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The distribution of Carabus hungaricus in Serbia has not been summarized so far. There are only a few published records of this beetle in Serbia. Due to new findings, it was necessary to update the distribution of this species in Serbia. Recent field research revealed that the largest suitable habitat is the Deliblato Sands in the South Banat District, where this species is common. Other important areas are located near the Romanian border in the vicinity of the city of Vršac and near Mužlja (in the neighborhood of the city of Zrenjanin, Central Banat District), where the northernmost population in Serbia is located. In addition to this overview of distribution, we propose monitoring methodology for C. hungaricus. We believe that regular monitoring can bring the necessary attention to the disappearing steppe habitats and that the results can help to manage them appropriately in the future.

The fig bark beetle (Hypoborus ficus Erichson, 1836) was recorded under the bark and in the wood of frost-damaged top branches of fig (Ficus carica L.) in two locations in Vrčin, in the vicinity of Belgrade, as well as in the "Jevremovac" Botanical Garden in Belgrade. Numerous specimens were found, and a total of 39 adults, as well as several larvae and pupae, were collected. The findings presented here are new data on the presence of the mentioned species of bark beetle on the territory of Serbia (the first precise data), after more than a century. This is another Mediterranean species that has been confirmed to have established populations in Serbia., {"references":["Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gültekin, L., Hlaváč, P., Korotyaev, B., Lyal, H. C. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sánchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhrovec, J., Trýzna, M., Velázquez de Castro, A. J., & Yunakov, N. N. (2017). Cooperative catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electrónicas SEA, 8. Amini, S., Nozari, J., Martinez, I., Hosseini, R., & Faccoli, M. (2020). Morphological and molecular identification of the Iranian bark and ambrosia beetles (Coleoptera, Curculionidae, Scolytinae). Zootaxa, 4852(3), 251-284. Avidov, Z., & Harpaz, I. (1969). Plant pests of Israel. Jerusalem: Israel Universities Press. 549 pp. Ćetković, A., Čubrilović, B., Plećaš, M., Popović, A., Savić, D., & Stanisavljević, L. (2012). First records of the invasive American wasp Isodontia mexicana (Hymenoptera: Sphecidae) in Serbia. Acta Entomologica Serbica, 17(1/2), 63-72. Fontana, P. (1925). Contribuzione alla fauna coleotterologica ticinese. Bolletino della Società Ticinese di Scienze Naturali, 20, 23-34. Gojšina, V., Páll-Gergely, B., Vujić, M., & Dedov, I. (2022). First record of the genus Xeropicta Monterosato, 1892 (Gastropoda: Eupulmonata: Geomitridae) in Serbia. Folia Malacologica, 30(1), 47-53. Hypoborus ficus Erichson, 1836 in GBIF Secretariat (2021). GBIF Backbone Taxonomy. Checklist dataset Retrieved from: https://doi.org/10.15468/39omei [Accssed on: 21.11.2022]. Isayeva, K. B. (2021). Bioecological peculiarities and economic significance of large fruit bark beetle (Scolytus mali Bechstein, 1805) and fig bark beetle (Hypoborus ficus Erichson, 1836) in the southern slopes of the Great Caucasus (Azerbaijan). Studia Biologica, 15(1), 61-66. Knížek, M. (2011). Scolytinae. In Löbl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol 7. Apollo Books, Stenstrup, Denmark. 373 pp. Langhoffer, A. (1915). Podkornjaci Hrvatske (Scolytidae Croatiae). Šumarski list, 3/4, 53-75. Marković, Č., & Stojanović, A. (1997). Prilog proučavanju faune sipaca (Coleoptera, Scolytidae) u Srbiji. Šumarstvo, 1, 40-51. Mifsud, D., & Knížek, M. (2009). The Bark Beetles (Coleoptera: Scolytidae) of the Maltese Islands (Central Mediterranean). Bulletin of the Entomological Society of Malta, 2, 25-52. Musolin, D. L., & Fujisaki, K. (2006). Chenges in ranges: trends in distribution of true bugs (Heteroptera) under conditions of the current climate warming. Russian Entomological Journal, 15(2), 175-179. Nadaždin, B., & Šeat, J. (2022). New data on Perillus bioculatus (Heteroptera: Pentatomidae) in Serbia: Do climate change and a new food source contribute to the true bug expansion? Acta Entomologica Serbica, 27(2), xx-xx. Sarikaya, O. (2013). Notes on bark and wood-boring beetles (Coleoptera: Bostrichidae; Curculionidae; Platypodinae and Scolytinae) of the sweetgum (Liquidambar orientalis Mill.) Forest Nature Protection Area, with a new record for Turkish fauna. Journal of Food Agriculture and Environment, 11, 2178-2185. Šeat, J., Nadaždin, B., & Šćiban, M. (2020). Acrosternum heegeri (Heteroptera: Pentatomidae) in Serbia. Acta Entomologica Serbica, 25(1), 77-81. Šeat, J., Vujić, M., & Nadaždin, B. (2019). New faunal data on true bugs (Heteroptera) in Serbia. Acta Entomologica Serbica, 24(1), 95-99. Skendžić, S., Zovko, M., Pajač Živković, I., Lešić, V., & Lemić, D. (2021). Effects of climate change on introduced and native agricultural invasive insect pests in Europe. Insects 12, 985. Stojanović, D. V., Jerinić-Prodanović, D., Kereši, T., Graora, D., & Marković, M. (2020). Choreutis nemorana (Hübner, 1799) (Lepidoptera: Choreutidae) in Serbia. Topola/Poplar, 206, 29-34. Tuncer, C., Knížek, M., & Hulcr, J. (2017). Scolytinae in hazelnut orchards of Turkey: clarification of species and identification key (Coleoptera, Curculionidae). ZooKeys, 710, 65-76. Vit, S., Alonso-Zaragaza, M. A., & Kníñek, M. (2022). Fauna Europaea: Scolytinae. Fauna Europaea version 2017.06 https://fauna-eu.org. Vujić, M., Đurić, M., & Tot, I. (2021a). Six new hoverfly species (Diptera: Syrphidae) in the fauna of Serbia. Kragujevac Journal of Science, 43, 149-155. Vujić, M., Ivković, S., Rekecki, T., Krstić, D., Stanković, V., Đurić, M., & Tot, I. (2021b). A first record of the alien mantis species Hierodula tenuidentata (Mantodea: Mantidae) in Serbia. Acta Entomologica Serbica, 26(1), 1-7. Witkowski, R., & Mazur, A. (2019). First record of fig bark beetle Hypoborus ficus Erichson (Curculionidae: Scolytinae) from Poland. Zootaxa, 4571(1), 143-144."]}

Key to the taxa of the genus Proleonhardella Jeannel, 1910 (modified after Jeannel 1924) (Figs 9���10) 1. Body more elongate, elliptical. Pronotum as wide as elytra, well constricted basally. Pronotal lateral margins well-rounded backwards. Mesosternal carina very low. Elytral punctuation rough and deep (subgenus Pholeuonillus Breit, 1913). Endogean, Mt Treskavica, southern Bosnia and Herzegovina....................................................................... P. (Pholeuonillus) adolfi (Reitter, 1911) ��� Body less elongate, oval. Pronotum narrower than elytra, weakly constricted basally. Pronotal lateral margins weakly rounded backwards. Elytral punctuation fine. Mesosternal carina elevated, angled (subgenus Proleonhardella Jeannel, 1910)....................................................................................... 2 2. Elytra shorter, more convex, less than twice as long as pronotum (Fig. 10A)................................. 3 ��� Elytra longer, less convex, more than twice as long as pronotum (Fig. 10B).................................. 6 3. Body of oval shape, longer (TL 1.8���2.2 mm). Antennae short, not reaching middle of body. Antennomere III not longer than antennomere V. Antennomere VIII globular in males (Fig. 10C). Cavernicolous, Mts Bjela��nica and Igman, central Bosnia and Herzegovina [P. (Proleonhardella) matzenaueri (Apfelbeck, 1907)]....................................................................................................... 4 ��� Body of subglobular shape, shorter (TL less than 1.8 mm). Antennae long, reaching middle of body. Antennomere III longer than antennomere V. Antennomere VIII slightly elongate in males (Fig. 10D)......................................................................................................................................... 5 4. Body of almost regular oval shape, longer (TL 2.0��� 2.2 mm). Elytra not widened towards middle. Cavernicolous, Mt Bjela��nica, central Bosnia and Herzegovina......................................................................................................... P. (Proleonhardella) matzenaueri matzenaueri (Apfelbeck, 1907) ��� Body narrower anteriorly, shorter (TL 1.8���2.0 mm). Elytra clearly widened medially. Cavernicolous, Mt Igman, central Bosnia and Herzegovina.......................................................................................................................................................... P. (Proleonhardella) matzenaueri ottonis M��ller, 1917 5. Body more elongate and convex, longer (TL 1.5 mm). Elytral punctuation finer and denser.Antennae shorter, with apical antennomeres thicker. Antennomere VIII barely longer than wide, antennomere IX as long as wide and antennomere X transverse in females. Cavernicolous, Mt Treskavica, southern Bosnia and Herzegovina............................................ P. (Proleonhardella) leonhardi (Breit, 1913) ��� Body less elongate and convex, shorter (TL 1.3 mm). Elytral punctuation stronger and less dense. Antennae longer, with apical antennomeres less thick. Antennomeres VIII, IX and X longer than wide in females. Cavernicolous, village of Trnovo, vicinity of the city of Sarajevo, central Bosnia and Herzegovina...................................................... P. (Proleonhardella) apfelbecki Jeannel, 1924 6. Body longer (TL 3.0��� 3.5 mm). Cavernicolous, Kamena Gora and vicinity of the towns of Prijepolje, Priboj and Pljevlja, southwestern Serbia and northern Montenegro............................................................................................................................................ P. (Proleonhardella) remyi Jeannel, 1934 ��� Body shorter (TL less than 2.435 mm)............................................................................................. 7 7. Elytra parallel in basal half (Fig. 10E).............................................................................................. 8 ��� Elytra narrowed in basal half (Fig. 10F)........................................................................................... 9 8. Pubescence long. Punctuation less fine. Apical antennomeres more widened. Pronotum less rounded. Apex of median lobe of aedeagus less bent ventrally. Elytral apex wide, obtuse and oblique. Cavernicolous, vicinity of the towns of Prijepolje and Priboj (southwestern Serbia) and Pljevlja (northern Montenegro).................................................. P. (Proleonhardella) hirtella Jeannel, 1934 ��� Pubescence short. Punctuation finer. Apical antennomeres less widened. Pronotum more rounded. Apex of median lobe of aedeagus more bent ventrally. Elytral apex attenuated. Cavernicolous, Mt Sjemeć, eastern Bosnia and Herzegovina......... P. (Proleonhardella) weiratheri (Reitter, 1913) 9. A1/A2 M 0.80. Mesosternal carina almost right-angled. Elytral lateral margins below humeral angles clearly sinuate. Maximum width of elytra prior to middle. Cavernicolous, village of Podromanija, near the town of Sokolac, eastern Bosnia and Herzegovina......................................................................................................................................... P. (Proleonhardella) neumanni (Apfelbeck, 1901) ��� A1/A2 M 0.76. Mesosternal carina obtuse-angled. Elytral lateral margins below humeral angles sub-parallel. Maximum width of elytra slightly after basal third. Cavernicolous, Mt Tara, western Serbia.................................................... P. (Proleonhardella) tarensis Ćurčić & Pavićević sp. nov., Published as part of ��ur��i��, Sre��ko, Pavi��evi��, Dragan, Vesovi��, Nikola, Vrbica, Maja, Kuraica, Milo��, Markovi��, ��or��e, Petkovi��, Matija, Lazovi��, Vladimir, Panteli��, Dejan & Bosco, Fabrizio, 2021, On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia, pp. 55-81 in European Journal of Taxonomy 782 (1) on pages 75-76, DOI: 10.5852/ejt.2021.782.1589, http://zenodo.org/record/5761502, {"references":["Jeannel R. 1910. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale 45 (1): 1 - 48.","Jeannel R. 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436.","Breit J. 1913. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Beitrage zur Blindkaferfauna von Bosnien und der Herzegowina. Entomologische Mitteilungen 2 (11): 351 - 358.","Jeannel R. 1934. Bathysciinae recueillis par M. M. Remy et R. Husson dans le Sandjak de Novi-Pazar et la Macedoine grecque. Revue francaise d'entomologie 1: 89 - 103.","Reitter E. 1913. Sechs neue von Herrn Weirather in Bosnien entdeckte Hohlensilphiden. Koleopterologische Rundschau 10: 153 - 157.","Apfelbeck V. 1901. Drei neue Hohlenkafer aus Bosnien. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 14 - 16."]}

Key to the leptodirine leiodid genera of the phyletic series of ��� Leonhardella ��� (modified after Gu��orguiev 1976) (Fig. 1) 1. Body of bathyscioid form, oval or ovoid, wide and short................................................................ 2 ��� Body of elliptical or pholeuonoid form............................................................................................ 6 2. Mesosternal carina absent (Fig. 1A). Cavernicolous, Mt Čvrsnica, southwestern Bosnia and Herzegovina.............................................................................................. Augustia Zariquiey, 1927 ��� Mesosternal carina present (Fig. 1B)................................................................................................ 3 3. Antennae short, not reaching middle of body................................................................................... 4 ��� Antennae much longer, reaching middle of body............................................................................. 5 4. Pubescence fine and recumbent, except in both lateral exterior border of elytra and apical elytral half, where long hairs occur. Antennae very short, barely exceeding pronotal base. Longer species (TL 2.5 mm). Mesosternal carina rounded. Basal lamina of tegmen of aedeagus without tooth. Cavernicolous, region of Kuči, vicinity of the city of Podgorica, eastern Montenegro..................................................................................................................................... Weiratheria Zariquiey, 1927 ��� Pubescence entirely normal and recumbent. Antennae somewhat longer, but not reaching middle of body. Shorter species (TL 1.85 mm). Mesosternal carina triangular. Basal lamina of tegmen of aedeagus with a pronounced tooth basally. Cavernicolous, vicinity of the town of Karystos, island of Euboea, southern Greece.......................................................................... Henrotiella Perreau, 1999 5. Pubescence short and recumbent. Anterior border of mesosternal carina without concavity (Fig. 1C). Protarsi weakly dilated in males. Shorter species (TL 1.3���3.5 mm). Cavernicolous and endogean, central, eastern and southern Bosnia and Herzegovina, southwestern and western Serbia and eastern Montenegro...................................................... Proleonhardella Jeannel, 1910 ��� Pubescence long and erect. Anterior border of mesosternal carina with a deep concavity (Fig. 1D). Protarsi not dilated in males. Longer species (TL 4.0��� 5.5 mm). Cavernicolous, western, southwestern and eastern Montenegro, southern Bosnia and Herzegovina and southern Croatia............................................................................................................................................. Blattochaeta Reitter, 1910 6. Body of elliptical form (Fig. 1E). Pronotum regularly or almost regularly arcuate, slightly narrower than elytra or as wide as elytra......................................................................................................... 7 ��� Body of pholeuonoid form (Fig. 1F). Pronotum campanuliform or strongly sinuated backwards, clearly narrower than elytra.............................................................................................................11 7. Body shorter (TL 1.8���2.0 mm). Antennae almost reaching middle of body. Elytra very attenuated apically, with sparse pubescence. Ventral border of mesosternal carina triangular and deeply grooved. Endogean and cavernicolous, southern Croatia and western Bosnia and Herzegovina....................................................................................................................................... Anisoscapha M��ller, 1917 ��� Body longer (TL 2.5���5.2 mm). Antennae reaching middle of body. Elytra regularly arcuate apically, with dense pubescence. Ventral border of mesosternal carina not grooved..................................... 8 8. Body shorter (TL 2.5���3.0 mm). Mesosternal carina not atrophied. Protarsi dilated in males (Fig. 1G)........................................................................................................................................... 9 ��� Body longer (TL 3.8���5.2 mm). Mesosternal carina atrophied posteriorly. Protarsi not dilated in males (Fig. 1H)............................................................................................................................... 10 9. Body very elongate and narrower. Pubescence long and erect. Median lobe more elongate, thin. Basal bulb small, with a short rounded basal projection (Fig. 1I). Paramerae distally widened. Cavernicolous, southern and southeastern Bosnia and Herzegovina and western Montenegro........................................................................................................................... Anillocharis Reitter, 1903 ��� Body less elongate and wider. Pubescence short and recumbent. Median lobe less elongate, wide. Basal bulb elongate, with a long sub-triangular basal projection (Fig. 1J). Paramerae distally narrow. Endogean and cavernicolous, Mts Bobija and Povlen, western Serbia........................................................................................................................................... Bozidaria Ćurčić & Pavićević gen. nov. 10. Body shorter (TL 3.8���4.6 mm). Pronotum as wide as elytra. Lateral pronotal margins regularly arcuate. Anterior border of mesosternal carina with no concavity. Paramerae with three setae. Endogean, central and southern Bosnia and Herzegovina.................... Pholeuodromus Breit, 1913 ��� Body longer (TL 5.0��� 5.2 mm). Pronotum slightly narrower than elytra. Lateral pronotal margins weakly sinuate in basal third. Anterior border of mesosternal carina with a deep concavity. Paramerae with four setae. Cavernicolous, southeastern Bosnia and Herzegovina and northern Montenegro................................................................................................................... Blattodromus Reitter, 1904 11. Antennomere VIII subglobular (Fig. 1K). Elytral punctuation strong and deep. Pubescence long, usually double, with bristle-like setae. Cavernicolous and endogean, western Serbia, southern, southeastern, central, western and eastern Bosnia and Herzegovina and northwestern Montenegro......................................................................................................... Pholeuonopsis Apfelbeck, 1901 ��� Antennomere VIII elongate (Fig. 1L). Elytral punctuation fine and shallow. Pubescence short, with no bristle-like setae......................................................................................................................... 12 12. Body shorter (TL 2.0 mm). Pronotum elongate, very narrow basally. Mesosternal carina low. Paramerae with two setae (Fig. 1M). Cavernicolous, vicinity of the town of Ključ, western Bosnia and Herzegovina................................................................................. Deelemaniella Perreau, 2002 ��� Body longer (TL 3.2���4.5 mm). Pronotum as long as wide or transverse, wider basally. Mesosternal carina high or atrophied. Paramerae with three setae (Fig. 1N)..................................................... 13 13. Mesosternal carina high. Protarsi somewhat dilated in males. Tibiae with no external spur. Cavernicolous, northwestern, western and northern Montenegro and southeastern and southern Bosnia and Herzegovina......................................................................... Leonhardella Reitter, 1903 ��� Mesosternal carina atrophied. Protarsi not dilated in males. Tibiae with external spur. Cavernicolous, Mt Durmitor, northern Montenegro................................. Tartariella Nonveiller & Pavićević, 1999, Published as part of ��ur��i��, Sre��ko, Pavi��evi��, Dragan, Vesovi��, Nikola, Vrbica, Maja, Kuraica, Milo��, Markovi��, ��or��e, Petkovi��, Matija, Lazovi��, Vladimir, Panteli��, Dejan & Bosco, Fabrizio, 2021, On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia, pp. 55-81 in European Journal of Taxonomy 782 (1) on pages 60-63, DOI: 10.5852/ejt.2021.782.1589, http://zenodo.org/record/5761502, {"references":["Gueorguiev V. B. 1976. Recherches sur la taxonomie, la classification et la phylogenie des Bathysciinae (Coleoptera Catopidae). Dissertationes Academiae Scientiarum et Artium Slovenicae, Ljubljana 19 (4): 91 - 147.","Zariquiey R. 1927. Nuevos Bathysciinae de los Balcanes (Col. Silphidae). Butlleti de l'Institucio Catalana d'Historia Natural 27: 152 - 158.","Perreau M. 1999. Nouveaux genres et nouvelles especes de Leptodirini (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104 (4): 399 - 406.","Jeannel R. 1910. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale 45 (1): 1 - 48.","Reitter E. 1910. Eine neue Bathyscia aus Dalmatien. Wiener entomologische Zeitung 29 (5 - 6): 164. https: // www. biodiversitylibrary. org / page / 12086245","Breit J. 1913. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Beitrage zur Blindkaferfauna von Bosnien und der Herzegowina. Entomologische Mitteilungen 2 (11): 351 - 358.","Jeannel R. 1911. Revision des Bathysciinae (Coleopteres Silphides): Morphologie, Distribution Geographique, Systematique. Libraire A. Schulz, Paris, France. https: // doi. org / 10.5962 / bhl. title. 63151","Jeannel R. 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436.","Curcic S. B., Schonmann H., Brajkovic M. M. & Curcic B. P. M. 2008 a. Serboleonhardella gen. n., based on Proleonhardella remyi Jeannel (Leptodirini, Leiodidae, Coleoptera), from a cave in southwestern Serbia. In: Makarov S. E. & Dimitrijevic R. N. (eds) Advances in Arachnology and Developmental Biology. Papers Dedicated to Prof. Dr. Bozidar Curcic: 463 - 473. Institute of Zoology, Faculty of Biology, University of Belgrade, Committee for Karst and Speleology, Serbian Academy of Sciences and Arts, Institute of Zoology, Bulgarian Academy of Sciences, Faculty of Life Sciences, University of Vienna & UNESCO MAB Committee of Serbia, Vienna-Belgrade-Sofia, Austria-Serbia-Bulgaria.","Apfelbeck V. 1901. Drei neue Hohlenkafer aus Bosnien. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 14 - 16.","Nonveiller G. & Pavicevic D. 1999. Tartariella durmitorensis n. gen. et n. sp., troisieme Leptodirini de la chaine Dinarique a moeurs hadesiennes (Coleoptera, Leiodidae). Bulletin de la Societe entomologique de France 104 (4): 307 - 318."]}

Bozidaria serbooccidentalis Ćurčić & Pavićević gen. et sp. nov. urn:lsid:zoobank.org:act: A2B51421-4CD9-4696-BF46-7EACBC395C4E Figs 2���3 Diagnosis The genus is currently monotypic and therefore a differential diagnosis for Bozidaria serbooccidentalis gen. et sp. nov. cannot be provided. Etymology The species is named after western Serbia, where its type locality and known localities are situated. Type material Holotype SERBIA ��� ♂; western Serbia, town of Ljubovija, Mt Bobija; alt. 1000 m; 19 Apr. 1980; Guido Nonveiller leg.; traps for endogean fauna baited with rotten meat; IZFB-21/1. Paratypes SERBIA ��� 1 ♂, 2 ♀♀; same collection data as for holotype; SBS- 21/1 to 21/3 ��� 1 ♀; same collection data as for holotype; IZFB-21/2 ��� 2 ♂♂, 2 ♀♀; same collection data as for holotype; CDP- 21/1 to 21/4 ��� 10 ♂♂, 14 ♀♀; western Serbia, town of Ljubovija, Mt Povlen, Debelo Brdo saddle, village of Gornje Ko��lje, Simina Jama Pit; 44��08���32.2��� N, 19��37���40.4��� E; 20 May���5 Nov. 2017; Milo�� Kuraica leg.; pitfall trapping; IZFB-21/3 to 21/26 ��� 2 ♂♂, 1 ♀; same locality as for preceding; 31 Dec. 2010; Iva Njunjić leg.; pitfall trapping; SBS- 21/4 to 21/6 ��� 18 ♂♂, 39 ♀♀; same collection data as for preceding; CDP-21/5 to 21/61 (Fig. 2). Description HABITUS. Body elliptical, TL R 2.51���2.80 mm (R 2.51���2.73 mm in males, R 2.67���2.80 mm in females), reddish-brown in colour, shiny, pubescent, with a fine punctuation (Fig. 2A). HEAD. Short, wide, slightly wider than long (HL/HW M 0.97), without eyes (Fig. 2A). Antennae long and slender, apically gradually widened and flattened, ending slightly after basal third of elytra in males or slightly prior to basal third of elytra in females. Antennomere II longer than antennomere I (A1/A2 M 0.795). A3/A2 M 0.67. A3/A5 M 1.24. Antennomeres IV���VI of similar length, of which IV narrowest and VI widest. Antennomere VII apically widened. A7/A6 M 1.56. Antennomere VIII slightly longer than half of antennomere VII (A7/A8 M 1.53), oval, somewhat elongate (A8LW M 1.52). Antennomere IX somewhat elongate, gradually widened distally (A9LW M 1.47). A9/A8 M 1.52 in males, M 1.42 in females. Antennomere X slightly longer than wide (A10LW M 1.24), more widened apically. Antennomere XI slender, ovoid, apically pointed, more elongate in males (A11LW M 2.45) than in females (A11LW M 1.97), as long as preceding two antennomeres combined or slightly shorter than the latter. Occipital carina present. Hairs yellow, erect. Microsculpture composed of small isodiametric meshes. THORAX. Pronotum transverse, widest sub-basally, almost twice as wide as long (PL/PW M 0.59) (Fig.2A). Lateral pronotal margins arcuate, rounded medially, sub-parallel prior to hind pronotal angles. Pronotal base more than twice as long as anterior pronotal margin (PB/AM M 2.13). Both anterior pronotal margin and pronotal base convex medially, the latter less pronouncedly. PL+EL/AL M 1.63 in males, M 1.89 in females. Fore angles prominent, obtuse, rounded, hind angles sharp, rounded, prominent, directed backwards. Microsculpture of pronotum composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 2B). Pronotal disc weakly convex. Mesosternal carina high, obtuse-angled, anterior margin strongly convex, posterior margin barely convex, almost straight, with hairs and teeth (Fig. 2C). Mesosternal carina with an apical tooth. Ventral border of mesosternal carina not grooved. ELYTRA. Elongate (EL/EW M 1.38 in males, M 1.40 in females), more than 2.5 times as long as pronotum (EL/PL M 2.65), obovoid, sub-parallel below humeral angles, weakly narrowed basally, rounded medially, narrowed apically (Fig. 2A). Apex rounded. Sutural striae absent. Scutellum small, triangular. Elytra widest between basal third and mid-length. Microsculpture composed of large isodiametric meshes. Hairs yellow, recumbent (Fig. 2D). Elytral disc convex. Pygidium not completely covered by elytra. LEGS. Elongate and thin, with hairs (Fig. 2A). Tibiae with a few spines laterally.Anterior tarsi tetramerous in both genders, somewhat dilated in males (P1LW M 1.825 in males, M 1.88 in females). ABDOMEN. Median lobe of aedeagus elongate, thin, sub-parallel, sub-apically rounded (Figs 2E, 3A). Apex elongate, triangular. Basal bulb relatively narrow, elongate, with a sub-triangular basal projection. Copulatory piece weakly chitinised, consisting of a basal phanera, median paired stripes and apical paired sclerotizations. Median lobe proximally straight, distally relatively curved, gradually narrowed distally in lateral view (Fig. 3B). Basal bulb narrow in lateral view. Parameres slender, thin, longer than median lobe, sub-terminally widened, terminally narrowed, apex slightly dilated, proximally arcuate and distally straight in lateral view, with three setae: one apical terminal, one apical inner and one sub-apical inner (Fig. 2F). Two apical parameral setae close-set. Parameral apices directed inwards. Parameres basally slightly curved, distally relatively straight in lateral view, sub-terminally widened, terminally narrowed in lateral view (Fig. 3B). GONOSTYLI. Almost straight, elongate, thin, with one apical seta, three inner setae and one outer seta (Fig. 2G). SPERMATHECA. Small, curved, widest in proximal third, apically sub-spherical (Fig. 2H). FEMALE ABDOMINAL STERNITE VIII. Large, transverse, setose both medially and distally, with a narrow, pointed anterior process. Microsculpture consisting of transverse polygonal meshes (Fig. 2I). Bionomy, distribution and type locality The type specimens were collected in traps for endogean fauna (cans) baited with rotten meat placed in the deep soil on Mt Bobija, near the town of Ljubovija, western Serbia, as well as by pitfall trapping with rotten meat as bait in the deep, totally dark parts of the Simina Jama Pit, village of Gornje Ko��lje, Debelo Brdo saddle, Mt Povlen, near the town of Ljubovija, western Serbia (Fig. 9). The type locality on Mt Bobija is located on its northern slope, at an altitude of 1000 m a.s.l., in a beech forest, close to several streams. The entrance of the Simina Jama Pit is situated at 920 m a.s.l., the total length of its investigated channels is 270 m, while its depth is 56 m. It starts with a 31-m long vertical passage, which splits into two horizontal channels ��� left and right (Anđelić et al. 2011). Beetle specimens were found at the end and in the middle of the left horizontal channel with a clay muddy substrate and rocks, on the vertical limestone walls and floor with a high level of humidity (presence of trickling water). The places where the specimens were found in the pit are shown in Fig. 4. It is assumed that the species is actually endogean, as is the case with some other leiodid taxa (e.g., Magdelainella spp.), which inhabit the soil beneath deeply sunken rocks and forest detritus, but can also be found in caves and pits (Pavićević et al. 2012)., Published as part of ��ur��i��, Sre��ko, Pavi��evi��, Dragan, Vesovi��, Nikola, Vrbica, Maja, Kuraica, Milo��, Markovi��, ��or��e, Petkovi��, Matija, Lazovi��, Vladimir, Panteli��, Dejan & Bosco, Fabrizio, 2021, On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia, pp. 55-81 in European Journal of Taxonomy 782 (1) on pages 63-66, DOI: 10.5852/ejt.2021.782.1589, http://zenodo.org/record/5761502, {"references":["Andelic M., Njunjic I., Jovanovic J. & Pavicevic D. 2011. Rezultati interdisciplinarnih istrazivanja Simine jame, selo Gornje Koslje, opstina Ljubovija. In: Calic J. (ed.) Proceedings of the 7 th Symposium on Karst Protection (Bela Palanka, Serbia, 21 - 22.5.2011): 143 - 148. Student Speleologic and Alpinistic Club & Municipality of Bela Palanka, Belgrade-Bela Palanka, Serbia.","Pavicevic D., Popovic M., Komnenov M. & Njunjic I. 2012. Diversity of arthropod fauna in caves and pits of Kamena Gora (Serbia) and its surroundings. In: Pavicevic D. & Perreau M. (eds) Fauna Balkana. Volume 1: 151 - 176. Department of Biology and Ecology, Faculty of Sciences, University of Novi Sad, Novi Sad, Serbia."]}