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Basalys villumi sp. nov. urn:lsid:zoobank.org:act: 4204B17E-4659-432C-9266-1C723B434E35 Figs 1F, 5F–H, 6A, Table 1 Diagnosis Body smooth; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical and longer than wide; F1 longest; F2–F11 similar in length; F2 conspicuously emarginate (Fig. 5D–E); scutellum subquadrate without posterior scutellar pits (Fig. 5F); fore wing bordered with small hairs, r-rs short but distinct, M+Cu nebulous (Fig. 1F); hind wing narrow, two thirds of fore wing length, with long hairs along posterior margin; petiole three times as long as wide (Figs 5D, 6A). Etymology We dedicate this species to Villum Fonden, who generously contributed to funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in the genitive case. Type material Holotype NHMD-608360, a complete male; paratype NHMD-608369, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 1.82 mm. Body highly glabrous. Head globular, as long as high (HeL = 0.33 mm), with sparse short hairs; eye round; toruli situated at middle of eye height; antenna 14-segmented; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical, elongate, distinctly longer than wide, with short hairs; F1 longest, F2–F11 subequal in length; F2 emarginate on anterior third; F12 as long as F1; tapering at apex (antennomeres length of holotype, in mm: Sc- 0.22 mm; P-0.09; F1-0.15; F2-0.09; F3-0.11; F4-0.10; F5-0.11; F6-0.10; F7-0.12; F8-0.10; F9-0.11; F10-0.12; F11-0.11; F12-0.14); mandibles weakly crossing at tip, not sickle or beak-shaped; occipital flange foveate, rather concave. MESOSOMA. Shorter than metasoma (MsL = 0.63 mm); pronotum not elongate, anterior part setose, epomia absent; mesoscutum slightly convex, smooth, notauli absent; anterior scutellar pit present, round; scutellum subquadrate, without posterior scutellar pits. Fore wing extending beyond metasoma (FwL = 1.78 mm), bordered with small hairs; Sc+R distinctly separated from anterior margin; marginal vein thickened along wing margin, longer than wide; r-rs short; basal vein curved, perpendicular to Sc+R, almost reaching M+Cu; M+Cu nebulous. Hind wing narrow, two thirds of fore wing length; bordered with long setae along posterior margin. Legs slender, covered with scattered setae; femur clavate; tibia weakly broadened apically; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, narrow, three times as long as wide (PtL = 0.21 mm; PtW = 0.07 mm), longitudinally striated; gaster ellipsoidal (GL = 0.61 mm; GH = 0.30 mm), glabrous; T2 and S2 longest, covering two thirds of gaster, anterior margin T2 straight; segments 3 and 4 ring-like, segment 5 longer than 3 and 4 combined. Female Unknown. Comments Using Nixon’s (1980) and Masner & García’s (2002) keys, Basalys villumi sp. nov. keys out to Basalys Westwood, 1833 because of the following characters: antenna with 14 segments, notauli absent, scutellum with anterior scutellar pit, wings fully developed, Sc+R separated from fore margin of wing, fore wing with distinct basal vein, basal margin of large tergite straight. The description of the species is also consistent with the diagnosis of Hou & Xu (2016). Basalys villumi sp. nov. differs from extant Basalys by having F1 longer than F2 (Nixon 1980; Hou & Xu 2016)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 76-77, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1980. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Diapriinae. Handbooks for the Identification of British Insects 8 (3 di): 1 - 55.","Masner L. & Garcia J. L. 2002. The genera of Diapriinae (Hymenoptera: Diapriidae) in the New World. Bulletin of the American Museum of Natural History 268: 1 - 138. https: // doi. org / c 7 zvsh","Hou Z. & Xu Z. 2016. First record of the genus Basalys Westwood, 1833 (Hymenoptera: Diapriidae) from China, with descriptions of two new species. Zoological Systematics 41 (3): 337 - 341. https: // doi. org / 10.11865 / zs. 201639"]}

Belyta knudhoejgaardi sp. nov. urn:lsid:zoobank.org:act: 618161BD-F876-422B-9C35-62BAD18B2457 Figs 1A, 2A–D, Table 1 Diagnosis Head nasiform; antennal shelf strongly projecting forward; scape as long as head; F1 longest; following flagellomere shorter but gradually lengthening (Fig. 2A); mesopleuron with one prominent longitudinal ridge (Fig. 2B); median propodeal keel simple (Fig. 2C); radial cell open, Rs fading before reaching fore margin; marginal vein hardly longer than r-rs (Fig. 1A); petiole 1.7 times as long as wide; gaster fusiform (Fig. 2B–C). Etymology We dedicate this species to the Knud Højgaard Foundation, who generously contributed to funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in genitive case. Type material Holotype NHMD-608408, a complete female; paratype NHMD-608400, a slightly damaged female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 4.8–5.3 mm. Body slightly pubescent, smooth and shiny. Head nasiform (HeL = 0.77– 0.89 mm), with sparse short hairs posteriorly; eye almond-shaped, glabrous; antennal shelf strongly prominent; antenna 15-segmented, with short heterogeneous setae; scape as long as head; pedicel thinner; F1 one and a half times as long as pedicel or other flagellomeres; F2–F13 gradually lengthening; F13 tapering at apex and slightly shorter than F1 (antennomere length of holotype, in mm: Sc-0.72; P-0.19; F1-0.29; F2-0.11; F3-0.12; F4-0.11; F5-0.12; F6-0.10; F7-0.11; F8-0.13; F9-0.13; F10-0.14; F11-0.15; F12-0.15; F13-0.28); mandibles simple. MESOSOMA. Almost as long as metasoma, flattened (MsL = 1.62–1.90 mm); pronotum with epomia extending from front coxa to posterodorsal pronotal corners; posterior margin of pronotum curved; mesoscutum with several hairs; notauli complete and deeply impressed; scutellum without posterior scutellar pits; mesopleuron with prominent longitudinal keel; propodeum with plicae, median propodeal keel simple. Fore wing hyaline, micro-pubescent, extending beyond metasoma (FwL = 2.74–3.78 mm); C, Sc+R, M+Cu, basal vein, marginal vein and r-rs pigmented; marginal vein shorter than its distance from basal vein; postmarginal vein barely longer than r-rs, pigmented on half length of radial cell; Rs nebulous proximally and pigmented distally from r-rs, projecting distally to postmarginal vein but fading before reaching wing margin; radial cell open. Hind wing hyaline, reduced and narrow (HwL =?– 2.39 mm). Legs slender, with hind femur and coxa enlarged, covered with scattered setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole glabrous, longitudinally striated and with transverse keels, 1.7 times longer (PtL = 0.50 mm) than wide (PtW = 0.30 mm); gaster fusiform, depressed (GL = 1.90–1.95 mm, GH = 0.48– 0.58 mm); S2 and T2 longest, covering two-thirds of gaster, T2 striated at junction with petiole; four ringlike segments distinguishable beyond large tergite, hypopygium bearing several long setae; ovipositor not exerted. Male Unknown. Comments Using Nixon’s (1957) key, Belyta knudhoejgaardi sp. nov. keys out to Belyta because of the following characters: macropterous, notauli present, mandibles of ordinary form, scutellum without a row of foveae along posterior margin, marginal vein shorter than its distance from basal vein, pronotum elongate, without a hollow on each side (= pronotal pits), epomia present, median keel of propodeum simple, petiole more than one and a half time as long as wide. The flattened body is generally accepted to diagnose Belyta spp. (Macek 1996) and is present in Belyta knudhoejgaardi, strengthening the placement of the species in Belyta. Following the key to European species of Belyta of Macek (1996), Belyta knudhoejgaardi sp. nov. keys out between B. subclausa Kieffer, 1916 and B. validicornis Thomson, 1858. It differs from the first by having a marked epomia and from the second by having the median propodeal keel simple (Macek 1996). The diagnostic character for Belyta knudhoejgaardi is the curiously marked longitudinal keel on the mesopleuron, which is absent in other species of Belyta. From the Eocene of Florissant is known B. mortuella Brues, 1910 (attributed to Belyta s. lat.), which differs from B. knudhoejgaardi by being shorter and having a less flattened body (Brues 1910)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 61-62, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Macek J. 1996. Revision of the European species of Belyta Jurine. Acta Musei Nationalis Prague, Series B, Historia Naturalis 51 (1 - 4): 29 - 39.","Brues C. T. 1910. The parasitic Hymenoptera of the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College 54 (1): 3 - 125. Available from https: // www. biodiversitylibrary. org / page / 30208802 [accessed 2 Jan. 2021]."]}

Pantoclis globosa sp. nov. urn:lsid:zoobank.org:act: 4BF21BF6-7925-4F2A-8A84-5D8C49BBD8D7 Figs 1D, 3D–H, Table 1 Diagnosis Eye glabrous; female antenna with: scape 5.5 times as long as wide, pedicel shorter than F1, F1 longest, F2–F13 as long as wide (Fig. 3D–E); male antenna with: scape 5.5 times as long as wide, pedicel rounded, F1 longest, emarginate on anterior third, F2–F12 subequal in length (Fig. 3G–H); mesosoma sparsely pubescent, shorter than metasoma; pronotum not elongate with epomia extending from pronotal shoulders to front coxa; notauli on mesoscutum complete and convergent posteriorly; propodeum with plicae; marginal vein shorter than radial cell and as long as three quarters of its distance to basal vein; radial cell closed; Rs inconspicuous proximally to r-rs (Fig. 1D); petiole as long as wide; gaster globous (Fig. 3F–G). Etymology The species name refers to the globular gaster of the species. The epithet is to be treated as an adjective. Type material Holotype NHMD-608414, a complete female but partially hidden by a milky coat; paratype NHMD- 608394, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 2.83 mm. Head oval, subtriangular in frontal view (HeL = 0.48 mm), bearing several short hairs, stouter on frons; eye oval, glabrous; antenna inserted on frontal shelf, at level with lower margin of eyes; toruli well separated; antenna bearing numerous short sensilla; scape shorter than head length, 5.5 times as long as wide; pedicel shorter than F1, longer than wide; 13 flagellomeres; F1 longest, two times as long as wide; F2–F13 as long as wide; F13 tapering at apex (antennomeres length of female holotype, in mm: Sc-0.44; P-0.14; F1-0.16; F2-0.10; F3-0.10; F4-0.09; F5-0.09; F6-0.09; F7-0.09; F8- 0.08; F9-0.09; F10-0.09; F11- 0.09; F12-0.09; F13-0.10); mandibles of ordinary form; occipital carina foveate. MESOSOMA. Smooth and shiny, shorter than metasoma (MsL = 0.97 mm), sparsely pubescent; pronotum not elongate with distinct and uninterrupted epomia; mesoscutum distinctly convex, notauli complete and convergent; anterior scutellar pit suboval, slightly narrower than apical distance between notauli; scutellum without posterior scutellar pit; propodeum with two prominent plicae; median propodeal keel simple. Fore wing hyaline, homogeneously micropubescent, extending beyond metasoma (FwL = 2.71 mm); C, Sc+R, M+Cu, basal vein, marginal vein, postmarginal vein and r-rs pigmented; Cu nebulous; marginal vein length around three quarters of its distance from basal vein; marginal vein shorter than radial cell; radial cell closed by Rs; Rs pigmented distally to r-rs, inconspicuous proximally. Legs slender, with short hairs; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole twice as long as wide (PtL = 0.27 mm; PtW = 0.12 mm), longitudinally striated, bearing long hairs dorsally; gaster globular, slightly longer than high (GL = 1.05 mm; GH = 0.67 mm), with sparse pubescence; S2 and T2 longest; four ring-like segments discernible beyond large tergite. Male BL = 3.50 mm; similar to female but antennae sexually dimorphic: scape shorter than head length (HeL = 0.53 mm), 5.5 times as long as wide, pedicel rounded, as long as wide, 12 flagellomeres elongate, cylindrical, longer than wide, F1 longest, with marked emargination on anterior third of segment, F2–F12 subequal in length, F12 tapering at apex (antennomeres length of male paratype, in mm: Sc- 0.38 mm; P-0.11; F1-0.25; F2-0.19; F3-0.19; F4-0.19; F5-0.18; F6-0.17; F7-0.16; F8-0.16; F9-0.16; F10-0.16; F11- 0.15; F12; 0.21). Remaining measurements: HeL = 0.53 mm; MsL = 1.24 mm; FwL = 2.84 mm; PtL = 0.32 mm; PtW = 0.15 mm; GL = 1.41 mm; GH = 0.82 mm. Comments Using Nixon’s (1957) key, Pantoclis globosa sp. nov. keys out to Pantoclis because of the following characters: antennae inserted on a frontal prominence, mandibles not forming a beak or sickle-shaped, notauli present, scutellum without foveae along posterior margin, marginal vein nearly as long as its distance from basal vein and presence of ring-like segments beyond the large tergite, marginal vein shorter than radial cell, epomia distinct, median propodeal keel simple, petiole at most one and a third times as long as its apical width. Pantoclis globosa also fits in Pantoclis according to the diagnosis of Hou et al. (2016). It differs from most extant representatives of Pantoclis by having the marginal vein much longer than the r-rs (Nixon 1957; Sharma 1980; Buhl 1998). Pantoclis longiscapa Chambers, 1974 has a similar fore wing configuration but differs by having F1 very long and F2 distinctly longer than the following flagellomeres (Chambers 1974). Pantoclis deperdita Brues, 1906, from the Eocene of Florissant has a shorter radial cell and marginal vein (Brues 1906) and is therefore considered as a different species from P. globosa., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 68-69, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Sharma S. K. 1980. On some new species of Belytinae and Diapriinae (Hymenoptera: Proctotrupoidea) from India. Oriental Insects 14 (1): 51 - 61. https: // doi. org / 10.1080 / 00305316.1980.10434583","Buhl P. N. 1998. New or little known Oriental and Australasian Belytinae (Hymenoptera: Diapriidae). Oriental Insects 32 (1): 41 - 58. https: // doi. org / 10.1080 / 00305316.1998.10433766","Chambers V. H. 1974. Taxonomic notes on the Belytinae, with a new species of Pantoclis Forster (Hym., Proctotrupoidea, Diapriidae). Journal of Entomology (B) 42 (2): 127 - 121. https: // doi. org / 10.1111 / j. 1365 - 3113.1974. tb 00064. x","Brues C. T. 1906. Fossil parasitic and phytophagous Hymenoptera from Florissant, Colorado. Bulletin of the American Museum of Natural History 22: 491 - 498. Available from http: // hdl. handle. net / 2246 / 1719 [accessed 2 Jan. 2021]."]}

Pantolyta chemyrevae sp. nov. urn:lsid:zoobank.org:act: 626E3F04-AB34-45C2-AA5A-82AD0A85CC8E Fig. 5A–C, Table 1 Diagnosis Eye round, extending at most a fourth of head length; scape as long as pedicel and first three flagellomeres combined; F1 two times as long as wide, narrower than pedicel; F2–F13 gradually widening towards apex (Fig. 5A–B); epomia present; anterior scutellar pit suboval, wider than shortest distance between notauli; plicae projecting posteriorly; fore wing not reaching middle of propodeum (brachypterous morph; Fig. 5A–B); petiole slightly longer than wide (Fig. 5C); gaster fusiform, elongate and tapering at apex; T2 striated dorsally at junction with petiole; ovipositor long, subequal in length to gaster (Fig. 5A). Etymology The species is dedicated to Vasilisa Chemyreva, Russian entomologist, in acknowledgment of her numerous publications on Diapriidae from the Palearctic region. The specific epithet is to be treated as a noun in the genitive case. Type material Holotype NHMD-608448, a complete female partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 2.43 mm. Head smooth, bare (HeL = 0.37 mm); eye round, small, less than a third of head length (ED = 0.11 mm); antenna bearing numerous short setae; toruli separated by shallow cleft; scape as long as pedicel and first three flagellomeres combined, at least as long as head; pedicel three times as long as wide; 13 flagellomeres; F1 two times as long as wide, narrower than pedicel; following flagellomeres gradually widening towards apex; F13 largest and longest of all (antennomeres length of holotype, in mm: Sc-0.46; P-0.16; F1-0.12; F2-0.08; F3-0.08; F4-0.08; F5-0.09; F6-0.08; F7-0.08; F8- 0.08; F9-0.08; F10-0.10; F11- 0.10; F12; 0.11; F13-0.19); mandibles simple, just crossing at tip; labial palpi three-segmented, first long, second narrowing at apex and third wider. MESOSOMA. Shorter than metasoma (MsL = 0.75 mm); pronotum not elongate, epomia present; notauli deep and complete, convergent posteriorly; anterior scutellar pit suboval, wider than shortest distance between notauli; mesopleuron bare, with epicnemial pit and carina present; propodeum with plicae projecting posteriorly, forming teeth, median propodeal keel simple. Fore wing reduced, not reaching middle of propodeum (FwL = 0.32 mm). Hind wing shorter (HwL = 0.17 mm). Legs slender with few unorganized erect setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, slightly longer than wide (PtL = 0.20 mm; PtW = 0.18 mm), with distinct longitudinal sculpture, and short hairs dorsally; gaster subcylindrical, elongate and tapering at apex (GL = 1.11; GH = ca 0.40 mm); T2 and S2 longest; T2 slightly striated dorsally at junction with petiole; T6 and T7 not clearly separated and forming long triangle; ovipositor exerted, slightly shorter than gaster (OL = 0.89 mm). Male Unknown. Comments Using Nixon’s (1957) key, Pantolyta chemyrevae sp. nov. keys out to Pantolyta Förster, 1856 because of the following characters: brachypterous, toruli rims without a distinct cleft, antennae 15-segmented, epomia on pronotum defined at pronotal collar, notauli present on mesoscutum, lower side of gaster slightly down curved. With the keys to species of Pantolyta provided in Chemyreva & Kolyada (2019) and Nixon (1957), P. chemyrevae keys out near P. stylata Kieffer, 1908 (= P. vernalis sensu Nixon 1957) but has the antennal shelf less prominent and a longer scape (Chemyreva & Kolyada 2019). Among the Baltic amber species, it differs from other Pantolyta species as follow: P. antiqua is macropterous and has F1 more than two times as long as pedicel; Pantolyta augustinusii has the toruli separated by a distinct cleft, the mesoscutum bearing long hairs and the plicae not projecting posteriorly; P. janzeni and P. perrichoti has a longer petiole, more than two times as long as wide; P. somnulenta is very similar to P. chemyrevae sp. nov. but is macropterous, has anterior scutellar pit narrower than P. chemyrevae and the plicae less produced posteriorly., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 72-73, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Chemyreva V. G. & Kolyada V. A. 2019. Review of the Pantolyta genus (Hymenoptera: Diapriidae: Pantolytini) from Russia, with description of a new species. Zoosystematica Rossica 28: 163 - 176. https: // doi. org / 10.31610 / zsr / 2019.28.1.163"]}

Cinetus elongatus sp. nov. urn:lsid:zoobank.org:act: A7DAC194-D7C5-41B4-B925-6BE4250CB42C Figs 1C, 3A–C, Table 1 Diagnosis Scape longer than head length, F1 longest flagellomere, deeply emarginated on anterior half, wider at apex of emargination (male; Fig. 3A–B); metapleural carina extending from hind coxa and forked anteriorly (Fig. 3C); marginal vein slightly shorter than its distance from basal vein, radial cell elongate, longer than marginal vein, postmarginal vein 1.5 times as long as radial cell (Fig. 1C); petiole 5.3 times as long as wide, with a few long setae (Fig. 3C). Etymology The species name derives from the general elongate aspect of the species. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-608402, a complete male but partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. 4.46 mm. Head higher than long, homogeneously pubescent (HeL = 0.54 mm); eye oval; antenna inserted on distinct shelf; toruli broadly separated; antenna with numerous short hairs; scape longer than head length; pedicel more than two times as short as shortest flagellomere, almost as wide as long; flagellomeres cylindrical, elongate, much longer than wide; F1 longest, with deep emargination basally, wider at apex of emargination; F2–F11 shorter, slightly decreasing in length; F12 longer than F7–F11, conical (antennomeres length of holotype, in mm: Sc-0.61; P-0.12; F1-0.41; F2-0.38; F3-0.36; F4-0.34; F5-0.35; F6-0.33; F7-0.31; F8-0.32; F9-0.29; F10-0.29; F11-0.26; F12-0.33); mandibles of ordinary form, bidentate, slightly crossing at tip. MESOSOMA. Pubescent, with short setae (MsL = 1.52 mm); pronotum reduced in length, epomia present; mesoscutum large, convex, notauli complete, convergent anteriorly, slightly diverging posteriorly but terminating in anterior scutellar pit; anterior scutellar pit deep, large, suboval with posterior margin convex; metapleural carina distinct, forked anteriorly; propodeum with plicae, median propodeal keel simple. Fore wing extending beyond metasoma, homogeneously micro-pubescent (FwL = 3.89 mm); C, Sc+R, basal vein, M+Cu, M, Cu, marginal vein, postmarginal vein, r-rs and Rs fully pigmented; marginal vein slightly shorter than its distance from basal vein; r-rs well developed; Rs closing radial cell in straight line; radial cell elongate, longer than marginal vein; postmarginal vein 1.5 times as long as radial cell. Hind wing shorter and narrower than fore wing; C pigmented; basal cell opened; three hamuli present. Legs slender, bearing numerous un-organized setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole 5.3 times as long as wide (PtL = 0.79 mm; PtW = 0.15 mm), longitudinally ribbed, with sparse very long setae; gaster ellipsoidal (GL = 1.61 mm; GH = 0.68 mm), pointed at apex, with sparse setae on first segments, more numerous at apex; T2 and S2 longest; four ring-like segments posterior to large tergite, genitalia externalized. Female Unknown. Comments Using Nixon’s (1957) key, Cinetus elongatus sp. nov. keys out near Leptorhaptus Förster, 1865 sensu Nixon (1957) because of the following characters: fourth antennal segment not modified, mandibles of ordinary form, scutellum without a row of foveae along posterior margin, marginal vein hardly shorter than the radial cell, petiole much longer than wide, notauli subparallel, terminates in scutellar hollow. Leptorhaptus is currently regarded as a synonym of Cinetus (Masner 1964). Cinetus elongatus displays all the diagnostic characters of Cinetus provided by Quadros & Brandão (2017), especially the large anterior scutellar pit, the posterior extremity of notauli directed toward a point within the anterior scutellar pit, the marginal vein shorter than radial cell and slightly shorter than its distance to basal vein. Among the Baltic amber species, it differs from other Cinetus species as follow: C. balticus has the scape as long as F1 (longer in Cinetus elongatus sp. nov.), the petiole shorter (ratio length/middle height: 4 vs 5.3) and the marginal vein longer than its distance from basal vein; Cinetus breviscapus sp. nov. has the scape shorter than F1, the petiole shorter (ratio length/middle height: 2.8), the marginal vein longer than its distance from basal vein and the postmarginal vein shorter than radial cell; C. inclusus has a shorter petiole (ratio length/middle height: 1.5) and a smaller body (1.9 mm)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 66-68, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Masner L. 1964. A comparison of some Nearctic and Palaearctic genera of Proctotrupoidea (Hymenoptera) with revisional notes. Casopis Ceskoslovenske Spolecnosti Entomologicke 61: 123 - 155.","Quadros A. L. & Brandao C. R. F. 2017. Genera of Belytinae (Hymenoptera: Diapriidae) recorded in the Atlantic dense ombrophilous forest from Paraiba to Santa Catarina, Brazil. Papeis Avulsos de Zoologia 57 (6): 57 - 91. https: // doi. org / 10.11606 / 0031 - 1049.2017.57.06"]}

Doliopria baltica sp. nov. urn:lsid:zoobank.org:act: 36E9B60D-6971-454F-AD9F-DD69D430C16B Figs 1G, 6B–D, Table 1 Diagnosis Head globular, with sparse long hairs on vertex; apical segment of maxillary palpi bearing one long seta (Fig. 6C); antenna 11-segmented; scape as long as pedicel + F1–F5 combined, scape with apical rim slightly notched; pedicel cylindrical, elongate; F1 narrower; F2–F6 widening; F7–F9 forming distinct club; F7–F8 two times as wide as long; F9 as wide as long (Fig. 6B); anterior scutellar pit suboval, not divided; scutellum subquadrate; propodeum punctuate and carinate (Fig. 6D); hind wing bordered with long setae along posterior margin and distal part of fore margin; petiole cylindrical, slightly longer than wide; gaster ellipsoidal; tergite 2 with anterior margin excised (Fig. 6D). Etymology The species name refers to the origin of the amber piece containing the specimen. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-608374, a complete female partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 1.20 mm. Body smooth and shiny, sparsely pubescent, hairs very long. Head globular, slightly higher than long with hairs posteriorly (HeL = 0.23 mm); eye oval, glabrous; maxillary palpi 5-segmented with one long seta on apical segment; antenna 11-segmented inserted on transverse shelf of low elevation; toruli separated by shallow cleft; scape as long as pedicel + F1–F5 combined, as long as head, with apical circumference slightly notched; pedicel as wide as scape, cylindrical; F1 narrower than pedicel and longer than wide; F2–F5 slightly shortening and widening; F6 two times as wide as long; F7–F9 wider than long, longer than previous flagellomeres, forming clava; F9 ovoid (antennomeres length of holotype, in mm: Sc-0.19; P-0.05; F1-0.03; F2-0.02; F3-0.02; F4-0.03; F5-0.03; F6-0.03; F7- 0.07; F8-0.07; F9-0.10); mandibles short, only crossing at tips. MESOSOMA. Slightly shorter than metasoma (MsL = 0.43 mm); pronotum with lateral posterior margin straight; mesoscutum large and slightly convex, without notauli or any other sulci; anterior scutellar pit deep, not divided, suboval; scutellum subquadrate; propodeum minutely punctuate in anterior part and carinate. Fore wing extending beyond metasoma (FwL = 0.77 mm), micropubescent and bordered with short setae; only Sc+R present, slightly separate from anterior margin of wing, distally ending in slight thickening of marginal vein. Hind wing length two thirds of length of fore wing (HwL = 0.55 mm), micropubescent; with long setae along posterior margin and distal part of costal margin; three hamuli present; basal cell open. Legs slender, with only hind coxa and femur stouter; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, narrow, longer than wide (PtL = 0.10 mm; PtW = 0.06 mm), longitudinally striated, bearing several long hairs; gaster ellipsoidal (GL = 0.44 mm; GH = 0.21 mm), not sharply pointed at apex, smooth and glabrous; T2 and S2 longest; T2 covering at least two thirds of gaster, its anterior margin medially excised for distance subequal to petiole length; ovipositor slightly exerted. Male Unknown. Comments In Masner & García’s (2002) keys to the genera of Diapriinae, Doliopria baltica sp. nov. keys out to Doliopria Kieffer, 1910 because of the following characters: frons unarmed, antenna 11-segmented, notauli absent, wings fully developed, Sc+R separate from anterior margin of wing, basal vein absent. The description of the specimen is consistent with the original diagnosis of the genus by Kieffer (1910) and the revised diagnosis of Masner & García (2002). Doliopria baltica differs from extant representatives of this genus as follow: D. americana Fouts, 1926 has a two-segmented club (Fouts 1926); D. brachyptera Ogloblin, 1960 has F9 more lengthened and is brachypterous; D. foersteri Ogloblin, 1960 has short hairs on eyes, flagellomeres longer than wide and lacks the anterior scutellar pit; D. equatoriana Ogloblin, 1960 has F2–F5 longer than wide (Oglobin 1960); D. collegii Ferrière, 1929 has F9 longer (Loiácono et al. 2013); D. flavipes Kieffer, 1910 has F5–6 as long as wide and the petiole shorter (Kieffer 1910); D. myrmecobia Kieffer, 1921 has a two-segmented club and the apical flagellomere sharply pointed at the apex (Loiácono et al. 2013)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 77-79, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Masner L. & Garcia J. L. 2002. The genera of Diapriinae (Hymenoptera: Diapriidae) in the New World. Bulletin of the American Museum of Natural History 268: 1 - 138. https: // doi. org / c 7 zvsh","Kieffer J. J. 1910. Beschreibung neuer sudamerikanischer im Zoologischen Museum zu Berlin aufbewahrter Diapriiden. Entomologische Rundschau 1: 46 - 48. Available from https: // www. biodiversitylibrary. org / page / 43669771 [accessed 2 Jan. 2021].","Fouts R. M. 1926. Notes on Serphoidea with descriptions of new species (Hymenoptera). Proceedings of the Entomological Society of Washington 28 (8): 167 - 179. https: // doi. org / 10.5281 / zenodo. 24228","Ogloblin A. A. 1960. Tres especies nuevas del genero Doliopria del Ecuador (Diapriidae, Hymenoptera). Revista de la Sociedad Entomologica Argentina 22: 69 - 76.","Loiacono M. S., Margaria C. B. & Aquino D. A. 2013. Diapriinae wasps (Hymenoptera: Diaprioidea: Diapriidae) associated with ants (Hymenoptera: Formicidae) in Argentina. Psyche 2013: 320590. https: // doi. org / 10.1155 / 2013 / 320590"]}

Pantolyta similis sp. nov. urn:lsid:zoobank.org:act: 1E2F7650-238A-4AC8-9900-38A6B9DFB59F Fig. 5D–E, Table 1 Diagnosis Head as high as long; eye small but functional; antennal shelf projecting forward and upward; antenna with homogeneous short setae; pedicel conical, with base narrower; F1 second longest; F2–F13 gradually widening; F13 longest and widest (Fig. 5D); epomia absent; median propodeal keel simple (Fig. 5E); fore wing not extending beyond petiole (brachypterous morph); petiole 1.3 times as long as wide; gaster fusiform, elongate; T2 and S2 covering anterior half of gaster; T2 with numerous fine and long striations medially at junction with petiole (Fig. 5D). Etymology The specific epithet refers to its similarity with the extant species Pantolyta macrocera (Thomson, 1858) and is to be treated as an adjective. Type material Holotype NHMD-608468, a complete female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 3.52 mm. Head glabrous, smooth, as high as long (HeL = 0.50 mm); eye round, small, less than a third of head length (ED = 0.18 mm); frontal prominence elongate; toruli facing upwards; antenna slightly but homogeneously pubescent; scape longer than head length; pedicel conical, thinner at base; F1 second longest; F2–F12 of subequal length, gradually widening; F12 as long as wide; F13 widest and longest, conical (antennomeres length, in mm: Sc- 0.58 mm; P-0.11; F1-0.17; F2-0.10; F3-0.10; F4- 0.11; F5-0.11; F6-0.11; F7-0.12; F8-0.11; F9-0.11; F10-0.11; F11-0.11; F12-0.12; F13-0.21); mandibles of ordinary form. MESOSOMA. Shorter than metasoma (MsL = 1.10 mm); pronotum not elongate, with straight posterior margin, epomia absent; mesoscutum convex, with notauli deep and complete; anterior scutellar pit large, rounded; scutellum sub-quadrate, without posterior scutellar pits; propodeum dorsally with prominent plicae, median keel simple. Fore wing very reduced, hardly reaching posterior margin of petiole (FwL = 1.04 mm). Hind wing reduced, shorter than fore wing (HwL = 0.52 mm). Legs slender, with only hind coxa widened; erect setae along hind tibia and tarsomere; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole 1.3 times as long as wide (PtL = 0.28 mm; PtW = 0.22 mm), longitudinally ribbed; gaster fusiform and elongate (GL = 1.64 mm; GH = 0.59 mm); S2 and T2 longest, covering anterior half of gaster; T2 striated medially at junction with petiole; three ring-like segments visible posteriorly to large tergite; apical sternites fused into long, smooth triangle; ovipositor exerted, shorter than gaster (OL = 0.74 mm). Male Unknown Comments The specimen was originally identified as the extant species Pantolyta macrocera (Thomson, 1858) by Buhl (2002) because of the brachypterous morph. However, we can observe some differences on the petiole and gaster. Pantolyta similis sp. nov. displays numerous fine longitudinal ribs dorsally on the petiole whereas there are fewer ribs in P. macrocera. Additionally, P. macrocera has shorter and stronger medial striations on T2 than Pantolyla similis (Chemyreva & Kolyada 2021: fig. 11g). Among the Baltic amber species, it differs from other Pantolyta species as follows: P. antiqua and P. somnulenta are macropterous and have a shorter petiole; Pantolyta augustinusii sp. nov. has the epomia present, whereas it is absent in Pantolyta similis sp. nov.; Pantolyta chemyrevae sp. nov. has the epomia present, the anterior scutellar pit more quadrate, the petiole more elongate and the striations on T2 shorter; P. janzeni and P. perrichoti has a longer petiole, more than two times as long as wide. Subfamily Diapriinae Haliday, 1833 Genus Basalys Westwood, 1833, Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 73-76, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Buhl P. N. 2002. On a Baltic amber collection of Platygastridae and Diapriidae (Hymenoptera). Entomologiske Meddelelser 70: 57 - 61.","Chemyreva V. G. & Kolyada V. A. 2021. Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystemica Rossica 30: 137 - 162. https: // doi. org / 10.31610 / zsr / 2021.30.1.137"]}

Spilomicrus succinalis sp. nov. urn:lsid:zoobank.org:act: C6DACEAE-5CBB-4BC5-A414-FC2458978B3D Figs 1H, 6E–H, Table 1 Diagnosis Head higher than long, minutely punctuate on vertex; scape with distinct punctuations; pedicel pearshaped, narrower at base; F1–F5 as long as wide; F6–F11 forming non-abrupt clava; F11 conical, longer than F10 (Fig. 6E, G–H); occipital flange foveate (Fig. 6F); pronotum laterally with small foveae along posterior margin (Fig. 6E); mesoscutum flat and smooth, with long hairs; notauli convergent, interrupting before posterior margin of mesoscutum; anterior scutellar pit bifoveate; posterior scutellar pits present (Fig. 6F); propodeum carinate; Sc+R pigmented, marginal vein wider than long, r-rs as long as marginal vein (Fig. 1H); first fore tarsomere bearing row of setae along inner margin; first hind tarsomere not longer than following two combined; petiole 1.5 times as long as wide; T2 with anterior margin wrinkled (Fig. 6F); two ring-like segments visible after T2. Etymology The species name derives from ‘Succinite’, a name given to Baltic amber due to its chemical composition. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-607131, a complete female; paratype NHMD-608344, a complete female; paratype NHMD-608354, a complete female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 1.96–2.65 mm. Head higher than long (HeL = 0.29–0.50 mm), with several hairs dorsally; head minutely punctuate on vertex; eye oval, higher than long; antennae inserted on shelf at level with lower half of eye; toruli distinct and slightly protruding; scape with distinct punctuations, 4.2 times as long as wide; pedicel pear-shaped, narrower at base; F1–F5 compact, as long as wide; following flagellomeres widening to form non-abrupt club; F11 wider and longer than F10, conical (antennomeres length of holotype, in mm: Sc-0.25; P-0.09; F1-0.05; F2-0.04; F3-0.04; F4-0.04; F5-0.05; F6-0.06; F7-0.06; F8-0.06; F9-0.08; F10-0.08; F11-0.10); mandibles simple, not forming beak; occipital flange foveate. MESOSOMA. Shorter than metasoma (MsL = 0.67–0.83 mm); pronotum not elongate, not visible in dorsal view, laterally with small foveae on posterior margin; mesoscutum wide, flat and smooth, with several long hairs; notauli present, deep, convergent posteriorly but not reaching posterior margin of mesoscutum; anterior scutellar pit bifoveate; scutellum with posterior scutellar pits present; propodeum carinate, with prominent median keel. Fore wing extending beyond gaster (FwL = 1.27–1.81 mm), hyaline and covered with micropubescence, rounded, bordered with long setae shortening toward wing apex; Sc+R pigmented and distinctly separated from wing margin; marginal vein thickening at apex of Sc+R and almost two times as wide as long; r-rs pigmented and elongate, almost as long as marginal vein. Hind wing half as long as fore wing; only C pigmented. Legs slender, with numerous long setae; first fore tarsomere bearing row of erect setae on inner margin; fore tibial spur long, curved; tibial spur formula 1-2-2; first hind tarsomere not longer than following two combined. METASOMA. Petiole longer than wide (PtL = 0.14–0.21 mm; PtW = 0.10–0.13 mm), covered with longitudinal carinae, glabrous; gaster beyond petiole broadly oval and narrowed at apex (GL = 0.87– 1.11 mm; GH = 0.23–0.43 mm), smooth and with few setae; T2 longest, covering two thirds of gaster, with anterior margin slightly wrinkled, overlapping petiole; S2 longest, as long as T2; segments 3 and 4 ring-like; segment 5 longer than 3 and 4 combined and sharply pointed at apex; ovipositor projecting as short stub. Male Unknown. Comments Using Nixon’s (1980) and Masner & García’s (2002) keys to the genera of Diapriinae, Spilomicrus succinalis sp. nov. clearly keys out to Spilomicrus because of the following characters: antenna with 13 segments, flagellum thickened towards apex, frons unarmed, ocellar triangle in the ocular zone, notauli present, scutellum with an anterior bifoveate pit, wings fully developed, fore wing with pigmented Sc+R, marginal vein not more than one and a half times as long as wide, anterior margin of syntergite straight, without a furrow or a cleft at base. Following the key to the British species of Nixon (1980) and to Eastern Palaearctic species of Chemyreva (2018), S. succinalis sp. nov. keys out near S. comatus Chemyreva, 2015. It differs from this species by having F11 longer than F10, the petiole less than two times as long as wide and lacking the long hairs on the postgena (Chemyreva 2015)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 79-81, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1980. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Diapriinae. Handbooks for the Identification of British Insects 8 (3 di): 1 - 55.","Masner L. & Garcia J. L. 2002. The genera of Diapriinae (Hymenoptera: Diapriidae) in the New World. Bulletin of the American Museum of Natural History 268: 1 - 138. https: // doi. org / c 7 zvsh","Chemyreva V. G. 2018. The Eastern Palaearctic parasitic wasps of the genus Spilomicrus Westwood, 1832 (Hymenoptera: Diapriidae). Far Eastern Entomologist 357: 1 - 20. https: // doi. org / 10.25221 / fee. 357.1","Chemyreva V. G. 2015. New and little known species of the genus Spilomicrus (Hymenoptera: Diapriidae) from the Easter Palaearctic. Zoosystematica Rossica 24 (2): 266 - 278. https: // doi. org / 10.31610 / zsr / 2015.24.2.266"]}

Cinetus breviscapus sp. nov. urn:lsid:zoobank.org:act: CCD7DD88-2504-4697-9B70-80EC819B1C24 Figs 1B, 2E–F, Table 1 Diagnosis Head with sparse long hairs; antenna with uniform pilosity; scape less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres cylindrical, elongate, longer than wide; F1 1.2 times as long as scape, deeply emarginated (Fig. 2F); anterior scutellar pits bean-shaped; propodeum with strong plicae; median propodeal keel simple; marginal vein longer than its distance from basal vein and as long as radial cell; postmarginal vein present on half radial cell; Rs directed toward base of wing; hind wing with basal cell open (Fig. 1B); hind tibia and tarsomeres with long recumbent hairs; petiole 2.8 times as long as high, with longitudinal ribs; gaster globose; T2 covering two thirds of gaster; S2 longer than T2 (Fig. 2E). Etymology The specific epithet is to be treated as an adjective and is composed of ‘ brevi- ’, meaning ‘short’, and ‘- scapus ’, a reference to the small size of the scape. Type material Holotype NHMD-300622, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 3.61 mm. Head with sparse long hairs (HeL = 0.47 mm); eye oval, glabrous; antennae inserted on transverse shelf; 14 antennomeres with uniform pilosity; scape elongate, less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres elongate, cylindrical; F1 1.2 times as long as scape, with a deep emargination covering almost half of segment; F2–F6 slightly shorter, of subequal length; F6–F12 decreasing in size but still much more longer than wide; F12 tapering at apex (antennomeres length, in mm, when possible: Sc-0.32; P-0.06; F1-0.39; F2-0.34; F3-0.32); mandibles of ordinary form, not forming a beak. MESOSOMA. Shorter than metasoma (MsL = 1.21 mm); pronotum without epomia, only visible in lateral view; mesoscutum smooth; notauli deep, diverging at posterior terminations toward a point outside scutellum; anterior scutellar pit bean-shaped; scutellum without posterior scutellar pits; mesopleuron with epicnemial pit, epicnemial bridge and subalar bridge present; propodeum with prominent plicae; median propodeal keel simple. Fore wing hyaline, uniformly micropubescent (FwL= 3.58 mm); venation complete with C, Sc+R, basal vein, M+Cu, marginal vein, r-rs and Rs pigmented; marginal vein very long, longer than its distance from basal vein and as long as radial cell; postmarginal vein fading after middle of radial cell; Rs proximally directed toward base of wing; M inconspicuous towards apex; Cu short and nebulous; radial cell closed. Hind wing narrow, with C pigmented and basal cell opened (HwL = 2.23 mm). Legs slender, covered with setae; tibial spur formula 1-2-2; hind tibia and tarsomeres with long recumbent hairs. METASOMA. Petiole elongate, almost three times as long as its middle height (PtL = 0.56 mm; PtW = 0.19 mm), with longitudinal ribs; gaster globose, oval, smooth (GL = 1.37 mm; GH = 0.71 mm); six segments identifiable; T2 longest, covering two thirds of gaster; other tergites reduced, decreasing in length until they become indistinguishable; S2 longest, longer than T2; following sternites very short; pygidium and hypopygium at least as long as previous segments; genitalia internalized. Female Unknown. Comments Using Nixon’s (1957) key, Cinetus breviscapus sp. nov. keys out to Cinetus Jurine, 1807 because of the following characters: mandibles of ordinary form, scutellum without a row of foveae posteriorly, marginal vein very long, notauli slightly divergent posteriorly. Following Nixon’s (1957) key to male species of Cinetus, Cinetus breviscapus keys out near C. aletes Nixon, 1957 because of its scape that is shorter than F1 but differs in having the first flagellomere less distinctly longer than the scape (ratio scape length/F1 length: 1.2 vs 1.43; Nixon 1957), the petiole longer (three times as long as wide vs “about two and a half times as long as wide”) and being bigger (body length ca 3.6 mm vs ca 2.2 mm). In Baltic amber, Cinetus breviscapus sp. nov. is considered as separate from Cinetus inclusus Maneval, 1938 and Cinetus balticus Szabó & Oehlke, 1986 because it clearly does not fit with their descriptions. The petiole of C. balticus is longer than that of C. breviscapus (ratio length/middle height: 4 vs 2.8) and the fore wing venation is different (C. breviscapus has a longer marginal vein, a longer radial cell, the postmarginal vein does not extend beyond the radial cell and Rs is conspicuous for almost all its length). Cinetus inclusus has a shorter petiole (ratio length/middle height: 1.5) and the scape as long as pedicel, F1 and half of F2 combined, unlike C. breviscapus whose scape is shorter than flagellomere 1.

Pantolyta augustinusii sp. nov. urn:lsid:zoobank.org:act: DD4A2A42-E41A-4731-B91D-AD9EFC205E50 Figs 1E, 4, Table 1 Diagnosis Eye not pubescent; toruli protruding; antenna distinctly pubescent; F1 with indistinct emargination; F1 half of scape length (Fig. 4A–B); mesoscutum bearing long hairs; median propodeal keel simple; fore wing radial cell closed; postmarginal vein extending little beyond radial cell; Rs nebulous proximally to r-rs, marginal vein 0.7 times as long as its distance from basal vein; r-rs distinct (Fig. 1E); petiole 2.4 times as long as wide (Fig. 4D). Etymology We dedicate this species to the Augustinus Foundation, who generously contributed to the funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in genitive case. Type material Holotype NHMD-300829, a complete male; paratypes NHMD-608391, a complete male; NHMD- 608406, a complete male but partially hidden by a milky coat; NHMD-608412, a complete male; NHMD-608404, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 2.51–3.47 mm. Head subtriangular in frontal view (HeL = 0.39–0.48 mm); eye large, not pubescent; several hairs on frons; antennae inserted on distinct but not prominent shelf; toruli well developed, as long as one sixth of scape length from base to apex, separated by distinct cleft; antennae bearing numerous short setae; scape almost six times as long as wide; pedicel slightly longer than wide; flagellomeres elongate, cylindrical; F1 half of scape length, with an indistinct emargination basally; F2–F11 subequal in length; F12 longer and tapering at apex (antennomeres length of holotype, in mm: Sc-0.38; P-0.08; F1-0.21; F2-0.19; F3-0.19; F4-0.19; F5-0.19; F6-0.20; F7-0.19; F8-0.20; F9-0.21; F10- 0.21; F11-0.21; F12-0.23); mandibles short, not forming a beak. MESOSOMA. Shorter than metasoma (MsL = 0.90–1.29 mm); pronotum visible in dorsal view; epomia present as a simple keel; pronotal shoulders rounded; mesoscutum smooth, bearing long hairs; notauli deeply impressed, anteriorly converging towards scutellum, posteriorly parallel just before transscutal sulcus; anterior scutellar pit large, deep; scutellum without posterior scutellar pits; mesopleuron bare, mesopleural pit present; propodeum with median keel simple. Fore wing hyaline, extending far beyond metasoma (FwL = 2.60–3.54 mm), covered with micropubescence; C, Sc+R, M+Cu, basal vein and Cu pigmented; basal and costal cells closed; marginal vein slightly widened, about two thirds as long as its distance from basal vein and twice as long as r-rs; postmarginal vein pigmented, extending slightly beyond radial cell; radial cell closed; Rs tubular distally to r-rs, nebulous proximally; M inconspicuous towards apex. Legs slender, hind femur slightly thicker; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, 2.4 times as long as its middle width (PtL = 0.27–0.37 mm; PtW = 0.11– 0.15 mm), with longitudinal ribs, long hairs laterally and ventrally; gaster fusiform (GL = 0.90–1.33 mm; GH = 0.42–0.58 mm), smooth; T2 longest, narrowing toward petiole; other segments much shorter. Female Unknown. Comments Using Nixon’s (1957) key to the genera of Belytinae, Pantolyta augustinusii sp. nov. keys out near Acropiesta Förster, 1856 because of the following characters: notauli present, second flagellomere not modified; scutellum without foveae; mandibles not forming a beak; marginal vein shorter than both radial cell and its distance from basal vein; pronotum with reduced epomia; large tergite markedly tapering towards petiole. Pantolyta augustinusii especially share with Acropiesta the antennal shelf prominent with toruli separated by a furrow, the 14-segmented antennae with F1 modified, the pronotal shoulders rounded, the notauli complete, the anterior scutellar pit large and quadrate, the propodeum with median keel simple, the fore wing venation with C, Sc+R, M+Cu, Rs, basal and marginal vein tubular, the petiole cylindrical (Macek 1998). However, due to their close similarities, Acropiesta has recently been synonymized under Pantolyta by Chemyreva & Kolyada (2021). Four other Pantolyta species are present in Baltic amber, two having been described under Acropiesta and not duly transferred by Chemyreva & Kolyada (2021): Pantolyta antiqua Buhl, 1999, Pantolyta janzeni (Buhl, 2002) comb. nov., Pantolyta perrichoti (Jouault & Nel, 2020) comb. nov. and Pantolyta somnulenta Maneval, 1938. Pantolyta antiqua has the marginal vein as long as its distance from basal vein whereas it is distinctly shorter in Pantolyta augustinusii (Buhl 1999); P. janzeni has a distinct emargination of F 1 in the male antenna, the marginal vein is shorter and the postmarginal vein is longer than in Pantolyta augustinusii (Buhl 2002); P. perrichoti has the median propodeal keel bifid (Jouault & Nel 2020) and P. somnulenta has the radial cell shorter than the marginal vein (Maneval 1938).

Myrmeleon formicarius(Linnaeus, 1767) The imago of M. formicarius is the largest neuropteran in Denmark with a forewing length of 35���40 mm (Figure 3 (a)). In addition to differing from M. bore in length, the wings are also wider, and M. formicarius can also be identified by its bright and broad spots on the pronotum (Figure 3 (b)), which are clearer and more distinct than those on M. bore (Nielsen, 2015). The dark brown larva has spots on the hind coxa and femora; it is the only antlion species in northern Europe with this trait. Furthermore, the larva is equipped with dolichasters between the base and first tooth of the mandibles. The ventral side of the head is pale, covered by several dark spots, and a pair of significantly larger spots. The larva has four-segmented labial palps. The distal half of the 8th abdominal sternite segment is similar to M. bore, being covered by only short, black spines and without setae. The 3rd instar of the larva is considerably larger than that of the other species in Denmark (Friheden 1973; Badano and Pantaleoni 2014; Nielsen 2015)., Published as part of Petersen, Nicholas H��sig, Nielsen, Ole Fogh & Vilhelmsen, Lars, 2022, Presence of populations of antlions (Neuroptera: Myrmeleontidae) in Denmark correlates with presence of aeolian sand, pp. 2831-2847 in Journal of Natural History 55 (45 - 46) on page 2837, DOI: 10.1080/00222933.2022.2028029, http://zenodo.org/record/6127595, {"references":["Nielsen OF. 2015. Danmarks netvinger. Stenstrup, Denmark: Apollo Booksellers; p. 195.","Friheden J. 1973. Morphological characteristics of North-European myrmeleontid larvae (Neuroptera). Entomol Scand. 4: 30 - 34. doi: 10.1163 / 187631273 X 00039.","Badano D, Pantaleoni RA. 2014. The larvae of European Myrmeleontidae (Neuroptera). Zootaxa. 3762: 1 - 71. doi: 10.11646 / zootaxa. 3762.1.1."]}

Keys to identification of antlions in Denmark Adults: 1. Wings with dark spots (Figures 1 (c) and 2).................................................. Euroleon nostras - Wings without dark spots (Figure 3 (a,c))..................................................................................... 2 2. Pronotum with large, bright spots (Figure 3 (b)). Length of forewings> 35 mm. Eltringsham���s organ in males absent.................................................. Myrmeleon formicarius - Pronotum with small, bright spots. Length of forewings Myrmeleon bore Larvae (Figure references to Friheden 1973 [F73]; Badano and Pantaleoni 2014 [BL14]): 1. Labial palp with three segments (F73, fig. 3). Freshly collected larva (BL14, fig. 29) greyish to pale brown. Ventral side of the head with a pair of median dark spots. No spots on the lower half of the ventral side of the head. Hind legs without dark spots on coxa and femur............................................................................................ Myrmeleon bore - Labial palp with four segments (F73, figs 2, 4, 5, 6)............................................................... 2 2. Mandibles with dolichasters (see below) between base and first tooth (F73, fig. 7). Distal half of 8th abdominal sternite with several short, black spines, without setae (F73, fig. 13; BL14, fig. 6C). Freshly collected larva (BL14, fig. 26) dark brown. Hind legs with two dark spots, one on coxa and one on femur................................................................................................................................ Myrmeleon formicarius - Mandibles without dolichasters between base and first tooth (F73, fig.). The distal half of 8th abdominal sternite with a few black spines and numerous bristles (F73, fig. 15; BL14, fig. 6I). Freshly collected larva reddish brown. Several oval and elongated dark spots on the ventral side of the head, both on the upper and lower half. Hind legs without dark spots on coxa and femur.......................................................... Euroleon nostras, Published as part of Petersen, Nicholas H��sig, Nielsen, Ole Fogh & Vilhelmsen, Lars, 2022, Presence of populations of antlions (Neuroptera: Myrmeleontidae) in Denmark correlates with presence of aeolian sand, pp. 2831-2847 in Journal of Natural History 55 (45 - 46) on pages 2835-2836, DOI: 10.1080/00222933.2022.2028029, http://zenodo.org/record/6127595, {"references":["Friheden J. 1973. Morphological characteristics of North-European myrmeleontid larvae (Neuroptera). Entomol Scand. 4: 30 - 34. doi: 10.1163 / 187631273 X 00039.","Badano D, Pantaleoni RA. 2014. The larvae of European Myrmeleontidae (Neuroptera). Zootaxa. 3762: 1 - 71. doi: 10.11646 / zootaxa. 3762.1.1."]}

Myrmeleon bore(Tjeder, 1941) The imago of M. bore has a forewing length of 25���33 mm (Figure 3 (c)). The wings are narrower than those of M. formicarius, which M. bore is easily confused with. Just like M. formicarius, M. bore also has bright spots on the pronotum, however, they are less distinct and might be completely absent. Adult males of M. bore has a small club-like organ called the Eltringham���s organ at the posterior margin of the hindwings, which helps distributing pheromones (Figure 3 (d)) (Elofsson and L��fqvist 1974; Lipovsek Delakorda et al. 2009; Nielsen 2015; Zhang et al. 2015). The larva is greyish to pale brown, the ventral side of the head is pale and with a pair of dark spots. The labial palp is three-segmented, and the hind legs are without dark spots. The distal half of the 8th abdominal sternite segment is similar to M. formicarius, being covered by only short, black spines and without setae (Friheden 1973; Badano and Pantaleoni 2014)., Published as part of Petersen, Nicholas H��sig, Nielsen, Ole Fogh & Vilhelmsen, Lars, 2022, Presence of populations of antlions (Neuroptera: Myrmeleontidae) in Denmark correlates with presence of aeolian sand, pp. 2831-2847 in Journal of Natural History 55 (45 - 46) on pages 2837-2838, DOI: 10.1080/00222933.2022.2028029, http://zenodo.org/record/6127595, {"references":["Elofsson R, Lofqvist J. 1974. The Eltringham organ and a new thoracic gland: ultrastructure and presumed pheromone function (Insecta, Myrmeleontidae). Zool Scripta. 3: 31 - 40. doi: 10.1111 / j. 1463 - 6409.1974. tb 00801. x.","Lipovsek Delakorda S, Pabst M, Devetak D. 2009. Morphology of the eyes and sensilla in the antlion larvae (Neuroptera: Myrmeleontidae). Life Sci. 2: 401 - 402. doi: 10.3217 / 978 - 3 - 85125 - 062 - 6 - 347.","Nielsen OF. 2015. Danmarks netvinger. Stenstrup, Denmark: Apollo Booksellers; p. 195.","Zhang QH, Zhou G, Hoover DR, Michaelson NJ, Bryant P, Margaryan A, Chauhan KR, Aldrich JR, Schneidmiller RG. 2015. Serendipitous, cross familial discovery of the first long-range chemical attractants for antlions (Neuroptera: Myrmeleontidae): (1 R, 2 S, 5 R, 8 R) - iridodial and Z, E-nepetalactol. Front Ecol Evol. 2: 1 - 7. doi: 10.3389 / fevo. 2014.00080.","Friheden J. 1973. Morphological characteristics of North-European myrmeleontid larvae (Neuroptera). Entomol Scand. 4: 30 - 34. doi: 10.1163 / 187631273 X 00039.","Badano D, Pantaleoni RA. 2014. The larvae of European Myrmeleontidae (Neuroptera). Zootaxa. 3762: 1 - 71. doi: 10.11646 / zootaxa. 3762.1.1."]}

Euroleon nostras(Geoffroy in Fourcroy, 1785) The imago of E. nostras is of the same size as M. bore, with a forewing length of 25��� 33 mm. The presence of several dark spots on the wings of E. nostras makes it easily recognisable, as they are absent from the other Danish antlion species (Figures 1 (c) and 2). E. nostras is without bright spots on the dorsal side of prothorax, and neither the males nor females have Eltringham���s organ (Nielsen 2015). The larva is reddish brown, the ventral side of the head is pale and with several spots, a pair of elongated dark markings near the median, with another pair of dark spots beneath. The labial palp is four-segmented, and the hind legs are without dark spots. The distal half of the 8th abdominal sternite is covered by two types of structures ��� a few black spines and numerous bristles (Friheden 1973; Badano and Pantaleoni 2014)., Published as part of Petersen, Nicholas H��sig, Nielsen, Ole Fogh & Vilhelmsen, Lars, 2022, Presence of populations of antlions (Neuroptera: Myrmeleontidae) in Denmark correlates with presence of aeolian sand, pp. 2831-2847 in Journal of Natural History 55 (45 - 46) on page 2837, DOI: 10.1080/00222933.2022.2028029, http://zenodo.org/record/6127595, {"references":["Nielsen OF. 2015. Danmarks netvinger. Stenstrup, Denmark: Apollo Booksellers; p. 195.","Friheden J. 1973. Morphological characteristics of North-European myrmeleontid larvae (Neuroptera). Entomol Scand. 4: 30 - 34. doi: 10.1163 / 187631273 X 00039.","Badano D, Pantaleoni RA. 2014. The larvae of European Myrmeleontidae (Neuroptera). Zootaxa. 3762: 1 - 71. doi: 10.11646 / zootaxa. 3762.1.1."]}

Genus Agapostemon Guérin-Méneville, 1844 Andrena (Agapostemon) Guérin-Méneville, 1844a: 448. Type species: Apis femoralis Guérin-Méneville, 1844, monobasic., Published as part of Sheffield, Cory S., Vilhelmsen, Lars & Bakker, Frederique, 2021, Taxonomy of the New World bee genus Agapostemon Guérin-Méneville - new names and synonymies (Hymenoptera: Halictidae), pp. 1-23 in European Journal of Taxonomy 751 (1) on page 6, DOI: 10.5852/ejt.2021.751.1375, http://zenodo.org/record/4792347, {"references":["Guerin-Meneville F. E. 1844 a. Iconographie du Regne Animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees de chaque genre d'animaux. Insectes. [1829 - 1838]. J. B. Bailliere, Paris. [text]. https: // doi. org / 10.5962 / bhl. title. 10331"]}

Agapostemon femoralis (Gu��rin-M��neville, 1844) stat. nov. Figs 3, 7D, 8D, 9B, 10B Andrena (Agapostemon) femoralis Gu��rin-M��neville, 1844a: 447 [♂]. Halictus cubensis Spinola, 1851: 203 [♂, not ♀] [synonymy of Ag. viridula by Engel 2004: 170]. Agapostemon semiviridis Cresson, 1865: 172 [♀] [synonymy of Ag. viridulus by Baker 1906: 274]. Agapostemon obscurata Cresson, 1869: 295 [♂, not ♀ as indicated]. syn. nov. Agapostemon obscuratus var. abjectus Cockerell, 1917b: 436 [♂, not ♀ as indicated] [synonymy of Ag. obscuratus by Roberts 1972: 513]. Agapostemon viridualus ��� Janjic & Packer, 2003: 109. Lapsus calami. Material examined Lectotype (designated here, Fig. 3) CUBA ��� ♂; Monchicourt leg.; RMNH. INS.1283531. Additional material CUBA ��� 1 ♀ (lectotype of Agapostemon semiviridis Cresson, 1865, designated by Cresson 1916: 109); ANSP 2788 (Fig. 4) ��� 1 ♂ (lectotype of Agapostemon obscurata Cresson, 1869, designated by Cresson 1916: 108); ANSP 2790 (Fig. 5) ��� 1 ♂ (holotype of Agapostemon obscuratus var. abjectus Cockerell, 1917); Cabanas [Caba��as]; 28 May [no year provided]; Palmer and Riley leg.; USNM 22938 (Fig. 6). Notes The lectotype ♂ of Halictus cubensis Spinola, 1851 (designated by Engel 2004: 170), from Cuba, Havana, Spinola Collection, MSNT, was not examined as the synonymy was not questioned. Remarks The name and description of Andrena femoralis is attributed to Gu��rin-M��neville (1844a), though the name (i.e., Andrena femoralis Guer.), type locality (i.e., Cuba) and first illustration appear on plate 83, figure 1 (incorrectly recorded as plate 73 by Van der Vecht 1957) for Cuvier���s (1836) work published in 1837 (Cuvier 1837). These images were later duplicated in Gu��rin-M��neville (1844b) as ���Insect��s, Pl[ate]. 73, Figure 1 ���. Banks (1909) commented on the dates of Gu��rin-M��neville���s Iconographie du Regne Animali, indicating that the volume dedicated to insects (i.e., Gu��rin-M��neville 1844a) is dated 1829���1838 with many references to other dates within the text, including 1844, the year most researchers have used in citing this work (e.g., Roberts 1972; Michener 2000, 2007, though Moure (1960) indicated 1845). However, as Cuvier���s original plates were published in 1837 (Cuvier 1837), Banks (1909) felt that those of the opinion ���that a named figure is valid without [accompanying] text must credit such names��� with the dates of the plate publication. However, Cowan (1971) more recently reviewed the issue of Gu��rin-M��neville���s works and concluded that the date relevant to the insects is 1844, and this decision is followed here. Van der Vecht (1957) examined material that he assumed was part of the type material for Ag. femoralis at the Rijksmuseum van Natuurlijke Historie, Leiden (now Naturalis Biodiversity Center, Leiden), though he also indicated the possibility of syntypes existing at other institutions; other specimens with the same collection information exist at Naturalis Biodiversity Center. Thus, to stabilize Ag. femoralis as the valid name of this Cuban taxon, the specimen Van der Vecht (1957) provided details on (i.e., fig. 3) is hereby selected as the lectotype. Incidentally, Ag. femoralis is the type species for the genus Agapostemon (Sandhouse 1936, 1943; Michener 1997). Dalla Torre (1896) was the first to treat Ag. femoralis as a synonym of Ag. viridulus (later followed by Friese 1902), though the assumption that the type material of Ag. viridulus was collected in Cuba was baseless. The female type material of Halictus cubensis Spinola was not an Agapostemon, but rather Augochlora regina Smith, 1853 (Augochlorini) (Engel 2004), so Engel (2004) designated the male as a lectotype, and placed it into synonymy with Ag. femoralis. However, as a result of this designation, Ag. cubensis Roberts became a junior secondary homonym of Ag. cubensis (Spinola), thus requiring the replacement name provided below. Though Roberts (1972) records the synonymy of Ag. semiviridis Cresson under Ag. viridulus as new, Baker (1906) had already treated it as a synonym. When described, Cresson (1865) indicated that it was potentially the female of Ag. viridulus., Published as part of Sheffield, Cory S., Vilhelmsen, Lars & Bakker, Frederique, 2021, Taxonomy of the New World bee genus Agapostemon Gu��rin-M��neville - new names and synonymies (Hymenoptera: Halictidae), pp. 1-23 in European Journal of Taxonomy 751 (1) on pages 9-15, DOI: 10.5852/ejt.2021.751.1375, http://zenodo.org/record/4792347, {"references":["Guerin-Meneville F. E. 1844 a. Iconographie du Regne Animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees de chaque genre d'animaux. Insectes. [1829 - 1838]. J. B. Bailliere, Paris. [text]. https: // doi. org / 10.5962 / bhl. title. 10331","Spinola M. 1851. Himenopteros. In: Gay C. (ed.) Historia Fisica y Politica de Chile, Zoologia [volumen 6]: 153 - 569. Maulde et Renou. https: // doi. org / 10.5962 / bhl. title. 16172","Engel M. S. 2004. On the identity of Halictus cubensis Spinola, 1851 (Hymenoptera: Halictidae). Entomological News 115: 169 - 170.","Cresson E. T. 1865. On the Hymenoptera of Cuba. Proceedings of the Entomological Society of Philadelphia 4: 1 - 200.","Baker C. F. 1906. Halictinae de Cuba. In: Earle F. S. (ed.) Primer Informe Anual de la Estacion Central Agronomica de Cuba: 253 - 274. La Universal de Ruiz y Hermano, Havana.","Cresson E. T. 1869. Notes on Cuban Hymenoptera, with descriptions of new species. Transactions of the American Entomological Society 2: 293 - 298. https: // www. jstor. org / stable / 25076211","Cockerell T. D. A. 1917 b. LIV. - Descriptions and records of bees. - LXXVIII. Annals and Magazine of Natural History Series 8 20 (120): 436 - 441. https: // doi. org / 10.1080 / 00222931709487033","Roberts R. B. 1972. Revision of the bee genus Agapostemon (Hymenoptera: Halictidae). The University of Kansas Science Bulletin 49: 437 - 590.","Janjic J. & Packer L. 2003. Phylogeny of the bee genus Agapostemon (Hymenoptera: Halictidae). Systematic Entomology 28: 101 - 123. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00204. x","Cresson E. T. 1916. The Cresson types of Hymenoptera. Memoirs of the American Entomological Society 1: 1 - 141. https: // doi. org / 10.5281 / zenodo. 26375","Van der Vecht J. 1957. On some Hymenoptera from the collection of Guerin-Meneville in the Leiden Museum. Zoologische Mededelingen 35: 21 - 31. Available from https: // repository. naturalis. nl / pub / 317915 [accessed 28 Apr. 2021].","Cuvier G. 1836. The Animal Kingdom, Arranged According to its Organization, Serving as a Foundation for the Natural History of Animals, and an Introduction to Comparative Anatomy. Vol. IV. G. Henderson, London. [text].","Cuvier G. 1837. The Animal Kingdom, Arranged According to its Organization, Serving as a Foundation for the Natural History of Animals, and an Introduction to Comparative Anatomy. Vol. IV. Insecta- Zoophytes. G. Henderson, London. [plates].","Guerin-Meneville F. E. 1844 b. Iconographie du Regne Animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees de chaque genre d'animaux. Tome II. Planches des Animaux invertebres. [1829 - 1844]. J. B. Bailliere, Paris. [plates].","Banks N. 1909. Dates of Guerin's Iconographie du Regne Animali. Entomological News 20: 396 - 397.","Michener C. D. 2000. The Bees of the World. Johns Hopkins University Press, Baltimore.","Michener C. D. 2007. The Bees of the World. Second Edition. Johns Hopkins University Press, Baltimore.","Moure J. S. 1960. Notes on the types of Neotropical bees described by Fabricius (Hymenoptera: Apoidea). Studia Entomologica 3: 97 - 160.","Cowan C. F. 1971. On Guerin's Iconographie: particularly the insects. Journal of the Society for the Bibliography of Natural History 6: 18 - 29. https: // doi. org / 10.3366 / jsbnh. 1971.6.1.18","Sandhouse G. A. 1936. The bees of the genus Agapostemon (Hymenoptera: Apoidea) occurring in the United States. Journal of the Washington Academy of Sciences 26: 70 - 83.","Sandhouse G. A. 1943. The type species of the genera and subgenera of bees. Proceedings of the United States National Museum 92: 519 - 619. https: // doi. org / 10.5479 / si. 00963801.3156.519","Michener C. D. 1997. Genus-group names of bees and supplemental family-group names. Scientific Papers of the Natural History Museum, The University of Kansas 1: 1 - 81. https: // doi. org / 10.5962 / bhl. title. 4052","Dalla Torre C. G. 1896. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 10. Guilelmi Engelmann, Leipzig [Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 10348","Friese H. 1902. Beitrag zur Apidenfauna der grossen Antillen. Zeitschrift fur systematische hymenopterologie und dipterologie 2: 196 - 201.","Fabricius J. C. 1775. Systema Entomologiae. Libraria Kortii, Flensburg and Leipzig [Flensbergi et Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 36510","Smith F. 1853. Catalogue of Hymenopterous Insects in the Collection of the British Museum. British Museum, London. https: // doi. org / 10.5962 / bhl. title. 20858"]}