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Centrophorus uyato (Rafinesque, 1810) Little Gulper Shark Synonymy. Dalatias nocturnus Rafinesque, 1810: 11, pl. 14, fig. 3 (Type locality: Sicily, Italy)— Cuvier, 1818: 455; Cuvier, 1829: 392; Cuvier, 1837: 246; Swainson, 1838: 129, 160; Swainson, 1839: 313; Duméril, 1865: 436; Jordan & Evermann, 1917: 77; Bigelow & Schroeder, 1948: 500. Spinax uyatus — Bonaparte, 1834: 49, fig. 2 (Italy); Bonaparte, 1846: 16 (Mediterranean); Böhlke, 1984: 158. Acanthias uyatus — Müller & Henle, 1839: 85 (Mediterranean); Gray, 1851: vii, 71 (Mediterranean); Steindachner, 1864: 27; Duméril, 1865: 439 (Algerian coast, Mediterranean); Barbosa du Bocage & de Brito Capello, 1866: 7, 21; Günther, 1870: 419 (Mediterranean); Canestrini, 1872: 40; Steindachner, 1875: 466; Moreau, 1881: 346 (Mediterranean); Réguis, 1882: 72; Rochebrune, 1883: 47 (Senegal and Gambia); Hilgendorf, 1884: 518; Duncan, 1891: 35 (Mediterranean); Moreau, 1892: 39; Seeley, 1895: 35 (Mediterranean); Parona, 1898: 38 (Ligurian Sea); Duncker, 1914: 291; de Buen, 1916: 303, figs (Mediterranean coast of Morocco); Landolt, 1947: 353. Centrophorus granulosus — Müller & Henle, 1839: 89, pl. 33 (Sicily, Italy); Barbosa du Bocage & de Brito Capello, 1864: 261 (Portugal); Vinciguerra, 1883: 18 (482) (Gulf of Genoa); Goode & Bean, 1896: 12, pl. 3, fig. 11 (W Atlantic, Mediterranean and adjacent areas); Boutan, 1926: 1 (Algeria); Dieuzeide, 1928a: 15, figs (Algeria); Dieuzeide, 1928b: 1314 (Algeria?); Andr & Canal, 1929: 511 (Algeria); Arcidiacono, 1931: 609 (Gulf of Squillace, Italy); Gruvel, 1931: 74 (State of Syria [Syria, Lebanon, Israel]); Ranzi, 1932: 240 (Naples, Italy); Belloc, 1934: 146, fig. (Western Sahara, Morocco, Canary Islands and Madeira); Ranzi, 1934: 343, 370 (Naples, Italy); Fowler, 1936: 73 (Mediterranean); Fowler, 1941 (in part): 231 (Mediterranean, inc. Italy); Šoljan, 1948: 66, figs (Adriatic Sea); Bigelow et al., 1955: 6 (Gulf of Mexico); Kirinčić & Lepetić, 1955: 24 (southern Adriatic Sea); Tortonese, 1956: 176, figs 94-95 (Italy); Cadenat, 1959: 748, fig. 1b (West Africa); Maurin, 1968: 82 (Morocco to Algeria); Maurin & Bonnet, 1970: 147 (Canary Islands to Cape Verde); Karrer, 1975: 64 (Namibia); Karlovac, 1976: 601, fig. 4 (Adriatic Sea); Guitart, 1979: 67, fig. 46 (Cuba); Bouchet et al., 1982: 577 (Tunisia); Zupanovic & El-Buni, 1982: 111 (Libya); Uyeno et al., 1983: 62, figs (Suriname); Compagno, 1984a: 37, figs (Atlantic and Indo-West Pacific); Gilat & Gelman, 1984: 263 (Levant Sea, Israel); McEachran & Branstetter, 1984: 130, fig. (NE Atlantic and Mediterranean); Muñoz-Chapuli, 1984: 9 (NE Atlantic);? Quéro, 1984: 43, fig.; Capapé, 1985: 97, fig. 1–7 (Eastern Atlantic); Jardas, 1985: 50 (Adriatic Sea); Anon, 1986: 93, fig. 22 (Atlantic Ocean); Bass et al., 1986 (in part): 50, fig. 5.1 (Walvis Bay and Mozambique); Golani, 1986: 23, fig. 2 (Levantine Sea, Israel); Muñoz-Chapuli & Ramos, 1989: 65, figs 1a, 3a, 4b and c, 5a, 6a, 7a (NE Atlantic and Mediterranean); Compagno et al., 1991: 54 (Hondeklip Bay to Namibia); Benli et al., 1993: 133, figs 1 and 2 (Sea of Marmara); Pisanty & Golani, 1995: 388 (Levantine Sea, Israel); Lanfranco, 1996: 6, pl. 4 (Malta); Aldebert, 1997: 284 (Gulf of Lion, France); Guallart, 1998: 1, figs (Balearic Sea); Bello, 1999: 69 (Adriatic Sea); Hernández-Hamón & Núñez, 1998: 107 (Pacific Columbia); Ungaro et al., 1999: 180 (Albania); Capapé et al., 2000: 129 (Languedoc coast, France); Bertrand et al., 2000: 5 (Gibraltar to Aegean Sea, Mediterranean); Golani & Pisanty, 2000: 71 (Levantine Sea, Israel); Baino et al., 2001: 234 (Alboran Island to Aegean Sea, Mediterranean); Guallart & Vicent, 2001: 135, fig. 4 (Balearic Sea, Spain); Bilecenoglu et al., 2002: 16 (Turkey); Schembri et al., 2003: 76, fig. 3c (Malta); McLaughlin & Morrissey, 2004: 481, fig. 3 (Cayman Trench, Jamaica); Moreno García, 2004: 214 (in part, not figs) (Mediterranean Sea); Sion et al., 2004: 155 (Ionian Sea, Greece); Golani, 2005: 11 (Levantine Sea, Israel); Lteif, 2015: 16, figs 9a, 14, (Lebanon); Serena, 2005: 27, figs, pl. I, 8 (Mediterranean Sea); Serét, 2005: 21 (Libya); Bessho, 2006: 28, figs 27–30, 33–35 (Japan, Namibia); Golani et al., 2006: 38, fig. (eastern Mediterranean); Hadjichristophorou, 2006: 163 (Cyprus); Megalofonou & Chatzispyrou, 2006: 67, fig. 4 (Crete, Greece); Mejía-Falla et al., 2007: 116 (Colombia); Psomadakis et al., 2009: 200 (Gulf of Naples, Italy); D’Onghia et al., 2010: 401 (Ionian Sea, Greece); Lipej & Dulĉić, 2010: 10 (Adriatic Sea); Damalas & Vassilopoulou, 2011: 145 (Aegean Sea, Greece); Colloca & Lelli, 2012: 12 (Lebanon); Costa et al., 2012: 7 (Portugal); Guijarro et al., 2012: 89 (Balearic Islands, Spain); Güven et al., 2012: 278 (Antalya Bay, Turkey); Iglésias, 2013: 38, pl. 16 (France and Cape Verde); Carneiro et al., 2014: 13 (Portugal); Farrugio & Soldo, 2014: 33 (Sicily, Italy and Tunisia); Veríssimo et al., 2014: 6 (Gulf of Mexico); Goren & Galil, 2015: 510 (Levant Sea, Israel); Barría et al., 2015a: 226 (Catalan Sea, Spain and Gulf of Lions, France); Barría et al., 2015b: 114 (Catalan Sea, Spain and Gulf of Lions, France); Carpenter & De Angelis, 2016: 1170, figs (Eastern Atlantic); Ramírez-Amaro et al., 2016: 639 (western Mediterranean); Cariani et al., 2017: 5 (Mediterranean);? Haroun et al., 2017: 84 (Egypt); Gajić, 2019: 101, fig. 14 (Croatia, Montenegro and Albania); Bariche & Fricke, 2020: 17, fig. 12 (Lebanon). ? Acanthias nigrescens Nardo, 1860: 70, 96 (Type locality: Venice, Italy). Entoxychirus uyatus — Gill, 1862: 498; Whitley, 1934: 199 (Australia). Acanthias ujatus — Döderlein, 1878: 30 (Sicily, Italy); Döderlein, 1881: 92 (Italy). Centrophorus uyatus — Goode & Bean, 1896: 508; Garman, 1906: 204; Garman, 1913: 9, 196. Squalus uyatus — Garman, 1899: 28. Squalus uyato — Garman, 1906: 204. Centrophorus bragancae Regan, 1906: 438 (Type locality: Cezimbra, Portugal)— Regan, 1908: 53 (coast of Portugal); Strand, 1908: 83. Centrophorus uyato — Fowler, 1936: 72, fig. 21 (Mediterranean and eastern Atlantic); Tortonese, 1938: 310 (Mediterranean); Poll 1951: 64, figs 33–34 (Angola and Namibia); Bigelow et al., 1953: 227, fig. 4 (Gulf of Mexico; Nice, Mediterranean Sea); Aksiray, 1954?: 233 (Turkish Seas); Bigelow et al., 1955: 5, 9; Springer & Bullis, 1956: 42 (Gulf of Mexico); Bigelow & Schroeder, 1957: 54, 66, 69, 72, 79–84, fig. 8e (Gulf of Mexico); Springer & Garrick, 1964: 81, 91; Krefft & Tortonese, 1973: 39 (NE Atlantic and Mediterranean); Bass et al., 1976: 31, figs 22, 24E, 24F, pl. 7 (southern Mozambique); Bridger, 1978: 26 (west of Ireland and Britain); Compagno, 1981: 8 [sharks] (Eastern Central Atlantic); Castro, 1983: 54, figs (Gulf of Mexico); Allu et al., 1984: 125 (Namibia); Compagno, 1984a: 45, figs (Atlantic, Indo-West Pacific); Compagno, 1984b: 9 [sharks] (Western Indian Ocean); McEachran & Branstetter, 1984: 132, fig. (NE Atlantic and Mediterranean); Lloris, 1986: 96, fig 24 (Namibia); Turón et al., 1986: 63 (Namibia); Fischer et al., 1987: 823 (Mediterranean); Compagno, 1988b: 603 (Comoro Islands); Compagno et al., 1989: 24, fig. (Hondeklip Bay, South Africa to Namibia);? Clark & Kristof, 1990: 277, fig. 9 (Caribbean); Springer, 1990: 11 (Gulf of Mexico, Mediterranean, and Natal, South Africa); Applegate et al., 1993: 35 (Atlantic Mexican waters); Gomon et al., 1994: 92, 94, figs 30, 31 (southern Australia); Last & Stevens, 1994: 60, figs, pl. 4 (fig. 8.5) (southern Australia); Meriç, 1995: 192 (Sea of Marmara, Turkey); Perry et al., 1995: 139 (Gulf of Mexico); Reiner, 1996: 22, fig. (Cape Verde); Bonfil, 1997: 105 (Veracruz, Mexico); Joseph, 1999: [unpaginated] (Sri Lanka); Cervigón & Alcalá, 1999 (Venezuela); Clarke, 2000: 377 (Rockall Trough, NE Atlantic); Baino et al., 2001: 234 (Alboran Island to Aegean Sea, Mediterranean); Graham et al., 2001: 551 (south-eastern Australia); Yearsley et al., 2001: 35, 360 (southern Australia); Bilecenoglu et al., 2002: 17 (Turkey); Daley et al., 2002: 53 (southern Australia); Ali & Saad, 2003: 58, fig. (Syria); Schembri et al., 2003: 77, fig. 3d (Malta); Serena, 2005: 28, figs, pl. I, 9 (Mediterranean Sea, NE Atlantic, Western Indian, Gulf of Mexico and Taiwan); Meriç et al., 2007: 31 (Turkey); White, 2008: 87, figs (southern Australia); Scacco et al., 2010: 39, fig. 3f (Mediterranean Sea); Castro, 2011: 81, figs 16a-e (north-western Atlantic); Davenport et al.: 2011: 557 (north-western Atlantic, USA); White et al., 2013: 36, fig. 2, 15, 16 (Atlantic); Veríssimo et al., 2014: 6, fig. 5 (Gulf of Mexico); Hipes, 2015: 1, fig. 11 (Gulf of Mexico, USA); Wienerroither et al., 2015: 834, fig. 2 (northern Norway); Farrag, et al., 2016: 481, fig. 2a (Egypt); Driggers et al., 2017: 52 (Gulf of Mexico); Haroun et al., 2017: 84 (Egypt); Lteif et al., 2017: 1491 (Lebanon); Biscoito et al., 2018: 471, fig. 7 (Madeira, Portugal); Carneiro et al., 2019: 36 (Portugal); Ehemann et al., 2019: 4 (Venezuela); Fernando et al., 2019: 231, figs 5e, 18d-f (Mutur, Sri Lanka); Follesa et al., 2019: 85 (Mediterranean); Psomadakis et al., 2019: 162, figs, pl. X (fig. 70) (Myanmar); Iglésias, 2020: 46, pl. 21 (France and Cape Verde); Ebert & Dando, 2021: 217, figs (NE Atlantic and Mediterranean); Kousteni et al., 2021: 1, figs 2 and 3 (Cypriot waters). Centrophorus machiquensis Maul, 1955: 5, figs 13–16 (Type locality: Madeira)— Krefft & Tortonese, 1973: 39 (Madeira); Ali & Saad, 2003: 57, fig. (Syria); Biscoito et al., 2018: 470, fig. 6 (Madeira, Portugal); Almeida & Biscoito, 2019: 99 (Canary Islands and Madeira); Carneiro et al., 2019: 36 (Portugal). Centrophorus ujato — Tortonese, 1956: 178, fig. 96 (Genova, Italy); Bini, 1967: 97, fig. (Italy); Sara, 1968: 1, figs 1–3 (west of Sicily); FAO, 1971: no pagination (Mediterranean Sea). Centrophorus spp. (granulosus group)— Forster et al, 1970 (in part): 388 (Western Indian Ocean). Centrophorus (forme) uyato-machiquensis — Cadenat & Blache, 1981: 58, figs. 36, 37 and 40 (NE Atlantic). Centrophorus spp. — Peyronel et al., 1984: 643 (Bay of Ajaccio, Corsica, France). Centrophorus cf. harrissoni (Undescribed gulper shark #2)— Kiraly et al., 2003: 2 (Puerto Rico, US Virgin Islands, Virginia, North Carolina and Florida Straits to Dry Tortugas). Centrophorus bragance — Hernández-Hamón & Núñez, 1998: 108 (as questionable synonym of C. granulosus). Centrophorus sp. (uyato ?)— Morón et al., 1998: 144 (Beruwela, Sri Lanka). Centrophorus sp. cf. uyato — Saad, et al., 2004: 430 (Syria). ? Centrophorus sp. (non uyato)— Serét, 2005: 21 (Libya). Centrophorus zeehaani White, Ebert & Compagno, 2008: 1, figs 8-10 (Type locality: South Australia)— Last & Stevens, 2009: 68, pl. 9.7, figs (southern Australia); Pethybridge et al., 2010: 1369 (Tasmania and Great Australian Bight, Australia); Pethybridge et al., 2011: 2743 (Tasmania and Victoria, Australia); Graham & Daley, 2011: 583 (southern Australia); Daley et al., 2012: 708 (southern Australia); White et al., 2013: 41 (southern Australia); Daley et al., 2015:127 (southern Australia); Wienerroither et al., 2015: 834, fig. 2 (northern Norway); Bineesh et al., 2016: 461 (Kollam, India). Centrophorus cf. uyato — McLaughlin & Morrissey, 2005: 1185, figs 2, 3 (Cayman Trench, Jamaica); Veríssimo et al., 2014: 7 (Gulf of Mexico); Serena et al., 2020: 502, 509 (Mediterranean); Bellodi et al., 2022: 2 (Mediterranean Sea). Centrophorus zeehani — Daley et al., 2012: fig. 2 (misspelling; southern Australia); Bineesh et al., 2016: 466 (misspelling; Kollam, India). Centrophorus ‘uyato’ — White et al., 2017: 86 (Eastern Atlantic); Almeida & Biscoito, 2019: 100 (Mediterranean Sea, Canary Islands and Madeira). Centrophorus cf. granulosus — Follesa et al., 2019: 85 (Mediterranean); FAO, 2018: unpaginated, fig. (Mediterranean Sea). ? Centrophorus granulosus — Parenti, 2019: 102 (Sicily). Material examined. Neotype: BMNH 2021.10.4.1 (eviscerated; GenBank accession ON167716), female 983 mm TL, between Gorgona and Capraia islands, Ligurian Sea, 43°19.8′ N, 9°56.1′ E, 180 m, 20 Dec. 2012. Other specimens: Australia: AMS I 44310 –001 (paratype of Centrophorus zeehaani; GenBank accession ON167706), adult male 826 mm TL, southwest of Coffin Bay, South Australia, 35°14′ S, 134°29′ E, 360–600 m, 28 July 2005; CSIRO CA 4104, adult male 843 mm TL, east of Gabo Island, Victoria, 37°40′ S, 150°15′ E, 504–508 m, 4 May 1984; CSIRO H 866–02, immature male 456 mm TL, CSIRO H 867–01, female 439 mm TL, east of Jervis Bay, New South Wales, 34°58′ S, 151°09′ E, 490–576 m, 10 Sep. 1986; CSIRO H 2268–02, adult male 800 mm TL, west of Bunbury, Western Australia, 33°03′ S, 114°25′ E, 701 m, 10 Feb. 1989; CSIRO H 6307–01 (skeletal parts), female 1027 mm TL, 12 July 2004, east of Flinders Island, Tasmania, ~ 40° S, ~ 149° E, 350–430 m; CSIRO H 6309–01 (skeletal parts; GenBank accession ON167708), adult male 865 mm TL, CSIRO H 6309–02 (skeletal parts), adult male 876 mm TL, CSIRO H 6309–04 (skeletal parts), adult male 906 mm TL, east of Flinders Island, Tasmania, ~ 40° S, ~ 149° E, 400–450 m, 1 Aug. 2004; CSIRO H 6310–04 (skeletal parts), female 970 mm TL, northeast of Flinders Island, Tasmania, 39°04′ S, 148°39′ E, 500–680 m, 24 Jul. 1986; CSIRO H 6311–01 (skeletal parts), female 655 mm TL, east of St. Helens, Tasmania, 41°27′ S, 148°44′ E, 850–860 m, 5 Jun. 1987; CSIRO H 6500-02, adult male 862 mm TL, east of Flinders Island, Tasmania, 40°15′ S, 148°45′ E, 329–512 m, 21 Aug. 2003; CSIRO H 6503–02 (skeletal parts), female 991 mm TL, CSIRO H 6503–03 (skeletal parts; GenBank accession ON167709), female 1023 mm TL, CSIRO H 6503–04 (skeletal parts; GenBank accession ON167710), female 987 mm TL, CSIRO H 6503–05 (skeletal parts), female 957 mm TL, northeast of Flinders Island, Tasmania, 39°20′ S, 148°45′ E, 370–420 m, 7 Apr. 2003; CSIRO H 6504–02, adult male 854 mm TL, CSIRO H 6504–03, female 817 mm TL, CSIRO H 6504–04, juvenile male 666 mm TL, CSIRO H 6504–05, adult male 861 mm TL, east of Jervis Bay, New South Wales, 35°12′ S, 150°58′ E, 320–500 m, July to Aug. 2003; CSIRO H 6628–01 (paratype of Centrophorus zeehaani), immature male 506 mm TL, CSIRO H 6628–02 (paratype of Centrophorus zeehaani; GenBank accession ON167705), immature male 645 mm TL, CSIRO H 6628–03 (paratype of Centrophorus zeehaani), adult male 875 mm TL, CSIRO H 6628–04 (paratype of Centrophorus zeehaani), adult male 910 mm TL, CSIRO H 6628–05 (holotype of Centrophorus zeehaani), adult male 893 mm TL, CSIRO H 6628–06 (paratype of Centrophorus zeehaani), adult male 852 mm TL, CSIRO H 6628–07 (paratype of Centrophorus zeehaani), adult male 906 mm TL, NMV A 29736 –001 (paratype of Centrophorus zeehaani), adult male 820 mm TL, southwest of Coffin Bay, South Australia, 35°14′ S, 134°29′ E, 360–600 m, 28 July 2005; CSIRO unreg. (DB 02/181), Great Australian Bight, adult male 857 mm TL; CSIRO unreg. (LJVC 880517), female 840 mm TL, unknown location; PMH095-11 (jaws only), female 104 cm TL, Albany, Western Australia. Eastern Atlantic (including Mediterranean): AMNH 78267, female 922 mm TL, AMNH 78269, female 937 mm TL, AMNH 78271, female 1016 mm TL, AMNH 78273, adult male 872 mm TL, AMNH 78277, adult male 895 mm TL, AMNH 78279, adult male 890 mm TL, between Tenerife and Gran Canaria, Canary Islands, Spain, 3 Oct. 1986; AMNH 78280, female 996 mm TL, AMNH 78282, female 1059 mm TL, AMNH 78283, female 1056 mm TL, AMNH 78284, adult male 832 mm TL, AMNH 78285, female 921 mm TL, AMNH 78286, female 1004 mm TL, AMNH 78291, female 1034 mm TL, AMNH 78292, female 980 mm TL, AMNH 78294, adult male 891 mm TL, between Tenerife and Gran Canaria, Canary Islands, Spain, 4 Oct. 1986; BMNH 1862.4.22.29, female 449 mm TL, Madeira, Portugal; BMNH 1864.7.18.1, female 894 mm TL, Madeira, Portugal; BMNH 1904.11.30.9- 10 (2 specimens), female 1036 mm TL, adult male 888 mm TL, west of Faro, Portugal, 662 m depth; BMNH 1904.11.30.11 (paralectotype of Centrophorus bragancae), juvenile male 488 mm TL, off Sesimbra, Portugal, 841 m depth; BMNH 1904.11.30.12 (lectotype of Centrophorus bragancae), female 467.5 mm TL, off Sesimbra, Portugal, 505 m depth; BMNH 2013.9.20.34, female 875 mm TL, BMNH 2013.9.20.35, female 930 mm TL, BMNH 2013.9.20.36, female 935 mm TL, BMNH 2013.9.20.37, female 731 mm TL, BMNH 2013.9.20.38, adult male 843 mm TL, BMNH 2013.9.20.39, adult male 865 mm TL, northeast Atlantic; CSIRO H 7471-01 (GenBank accession ON167715), adult male 853 mm TL, east of Corsica, France, 42°05.8′ N, 9°44.5′ E, 550–565 m, 30 May 2012; CSIRO H 7472-01, juvenile male 440 mm TL, southeast of Corsica, France, 41°47.9′ N, 9°30.5′ E, 466–480 m, 31 May 2012; ERB 1288, male 770 mm TL, Concarneau fish-market, France, 9 Sep. 2000; ERB 0763, male, 850 mm TL, Concarneau, France, 13 May 2009; HUJ 10885, female 760 mm TL, Hadera, Israel, 13 Sep. 1982; HUJ 11339, adult male 796 mm TL, Haifa Bay, Israel, 1 Mar. 1983; HUJ 17029, female 476 mm TL, Haifa, Israel, 16 Mar. 1983; HUMZ 151304, juvenile male 578 mm TL; HUMZ 151306, female 978 mm TL, Namibia; HUJ 21135 (GenBank accession ON167718), female 753 mm TL, Haifa fishing port, Israel, 29 July 2013; MNHN-IC 1905- 0568, juvenile male 454 mm TL, Portugal, 460 m depth, June 1903; MNHN-IC 1969-0269, juvenile male 499 mm TL, southwest of Monrovia, Liberia, 6°08′ N, 10°56′ W, 400 m depth, 27 Apr. 1964; MNHN-IC 0000-1224, female 537 mm TL, Naples, Italy; MNHN-IC 2005-0169, adult male 892 mm TL, west of Ireland, 28 Apr. 2005; NMW 15009, female 889 mm TL, Nice; NMW 63020, adult male 880 mm TL, Nice, France, 1902, Published as part of White, William T., Guallart, Javier, Ebert, David A., Naylor, Gavin J. P., Mo, Ana Veríssi-, Cotton, Charles F., Harris, Mark, Serena, Fabrizio & Iglésias, Samuel P., 2022, Revision of the genus Centrophorus (Squaliformes: Centrophoridae): Part 3 - Redescription of Centrophorus uyato (Rafinesque) with a discussion of its complicated nomenclatural history, pp. 1-51 in Zootaxa 5155 (1) on pages 15-41, DOI: 10.11646/zootaxa.5155.1.1, http://zenodo.org/record/6669082, {"references": ["Rafinesque, C. S. (1810) Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia, con varie osservazioni sopra i medisimi. Sanfilippo, Palermo, 1, 3 - 69. https: // doi. org / 10.5962 / bhl. title. 104418", "Cuvier, G. (1818) Dictionnaire des sciences naturelles, dans lequel on traite meithodiquement des diffeirens e \u02c6 tres de la nature, consideireis soit en eux-me \u02c6 mes, d'apreIs l'eitat actuel de nos connoissances, soit relativement aI l'utilitei qu'en peuvent retirer la meidecine, l'agriculture, le commerce et les artes. Suivi d'une biographie des plus ceileIbres naturalists. Volume 12. Strasbourg, F. G. Levrault, 564 pp.", "Cuvier, G. (1829) Le Regne Animal, distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee. Edition 2. 2, 406 pp.", "Cuvier, G. (1837) The animal kingdom, arranged according to its organization, serving as a foundation for the natural history of animals: and an introduction to comparative anatomy. Volume 2. G. Henderson, London, 412 pp.", "Swainson, W. (1838) On the natural history and classification of fishes, amphibians, & reptiles, or monocardian animals. A. Spottiswoode, London, 1, 368 pp. https: // doi. org / 10.5962 / bhl. title. 62140", "Swainson, W. (1839) On the natural history and classification of fishes, amphibians, & reptiles, or monocardian animals, Volume 2. Spottiswoode & Co., London, 452 pp. https: // doi. org / 10.5962 / bhl. title. 62140", "Dumeril, A. H. A. (1865) Histoire naturelle des poissons ou ichthyologie generale. Tome Premier. I. Elasmobranches. Plagiostomes et Holocephales ou Chimeres, 720 pp.", "Jordan, D. S. & Evermann, B. W. (1917) The genera of fishes, from Linnaeus to Cuvier, 1758 - 1833, seventy-five years, with the accepted type of each. A contribution to the stability of scientific nomenclature. Leland Stanford Jr. University Publications, University Series, 27, 1 - 161.", "Bigelow, H. B. & Schroeder, W. C. (1948) Cyclostomes. Sharks. In: Fishes of the Western North Atlantic. Memoirs of the Sears Foundation of Marine Research, 1, 29 - 576.", "Bonaparte, C. L. (1834) Iconografia della fauna italica per le quattro classi degli animali vertebrati. Tomo III. Pesci. Roma. Fasc. 6 - 11, puntata 29 - 58. https: // doi. org / 10.5962 / bhl. title. 70395", "Bonaparte, C. L. (1846) Catalogo metodico dei pesci europei. Atti della Settima Adunanza degli Scienziati Italiani 7 a Adunanza, Napoli, Part 2, 95 pp. https: // doi. org / 10.5962 / bhl. title. 59507", "Bohlke, E. B. (1984) Catalog of type specimens in the ichthyological collection of the Academy of Natural Sciences of Philadelphia. Special Publication, Academy of Natural Sciences of Philadelphia, 14, 1 - 216", "Muller, J. & Henle, F. G. J. (1839) Systematische Beschreibung der Plagiostomen. Plagiostomen, Berlin, pp. 29 - 102. https: // doi. org / 10.5962 / bhl. title. 6906", "Gray, J. E. (1851) List of the specimens of fish in the collection of the British Museum. Part I. - Chondropterygii. London, 160 pp.", "Steindachner, F. (1864) Ichthyologische Notizen. Anzeiger der Kaiserlichen Akademie der Wissenschaften, MathematischNaturwissenschaftlichen Classe, 1 (5), 1 - 70. https: // doi. org / 10.5962 / bhl. title. 16269", "Gunther, A. (1870) Catalogue of the fishes in the British Museum. Catalogue of the Physostomi, containing the families Gymnotidae, Symbranchidae, Muraenidae, Pegasidae, and of the Lophobranchii, Plectognathi, Dipnoi, Ganoidei, Chondropterygii, Cyclostomata, Leptocardii, in the British Museum. Volume 8, 549 pp.", "Canestrini, G. (1872) Pesci d'Italia. Parts II. In: Cornalia, Fauna d'Italia, 1870 - 74, vol. 3. F. Vallardi, Milano, pp. 37 - 208.", "Steindachner, F. (1875) Ichthyologische Beitrage (II) - I. Die Fische von Juan Fernandez in den Sammlungen des Wiener Museums. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Klasse, 71, 443 - 479.", "Moreau, E. (1881) Histoire naturelle des poissons de la France. Vol. 1, 480 pp. https: // doi. org / 10.5962 / bhl. title. 12541", "Reguis, J. M. F. (1882) Essai sur l'Histoire Naturelle des Vertebres de la Provence et des Departements Circonvoisins, 425 pp. https: // doi. org / 10.5962 / bhl. title. 8200", "Rochebrune, A. T. de. (1883) Faune de la Senegambie. Poissons. Actes de la Societe Linneenne de Bordeaux, 36 (6), 20 - 190. https: // doi. org / 10.5962 / bhl. title. 50898", "Hilgendorf, F. (1884) Bericht uber die Leistungen in der Ichthyologie wahrend des Jahres 1883. Archiv fur Naturgeschichte, 50, 435 - 521.", "Duncan, P. M. (1891) Cassell's Natural History. Cassell & Company Ltd., Vol. 5., 384 pp. https: // doi. org / 10.5962 / bhl. title. 31259", "Moreau, E. (1892) Manuel d'Ichthyologie francaise. Paris, 650 pp. https: // doi. org / 10.5962 / bhl. title. 8444", "Seeley, H. G. (1895) Pisces. In: Duncan, P. M. (Ed.), Cassell's natural history. Volume 5. Cassell and Company, Ltd., London, Paris and Melbourne, pp. 1 - 149.", "Parona, C. (1898) La pesca marittima in Liguria. Bollettino dei Musei di Zoologia e Anatomia Comparata della R. universita di Genova, 66, 3 - 68.", "Duncker, G. (1914) Generalindex zu Franz Steindachners Ichthyologischen Mitteilungen, Notizen und Beitragen. Mitteilungen aus den Naturhistorischen Museum in Hamburg, 31 (2), 285 - 352.", "de Buen, F. (1916) Sobre la presencia y caracterizacion del Acanthias uyatus Mull. Henl., de la costa mediterranea de Marruecos. Boletin de la Real Sociedad Espanola de Historia Natural, 16, 303 - 305.", "Landolt, H. H. (1947) Ueber den Zahnwechsel bei Selachiern. Revue Suisse de Zoologie, 54 (19), 305 - 368.", "Barbosa du Bocage, J. V. & de Brito Capello, F. (1864) Apontamentos para a ichthyologia de Portugal. Peixes Plagiostomos, Primeira Parte Esqualos, 40 pp.", "Vinciguerra, D. (1883) Resultati ittiologici delle crociere del Violante. Annali del Museo Civico di Storia Naturale di Genova, 18, 465 - 590.", "Goode, G. B. & Bean, T. H. (1896) Oceanic ichthyology, a treatise on the deep-sea and pelagic fishes of the world, based chiefly upon the collections made by the steamers Blake, Albatross, and Fish Hawk in the northwestern Atlantic, with an atlas containing 417 figures. Special Bulletin U. S. National Museum, 2, 553 p. https: // doi. org / 10.5962 / bhl. title. 48521", "Boutan, L. (1926) Etude sur le Centrophorus granuleux. Bulletin des travaux publies par la Station d'aquiculture et de peche de Castiglione, Alger, 1, 1 - 146.", "Dieuzeide, R. (1928 a) Etude histologique de la peau du Centrophore granuleux (Centrophorus granulosus Mull. et Henl.). Bulletin des travaux publies par la Station d'aquiculture et de peche de Castiglione, Alger, 1, 15 - 24.", "Dieuzeide, R. (1928 b) Sur la transformation et la disparition de certains denticules de la peau du Centrophore (Centrophorus granulosus Mull. et Henl.). Comptes Rendus de l'Academie des Sciences, 186, 1314.", "Andr, M. & Canal, H. (1929) Contribution l'etude des huiles d'animaux marins. Recherches sur l'huile de centrophore granuleux (Centrophorus granulosus, Muller et Henl). tude pecime des matires insaponificables pecimen de l'oeuf, du foie du foetus et du foie des animaux adultes. Bulletin de la Societe chimique de France, 45, 511 - 524.", "Arcidiacono, F. (1931) La pesca del pesce vacca Hexanchus griseus L.) nella marina di Riposto. Bollettino di pesca, piscicoltura e idrobiologia, 7, 608 - 612.", "Gruvel, A. (1931) Les Etats de Syrie. Richesses marines et fluviales. Exploitation actuelle - Avenir. Societe d'Editions Geographiques, Maritimes et Coloniale, 453 pp.", "Ranzi, S. (1932) Le base fisio-morfologiche dello sviluppo embrionale dei Selaci. Pt. I. Pubblicazioni della Stazione zoologica di Napoli, 12, 209 - 290.", "Belloc, G. (1934) Catalogue des pecime comestibles du Maroc et de la Cote occidentale d'Afrique (du Cap Sorel au Cap Vert), Premiere partie - Poissons cartilagineux. Revue des Travaux de l'Institut des Peches Maritimes, 7, 117 - 193.", "Ranzi, S. (1934) Le basi fisio-morphologiche dello sviluppo embrionale dei Selaci. Pts. II e III. Pubblicazioni della Stazione zoologica di Napoli, 13, 331 - 437.", "Fowler, H. W. (1936) The marine fishes of West Africa based on the collection of the American Museum Congo expedition, 1909 - 1915. Part 1. Bulletin of the American Museum of Natural History, 70, 1 - 605.", "Fowler, H. W. (1941) Contributions to the biology of the Philippine archipelago and adjacent regions. The fishes of the groups Elasmobranchii, Holocephali, Isospondyli, and Ostariophysi obtained by the United States Bureau of Fisheries Steamer \" Albatross \" in 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Bulletin of the United States National Museum, 100, 1 - 879.", "Soljan, T. (1948) Fishes of the Adriatic (Ribe Jadrana). Fauna et flora Adriatica, Volume 1, Pisces. Nakledni Zavod Hrvatske, Zagreb.", "Bigelow, H. B., Schroeder, W. C. & Springer, S. (1955) Three new shark records from the Gulf of Mexico. Breviora, 49, 1 - 12.", "Kirincic, J. & Lepetic, V. (1955) Recerches sur l'ichthyobenthos dans les profounders de l'Adriatique meridionale et possibilite d'exploration au moyen des palangres. Acta Adriatica, 7, 1 - 113.", "Tortonese, E. (1956) Leptocardia, Ciclostomata, Selachii. Fauna d'Italia, vol. 2. Calderini, Bologna, 334 pp.", "Cadenat, J. (1959) Note d'Ichtyologie pec-africaine. XXII. Centrophorus lusitanicus Bocage et Capello 1864 (Selacien Squalidae), espece valable differente de C. granulosus. Bulletin de l'Institut Francais D'Afrique Noire, 21, 743 - 746.", "Maurin, C. (1968) Ecologie ichthyologique des fonds chalutables Atlantiques (de la baie ibero-marocaine a la Mauritanie) et de la Mediterranee occidentale. Revue des Travaux de l'Institut des Peches Maritimes, 32, 1 - 136.", "Maurin, C. & Bonnet, M. (1970) Poissons des cotes nord-ouest africaines (campagnes de la \" Thalassa \" 1962 et 1968). Revue des Travaux de l'Institut des Peches Maritimes, 34, 125 - 170.", "Karrer, C. (1975) Uber fische aus dem Sudostatlantik (Teil 2). Mitteilungen aus dem Zoologischen Museum Berlin, 51, 63 - 82.", "Karlovac, O. (1976) Jadranski morski psi iz porodice Squalidae. Pomorski Zbornik, 14, 595 - 610.", "Guitart, D. J. (1979) Sinopsis de los Peces marinas de Cuba. Tomo 1. Editorial Cientifico - Tecnica, Havana, Cuba.", "Bouchet, C., Capape, C., Chadli, A. & Baouendi, A. (1982) Mise en evidence d'une secretion holocrine dans l'uterus gravide a terme d'un requin-chagrin Centrophorus granulosus (Schneider, 1801) (Pisces, Squalidae). Archives de l'Institut Pasteur de Tunis, 53, 577 - 586.", "Zupanovic, S. & El-Buni, A. A. (1982) A contribution to demersal fish studies off the Libyan coast. Socialist People's Libyan Arab Jamahiriya, General Fisheries Authority, Marine Research Centre, Bulletin, 3, 77 - 122.", "Uyeno, T., Matsuura, K. & Fujii, E. (1983) Fishes trawled off Suriname and French Guiana. Japan Marine Fishery Resource Research Centre, Tokyo.", "Compagno, L. J. V. (1984 a) FAO Species Catalogue. Vol. 4, Sharks of the World. An annotated and illustrated catalogue of shark species known to date. FAO Fisheries Synopsis No. 125. vol. 4, pt. 1 (noncarcharhinoids), pp. viii, 1 - 250, pt. 2 (Carcharhiniformes), pp. 251 - 655.", "Gilat, E. & Gelman, A. (1984) On the sharks and fishes observed using underwater photography during a deep-water cruise in the Eastern Mediterranean. Fisheries Research, 2, 257 - 271. https: // doi. org / 10.1016 / 0165 - 7836 (84) 90029 - 8", "McEachran, J. D. & Branstetter, S. (1984) Squalidae. In: Whitehead, P. J. P., Bauchot, M. - L., Hureau, J. - C., Nielsen, J. & Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and the Mediterranean / Poissons de l'Atlantique du Nord-Est et de la Mediterranee. Vol. I. UNESCO, Paris, pp. 128 - 147.", "Munoz-Chapuli, R. (1984) Ethologie de la reproduction chez quelques requins de l'Atlantique nord-est. Cybium, 8, 1 - 14.", "Quero, J. C. (1984) Les poissons de mer des peches Francaises. Jacques Grancher, Paris.", "Capape, C. (1985) Nouvelle description de Centrophorus granulosus (Schneider, 1801) (Pisces, Squalidae). Donnees sur la biologie de la reproduction et le pecim alimentaire des pecimens des cotes tunisiennes. Bulletin de l'Institut National Scientifique et Technique d'Oceanographie et de Peche de Salommbo, 12, 97 - 141.", "Jardas, I. (1985) Pregled riba (sensu lato) Jadranskog mora (Cyclostomata, Selachii, Osteichthyes) s obzirom na taksonomiju I utvrdeni broj. Biosistematika, 11, 45 - 74.", "Anon. (1986) Monografiias de zoologiia marina. Volume 1. Instituto de Ciencias del Mar, Barcelona, Consejo Superior de Investigaciones Cientiificas, Madrid.", "Bass, A. J., Compagno, L. J. V. & Heemstra, P. C. (1986) Family no. 5: Squalidae, pp. 49 - 62. In: Smith, M. M. & Heemstra, P. C. (eds). Smith's Sea fishes. Berlin: Springer-Verlag.", "Golani, D. (1986) On deep-water sharks caught off the Mediterranean coast of Israel. Israel Journal of Zoology, 34, 23 - 31.", "Munoz-Chapuli, R. & Ramos, F. (1989) Review of the Centrophorus sharks (Elasmobranchii, Squalidae) of the eastern Atlantic. Cybium, 13, 65 - 81.", "Compagno, L. J. V., Ebert, D. A. & Cowley, P. A. (1991) Distribution of offshore demersal cartilaginous fish (class Chondrichthyes) off the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11, 43 - 139. https: // doi. org / 10.2989 / 025776191784287664", "Benli, H. A., Cihangir, B. & Bizsel, K. C. (1993) A new record for the Sea of Marmara; (Family: Squalidae) Centrophorus granulosus (Bloch & Schneider, 1801). Doda, Turkish Journal Of Zoology, 17, 133 - 135.", "Pisanty, S. & Golani, D. (1995) Vertical distribution of demersal fish on the continental slope of Israel (Eastern Mediterranean). In: Armantrout, N. B. (Ed.), Condition of the world's aquatic habitats. Proceedings of the World Fisheries Congress, Theme 1. Oxford & IBH Publishing Co. Pvt. Ltd, New Delhi.", "Lanfranco, G. G. (1996) The fish around Malta (Central Mediterranean). Progress Press Co. Ltd. (Malta), 132 pp.", "Aldebert, Y. (1997) Demersal resources of the Gulf of Lions (NW Mediterranean). Impact of exploitation on fish diversity. Vie et Milieu - Life and Environment, 47, 275 - 284.", "Guallart, J. (1998) Contribucion al conocimiento de la biologia y la taxonomia del tiburon batial Centrophorus granulosus (Bloch y Schneider, 1801) (Elasmobranchii, Squalidae) en el mar balear (Mediterraneo Occidental). PhD Thesis. Universitat de Valencia, Valencia, 291 pp.", "Bello, G. (1999) The chondrichthyans of the Adriatic Sea. Acta Adriatica, 40, 65 - 76. https: // doi. org / 10.1093 / mollus / 65.2.233", "Hernandez-Hamon, H. & Nunez, S. R. (1998) Nuevo registro para el caribe sur Colombiano del tiburon quelvacho Centrophorus granulosus (Schneider) (Pisces: Centrophoridae). Boletin de Investigaciones Marinas y Costeras, 27, 107 - 112. https: // doi. org / 10.25268 / bimc. invemar. 1998.27.0.336", "Ungaro, N., Marano, C. A., Marsan, R., Martino, M., Marzano, M. C., Strippoli, G. & Vlora, A. (1999) Analysis of demersal species assemblages from trawl surveys in the South Adriatic Sea. Aquatic Living Resources, 12, 177 - 185. https: // doi. org / 10.1016 / S 0990 - 7440 (00) 88469 - 7", "Capape, C., Tomasini, J. A. & Quignard, J. P. (2000) The pleurotremate elasmobranchs from the Languedoc coast (southern France): biological and demographic observations. Vie et Milieu - Life and Environment, 50, 123 - 133.", "Golani, D. & Pisanty, S. (2000) Biological aspects of the Gulper shark, Centrophorus granulosus (Bloch and Schneider, 1801), from the Mediterranean coast of Israel. Acta Adriatica, 41, 71 - 78.", "Guallart, J. & Vicent, J. J. (2001) Changes in composition during embryo development of the gulper shark, Centrophorus granulosus (Elasmobranchii, Centrophoridae): an assessment of maternal-embryonic nutritional relationships. Environmental Biology of Fishes, 61, 135 - 150. https: // doi. org / 10.1023 / A: 1011080429625", "Bilecenoglu, M., Taskavak, E., Mater, S. & Kaya, M. (2002). Checklist of the marine fishes of Turkey. Zootaxa, 113 (1), 1 - 194. https: // doi. org / 10.11646 / zootaxa. 113.1.1", "Schembri, T., Fergusson, I. K. & Schembri, P. J. (2003) Revision of the records of shark and ray species from the Maltese Islands (Chordata: Chondrichthyes). The Central Mediterranean Naturalist, 4, 71 - 104.", "McLaughlin, D. M. & Morrissey, J. F. (2004) New records of elasmobranchs from the Cayman Trench, Jamaica. Bulletin of Marine Science, 75, 481 - 485.", "Sion, L., Bozzano, A., D'Onghia, G., Capezzuto, F. & Panza, M. (2004) Chondrichthyes species in deep waters of the Mediterranean Sea. Scientia Marina, 68 (Suppl. 3), 153 - 162. https: // doi. org / 10.3989 / scimar. 2004.68 s 3153", "Golani, D. (2005) Checklist of the Mediterranean fishes of Israel. Zootaxa, 947 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 947.1.1", "Lteif, M. (2015) Biology, distribution and diversity of cartilaginous fish species along the Lebanese coast, eastern Mediterranean. Universite de Perpignan via Domitia, France, 262 pp.", "Serena, F. (2005) Field identification guide to the sharks and rays of the Mediterranean and Black Sea. FAO Species Identification Guide for Fishery Purposes. FAO, Rome.", "Seret, B. (2005) Chondrichthyan fishes of Libya: proposal for a research programme. United Nations Environment Programme, Mediterranean Action Plan, Regional Activity Centre for Specially Protected Areas, Tunisia, 31 pp.", "Bessho, M. (2006) Taxonomic revision of the genus Centrophorus (Centrophoriformes: Centrophoridae) in Japanese waters. Unpublished Masters Thesis, Graduate School of Fisheries Science, Hokkaido University, 85 pp.", "Golani, D., Ozturk, B. & Basuta, N. (2006) Fishes of the eastern Mediterranean. Turkish Marine Research Foundation, Istanbul, Turkey, 260 pp.", "Hadjichristophorou, M. (2006) Chondrichthyes in Cyprus. In: Basusta, N., Keskin, C., Serena, F. & Seret, B. (Eds.), The proceedings of the workshop on Mediterranean cartilaginous fish with emphasis on Southern and Eastern Mediterranean. Turkish Marine Research Foundation, Istanbul-Turkey.", "Megalofonou, P. & Chatzispyrou, A. (2006) Sexual maturity and feeding of the gulper shark, Centrophorus granulosus, from the eastern Mediterranean Sea. Cybium, 30, 67 - 74. https: // doi. org / 10.26028 / cybium / 2006 - 304 supp- 009", "Mejia-Falla, P. A., Navia, A. F., Mejia-Ladino, L. M., Acero, A. P. & Rubio, E. A. (2007) Tiburones y rayas de Colombia (Pisces: Elasmobranchii): lista actualizada, revisada y comentada. Boletin de Investigaciones Marinas y Costeras, 36, 111 - 149.", "Psomadakis, P. N., Maio, N. & Vacchi, M. (2009) The chondrichthyan biodiversity in the Gulf of Naples (SW Italy, Tyrrhenian Sea): an historical overview. Cybium, 33, 199 - 209. https: // doi. org / 10.26028 / cybium / 2009 - 333 - 003", "D'Onghia, G., Maiorano, P., Sion, L., Giove, A., Capezzuto, F., Carlucci, R. & Tursi, A. (2010) Effects of deep-water coral banks on the abundance and size structure of the megafauna in the Mediterranean Sea. Deep Sea Research Part II Topical Studies in Oceanography, 57, 397 - 411. https: // doi. org / 10.1016 / j. dsr 2.2009.08.022", "Lipej, L. & Dulcic, J. (2010) Checklist of the Adriatic Sea fishes. Zootaxa, 2589 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 2589.1.1", "Damalas, D. & Vassilopoulou, V. (2011) Chondrichthyan by-catch and discards in the demersal trawl fishery of the central Aegean Sea (Eastern Mediterranean). Fisheries Research, 108, 142 - 152. https: // doi. org / 10.1016 / j. fishres. 2010.12.012", "Colloca, F. & Lelli, S. (2012) Report of the FAO EastMed support to the fishing trials carried out off the South Lebanese Coast. EastMed Technical Documents 14. FAO, Rome, 38 pp.", "Costa, F. O., Landi, M., Martins, R., Costa, M. H., Costa, M. E., Carneiro, M., Alves, M.

Family Dasyatidae Dasyatidae (stingrays) are the dominant group of rays in the coastal tropical Indo-West Pacific. While stingray taxonomy is problematic, recent revisions have vastly improved knowledge of this family (Last et al. 2016a). Dasyatidae are amongst the most commonly landed ray groups in Bangladesh, with high demand for consumption and accessories (Last et al. 2016a; Haque et al. 2018). This report identified seven species of Dasyatidae rays in addition to the 24 species previously reported in Bangladesh (Roy et al. 2015)., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 213, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Last, P., Naylor, G., Seret, B., White, W., de Carvalho, M. & Stehmann, M. (2016 a) Rays of the World. CSIRO publishing, Melbourne, ix + 790 pp.","Haque, A. B., Biswas, A. & Latifa, G. A. (2018) Observations of shark and ray products in the processing centres of Bangladesh, trade in CITES species and conservation need. TRAFFIC Bulletin, 30 (1), 6 - 14.","Roy, B. J., Singha, N. K., Rhaman, M. G. & Ali, A. H. M. H. (2015) Status and recorded of sharks and rays in the Bay of Bengal of Bangladesh Region. Brazilian Journal of Biological Sciences, 2 (4), 343 - 367."]}

Family Narcinidae One specimen of Narcine was recorded which appears to be consistent with N. sp. of Psomadakis et al. (2020). This appeared to be morphologically similar to N. brevilabiata and N. atzi. Further specimens are required to confirm the species identification therefore this species is tentatively identified as Narcine sp. (Figure 14). (sensu Psomadakis et al. 2020)., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 224, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Psomadakis, P., Thein, H., Russell, B. C. & Tun, M. T. (2020) Field identification guide to the living marine resources of Myanmar. FAO species identification guide for fishery purposes. FAO, Rome, 840 pp."]}

Genus Neotrygon The Neotrygon kuhlii species complex has recently been expanded (Last et al. 2016). These taxonomic revisions, however, did not consider Bay of Bengal specimens in detail. To date, two species have been recorded from Bangladesh; N. indica and N. kuhlii (Haque et al. 2019a; Hoq et al. 2011). In this present study, a total of 649 specimens were examined, and five sampled for genetic analysis. Morphological differences were observed, including the dorsal coloration and tail banding. Several specimens were most consistent with N. caeruleopunctata, but have been preliminarily identified as N. cf. caeruleopunctata (Figure 5) due to possible genetic differences in the Bay of Bengal specimens. It is likely that previous records of N. kuhlii represent this species, with N. kuhlii now considered to occur only from the Solomon Islands, and adjacent island groups (Last et al. 2016a). All genetically sampled specimens were labelled as Neotrygon Bay of Bengal variants., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 218, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Haque, A. B., Das, S. A. & Biswas, A. R. (2019 a) DNA analysis of elasmobranch products originating from Bangladesh reveals unregulated elasmobranch fishery and trade on species of global conservation concern. PloS one, 14 (9), e 0222273. https: // doi. org / 10.1371 / journal. pone. 0222273","Hoq, M. E., Haroon, A. Y. & Hussain, M. G. (2011) Shark fisheries in the Bay of Bengal, Bangladesh: Status and potentialities. Support to Sustainable Management of the BOBLME Project, Bangladesh Fisheries Research Institute, 76 pp.","Last, P., Naylor, G., Seret, B., White, W., de Carvalho, M. & Stehmann, M. (2016 a) Rays of the World. CSIRO publishing, Melbourne, ix + 790 pp."]}

Genus Telatrygon Three specimens of a sharp-snouted stingray were preliminarily identified as Telatrygon crozieri,based on morphology. Samples were not obtained for genetic analysis and so further work is needed to confirm the identification and to distinguish T. crozieri from the recently described T. biasa recorded from Indonesia (Last et al. 2016d)., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 221, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Last, P. R., White, W. T. & Naylor, G. (2016 d) Three new stingrays (Myliobatiformes: Dasyatidae) from the Indo-West Pacific. Zootaxa, 4147 (4), 377 - 402. https: // doi. org / 10.11646 / zootaxa. 4147.4.2"]}

Family Mobulidae Two species of Mobula have previously been recorded from Bangladesh; Mobula mobular and M. kuhlii ( Hoq et al. 2011; Roy et al. 2014a ). In this study, M. birostris, M. tarapacana, M. eregoodoo and M. thurstoni (Figure 13) were recorded following examination of 97 mobulid specimens. Genetic analysis confirmed species identification for M. tarapacana, M. kuhlii and M. mobular, and the presence of either M. alfredi or M. birostris, but the NADH2 sequences alone cannot distinguish between these two species due to overlapping sequence polymorphism (G. Naylor, unpubl. data). This study confirms the presence of at least six mobulid species in Bangladesh; M. alfredi or M. birostris, M. eregoodoo, M. kuhlii, M. mobular, M. tarapacana and M. thurstoni where M. thurstoni has been reported for the first time with possible presence of M. alfredi., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 223, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Hoq, M. E., Haroon, A. Y. & Hussain, M. G. (2011) Shark fisheries in the Bay of Bengal, Bangladesh: Status and potentialities. Support to Sustainable Management of the BOBLME Project, Bangladesh Fisheries Research Institute, 76 pp.","Roy, B. J., Alam, M. F., Rhaman, M. G., Singha, N. K. & Akhtar, A. (2014 a) Landing trends, species composition and percentage composition of Sharks and Rays in Chittagong and Cox's Bazar, Bangladesh. Global Journal of Science Frontier Research, 14, 070403 - 079999."]}

Family Triakidae Three houndshark species have been previously recorded from Bangladesh, i.e. Mustelus manazo, M. mosis and Iago garricki (Hoq et al. 2011; Habib et al. 2020).A total of 31 specimens of a second species of the Iago omanensis complex (which includes the poorly defined I. mangalorensis, if considered valid (Sen et al. 2020; Weigmann 2016; Maduna & Bester-van der Merwe 2017)) were recorded in this study. There is a possibility the recorded Iago sp. could be a new species to science as two separate species (Iago sp. A and Iago sp. B) have been theorised to be present in the Indian waters (Bineesh et al. 2017). This species is tentatively identified as Iago cf. omanensis (Figure 18) pending further taxonomic investigation. It brings the total number of triakid species in Bangladesh to four, with the taxonomy of the Mustelus species in this region being investigated by one of the authors (WW)., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 226, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Hoq, M. E., Haroon, A. Y. & Hussain, M. G. (2011) Shark fisheries in the Bay of Bengal, Bangladesh: Status and potentialities. Support to Sustainable Management of the BOBLME Project, Bangladesh Fisheries Research Institute, 76 pp.","Habib, K. A., Neogi, A. K., Nahar, N., Oh, J., Lee, Y. H. & Kim, C. G. (2020) An overview of fishes of the Sundarbans, Bangladesh and their present conservation status. Journal of Threatened Taxa, 12 (1), 15154 - 15172. https: // doi. org / 10.11609 / jott. 4893.12.1.15154 - 15172","Sen, S., Dash, G., Valappil, A. K., Kizhakudan, S. J. & Chakraborty, R. (2020) Occurrences of Intersexual Hound Sharks, Iago cf. omanensis (Triakidae: Carcharhiniformes) from North-western Bay of Bengal. Thalassas: An International Journal of Marine Sciences, 36, 525 - 534. https: // doi. org / 10.1007 / s 41208 - 020 - 00220 - 0","Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88 (3), 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874","Maduna, S. N. & Bester-van der Merwe, A. E. (2017) Molecular research on the systematically challenging smoothhound shark genus Mustelus: a synthesis of the past 30 years. African Journal of Marine Science, 39 (4), 373 - 387. https: // doi. org / 10.2989 / 1814232 X. 2017.1394365","Bineesh, K. K., Gopalakrishnan, A., Akhilesh, K. V., Sajeela, K. A., Abdussamad, E. M., Pillai, N. G. K., Basheer, V. S., Jena, J. K. & Ward, R. D. (2017) DNA barcoding reveals species composition of sharks and rays in the Indian commercial fishery. Mitochondrial Dna Part A, 28 (4), 458 - 472. https: // doi. org / 10.3109 / 19401736.2015.1137900"]}

Genus Pastinachus The current study confirmed two Pastinachus ater (Figure 7) specimens in Bangladesh and 18 specimens of Pastinachus solocirostris (Figure 8). It is assumed that, P. ater has been previously reported from Bangladesh as P. sephen (Roy et al. 2015), though a recent taxonomic study found that P. sephen is only found in the Western Indian Ocean (Red Sea to Pakistan) (Last & Manjaji-Matsumoto 2010). However, four specimens were morphologically very close to P. sephen and referred to as P. cf. sephen (Figure 9) needing further taxonomic work. We also recorded two specimens of a third species of Pastinachus, which are morphologically consistent with P. gracilicaudus. Further taxonomic examination is required, however, as differences were found between the NADH2 sequences of the Bangladesh as compared to Borneo specimens. This species is tentatively referred to as Pastinachus cf. gracilicaudus (Figure 10)., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 219, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Roy, B. J., Singha, N. K., Rhaman, M. G. & Ali, A. H. M. H. (2015) Status and recorded of sharks and rays in the Bay of Bengal of Bangladesh Region. Brazilian Journal of Biological Sciences, 2 (4), 343 - 367.","Last, P. R. & Manjaji-Matsumoto, B. M. (2010) Description of a new stingray, Pastinachus gracilicaudus sp. nov. (Elasmobranchii: Myliobatiformes), based on material from the Indo - Malay Archipelago. In: Last, P. R., White, W. T. & Pogonoski, J. J. (Eds.), Descriptions of New Sharks and Rays from Borneo. CSIRO Marine and Atmospheric Research Paper 032. CSIRO Marine and Atmospheric Research, Hobart, pp. 115 - 127."]}

Family Rhinobatidae Rhinobatos ranongensis (Figure 15), which was recently described from the Bay of Bengal and Arabian sea (Last et al. 2019), was recorded from> 300 specimens from a single site. With the record of this Rhinobatos species, a total of at least eight species of Rhinopristiformes, including giant guitarfish (Glaucostegidae), guitarfish (Rhinobatidae) and wedgefish (Rhinidae), have been confirmed in the territorial waters of Bangladesh (Haque in review). Although not a new record, one specimen from one site and one genetic sample collected from the processed meat of an unknown species was confirmed to be Rhinobatos lionotus representing the first verified report from Bangladesh., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 225, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Last, P. R., Seret, B. & Naylor, G. J. (2019) Description of Rhinobatos ranongensis sp. nov. (Rhinopristiformes: Rhinobatidae) from the Andaman Sea and Bay of Bengal with a review of its northern Indian Ocean congeners. Zootaxa, 4576 (2), 257 - 287. https: // doi. org / 10.11646 / zootaxa. 4576.2.3"]}

Genus Urogymnus Urogymnus granulatus (Figure 11) was reported for the first time from Bangladesh, based on three specimens from one study site. NADH2 sequence analysis from one of the three specimens supported this identification. This confirms that four species of Urogymnus occur in Bangladeshi waters; U. asperrimus, U. granulatus (likely previously misidentified as U. Africana) ( Hoq et al. 2011; Roy et al. 2015), U. lobistoma and U. polylepis, especially around mangrove areas in the southwestern region. Urogymnus polylepis (Figure 12) has been recorded in news articles but has not yet been added to checklists. Hence, while this is not considered a new record for Bangladesh, it is the first verified record from 52 specimens., Published as part of Haque, Alifa Bintha, White, William T., Cavanagh, Rachel D., Biswas, Aparna Riti & Hossain, Nazia, 2021, New records of elasmobranchs in the Bay of Bengal, Bangladesh: further taxonomic research is essential, pp. 211-230 in Zootaxa 5027 (2) on page 223, DOI: 10.11646/zootaxa.5027.2.4, http://zenodo.org/record/5448230, {"references":["Hoq, M. E., Haroon, A. Y. & Hussain, M. G. (2011) Shark fisheries in the Bay of Bengal, Bangladesh: Status and potentialities. Support to Sustainable Management of the BOBLME Project, Bangladesh Fisheries Research Institute, 76 pp.","Roy, B. J., Singha, N. K., Rhaman, M. G. & Ali, A. H. M. H. (2015) Status and recorded of sharks and rays in the Bay of Bengal of Bangladesh Region. Brazilian Journal of Biological Sciences, 2 (4), 343 - 367."]}