A. fastigata (Fig. 31A)]; hypandrium in ventral view, narrowed laterally towards the apex in A. festiva (Fig. 33C) and A. lanei (Fig. 41C) [versus hypandrium expanded laterally towards the apex in A. fastigata (Fig. 31C)]; aedeagal lobe in ventral view with apex acute in A. festiva (Fig. 33C) and A. lanei (Fig. 41C) [versus aedeagal lobe circular, with apex rounded in A. fastigata (Fig. 31C)] (see “differential diagnosis” under each species or key). Comments. The Holotype specimen of R. festiva is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue. 10021709. Biology. Label data suggest that A. festiva visits flowers of Coffea arabica (Rubiaceae) and Taraxacum officinale (Asteraceae). Geographical range. Argentinomyia festiva (n= 20) is distributed in Argentina * (Bermberg), Bolivia * (Cochabamba, La Paz), both slopes of Central Cordillera in Colombia * (Antioquia) and Ecuador (Tungurahua, Pichincha), Eastern slope of Eastern Cordillera in Colombia (Cundinamarca) and Perú * (Huanuco) (Fig. 94D). The species is endemic to the Northern Andes domain, distributed at middle and high altitudes (1500–2810 m) in the provinces: Cauca (2100–2810 m), Magdalena (1500 m), North Andean Páramo (1800 m). Non-type material examined. ARGENTINA. Bermberg, -34,663428, -58,841536, 30 m, 4.x.1927, R.C. Shannon (1♀, USNM ENT 01443817). BOLIVIA. Cochabamba, Campero, 6km al sur de la Alquile a Sucre, - 18,251 667, -65,218333, 2428 m, 1.iv.1999, P. Hibb (1♀, USNM ENT 00023642); La Paz, NE de Santa Fe Colonia Luayza, Entel (Telephone) Antenna, -15,77648, -67,595115, 1900 m, 13.iv.2001, A. Norrbom (3♁, USNM ENT 00055173, 00055171, 00055172); Nor Yungas arriba Coroico, Cerro Uchumachi, lower slope 2°, forest and shrubs, -16,197498, -67,71861, 2048 m, 4.v.2001, A. Norrbom (3♁, USNM ENT 00056122–23, 00055190). COLOMBIA. Antioquia, La Ceja, Alto de La Unión, en Forest secundario, 6.016132, -75.370182, 2500 m, Net, 10.ix.2008, A. L. Montoya (1♁, CEUA 47452); Rionegro, Cabeceras, Finca La Escondida, 6.140929, -75.411164, 2100 m, 17.v.2013, D. Caraballo, Manual (1♁, CEUA 114295); Caldas, Vereda La Clara, 6,051 67, -75,62194, 1840 m, Van SomerenRydon (VSR) Fish, 6-8.xii.2010, Y. Durango, H. Areiza, D. Carvajal (3♁, TdeA 1990-1992); Cundinamarca, Silvania, Vereda Azafranal, 4.342730, -74.404917, 1500 m, Net, en Coffea arabica, 6.xii.2009, N. Flores (1♀, UNAB); ECUADOR. Pichincha, Malchigui, 0.045695, -78.350088, 2643 m, 10.iv.2011, S. López (1♁, QCAZ 103706); -0.069732, -78.459566, 2810 m, 27.i.2000, in top of flower Taraxacum officinale, C. Enríquez (1♁, QCAZ 103705); Yaruqui, -0.167036, -78.318787, 2583 m, 23.x.2005, J.M. Viera (1♁, QCAZ 103704); 6.i.1986, G. Correa (1♁, QCAZ 103707). PERÚ. Huanuco, NE Huanuco, -9,337663, -76,24039, 2500 m, 17.ix.1954, W.M. Mathes (1♀, CAS)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 60-62, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898"]}

Argentinomyia nigrans (Fluke, 1945) (Figs 53, 54 and 92D) Rhysops nigrans Fluke, 1945: 8. Type locality: Brazil. Santa Catarina. Nova Teutonia. HOLOTYPE Female AMNH (Examined). Refs.: Fluke, 1945: 26 (Fig. 8, head, lateral view, female); 1957: 279 (Figs 111–112, male genitalia). Rhysops nigrans. Fluke, 1957: 266 Rhysops nigrans. Thompson et al. 1976: 44 Argentinomyia nigrans. Montoya, 2016: 461 Rhysops funerea Hull, 1949: 74. Type locality: Brazil: Santa Catarina: Nova Teutonia. HOLOTYPE Male Hull collection, examined from photographs). n. syn. Rhysops funereus. Thompson et al 1976: 43 Type material. HOLOTYPE. Adult Female, BRAZIL. Santa Catarina, Nova Teutonia, 857 m, 1.vi.1936, F. Plaumann (AMNH). ALLOTYPE. BRAZIL Label with the same data as Holotype except 20.vii.1937 (1♁, AMNH). PARATYPE. BRAZIL. Santa Catarina, Bom Retiro, 845 m, 21.i.1929, F. Plaumann (1♁, 3♀, WIRC; 1♁, AMNH; 1♀, USNM ENT 00058843);... vi and vii.1936 and 1937, F. Plaumann (10♀, AMNH). METATYPE: BRAZIL. Santa Catarina, Nova Teutonia, 530 m, 21.vi.1936, F. Plaumann (3♀, AMNH). Length (n= 5): Body, 9.5–10.1 mm; Wings, 6.2–7.1 mm. Diagnosis. Face perpendicular without prominent grooves above the tubercle; wing only tinged with brown; abdomen black with small yellow to red triangular maculae on basal corners of the 2 nd tergum, reaching over the sides and 1/2 of segment length; femora and tarsi black, surstylus with dorsal and ventral margins similar (squarelike); hypandrium narrowed laterally towards the apex; aedeagal lobe circular, apex rounded. Redescription. MALE. Head (Fig. 53A): Face nearly straight, tubercle low with two very faint transverse depressions; white pollinose, shining on a broad mid-vitta, around the oral edges, a narrow band dorsal the gena from the eyes to the oral edge, an arc dorsal the antennae and thinly pollinose from antennae down the slopes of the face almost to the flat area opposite tubercle, pile white. Frontal triangle dark with a broad opaque band just ventral the ocelli, gradually fading into white pollen in the depression, pile black. Gena shining black with thin, more whitish pollen, pile white. Ocellar triangle semi-shining black with black pile. Occiput whitish pollinose and white pilose on lower two-thirds, black pilose on dorsal 1/3. Antennae brownish-black, short, ratio 1.0:1.3:1.9, reaching to the tubercle, scape shorter than basoflagellomere, basoflagellomere a little shorter than scape and pedicel, reddishyellow but darker on outer sides and dorsal on basoflagellomere, arista yellow at base and dark on apical 1/3, pilose. Thorax (Figs 53B–C): Black, mesonotum semiopaque on disc, more shining alongside, pile longer and brownishyellow; with two median white-pollinose vittae on anterior half, pile whitish and very short, dark brown alongside beyond the notopleural suture. Pleura lightly white pollinose, white pile; notopleural tubercle prominent. Scutellum blue-black, shining, slightly rugose, pile short and pale; fringe thin and white. Wing (Fig. 53C): Tinged with brown, stigma elongate brown, microtrichose, except cell c bare basal 1/2, cells bm bare on basal 1/3 and r only slightly bare basally, cell cua bare on anterior 1/3 or more, very light clouds at the fork of second and third veins and on anterior cross-vein, calypter wholly brown; plumula white; halter yellow, capitulum slightly brown. Legs (Fig. 53C): Dark brown to black; femur reddish-yellow apically; tibia reddish yellow on basal 1/3; pile all black, even the mat under the metatarsus. Abdomen (Fig. 53B): Semi-shining black to opaque, yellowish-red triangles on basal corners of the 2 nd tergum reaching over the sides and 1/2 of segment length, pile longer alongside, short, black and appressed on discs of tergites; sterna shining black, yellow and long pilose; male genitalia: surstylus in lateral view (Fig. 54A) with dorsal and ventral margins similar (square-like); hypandrium in ventral view (Fig. 54C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 54C) circular, apex rounded. FEMALE (Figs 53D–F). Similar to male except for usual sexual dimorphism and differing in the frontal triangle pollen whitish along the eyes, brownish elsewhere. Taxonomic notes. Argentinomyia nigrans is similar in appearance to A. plaumanni sp. nov. differing from it by having the wing tinged with brown (Figs 53 B-C, E-F); 2 nd tergum with a pair of triangular yellow maculae on basal corners reaching over the sides and 1/2 of segment length, others abdominal segments without maculae (Figs 53B, E). In A. plaumanni sp. nov. the wing slightly yellowish (Figs 61 B-C, E-F); 2 nd tergum with a pair of basolateral elongate yellow maculae reaching the apical 3/4 of the segment length; 3 rd tergum with a pair of basal elongated maculae reaching apical 2/3, but not reaching lateral margin; 4 th and 5 th terga black (Figs 61B, E). Based on males, A nigrans differs from A. plaumanni sp. nov. in having the surstylus in lateral view (Fig. 54A) with dorsal and ventral margins similar (square-like) [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. plaumanni sp. nov. (Fig. 62A)] (see “diagnosis” under each species or key). Remarks. Hull (1949) recognized that A. funerea is similar to A. nigrans and established that A. nigrans differs from it by the combination of the following characters: the male with an obscure yellowish macula on 2 nd tergum, more indistinct than in the female, tibia reddish yellow on basal 1/3, wings only tinged with brown instead of being deeply brownish as in A. funerea. Cooper & Cumming (1993) did not register a type specimen for A. funerea housed at the CNC. The examination of the type specimen from photographs provided by the CNC revealed that indeed it is housed there and corresponds to a specimen of A. nigrans (Figs 54D–G), therefore, A. funerea is formally synonymized under A. nigrans. Comments. The Holotype specimen of R. nigrans is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue. 10021029. Biology. Label data suggest that A. nigrans visits flowers of “Granadilla” Passiflora edulis (Passifloraceae). Geographical range. Argentinomyia nigrans (n= 194) is the abundant species in collections, distributed in Argentina * (Misiones), Serra de Fartura and Serra de Mantiqueira in Brazil (Minas Gerais, Paraná, Rio Grande do Sul, Santa Catarina, S„o Paulo) and Eastern slope of Western Cordillera of Colombia * (Boyacá) (Fig. 92D). The species is present at low and middle altitudes (233–2150 m) in the following biogeographical domains and provinces: Northern Andes: Magdalena (871 m); Paraná (233–2150 m): Araucaria Forest (914–1200 m), Atlantic Forest (1662–2150 m) and Paraná Forest (233–857 m). Non-type material examined. ARGENTINA. Misiones, Iguazú, -25.655778, -54.471099, 233 m, 4-10.x.1927, R.C. and E.M. Shannon (1♀, USNM ENT 01443753). BRAZIL. Minas Gerais, 16Km a SE de Itamonte Pq Nac Itamonte, -22,368504, -44,778372, 2150, 26.v.2011, M. G. Hermes (3♀, CEUFLA); Paraná, Curitiba, -25,424429, -49,265382, 914 m, 1975-1976, Malaise trap, H. S. Telford (13♁, 11♀, DZUP 46593; 46579; 46600; 46589; 46602; 46603; 46604; 46598; 46578; 46575; 46576; 46567; 46573; 46571; 46594; 46583; 46584; 46566; 46580; 46587); 934 m, 28.x.1975, A. Sakakibara (2♀, DZUP 4657046595); Refugio de vida silvestre dos campos de Palmas, - 26,533 807, -51,599362, 1200 m, A. Couto (4♀, DZUP); Curitiba, Cto Politécnico, -25,436585, -49,255367, 962 m, 21.vi.2005, M. G. Hermes (1♀, CEUFLA); Rio Grande do Sul, Bar„o de Cotegipe, -27,6208, -52,3797, 687 m, 15.i.1967, F. Giacomel (2♀, DZUP 46562-63) (Marinoni et al. 2007); Santa Catarina, Nova Teutonia, -27,183334, -52,383334, 500 m, 1.vi.1964, F. Plaumann (35♁, 44♀, USNM ENT 01406596; 01406443; 01406435; 01406436; 01384750; 01406454; 01406585; 01406446; 01406450; 01406425; 01406452; 01406580; 01406440; 01406442; 01406579; 01406611; 01406437; 01406453; 01443775; 01406605; 01384767; 01384677; 01384330; 01384727; 01384699; 0134683; 01384477; 01384441; 01384720; 01384806; 01384855; 01384783; 01384877; 01384743; 01384670; 01384891; 01384820; 01384835; 01383238; 01383141; 01443763; 01384907; 01384666; 01383952; 01384872; 01383522; 01384886; 01384694; 01384662; 01383987; 0138772; 01384640; 01383363; 01383681; 01384771; 01406600; 01406606; 01406448; 0146595; 01406591; 01406449; 01406610; 01406415; 01406441; 01406590; 01406438; 01406601; 01443758; 01443762; 01443752); 6.i.1969, F. Plaumann (3♁, 5♀, AMNH); 400 m, 3.v.1937, F. Plaumann (9♁, 11♀, BMNH); 3.v.1937, F. Plaumann (1♁, 1♀, FMNH); Bom Jardim da Sierra, - 28,137 522, -49,966651, 1089 m, 5.vi.2006, A.J.C. Aguiar, A. Martines, L.R.R. Faria Jr (1♀, CEUFLA); São Paulo, Campo do Jardão, -22,742928, -45,596348, 1662 m, 1.i.1954, J. Lane (6♁, 1♀, WIRC). COLOMBIA. Boyacá, Buena Vista, Vereda Patiño, 5.650832, -74.286905, 871 m, 8.iv.2004, W. Avila, Net en “Granadilla” Passiflora edulis (1♀, UNAB)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 92-94, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Montoya, A. L. (2016) 36 Family Syrphidae. In: Wolff, M., Nihei, S. S. & de Carvalho, C. J. B. (Eds.), Catalogue of Diptera of Colombia. Zootaxa, 4122 (1), 457 - 537. https: // doi. org / 10.11646 / zootaxa. 4122.1.39","Cooper, B. E. & Cumming, J. M. (1993) Diptera types in the Canadian National Collection of Insects. Part 2. Brachycera (exclusive of Schizophora). Research Branch Agriculture Canada Publication 1896 / B. Research Branch Agriculture Canada, Ottawa, 105 pp. [also published electronically at http: // www. nadsdiptera. org / Catalogs / CNCtypes / Part 2. htm]","Marinoni, L., Morales, M. N. & Spaler, I. (2007) Illustrated clave for the genera of Syrphinae (Diptera, Syrphidae) of occurrence in the South of Brasil. Biota Neotropical, 7, 145 - 160. https: // doi. org / 10.1590 / S 1676 - 06032007000100019"]}

Argentinomyia berthae (Lima, 1946) Proposed standard English name: Berthae’s long-antennae flower fly. (Figs 18, 19 and 92B) Rhysops berthae Lima, 1946:155. Type locality: Brazil, Rio de Janeiro. HOLOTYPE Male ENA. Refs.: Lima, 1946:155 (Fig. 1, head; Fig. 2, abdomen; Fig. 3, wing) Rhysops berthae. Thompson et al. 1976, 42 Argentinomyia longicornis. Miranda (2017) misidentification NEOTYPE. Adult male, pinned, deposited at the Instituto Nacional de Pesquisas da Amazônia. Original label: “ BRAZIL, Amazonas, Igarapé Acará, Reserva Florestal Adolpho Ducke” / “ -2.963343, -59.922834, 110 m, Malaise suspensa sobre lâmina d’água” / “ 30.ix.2001, Vidal, Jo„o Leg.” “ NEOTYPE / Argentinomyia berthae / Lima, 1946 ” [blue except the first line] (INPA-DIP 000095). Length (n= 2): Body, 6.1–6.5 mm; Wings, 5.8–6.1 mm. Diagnosis. Face with tubercle low, one transversal groove; frontal triangle with a transversal brownish pollinose fascia, pile black; basoflagellomere longer, a little longer than scape and pedicel together; scutum blue metallic without pollinosity, except white pollinose on lateral sides; metafemur brown except yellow on basal 1/4; abdomen narrowed toward the base, with two pairs of oval maculae on 2 nd to 3 rd terga; surstylus with dorsal margin slightly concave and ventral margin slightly convex; hypandrium narrowed laterally towards the apex; aedeagal lobe circular, apex rounded. Redescription. MALE. Head (Fig. 18A): Face black, sides of face whitish-pollinose and pilose; tubercle low with a single transversal groove. Frontal triangle with a transversal brownish pollinose fascia, pile black. Gena orange, covered with white pollen. Ocellar triangle opaque, with a brown cross band which is narrow in the anteromedial half, without a point projecting downwards in the middle, yellow whitish pilose. Occiput white pollinose and pilose. Antennae yellow, short, ratio 1.5:1.0:2.3; basoflagellomere brown dorsally, longer than pedicel; arista yellow at base and dark on apical third, pilose. Thorax (Figs 18C): Black aeneous, scutum blue metallic without a pollinose pattern, except white pollinose on lateral sides of notopleuron and postpronotum. Pleura shining aeneous, white pollinose and pilose, sparse. Scutellum shining greenish-brassy to aeneous, yellow pilose. Wing (Fig. 18C): Hyaline, stigma yellow, membrane microtrichose, except cell c bare on basal 1/4, cells bm bare on basal 1/2 and cell r bare on basal 1/2, cell cua bare on anterior 1/3; calypter wholly white; plumula yellow; halter yellow, capitulum yellow. Legs (Fig. 18C): Yellow, pro- and mesolegs light yellow; pro- and mesofemora yellow, metafemur brown except yellow on basal 1/4; metatibia brown, pro- and mesocoxae yellow, metacoxa brown; pile yellow. Abdomen (Figs 18C): Abdomen narrowed toward the base, 2 nd tergum with a pair of lateral rounded maculae, covered by the entire lateral margin but attenuated posteriorly; 3 rd tergum with a pair of elongate, slightly oblique, yellow maculae, reaching the lateral margin and slightly touching each other toward the middle; 4 th and 5 th terga black; yellow pilose, with blackish pile on black areas, male genitalia: surstylus in lateral view (Fig. 19A) with dorsal margin slightly concave and ventral margin slightly convex; hypandrium in ventral view (Fig. 19C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 19C) circular, apex rounded. FEMALE (Figs 13D–F). Similar to male except for usual sexual dimorphism and differing in the frontal triangle with a transversal black macula, 4 th tergum with a pair of oblique, yellow maculae. Taxonomic notes. Argentinomyia berthae is similar in appearance to A. festiva and A. lanei, differing from them by the following combination of characters: Face with tubercle low, one transversal groove; frontal triangle with a transversal brownish pollinose fascia, pile black (Figs 18A, C, D, F); basoflagellomere longer, a little longer than scape and pedicel together (Figs 18A, C, D, F); scutum blue metallic without pollinosity, except white pollinose on lateral sides (Figs 18B, E, F); metafemur brown except yellow on basal 1/4 (Figs 18B, D, F); abdomen narrowed toward the base, with two pairs of oval maculae on 2 nd to 3 rd terga (Figs 18B, E, F-G). Argentinomyia festiva differs from A. berthae by the presence of a golden and very thick facial pubescence, frontal triangle yellow pilose, face with dorsal part forked by a broad dash of pollen reaching to the tubercle base (Figs 32A, C, D-F); metafemur black, only yellow on basal 3/4 (Figs 32A, C, D-F); abdomen with three pairs of yellow vittae on 2 nd to 4 th terga of male, 3 rd tergum with a pair of elongate, slightly oblique, yellow maculae, reaching the lateral margin and slightly touching each other toward the middle (Figs 32B, E). In A. lanei the female frontal triangle with two rounded maculae (Figs 40 D-F); mesonotum punctulate, shining bronze to aeneous, with brownish vittae on anterior half (Figs 40B, E); pro- and mesolegs light orange-brown; metafemur brown except for yellow on basal 1/2 and the tip (Figs 40A, C, D-F); abdomen with a pair of oblique maculae on 3 rd to 4 th terga (Figs 40B, E). Based on males, A. berthae differs from A. festiva and A. lanei in having the surstylus in lateral view (Fig. 19A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length, elongated, at least two times longer than broad in A. festiva (Fig. 33A) and shorter than broad in A. lanei (Fig. 41A)]; aedeagal lobe in ventral view (Fig. 16C) circular, apex rounded [versus aedeagal lobe, with acute apex in A. festiva (Fig. 33C) and A. lanei (Fig. 41C)] (see “differential diagnosis” under each species or key). Remarks. Argentinomyia berthae was originally described from a unique male, collected in Rio de Janeiro, Brazil by Dr. Bertha Luz, Nº9790 and deposited in the Coleҫao da Escola Nacional de Agronomia (ENA, now Colección Costa Lima, Universidad Federal Rural de Río de Janeiro) (Lima 1946). However, attempts to trace the type material at this institution failed, suggesting that the original material was destroyed, or lost (Dr. Francisco Racca Filho of Colección Costa Lima de la Universidad Federal Rural de Río de Janeiro and Dr. José Pujol Luz of Universidade de Brasília: personal communication). Another relevant issue about A. berthae is that despite the original description was brief but illustrated the frontal view of head as well as the dorsal habitus of abdomen and wing (Lima 1946, figs 1-3), the lack of a detailed diagnosis resulting in uncertain status, with the erroneous identification of this species as the widespread A. longicornis by Miranda (2017; http://syrphidae.inpa.gov.br/syrphidae/index.php/generos/syrphi?id=63). This taxonomic confusion could be caused by the scarcity of specimens of this rare species in collections as well as not being included in a key since its description; indicating indeed the difficulty to recognize it (ICZN 1999, Art. 75.3.1). Therefore, since the name-bearing original type specimen was confirmed to be lost as well as the uncertain taxonomic status, the Neotype designation is deemed needed to ensure the stability of this taxon (ICZN 1999, Art. 75.3.4). We selected Neotype specimen that are consistent with the original description and illustrations provided by Lima (ICZN 1999, Art. 75.3.5). To fulfill the qualifying criteria, we provide unique identifying information associated with the Neotype, including redescription, taxonomic notes, habit photographs as well as the illustration of male genitalia, with the inclusion in a new key as well as a comparative diagnosis to clarifying the taxonomic status (ICZN 1999, Art. 75.3.1–75.3.3). We designated a Neotype for A. berthae, using a male specimen collected in Brazil (Amazonas, Igarapé Acará, Reserva Florestal Adolpho Ducke), in a location closer to the natural location of the original type (ICZN 1999, Art. 75.3.6). The Neotype is deposited in the Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil (ICZN 1999, Art. 75.3.7). Non-type material examined. BRAZIL. Rio de Janeiro, -22.763426, -43.688251, 27 m, i.1939, R.C. Shannon Leg. (1♀, USNM ENT 01443843; headless). VENEZUELA. Amazonas, T. F. Amazonas, Basecamp, Cerro de la Neblina, T.F.A., 0.804185, -66.006645, 2645 m, Malaise trap over a small stream at the side of basecamp, 20- 24.iii.1984, O. Flint and J. Louton (1♀, USNM ENT 01443843) ”. Geographical range. Argentinomyia berthae (n= 2) is distributed in Brazil (Amazonas and Rio de Janeiro) and Venezuela * (Cerro de la Neblina) (Fig. 92B). The species is present at low and middle altitudes (27 and 2655 m) in the following biogeographical domains and provinces: Amazonian: Imerí (2655 m); Paraná: Atlantic Forest (27 m)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 38-41, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Lima, A. M. da C. (1946) Nova especie do genero Rhysops Williston (Diptera: Syrphidae). Boletim da Sociedade Brasileira de Agronomia, 1946, 155 - 156.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature, 4 th ed. International Trust for Zoological Nomenclature (The Natural History Museum), London, 306 pp. [http: // www. iczn. org / iczn / index. jsp]"]}

Argentinomyia tropica (Curran, 1937) (Figs 86, 87 and 91A) Melanostoma tropicum Curran, 1937: 3. Type locality: Brazil. S„o Paulo, Campos do Jord „o. HOLOTYPE Male AMNH (Examined) Melanostoma tropicum. Frey, 1946: 157 Rhysops tropicus. Thompson et al. 1976: 44 Argentinomyia tropica. Montoya 2016: 461 Argentinomyia tropica. Montoya et al. 2017: 395 Argentinomyia tropica. Marín-Armijos et al. 2017: 168 Argentinomyia tropica. Miranda, 2017 (Distributional record from the Brazilian Amazon) Type material. HOLOTYPE. Adult Male, BRAZIL. S„o Paulo, Campos do Jard„o, 179 m, 23.i.1936, F. Lane (AMNH). PARATYPE. Adult Male. BRAZIL. Label same data as Holotype (AMNH). Length (n= 4): Body, 8.6–10.1 mm; Wings, 8.6–9.0 mm. Diagnosis. Face less produced ventrally, the sides thickly coated with cinereous yellow pollen; legs reddish, metafemur brown, only reddish or yellowish on basal 1/2, metatibia yellow on basal and apical 1/4; abdomen with three pairs of reddish maculae, on 3 rd and 4 th terga extending from the base to apical 1/4 and 1/3; surstylus with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; aedeagal lobe oval, apex rounded. Redescription. MALE. Head (Fig. 86A): Face less produced ventrally, tubercle low but distinct, narrowly separated from the oral anterior margin; sides of face thickly cinereous yellow pollinose, the median 1/5 shining black and not at all ridged or grooved, pile pale yellowish. Gena cinereous shining with thin, more whitish pollen, pile white. Ocellar triangle brownish-yellow pollinose with a shining, roughened area dorsal the antennae, pile black; vertical triangle thinly brownish pollinose and blackish pilose. Occiput cinereous pollinose, pile whitish, the dorsal occipital cilia fine and black. Antennae reddish, short, ratio 1.4:1.2:1.5, basoflagellomere brown dorsal and apically; basoflagellomere 1/2 longer than broad as long as the scape and pedicel. Antennae reaching to a little ventral the middle of face; arista brown, pilose. Thorax (Figs 86B–C): Aeneous, mesonotum shining, yellow pilose; with a pair of very broad, narrowly separated pale pollinose vittae in the middle on anterior half and some lights with thin brownish pollen. Pleura thinly grayish-brown pollinose, yellow pilose. Scutellum shining, with long yellowish to black hairs, with a few shorter yellow hairs intermixed; fringe long and yellow. Wing (Fig. 86C): Cinereous hyaline with orange-yellow tinge, stigma orange-yellow; microtrichose, except cell c bare basal 1/5, cells bm, r and cua bare on basal 1/3; calypter wholly white-yellowish. Legs (Fig. 86C): Reddish to orange, pro- and mesolegs yellow; metafemur brown, except reddish on almost the basal 1/2, pile reddish, black on brown portions and apical segments of tarsus. Abdomen (Fig. 86B): Long and slightly spatulate, being narrowest at the end of the 2 nd tergum and widest at the end of 3 rd; black with three pairs of reddish maculae shining with the sides and anterior border obscurely reddish, on 2 nd tergum opaque black with the lateral 1/6 reddish, on 3 rd tergum opaque black with maculae extending from the base to the apical 1/4, gently tapering posteriorly and broadly separated from the lateral margins by a sub-shining vitta; maculae on 4 th tergum extend to the apical 1/3 and the posterior margin is broadly subshining; 5 th tergum very short and wholly shining; pile short and black, long and yellowish on sides of 1 st and 2 nd and basal 1/2 of 3 rd tergum; sterna metallic brownish red with black pile; genitalia brownish-red and with thin yellowish-brown pollen; male genitalia: surstylus in lateral view (Fig. 87A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium in ventral view (Fig. 87C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 87C) oval, apex rounded. FEMALE (Figs 86D–F). Similar to male except for usual sexual dimorphism and differing in frontal triangle blue-black with a broad transverse golden pollinose band in the depression, black pilose, 5 th tergum includes a pair of fasciate maculae. Female of A. tropica is similar to A. luculenta differing in the face more perpendicular and less produced below. Frontal triangle blue-black with a broad transverse golden pollinose band in the depression, black pilose. Thorax with short yellowish pile, on a background of thin brownish pollen. Taxonomic notes. Argentinomyia tropica is a black species, face less produced ventrally, the sides thickly coated with cinereous yellow pollen (Figs 86A, C, D, F); legs reddish, metafemur brown, only reddish or yellowish on basal 1/2, metatibia yellow on basal and apical 1/4 (Figs 86A, C, D, F); abdomen with three pairs of reddish maculae, on 3 rd and 4 th terga extending from the base to apical 1/4 and 1/3 (Figs 86B, E). In A. luculenta the face is more perpendicular and less produced ventrally; frontal triangle blue-black with a broad transverse golden pollinose band in the depression, pile black (Figs 47A, C, D, F); the pile of thorax short and yellowish on a background of thin brownish pollen (Figs 47B, E); metafemur yellow on basal 1/2 (Figs 47A, C, D, F); abdominal maculae rectangular (Figs 47B, E), slightly similar to the female of A. tropica, except on 3 rd and 4 th terga, where they reach apical 1/3 and 1/2 (Figs 47E), respectively. Based on males, A. tropica differs from A. luculenta in having the aedeagal lobe in ventral view (Fig. 87C) oval, with apex rounded [versus aedeagal lobe with apex acute in A. luculenta (Fig. 48C)] (see “diagnosis” under each species or key). Comments. The Holotype specimen of M. tropicum is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue.10020983. Biology. Label data and field observations suggest that A. tropica visits flowers of Libanothamnus humbertii (Asteraceae). Geographical range. Argentinomyia tropica (n= 50) is distributed in Argentina (Tucumán), Serra da Mantiqueira and Serra Paranapiacaba in Brazil (Brazilian Amazon, Sao Paulo, Paraná), Eastern slope of West and Eastern Cordilleras in Colombia * (Antioquia), both slopes of Central Cordillera in Colombia and Ecuador * (Tungurahua) and Cordillera de Mérida in Venezuela * (Merida) (Fig. 91A). The species is present at low, middle and high altitudes (179–3600 m) in the following biogeographical domains and provinces: Northern Andes (231–3600 m): Cauca (3600 m), Guajira (3500 m), Magdalena (2034–3150 m), North Andean Páramo (2300 m), Sabana (231 m); Paraná (179–1274 m): Araucaria Forest (867–1274 m), Atlantic Forest (179–761 m), Paraná Forest (826 m); Chacoan: Chaco (526–1426 m). Non-type material examined. ARGENTINA. Tucumán, Burruyacu, Quebrada La Toma de Tafí Viejo, -26.499425, -64.741942, 526 m, 21.xii.1950, R.A. Golbach (3♁, WIRC); Same data, except: Villa Nougués, - 26.849763, -65.382689, 1426 m, i.1929, R.K. (2♀, WIRC). BRAZIL. Paraná, Ponta Grossa, -25.187256, - 50.143897, 867 m, ix.1945, P. Machado (1♀, WIRC). São Paulo, Campos do Jord „o, 22.742928, -45.596348, 761 m, 10.i.1936, F. Lane (2♁, 1♀, WIRC); Ribeirao Grande, Fazenda Intermontes, -24.197324, -48.424401, 826 m, 6.iv.2006, S. B. Vosgueritchian (1♀, CEUFLA). COLOMBIA. Antioquia, Andes, Santa Rita, Páramo de Santa Rita, 5,549 9389, -75,9912444, 3600 m, Net, 5.xii.2009, M. Wolff (1♀, CEUA 69642); Van Someren-Rydon (VSR) Fish, 12.viii.2010, GEUA (1♀, CEUA 87052); Belmira, Páramo Santa Inés Belmira, 6,647 5556, -75,6713333, 3150 m, Net, 1.x.2011, N. Uribe (1♁, CEUA 93084); Bello, San Félix, Las Baldías, 6,330 58, -75,645620, 3150 m, Net, 16.viii.2015, A.L. Montoya (2♀, CEUA 93082-83); …, 17.ii.2013 (1♀, CEUA 93135); Girardota, Vereda La Honda, 6,347 375, -75,4359944, 2034 m, Net, 26.ix.2009, N. Uribe (2♀, CEUA 69474–75); …, Vereda El Palmar, Secondary Forest, 6,343 502, -75,435102, 2200 m, Van Someren-Rydon (VSR) Víscera, 23.vii.2009, N. Uribe; G. Valencia (1♀, CEUA 47457); Guarne, Vereda Piedras Blancas, Parque Ecológico Piedras Blancas, cerca de quebrada, 6,291 588, -75,493345, 2200 m, Net, 5.v.2009, A.L. Montoya (1♀, CEUA 47449); Medellín, Santa Elena, Parque Ecológico Piedras Blancas, 6,295 895, -75,499891, 2470 m, Net, 15.ii.2009, A.L. Montoya (1♀, CEUA 47449); San José de la Montaña, Vereda El Congo, Sector La Laguna, 6,763 791, -75,701738, 3117 m, Páramo, Net, 21- 30.vi-5.iv.2017, A. L. Montoya; A. F. Sepúlveda (2♀, CEUA 98096, 98113); Sonsón, Vereda Nori, Cerro Nori, 5,809 667, -75,269028, 2853 m, Forest, Malaise canopy, 31.iii-7.iv.2018, A.L. Montoya; J. P. Carmona (1♀, CEUA 103433). Caquetá, Florencia, Las Brisas, 1,753 889, -75,747639, 2040 m, Malaise, Bosque Piso, 3er Túnel C 8, 1- 15.ii.2017, Y. Ramos-Pastrana (7#m, LEUA). ECUADOR. Tungurahua, Baños, -1.396949, -78.421669, 2300 m, 21.vii.1939, W. C. MacIntyre (1♁, AMNH). VENEZUELA. Mérida, Páramo Mucubaji, Laguna Negra, 8,797 016, -70,828651, 3500 m, ex flowers of Libanothamnus humbertii (89V27) 28-31.x.1989, A.L. Norrbom (1♀, USNM ENT)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 138-140, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Curran, C. H. (1937) The Neotropical species of Melanostoma and allies (Syrphidae: Diptera). American Museum Novitates, 926, 1 - 4.","Frey, R. (1946) Ubersicht der Gattungen der Syrphiden-Unterfamilie Syrphinae (Syrphinae + Bacchinae). Notulae Entomologicae, 25, 152 - 172.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Montoya, A. L. (2016) 36 Family Syrphidae. In: Wolff, M., Nihei, S. S. & de Carvalho, C. J. B. (Eds.), Catalogue of Diptera of Colombia. Zootaxa, 4122 (1), 457 - 537. https: // doi. org / 10.11646 / zootaxa. 4122.1.39","Montoya, A. L., Ricarte, A. & Wolff, M. (2017) Two new species of Quichuana Knab (Diptera: Syrphidae) from the Paramo ecosystems in Colombia. Zootaxa, 4244 (3), 390 - 402. https: // doi. org / 10.11646 / zootaxa. 4244.3.7","Marin-Armijos, D., Quezada-Rios, N., Soto-Armijos, C. & Mengual, X. (2017) Checklist of the flower flies of Ecuador (Diptera, Syrphidae). ZooKeys, 691, 163 - 199. https: // doi. org / 10.3897 / ZooKeys. 691.13328"]}

Argentinomyia occidentalis Montoya & Wolff sp. nov. Proposed standard English name: Western long-antennae flower fly. (Figs 5B, 7F, 52, 53, 90B) Type material. HOLOTYPE. Adult Male, pinned, deposited at Colección Entomológica Universidad de Antioquia. Original label: “ COLOMBIA, Antioquia, Urrao, Páramo del Sol, 6,455 8333, -76,0939” / “ 3638-3800 m, 11.iii.2009, Leg J. Duque ”. “ HOLOTYPE / Argentinomyia occidentalis / Montoya & Wolff 2023” (CEUA 47448, dissected; left antenna broken and attached on specimen label). Identified as Argentinomyia sp. 5 by Montoya 2010. PARATYPE. COLOMBIA. Antioquia, Andes, Santa Rita, Páramo de Santa Rita, Laguna Santa Rita, 5.5499389, -75.9912444, 3611-3638 m, 3-5.iii.2010, M. Wolff Leg. (1♀, CEUA 47447). Length (n= 2): Body, 8.1 mm; Wings, 7.5 mm. Diagnosis. Abdomen with a pair of thin maculae (diamond-shape) covered by gray pollen on 3 rd to 4 th terga of male, the same type of macula on 2 nd to 5 th terga of female, maculae on all segments isolated from base and apex. Surstyle bent back as a Z in shape, no more than two times longer than wide; aedeagal lobe with acute apex. Description. MALE. Head (Fig. 57A): Face shining black and somewhat brassy, golden pollinose and pilose. Gena black, golden pollinose and pilose. Frontal triangle golden pollinose, black pilose. Ocellar triangle black, brown pilose. Occiput golden pollinose. Antenna black, short, ratio 1.0:1.3:2.5, basoflagellomere at least twice as long as the pedicel and scape combined, black pilose; scape slightly curved basally, shorter than the basoflagellomere; arista black, slightly pilose. Thorax (Figs 57B–C): Metallic brownish-black, mesonotum with two faint yellow-whitish pollinose vittae on anterior half; mesonotum yellowish pilose, with lateral sides yellowish pollinose, yellow and black pile intermixed. Pleura shining black, silver dusted, except on bare area of anterior anepisternum, yellowgolden pilose. Scutellum shining, slightly rugose on apical half, pile yellow in addition to long marginal black pile. Wing (Fig. 57C): Hyaline, stigma brownish, microtrichose; calypter wholly golden yellowish; plumula, halter and capitulum yellowish. Legs (Figs 6F, 57C): Yellow to brown, yellow and black pilose, femora brown basally, except orange on apical 1/4; pro- and mesotibiae orange with a dark median ring; tarsi brown; metatibia extensively brown, only slightly orange on basal and apical 1/4. Abdomen (Figs 57B, 58D): Black metallic, bronze-green, 1 st to 2 nd terga black shining, with long lateral yellowish pile; 3 rd to 4 th terga with a pair of thin maculae cover with gray pollen, diamond shape, golden pilose, sterna metallic bronze-green with long yellow pilosity; male genitalia: surstylus yellow, in lateral view (Fig. 58A) bent back as a Z in shape, no more than two times longer than wide; hypandrium in ventral view (Fig. 58C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 58C) with acute apex. FEMALE (Figs 5B, 7F, 57D–F, 58E). Similar to male except for usual sexual dimorphism and differing in the abdomen with a pair of thin maculae covered with gray pollen on 2 nd to 4 th terga. Taxonomic notes. Argentinomyia occidentalis sp. nov. is similar to A. rex having black antennae, pro- and mesotibiae orange with a dark median ring, but differs from it by the following combination of characters: Abdomen with a pair of thin maculae (diamond-shape) covered by gray pollen on 3 rd to 4 th terga of male (Fig. 57B), the same type of macula on 2 nd to 5 th terga of female, maculae on all segments isolated from base and apex (Fig. 57E). In A. rex the abdomen with lineate yellow maculae (Figs 68B, E). Based on males, A. occidentalis sp. nov. differs from A. rex in having the surstylus in lateral view (Fig. 58A, D) bent back as a Z in shape, no more than two times longer than wide [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length, elongated, three to four times longer than broad in A. rex (Fig. 69C)]; aedeagal lobe in ventral view (Fig. 58C, D) with acute apex [versus aedeagal lobe circular with apex rounded in A. rex (Fig. 69C)] (see “diagnosis” under each species or key). Etymology. The name Argentinomyia occidentalis (nominative, adjective feminine) is derived from the Latin word occidens, occidentis (Brown 1956: 567) which refers to the western. The name is given in allusion to the western distribution in Colombia. Remarks. The type material was collected in two sampling events in the two Páramo complexes of Farallones de Citará and Páramo de Frontino, in the Western cordillera in Antioquia, Colombia (58F–G) in February 2009 and March 2010. A. occidentalis sp. nov. is predominantly distributed in mountainous areas and seems to be confined to high elevations between 3611-3800 m, where occurs in sympatry with Quichuana citara Montoya & Wolff 2017, A. altissima, Meropidia neurostigma Hippa, Platycheirus (Tuberculanostoma) antennatum (Fluke) and Styxia eblis (Hull). For a complete discussion about topographic characteristics and endemism in these type localities of the Western cordillera in Colombia see Montoya et al. (2017). Geographical range. Argentinomyia occidentalis sp. nov. (n= 2) is confined to the eastern slope of the Western Cordillera and is known only from two localities in Colombia (Antioquia) (Fig. 90B). The species is present at high altitudes (3611-3800m) in the Northern Andes domain and Cauca province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 98-101, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Montoya, A. L., Ricarte, A. & Wolff, M. (2017) Two new species of Quichuana Knab (Diptera: Syrphidae) from the Paramo ecosystems in Colombia. Zootaxa, 4244 (3), 390 - 402. https: // doi. org / 10.11646 / zootaxa. 4244.3.7"]}

Argentinomyia aurifacies Thompson & Montoya sp. nov. Proposed standard English name: Golden-faced long-antennae flower fly. Argentinomyia DR sp. 1 Thompson, unpublished (Figs 14, 15 and 95) Type material. HOLOTYPE. Adult Male, pinned, deposited at National Museum of Natural History. Original label: “ DOMINICAN REPUBLIC, La Compartición, Pico Duarte ” / “ P. N. Armando Bermúdez / 19,025 337, -70,99825, 3087m ” / “RD-255 / 1.vii.2004, D. Pérez “ HOLOTYPE / Argentinomyia aurifacies / Thompson & Montoya 2023” (USNM 00038437). PARATYPES. DOMINICAN REPUBLIC. La Vega, La Compartición, Pico Duarte ” / “ P. N. Armando Bermúdez / 19,025 337, -70,99825, 3087m ” / “RD-255 / 1.vii.2004, D. Pérez (3♀ USNM ENT 00038440- 42); …, La Vega, Sabana Kelly (= Quéliz) / Valle Nuevo 18,750 116, -70,616826, 2294 m, / 2.iv.2003, D. Pérez, R. Bastardo, B. Hierro (4♀, USNM ENT 00038431-34); San Juan Sierra Neiba, Trail to Sabana del Silencio, 11km SSW de El Cercado / 18,666 667, -71,500000, 1700-1900 m, 10.vii.2006, N.E. Woodley (1♁, USNM ENT 01443646; Genitalia dissected). Length (n= 5): Body, 7.4–8.1 mm; Wings, 5.8– 4.5 mm. Diagnosis. Face extensively covered by golden pollinose and pilose; female frontal triangle with a transversal golden-brownish pollinose band; metafemur yellow, only obscured on basal 2/6 and apical 1/6 or little more; surstylus in lateral view with dorsal and ventral margins approximately of the same width in the whole length. Description (Holotype). MALE. Head (Fig. 14A): Face entirely gold, white pollinose, white to golden pilose; tubercle low, bare, area above the tubercle white pollinose. Frontal triangle silver pollinose laterally, golden pilose. Frontal triangle golden-grey pollinose, golden pilose. Gena black. Occiput black pilose dorsally, yellow grey ventrally.Antenna orange, yellowish, darker toward the end and dorsal on basoflagellomere, short, ratio 1.2:1.0:1.8, basoflagellomere as long as the scape and pedicel together, apically rounded; pile black, arista brown. Thorax (Figs 14B–C): Aeneous grey, yellow pilose and brownish pollinose, with three brownish vittae, a median slender brown vitta extending along the thorax and ending at the scutellum, in addition to lateral wider brownish vittae, widening towards the pronotum and narrowed posteriorly near the pre-scutellum; pleura shining aeneous, grey pollinose, pile yellowish and sparse; scutellum shining brassy to aeneous, golden pilose. Wing (Fig. 14C): Very lightly tinged with brown, stigma brownish, membrane microtrichose, except cell c bare on basal 1/2, cells bm, r and cua bare on basal 1/5, respectively, the apical margin of the wing to just past the end of the 3rd vein brownish, giving the impression of an elongated brown macula; calypter wholly brown; plumula white; halter white, capitulum white. Legs (Fig. 14C): Yellow, metafemur slightly obscured on apical 1/5 dorsally; first pro- and mesotarsi yellow, other segments brown, metatarsus brown; coxae yellow. Abdomen (Fig. 14B): Black, parallel-sides, 1 st tergum shining metallic aeneous-black; 2 nd tergum with shallow, diffuse, bluish metallic maculae; 3 rd and 4 th terga with rectangular maculae, extending to the apical 2/3, respectively; 5 th tergum with a pair of small lateral rounded maculae, reaching the 1/2 of the segment length; pile yellow, with some black pile in the anterior edge of each segment; male genitalia: surstylus in lateral view (Fig. 15A) with dorsal and ventral margins approximately of the same width in the whole length; hypandrium in ventral view (Fig. 15C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 15C) circular, apex rounded. FEMALE (Figs 14D–F): Similar to male except: for usual sexual dimorphism, but differing in thorax comparatively more bluish, lateral vittae slender; frontal triangle with a transversal golden-brownish pollinose band; abdomen mostly bluish, with a pair of apically rounded maculae on 3 rd to 4 th terga, reaching to the apical 1/2 of each segment and there is an additional small lateral rounded macula on 5 th tergum reaching apical 1/2. Taxonomic notes. Argentinomyia aurifacies sp. nov. is similar in appearance to A. taina sp. nov. and A. praeusta but is easily distinguished from them by the face extensively covered by golden pollinose and pilose (Fig. 14 A-B, D- E); female frontal triangle with a transversal golden-brownish pollinose band (Figs 14 D-E); metafemur yellow, only obscured on basal 2/6 and apical 1/6 or little more (Figs 14A, C, D-F); 3 rd and 4 th terga with rectangular maculae, extending to the apical 2/3, respectively; 5 th tergum with a pair of small lateral rounded maculae, reaching the 1/2 of the segment length (Figs 14B, E). In A. praeusta the female frontal triangle with a medial white pollinose vitta (Figs 65 D-E); metafemur obscured on apical 1/3 or little more (Figs 65A, C, D-F); 3 rd and 4 th terga with maculae extending to the apical 1/2; 5 th tergum without a pair of small lateral rounded maculae (Figs 65B, E). In A. taina sp. nov. the frontal triangle with transversal golden-brownish pollinose band touching slightly the sides of the eyes (Figs 77 D-E); metatibia mostly brownish apically, yellow basally and slightly obscured on apical 1/5 dorsally (Figs 77C, F); 3 rd and 4 th terga with maculae extending to the apical 2/3; 5 th tergum with a pair of small lateral rounded maculae (Figs 77B, E). Based on males, Argentinomyia aurifacies sp. nov. differs from A. praeusta and A. taina sp. nov. in having the surstylus in lateral view (Fig. 15A) with dorsal and ventral margins approximately of the same width in the whole length [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. praeusta (Fig. 66A); surstylus very elongated and widened dorsally, with a small extension at the ventral margin in A. taina sp. nov. (Fig. 78A) ]; hypandrium in ventral view (Fig. 15C) narrowed laterally towards the apex [versus hypandrium expanded laterally towards the apex in A. praeusta (Fig. 66C) and A. taina sp. nov. (Fig. 78C)] (see “diagnosis” under each species or key). Etymology. The specific epithet “ aurifacies ” is an adjective derived from the Latin words “ aureus ”, “ aureo ”, denoting golden (Brown 1956: 28) and the Latin “ facies ”, countenance, external form, figure, or surface (Brown 1956: 313). This name refers to the golden face of this species. Remarks. Our study indicates that Argentinomyia aurifacies sp. nov., A. taina sp. nov. and A. praeusta occur sympatrically throughout middle and high elevations in the Dominican Republic. Geographical range. Argentinomyia aurifacies sp. nov. (n= 9) is exclusively known from the Cordillera Central in the Dominican Republic (La Compartición, La Vega, Sabana Kelly (= Quéliz) / Valle Nuevo, San Juan Sierra Neiba) (Fig. 95). The species is endemic to the West Indies domain, distributed at middle and high altitudes (1700–3087 m) in the Hispaniola province. Argentinomyia aurifacies sp. nov. and A. taina sp. nov. seems to occur in a restricted range and are rather rare in collections when compared to A. praeusta, which is widespread and abundant.

Argentinomyia taina Thompson & Montoya sp. nov. Proposed standard English name: Taina’s long-antennae flower fly. Argentinomyia DR sp. 2 Thompson, unpublished (Figs 77, 78 and 95) Type material. HOLOTYPE. Adult male, pinned, deposited at National Museum of Natural History. Original label: “ DOMINICAN REPUBLIC. Neyba Mountain Range, San Juan Serra Neiba ” / “ Trail to Sabana del Silencio, 11km SSW de El Cercado, 18,666 667, -71,500000” / “ 1700–1900 m, 10.vii.2006, N.E. Woodley ”. “ HOLOTYPE / Argentinomyia taina sp. nov. / Thompson & Montoya 2023” (USNM ENT 01443645). PARATYPES. DOMINICAN REPUBLIC. La Vega, Sabana Kelly (= Quéliz) / Valle Nuevo 18,750 116, -70,616826, 2294 m, / 2.iv.2003, D. Pérez, R. Bastardo, B. Hierro (2♀, USNM ENT 00038429, 01443644); …, La Compartición, Pico Duarte, P. N. Armando Bermúdez / 19,025 337, -70,99825, 3087 m, RD-255 / 1.vii.2004, D. Pérez (1♁, USNM ENT 00038438; Genitalia dissected). Length (n= 3): Body, 8.5–9.3 mm; Wings, 7.5–7.6 mm. Diagnosis. Frontal triangle with transversal golden-brownish pollinose band touching slightly the sides of the eyes; scutum with three brownish vittae, a median wider brown vitta extending along the thorax, in addition to lateral slender brownish vittae; metatibia mostly brownish apically, yellow basally and slightly obscured on apical 1/5 dorsally; 3 rd and 4 th terga with maculae extending to the apical 2/3; 5 th tergum with a pair of small lateral rounded maculae; surstylus in lateral view very elongated and widened dorsally, with a small extension at the ventral margin. Description. MALE. Head (Fig. 77A): Face black, white pollinose, white pilose; tubercle low, bare, area above the tubercle white pollinose; frontal triangle silver pollinose laterally, black pilose; frontal triangle brownishgrey pollinose, black pilose; gena black; occiput black pilose dorsally, yellow hoary ventrally. Antenna yellowishorange, short, ratio 1.0:1.3:1.4, basoflagellomere as long as the scape and pedicel together, darker toward the end and dorsal on basoflagellomere, pile black, apically rounded; arista brown. Thorax (Figs 77B–C): Aeneous grey, yellow-brownish pilose and pollinose, with three brownish vittae, a median wider brown vitta extending along the thorax, in addition to lateral slender brownish vittae. Pleura shining aeneous, grey pollinose, pile yellowish and sparse. Scutellum shining brassy to aeneous, golden pilose. Wing (Fig. 77C): Lightly tinged with brown, stigma brownish, membrane microtrichose, except cell c bare on basal 1/3, cells bm, r and cua bare on basal 1/3, the apical margin of the wing to just past the end of third vein brownish, giving the impression of an elongated brown macula; calypter wholly brown; plumula white; halter white, capitulum white. Legs (Fig. 77C): Yellow, pro- and mesofemora yellow, metafemur obscured on basal 2/6 and apical 1/6 or little more; pro- and mesotibiae yellow, metatibia brown, only yellow on basal 1/7; 1 st protarsus yellow on basal 1/3, other segments brown, 1 st mesotarsus extensively, other segments brown, metatarsus brown; coxae golden-brown. Abdomen (Fig. 77B): Black, parallelsides, 1 st tergum shining metallic aeneous-black; 2 nd tergum with a shallow, diffuse, bluish metallic maculae; 3 rd and 4 th terga with rectangular maculae, medio-apically rounded and extending to the apical 1/2, respectively; 5 th tergum without maculae; pile yellow; male genitalia: surstylus in lateral view (Fig. 78A) very elongated and widened dorsally, with a small extension at the ventral margin, shorter than broad; hypandrium in ventral view (Fig. 78C) expanded laterally towards the apex; aedeagal lobe in ventral view (Fig. 78C) circular, apex rounded. FEMALE (Figs 77D–F). Similar to male except for usual sexual dimorphism, but differing in the frontal triangle with transversal golden-brownish pollinose band touching slightly the sides of the eyes. Thorax comparatively more bluish, lateral vittae slender. Abdomen bluish, with a pair of apically rounded maculae on 3 rd to 4 th terga, reaching to the apical 1/2 of each segment, there is an additional small rounded macula on 5 th tergum reaching the apical 1/2. Taxonomic notes. Argentinomyia taina sp. nov. is similar to A. praeusta and A. aurifacies differing by the following combination characters: the frontal triangle with a transversal golden-brownish pollinose band, touching slightly the sides of the eyes; scutum with three brownish vittae, a median wider brown vitta extending along the thorax, in addition to lateral slender brownish vittae (Figs 77A, C, D, F); metatibia mostly brownish apically, yellow basally and slightly obscured on apical 1/5 dorsally (Figs 77A, C, D, F); 3 rd and 4 th terga with maculae extending to the apical 2/3; 5 th tergum with a pair of small lateral rounded maculae (Figs 77B, E). In A. aurifacies sp. nov. the face extensively covered by golden pollinose and pilose; female frontal triangle with a transversal golden-brownish pollinose band (Figs 14A, C, D, F); metafemur yellow, only obscured on basal 2/6 and apical 1/6 or little more (Figs 14A, C, D, F); 3 rd and 4 th terga with rectangular maculae, extending to the apical 2/3, respectively; 5 th tergum with a pair of small lateral rounded maculae, reaching the 1/2 of the segment length (Figs 14B, E). In A. praeusta the female frontal triangle with a medial white pollinose vitta (Figs 65A, C, D, F); metafemur obscured on apical 1/3 or little more (Figs 65A, C, D, F); 3 rd and 4 th terga with maculae extending to the apical 1/2; 5 th tergum without a pair of small lateral rounded maculae (Figs 65B, E). Based on males, A. taina sp. nov. differs from A. aurifacies sp. nov. and A. praeusta in having the surstylus in lateral view (Fig. 78A) very elongated and widened dorsally, with a small extension at the ventral margin [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. praeusta (Fig. 66A); surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. aurifacies sp. nov. (Fig. 15A)]; hypandrium in ventral view expanded laterally towards the apex in A. taina sp. nov. (Fig. 78C) and A. praeusta (Fig. 66C) [versus hypandrium narrowed laterally towards the apex in A. aurifacies sp. nov. (Fig. 15C)] (see “diagnosis” under each species or key). Etymology. Noun in apposition, the name “ taina ” refers to the extinct indigenous civilization who inhabited the Islands of the West Indies. Geographical range. The known range of Argentinomyia taina sp. nov. (n= 4) is restricted to the Cordillera Central in the Dominican Republic (San Juan Sierra Neiba, La Vega, Sabana Kelly (= Quéliz) / Valle Nuevo, La Compartición, Pico Duarte) (Fig. 95). The species is endemic to the West Indies domain at middle and high altitudes (1700–3087 m) in the Hispaniola province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 126-128, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Argentinomyia praeusta (Loew, 1866) (Figs 7E, 65, 66 and 95) Syrphus praeustus Loew, 1866: 155. Type locality: Cuba. HOLOTYPE Female MCZ. Rhysops praeustus. Thompson et al. 1976: 44 Rhysops praeustus. Thompson 1981: 105 (Synonyms and taxonomic notes) Rhysops quadrimaculata Hull, 1944: 28. Type locality: Cuba. HOLOTYPE Male MCZ (Examined from Photography). Refs.: Fluke, 1945: 9 (redesc.) 26 (Fig. 12, head, lateral view, male) 28 (Fig. 41, male abdomen) Rhysops quadrimaculata. Fluke, 1945: 9 Rhysops quadrimaculatus. Thompson et al. 1976: 44 Type material. HOLOTYPE. Adult Female, CUBA. Nonspecific locality provided. (MCZ Cat 23785). PARATYPE. CUBA. (1♀, MCZ Cat 4162). Wright. DOMINICAN REPUBLIC. Loma Vieja, S. Constanza, 1200 m, viii.1938 (1♁, CNC Diptera). Length (n= 7): Body, 5.4–6.7 mm; Wings, 4.2–4.6 mm. Diagnosis. Female frontal triangle with a medial white pollinose vitta; metafemur yellow with a black apical ring, metatibia brownish; 3 rd and 4 th terga with maculae extending to the apical 1/2; 5 th tergum without a pair of small lateral rounded maculae; surstylus with dorsal margin slightly concave and ventral margin slightly convex; hypandrium expanded laterally towards the apex; aedeagal lobe circular, apex rounded. Redescription. MALE. Head (Fig. 65A): Face brilliantly shining, metallic black, with perhaps a faint bluish luster, mid-vitta, violet, margins are lined by a thin vitta of whitish pubescence. Frontal triangle bluish-black, grey dusted on dorsal half and with a narrow thin line of grey-white pubescence running down the eyes margin which expands anteriorly into a small, triangular spot on dorsal part of the face and again on lower part of the face opposite the tubercle, the lower half of face sparsely white pubescent and diagonally striate opposite the tubercle; the middle of the face is cut by four shallow grooves, the outer ones less distinct; Pile of the face and frontal triangle light brownish-yellow, of vertex darker; the vertical triangle small, the occiput not protruding beyond the eyes on dorsal 1/3. Antennae yellowish, short, ratio 1.2:1.3:16, darker toward the end and dorsal on basoflagellomere; scape and pedicel nearly equal in length; light brownish-yellow; arista yellow at base and dark on apical third, with pile short. Thorax (Figs 65B–C): Very convex, shining metallic black; with three brownish pollinose vittae, the median run halfway down the posterior part of mesonotum, there are suggestions of the fourth pair of such vittae lying some distance from the others on lateral part of mesonotum. There is a low but well-developed, rounded bump in the middle of the anterior lateral corners of the thorax lying diagonally from the humeri. Pleura grey shining, whitish pollinose, yellow whitish pilose. Scutellum broadly rounded, brilliantly metallic, with two or three subterminal, transverse, faint, ripple-like depressions; fringe long. Wing (Fig. 65C): Very lightly tinged with brown, stigma brownish, microtrichose, except cell c bare on basal 2/3, cells bm and r bare on basal 2/3, cell cua bare on anterior 3/5, the apical margin of the wing to just past the end of third vein marginate with brown, giving the impression of an elongated brown macula; calypter yellow, border and fringe dark brown; plumula white; halter yellow, capitulum yellowish. Legs (Figs 7E, 65C): Almost wholly pale brownish-yellow; metafemur obscured on apical 1/3 or little more; metatibia and metatarsus obscurely dark except the indefinite base, metatarsus and three basal tarsomeres yellow, another tarsus dark brown. Abdomen (Fig. 65B): Long and slender, rather flattened, sides almost parallel, end of 3 rd tergum barely wider than the base of abdomen; 1 st tergum shining metallic black, 2 nd shining on basal 2/5 with a brassy or golden, brown appearance; opaque, dark sepia-brown on almost all of the remaining segments and quite to their posterior margin. Located near the base upon sides of 2 nd tergum, not reaching the anterior corners and covering about 1–1/2 the segment length there are shallow, diffuse, brownish-yellow maculae; in some specimens, it extends inward for a greater distance. 3 rd tergum with a pair of large, square, light brownish-yellow maculae occupying the base, the entire anterior corners and the sides for 1/2 the segment length; their medial surfaces are just a little cut-away and diminished on posterior surfaces; the remainder segments opaque, dark sepia. 4 th tergum with a pair of similar maculae, slightly smaller, barely darker and a very little closer together; the remainder of that segment shining black. The pile upon abdomen seems to be practically absent on 3 rd and 4 th terga. The sparse lateral pile of 2 nd is pale; male genitalia: surstylus in lateral view (Fig. 66A) with dorsal margin slightly concave and ventral margin slightly convex; hypandrium in ventral view (Fig. 66C) expanded laterally towards the apex; aedeagal lobe in ventral view (Fig. 66C) circular, apex rounded. FEMALE (Figs 65D–F). Similar to male except for usual sexual dimorphism and differing in Abdomen opaque black, more shining on sides with a pair of quadrate yellow maculae in the basal marginal corners of 3 rd and 4 th terga, slightly wider on margin and reaching nearly to the apical 1/2. The female of A. praeusta is similar in appearance to A. jamaicensis sp. nov., differing by the following combination characters: Female frontal triangle with a medial white pollinose vitta; metafemur obscured on apical 1/3 or little more; 3 rd and 4 th terga with maculae extending to the apical 1/2; 5 th tergum without a pair of small lateral rounded maculae; metafemur yellow with a black apical ring, metatibia brownish. Taxonomic notes. Argentinomyia praeusta differs from A. aurifacies sp. nov. and A. taina sp. nov. by having the female frontal triangle with a medial white pollinose vitta (Figs 65D, F); metafemur yellow with a black apical ring, metatibia brownish (Figs 65A, C, D, F); 3 rd and 4 th terga with maculae extending to the apical 1/2; 5 th tergum without a pair of small lateral rounded maculae (Figs 65B, E). In Argentinomyia aurifacies sp. nov. the face extensively covered by golden pollinose and pilose (Figs 14A, C, D, F); female frontal triangle with a transversal goldenbrownish pollinose band (Figs 14D, F); metafemur yellow, only obscured on basal 2/6 and apical 1/6 or little more (Figs 14A, C, D, F). In Argentinomyia taina sp. nov. the frontal triangle with transversal golden-brownish pollinose band touching slightly the sides of the eyes (Figs 77A, C, D, F); metatibia mostly brownish apically, yellow basally and slightly obscured on apical 1/5 dorsally (Figs 77A, C, D, F); 3 rd and 4 th terga with maculae extending to the apical 2/3; 5 th tergum with a pair of small lateral rounded maculae (Figs 77B, E). Based on males, A. praeusta differs from A. aurifacies sp. nov. and A. taina sp. nov. in having the surstylus in lateral view (Fig. 66A) with dorsal margin slightly concave and ventral margin slightly convex [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. aurifacies sp. nov. (Fig. 15A); surstylus very elongated and widened dorsally, with a small extension at the ventral margin in A. taina sp. nov. (Fig. 78A)]; hypandrium in ventral view expanded laterally towards the apex in A. praeusta (Fig. 66C) and A. taina sp. nov. (Fig. 78C) [versus hypandrium narrowed laterally towards the apex in A. aurifacies sp. nov. (Fig. 15C)] (see “diagnosis” under each species or key). Remarks. Thompson (1981) compared the type of Rhysops praeusta (MCZ: https://mczbase.mcz.harvard.edu/ name/ Syrphus %20praeustus) and R. quadrimaculata (MCZ: https://mczbase.mcz.harvard.edu/name/ Rhysops %20q uadrimaculata) and found that they corresponded to opposite sexes of the same species (conspecific) with the pile of male brownish, abdomen narrower, with large, quadrate maculae, sides of 2 nd tergum shiny. He concluded that these are typical of sexual dimorphism found in other species of the genus, therefore, formally synonymized Rhysops quadrimaculatus under Rhysops praeustus. Geographical range. Argentinomyia praeusta (n= 24) is endemic to the West Indies and widely distributed in Cuba (Camagüey), Serra de Bahruco / Massif de la Selle and the Cordillera Central, Cordillera Septentrional in the Dominican Republic* (Independencia, La Vega, La Vega, Pedernales) and the Blue Mountains in Jamaica * (Portland Hardward) (Fig. 95). The species is endemic to the West Indies domain at middle altitudes (1100–2400 m), widely distributed in the provinces: Cuban (1100 m), Jamaica (1120 m) and Hispaniola (1200–2400 m). Non-type material examined. CUBA. Camagüey, Nonspecific locality provided. 21,521 757, -77,781167, 1100 m, Wright (1♁, MCZ). DOMINICAN REPUBLIC. Independencia, Road 47, between Los Pinos and Ángel Félix, 18,743 411, -71,675054, 700 m, 20.vi.2005, N.E. Woodley (1♀, USNM ENT 01443771); near Serra de Baoruco National Park, 3.5km El Aguacate, Montane meadow, 18,166 667, -71,416667, 2210 m, 30.ix.1991, C. Young; S. Thompson; R. Davidson; J. Rawlins (1♁, USNM ENT 01443776); La Vega, Sabana Kelly (= Quéliz) Valle Nejeno, 18,750 116, -70,616826, 2294 m, 2.iv.2003, D. Pérez, R. Bastardo; B. Hierro (2♁, 6♀, USNM ENT 00038429–36); Trail to Peak and Centro SOECI, Pico Diego de Campo, 19,580 523, -70,740457, 918 m, RD-184, 4.ii.2003, D. Pérez, R. Bastardo; A. Marmolejos (3♁, 3♀, USNM ENT 00038437–42); La Compartición, Pico Duarte, P. N. Armando Bermúdez, 19,025 337, -70,99825, 3087 m, RD-255, 7.i.2004, D. Pérez (2♁, USNM ENT 00038439; 0003843); Pedernales, 9.7km NE, los Arroyos, 18,823 174, -70,832299, 2070 m, 15.vii.1990, J. Rawlins, C.W. Young; S.A. Thompson (1♀, USNM ENT 01443777). JAMAICA. Portland Hardward Gap, 18,068 139, - 76,636 631, 1120 m, 5.vi.1965, T.H: Farr (1♀); 19.vii.1966, Howden; Becker (1♀, CNC DIPTERA 112214)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 109-112, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Loew, H. (1866) Diptera Americae septentrionalis indigena. Centuria sexta. Berliner Entomologische Zeitschrift, 9 (1865), 127 - 186.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Thompson, F. C. (1981) The Flower Flies of the West Indies (Diptera, Syrphidae). Memoirs of the Entomological Society of Washington, 9, 1 - 200.","Hull, F. M. (1944) Studies on syrphid flies in the Museum of Comparative Zoology. Psyche, 51, 22 - 45.","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29."]}

Argentinomyia jalcaensis Thompson & Montoya sp. nov. Proposed standard English name: Jalca’s long-antennae flower fly. Tuberculanostoma 08-08 Thompson, unpublished (Figs 37, 38 and 91C) Type material. HOLOTYPE. Adult male, pinned, deposited at Natural History Museum of Denmark, Copenhagen. Original label: “ PERÚ, Santana, Áncash, 22: 35 km southeast of Huaraz, Cerro Cahuish” / “ -9.383915, -77.651629, 4100 m, Quabrado Pucaudao” / “ 15-18.ii.1987, O. Karsholt ”. “ HOLOTYPE / Argentinomyia jalcaensis / Thompson & Montoya 2023”. Length (n= 1): Body, 6.5 mm; Wings, 5.4 mm. Diagnosis. Generally metallic bluish-black fly. Face slightly produced; oral opening about 1.5 times as long as wide; antenna black; scutum yellow pilose; halter black; legs except pro- and mesotibiae yellow on apical 1/5 and basal 1/3, yellow pilose. Hypandrium acuted laterally towards the apex; apex of hypandrium (superior lobes) no feline claw shape. Description. MALE. Head (Fig. 37A): Face only slightly produced anteriorly, about 1/2 as long anteriorly as high dorsally, sparsely white pollinose laterally, shiny on medial 2/3, white pilose laterally; oral opening about 1.5 times as long as wide; gena black, shiny, silvery-white pollinose, yellow pilose; tubercle distinct, only slightly dorsal to oral margin; lunule black; frontal triangle broad, sparsely white pollinose laterally, shiny anteromedially, black pilose; occiput silvery-white pollinose and yellow pilose and pollinosity, sparser on dorsal 1/4, with some longer black cilia on dorsal 1/4. Antenna black, short, about half as long as face, ratio 1.0:1.0:1.6; basoflagellomere oval, slightly longer than broad; arista about 1.2 times as long as basoflagellomere. Thorax (Figs 37B–C): Postpronotum sparsely white pollinose; scutum shiny, long yellow pilose; scutellum shiny, yellow pilose; pleuron sparsely white pollinose, yellow pilose. Wing (Fig. 37C): Hyaline, stigma brownish yellow, microtrichose; calypter white; plumule white; halter black, capitulum white. Legs (Fig. 37C): Black, except pro- and mesotibiae yellow on apical 1/5 and basal 1/3, yellow pilose; coxae and trochanters silvery-white pollinose, white pilose; profemur without bristles; metatarsus with apical tarsomeres black pilose. Abdomen (Fig. 37B): Subshiny, dull black laterally in form of large elongate areas on lateral 1/4 reaching to apical 1/4, dull black pollinose medially and apically, long yellow pilose except shorter medially; sterna shiny, yellow; male genitalia: Black, small, yellow pilose, surstylus in lateral view (Fig. 38A) shorter; hypandrium in ventral view (Fig. 38C) acuted laterally towards the apex; apex of hypandrium (superior lobes) no feline claw shape, with a lateral rounded extension (Fig. 38C); aedeagal lobe in ventral view (Fig. 38C) circular, apex rounded. FEMALE. Unknown. Etymology. The specific epithet “ jalcaensis ” is derived from the ecoregion name where the species was found in the Peruvian Andes and the Latin suffix - ensis denoting place, locality, country, or belonging to, pertaining to (Brown 1956: 45, 303). The “ Jalca ” is defined in its broadest sense as the shrubby or prairie, humid or sub-humid mountain range, located between 4,800 to 6,768 meters above sea level (Height of Huascarán) (Sonter et al. 2018). Taxonomic notes. Argentinomyia jalcaensis sp. nov. is similar in appearance to A. opaca, from which it differs in having the following combination of characters: Generally metallic bluish-black. Face slightly produced; oral opening about 1.5 times as long as wide; antenna black (Figs 37A, C, D-F); scutum yellow pilose; halter black (Figs 37 B-C, E-F); legs except pro- and mesotibiae yellow on apical 1/5 and basal 1/3, yellow pilose (Figs 37A, C, DF). Argentinomyia opaca is characterized by the face perpendicular with a well-rounded tubercle, deeply concave dorsally (Figs 59A, C, D-F); scutum black pilose; halter white (Figs 59 B-C, E-F); legs black, black pilose (Figs 59A, C, D-F). Based on males, A. jalcaensis sp. nov. differs from A. opaca in having the hypandrium in ventral view (Fig. 38C) acuted laterally towards the apex [versus hypandrium narrowed laterally towards the apex in A. opaca (Fig. 60C)]; apex of hypandrium (superior lobes) no feline claw shape, with a lateral rounded extension (Fig. 38C) [versus apex of hypandrium feline claw shape in A. opaca (Fig. 60C)] (see “differential diagnosis” under each species or key). Distribution. Argentinomyia jalcaensis sp. nov. (n= 1) is exclusively distributed in the highlands (4100 m) of the Cordillera Occidental and Central Sur in Perú, which corresponds to the Central Andes domain, Puna province (Fig. 91C). The species is restricted to high-elevation Jalca grassland of Cerro Cahuish in Perú, which raises the need for further expeditions to neighboring areas of the type locality, in order to discover and describe the female specimens as well as to know its population dynamic and conservation status. Conservation status. Unfortunately, the region where A. jalcaensis sp. nov. inhabits is near the Pierina gold and silver open-pit mine, located in the Callejón de Huaylas in the Peruvian Andes (Cordillera Negra of northcentral Perú), approximately 10 kilometers directly north of Huaraz at a height of 4100 meters. Since mining has the potential to produce serious and specific threats to biodiversity, with intensive land use that severely fragments and declines the extension and quality of the habitat due to deforestation (see Sonter et al. 2018; Montoya et al. 2021), with the loss of connectivity by the reduction of functional conservation corridors, we consider that populations of A. jalcaensis sp. nov. is probably “Near Threatened”, “Endangered” or “Vulnerable”, since it occupies small range, threatened by several human threats. In this sense, we suggest that this species should be studied and its level of threat evaluated, taking into account the conservation criteria of the International Union for Conservation for Nature—IUCN, considering the restricted distribution to a unique habitat such as the Jalca, where the impact of gold and silver mining could cause its disappearance in the coming years, making the need to encourage conservation efforts as has suggested for species Copestylum enriquei Montoya, Parada-Marín & Ramos-Pastrana, 2022 in the Amazonian region (Montoya et al 2022).

Argentinomyia catabomba (Williston, 1891) (Figs 24, 25 and 96B) Melanostoma catabombum Williston, 1891: 12. Type locality: Mexico. Guerrero, Omiltemi, 8000ft and Amula, 6000ft. SYNTYPE 6 Males and 3 Females BMNH (Examined). LECTOTYPE Male BMNH here designated. Refs.: Fluke, 1945: 5 (Redescription), Fig. 2 (head) Rhysops catabomba. Curran, 1937: 2 (key) Rhysops catabomba. Fluke, 1945: 5 Rhysops catabomba. Thompson et al. 1976: 42 Melanostoma catabombum. Papavero & Ibañez-Bernal, 2003: 166 Argentinomyia catabomba. Mengual, 2020: 7 Melanostoma melanocerum Williston, 1891: 13. Type locality: Costa Rica. Rio Sucio SYNTYPE 2 Females BMNH. LECTOTYPE Female BMNH here designated. (Examined). n. syn. Rhysops melanocerum. Williston, 1907: 2 Rhysops melanocera. Curran, 1937: 2 (key) Type material. LECTOTYPE. Adult Male, MEXICO. Guerrero, Omiltemi, 2438 m, 1.vii.1903, H. H. Smith (BMNH, Previously SYNTYPE). “ LECTOTYPE / Melanostoma catabombum / det. A.L. Montoya 2023”. PARALECTOTYPES. Label with the same data as LECTOTYPE except (5♁, 3♀, BMNH Previously SYNTYPE). “ PARALECTOTYPE / Melanostoma catabombum / det. A.L. Montoya 2023”. MEXICO. Guerrero, Xucumanatlan, 1900 m, 1.vii.1903, H. H. Smith (8♁, BMNH, Previously SYNTYPE). “ PARALECTOTYPE / Melanostoma catabombum / det. A.L. Montoya 2023”. MEXICO. Guerrero, Amula (Note: Modern name of Amula is Almolonga), 1829 m, 1.vii.1903, H. H. Smith (3♁, BMNH, Previously ALLOTYPE). “ PARALECTOTYPE / Melanostoma catabombum / det. A.L. Montoya 2023”. Label with the same data as LECTOTYPE except (2♁, 1♀, AMNH 19161, 19255, in the USNM ENT, Previously COTYPE). “ PARALECTOTYPE / Melanostoma catabombum / det. A.L. Montoya 2023”. LECTOTYPE. Adult Female, COSTA RICA. Rio Sucio (Previously SYNTYPE). “ LECTOTYPE / Melanostoma melanocerum / det. A.L. Montoya 2023”. PARALECTOTYPE. COSTA RICA. Rio Sucio (1♀, BMNH, Previously SYNTYPE). “ PARALECTOTYPE / Melanostoma melanocerum / det. A.L. Montoya 2023”. Length (n= 8): Body, 8–10 mm; Wings, 7.5–8.1 mm. Diagnosis. Scape and basoflagellomere of equal length; wing hyaline, almost bare on basal 2/3 or more; protarsus dark; abdomen with rounded oblique yellow maculae on 3 rd to 4 th terga; surstylus brown with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad; hypandrium narrowed laterally towards the apex; aedeagal lobe with apex acute. Redescription. MALE. Head (Fig. 24A): Face black, sides of the frontal triangle and face thickly covered with uniform yellow golden pollen, leaving a linear spot in the median dorsal area of the antennae and a broad ventromedial vitta shining, somewhat metallic black. Gena semi-shining. Frontal triangle black pilose Ocellar triangle blue metallic with thin brownish pollen, pile black. Occiput with yellow hairs dorsally, long pile black, pollen and pile paler ventral. Antennae brownish orange, short, ratio 1.4:1.0:1.6, basoflagellomere largely blackish dorsal; scape about as long as the oval basoflagellomere, distinctly longer than pedicel; arista yellow-orange at base and dark on apical third. Thorax (Fig. 24B–C): Bronze, mesonotum shining, yellow pilose; with three brownish vittae on anterior half, the median one slender, side broadly brownish pollinose, pile yellow-brownish. Pleura of a similar colour in middle, more steel-blue ventral, posterior margin of anepisternum slightly reddish, grey pollinose. Scutellum shining steel-blue, yellowish pilose; fringe long and yellow. Wing (Fig. 24C): Diluted reddish, stigma brown, with a narrow cloudy-blackish macula at the tip, cells dm, cua 1 bare on basal 2/3 or more, cell cua bare on anterior 1/3; calypter white, border and fringe brownish; plumula white, halter orange yellowish, capitulum slightly yellowish. Legs (Fig. 24C): Yellow to reddish; pro- and mesofemora brownish basally; pro- and mesotarsi black, except basal segments reddish yellow; metafemur brown except yellow apically; metatibia brown except yellow on basal 1/4; basal two segments of tarsi yellow. Abdomen (Fig. 24B): Narrow, its greatest width at the distal end of the 3 rd tergum; opaque black; 1 st tergum and hypopygium shining metallic; 2 nd tergum metallic on antero-lateral margin, with lateral yellow maculae that run alongside and is narrow at apex; 3 rd and 4 th terga each with a pair of moderately large, rounded yellow maculae anteriorly, reaching the lateral margin; pile long and light yellow laterally; sterna blackish, yellow pilose; male genitalia: cercus yellow, surstylus brown, in lateral view (Fig. 25A) with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad; hypandrium in ventral view (Fig. 25C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 25C) with apex acute. FEMALE (Figs 24D–F). Similar to male except for usual sexual dimorphism and differing in the frontal triangle and sides of face whitish-pollinose, with an opaque black cross-band, in addition to a pointed band of pollen extending medially. Thorax and scutellum moderately shining, greenish-blue; mesonotum with three darker vittae, the median one linear. Abdomen subopaque, black, 1 st to 5 th terga more shining; 2 nd tergum with a yellow macula on each side reaching anterior margin; the oval maculae of 3 rd and 4 th terga not touching lateral margin. Female of A. catabomba can be confused with A. CR-14, but differs from it by the pro- and mesofemora brownish basally; metafemur brown except yellow apically; pro- and mesotarsi with the three apical tarsomeres yellow. 2 nd tergum metallic on anterolateral margin, with lateral yellow maculae that run alongside and is narrow at the apex. 3 rd and 4 th terga each with a pair of moderately large, rounded yellow maculae anteriorly. Taxonomic notes. Argentinomyia catabomba is characterized by the scape and basoflagellomere of equal length (Figs 24A, C, D-F); protarsus dark (Figs 24A, C, D-F); abdomen with rounded oblique yellow maculae on 3 rd to 4 th terga (Figs 24B, E). Argentinomyia catabomba is similar to the South American species A. lanei and A. berthae differing from them by the wing hyaline with costal cell bare on basal 3/4 or more and protarsus dark (Figs 24 BC, E-F). A. catabomba can be confused with Argentinomyia CR-14, a species with pro- and mesofemora orange; metafemur is brown, only yellow on basal 1/3; pro- and mesotarsi with the three apical tarsomeres black (Figs 24A, C, D-F); 2 nd tergum orange on basolateral 2/3, 3 rd and 4 th terga with oval macula extending on lateral margin (Figs 24B, E). Based on males, A. catabomba differs from A. berthae and A. lanei in having the surstylus in lateral view (Fig. 25A) with dorsal and ventral margins approximately of the same width in the whole length [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. berthae (Fig. 19A); surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. lanei (Fig. 41A)]; aedeagal lobe in ventral view, with acute apex in A. catabomba (Fig. 25C) and A. lanei (Fig. 41C) [versus aedeagal lobe circular, apex rounded in A. berthae (Fig. 19C)] (see “differential diagnosis” under each species or key). Remarks. According to FC Thompson (personal unpublished notes), Fluke (1945) keys out A. catabomba together with A. scitula (Williston, 1888) and separated it by the “front tarsi wholly yellow”. However, FC Thompson, pointed out that this character is in conflict with Williston’s original description, who clearly indicated the “basal portion of protarsus is yellow” in A. scitula, then it is probable that Fluke meant to say protarsus is dark in A. catabomba, with only the basal segments reddish yellow (Curran 1937). A female specimen collected in Guatemala was misidentified as Rhysops vittithorax by Hull (CNC DIPTERA 112215) which is a synonym of A. longicornis (Walker, 1836) therefore we corrected the identification as A. catabomba. When examining and comparing the syntype of Melanostoma melanocerum (Williston) (2 females) with the syntype of Melanostoma catabombum Williston (1 male) deposited in the BMNH, we found that they correspond to opposite sexes of the same species since both female and male have the same patterns of pollinosity on thorax and abdominal maculae are similar, with only the sexual dimorphism found in most species of the genus. Thus, based on examination of the syntype and due to the morphological evidence presented above, Melanostoma melanocerum is synonymized with A. catabomba. Finally, Williston, 1891: 12 described Melanostoma catabombum based on six males and three females collected in Mexico, Guerrero, Omiltemi, 8000ft and Amula, 6000ft. The nine specimens comprise the type series deposited in the collection of the AMNH and BMNH labelled as “ Syntype ”, “ Allotype ” and “Cotype”. Curran (1937) and Fluke (1945) keyed out and provided a redescription to the species A. catabomba but did not select or designate a Lectotype. Therefore, to avoid assuming which specimen is the Holotype and objectively establish the identity of this taxon and consequently stabilize the nomenclature and universal interpretation of the name, one male specimen from the BMNH labelled “ Syntype ” Melanostoma catabombum / Williston, 1891 / “ MEXICO. Guerrero, Omiltemi / 2438 m, 1.vii.1903, H. H. Smith” is hereby selected and designated as the Lectotype. The remaining syntypes in the BMNH labelled “ Syntype ” Melanostoma catabombum / Williston, 1891 / “ MEXICO. Guerrero, Xucumanatlan, 1900 m, 1.vii.1903, H. H. Smith and MEXICO. Guerrero, Amula 1829 m, 1.vii.1903, H. H. Smith” as well as the ones in the AMNH labelled “ Syntype ” Melanostoma catabombum / Williston, 1891 / “ MEXICO. Guerrero, Amula 1829 m, 1.vii.1903, H. H. Smith” are consequently designated as Paralectotypes to ensure consistent interpretation of this name and have been so labelled. The arguments for selecting this specimen were: (1) it is in almost perfect condition and (2) corresponds exactly to the same type locality. Below the above criteria, Lectotype and Paratype are designated for Melanostoma melanocerum from two female Syntype specimens housed in the BMNH and labelled “ COSTA RICA. Rio Sucio”. Geographical range. Argentinomyia catabomba (n= 66) is distributed in Costa Rica * (Cartago, Limón, Puntarenas, San José), Guatemala * (Antigua, Baja Verapaz, Panajachel), El Salvador * (San Salvador), Mexico (Chiapas, Guerrero, Mexico D.F., Oaxaca, Xalapa) and Panamá * (Chiriqui) (Fig. 96B). The species is endemic to the Mesoamerican domain, distributed at low and middle altitudes (94–2510 m) in the provinces: Chiapas Highlands (1550–2510 m), Pacific Lowlands (1900–2438 m), Puntarenas-Chiriquí (1200 m), Serra Madre del Sur (1829 m), Trans-Mexican Volcanic Belt (2463 m) and Veracruzan (94 m). Non-type material examined. COSTA RICA. Cartago, 9.852518, -83.923886, 1390 m, xi.1965, N.L.H. Krauss (1♁, USNM ENT 00036957); Rio Grande de Orosi, desde Administración hasta Sendero La Pava, 9,797 784, -83,925756, 1150–1600 m, 2.i.1997, R. Guzmán (2♁, INBioCRI 002537010–11); Limón, P.I.I A. Valle del Silencio, alrededor Refugio Sendero Circular, 9,297 345, -83,043913, 2450 m, 5.x.2003, R. Gonzales (1♁, INBioCRI 0003754850); …, 2420 m, 6.vi.1996, R. Villalobos (1♁, INBioCRI 000399294); Puntarenas, Quarry (Tajo) near Las Alturas Station, 8,856 123, -82,850287, 1064 m, 16.vii.1995, N. Norrbom (1♀, USNM ENT 00036947); Estación Altmira, 1 km S del Cerro Biolley, 1450 m, 28.vii.1995, R. Villalobos (1♀, INBioCRI 002322716); San José, F. Cementerio de la Maquina P.N. Chirripó, 2100–2500 m, 3.ii.1993, F. Quesada and M. Segura (2♁, INBioCRI 001208818; 001305772). GUATEMALA. Antigua, 14.558571, -90.729523, 1550 m, 23.vi.1923, E. G. Smith (1♀, CNC DIPTERA 112215) (Identified as Rhysops vittithorax by Hull and Rhysops sp. 19 by Vockeroth); Baja Verapaz, Cerca Purulhá, camino Pantín, 15,207 742, -90,176891, 1500 m, 22.ix.2005, José Monzón (2♁,Colección J. Monzón); Salamá, Camino Paten a Santa Rosa, 15,062 189, -90,353998, 1700 m, 15.vi.2007, José Monzón (2♁, Colección J. Monzon); Sololá, Panajachel, 14,740 962, -91,151845, 1750 m, 19.viii.1963, D.Q Cavagnaro, M.E. Irwen (1♀, CAS). EL SALVADOR. San Salvador, 13,696 903, -89,254923, 893 m, 8.vii.1963, D.Q Cavagnaro, M.E. Irwen (1♁, CAS). MEXICO. Chiapas, San Cristóbal de las Casas, L. C. Huitepec, 16.736812, -92.637332, 2510 m, 26.ii.2011, P. Sagot (12♁, 11♀, ECO-TAP-E); Zinacantan, Muktavits (Cerro Huitepec) 16,744 909, -92,682661, 2804 m, 15.vii.1976, D.E and J.A. Breedloe (1♁, 1♀, CAS); Mexico, Parque, along Lakes of Zempoala, 19.053873, -99.311982, 2463 m, 10-11.viii.1989, A.L. Norrbom, Sweeping on Dahlia imperiales (89 M12) (1♁, USNM ENT 00036950); D.F, L. Conradt (1♀, USNM ENT 00036958); Oaxaca, Km 188 in Valle Nacional, Road 29, 17.767761, -96.323867, 94 m, 29.vii.1992, M. Wood (1♀, CNC DIPTERA 112220); Xalapa, 19,543 487, -96,909959, 1450 m, 22.ii.1998, A. Freidberg (1♀, USNM ENT 01443748). PANAMÁ. Chiriqui, 2Km Santa Clara, Finca Hartmann S. 8.844832, -82.760761, 1200 m, 27.vi.1981, B. Gill (1♁, CNC DIPTERA 112216) (Identified as Rhysops sp. 19 by Vockeroth)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 46-49, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Williston, S. W. (1891 - 1892) Fam. Syrphidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Zoologia- Insecta-Diptera, 3, 57 - 72 (December 1891), 73 - 78 (May 1892).","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Curran, C. H. (1937) The Neotropical species of Melanostoma and allies (Syrphidae: Diptera). American Museum Novitates, 926, 1 - 4.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Papavero, N. & Ibanez-Bernal, S. (2003) Contribution to a history of Mexican Dipterology. Part II. The Biologia CentraliAmericana. Acta Zoologica Mexicana, 88, 143 - 232. https: // doi. org / 10.21829 / azm. 2003.88881793","Mengual, X. (2020) Phylogenetic relationships of the bacchine flower flies (Diptera: Syrphidae) based on molecular characters, with a description of a new species of Melanostoma (Schiner, 1860). Contributions to Zoology, 82 (2), 210 - 244. https: // doi. org / 10.1163 / 18759866 - 20191410","Williston, S. W. (1907) Class I, Hexapoda. Order IV, Diptera. Dipterological notes Journal of the New York Entomological Society, 15, 1 - 2.","Williston, S. W. (1888) Diptera Brasiliana, ab H. H. Smith Collecta. Transactions of the American Entomological Society, 15, 243 - 292."]}

Identification key to the Neotropical genera of Melanostomini and Bacchini A key to the genera of Neotropical Melanostomini and Bacchini was prepared based on characters proposed by Fluke (1945, 1957), Thompson (1999a), Thompson (2006), Thompson et al. (2010), Huo (2014), Thompson & Skevington (2014), Young et al. (2016), Miranda & Moran (2017), Ramage et al. (2018) and Mengual (2020). 1. Postpronotum pilose (MCAD Fig. 30); male abdomen with four unmodified pregenital segments; 5 th tergum usually not visible in dorsal view (MCAD Figs 1, 4); female abdomen tergum 6 th hardened or membranous, visible, if tergum 6 th and onward visible, then these are membranous; epiproct as two plates that may be connected to the base, or as a single plate, or as a band with two projections (subfamilies Eristalinae, Microdontinae and Pipizinae).... other flower flies (“MCAD key couplet 2”) - Postpronotum bare; male abdomen with five unmodified pregenital segments; 5 th tergum visible in dorsal view (MCAD Figs 53–61); female abdomen tergum 6 th hardened, visible; epiproct divided into two plates that may be connected to the base (subfamily Syrphinae)................................................................................. 2 2. Face and/or scutellum partially yellow or yellowish-brown in background color; phallus (aedeagus) two-segmented........................................................................................ other Syrphinae genera - Face and scutellum entirely black in background color (some species with partly pale face or scutellum); phallus (aedeagus) simple, unsegmented (Bacchini and Melanostomini)......................................................... 3 3. Face usually straight, oral margin and antennal bases in line, without tubercle, flat, straight or convex; abdomen petiolate, distinctly narrower than thorax (Fig. 4A, MCAD Figs 59, 60); male: 2 nd tergum narrow, narrower than 3 rd and 4th terga (Mengual et al. 2018, Fig. 18); female abdominal tergum 6 th, not noticeable nor visible without dissection, tergum 6 th internal to the 5 th, tergum 5 th 1/10 of the length of tergum 4 th, almost inconspicuous (Miranda & Moran 2017, Figs 59, 80 and 90)................................................................................................ Leucopodella Hull - Face with tubercle; abdomen parallel-sided or narrowly oval (MCAD Figs 55, 58); other characters variable; female tergum seventh visible without dissection........................................................................ 4 4. Antennal cavities broadly confluent (MCAD Fig. 24); face narrow, with distinctive, rounded facial tubercle; metathoracic pleuron with several fine subappressed pile ventral to spiracle; katepisternum with pile patches broadly separated posteriorly, but joined anteriorly; abdomen oval, wide and flat, clearly broader than scutum, usually with yellow-orange maculae on 2 nd to 4 th terga; surstyli elongated, one-lobed, digitiform or pointed, generally widened apically, postgonites small, squarish or triangular in shape (Fluke 1957: 278, Figs 119–122; Borges & Pamplona 2003); female tergum 5 th 1/3 of the length of tergum 4 th, tergum 6 th 1/3 of the length of 5 th (Miranda & Moran 2017, Figs 66, 84 and 97)................. Xanthandrus Verrall - Antennal cavities broadly separated (MCAD Fig. 23); metathoracic pleuron bare; katepisternum with pile patches usually broadly separated throughout; abdomen variable, slender and parallel-sided, as broad as scutum; other characters variable...................................................................................................... 5 5. Metasternum greatly reduced to a small diamond-shaped area (Figs 3A, 5J; Andersson 1970: 273, Fig. 1C, 3C; Shatalkin 1975: 118, Fig. 11; Thompson et al 2010: 771, Fig. 34; Haarto and Ståhls 2014: 95, Figs 1A; Thompson & Skevington 2014); antenna short, with scape and basoflagellomere as long as wide; face not produced below, with small tubercle, facial pollinosity neither punctate nor rippled (MCAD Fig. 28); male abdomen nearly parallel-sided, from two to five times as long as greatest width, female with abdomen varying from nearly parallel-sided to oval, from 1.7 to 2.5 times as long as greatest width; usually with triangular or subquadrate shaped yellow maculae; male genitalia with surstyli simple, one-lobed, elongate, straight and thin, with a small basomedial tubercle; postgonites (apex of hypandrium) massive, irregular or squarish in shape; female tergum 5 th 1/4 of the length of tergum 4 th, tergum 6 th 1/5 of the length of tergum 5 th......................... Melanostoma Schiner [Note: A new species described from Cameroon by Mengual (2020) does not have the reduced metasternum] - Metasternum entire, not reduced (Fig. 3B; MCAD Fig. 35; Haarto and Ståhls 2014: 95, Fig. 1B; SNLSA Fig. 12C); face variable, almost straight in profile with weak tubercle or moderately to strongly produced forward ventrally, sometimes with pollinosity forming punctuate or rippled pattern, other characters variable........................................ 6 6. Face produced forward into a snout, with a very prominent and abrupt tubercle closer to the antennal base than to the oral opening apex (Fig. 4N, Fluke 1943: 427, Figs 1–7); face sparsely pollinose and without punctate shiny (bare) maculae (Fig. 4N); head as long as or longer than high, wider than thorax; male eyes dichoptic or holoptic; antenna elongate, as long as or longer than face; basoflagellomere elongate, four or more times as long as broad, near equal to scape and pedicel together (Fig. 4N); notopleura without a distinct tubercle; subscutellar fringe short, thin and white; prolegs simple in male, sometimes with a curled tibial seta (Fig. 4N, Fluke 1943: 427, Figs 8–9); male genitalia with surstyli curved; superior lobes transverse and elongated; ejaculatory apodeme elongate and knobbed toward the apex (Fluke, 1957: 278, Figs 124–125; Shatalkin 1975: 118, Fig. 12); small species, shining black or blue metallic, 7mm or less.............. Platycheirus (Tuberculanostoma) Fluke - Face straight or slightly produced anteriorly, with a low and indistinct tubercle, frequently densely pollinose and with small shiny punctate-like bare maculae; head higher than long; male eyes face not greatly produced anteriorly; basoflagellomere oval or slightly elongate, never more than twice as long as broad; notopleura with a distinct tubercle; subscutellar fringe complete; male sometimes with legs modified, either broadened or with special bristles, pile tufts, modified hairs or none; frequently medium to large flies, sometimes with pale to metallic colored markings......................................... 7 7. Face slightly receding to rather strongly produced below, with obscure to prominent tubercle, densely pollinose and sometimes with pollinosity forming a distinctive bare punctum or rippled pattern (Fig. 4M); male protibia usually with long black bristles posteriorly, sometimes protibia and/or tarsus expanded, usually with specialized setae (Fig. 4M); abdomen nearly parallel-sided in males, narrowly oval in the female; with yellow to orange or silver-gray pollinose on bluish submetallic macula, generally with quadrate markings; surstyli bifid or bilobed with a long, slightly curved lateral lobe and a short, stout basomedial lobe, superior lobes usually slender, recurved and crescent-shaped (Vockeroth 1990: Fig. 83; Thompson & Skevington 2014; Young et al. 2016); female tergum 5 th 1/3 of the length of tergum 4 th, tergum 6 th 1/5 of the length of tergum 5 th (Miranda & Moran 2017, Figs 70, 71, 86 and 100).......................................................... Platycheirus (Carposcalis) - Face straight in profile, not produced anteriorly, usually uniformly pollinose, sometimes with shiny (bare) punctate maculae; male legs slender, without bristles, pile tufts, or modified hairs; abdomen frequently with pale-colored maculae; surstyli elongate, one-lobed, postgonites massive, irregular, elongate, triangular, or squarish in shape......................... 8 8. Antennae short, scape broader than long, scape nearly equal to pedicel, basoflagellomere large, slightly oval and apically rounded; face slightly receding to perpendicular with a well-rounded tubercle, never with transverse grooves dorsally along tubercle neither broadly (bare) punctuate (Fig. 6D); mesocoxa pilose posteriorly (Fig. 6E, SNLSA Figs 12–13); abdomen elongated or with parallel sides, with four to five pairs of large rounded to triangular markings on terga, always with small macula on 5 th tergum; male genitalia greatly enlarged and strongly sclerotized, apex of hypandrium (superior lobes) and cerci elongated, surstyli three to four times longer than wide (Fig. 6G, Fluke 1957: 278, Fig. 123; SNLSA Figs 14A–D); female tergum 5 th 1/4 of the length of tergum 4 th, tergum 6 th 1/5 of the length of tergum 5 th (SNLSA Figs 13E)...... Talahua Fluke [Note: Montoya & Wolff (2020) provided a complete redescription for Talahua, differential diagnosis, images of the habitus and illustrations of male genitalia] - Antenna elongate, scape longer than broad; basoflagellomere oval or elongate; face straight in profile (Figs 4, 6 – 9, MCAD Fig. 22), facial tubercle low dorsally, not distinct (Figs 4, 6 – 9); mesocoxa bare posteriorly; male genitalia normal size, apex of hypandrium (superior lobes) triangular to rectangular, irregular in shape and cercus short; female tergum 5 th 1/3 of the length of tergum 4 th, tergum 6 th 1/4 of the length of tergum 5 th (Figs 5, 6 and 9)................. Argentinomyia Lynch-Arribálzaga, Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 12-13, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Thompson, F. C. (1999 a) A key to the genera of the flower flies (Diptera, Syrphidae) of the Neotropical Region including descriptions of new genera and species and a glossary of taxonomic terms used. Contributions on Entomology International, 3, 321 - 378.","Thompson, F. C., Rotheray, G. E. & Zumbado, M. A. (2010) Family Syrphidae. In: Brown, B. E. (Ed.), Manual of Diptera of Central America. NRC Press, Ottawa, pp. 763 - 792.","Huo, K. K. (2014) Spazigasteroides a new genus of China with black face and scutellum in the Syrphini (Diptera: Syrphidae). Zootaxa, 3755 (3), 230 - 240. https: // doi. org / 10.11646 / zootaxa. 3755.3.2","Thompson, F. C. & Skevington, J. H. (2014) Afrotropical flower flies (Diptera: Syrphidae). A new genus and species de Kenya, with a review of the melanostomine group of genera. Zootaxa, 3847 (1), 97 - 114. https: // doi. org / 10.11646 / zootaxa. 3847.1.5","Young, A. D., S. A. Marshall, & Skevington, J. H. (2016) Revision of Platycheirus Lepeletier and Serville (Diptera: Syrphidae) in the Nearctic north of Mexico. Zootaxa, 4082 (1), 1 - 317. https: // doi. org / 10.11646 / zootaxa. 4082.1.1","Miranda, G. F. G. & Moran, K. (2017) The female abdomen and genitalia of Syrphidae (Diptera). Insect Systematics & Evolution, 48, 157 - 201. https: // doi. org / 10.1163 / 1876312 X- 48022153","Ramage, T., Charlat, S. & Mengual, X. (2018) Flower flies (Diptera, Syrphidae) of French Polynesia, with the description of two new species. European Journal of Taxonomy, 448, 1 - 37. https: // doi. org / 10.5852 / ejt. 2018.448","Mengual, X. (2020) Phylogenetic relationships of the bacchine flower flies (Diptera: Syrphidae) based on molecular characters, with a description of a new species of Melanostoma (Schiner, 1860). Contributions to Zoology, 82 (2), 210 - 244. https: // doi. org / 10.1163 / 18759866 - 20191410","Mengual, X., Miranda, G. F. G. & Thompson, F. C. (2018) Unraveling Ocyptamus and the Baccha legacy (Diptera: Syrphidae): redefinition of groups and new species descriptions. Zootaxa, 4461 (1), 1 - 44. https: // doi. org / 10.11646 / zootaxa. 4461.1.1","Borges, Z. M. & Pamplona, D. M. (2003) Revision of the Neotropical Xanthandrus Verral (Diptera, Syrphidae). Revista Brasileira de Entomologia, 47, 155 - 167. https: // doi. org / 10.1590 / S 0085 - 56262003000200002","Andersson, H. (1970) Taxonomic notes on the genera Platycheirus and Melanostoma (Dipt., Syrphidae) with lectotype designations. Entomologica Scandinavica, 1, 236 - 240. https: // doi. org / 10.1163 / 187631270 X 00087","Shatalkin, A. I. (1975) A taxonomic analysis of the hoverflies (Diptera, Syrphidae). Entomologicheskoe Obozrenie, 54, 894 - 909.","Haarto, A. & Stahls, G. (2014) When mtDNA COI is misleading: congruent signal of ITS 2 molecular marker and morphology for North European Melanostoma Schiner, 1860 (Diptera, Syrphidae). ZooKeys, 431, 93 - 134. https: // doi. org / 10.3897 / ZooKeys. 431.7207","Fluke, C. L. (1943) A new genus and new species of Syrphidae (Diptera) from Ecuador. Annals of the Entomological Society of America, 36, 425 - 431. https: // doi. org / 10.1093 / aesa / 36.3.425","Vockeroth, J. R. (1990) Revision of the Nearctic species of Platycheirus (Diptera, Syrphidae). The Canadian Entomologist, 122, 659 - 766. https: // doi. org / 10.4039 / ENT 122659 - 7","Montoya, A. L. & Wolff, M. (2020) Description of six new large species of Argentinomyia Lynch-Arribalzaga, 1891 and redescription of Talahua fervida (Fluke, 1945) (Diptera, Syrphidae, Syrphinae). ZooKeys, 929, 19 - 51. https: // doi. org / 10.3897 / zookeys. 929.37666"]}

Montoya, Augusto L., Wolff, Marta (2023): Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species. Zootaxa 5234 (1): 1-157, DOI: https://doi.org/10.11646/zootaxa.5234.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5234.1.1

Argentinomyia octomaculata (Enderlein, 1938) Proposed standard English name: Eight-spotted long-antennae flower fly. Allograptina octomaculata Enderlein, 1938: 226. Type locality: MEXICO. HOLOTYPE Female ZMHU (non-examined). Refs.: Thompson 1999a: 375, Fig. 13–15; Thompson 1999b: 51 (Species incertae sedis status); Thompson et al. 2010, 779, couple 84 (key to Argentinomyia octomaculata) Allograptina octomaculata. Vockeroth 1969: 138 Allograptina octomaculata. Thompson et al. 1976: 39 Rhysops octomaculatus. Thompson 1999a: 338 (taxonomic status notes) Type material. HOLOTYPE. Adult female, MEXICO. No precise locality was given. Deppe S. (ZMB 4012). Length. Body, 7.5–8.3mm; Wings, 7.2–7.7mm. Diagnosis. Face, scutellum and pleuron partially pale in color, usually yellow or yellowish-brown in ground color; face with three to four transversal grooves, calypter wholly brown; all coxae yellow; abdomen with a pair of long, apically rounded oblique maculae on 2 nd to 4 th terga. Redescription. FEMALE. Head (Adapted from Enderlein, 1938 and Thompson 1999a). Face orange, perpendicular, tubercle low, covered with yellow golden pilose, leaving a lineal spot in the middle dorsal antennae and a broad median vitta ventral shining, somewhat metallic yellow. Frontal triangle whitish pollinose, with a broad opaque band just ventral ocelli, gradually fading into white pollen in depression, pile brownish. Gena orange more thinly yellow pollinose, yellow pilose. Ocellar triangle grey metallic, black pile. Occiput smooth, yellow pilose. Antennae yellow-brownish, scape and pedicel about as long as basoflagellomere darkened on dorsal 1/2 and at apex; arista reddish, darker toward the tip, pilose. Thorax: Grey to yellow-orange, mesonotum grey-blue semishning, grey pollinose, yellow-golden pilose; with two medians gray pollinose vittae on anterior half. Pleura yellowish-white pollinose, yellow pilose, yellow on postpronotum and supracoxa, the middle region of anepimeron, dorsal surface of katepisternum, katatergum, anatergum and postalar callus. Scutellum yellow shining, yellow-golden pollinose, with long yellow hairs and short yellow pile intermixed; fringe short and yellow. Wing: Hyaline yellow ocher, slightly brownish on apex, stigma yellow ocher, mostly bare, costal cell bare, cells dm and r bare, cell cua bare on anterior 1/3; almost bare on basal 2/3 or more; calypter wholly brown; plumula yellow, halter yellow-orange, capitulum orange. Legs: Pro- and metalegs brown-yellow, yellow pilose; metafemur brownish except yellowish on underside near the base and dorsal apex; metatibia yellowish on basal 1/6; tarsus brownish, pile black, with a few yellow hairs toward femur bases; metabasitarsomere long, five to six times longer than broad, all coxae yellow. Abdomen: Oval, 1 st tergum brown with base and lateral sides yellow; 2 nd tergum brownish-orange, elongate, apically rounded, lateral maculae along the outer portion of tergum; 3 rd and 4 th terga with two elongated oval maculae, reaching the apical 2/3 of tergum respectively; 5 th tergum with slightly smaller maculae that begin at the base; sterna yellowish, black pilose. Male. Unknown. Taxonomic notes. Argentinomyia octomaculata is similar to A. festiva and A. catabomba, differing by the following combination of characters: Face, scutellum and pleuron partially pale in color, usually yellow or yellowishbrown in ground color; face with three to four transversal grooves; abdomen with a pair of long, apically rounded maculae on 2 nd to 4 th terga. A. octomaculata is also similar to A. CR-21 and A. humboldti sp. nov. differing from them by the facial tubercle yellow; calypter wholly brown; all coxae yellow; abdomen with a pair of long, apically rounded oblique maculae on 2 nd to 4 th terga (see “diagnosis” under each species or key). Geographical range. Argentinomyia octomaculata is so far only known from Mexico (Mexico D.F.)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on page 101, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Enderlein, G. (1938) Beitrag zur Kenntnes der Syrphiden. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1937, 192 - 237.","Thompson, F. C. (1999 a) A key to the genera of the flower flies (Diptera, Syrphidae) of the Neotropical Region including descriptions of new genera and species and a glossary of taxonomic terms used. Contributions on Entomology International, 3, 321 - 378.","Thompson, F. C. (1999 b) Data Dictionary and standards [for fruit fly systematic information database]. Myia, 9, 49 - 63.","Thompson, F. C., Rotheray, G. E. & Zumbado, M. A. (2010) Family Syrphidae. In: Brown, B. E. (Ed.), Manual of Diptera of Central America. NRC Press, Ottawa, pp. 763 - 792.","Vockeroth, J. R. (1969) A revision of the genera of Syrphini (Diptera, Syrphidae). Memoirs de la Entomological Society of Canada, 62, 1 - 176. https: // doi. org / 10.4039 / ENTM 10162 FV","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898"]}

Argentinomyia lanei (Fluke, 1936) Proposed standard English name: Lanei’s long-antennae flower fly. (Figs 40, 41 and 94C) Melanostoma lanei Fluke, 1936: 61. Type locality: Brazil. S„o Paulo, S„o Paulo. HOLOTYPE Male AMNH (Examined). Refs.: Fluke, 1936: 61 (Fig. 4, male abdomen, Fig. 5, head, lateral view, male); 1945: 26 (Fig. 14, head, lateral view, female); 1957: 277 (Figs 103–104, male genitalia); Marinoni & Thompson, 2004: 567 Rhysops lanei. Curran, 1937: 2 (key) Rhysops lanei. Fluke, 1957: 266 Rhysops lanei. Thompson et al. 1976: 43 Argentinomyia lanei. Marinoni & Thompson, 2004: 567 Argentinomyia lanei. Thompson & Skevington, 2014: 98 Rhysops lachrymellea Hull, 1942: 10. Type locality: Brazil. S„o Paulo. HOLOTYPE Male CNC Rhysops lachrymellea. Thompson et al. 1976: 43 Rhysops lachrymellea. Cooper & Cumming, 1993: 70 Type material. HOLOTYPE. Adult Male, BRAZIL. São Paulo, 800 m, 1.i.1930, J. Lane (1♁, AMNH). ALLOTYPE, label with the same data as Holotype: (1♀, Hull’s collection). PARATYPE. Same data as Holotype (1♁, Collection de Institute São Paulo, Brazil); Same data as Holotype: (2♀, AMNH). “ PARATYPE / Rhysops / lachrymellea ”. Length (n= 4): Body, 6.3–7.4 mm; Wings, 5.9–6.7 mm. Diagnosis. Female frontal triangle with two rounded maculae; face with low tubercle, one transversal groove; mesonotum shining bronze to aeneous, reticulated with brown vittae on anterior half; metafemur at least 1/4 yellow basally; abdomen narrowed toward the base, with a pair of oblique maculae on 3 rd to 4 th terga; surstylus with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad; hypandrium narrowed laterally towards the apex; aedeagal lobe with apex acute. Redescription. MALE. Head (Fig. 40A): Face black, covered with white pollen which is slightly brownish alongside; mid-vitta metallic black, bare, violet margined, reaches from oral edge to just dorsal the flat tubercle and then forks so that there is a median dash of pollen extending from antennae almost to the tubercle; a large shining spot dorsal the antennae, with a single transverse groove; pile of the head all pale except a few black hairs on vertical triangle. Gena black, covered with white pollen which is slightly brownish. Ocellar triangle black with thin yellowish pollen, pile yellow. Occiput with yellow hairs dorsal, pollen and pile paler ventral. Antennae yellow, short, ratio 1.5:1.0:1.8, basoflagellomere brown dorsal; pedicel and scape subequal, together with about the length of basoflagellomere, black pilose; arista yellow at base and dark on apical 1/3, pilose. Thorax (Figs 40B–C): Punctulate, dull grey except on sides which are more aeneous, yellow pilose; with three brownish vittae on anterior half, a median slender brown vitta extending along the thorax, on each side a wider brownish vitta. Pleura shining aeneous, posterior margin of anepisternum somewhat reddish, pollen whitish and sparse. Scutellum punctulate, shining greenish-brassy to aeneous. Wing (Fig. 40C): Hyaline, stigma slightly yellowish, microtrichose, except cell c bare basal 1/2, cells bm and r almost bare, cell cua bare on anterior 1/3; calypter wholly yellow-whitish; plumula white; halter yellow, capitulum orange to reddish. Legs (Fig. 40C): Pro- and mesolegs light orange-brown; metafemur brown except for yellow on basal 1/2 and the tip; metatibia black to brownish; tarsus infuscate; pile pale, no peculiar hairs present. Abdomen (Fig. 40B): Slender, narrowed toward the base, semi-shining black with three pairs of elongate, slightly oblique, yellow maculae, large lateral maculae on 2 nd tergum separated by a 1/3 of segment’s width and covering entire lateral margin but attenuated posteriorly; on 3 rd tergum larger, more narrowly separated, oblong and somewhat oblique; the pair that posteriorly diverge and turn out to the lateral margin in the middle of the segment, basal corners of this segment are dark; the 4 th tergum with quite similar maculae which, however, do not reach lateral margin; sterna yellowish-orange, sterna 4 th yellow basally and brown on apical 3/4; pile short and yellow; cercus and surstyle yellowish to reddish; male genitalia: surstylus in lateral view (Fig. 41A) with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad; hypandrium in ventral view (Fig. 41C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 41C) with apex acute. FEMALE (Figs 40D–F). Similar to male except for usual sexual dimorphism and differing in frontal triangle more brownish pollinose and there is a definite dark brown transverse band just in front of median ocellus. There are indications of small yellow maculae on 5 th tergum. Scutellum apex with a pair of slender bristle yellow. Taxonomic notes. Argentinomyia lanei is similar to A. festiva and A. fastigata, distinguished from its congeners by the combination of characters: Female frontal triangle with two rounded maculae; face with low tubercle, one transversal groove (Figs 40 D-F); mesonotum shining bronze to aeneous, reticulated with brown vittae on anterior half (Figs 40B, E); metafemur at least 1/4 yellow basally (Figs 40A, C, D-F); abdomen narrowed toward the base, with a pair of oblique maculae on 3 rd to 4 th terga (Figs 40B, E). Based on males, A. lanei differs from A. fastigata and A. festiva in having the surstylus in lateral view (Fig. 41A), with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length, elongated, at least two times longer than broad in A. festiva (Fig. 33A); surstylus with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad in A. fastigata (Fig. 31A)]; hypandrium in ventral view, narrowed laterally towards the apex in A. lanei (Fig. 41C) and A. festiva (Fig. 33C) [versus hypandrium expanded laterally towards the apex in A. fastigata (Fig. 31C)]; aedeagal lobe in ventral view, with acute apex in A. lanei (Fig. 41C) and A. festiva (Fig. 33C) [versus aedeagal lobe circular, apex rounded in A. fastigata (Fig. 31C)] (see “differential diagnosis” under each species or key). Comments. The Holotype specimen of R. lanei is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue.10023898. Geographical range. Argentinomyia lanei (n= 58) is distributed in Argentina * (Tucumán), Serra de Fartura in Brazil (Minas Gerais, Paraná, Santa Catarina, S„o Paulo) and the Eastern slope of Central cordillera in Ecuador * (Azuay) (Fig. 94C). The species is present at low and middle altitudes (400–2650 m) in the following biogeographical domains and provinces: Northern Andes: Cauca (2650 m); Paraná (400–1274 m): Araucaria Forest (934–1274 m), Atlantic Forest (800 m), Paraná Forest (400–857 m); Chacoan (745–1114 m): Cerrado (745 m), Chaco (1114 m). Non-type material examined. ARGENTINA. Tucumán, Burruyacu, Villa Padre Monti, -26.482644, - 64.966915, 1114 m, i.17-ii.7-1948, R. Golbach, Det. C.L. Fluke, 1951 (1♁, 1♀, FSCA, C.L. Fluke, 1951). BRAZIL. Minas Gerais, Passos, -20,7189, -46,6097, 745 m, 11.i.1961, C. Elias (1♀, DZUP 45574, Det. H. S. Telford); Paraná, Curitiba, -25,4278, -49,2731, 934 m, 1.4.1976, H. S. Telford (1♁, DZUP 45570, Det. F.C. Thompson); Curitiba, -25,4278, -49,2731, 934 m, 24.02.1975, H. S. Telford (1♀, DZUP 45573, Det. L. Marinoni); Guarapuava, Estância Santa Clara, -25,3833, -51,45, 1098 m, 8.11.1986, PROFAUPAR (1♀, DZUP 43072, Det. F.C. Thompson); Refugio de vida silvestre dos campos de Palmas, -26,533807, -51,599362, 1200 m, A. Couto (4♁, DZUP); Santa Catarina, Nova Teutonia, -27,183334, -52,383334, 530 m, 1ix.1948, A. Maller (1♀, AMNH); 530, 1.ii.1952, F. Plaumann (1♁, CNC DIPTERA 112201, Det. J.R. Vockeroth); 6.1.1969, F. Plaumann (1♁, CNC DIPTERA 112202– 03, Det. J.R. Vockeroth); 857 m, 1.vii.1964, F. Plaumann (23♁, 7♀, USNM ENT 01384856; 01406258; 01462446; 01406287; 01406220; 01406269; 01384740; 01406373; 01384707; 01384635; 01406325; 01406369; 01406305; 01384874; 01384503; 01384319; 01406288; 01384776; 01406240; 01384697; 01406225; 01406268; 01406379; 01443846; 01406386; 01406340; 01406296; 01406207; 01406360; 01406291; 01406318); 400 m, 29.vii.1938, F. Plaumann (5♁, 6♀, BMNH); 857 m, 6.i.1973, F. Plaumann (1♁, Field Museum Natural History 1975; D. L. Pearson Colen Acc Z-15, 289). ECUADOR. Azuay, San José de Raranga, -3,221361, -78,957493, 2650m (CNC)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 70-72, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Fluke, C. L. (1936) New Syrphidae (Diptera) from Brazil and Cuba. Journal of the Kansas Entomological Society, 9, 59 - 65.","Marinoni, L. & Thompson, F. C. (2004) Flower flies of southeastern Brasil (Diptera: Syrphidae). Part I. Introduction and new species. Studia Dipterologica, 10, 565 - 578. Available from: https: // corpora. tika. apache. org / base / docs / govdocs 1 / 093 / 093859. pdf (accessed 13 January 2023)","Curran, C. H. (1937) The Neotropical species of Melanostoma and allies (Syrphidae: Diptera). American Museum Novitates, 926, 1 - 4.","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Thompson, F. C. & Skevington, J. H. (2014) Afrotropical flower flies (Diptera: Syrphidae). A new genus and species de Kenya, with a review of the melanostomine group of genera. Zootaxa, 3847 (1), 97 - 114. https: // doi. org / 10.11646 / zootaxa. 3847.1.5","Hull, F. M. (1942) Some new species of Syrphidae. Journal of the Kansas Entomological Society, 15, 10 - 12.","Cooper, B. E. & Cumming, J. M. (1993) Diptera types in the Canadian National Collection of Insects. Part 2. Brachycera (exclusive of Schizophora). Research Branch Agriculture Canada Publication 1896 / B. Research Branch Agriculture Canada, Ottawa, 105 pp. [also published electronically at http: // www. nadsdiptera. org / Catalogs / CNCtypes / Part 2. htm]"]}

Argentinomyia plaumanni Thompson & Montoya sp. nov. Proposed standard English name: Plaumanni’s long-antennae flower fly. Argentinomyia 74–06 Thompson, unpublished (Figs 61, 62 and 93D) Type material. HOLOTYPE. Adult male, pinned, deposited at National Museum of Natural History. Original label: “ BRASIL, Santa Catarina, Nova Teutonia” / “ -27,183334, -52,383334, 857m 10.i.1965, F. Plaumann ” “ HOLOTYPE / Argentinomyia plaumanni / Thompson & Montoya 2023” (USNM ENT 00036936). Identified as Argentinomyia 74–06 by Thompson 1974. PARATYPES. BRAZIL. Labels: same data as Holotype (7♁, USNM ENT 00036934; 00036931; 00036932; 00036937; 00036933; 00036935; 0036938); BRASIL. Curitiba, 3.i.1954, N.L.H. Krauss (1♁, USNM ENT 01443774, dissected). Length (n= 5): Body, 8.6–9.1 mm; Wings, 7.2–7.4 mm. Diagnosis. Legs black to brownish; except femur yellow on apical 1/4; pro-, meso- and metatibia yellow on basal 1/3, 1/4 and 1/2, respectively, tarsus black; black pilose; abdomen black, 2 nd tergum with a pair of basolateral elongate yellow maculae reaching the apical 3/4 of the segment length; 3 rd tergum with a pair of basal elongated maculae reaching apical 2/3, but not reaching lateral margin; 4 th and 5 th terga black; surstylus with dorsal margin slightly concave and ventral margin slightly convex. Description. MALE. Head (Fig. 61A): Face perpendicular with four faint transverse depressions, tubercle elongate, broad and low; mid-vitta shining black, sides white pollinose and pilose. Frontal triangle shiny, coppery pollinose, black pilose. Gena shining, white pollinose. Ocellar triangle black, black pilose; occipital pile fine, short and black, white pollinose. Antennae orange, short, ratio 1.2:1.0:2.0; scape darker, basoflagellomere darkened dorsally, scape only very little longer than pedicel which is wider than long, basoflagellomere short, little longer than the scape and pedicel together; arista yellow at base and dark on apical third. Thorax (Figs 61B–C): Aeneous, scutum with two very faint pollinose white vittae on anterior half, yellow-golden pilose. Pleura lightly coated with brownish to rusty-colored pollen, yellow pilose. Scutellum black, with long golden pile, in addition to some brown pile in the margin. Wing (Fig. 61C): Slightly yellowish, stigma yellow, microtrichose, except bare on basal 1/4 of cell bm, cell cua bare basally; calypter wholly whitish; plumula yellow; halter yellow, capitulum yellow. Legs (Fig. 61C): Black to brownish; except femur yellow on apical 1/4; pro-, meso- and metatibia yellow on basal 1/3, 1/4 and 1/2, respectively, tarsus black., black pilose. Abdomen (Fig. 61B): Narrow with nearly parallel sides, shining black, 2 nd tergum with a pair of basolateral elongate yellow maculae reaching the apical 3/4 of the segment length; 3 rd tergum with a pair of basal elongated maculae reaching apical 2/3, but not reaching lateral margin; 4 th and 5 th terga black, black pilose, sterna yellow pilose; male genitalia: surstylus in lateral view (Fig. 62A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium in ventral view (Fig. 62C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 62C) circular, apex rounded. FEMALE. Unknown. Taxonomic notes. Argentinomyia plaumanni sp. nov. is similar to A. nigrans differing by the following combination characters: Legs black to brownish; except femur yellow on apical 1/4; pro-, meso- and metatibia yellow on basal 1/3, 1/4 and 1/2, respectively, tarsus black, black pilose (Figs 61A, C, D, F); 2 nd tergum with a pair of basolateral elongate yellow maculae reaching the apical 3/4 of the segment length (Figs 61B, E). In A. nigrans the femora and tarsi black (Figs 53A, C, D, F); abdomen black with small yellow to red maculae on basal corners of the 2 nd tergum, reaching over the sides and 1/2 of segment length (Figs 53B, E). Based on males, A. plaumanni sp. nov. differs from A nigrans in having the surstylus in lateral view (Fig. 62A) with dorsal margin slightly concave and ventral margin slightly convex [versus surstylus with dorsal and ventral margins similar (square-like) in A nigrans (Fig. 54A)] (see “diagnosis” under each species or key). Etymology. The specific epithet of this species is a Latinized noun in the genitive case of the last name ‘Plaumann’ in honor of Fritz Plaumann, who collected the type series in Nova Teutonia. Geographical range. Argentinomyia plaumanni sp. nov. (n= 9) is exclusively known from southeastern Brazil (Santa Catarina and Paraná) (Fig. 93D). The species is endemic to the Paraná domain at low altitudes (857 m) in the Paraná Forest province.

Argentinomyia maculata (Walker, 1852) (Figs 49, 50 and 93C) Paragus maculatus Walker, 1852: 220. Type locality: Brazil, Santa Catarina, Nova Teutonia. HOLOTYPE Female BMNH (Examined) (Previously SYNTYPE) Rhysops maculatus. Thompson et al. 1976: 43 Argentinomyia maculatus. Navarro et al. 2009: 520 Rhysops minuscula Fluke, 1945: 11. Type locality: Brazil. Santa Catarina, Nova Teutonia. HOLOTYPE Male AMNH (Examined). Refs.: Fluke, 1945: 27 (Fig. 16, head, frontal view, male, Fig. 17, antenna, lateral view) 29 (Fig. 47, male abdomen); 1957: 279 (Figs 109–110, male genitalia) Rhysops minuscula. Fluke, 1957: 266 Melanostoma discretum Frey, 1946: 157. Type locality: Brazil. Santa Catarina, Nova Teutonia. HOLOTYPE Male ZMHU Type material. HOLOTYPE. Adult Female, BRAZIL. Santa Catarina, Nova Teutonia (BMNH, Previously SYNTYPE). “ SYNTYPE / Paragus maculatus / Walker, 1852 ”. HOLOTYPE Adult Male, BRAZIL. Santa Catarina, Nova Teutonia, 300-500 m, 2.v.1937, F. Plaumann (BMNH) “ HOLOTYPE / Rhysops / minuscula ”. PARATYPE. BRAZIL. Label with the same data as HOLOTYPE except: 3.v.1937 and 1.xi.1938, F. Plaumann (11♁, 1♀, BMNH); 24.vi.1937 and 29.vii.1938 (15♀, BMNH; 1♁, 1♀, AMNH); 500 m, 1964-1974 (1♁, 2♀, USNM ENT 00058844); 2.vi.1937 (1♀, AMNH) “ ALLOTYPE / Rhysops / minuscula ”; 3.vi-1.ix.1937 and 1938; vi-x.24.1937 and 1938 (11♁, 15♀, AMNH; 3♁, 3♀, WIRC) “ PARATYPE / Rhysops / minuscula ”. Length (n= 5): Body, 5.4–6.1 mm; Wings, 3.8–4.5 mm. Diagnosis. Small species, female frontal triangle with two rounded brown pollinose maculae; 3 rd to 4 th terga with triangular maculae; metatibia mostly yellow; metafemur yellow with a black ring or smudge on apical margin; surstylus in lateral view with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium in ventral view expanded laterally towards the apex; aedeagal lobe in ventral view with apex acute. Redescription. MALE. Head (Fig. 49A): Face black, very highly polished blue-black; sides of face black, whitish on gena and lower occiput; mid-vitta highly polished with a narrow gray pollinose vitta, which reaches from the base of antennae almost to the first depression; this pollinose vitta connects with a narrow vitta, which follows downsides of the face to the oral margin and connects with a broad patch of pollen, which in turn connects to a narrow strip along the eyes, broadening on frontal triangle with a narrow point projecting toward the antennae, but not quite reaching them; tubercle low and with two faint transverse depressions; side of the face between tubercle and the eyes is a flattened rugose patch which has about three ridges on dorsal half of patch. Frontal triangle pollen brownish-black but bordered with gray pollen. Ocellar triangle dark brown with coppery reflections; pile all black. Occiput metallic blackish-black, glossy, whitish pollinose and white pilose, only black pilose on dorsal 1/3. Antennae yellow, short, ratio 1.2:1.1:1.8, scape and pedicel about equal in length, pedicel about half as long as scape or basoflagellomere, yellow in color, basoflagellomere black dorsal and at the apex; arista yellow at base and dark on apical 1/3, pilose. Thorax (Figs 49B–C): Brown to black metallic, mesonotum dark, black pilose with paler hairs on disc; with two faints brownish pollinose vittae on anterior half. Pleura shining, very lightly pollinose, especially around propleura, pile all yellowish-brown. Scutellum highly polished with a greenish-black hue, pile light brown. Scutellum with slightly longer hairs; disc with about five faint transverse depressions; notopleura tubercle very prominent. Wing (Fig. 49C): Hyaline, stigma light brownish, microtrichose, except cell c bare on basal 1/3, cell bm bare on basal 1/3 and r bare on basal 1/4 or more, cell cua microtrichose; calypter wholly dark brown; plumula light golden; halter yellowish red, capitulum slightly brownish. Legs (Fig. 49C): Yellow; pro- and mesofemora and tibiae yellow; metafemur yellow on basal 2/3, with a darker smudge on apical margin, pile sparse, long, black on dark areas; metatibia yellow on middle 1/4; tarsus yellow. Abdomen (Fig. 49B): Somewhat spatulate, opaque to dull black but shining alongside; with two pairs of yellow triangular maculae located in the anterior corners of 3 rd and 4 th terga; yellow pilose and rather elongate alongside the 2 nd tergum; sterna polished, the incisure between the 2 nd and 3 rd sternites yellow; male genitalia: polished black, with black pile; surstylus in lateral view (Fig. 50A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium in ventral view (Fig. 50C) expanded laterally towards the apex; aedeagal lobe in ventral view (Fig. 50C) with apex acute. FEMALE (Figs 49D–F). Similar to male except for usual sexual dimorphism and differing in the pollen on face whitish, as is also the pile. Frontal triangle is wide with two large dark brown pollinose maculae, which are narrowly connected and are located in front of the ocelli; a narrow band of white pollen separates the dark spots from the ocelli and a narrow brown vitta runs from the median ocellus to highly polished lunule. Mesonotum more shining blackish, with vittae bluish in color and pile all white. The abdomen is more oval, semi-shining on disc, highly polished alongside; yellow spots often obscured. Taxonomic notes. Argentinomyia maculata is similar in appearance to A. pollinosa differing from them by the female frontal triangle with two rounded brown pollinose maculae (Figs 49D, F); metafemora yellow with a black ring or smudge on apical margin; metatibiae mostly yellow (Figs 49A, C, D, F); 3 rd to 4 th terga with triangular maculae (Figs 49B, E). In A. pollinosa the female frontal triangle with a complete transverse black fascia (Figs 63D, F); metafemora yellow except the apical 1/5 narrowly brown; metatibiae yellow on basal 1/4 (Figs 63A, C, D, F); 3 rd to 4 th terga with quadrate maculae (Figs 63B, E). Based on males, A. maculata differs from A. pollinosa in having the surstylus in lateral view (Fig. 50A) with dorsal margin slightly concave and ventral margin slightly convex [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. pollinosa (Fig. 64A)]; aedeagal lobe in ventral view (Fig. 50C) with apex acute [versus aedeagal lobe circular, with apex rounded in A. pollinosa (Fig. 64C)] (see “diagnosis” under each species or key). Comments. The Holotype specimen of R. minuscula is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue. 10021028. Geographical range. Argentinomyia maculata (n= 153) is the second most abundant species in collections, distributed in Argentina * (Tucumán) and Serra da Fartura and Serra do Mar in Brazil (Paraná, Santa Catarina) (Fig. 93C). The species is present at low and middle altitudes (400–1274 m) in the following biogeographical domains and provinces: Paraná (400–1274 m): Araucaria Forest (895–1274 m), Paraná Forest (400–500 m); Chacoan: Chaco (526–915 m). Non-type material examined. ARGENTINA. Tucumán, Burruyacu, Quebrada La Toma, -26.593592, - 64.811185, 526 m, 21.xii.1950, R. A. Golbach (2♁, 1♀, WIRC); Villa Padre Monti, -26.488779, -64.971602, 915 m, 5.ii.1948, R.A. Golbach (1♁, 2♀, WIRC). BRAZIL. Paraná, Refugio de vida silvestre dos campos de Palmas, -26,533807, -51,599362, 1200 m, A. Couto (4♁, DZUP); Guarapuava, Estância Santa Clara, -25,3833, -51,45, 1098 m, 10.vi.1986, PROFAUPAR (1♀, DZUP 43075, Det. F.C. Thompson); Ponta Grossa, Pq Estadual de Vila Velha-IAPAR, -25,246336, -50,02104, 895 m, 15.xi.2000, N. G. Ganho and R. C. Marinoni (1♀, DZUP 42303, Det. L. Marinoni); Antonina, Reserva Biológica Estadual Sapitanduva, -25,4667, -48,8333, 60 m, 11.iii.1986, PROFAUPAR (1♁, DZUP 43074, Det. F.C. Thompson); Santa Catarina, Nova Teutonia, -27,183334, -52,383334, 500 m, 1.vii.1964, F. Plaumann (1♁, 2♀, USNM ENT 00058844; 00058844); 10.v.1937, F. Plaumann (11♁, 15♀, AMNH); (3♁, 3♀, WIRC); 5.ii.1937, F. Plaumann (1♀, AMNH); 2.iii.1937, F. Plaumann (27♀, BMNH); 6.viii.1937, F. Plaumann (1♁, AMNH); 29.vii.1938, F. Plaumann (8♁, BMNH); 6.i.1964, F. Plaumann (1♁, CNC DIPTERA 112208, Det: J.R. Vockeroth); Same data, except: (22♁, 12♀, USNM ENT 01406354; 01406293; 01406278; 01406326; 01406193; 01406346; 01406402; 01406372; 0140233; 01406199; 01406303; 01406323; 01406378; 01406194; 01406339; 01406343; 014006381; 01406263; 01406259; 01406230; 01444377801406257; 01406341; 01406261; 01406301; 01406336; 01406254; 01406327; 01406310; 01406355; 01406210; 01406290; 0140639; 01406266; 01406397); 11.i.1969, F. Plaumann (1♁, 1♀, CNC DIPTERA 112209-10, Det: J.R. Vockeroth); 7.i.1971, F. Plaumann (1♁, 3♀, AMNH)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 85-88, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Walker, F. (1852) Diptera. In: Saunders, W. W. (Ed.), Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq., F. R. S., F. L. S., & c. Vol. 1. Van Voorst, London, pp. 157 - 414.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Navarro, F. R., Cuezzo, F., Goloboff, P. A., Szumik, C., Lizarralde De Grosso, M. & Quintana, M. G. (2009) Can insect data be used to infer areas of endemism? An example from the Yungas of Argentina. Revista Chilena de Historia Natural, 82 (4), 507 - 522. https: // doi. org / 10.4067 / S 0716 - 078 X 2009000400006","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Frey, R. (1946) Ubersicht der Gattungen der Syrphiden-Unterfamilie Syrphinae (Syrphinae + Bacchinae). Notulae Entomologicae, 25, 152 - 172."]}

Argentinomyia altissima (Fluke, 1945) Proposed standard English name: High elevations long-antennae flower fly. (Figs 12, 13 and 90C) Melanostoma altissimum Fluke, 1945: 20. Type locality: Ecuador. Imbabura, Cuicocha, 3200m. HOLOTYPE Male AMNH (Examined). Refs.: Fluke, 1945: 29 (Fig. 48, female abdomen, Fig. 49, male abdomen); 1957: 276 (Figs 95–96, male genitalia); Ángel-Villarreal et al. 2021: 5 (distrib. record) Rhysops altissimum. Fluke 1957: 266 Rhysops altissimus. Thompson et al. 1976: 42 Argentinomyia altissimus. Thompson & Skevington, 2014: 98 Argentinomyia altissima. Montoya, 2016: 459 Argentinomyia altissima. Montoya et al. 2017: 395 Argentinomyia altissima. Marín-Armijos et al. 2017: 168 Type material. HOLOTYPE. Adult Male, ECUADOR. Cuicocha, Imbabura, 3200 m, 27.v.1939, F.M. and H. Brown (AMNH). PARATYPE. Two adult females. ECUADOR. Deposited at AMNH. Label the same data as HOLOTYPE. Length (n= 2): Body, 8.2–8.9 mm; Wings, 7.7–8.1 mm. Diagnosis. Facial pile mostly black; legs black, pro- and mesotibiae mostly brownish with a dark median ring; abdominal maculae almost as wide as long, maculae on 3 rd tergum of male quadrate and on female triangular; surstylus very elongated and widened ventrally, with a median-sized extension in the dorsal margin; aedeagal lobe hearth shape, with the apical margin concave. Redescription. MALE. Head (Fig. 12A): Face black, nearly perpendicular, tubercle low yet broad, mid-vitta broad, shining and bare laterally. Frontal triangle shining black dorsally, the narrowband ventral to the antennae mostly black, yellow only on ventrolateral part of the frontal triangle. Gena shining black with thin, more whitish pollen, pile white. Ocellar triangle black with thin yellowish pollen, pile black. Occiput with yellow hairs dorsal, long cilia black, pollen and pile paler ventral. Antennae reddish yellow, short, ratio 1.4:1.0:1.8, basoflagellomere about as long as scape and pedicel together, scape darker, basoflagellomere darkened dorsally and orange ventrally, basoflagellomere apically rounded, arista long and black, bare. Thorax (Figs 12B–C): Black, mesonotum shining obscured by brown pollen, with two median whitish pollinose vittae on anterior half, pile long and mostly black; pleura coated with brownish pollen which is whitish posteriorly, yellow pilose, anterior anepimeron black pilose. Scutellum with long black hairs with some very short golden hairs intermixed, fringe long and yellow. Wing (Fig. 12C): Hyaline, stigma yellow, membrane microtrichose, except bare on basal 1/6 of bm and 1/5 of the cua; calypter whitish, border and fringe brown; plumula white; halter yellow, capitulum brown. Legs (Fig. 12C): Mostly black, pro- and mesotibiae extensively brownish with a dark median ring, brownish pilose. Abdomen (Fig. 12B): Abdominal maculae wider, almost quadrate, reaching less than 1/2 length of 3 rd and 4 th terga; on 2 nd tergum shows evidence of narrow elongate maculae restricted to the lateral edge; sterna yellowish, yellow pilose; male genitalia: surstylus in lateral view (Fig. 13A) very elongated and widened ventrally, with a median sized extension in the dorsal margin; hypandrium in ventral view (Fig. 13C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 13C) hearth shape, with the apical margin concave. FEMALE (Figs 12D–F, 13D–E). Similar to male except: for usual sexual dimorphism, frontal triangle with a transverse band on lower 1/2, a distinct depression in the middle. Mesonotal pile sparse and much shorter than in male. Argentinomyia altissima differs from A. bolivariensis only in the darker legs and triangular abdominal maculae; on 2 nd tergum with a pair of elongate oval maculae; 3 rd and 4 th terga with triangular maculae; 3 rd tergum with inner margins parallel, their outer margins converging, separated from lateral margins; smaller maculae on 5 th tergum. Female A. bolivariensis is characterized by abdominal maculae often absent but when present, they show clearly reddish dashes converging posteriorly. Taxonomic notes. Argentinomyia altissima is remarkably similar in morphology to A. bolivariensis, differing by the following combination of characters: facial pile of male mostly black (Fig. 13A, C, D-F); legs black, pro- and mesotibiae mostly brownish with a dark median ring (Fig. 13 B-C, E-F); abdominal maculae almost as wide as long, maculae on 3 rd tergum of male quadrate (Fig. 13B), on female triangular (Fig. 13E). In A. bolivariensis, the face is even brown-yellowish microtrichose except near antennae (Fig. 20A, C, D-F); pro- and mesotibiae without a dark median ring (Fig. 20 B-C, E-F); abdominal maculae slenderer, elongated, about two and 1/2 times as long as wide on 3rd tergum (Fig. 21B); female abdomen generally black and covered by grey pollinosity, at most with small reddish maculae (Fig. 21E). Based on males, A. altissima differs from A. bolivariensis in having the surstylus in lateral view (Fig. 13A) very elongated and widened ventrally, with a median-sized extension in the dorsal margin [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. bolivariensis (Fig. 21A)]; aedeagal lobe in ventral view (Fig. 13C) hearth shape, with the apical margin concave [versus aedeagal lobe with acute apex in A. bolivariensis (Fig. 21C)] (see “diagnosis” under each species or key). Remarks. Argentinomyia altissima, A. bolivariensis, A. rex and A. opaca are found sympatrically throughout the highlands of the northwestern Tropical Andes, where appear to be associated with high Andean Forest and Páramos habitats in Northwestern Colombia based on exclusivity and fidelity criteria (Montoya et al. 2021). Comments. The Holotype specimen of M. altissima is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue. 10021682. Biology. Ángel et al. (in press) analyzed the stomach contents of several specimens of A. altissima from three Andean ecosystems in Colombia, finding that the main source of pollen for this species belongs to the family Asteraceae (n= 3034 pollen grains, 87%), followed by Sapindaceae (n= 191, 6%), Rosaceae (n= 150, 4%), Onagraceae (n= 65, 2%) and Loranthaceae (n= 24, 1%). The palynological analysis showed that 60.4% of the pollen grains consumed by A. altissima belong to the species Baccharis latifolia (n= 2111, Asteraceae), followed by the Asteroideae group (n= 599, 17.1%, Asteraceae), Taraxacum-Hyppochaeris (n= 317, 9.1%, Asteraceae), Allophylus sp. (n= 191, 5.5%, Sapindaceae), Cotoneaster pannosus (n= 150, 4.3%, Rosaceae), Fuchsia sp. (n= 65, 1.9%, Onagraceae), Gaiadendron punctatum (n= 24, 0.7%, Loranthaceae), Croton mutisianus (n= 13, 0.4%, Euphorbiaceae), Vibrurnum sp. (n= 12, 0.3%, Adoxaceae), Senecio madagascariensis (n= 5, 0.1%, Asteraceae), Piper sp. (n= 3, 0.1%, Piperaceae), Brassica – Capsella (n= 2, 0.1%, Brassicaceae) and Bidens pilosa (n= 2, 0.1%, Asteraceae). Field observations and label data also reveal that A. altissima visits flowers of Ageratina tinifolia, Baccharis sp., Diplostephium rosmarinifolium, Espeletia occidentalis, Pentacalia ledifolia (Asteraceae) and Buddleja coriacea (Scrophulariaceae). Specimens collected using canopy Malaise trap suggest that A. altissima flies in the higher tree canopy (height> 4 m). Geographical range. Argentinomyia altissima (n= 108) is a common species, present throughout the western slope of Western Cordillera in Colombia (Antioquia), both slopes of Central Cordillera in Colombia * (Antioquia, Cundinamarca, Tolima) and Ecuador (Cotopaxi, Cuicocha, Pichincha), the western slope of Eastern Cordillera in Colombia (Meta) and Eastern slope of Cordillera Oriental in Perú * (Cuzco) (Fig. 90C). The species is present at middle and high altitudes (2200–3610 m) in the Northern Andes domain and the provinces: Cauca (2200–3610 m), Magdalena (2200–3150 m) and North Andean Páramo (2700 m). Non-type material examined. COLOMBIA. Antioquia, Andes, Cerro Caramanta, Páramo Laguna de Santa Rita, 5.550249, -75.988733, 3610 m, Net, 3-5.iii.2009, M. Wolff (1♁, 1♀, CEUA 47446, 69642); Bello, San Félix, Las Baldías, 6,330 58, -75,645620, 3150 m, Páramo, Net, 16.vii.2016, A. L. Montoya (2♁, CEUA 93080-81); …, Belmira, Páramo Santa Inés, Cabaña Cabildo Verde, El Morro-Alto de La Gallina, 6,657 8611, -75,6715, 3301 m, Van Someren-Rydon (VSR) Fish, 5-8.x.2011, GEUA (2♁, CEUA 43353, 43362); …, 6,647 5556, -75,6713333, 3241 m, Páramo,Arbusto, ex Espeletia occidentalis, Net, 2-5.v.2008, C. Bota (1♁, CEUA 47446); …, ex Pentacalia ledifolia, Net, 4-14.ii.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona (2♁, CEUA 95300); ex Ageratina tinifolia, Net, 4- 14.ii.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona (1♁, CEUA 95379); Net, 2-13.xii.2017, C. Rodríguez; A. Echeverry (2♁, CEUA 98311, A9838); Net, 16-27.ix.2017, C. Rodríguez; A. Rúa (1♁, CEUA 98347); …, Malaise, 2-13.xii.2017, C. Rodríguez; A. Echeverry (1♁, CEUA 98402); 5-8.x.2011, GEUA (1♁, CEUA 43354); …, Malaise canopy, 4-14.xii.2016, A. L. Montoya; J. Torres; J. P. Carmona (4♁, CEUA 95110, 95195, 95456); 25.iii-5.iv.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona; 2-13.xii.2017, C. Rodríguez; A. Echeverry (2♁, CEUA 103346, 95584); 21- 30.vi-5.iv.2017, C. Rodríguez; J. P. Carmona (1♀, CEUA 97994); 6,634 233, -75,658654, 3020 m, Forest, Malaise, 2-13.xii.2017, C. Rodríguez; A. Echeverry (1♁, CEUA 98294); 6,630 476, -75,663106, 2924 m, Grassland, Malaise, 25.iii-5.iv.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona (6♁, CEUA 95520, 95528); …, Belmira, Páramo Santa Inés, Vereda Río Arriba, Sector Laguna de Sabanas, 6,645 901, -75,670441, 3200 m, Páramo, Net, 16-27.ix.2017, A. L. Montoya; Y. Cardona (1♀, CEUA 98219); 6,637 851, -75,662949, 2879 m, Grassland, Malaise, 2-13.xii.2017, J. Torres-Toro; J. P. Carmona (2♁, 1♀, CEUA 98395); Malaise, 2-13.xii.2017, J. Torres-Toro; J. P. Carmona (1♀, CEUA 103644); 16-27.ix.2017, A. L. Montoya; Y. Cardona (1♁, 2♀, CEUA 98201, 98206, 98208); 25.iii-5.iv.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona (1♁, 1♀, CEUA 95558); Malaise, 16-27.ix.2017, A. L. Montoya; Y. Cardona (2♁, CEUA 98317, 98372); …, Net, 2-13.xii.2017, J. Torres-Toro; J. P. Carmona (1♀, CEUA 98363); ex Diplostephium rosmarinifolium, Net, 4-14.ii.2017, A. L. Montoya; C. Rodríguez; J. D. Carmona (4♁, CEUA 95109); San José de la Montaña, Vereda El Congo, Sector La Laguna, 6,763 791, -75,701738, 3117 m, Páramo, Malaise canopy, 4-14.ii.2017, C. Henao; A. F. Sepúlveda (2♁, CEUA 95189); Malaise, 21- 30.vi-5.iv.2017, A. L. Montoya; A. F. Sepúlveda (2♀, CEUA 98088); 2-13.xii.2017, C. Henao; A. F. Sepúlveda (1♀, CEUA 98337); 25.iii-5.iv.2017, M. Wolff; C. Henao; A. F. Sepúlveda (2♀, CEUA 98188); …, 21- 30.vi-5.iv.2017, A. L. Montoya; A. F. Sepúlveda (1♀, CEUA 98023); …, 2-13.xii.2017, C. Henao; A. F. Sepúlveda (1♁, CEUA 98330); …, 2- 13.xii.2017, C. Henao; A. F. Sepúlveda (2♁, ♀1♀, CEUA 98331); 25.iii-5.iv.2017, M. Wolff; C. Henao; A. F. Sepúlveda (2♁, CEUA 97930); Pan trap blue, 4-14.xii.2016, C. Henao; A. F. Sepúlveda (1♀, CEUA 95437); …, Pan trap white, 25.iii-5.iv.2017, M. Wolff; C. Henao; A. F. Sepúlveda (1♀, CEUA 97963); …, Net, 4-14.ii.2017, C. Henao; A. F. Sepúlveda (2♀, CEUA 95196, 95429); …, 25.iii-5.iv.2017, M. Wolff; C. Henao; A. F. Sepúlveda (2♁, 3♀, CEUA 95589, 95594); …, 21- 30.vi-5.iv.2017, A. L. Montoya; A. F. Sepúlveda (4♁, 5♀, CEUA 98077, 98093); …, 2-13.xii.2017, C. Henao; A. F. Sepúlveda (1♀, CEUA 98339); …, 6,768 592, -75,708826, 3012 m, Forest, Pan trap blue, 25.iii-5.iv.2017, M. Wolff; C. Henao; A. F. Sepúlveda (1♀, CEUA 97962); 6,768 593, -75,712085, 2995 m, Grassland, Net, 4-14.ii.2017, C. Henao; A. F. Sepúlveda (1♁, CEUA 95416); 21- 30.vi-5.iv.2017, A. L. Montoya; A. F. Sepúlveda (1♀, CEUA 98047); Sonsón, Vereda San Francisco, Cerro Las Palomas A, 5,725 972, -75,24925, 3364 m, Páramo, Net, 23.vi-2.vii.2018, J. P. Carmona; J. Sauceda; J. Vallejo (1♁, CEUA 103406); …, Vereda Nori Cerro Nori, 5,812 869, -75,268513, 2755 m, Grassland, Malaise canopy, 31.iii.-7.iv.2018, A.L. Montoya; J. P. Carmona (1♀, CEUA 103429); 5,812 861, -75,268444, 3045 m, Páramo, Net, 31.iii.-7.iv.2018, A.L. Montoya; J. P. Carmona (4♁, CEUA 103461); Medellín, Corregimiento de Santa Elena, Vereda Piedras Blancas, en Robledal, 6.291588, -75.493345, 2200 m, Net, 26.ix.2009, C. Bota (1♁, CEUA 47453); Cundinamarca, Bogotá, 4.592901, -74.081040, 2700 m, 25.iii.1937, Osorno (1♁, CNC DIPTERA);.. 25.iii.1937, B. Guerra (2♁, USNM ENT 01443844; 01443835); Tolima, Murillo, Robledal, 4.870242, -75.177614, 3000 m, 6.x.2012, C. Bota, Net (1♁, CEUA 73203); Meta, Meta Dist. 1932, B. Guerra (1♁, USNM ENT 001443836). ECUADOR. Cotopaxi, - 0.773272, -78.549621, 3200 m, 16.i.1993, A. Salazar (1♁, QCAZ 103717); Pichincha, Guayllabamba, -0.059052, -78.341446, 2200 m, en Baccharis sp. (Asteraceae) 27.i.2000, P. Jiménez (1♁, QCAZ 103715); Pichincha, Quito Yuruqui, -0.231633, -78.539767, 2900 m, en Baccharis sp. (Asteraceae) 1.vi.1986, G. Correa (1♁, QCAZ 103719); - 0.231633, -78.539767, 2870 m, en Buddleja coriacea, G. Paz and Miño (3♁, QCAZ 103716-18 &..3720); Quito, -0.180464, -78.465438, 2800 m, 8.vii.1984, L. Coloma (1♁, QCAZ 103721); Tungurahua, 6-12 mi SW. Baños, 2500-3000 m, 13.ii.1955, E.I. Schlinger and E.S. Ross (1♁, USNM ENT 01443834). PERÚ. Cuzco, Pumapaccha, -13,515778, -71,972709, 3426 m, Jared Shorma (iNaturalist catalogue number 37168068; https://www.inaturalist. org/observations/37168068)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 28-31, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Angel-Villarreal, S. L., Bogota-Angel, R. G. & Montoya, A. L. (2021) Comunidades de sirfidos (Diptera) asociadas a coberturas influenciadas por actividades antropicas en los cerros orientales de Bogota, Colombia. Caldasia, 43 (1), 161 - 171. https: // doi. org / 10.15446 / caldasia. v 43 n 1.82464","Fluke, C. L. (1957) A study of male genitalia of the Melanostomatini (Diptera: Syrphidae). Wisconsin Academy of Sciences, Arts and Letters, 46, 261 - 279.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Thompson, F. C. & Skevington, J. H. (2014) Afrotropical flower flies (Diptera: Syrphidae). A new genus and species de Kenya, with a review of the melanostomine group of genera. Zootaxa, 3847 (1), 97 - 114. https: // doi. org / 10.11646 / zootaxa. 3847.1.5","Montoya, A. L. (2016) 36 Family Syrphidae. In: Wolff, M., Nihei, S. S. & de Carvalho, C. J. B. (Eds.), Catalogue of Diptera of Colombia. Zootaxa, 4122 (1), 457 - 537. https: // doi. org / 10.11646 / zootaxa. 4122.1.39","Montoya, A. L., Ricarte, A. & Wolff, M. (2017) Two new species of Quichuana Knab (Diptera: Syrphidae) from the Paramo ecosystems in Colombia. Zootaxa, 4244 (3), 390 - 402. https: // doi. org / 10.11646 / zootaxa. 4244.3.7","Marin-Armijos, D., Quezada-Rios, N., Soto-Armijos, C. & Mengual, X. (2017) Checklist of the flower flies of Ecuador (Diptera, Syrphidae). ZooKeys, 691, 163 - 199. https: // doi. org / 10.3897 / ZooKeys. 691.13328","Montoya, A. L., Parra, J. L. & Wolff, M. (2021) Structure and diversity of hoverflies (Diptera: Syrphidae) in northwestern Colombian Paramos: towards the identification of bioindicator species in the Tropical Andes. Journal Insect Conservation, 25 (4), 1 - 20. https: // doi. org / 10.1007 / s 10841 - 021 - 00346 - 3"]}

Argentinomyia testaceipes Lynch-Arribálzaga, 1891 (Figs 1A–B, 81, 82 and 94A) Argentinomyia testaceipes Lynch-Arribálzaga, 1891: 200 (41). Type locality: Argentina. Prov. Buenos Aires, Chacabuco. HOLOTYPE Female MACN (non-examined). Refs.: Lynch-Arribálzaga, 1891:199 (1892c: 41): 268 (Fig. 4, habit) Argentinomyia testaceipes. Thompson et al. 1976: 57 Argentinomyia testaceipes. Thompson 1999: 338 (Synonyms and status notes) Argentinomyia testaceipes. Reemer & Ståhls 2014: 151 Rhysops lopesi Fluke, 1945: 6. Type locality: Brazil. S„o Paulo, S„o José dos Campos. HOLOTYPE Female AMNH (Examined). Refs.: Fluke, 1945: 26 (Fig. 5, head, lateral view, female) Rhysops lopesi. Thompson et al. 1976: 43 Type material. HOLOTYPE. Adult Male, ARGENTINA. Prov. Buenos Aires, Chacabuco, 173m (MACN). PARATYPE, BRAZIL. São Paulo, São José dos Campos, 53 m, 9.x.1933, H. S. Lopes (1♀, AMNH). Length (n= 3): Body, 7.3–8.5 mm; Wings, 7.1–7.5 mm. Diagnosis. Face with a low tubercle and with no prominent depressions dorsally; antennae elongate, the scape nearly four times as long as pedicel and about twice as long as basoflagellomere, arista thick and pilose; frontal prominence greatly produced; female frontal triangle wide, male dichoptic; sides of the face evenly white pubescent, except for a median vitta, which is broadest over tubercle, not punctuated; mesonotum highly polished, short golden pilose; without a pollinose vitta on anterior half; metafemur without long yellow pile, metatibia yellow; abdomen metallic, without orange maculae; surstylus with dorsal and ventral margins approximately of the same width in the whole length; apex of hypandrium (superior lobes) short, as long as wide. Redescription. MALE. Head (Fig. 81A): Face shining black, white pile except for some brownish hairs on dorsal half of frontal triangle. Face with a very low elongate tubercle, only slightly concave ventral antennae, with no distinct transverse depressions, although there are two very faint depressed lines immediately above the tubercle. Face entirely covered with white pollen except for a median vitta, which is broadest over the tubercle; oral tips somewhat produced ventrally, a shining area between eyes and oral tips. Frontal triangle highly polished with coppery reflections, extremely wide; ocelli on a slightly raised triangle, pile white ventral, more brownish in a wide band across vertex. Gena heavily coated with white pollen and white pile; side of the tubercle is a large, rectangular, flat, slightly rugose area, although the wrinkles show only on dorsal edge; brown, pile also appears on rather wide dorsal part of the occiput, rest of occiput with white pollen and pile. Antennae yellowish, inserted on a raised base, long, ratio 3.0:0.75:1.5, scape extremely long, black pile; pedicel dark brown, about 1/4 as long as scape; basoflagellomere black and about 1/2 as long as the scape; arista thick, about as long as the third segment, yellow at base and dark on apical 1/3, bare. Thorax (Figs 81B–C): Black, mesonotum highly polished, short golden pilose; without pollinose vittae on anterior half, pile yellow. Pleura shiny black, white pilose, white pollinose. Pleura shiny bluish cast, covered with white pollen and rather long white pile. Scutellum polished, but with two rugose depressions near the tip, pile entirely white; fringe with short golden pile; notopleura with only a slight indication of tubercle. Wing (Fig. 81A): Hyaline, stigma diluted yellow, microtrichose, except cell c bare basal 1/6, cells bm and r only slightly bare basally, cell cua pilose, calypter wholly yellowish-white; plumula white, halter yellowish-white, capitulum white. Legs (Fig. 81A): Yellow; pro- and mesofemora brown on basal 1/2; metafemur brown on basal 2/7 and apical 1/7; pro- and mesotibiae yellow, metatibia yellow with a broad black ring on apical 1/3, pile everywhere yellow, tarsomeres orange-brownish, yellow pilose, except a few black pile in the metatarsi. Abdomen (Fig. 81B): Everywhere polished with dark brown to black reflections, no evidence of yellow maculae anywhere, pile all white except for some darker hairs down the middle of the 4 th tergum; sterna black shining, yellow pilose; male genitalia: surstylus in lateral view (Fig. 82A) with dorsal and ventral margins approximately of the same width in the whole length, shorter than broad; hypandrium in ventral view (Fig. 82C) narrowed laterally towards the apex; aedeagal lobe in ventral view (Fig. 82C) circular, apex rounded. FEMALE (Figs 81C–F). Similar to male except for usual sexual dimorphism and differing in frontal prominence greatly produced and legs yellow. The female of A. testaceipes is similar to A. norrbomi sp. nov. but differs from it by the face with a low tubercle, with no distinct transverse grooves above the tubercle. Frontal prominence greatly produced. Legs yellow. Abdomen metallic, without orange maculae. In A. norrbomi sp. nov. the face with four or more distinct transverse grooves above the tubercle. Frontal prominence not greatly produced. Legs yellow, except metafemur black on apical 1/2. Abdomen with orange maculae. Taxonomic notes. Argentinomyia testaceipes is differentiated from A norrbomi. Sp. nov. by the following combination of characters: face without transverse grooves above the tubercle; frontal prominence greatly produced (Figs 81A, D); metafemur yellow (Figs 81A, C, D, F); abdomen metallic, without orange maculae (Figs 81B, E). In A. norrbomi sp. nov. the face with four or more distinct transverse grooves above the tubercle; frontal prominence not greatly produced (Figs 55A, D); legs yellow, except metafemur black on apical 1/2 (Figs 55A, C, D, F); abdomen with orange maculae (Figs 55B, E). Based on males, A. testaceipes differs from A. norrbomi sp. nov. in having the surstylus in lateral view (Fig. 82A) with dorsal and ventral margins approximately of the same width in the whole length [versus surstylus with dorsal margin slightly concave and ventral margin slightly convex in A. norrbomi sp. nov. (Fig. 56A)]; apex of hypandrium (superior lobes) in ventral view (Fig. 82C) short, as long as wide [apex of hypandrium elongated, about 3× longer than wide in A. norrbomi sp. nov. (Fig. 56C)] (see “diagnosis” under each species or key). Comments. The Holotype specimen of Rhysops lopesi is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue. 10021027 and non-type specimen in the USNM: http://syrphidae.myspecies.info/taxonomy/term/117. Geographical range. Argentinomyia testaceipes (n= 8) is distributed in the Serra Ambato in Argentina (Buenos Aires, Puerto Catamarca), Brazil (Paraná, S„o Paulo) and the Eastern slope of Cordillera Azul in Perú * (Previsto) (Fig. 94A). The species is present at low and middle altitudes (53–1274 m) in the following biogeographical domains and provinces: Paraná (53–1274 m): Araucaria Forest (895–1274 m) and Atlantic Forest (53–750 m); Chacoan (173–1226 m): Chaco (1226 m) and Pampa (173 m). Non-type material examined. ARGENTINA. Puerto Catamarca, Cuesta del Clavillo-IBP Program Frst, - 27,978 958, -65,74794, 1226 m, J.L. Neff (1♀, USNM ENT 01443766; http://syrphidae.myspecies.info/taxonomy/ term/117). BRAZIL, Paraná, Ponta Grossa, Pq Estadual de Vila Velha-IAPAR, -25,246336, -50,02104, 895 m, 7.v.2007, M.N. Morales (1♀, CEUFLA); Refugio de vida silvestre dos campos de Palmas, -26,533807, -51,599362, 1200 m, A. Couto (1♁, 1♀, DZUP); São Paulo, Barueri, -23.511369, -46.872942, 750 m, 12.viii.1957, K. Lenko (1♁, USNM ENT 01443767). Identified as Rhysops lopesi by Thompson (1971). PERÚ. Previsto, -9.0901948, - 75.737630, 800 m, 24.vi.1967, J. Schunke (1♀, BMNH)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 131-133, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Lynch-Arribalzaga, F. (1891 - 1892) Dipterologia Argentina, Syrphidae. Anales de la Sociedad Cientifica Argentina, 32, 80 - 99 + 118 - 131 + 194 - 202 + 247 - 256 + 307 - 314; 133 + 151 - 158 + 111 - 121 + 189 - 199 + 236 - 253; 134 + 133 - 146 + 173 - 192 + 242 - 280.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29."]}

Argentinomyia ivani Montoya sp. nov. Proposed standard English name: Ivan’s long-antennae flower fly. (Figs 35, 36 and 91A) Argentinomyia sp. COL-21 (11) Montoya 2019, unpublished Type material. HOLOTYPE. Adult Male, pinned, deposited at Colección Entomológica Universidad de Antioquia. Original label: “ COLOMBIA, Antioquia, Belmira, Páramo Santa Inés, Cabaña Cabildo Verde, El Morro, Alto de La Gallina, Páramo ” / “Malaise Canopy #2, 6,632 639, -75,645267, 3170–3200m ” / “ 4-14.ii.2017, A. L. Montoya Leg ”. “ HOLOTYPE / Argentinomyia ivani Montoya 2023 ”. Identified as Argentinomyia sp. 21 by Montoya 2019 (CEUA 95457, dissected). Length (n= 1): Body, 8.4–8.9 mm; Wings, 7.2–7.3 mm. Diagnosis. Face blue metallic; wing slightly brownish on anterior margin, cell r covered by a brownish translucent macula; 3 rd and 4 th terga with a pair of rectangular yellow maculae extending from the base to the apical 2/3 of the segment length, apically rounded and reaching the lateral margin; surstylus with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium laterally expanded towards the apex; aedeagal lobe circular, apex rounded to oval. Description. MALE. Head (Fig. 35A): Face blue metallic, whitish-grey pollinose and pilose. Facial tubercle at least with two slightly pronounced transversal grooves. Gena black, whitish pollinose and pilose. Frontal triangle black pollinose and pilose. Ocellar triangle black with black pile. Occiput white pollinose, black pilose on dorsal 2/3 and white pilose on ventral 1/3. Antenna dark reddish-brown, short, ratio 1.0:1.3:2.5, basoflagellomere as long as the pedicel and scape combined; basoflagellomere blackish dorsal and reddish-orange ventral; arista brown. Thorax (Figs 35B–C): Mesonotum, scutellum and pleura black-blue semiopaque, golden yellow pilose. Scutum with two whitish vittae on anterior half, reaching beyond the notopleural suture, golden pilose, except black pile in the postpronotum, notopleuron and postalar callus. Pleura bluish pollinose, yellowish pilose, except anterior anepisternum and katerpimeron with black pile. Scutellum shining, slightly rugose on apical half, pile golden in addition to long marginal black pile. Wing (Fig. 35C): Slightly brownish on anterior margin, stigma brownish, microtrichose, except cell bm bare on basal 1/2, r bare on basal 1/4 and covered by a brownish translucent macula, cell cua bare on anterior 1/3; calypter wholly brown; plumula yellowing; halter orange-brown, capitulum white. Legs (Fig. 35C): Brown to black, black pilose, 1 st and 2 nd metatarsi orange-yellow, others black. Abdomen (Fig. 35B): Black, 1 st- 2 nd terga black shining, with long lateral yellowish pile; 3 rd and 4 th terga with a pair of rectangular yellow maculae extending from the base to the apical 2/3 of the segment length, apically rounded and reaching the lateral margin; male genitalia: surstylus in lateral view (Fig. 36A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; hypandrium in ventral view (Fig. 36C) laterally expanded towards the apex; aedeagal lobe in ventral view (Fig. 36C) circular, apex rounded to oval. FEMALE. Unknown. Taxonomic notes. Argentinomyia ivani sp. nov. is morphologically most similar to A. belmira sp. nov., both having the 1 st and 2 nd metatarsi orange-yellow, strongly contrasting with the dark color of other segments. Argentinomyia ivani sp. nov. differs by the following combination of characters: Face blue metallic (Figs 36A, C, D-F); wing slightly brownish on anterior margin, cell r covered by a brownish translucent macula (Figs 36 B-C, E-F); 3 rd and 4 th terga with a pair of rectangular yellow maculae extending from the base to the apical 2/3 of the segment length, apically rounded and reaching the lateral margin (Figs 36B, E). In Argentinomyia belmira sp. nov. the face black and somewhat brassy (Figs 16A, C, D-F); wing slightly brownish (Figs 16 B-C, E-F); 3 rd and 4 th terga almost entirely yellow, with a pair of wider yellow maculae from the base to the apical margin of the segment, rounded on lateral edge and with a thin median black stripe and narrow posterior black margin; the 5 th tergum almost entirely yellow with only a small posteromedian black triangle, black pilose in the black areas and on medio-apical region of the 3 rd to the 5 th terga (Figs 16B, E). Based on males, A. ivani sp. nov. differs from A. belmira sp. nov. in having the hypandrium in ventral view (Fig. 36C) laterally expanded towards the apex [versus hypandrium narrowed laterally towards the apex in A. belmira sp. nov. (Fig. 17C)] (see “differential diagnosis” under each species or key). Etymology. The name Argentinomyia ivani sp. nov. is a patronym given in memory of the uncle of the first author, Iván Montoya (Pharmacist for more than 40 years), who after my beloved father Augusto Montoya passed away, provided me support, guidance and fruitful teachings that will remain forever in my mind and heart. For him, this beautiful species that flies high as his unforgettable smile and admirable way of being, with open hands to serve those who need it. Geographical range. Argentinomyia ivani sp. nov. (n= 1) is distributed in the Central Cordilleras in Colombia (Fig. 91A). The species is endemic to the Tropical Andes, distributed at the middle and high altitudes (3170–3200 m) in the Northern Andes domain and Magdalena province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 64-66, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Argentinomyia spinifemorata Montoya sp. nov. Proposed standard English name: Spiny femur long-antennae flower fly. (Figs 6E, 76 and 90B) Type material. HOLOTYPE. Adult female, pinned, deposited at Museo Entomológico Francisco Luis Gallego. Original label: “ COLOMBIA, Santa Elena” / “en Barbecho, 6.210509, -75.498419 ” / “ 2546 m, x.1965, R. Vélez ”. “ HOLOTYPE / Argentinomyia spinifemorata / Montoya 2023” (MEFLG 27270). Length (n= 1): Body, 5.8 mm; Wings, 5.3 mm. Diagnosis. Sides of face orange golden pollinose and pilose, tubercle with two transversal grooves, with a median dash of pollen extending from antennae almost to the tubercle; frontal triangle with a brown velvet cross band golden pilose; scutum aeneous grey, punctulate, with three brownish vittae on anterior half, a median slender brown vitta extending along the thorax, in addition to a lateral wider brownish vitta, widening towards the pronotum and narrow near the pre-scutellum; metafemur swollen, with ventral spines reaching the anterior 1/2; terga 3 rd to 5 th with basal transversal fascia. Description. FEMALE. Head (Fig. 76A): Face black, sides of face orange golden pollinose and pilose; tubercle low with at least two transversal grooves, with a median dash of pollen extending from antennae almost to the tubercle. Gena black, covered with yellow pollen and pile. Ocellar triangle opaque, with a brown velvet cross band golden pilose. Occiput golden pollinose and pilose. Antennae, yellow, short, ratio 1.0:1.2:2.6; basoflagellomere brown dorsal, longer, a little longer than scape and pedicel together; arista yellow at base and dark on apical third, slightly pilose. Thorax (Figs 76B–C): Aeneous grey, punctulate, yellow pilose and pollinose; with three brownish vittae on anterior half, a median slender brown vitta extending along the thorax, in addition to a lateral wider brownish vitta, widening towards the pronotum and narrow near the pre-scutellum. Pleura shining aeneous, grey pollinose, pile whitish and sparse. Scutellum punctulate, shining greenish-brassy to aeneous. Wing (Fig. 76C): Slightly yellowish, stigma orange-brownish, microtrichose, except cell c bare on basal 4/5, cells bm and r bare on basal 4/5, respectively, cell cua bare on anterior 1/2, basicosta orange, orange pilose, except for some black pile; calypter brown, border and fringe yellow; plumula yellow; halter yellow, capitulum yellow. Legs (Figs 6E, 76C, E): yellow, except the metafemur brown, only slightly yellow apically, metafemur swollen, with ventral spines reaching the anterior 1/2; basal three segments of pro-, meso- and metatarsi yellow, the apical ones black; pile yellow, coxae brown. Abdomen (Figs 76B, D): Abdomen narrowed toward the base, 2 nd tergum with a pair of lateral rounded maculae, covering the entire lateral margin but attenuated posteriorly; 3 rd to 5 th terga with basal transversal fascia, reaching apical 1/4, 1/4 and 1/3 respectively, pile yellow. MALE. Unknown. Etymology. The epithet “ spinifemorata ” is an adjective derived from the Latin “ spinei ”, spine, thorn, backbone (Brown 1956: 738) and the Latin “ femur ”, thigh: femur, femoral (Brown 1956: 468) and the Latin suffix - atus, provided with, having the nature of, pertaining to. It refers to the metafemur with ventral spines of this species. Remarks. Argentinomyia spinifemorata sp. nov. is unique within Melanostomini, characterized by having the metafemur swollen, with ventral spines reaching the anterior 1/2 (Figs 6E, 76C, E). Within Syrphinae, the metafemur of Salpingogaster Schiner, 1868 (tribe Syrphini) has a distinct row of ventral spinose bristles on apical 1/2, but the metafemur is not swollen. The presence of enlarged metafemur and ventral spines is common in groups of Eristalinae, including several variations. In Sterphus Philippi (Milesiini, Xylotina) the metafemur has a line of strong ventral spiny setae, meanwhile, in Spilomyia Meigen (Milesiini, Milesiina) the metafemur has a single small ventroapical spine (calcar, apicoventral extension). For its part, in Lepidomyia Loew, 1864 (Brachyopini, Brachyopina) the metafemur has strong ventral (small) spiny setae in the pro-, meso- and metafemur (Thompson 1999; Thompson et al. 2010). A marked sexual dimorphism in the shape of metafemur has also been recorded in Brachypalpus Macquart and Chalcosyrphus Curran (Eristalinae, Milesiini, Xylotina), where the female metafemur is less swollen, ventrally with rows of reduced spines, more developed in males than in females (Jeonga et al. 2017). These dimorphic features play a key role in mating, through which the male can hold the female during mating and/or elicit tactile responses which are critical to the cascade of cues and counter cue that lead to successful mating (Rotheray & Gilbert 2011). In this context, a sexual dimorphism could be expected for A. spinifemorata sp. nov., in such a case, the male conspecific may have a very swollen metafemur and strong ventral spines, compared to those evidenced in the female. However, we are aware that this appreciation will remain speculative until the male is discovered and described, considering also the fact that the type specimen was collected 76 years ago and despite exhaustive sampling efforts being carried out near the type locality, non-additional specimens have been found, which could be probably due to the rarity of this species (for a complete discussion of rarity see Montoya et al. 2022) or its potential local extinction due to the increase in deforestation in the Andean region of Colombia in the last years. However, our remarkable discovery opens new questions about how these characteristic posterior leg modifications originated and evolved within Syrphinae, particularly in Melanostomini. In this sense, more intensive sampling efforts and targeted fieldwork are needed to unravel the many issues surrounding this rare species. Taxonomic notes. Although A. spinifemorata sp. nov. is described from a single female (Figs 6E, 76), we evidenced that the species differs significantly from the A. berthae (Fig. 18D – F), A. fastigata (Fig. 30D – F), A. festiva (Fig. 32D – F) and A. lanei (Fig. 40D – F) by the unique metafemur swollen, with ventral spines reaching the anterior 1/2. The apparently restricted distribution raises the need for further expeditions to neighboring areas of the type locality in order to increase the knowledge about the species distribution and population dynamics as well as find male specimens to be described. Geographical range. Argentinomyia spinifemorata (n= 1) is known only from the Holotype, which was collected in a forest area of Medellín (Colombia) at an altitude of 2546 m (Fig. 90B). The species is endemic to the Northern Andes domain at middle altitudes (2546 m) in the Magdalena province.

Argentinomyia jamaicensis Montoya sp. nov. Proposed standard English name: Jamaica’s long-antennae flower fly. (Figs 39 and 95) Rhysops sp. 10 of Vockeroth 1973, unpublished Type material. HOLOTYPE. Adult female, pinned, deposited at Canadian National Collection of Insects, Arachnids and Nematodes, Canada. Original label: “ JAMAICA, 40000´, Hardward Gap, 18.068139, -76.636631, 1120m ” / “ 19.vii.1966, Howden & Becker” “ HOLOTYPE / Argentinomyia jamaicensis / Montoya 2023” (CNC DIPTERA 112271). Length (n= 1): Body, 4.5 mm; Wings, 3.6 mm. Diagnosis. Small species, spherical head, with a narrow band of pollen extending near to the anterior edge of the ocellar triangle; scape shorter than basoflagellomere; wings extensible hyaline; abdominal maculae metallic blue; legs dark brown, with only the apical 1/6 white-yellowish. Argentinomyia jamaicensis sp. nov. and A. praeusta share the condition of having the female frontal triangle with a medial white pollinose vitta (see “differential diagnosis and taxonomic notes” under each species or key). Description. FEMALE. Head (Fig. 39A): Face brilliantly shining, metallic blue-black, with a faint bluish luster, mid-vitta, margins are demarcated by a thin vitta of whitish pubescence. Frontal triangle bluish-black, darkgrey dusted on dorsal half and with a narrow thin line of grey-white pubescence running down the eyes margin which expands anteriorly into a small, lineal spot on dorsal part of the face and again on lower part of the face opposite the tubercle. Lower half of face sparsely white pubescent and four transversal grooves dorsal the tubercle; pile of the face and frontal triangle light brownish-yellow, of vertex darker; vertical triangle small. Antenna light brownish-yellow, short, ratio 1.1:1.3:1.5, scape and pedicel nearly equal in length; arista white at base and dark on apical third, with short pile. Thorax (Figs 39B–C): Very convex, shining metallic blue-black; with three brownish pollinose vittae, the median run halfway down the posterior part of mesonotum, there are suggestions of a fourth pair of such vittae lying some distance from the others on lateral part of mesonotum. There is a low but well-developed, very rounded bump in the middle of the anterior lateral corners of the thorax lying diagonally from the notopleuron. Pleura blue shining, grey-whitish pollinose, yellow whitish pilose. Scutellum broadly rounded, brilliantly metallic, with two or three subterminal, transverse, faint, ripple-like depressions; fringe long. Wing (Fig. 39C): Hyaline; stigma yellowish-white; calypter wholly yellowish-white; plumula white; halter white, capitulum white. Legs (Fig. 39C): Almost wholly dark brownish; femur and tibia with only the apical 1/6 white-yellowish. Abdomen (Fig. 39B): Bluish-black, metallic, with a complete black macula reaching 2/3 of tergum, anteriorly black and divided by a median-lineal blue macula, pile sparser, short, white; sterna bluish-black, yellow pilose. MALE. Unknown. Etymology. The specific epithet is derived from the country’s name where the species was found, “ Jamaica ” and the Latin suffix - ensis denoting place, locality, country, or belonging to, pertaining to (Brown 1956: 45, 303). The name should be treated as a noun in apposition. Remarks. Argentinomyia jamaicensis is exclusively known from the highlands of Blue Mountain in Jamaica. The apparently restricted distribution raises the need for further expeditions to neighboring areas of the type locality, in order to find and describe the male specimens. A. jamaicensis occur in sympatry with A. praeusta in the middle elevations of Jamaica. Distribution. Argentinomyia jamaicensis sp. nov. (n= 1) is only known from the type locality in the Blue Mountain in Jamaica (Hardward Gap) (Fig. 95). The species is endemic to the West Indies domain and distributed at middle elevations (1120 m) in the Jamaica province., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 69-70, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Argentinomyia scitula (Williston, 1888) (Fig. 73) Melanostoma scitulum Williston, 1888: 264. Type locality: BRAZIL. Mato Grosso, Chapada. SYNTYPE 2 Male AMNH (Examined). Refs.: Fluke, 1945: 7 (redesc.) 29 (Fig. 43, male abdomen, Fig. 54, wing) Rhysops scitulum. Williston, 1907: 2 Rhysops scitula. Curran, 1937: 2 (key) Rhysops scitula. Fluke, 1945: 7 Rhysops scitulus. Thompson et al. 1976: 44 Type material. HOLOTYPE. Adult male, BRAZIL. Mato Grosso, Chapada, 15°27’43.51”S, 55°44’54.20”O, 716 m (AMNH). Length. Body, 7.2–8.0mm; Wings, 5.7–6.4mm. Diagnosis. Argentinomyia scitula is similar to Argentinomyia longicornis differing by the following combination characters: Basoflagellomere as long as the scape and pedicel together; wings with two short, transverse maculae in front on median half, microtrichose; protarsus are wholly yellow; abdomen with a basal pair of elongate yellow maculae on 3 rd tergum reaching the apical 1/3, a similar smaller pair on 4 th tergum separated from the base of tergum and occupying 1/2 its length. Redescription. MALE. Head (Adapted from Williston 1888 and Fluke 1945). Sides of the face with a slender vitta and spot of pollen, not punctulate. Antennae brownish, orange ventrally, less elongated, the three segments of nearly equal length, basoflagellomere not twice as long as wide. Thorax: Black, mesonotum shining, with two faint whitish-pollinose vittae on anterior half, yellow pilose, the side brownish pollinose, pile brownish. Pleura shining black with a light coating of white to brownish pollen, mesopleura with brownish metallic reflections, yellow pilose. Scutellum shining, slightly rugose on apical 1/2, pile long and yellow, fringe long and yellow. Wing (Fig. 73): Hyaline, stigma light brownish, elongate, anterior cross-vein (r-m) clouding; calypter wholly dark brown; plumula light brown; halter yellowish-red, capitulum slightly brownish. Legs: Yellow, pro- and mesotibiae yellow; pro- and mesofemora dark brown, yellow at the anterior 1/4, femur narrowly yellow at the base; pro- and metafemora only pale brown on basal 1/2 but may be quite dark on fully matured specimens. Abdomen (Fig. 73): narrow, of nearly equal width throughout; with a basal pair of elongate yellow maculae on 3 rd tergum reaching the apical 1/3, a similar smaller pair on 4 th tergum separated from the base of tergum and occupying 1/2 its length. Remarks. Fluke (1945) was unable to recognize A. scitula (Williston) among the numerous specimens he studied from southern Brazil. Fluke reviewed a male from the AMNH (Fig. 73) pointing out that the opacity of the anterior transverse vein (r-m) is quite characteristic, but it is not entirely reliable as other species such as A. crenulata also have a slightly brown macula along this vein. These characteristics were verified and corroborated during the visit to the AMNH. The Holotype in the AMNH is a specimen collected in Chapada, Rio Grande do Sul, Brazil. Apparently, A. scitula could be a valid species, but as the Holotype is a headless, teneral and undissected male, there were nonadditional specimens found, although extensive material from Brazil was examined, therefore, the identity of the species remains doubtful and a more in-depth morphological analysis will be necessary to clear up its validity. Comments. The Holotype specimen of M. scitulum is currently in the AMNH. Photos are available on their website AMNH: https://sci-web-001.amnh.org/imulive/iz.html?#details=ecatalogue.10020982. Geographical range. Argentinomyia scitula is distributed in Brazil (Rio Grande do Sul)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 121-122, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Williston, S. W. (1888) Diptera Brasiliana, ab H. H. Smith Collecta. Transactions of the American Entomological Society, 15, 243 - 292.","Fluke, C. L. (1945) The Melanostomatini of the Neotropical Region (Diptera, Syrphidae). American Museum Novitates, 1272, 1 - 29.","Williston, S. W. (1907) Class I, Hexapoda. Order IV, Diptera. Dipterological notes Journal of the New York Entomological Society, 15, 1 - 2.","Curran, C. H. (1937) The Neotropical species of Melanostoma and allies (Syrphidae: Diptera). American Museum Novitates, 926, 1 - 4.","Thompson, F. C, Vockeroth, J. R. & Sedman, Y. S. (1976) Family Syrphidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Edanee, S \" o Paulo, pp. 1 - 195. https: // doi. org / 10.5962 / bhl. title. 49898"]}

Argentinomyia humboldti Montoya & Wolff sp. nov. Proposed standard English name: Humboldt’s long-antennae flower fly. (Figs 34 and 90D) Type material. HOLOTYPE. Adult female, pinned, deposited at Colección Entomológica Universidad de Antioquia. Original label: “ COLOMBIA, Antioquia, Belmira, Páramo Santa Inés” / “Cabaña Cabildo Verde, El Morro, Alto de La Gallina ” / “ 6,632639, -75,645267, 3170–3200m ” / “ 21-30.vi.2017, Net, C. Rodríguez; J. P. Carmona ”. “ HOLOTYPE / Argentinomyia humboldti / Montoya & Wolff 2020 ” (CEUA 98063). Identified as Argentinomyia sp. 16 by Montoya 2019. PARATYPES. COLOMBIA. Label with the same data as Holotype (1♀, CEUA 98064); Valle del Cauca, Jamundí, Corregimiento Meseta (Cabecera) La Selva, 3,167 630, -76,764164, 2200 m, Malaise trap, 27.v.2004, S. Sarria; M. Lasso, Collection sample·#5166 (2♀, IavH); ECUADOR. Napo Province, Yanayacu Biological Station, -0.598967, -77.891553, 2145 m, Net on flowers of Miconia sp., 21.viii.2022, X. Mengual (1♀, INABIO). Length (n= 3): Body, 7.4–7.9 mm; Wings, 6.8–7.3 mm. Diagnosis. Facial tubercle brown, face golden pilose and pollinose, facial pubescence not punctuate; calypter wholly yellowish; scutellum and pleuron partially yellow ground color, yellow-golden pilose and pollinose in these areas; wing bare basally; coxae brown; metabasitarsus yellow-orange, usually strongly contrasting with the dark color of the tibiae and the rest of the tarsi; abdomen with a pair of long, apically rounded rectangular to triangular maculae on 2 nd to 3 rd terga. Description. FEMALE. Head (Fig. 34A): Face black, golden-yellow pollinose and pilose. Tubercle low, bare, area above the tubercle yellow pollinose. Frontal triangle yellow-silver pollinose, golden pilose. Frontal triangle brown-velvety pollinose, golden pilose. Gena black. Occiput black pilose dorsally, yellow hoary ventrally. Antenna orange, short, ratio 1.0:1.5:2.4, basoflagellomere as long as the scape and pedicel together, yellowish, darker toward the end and dorsal on basoflagellomere, pile black, apically rounded; arista brown, bare. Thorax (Figs 34B–C): Grey to yellow-orange, mesonotum yellow semishining, grey to golden pollinose, yellow-golden pilose. Pleura yellow pollinose, yellow pilose, postpronotum, the middle region of anepimeron, dorsal surface of katepisternum, katatergum, anatergum and postalar callus yellow golden pilose and pollinose. Scutellum grey shining, yellowgolden pollinose, with long yellow hairs and short yellow pile intermixed; fringe short and yellow. Wing (Fig. 34C): Slightly yellowish, stigma brownish, mostly bare, costal cell bare on basal 1/2, cells dm and r bare, cell cua bare on anterior 3/4, apical border slightly brownish; calypter wholly brown; plumula yellow; halter yellow, capitulum yellow. Legs (Fig. 34C): Slightly thickened, yellow, pro- and mesolegs yellow, metafemur brown, except yellow on apical 1/2, metatibiae and tarsi brown, brown pilose on brown areas; coxae brown. Abdomen (Fig. 34B): Black, yellow maculae, 1 st tergum laterally yellow, extending the maculae to the lateral area of the 2 nd tergum in the whole length, 3 rd tergum with a pair of rectangular maculae, apically rounded and extending only the apical 1/2 of segment length, in addition to slightly lateral maculae, 4 th tergum with short triangular maculae, extending only the apical 1/2 of segment length, 5 th with basolateral small maculae, in addition to slightly lateral maculae. MALE. Unknown. Etymology. The name of this beautiful species, Argentinomyia humboldti sp. nov. is given in honor of Alexander von Humboldt on the 250 th anniversary of his birth. Humboldt was an enthusiastic Prussian naturalist who collected many elements of the American fauna and flora in South and Central America, including Ecuador, Colombia and Venezuela. His expeditions through South America provided biological material for several dipterologists, contributing to the knowledge of many Neotropical organisms in the early eighteenth century. The specific epithet is to be treated as a noun in the genitive case. Biology. Label data suggest that A. humboldti sp. nov. visits flowers of Miconia sp. (Melastomataceae). Geographical range. Argentinomyia humboldti sp. nov. (n= 4) is distributed in Colombia (Antioquia and Valle del Cauca) and Ecuador (Napo) (Fig. 90D). The species is endemic to the Northern Andes domain, distributed throughout middle and high altitudes (2200–3200 m) in the provinces: Cauca (2200 m), Magdalena (3170–3200 m) and North Andean Páramo (2145 m)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 63-64, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993, {"references":["Montoya, A. L. & Wolff, M. (2020) Description of six new large species of Argentinomyia Lynch-Arribalzaga, 1891 and redescription of Talahua fervida (Fluke, 1945) (Diptera, Syrphidae, Syrphinae). ZooKeys, 929, 19 - 51. https: // doi. org / 10.3897 / zookeys. 929.37666"]}

Argentinomyia norrbomi Montoya sp. nov. Proposed standard English name: Norrbom’s long-antennae flower fly. (Figs 4J, K, 8B, 9A, 55, 56 and 93D) Type material. HOLOTYPE. Adult male, pinned, deposited at National Museum de Natural History. Original label: “ BRAZIL, Paraná ” / “Palmas, Linha Alegría, Fazenda Cerro Chato, -26.50252, -51.67033, 1224 m, Malaise en la cima de campo; 3-4.iii.2015, M. Savaris; Allen L. Norrbom ” “ HOLOTYPE / Argentinomyia norrbomi / Montoya 2023” (USNM ENT 01443832). PARATYPE. BRAZIL. Label with the same data as Holotype (7♁, 5♀, USNM ENT 01443831, 6 ♁, 1♀, DZUP). Length (n= 3): Body, 5.8–6.2 mm; Wings, 4.3–4.7 mm. Diagnosis. Face with four or more distinct transverse grooves above the tubercle; frontal prominence is not greatly produced; male eyes widely dichoptic; legs yellow, except metafemur black on apical 1/2; abdomen with a pair of basal maculae on 2 nd and 3 rd terga extended only the apical 1/2 and 1/4, respectively; surstylus with dorsal margin slightly concave and ventral margin slightly convex; apex of hypandrium (superior lobes) in ventral view elongated, about 3× longer than wide. Description. MALE. Head (Figs 4J, K, 9A, 55A): Face black, entirely covered with white pollen and pile except for a median vitta; tubercle low and with five very faint depressed lines immediately above tubercle. Frontal triangle polished with coppery reflections, extremely wide and with white pile. Frontal prominence is not greatly produced; eyes widely dichoptic; occiput white pilose and pollinose. Antenna orange, long, ratio 3.0:1.0:1.5, scape extremely long, about as long as the pedicel and basoflagellomere; arista yellow at base and dark on apical 1/3, bare. Thorax (Figs 55B–C): Black, mesonotum highly polished, short white pilose; with a coppery pollinose vitta on anterior half, pile white. Pleura shiny black, yellow pilose and pollinose. Scutellum black, polished, with two rugose depressions near the tip, pile entirely yellow. Wing (Fig. 55C): Hyaline, slightly yellowish, stigma yellow, microtrichose, except cell c bare basal 2/3, cells bm and r only slightly bare basally, cell cua bare; calypter wholly yellow; plumula yellow; halter yellow, capitulum yellow. Legs (Fig. 55C): yellow, except metafemur black on apical 1/2; only metafemur brownish on apical 1/2; tarsomeres orange brownish, orange pilose. Abdomen (Fig. 55B): Black, parallel-sided, 2 nd tergum black, 3 rd tergum with a basal pair of elongate yellow maculae reaching the apical 5/6; 4 th tergum with a pair of basal maculae occupying 1/3 the tergum length; pile all yellow; male genitalia: surstylus in lateral view (Fig. 56A) with dorsal margin slightly concave and ventral margin slightly convex, shorter than broad; apex of hypandrium (superior lobes) in ventral view (Fig. 56C) elongated, about 3× longer than wide; aedeagal lobe in ventral view (Fig. 56C) circular, apex rounded. FEMALE (Figs 8B, 9A, 55D–F). Very similar to male but the frontal triangle is wider and greyish pollinose, abdomen with a pair of basal maculae on 2 nd and 3 rd terga extended only the apical 1/2 and 1/4, respectively. Female of A. norrbomi sp. nov. is similar to A. testaceipes but differs from it by the face with four or more distinct transverse grooves above the tubercle. Frontal prominence not greatly produced. Legs yellow, except metafemur black on apical 1/2. Abdomen with orange maculae. In A. testaceipes the face with a low tubercle, with no distinct transverse grooves above the tubercle. Frontal prominence greatly produced. Legs yellow. Abdomen metallic, without orange maculae. Taxonomic notes. Argentinomyia norrbomi sp. nov. is differentiated from A. testaceipes by the following combination of characters: Face with four or more distinct transverse grooves above the tubercle; frontal prominence not greatly produced (Figs 55A, D); legs yellow, except metafemur black on apical 1/2 (Figs 55A, C, D, F); abdomen with orange maculae (Figs 55B, E). In A. testaceipes face without transverse grooves above the tubercle; frontal prominence greatly produced (Figs 81A, D); metafemur yellow (Figs 81A, C, D, F); abdomen metallic, without orange maculae (Figs 81B, E). Based on males, A. norrbomi sp. nov. differs from A. testaceipes in having the surstylus in lateral view (Fig. 56A) with dorsal margin slightly concave and ventral margin slightly convex [versus surstylus with dorsal and ventral margins approximately of the same width in the whole length in A. testaceipes (Fig. 82A)]; apex of hypandrium (superior lobes) in ventral view (Fig. 56C) elongated, about 3× longer than wide [apex of hypandrium short, as long as wide in A. testaceipes (Fig. 82C)] (see “diagnosis” under each species or key). Etymology. The specific epithet of this species is a Latinized noun in the genitive case of the last name ‘Norrbom’ in honor of Allen L. Norrbom, a research entomologist at the USDA-ARS-Systematic Entomology Laboratory, who collected the type series of this species in Palmas, Linha Alegría, Fazenda Cerro Chato. The species is dedicated to him in gratitude for his hospitality and support during my internship (ALM) at the Smithsonian Institution in 2018. Remarks. I (ALM) received the specimens of A. norrbomi sp. nov. from Allen L. Norrbom and the first impression was that they corresponded to A. testaceipes. However, when studied more thoroughly noticed the presence of yellow legs, except for a metafemur black on apical 1/2 and abdomen with elongated yellow maculae reaching the apical 5/6. The characteristic that most caught the attention was the male genitalia with elongated superior lobes. In consequence, when comparing it with specimens of A. testaceipes, it was concluded that this very special and marvelous specimen turned out to be a previously undescribed species. Geographical range. Argentinomyia norrbomi sp. nov. (n= 28) is exclusively distributed in Serra da Fartura in Brazil (Paraná) (Fig. 93D). The species is endemic to Paraná domain (1224 m), occurring in sympatry with A. plaumanni sp. nov. throughout the middle elevations of southeastern Brazil, particularly in the Atlantic Forest where they appear to be widely distributed, but rare and less abundant (Paraná Forest province)., Published as part of Montoya, Augusto L. & Wolff, Marta, 2023, Taxonomic revision of the Neotropical genus Argentinomyia Lynch-Arribálzaga, 1891 (Diptera: Syrphidae), with description of 16 new species, pp. 1-157 in Zootaxa 5234 (1) on pages 96-98, DOI: 10.11646/zootaxa.5234.1.1, http://zenodo.org/record/7609993

Ptecticus benecki sp. nov. Figs. 14, 16, 18, 20���21, 24, 26, 28���29, 32���34, 38���39, 42, 44, 46, 48���49, 55���57, 58���61, 66, 68, 70, 72. Diagnosis. Yellow species, except mesonotum and pleura light brown (Fig. 24). Antenna yellow in male and female, (Figs. 14, 16). R 2+3 originating at a distance approximately the longitude of r-m (Figs. 28, 29). Abdomen with segments 1���3 yellow with a basal brown band and 4���7 dark brown (Figs. 38, 39). Terminalia yellow with cerci dark brown (Figs. 39, 48). Material examinated. HOLOTYPE: ♂, body length 10.0 mm, wing 10.0 mm. Colombia, Antioquia, San Vicente Ferrer, Vda. Chaparral, Finca La Mosca, organal. 6.266579, -74.360599, 2150m, Guano Carollia perspicillata 03.iii.2019. C. Bota; J. Torres Toro; M. Wolff. Manual. Emergence adults 12.iv.2019. CEUA-88649 (imago + puparium). PARATYPES: 52 males: body length 8.5���11.5mm, wing 8.0���10.0 mm; 57 females: body length 7.0���11.5 mm, wing 6.5���11.0 mm; puparia: length 9.0���12.0mm. Same locality. Catalogue numbers: CEUA-62578 (♀), 62579 (♀), 88662 (♀), 88664 (2♂, 2♀), 88665 (5♂, 4♀), 88666 (3♂, 2♀), 88667 (3♂, 3♀), 88668 (1♂, 1♀), 88669 (1♂, 2♀), 88670 (2♂), 88671 (♀ dissected), 118122 (3♂, 2♀), 118131 (♂), 118132 (♀ dissected), 118133 (♀), 118134 (♀ dissected), 118135 (♂), 118136 (♀), 118137 (♀), 118138 (♀), 118139 (♂), 118140 (♂), 118141 (♂), 118142 (2♂, 1♀), 118143 (♀), 118144 (1♂, 3♀), 118145 (2♂), 118147 (♂), 118148 (3♀), 118149 (♂), 118152 (♂), 118153 (♀), 118154 (♀), 118155 (♀), 118156 (♀), 118157 (♂), 118158 (♀), 118159 (♀), 118160 (♂), 118161 (1♂, 2♀), 118162 (♂), 118163 (♂), 118164 (2♂, 1♀), 118165 (♀), 118166 (♂), 118167 (1♂, 1♀), 118168 (♀) 118169 (♂), 118170 (♂), 118171 (2♀), 118172 (♂), 118173 (3♀), 118174 (♂), 118175 (1♂, 1♀), 118176 (♂), 118177 (1♂, 1♀), 118178 (♂), 118179 (♂), 118180 (1♂, 2♀), 118186 (♀), 118192 (♀). Description. Male. Body length 10.0 mm, wing 10.0 mm. Head (Fig. 14, 18) almost globular, hairs yellow, only a black tuft behind ocellar tubercle mainly erect, longest hairs slightly longer than length of scape. Frontal callus yellow whitish, markedly prominent, oval, wider than long, almost as long as scape in profile, drop-shaped in frontal view. Frons tapered towards frontal callus, very narrow immediately above it, yellowish white. Ocellar tubercle blackish. Vertex yellowish (Fig. 18). Occiput black. Antenna yellowish, scape slightly longer than broad at distal margin, pedicel with usual subtriangular projection on inner surface not sharpened, flagellar complex round (Figs. 20, 21), subapical arista like terminal flagellomere brownish. Face yellowish white, proboscis yellowish brown. Thorax (Figs. 24, 26) light brown, sometimes appearing grayish, hairs conspicuous golden and black, scutum hairs generally shorter than frontal hairs; propleuron yellow, with golden hairs; notopleuron yellowish with golden hairs; katatergite yellowish-white with golden hairs; mediotergite yellow with golden hairs; anepisternum yellowish-white extensively bare; katepisternum yellowish-white with golden hairs; anepimeron yellow with golden hairs; laterotergite yellowish-white with a small dark and sclerotized tubercle; scutellum grayish-brown triangular. Wings (Fig. 28) hyaline, shining with the incidence of light, with bare spaces in alula cup and medial cells, pterostigma yellow. Cell r 2+3 low and long, almost 6 times longer than tall, vein R 2+3 arising after and almost at 3/4 of r-m length, almost parallel to R 1, longer than R 4. Crossvein r-m straight, longer than distal margin of discal cell, discal cell with upper posterior margin almost straight, Vein M 3 slightly parallel to M 2 with apical fourth reduced. Wing fold in cup cell pigmented. Halter stem yellowish basally to brown apically, halter knob yellowish brown. Posttegula with a bunch of golden hairs. Legs (Figs. 32���34). Anterior legs hairs mostly yellow. Anterior femur, tibia and basitarsus yellow, second to fifth tarsi dark brown. Mid and hind legs hairs mostly yellow. Mid and hind femur, tibia, and basitarsus yellow, second tarsus mostly yellow, third to fifth brown, darker in hind last two tarsi. Abdomen (Figs. 38, 39) petiolated, yellowish, Tergites 1 to 3 yellow with transverse yellowish brown bands mainly on the basal half, wider along lateral margin of tergites 1 and 2, the remain visible tergites completely shine brown, the venter yellow in segments 1, 2 and apical half of 3, otherwise brown, Terminalia (Fig. 42, 44, 46) yellowish brown. Epandrium arched, slightly longer than wide, proctiger triangular, cerci yellowish brown inserted in the middle of proctiger. Genital capsule rounded, gonostylus as figure 48, posterior sclerotized portion of phallus hook-like as figure 49. Female. Body length 7.0���11.5 mm, wing 6.5���11.0 mm (fig. 29). Differs from male as follows: Upper frons almost parallel, yellow with two narrow lateral triangular dark bands ending before ocellar triangle (Fig. 16). Terminalia (Fig. 55), genital fork with the apex of anterior margin triangular subacute (figs. 56, 57). Puparium. (Figs. 58���61, 66, 68, 70, 72). Length Holotype: 10.5 mm; Paratypes: 9.0���12.0mm, including head. Head subtriangular, as long as one tenth of entire puparium, longer than wide. Labrum subrectangular rounded at apex. Antenna short, black, rising dorsolateral in anterior part of head at the middle height of labrum. Eyes prominent, arising in middle of head capsule, lateral setae. Molar area (mo) scarcely visible, ventrocraneal sulcus slightly evident. Chaetotaxy: 2 pairs of labral setae (Lb), 2 pairs of clypeofrontal setae (Cf), 2 pairs of dorsolateral setae (Dl), 1 pair of lateral setae (L), 2 pairs of ventrolateral setae (Vl), usually with 3 pairs of ventral setae (V), (V 3) could be absent or difficult to see and 1 pair of ventroapical (Va) setae in the labral-molar area. Thorax. Prothorax subrectangular, with anterior spiracles prominent; mesothorax rectangular with curved side edges; metathorax rectangular with curved side edges. Chaetotaxy: Prothorax with 2 pairs of anterodorsal setae (Ad), 2 pairs of dorsal setae (D), 1 pair of dorsolateral (Dl), 1 pair of ventrolateral (Vl), and 2 pairs of ventral setae (V), one simple (V 1) and one tripartite (V 2). Mesothorax with 3 pairs of anterodorsal setae (Ad), 1 pair of dorsolateral (Dl), 1 pair of ventroleteral setae (Vl), and 2 pairs of ventral setae (V), one simple (V 1) and one tripartite (V 2). Metathorax with 3 pairs of dorsal setae (D), 1 pair of lateral setae (L), 1 pair of ventrolateral setae (Vl), and 2 pairs of ventral setae (V), one simple (V 1) and one bipartite (V 2). Abdomen. Chaetotaxy in segments 1 st to 7 th 3: pairs of dorsal setae (D), 1st segment with 2 pairs of dorsolateral setae (Dl), remain segments with 1 pair of dorsolateral setae (Dl), 1 pair of lateral setae (L) only in segments 6 th and 7 th, 1 pair of ventrolateral setae (Vl) and 3 pairs of ventral setae (V). Segments 2 nd to 5th with tiny lateral spiracular horn in the middle. Segments 1 to 5 present an elevated wart at each side located at the level of the dorsolateral seta (D1), with a diameter usually bigger than the structural units forming the surrounding of cubital vein + posterior branch of cubital vein; CuP=posterior branch of cubital vein; cup=posterior cubital cell; d=discal cell; h=humeral crossvein; M=medial vein, or media; M 1 =first branch of media; m 1 =first medial cell; M 1+2 =fused first and second branch of media; M 2 =second branch of media; m 2 =second medial cell; M 3 =third branch of media; m 3 =third medial cell; M 3+4 =fused third and fourth branch of media; M 4 =fourth branch of media; m 4 =fourth medial cell; m���cu=medial���cubital crossvein; m���m=medial crossvein; R 1 =anterior branch of radius; r 1 =first radial cell; R 2+3 =second branch of radius; r 2+3 =second + third radial cell; r 4 =fourth radial cell; R 4 =upper branch of third branch of radius; r 5 =fifth radial cell; R 5 =lower branch of third branch of radius; r���m=radial���medial crossvein; Rs=radial sector; sc=subcostal cell; Sc=subcostal vein. Scale bar 1.0 mm. cuticle (Fig. 72) in dorsal view. Segment 6 th with linear external patch. Segment 8 th semicircular in shape, exhibits posterior spiracle (Ps) dorsally just before apex with anal slit (asl) much shorter than segment 8 th, surrounded by two perianal grooves; chaetotaxy: 1 pair of dorsal setae (D), 1 pair of lateral setae (L), 4 pairs of ventral setae (V), 1 pair of subapical setae (Sa), and 2 pairs of apical setae (Ap). Etymology: The epithet of this species is named after Dr. Mark Benecke, amazing forensic scientist and friend. Comments. Ptecticus benecki appears to be a species close to the Mexican species Ptecticus comstocki, based on couplet 30, page 24 of James & McFadden (1982), both share the yellow coloration of the hind tibia, sometimes appearing slightly very light brown, but never dark or black (at least in the male). In both species, the tibiae have mainly yellow pilosity with some black hairs that may be inconspicuous and do not surround tibiae in its basal half. P. benecki can be separated from P. comstocki by i) the abdominal pattern, in P. comstocki yellow abdomen with brown bands present in tergites 2 to 5, while in P. benecki the abdomen is also yellow, but the dark brown bands are present in tergites 1 to 3, the 4 and remaining completely dark (Fig. 38, 39); and ii) the genitalia, in P. comstocki the proctiger are short and rounded (illustrated in figure 65 page 50 in James & McFadden 1982), while in P. benecki proctiger is more prominent with subtriangular appearance (Fig. 44, 46)., Published as part of Torres-Toro, Juliana, Pujol-Luz, Jos�� Roberto & Wolff, Marta, 2022, Two new species of Ptecticus Loew, 1855 (Diptera: Stratiomyidae), from bat guano in a Colombian cave, pp. 61-88 in Zootaxa 5116 (1) on pages 67-73, DOI: 10.11646/zootaxa.5116.1.3, http://zenodo.org/record/6364301, {"references":["James, M. T. & McFadden, M. W. (1982) The Sarginae (Diptera, Stratiomyidae) of Middle America. Melanderia, 40, vii - x + 1 - 50."]}

Ptecticus pseudospatulatus sp. nov. Figs. 15, 17, 19, 22���23, 25, 27, 30���31, 35���37, 40���41, 43, 45, 47, 50���51, 52���54, 62���65, 67, 69, 71, 73, 74���79, 80���83. Diagnosis. Dark species except abdomen (Figs. 25, 27). Mesonotum dark brown and pleura dark with anepimeron bright yellow. Antenna light brown (Fig. 17). Vein R 2+3 originating very close to r-m especially in male (Fig. 30). Anterior legs with all tarsi appearing dark, middle and hind with only tarsi 4, 5 appearing dark (Fig. 35���37). Abdomen with tergites 1���4 yellow, 2���3 stripe marked with dark brown, and 5���7 completely dark brown (Figs. 40, 41). Terminalia completely dark with cerci very long (Figs. 41, 45, 47). Material examinated. HOLOTYPE: ♂, Colombia, Antioquia San Vicente Ferrer, Vda. Chaparral, Finca. La Mosca, organal. 6.266579, -74.360599, 2150m, Guano Carollia perspicillata 15.ii.2020. J. Torres-Toro; C. HenaoSep��lveda; V. Ossa. Manual. Emergence adults 22.iii.2020. CEUA-88654. (imago + puparium). PARATYPES. 20 males: body length 9.5���13.5 mm, wing 7.0���11.0 mm; 21 females: 9.5���13.0 mm, wing 8.5���12.0mm; puparia: length 9.5���14.0 mm. Same locality. Catalogue numbers: CEUA-88652 (♀), 88653 (♂), 88655 (♂), 88656 (♀), 88657 (♀ dissected), 88658 (♀), 88659 (♂), 88660 (♂), 88661 (♂), 88672 (♀ dissected), 88674 (♀), 118121 (2♂, 2♀), 118123 (♂, 2♀), 118124 (2♂, 3♀), 118125 (♀), 118126 (♂), 118127 (♀), 118128 (♂), 118130 (♀), 118146 (♂), 118150 (♂), 118151 (♀ dissected), 118181 (♀), 118182 (♀), 118183 (♀), 118184 (♀), 118185 (♂), 118187 (♀), 118188 (♂), 118189 (♂), 118190 (♂), 118191 (♀), 118193 (♂), 118194 (♂), 118195 (♂). Other examined material: one specimen atypically small, body length 8.0 mm, wing 7.0 mm, catalogue number CEUA-118129 (♀). Description. Male. Body length 12.0 mm, wing 12.5 mm. Head (Figs. 15, 19) almost globular. Head with hairs black at the vertex, upper frons and scattered at frontal callus, golden hairs at upper half of the occiput, scattered hairs black on frons, otherwise yellow, mainly erect, longest hairs reaching length of scape. Frontal callus prominent, oval, wider than long, almost as long as scape in profile, triangle drop-like in frontal view, yellow. Frons tapered towards frontal callus, slightly narrow immediately above it, yellowish-light brown. Ocellar tubercle black with a posterior dark brown stripe towards vertex. Vertex with a dark brown transverse stripe from ocellar tubercle to occiput. Occiput black. Antenna brownish, scape slightly longer than broad at distal margin, pedicel with usual subtriangular projection on inner surface slightly rounded, flagellar complex nearly round from outer view (Figs. 22, 23), subapical arista like terminal flagellomere black. Face yellowish brown, proboscis yellowish brown. Thorax (Figs. 25, 27) brown to dark brown, thoracic hairs conspicuous, mostly black, scutum hairs generally shorter than frontal hairs; propleuron brown with golden hairs; notopleuron brownish with golden hairs katatergite brown with golden hairs; mediotergite yellowish brown with golden hairs; anepisternum blackish extensively bare, katepisternum blackish with whitish hairs, anepimeron bright yellow with golden hairs, scutellum yellowish brown, triangular. Wing (Fig. 30) almost hyaline shining, alula completely bare, pterostigma yellow. Cell r 2+3 low and long, almost 8 times longer than tall, vein R 2+3 arising after and approximately at 1/2 of r-m length, noticeably fused to R 1, so, shorter than R 4. Crossvein r-m almost straight, longer than distal margin of discal cell, discal cell with upper posterior margin almost straight, vein M 3 slightly parallel to M 2 with apical fourth reduced. Wing fold in cup cell pigmented. Halter stem yellowish brown basally to brown apically, halter knob darker brown dorsally. Posttegula with a bunch of black hairs. Legs (Figs. 35���37). Hairs mostly yellow. All legs yellowish to light brown, darker in the apical half of femora and tibia; anterior legs with all tarsi gradually dark to the apex, middle and hind legs with the apical tarsomeres appearing darker because the black hairs in all the dorsal extension. Abdomen (Fig. 40, 41) markedly petiolated, with tergites 1���4 yellow, 2���3 marked with dark brown, tergites and sternites 5���7 completely dark brown-black. Terminalia (Fig. 50) completely dark brown. Epandrium straight, proctiger rounded (Figs. 43, 45, 47), cerci strongly sclerotized. Genital capsule rounded and gonostylus as in figure 45, posterior sclerotized portion of phallus hook-like shaped as in figure 51. Female. Body length 9.5���13.0 mm, wing 8.5���12.0mm (Fig. 31). Differs from male in the following: upper frons almost parallel, yellowish brown with two lateral triangular dark bands reaching the sides of ocellar triangle (Fig. 17). Terminalia (Fig. 52), genital fork with the apex of anterior margin rounded (Figs. 53, 54). Puparium. (Figs. 62���65, 67, 69, 71, 73). Length Holotype: 11.0 mm; Paratypes: 9.5���14.0 mm comprising head. Nearly identical of P. benecki sp. nov. except the following few characters: Head. as long as one eleventh of entire puparium. ventrocraneal sulcus well developed. Chaetotaxy: 3 pairs of lateral setae (L) 3 pairs of ventral setae (V). Thorax. chaetotaxy: Prothorax, 1 pair of ventral setae (V) tetrapartite. (V 2). Abdomen. A tiny, sclerotized tubercle resembling a scar with size equivalent to the structural units of the cuticle (Fig. 73) at the level of the dorsolateral seta (D1). Additionally, the general coloration in P. spatulatus is darker and with the longitudinal dark sections wider than in P. benecki sp. nov. Mature larvae (final instar, wander). (Figs. 74���79). The last instar of Ptecticus pseudospatulatus share with the puparia almost all characters as the design of the integument, size, location of setae and the presence of a dark cuticular wart of similar size to the surrounding elements at the level of the lateral bristles, as well as the formation of the suture in the form of a ���Y��� where hatching occurs. Both differ behaviorally in that the last larval instar presents active movements on itself, sideways and is ���fast��� when moving forward to seek shelter; morphologically, the mature larva is slightly softer and, of course, presents abdominal spiracles, general appearance is bright in colors (Figs. 74���79). The pupa, on the other hand, is completely rigid, evidently darker and, even though tiny, with the presence of pupal respiratory horns. As Schremmer (1986) specifies, for Sarginae and Hermetiinae ���the penultimate instar in relation to the last or ���pre-pupa���, may look like larvae of different species���, unlike other mainly aquatic subfamilies. The mature larva of Ptecticus pseudospatulatus differs strongly from an earlier instar that we fixed (two specimens length 11.5 mm and 12.0 mm; possibly from the penultimate instar five), In our specimens���in that earlier stage���there is an absence of integument design, general setation is dense and conspicuous without differentiation of dorsal, lateral or ventral setae on thorax and abdomen, the appearance of the body is remarkably soft to touch, the cephalic capsule is dark, elongated and the eyes are compact instead of voluptuous (Figs. 80���83). Eight larvae, six mature: (CEUA-88663, CEUA-88650). Etymology. The epithet of this species refers to similarity with the species P. spatulatus McFadden. Comments. Ptecticus pseudospatulatus have genitalia, very similar to the Mexican species Ptecticus spatulatus according to McFadden (fig. 62 in James & McFadden 1982). As mentioned above, the external morphology of P. spatulatus was examined from holotype pictures [CNC1173628] (CNC 2021) and differs in the following characters: i) P. spatulatus ��� superior front hairs are yellow (Fig. 11), whereas in P. pseudospatulatus are black (Fig. 15); ii) the abdominal pattern of P. spatulatus is characterized by being obscure in almost the entire extension of the tergites from the second (Figs. 9, 10), whereas in P. pseudospatulatus from tergites 2 to 4 are limited to the middle region or the anterior central margin (Figs. 40, 41); iii) in both species, the R 2+3 vein emerges equally posterior to r-m, but in Ptecticus pseudospatulatus much closer to r-m (Fig. 30) than in P. spatulatus (Fig. 8), also vein r-m in P. pseudospatulatus is almost straight and in P. spatulatus obliquous; iv) in the posterior legs of P. spatulatus the tibias are dark in 2/3 of the extension and all the tarsomeres are yellow (Fig. 9), whereas in P. pseudospatulatus the tibias are lighter (possibly 1/2) and the tarsomeres 4 and 5 are darker at least dorsally (Fig. 37). Finally, the Table 2, shows a comparison between the known Colombian species., Published as part of Torres-Toro, Juliana, Pujol-Luz, Jos�� Roberto & Wolff, Marta, 2022, Two new species of Ptecticus Loew, 1855 (Diptera: Stratiomyidae), from bat guano in a Colombian cave, pp. 61-88 in Zootaxa 5116 (1) on pages 74-85, DOI: 10.11646/zootaxa.5116.1.3, http://zenodo.org/record/6364301, {"references":["Schremmer, F. (1986) Die polymetabole Larval-Entwicklung der Waffenfliegenart Hermetia illucens. Ein Beitrag zur Metamorphosse der Stratiomyidae. Annalen des Naturhistorischen Museums in Wien, 88 - 89, 405 - 429.","James, M. T. & McFadden, M. W. (1982) The Sarginae (Diptera, Stratiomyidae) of Middle America. Melanderia, 40, vii - x + 1 - 50.","Schiner, J. R. (1868) Diptera. In: Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859, unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil 2. 1 (B). Kaiserlich-koniglichen Hof- und Staatsdruckeri in comission bei Karl Gerold's Sohn, Wien, VI + 388 pp.","Enderlein, G. (1914) Dipterologische Studien. IX. Zur Kenntnis der Stratiomyiiden mit 3 astiger Media und ihre Gruppierung. A. Formen, bei denen der 1. Cubitalast mit der Discoidalzelle durch Querader verbunden ist oder sie nur in einem Punkte beruhrt (Subfamilien: Geo-sarginae, Analcocerinae, Stratiomyiinae). Zoologischer Anzeiger, 43 (13), 577 - 615.","James, M. T. (1938) New and little-known Neotropical Stratiomyidae (Diptera). Revista de Entomologia, Rio de Janeiro, 8 (1 - 2), 196 - 203."]}

We present the first report of Nasutitermes guayanae feeding on human bone remains found in an urban area of the municipality of Florencia, Caquetá, Colombia, in the Colombian Amazon piedmont. The record indicates an expansion in the diet of these termites. The observation suggests that the association of N. guayanae with decomposing bodies may be a possible tool for the estimation of postmortem intervals. RESUMEN Presentamos el primer reporte de Nasutitermes guayanae alimentándose de restos óseos humanos encontrados en un área urbana del município de Florencia, Caquetá, Colombia, en el Piedemonte Amazónico colombiano. El registro indica una expasión en la dieta de esas térmitas. La observación sugiere que la asociación de N. guayanae con cuerpos en descomposición puede ser una posible herramienta para la estimación de intervalos postmortem.