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The life cycle, morphology, and bionomy of Photinus extensus Gorham, 1881, an endemic species of Mexico, are described. Redescriptions of adults (male and female) are also presented. Larvae were reared to the adult stage from eggs laid by females collected at the El Pedregal de San Ángel Ecological Reserve, south of Mexico City. The activity period of adults of P. extensus begins at the end of July and finishes by the end of August. Females lay between 3 and 198 eggs. Larvae hatch from the eggs after a period of 11 to 71 days, undergo 6 larval instars and a pupal stage in an annual cycle. Morphological characters of the sixth larval instar of P. extensus are compared with those of several other genera assigned to the tribe Photinini. Knowledge of the natural history of firefly larvae is relevant since most species do not feed as adults and therefore depend on resources acquired during the larval stage.

Cenophengus kikapu sp. nov. ( Figs. 7–11) Etymology. The term kikapu alludes to the Kikapu tribe that lives in Coahuila, Mexico. Diagnosis. Cenophengus kikapu is similar in appearance to Cenophengus sonorensis Zaragoza 2008, but can be recognized by the following characters. In Cenophengus sonorensis the head is almost as wide as the pronotum, whereas in C. kikapu the head is wider than the pronotum. In addition the terminal maxillary palpomere is as long as the preceding three combined in C. sonorensis, whereas in C. kikapu it is longer than the preceding three combined. Description, male. Total body length 6.4 mm; total body width 0.8 mm. Dark brown body, except for the pronotum, legs and the last two abdominal segments that are amber colored (Figs. 7, 8). Head. Surface concave, wider (0.86 mm) than long (0.52 mm), measured dorsally to exterior margins almost as wide (0.86 mm) as pronotum (0.8 mm), integument chagrined, coarsely punctuate, each puncture bearing an amber seta; interantennal distance (0.09 mm), half of the length of 1 st antennomere; large eyes, hemispherical, prominent, finely faceted, longer (0.45 mm) than wide (0.36 mm); interocular distance equal to length of eyes; short antennae (1.58 mm) barely reaching pronotal posterior margin; 1 st antennomere (0.15 mm) longer than next two combined, 3 rd cup-shaped, 4 th in length 0.12, 5 th to 11th about equal in length (0.15 mm), 12 th (terminal) lanceolate (0.2 mm), antennal rami lanceolate, two times longer than respective antennomere; anterior border of frons concave; clypeus bilobed; terminal maxillary palpomere robust securiform, as long as preceding three combined (0.3 mm); terminal labial palpomere spindle-shaped, five times longer than preceding (0.1 mm). Thorax. Pronotum longer (1.07 mm) than wide (0.8 mm); tegument chagrined, coarsely punctuate; each puncture bearing an amber seta, convex disc, one longitudinal excavation on each side of midline, anterior border concave, posterior almost straight with a middle notch, lateral margins almost parallel, anterior and posterior angles rounded; scutellum almost quadrangular, posterior border truncated, shiny integument, thickly dotted, each puncture with a yellowish seta; elytra almost three and a half times as long (1.95 mm) as wide (0.53 mm), convex; hindwings with radial cell closed, r3 vein (r3) absent, r4 vein (r4) developed, the posterior radial vein (RP) reduced (length less than half the size of the vein MP1+2), medial field containing six main veins: MP3, MP4, CuA1, CuA2, CuA3+4, and AA 3+4; CuA and AA well marked and cubitoanal cell closed, AP3+4 long.; 1 st and 2 nd tarsomeres of pro-, meso- and metathoracic legs about equal in length. Abdomen. Integument shiny, dotted and silky, penultimate sternite sinuate, last sternite cleft. Aedeagus: with three teeth at the apex of paramere (Figs. 9–11). Immatures and females. Unknown. Distribution: Coahuila, Mexico (Fig. 6). Type material. Holotype (male) (COL-TIP-03587): “ Mexico: El Cañón, Cuatro/ Ciénegas, Coahuila, Col. MTO/y UOGV / 21-feb-2012 col./ nocturna luz blanca”. Paratypes (COL-TIP-03588, COL-TIP-03589): two males, same data. Deposited at CNIN.

Cenophengus tupae sp. nov. ( Figs. 23–27) Etymology. The term Tu-pae means mud in the Pame language (Xi’úi), which is spoken in Tamosopo, San Luis Potosí, Mexico. This term refers to the color of the body in the new species. Diagnosis. Cenophengus tupae is similar in appearance to Cenophengus wittmeri Zaragoza, 1984, but can be distinguished by its shorter size, interocular distance and the terminal maxillary palpomere. In C. tupae interocular distance is 2.5 times longer than eye width, whereas in C. wittmeri it is twice longer than eye width. The terminal maxillary palpomere is shorter than the preceding three combined in C. tuape, whereas in C. longicollis it is as long as the preceding three combined. Additionally in C. tupae the antennal rami is three times as long as respective antennomere, whereas in C. wittmeri it is twice longer than the respective antennomere. Description, male. Total body length 5.2 mm; total body width 0.6 mm. Brown body, except for the antennae and a stripe on the pronotum that are amber colored (Figs. 23, 24). Head. Surface concave, wider (0.6 mm) than long (0.4 mm), measured dorsally to exterior margins almost as wide (0.6 mm) as the pronotum (0.62 mm), integument chagrined, coarsely punctuate, each puncture bearing an amber seta; interantennal distance a third of 1 st antennomere length (0.05 mm); medium sized eyes, hemispherical prominent, finely faceted, longer (0.3 mm) than wide (0.25 mm); interocular distance greater than the eyes length (0.4 mm); long antennae (1.6 mm) extending beyond pronotal posterior margin; 1 st antennomere (0.16 mm) longer than the next two combined, 3 rd cup-shaped, 4 th in length (0.12 mm), 5 th to 11 th about equal in length (0.15 mm), 12 th (terminal) lanceolate (0.15 mm); antennal rami lanceolate, three times longer than respective antennomere; anterior border of frons concave; clypeus bilobed; terminal maxillary palpomere securiform, as long as the preceding two combined (0.25 mm); terminal labial palpomere spindle-shaped, three times longer than the preceding. Thorax. Pronotum longer (0.72 mm) than wide (0.6 mm); tegument chagrined, coarsely punctuate; each puncture bearing an amber seta, convex disc, with two longitudinal excavations one on each side of the midline, anterior border concave, the posterior almost straight with a middle notch, lateral margins almost parallel, anterior angles rounded and posterior angles acute; scutellum almost quadrangular, posterior border truncated, shiny integument, thickly dotted, each puncture with an amber colored seta; elytra almost four times as long (1.9 mm) as wide (0.46 mm), convex; hindwings with radial cell closed, r3 vein present, r4 vein developed, the posterior radial vein (RP) reduced (length less than half the size of the vein MP1+2), medial field containing six main veins: MP3, MP4, CuA1, CuA2, CuA3+4, and AA 3+4; CuA and AA well marked and cubitoanal cell closed, AP3+4 long; 1 st and 2 nd tarsomeres of prothoracic legs about equal in length, 1st tarsomere of meso- and metathoracic legs longer than 2nd. Abdomen. Integument shiny, punctured, densely setose, penultimate sternite with posterior margin sinuate, last sternite cleft. Aedeagus: with one tooth at the apex of paramere (Figs. 25–27). Immatures and females. Unknown. Distribution: San Luis Potosí, Mexico (Fig. 17). Type material. Holotype (male) ( COL-TIP-03594): Mexico, San Luis Potosí,/ Tamasopo. Cerro al noroeste/ del cafetal, 01-06-15, N 21°55.47’ / W 99°24.95’ Col./ Jessica Ríos. Paratype ( COL-TIP-03595): male, same data. Deposited at CNIN., Published as part of Vega-Badillo, Viridiana, Zaragoza-Caballero, Santiago & Ríos-Ibarra, Jessica Jazmín, 2021, New species of the genus Cenophengus LeConte 1881 (Coleoptera; Phengodidae) from Mexico and Guatemala, pp. 223-238 in Zootaxa 5023 (2) on page 233, DOI: 10.11646/zootaxa.5023.2.3, http://zenodo.org/record/5225824, {"references":["Zaragoza Caballero, S. (1984) Descripcion de dos especies nuevas y nuevos registros de Cenophengus (Coleoptera: Phengodidae; Mastinocerini). Anales del instituto de Biologia de la Universidad Nacional Autonoma de Mexico, Serie Zoologia, 55 (1), 203 - 208."]}

Cenophengus mumui sp. nov. ( Figs. 18–22) Etymology. The term Mumui means sand in the Pame language (Xi’úi) which is spoken in Tamosopo, San Luis Potosí, Mexico. This term refers to the sandy color of the body in the new species. Diagnosis. Cenophengus mumui is similar in appearance to Cenophengus munizi Zaragoza-Caballero 2008. These two species can be differentiated by the shape and color of the head, which is square and brown in the new species, while C. munizi exhibits a rectangular shaped head, which is amber colored like the rest of the body. Additionally, in C. mumui the antennal rami are 1.5 times as long as respective antennomere, whereas in C. munizi they are twice as long as respective antennomere. Description, male. Total length 3.5 mm; total width 0.58 mm. Yellow body, except for the head which is brown colored (Figs. 18, 19). Head. Surface concave, wider (0.61 mm) than long (0.49 mm), measured dorsally to exterior margins almost wider (0.61 mm) than the pronotum (0.58 mm), integument smooth, coarsely punctuate, each puncture bearing an amber seta; interantennal distance longer than half of 1 st antennomere length, (0.1 mm); small eyes, hemispherical, prominent, finely faceted, longer (0.2 mm) than wide (0.15 mm); interocular distance greater than the eyes length (0.36 mm); antennae (1.8 mm) extending beyond pronotal posterior margin; 1 st antennomere (0.15 mm) as long as next two combined, 3 rd cup-shaped, 4 th in length (0.12 mm), 5 th to 11 th about equal in length (0.17 mm), 12 th (terminal) lanceolate (0.25 mm), antennal rami lanceolate, one and a half times longer than respective antennomere; anterior border of frons concave; clypeus bilobed; terminal maxillary palpomere robust and securiform, as long as the preceding three combined (0.16 mm); terminal labial palpomere cylindrical (0. 05 mm) and two times longer than the preceding. Thorax. Pronotum as long (0.58 mm) as wide (0.56 mm); tegument smooth, coarsely punctuate; each puncture bearing an amber seta, convex disc, with one longitudinal excavation on each side of the midline, anterior border concave, posterior almost straight with a middle notch, lateral margins almost parallel, anterior and posterior angles rounded; scutellum almost quadrangular, posterior border truncated, shiny integument, thickly dotted, each puncture with an amber seta; elytra almost four times as long (1.62 mm) as wide (0.37 mm), convex; hindwings with radial cell closed and slightly defined, r3 and r4 veins absent, the posterior radial vein (RP) reduced (length less than half the size of the vein MP1+2), those of the anterior anal and posterior anal sectors, absent, only AA well marked, AP3+4 short.; 1 st and 2 nd tarsomeres of pro-, meso- and metathoracic legs about equal in length. Abdomen. Integument shiny, punctured, densely setose, penultimate sternite with posterior margin sinuate, last sternite cleft. Aedeagus: with three teeth at the apex of paramere (Figs. 20–22). Immatures and females. Unknown Distribution: San Luis Potosí, Mexico (Fig. 17). Type material. Holotype (male) ( COL-TIP-03596): Mexico, San Luis Potosí,/ Tamasopo. Cerro al noroeste/ del cafetal, 01-06-15, N 21°55.47’ / W 99°24.95’ Col./ Jessica Ríos. Deposited at CNIN., Published as part of Vega-Badillo, Viridiana, Zaragoza-Caballero, Santiago & Ríos-Ibarra, Jessica Jazmín, 2021, New species of the genus Cenophengus LeConte 1881 (Coleoptera; Phengodidae) from Mexico and Guatemala, pp. 223-238 in Zootaxa 5023 (2) on page 231, DOI: 10.11646/zootaxa.5023.2.3, http://zenodo.org/record/5225824, {"references":["Zaragoza-Caballero, S. (2008) Cenophengus en Mexico (Coleoptera: Phengodidae: Mastinocerinae): descripcion de cuatro nuevas especies. Dugesiana, 15 (2), 153 - 158."]}

Cenophengus xiinbali sp. nov. (Figs. 28–32) Etymology. The term “Aj xíinbal” means traveler in the Maya language. This species is named in honor of the South American migrant people who have gone missing or perished in the pursuit of a better life. Diagnosis. Cenophengus xiinbali is similar in appearance to Cenophengus longicollis Wittmeri 1976, but can be distinguished by the interocular distance and terminal maxillary palpomere. In C. xiinbali the interocular distance is 3.5 times longer than eye width, whereas in C. longicollis it is three times longer. The terminal maxillary palpomere is as long as the preceding three combined in C. xiinbali, whereas in C. longicollis it is longer than the preceding three combined. Description, male. Total body length 8.3 mm; total body width 0.93 mm. Brown body, except for the pronotum, legs and last two abdominal segments, which are orange colored (Figs. 28, 29). Head. Surface concave, longer (0.8 mm) than wide (0.73 mm), measured dorsally to exterior margins thinner (0. 73 mm) than pronotum (0.93 mm), integument chagrined, coarsely punctuate, each puncture bearing an amber seta; interantennal distance half-length of 1 st antennomere (0.12 mm); small sized eyes, hemispherical, finely faceted, almost as long (0.33 mm) as wide (0.12 mm); interocular distance greater than eyes length (0.6 mm); antennae (2.4 mm) extending beyond pronotal posterior margin; 1 st antennomere (0.21 mm) as long as next two combined, 3 rd cup-shaped, 4 th in length (0.1 mm), 5 th to 11 th about equal in length (0.21 mm), 12 th (terminal) lanceolate (0.26 mm), antennal rami lanceolate, ramus two times longer than respective antennomere; anterior border of frons concave; clypeus bilobed; terminal maxillary palpomere securiform, as long as the preceding three combined (0.3 mm); terminal labial palpomere spindle-shaped, three times longer than the preceding (0.06 mm). Thorax. Pronotum longer (1.14 mm) than wide (0.93 mm); tegument chagrined, coarsely punctuate; each puncture bearing an amber seta, convex disc, with one longitudinal excavation on each side of the midline, anterior border concave, posterior border almost straight with a middle notch, lateral margins almost parallel, anterior angles rounded and posterior angles acute; scutellum almost quadrangular, posterior border truncated, shiny integument, thickly dotted, each puncture with a black seta; elytra almost four and a half times as long (2.68 mm) as wide (0.64 mm), convex; hindwings with radial cell closed, r3 vein absent, r4 vein reduced, the posterior radial vein (RP) reduced (length less than half the size of the vein MP1+2), medial field containing six main veins: MP3, MP4, CuA1, CuA2, CuA3+4, and AA 3+4; CuA and AA well marked and cubitoanal cell closed, AP3+4 long.; 1 st and 2 nd tarsomeres of prothoracic legs about equal in length, 1st tarsomere of meso- and metathoracic legs longer than 2nd. Abdomen. Integument shiny, punctured, densely setose, penultimate sternite with posterior margin sinuate, last sternite cleft. Aedeagus: with three teeth at the apex of paramere (Figs. 30–32). Immatures and females. Unknown Distribution: Puerta Parada, Guatemala (Fig. 17). Type material. Holotype (male) ( COL-TIP-03597): “ Guatemala: Guatemala Dept./ Puerta Parada Km 14.5 carr. a/ El Salvador 1840 m alt./ 8-15/VI/2013 Col. J.C Schster ”. Paratype (COL-TIP-03598): male, same data. Deposited at CNIN., Published as part of Vega-Badillo, Viridiana, Zaragoza-Caballero, Santiago & Ríos-Ibarra, Jessica Jazmín, 2021, New species of the genus Cenophengus LeConte 1881 (Coleoptera; Phengodidae) from Mexico and Guatemala, pp. 223-238 in Zootaxa 5023 (2) on pages 233-236, DOI: 10.11646/zootaxa.5023.2.3, http://zenodo.org/record/5225824

Cleicosta equatoreana sp. nov. (Figs. 2 I-L) Type material: Holotype (♂ NMNH): " ECUADOR: Sucumbios / Sacha Lodge, 270 m / 00.5°S, 76.5°W / 04-14 May 1994 / P. Hibbs, Malaise""Michael Ivie Collection". Diagnosis: Cleicosta equatoreana sp. nov., is similar to C.monaguense sp.nov., however they differ in their interantennal distance. In C. equatoriana sp. nov., it is equal to antennal fossa width; in C. monaguense sp. nov., interantennal distance is greater than fossa width. Additionally, in C. equatoreana sp. nov., the terminal maxillary palpomere is 1.5 times longer than the preceding three combined, whereas in C. monaguense sp. nov., the terminal palpomere is equal in length as the preceding three combined. Description: Male: Body length 2.4, maximum width 0.38. Body color light brown. Head: Surface concave; wider (0.42) than long (0.32); at eye level, wider (0.42) than pronotum; integument smooth, coarsely punctuate, each puncture bearing an amber seta; interantennal distance (0.06) equal to antennal fossa width (0.07); eyes large, hemispherical, finely facetted, prominent, longer (0.18) than wide (0.08); interocular distance (0.24) 3 times greater than eye width; antennae short (0.81), barely reaching pronotal posterior border; 1 st antennomere longer (0.1) than next 2 combined, 3 rd cup-shaped, small (0.03), 4 th 0.07 in length, 5 th to 10 th about equal in length (0.08), 11 th 0.06 in length, 12 th (terminal) bullet-shaped with apex acute (0.06); antennal rami lanceolate, twice as long as respective antennomere; labrum bilobed; terminal maxillary palpomere robust, spindle-shaped (0.12), 1.5 times longer than preceding 3 combined; terminal labial palpomere spindle-shaped (0.04), 4 times longer than preceding one. Thorax: Pronotum wider (0.38) than long (0.31); integument smooth, coarsely punctuate, each puncture bearing an amber seta; disc convex, anterior margin almost straight, posterior margin convex with small notch; scutellum spatulate, posterior margin with small notch on posterior margin, integument shiny, densely punctuate, each puncture bearing an amber seta; elytra short, 3 times longer (1.02) than wide (0.32); posterior wings (Fig. 2L) with posterior medium vein (MP 1 + 2) distinct, posterior radial vein (RP) absent, radial cell (CR) open; cubital-anal wing area with undivided anterior anal vein (AA) and posterior anal vein (AP) visible; 1 st and 2 nd tarsomeres of all legs about equal in length. Abdomen: Integument shiny, punctuate, with silky appearance due to dense setosity; penultimate sternite with posterior margin emarginate. Female and immatures: Unknown. Distribution: Sucumbios, Ecuador (Fig. 3). Etymology: Specific epithet alludes to the type locality.

Cleicosta monaguense sp. nov. (Figs. 2 E-H) Type material: Holotype (♂ MIZA): " VENEZUELA: Monagus [Monagas], 700 m / Caripe, Cueva #87-82/ Guacharo, 20-30 July 1987 / Forest over coffee/ S & J Peck, Malaise FIT""From the Michael Ivie Collection". Paratypes: " VENEZ: Monagus, 700 m / Caripe, Cueva #87-82/ Guacharo, 20-30 July 87/ Forest over coffee/ S & J Peck, Malaise" "Michael Ivie Collection" (3♂ CNIN, 3♂ MTEC). Diagnosis: Cleicosta monaguense sp. nov., can be separated from the similar C. breviplumata comb. nov., by its greater interantennal distance which equals twice the antennal fossa width (interantennal distance equals antennal fossa width in C. breviplumata comb. nov.). The interocular distance is three times greater than eye width in C. monaguense sp. nov., and in C. breviplumata comb. nov., it is four times greater. Additionally, in C. monaguense sp. nov., the first tarsomere is shorter than the second in all legs, whereas in C. breviplumata comb. nov., the first and second tarsomeres are equal in length in all three pairs of legs. Description: Male: Body length 3.50, maximum body width 0.46. Light brown body. Head: Surface concave, wider (0.53) than long (0.41); at eye level, wider (0.53) than pronotum (0.46); integument smooth thick and roughly dotted densely and coarsely punctuate, each puncture bearing a brown seta, interantennal distance (0.12) equal to twice antennal fossa width (0.07); eyes large, hemispherical, finely facetted, prominent, longer (0.22) than wide (0.01); interocular distance (0.31) 3 times greater than eye width; antennae short (1.17), barely reaching pronotal posterior margin, 1 st antennomere (0.10) longer than next 2 combined, 3 rd cup-shaped and short (0.03), 4 th in length 0.10, 5 th to 10 th about equal in length (0.11), 11 th measuring 0.09, 12 th (terminal) bullet-shaped with apex acute (0.12); antennal rami lanceolate, 1.5 times as long as respective antennomere; labrum bilobed; terminal maxillary palpomere robust, spindle-shaped, as long as preceding 3 combined (0.15); terminal labial palpomere spindle-shaped (0.07), 6 times as long as preceding one. Thorax: Pronotum wider (0.46) than long (0.42), integument smooth, densely and coarsely punctuate; each puncture bearing a brown seta; disc convex, anterior margin almost straight, anterior angles rounded, lateral margins slightly curved, posterior margin curved, posterior angles acute; scutellum spatulate, with small notch on posterior margin, integument shiny, densely punctuate, each puncture bearing a yellow seta; elytra short, 2.5 times longer (0.95) than wide (0.33); posterior wings (Fig. 2H) with posterior medium vein (MP 1 + 2) long and distinct, posterior radial vein (RP) absent, radial cell (CR) open; wing cubital-anal area with anterior anal vein (AA) (divided into AA 1 + 2 and AA 3 + 4) and posterior anal vein (AP) distinct, cubital-anal vein (CuA) divided into MP 4,MP 3a and MP 3b mid-posterior veins;1 st tarsomere of all legs shorter than 2 nd. Abdomen: Integument shiny, punctuate, with silky appearance due to dense setosity; penultimate sternite with posterior margin sinuate, last sternite with posterior margin notched; pygidial posterior margin straight. Female and immatures: Unknown. Distribution: Monagas, Venezuela (Fig. 3). Etymology: Specific epithet alludes to the type locality.

Cleicosta gen. nov. Type species: Cenophengus breviplumatus Wittmer, 1976. Diagnosis: Cleicosta gen. nov., is morphologically similar to Cenophengus LeConte, 1881: both genera exhibit clearly separated tentorial pits, vertical frons and simple tarsomeres. Cleicosta gen. nov., differs from Cenophengus in the arrangement of the gular sutures: parallel anteriorly in the new genus and divergent anteriorly in Cenophengus. Additionally, in Cleicosta gen. nov., pronotum is subquadrate in shape and elytra are short (reaching only first or second abdominal segment). In Cenophengus the pronotum is rectangular, and elytra long, last 3 tergites exposed. Other important characteristics in Cleicosta gen. nov., are: aedeagus with lateral lobes parallel, elongate, narrowed medially to toothless apex; in Cenophengus with lateral lobes parallel,with apical teeth. Head: Wider than long (Fig. 1A), fully exposed; integument smooth, glossy, coarsely punctuate: antennae 12-articulated serrated, extending slightly beyond pronotal posterior margin; antennomeres 4 th to 11 th with lanceolate rami,1.5 to twice times longer than respective antennomere; frons vertically produced; interantennal distance nearly equal to first antennomere length; eyes finely facetted, hemispherical, variable in length; labrum 3 times wider than long; mandibles simple, thin, falcate; maxillary palpi 4-segmented; terminal palpomere bullet-shaped with apex acute, longer than preceding 3; 3 rd palpomere shorter than 2 nd; labial palpi 2-segmented; terminal palpomere fusiform, 5 times longer than preceding one; tentorium with 2 distinctly separated pits; gular sutures parallel anteriorly (Fig. 1B). Thorax: Pronotum subquadrate (Fig. 1C); anterior angles rounded, posterior angles acute, lateral margins rounded; integument smooth, glossy, densely punctured (Fig. 1D); prosternal anterior margin almost straight; sternal suture complete. Elytra short, reaching 1 st or 2 nd abdominal segment, 3 to 4 times longer than wide, subparallel, apex rounded. Posterior wings with posterior medium vein (MP 1 + 2) always present; posterior radial vein (RP) absent; radial cell open; venation otherwise variable. Length of legs gradually increasing from pro- to metathoracic legs; tarsi simple; in all pairs of legs, length of 4 th tarsomere equal to half the length of 5 th; claws simple. Abdomen: Integument shiny, punctured, densely setose; penultimate sternite with posterior margin sinuate; last sternite deeply notched.Aedeagus with median lobe cylindrical, with apex rounded; lateral lobes elongate, parallel externally, narrowed medially to toothless apex (Figs. 1 E-G). Etymology: The new genus is named Cleicosta in recognition of the outstanding work of Dr. Cleide Costa, on her eightieth birthday. Her first name was abbreviated as "Clei" in order to avoid homonymy with the elaterid genus Cleidecosta Johnson, 2002. Gender feminine.

Tribe Deltochilini Lacordaire, 1856 Canthon ateuchiceps Bates, 1881. Source: Deloya & Morón 1994 Canthon cyanellus LeConte, 1859. Source: Delgado 1997 Canthon (Glaphyrocanthon) corporaali Balthasar, 1939. Source: Rivera–Cervantes & Halffter 1999 Canthon (Glaphyrocanthon) leechi (Martínez, Halffter & Halffter, 1969). Source: Rivera-Cervantes & Halffter 1999 Canthon humectus (Say, 1832). Source: Deloya et al. 1993 Canthon indigaceus LeConte, 1866. Source: Deloya et al. 1993 Canthon viridis Beauvois, 1805. Source: Bates 1886 - 1889 Deltochilum (Deltochilum) scabriusculum Bates, 1887. Source: Bates 1886 –1889 Deltochilum gibbosum Bates, 1887. Source: Deloya et al. 1993 Deltochilum tumidum Howden, 1966. Source: Deloya et al. 1993 Pseudocanthon perplexus LeConte, 1847. Source: Deloya & Morón 1994, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 42, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Bates, H. W. (1881) Adephaga. In: Biologia Centrali-Americana. Insecta: Coleoptera. Fol. I. Part. 1. published for the editors by R. H. Porter, London, Available from: http: // www. sil. si. edu / DigitalCollections / bca / navigation / bca _ 12 _ 01 _ 01 / bca _ 12 _ 01 _ 01. cfm? start = 17 (accessed 27 August 2017)","Deloya, C. & Moron, M. A. (1994) Coleopteros Lamellicornios del Distrito de Jojutla, Morelos, Mexico (Melolonthidae, Scarabaeidae, Trogidae y Passalidae). Listados Faunisticos de Mexico (F). Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Mexico, 49 pp.","Delgado, L. (1997) Distribucion estatal de la diversidad y nuevos registros de Scarabaeidae (Coleoptera) mexicanos. Folia Entomologica Mexicana, 99, 37 - 56.","Rivera-Cervantes, L. E. & Halffter, G. (1999) Monografia de las especies mexicanas de Canthon del subgenero Glaphyrocanthon (Coleoptera: Scarabaeinae). Acta Zoologica Mexicana, Nueva Serie, 77, 23 - 150.","Deloya, C., Burgos, A., Blackaller, J. & Lobo, J. M. (1993) Los Coleopteros Lamellicornios de Cuernavaca, Morelos, Mexico (Passalidae, Trogidae, Scarabaeidae, y Melolonthidae). Boletin de la Sociedad Feracruzana de Zoologia, 3 (1), 15 - 55.","Bates, H. W. (1886 - 1890) Pecticornia and Lamellicornia. In: Biologia Centrali-Americana, Insecta. Coleoptera. Fol. II. Part 2. published for the editors by R. H. Porter, London. Available from: http: // www. sil. si. edu / DigitalCollections / bca / navigation / bca _ 12 _ 02 _ 02 / bca _ 12 _ 02 _ 02. cfm? start = 19 (accessed 8 November 2017)"]}

Tribe Omaliini MacLeay, 1825 Omalium meximontanum Thayer, 2003. Source: Márquez 2006 Phloeonomus centralis Blackwelder, 1944. Source: Burgos & Trejo-Loyo 2001; Caballero-Pérez 2003; Márquez 2001, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 33, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin Sociedad Entomologica Aragonesa, 38, 181 - 198.","Burgos S., A. & Trejo-Loyo, A. G. (2001) Lista preliminar de los coleopteros registrados para el estado de Morelos, Mexico. In: Navarrete Heredia, J. L., Fierro-Lopez, H. E. & Burgos-Solorio, A. (Eds.), Topicos sobre Coleoptera de Mexico. Universidad de Guadalajara y Universidad Autonoma del Estado de Morelos, Guadalajara, Jalisco, pp. 69 - 95.","Caballero-Perez, U. (2003) Staphylinidae necrofilos (Insecta: Coleoptera) de la Sierra de Huautla, Morelos. Bachelor's thesis. Universidad Nacional Autonoma de Mexico, Facultad de Estudios Superiores Iztacala, Mexico, 110 pp.","Marquez, J. (2001) Especies necrofilas de Staphylinidae del municipio de Tlayacapan Morelos. Folia Entomologica Mexicana, 40 (1), 93 - 131."]}

Tribe Exosternini Bickhardt, 1914 Phelister sp. Source: Burgos & Trejo-Loyo 2001 Pseudister rufulus (Lewis, 1888). Source: Burgos & Trejo-Loyo 2001; Navarrete-Heredia & González-Estrada 2003, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 30, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Burgos S., A. & Trejo-Loyo, A. G. (2001) Lista preliminar de los coleopteros registrados para el estado de Morelos, Mexico. In: Navarrete Heredia, J. L., Fierro-Lopez, H. E. & Burgos-Solorio, A. (Eds.), Topicos sobre Coleoptera de Mexico. Universidad de Guadalajara y Universidad Autonoma del Estado de Morelos, Guadalajara, Jalisco, pp. 69 - 95.","Lewis, G. (1888) Histeridae. In: Biologia Centrali-Americana. Insecta: Coleoptera. Fol. II. Part. 1. published for the editors by R. H. Porter, London. Available from: http: // www. sil. si. edu / DigitalCollections / bca / navigation / bca _ 12 _ 02 _ 01 / bca _ 12 _ 02 _ 01. cfm? start = 198 (accessed 27 August 2017)","Navarrete-Heredia J. L. & Gonzalez-Estrada, D. (2003) Las especies de Histeridae (Coleoptera) de la Coleccion Entomologica del Centro de Estudios en Zoologia, Universidad de Guadalajara (Mexico). Boletin de la Sociedad Entomologica Aragonesa, 33, 125 - 129."]}

Subfamily Saprininae Blanchard, 1845 Xerosaprinus psyche (Casey, 1916). Source: Burgos & Trejo-Loyo 2001; Navarrete-Heredia & González-Estrada 2003, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 30, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Burgos S., A. & Trejo-Loyo, A. G. (2001) Lista preliminar de los coleopteros registrados para el estado de Morelos, Mexico. In: Navarrete Heredia, J. L., Fierro-Lopez, H. E. & Burgos-Solorio, A. (Eds.), Topicos sobre Coleoptera de Mexico. Universidad de Guadalajara y Universidad Autonoma del Estado de Morelos, Guadalajara, Jalisco, pp. 69 - 95.","Navarrete-Heredia J. L. & Gonzalez-Estrada, D. (2003) Las especies de Histeridae (Coleoptera) de la Coleccion Entomologica del Centro de Estudios en Zoologia, Universidad de Guadalajara (Mexico). Boletin de la Sociedad Entomologica Aragonesa, 33, 125 - 129."]}

Tribe Scydmaenini Leach, 1815 Scydmaenus brunneus Schaum, 1841. Source: Burgos & Trejo-Loyo 2001; Navarrete-Heredia & Newton 2013 Ascydmus sp. Source: Burgos & Trejo-Loyo 2001; Márquez 1994 Neladius sp. Source: Burgos & Trejo-Loyo 2001; Márquez 1994, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 37, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Burgos S., A. & Trejo-Loyo, A. G. (2001) Lista preliminar de los coleopteros registrados para el estado de Morelos, Mexico. In: Navarrete Heredia, J. L., Fierro-Lopez, H. E. & Burgos-Solorio, A. (Eds.), Topicos sobre Coleoptera de Mexico. Universidad de Guadalajara y Universidad Autonoma del Estado de Morelos, Guadalajara, Jalisco, pp. 69 - 95.","Navarrete-Heredia, J. L. & Newton, A. F. (2013) Staphylinidae of Mexico: checklist updates and papers published between 2002 - 2013. Dugesiana, 20 (2), 127 - 139.","Marquez, J. & Navarrete, J. L. (1994) Especies de Staphylinidae (Insecta: Coleoptera) asociadas a detritos de Atta mexicana (F. Smith) (Hymenoptera: Formicidae) en dos localidades de Morelos, Mexico. Folia Entomologica Mexicana, 91, 31 - 46."]}

Subtribe Calopterina Green, 1949 Calopteron bifasciatum Gorham, 1880. Source: Burgos & Trejo-Loyo 2001; Zaragoza-Caballero et al. 2003 Calopteron flavipes Blanchard, 1837. Source: CNIN Calopteron jimenezi Dugès, 1878. Source: Zaragoza-Caballero et al. 2003 Calopteron reticulatum (Fabricius, 1775). Source: Burgos & Trejo-Loyo 2001; Zaragoza-Caballero et al. 2003 Calopteron terminale (Say, 1823). Source: CNIN Calopteron obliquum Gorham, 1882. Source: Burgos & Trejo-Loyo 2001 Idiopteron sp. Source: CNIN Leptoceletes sp. Source: CNIN, Published as part of Zaragoza-Caballero, Santiago, Rodríguez-Mirón, Geovanni M., Vega-Badillo, Viridiana, González-Ramírez, Mireya, Zurita-García, Martín L., Domínguez-León, Daniel E., López-Pérez, Sara, Gutiérrez-Carranza, Ishwari G., Cifuentes-Ruiz, Paulina, Pérez-Hernández, Cisteil X., Valle, Enya Ramírez Del & Gutiérrez, Nayeli, 2019, A checklist of the Coleoptera (Insecta) from Morelos, Mexico, pp. 1-122 in Zootaxa 4580 (1) on page 59, DOI: 10.11646/zootaxa.4580.1.1, http://zenodo.org/record/2637573, {"references":["Burgos S., A. & Trejo-Loyo, A. G. (2001) Lista preliminar de los coleopteros registrados para el estado de Morelos, Mexico. In: Navarrete Heredia, J. L., Fierro-Lopez, H. E. & Burgos-Solorio, A. (Eds.), Topicos sobre Coleoptera de Mexico. Universidad de Guadalajara y Universidad Autonoma del Estado de Morelos, Guadalajara, Jalisco, pp. 69 - 95.","Zaragoza-Caballero, S., Noguera, F. A., Chemsak, J. A., Gonzalez-Soriano, E., Rodriguez-Palafox, A., Ramirez-Garcia, E. & Ayala, R. (2003) Diversity of Lycidae, Phengodidae, Lampyridae, and Cantharidae (Coleoptera) in a tropical dry forest region in Mexico: Sierra de Huautla, Morelos. The Pan-Pacific Entomologist, 79 (1), 23 - 37."]}