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© 2020 The Middle East, specially the Zagros region, lies in a strategic position as a crossroads between Africa, Europe and eastern Asia. The landscape of this region that prevailed around the Neanderthal and anatomically modern human occupations is not well known. Only a few sites have been studied in detail in this area, often providing only a faunal list. These reveal that Neanderthals and anatomically modern humans lived in a landscape mainly composed of dry steppes. Here we extend the data obtained from Kaldar Cave through a systematic study of the rodent assemblage. The site provided evidence of a Pleistocene occupation attested by lithic tools associated with the Middle and Upper Palaeolithic, but it was also occupied during the Holocene, as evidenced by Neolithic artefacts. First excavations have revealed small vertebrates in Layer 4 (sub-layer 5 and 5II), belonging to the Upper Palaeolithic, and Layer 5 (sub-layers 7 and 7II), belonging to the Middle Palaeolithic. The rodent assemblage of Kaldar Cave is mainly composed of six arvicoline, two cricetine, one glirid, one dipodid, one gerbilline and two murine species. This assemblage shows that during the Late Pleistocene the environment around the site was mainly composed of open dry steppes, as indicated by the most abundant taxa, Microtus, Ellobius and Meriones. However, murine species indicate the presence of a vegetation cover. The palaeoclimatic conditions are characterized by lower temperatures and also less precipitation than at present. The results obtained with the rodent assemblages show that there is no major palaeoenvironmental or palaeoclimatic change that would explain the cultural shift between Layer 5 (Middle Palaeolithic) and Layer 4 (Upper Palaeolithic).

Along with the early age obtained for the cultural remains attributed to anatomically modern humans from Kaldar Cave, the archaeological assemblages recovered from both Kaldar and Gilvaran Cave located in the Khorramabad Valley (Iran), have yielded charcoal remains that allow the identification of Prunus spp. These remains correspond to the Middle and Upper Palaeolithic, which are the earliest finds attesting to the presence of this taxa in the area. Our anatomical observation of the samples revealed the presence of Prunus spp. (plums) and Prunus cf. amygdalus (cf. almond). This also reflects specific plant communities in the area, characteristic of open forest growing in cool, dry conditions. These results provide new insights into the arboreal cover in this area during an Upper Pleistocene period. Furthermore, anthracological evidence together with other contextual materials provides new clues to assess how Neanderthals and early modern humans adapted to their surrounding landscape, and their relationship with their environment in this region and beyond.

Filiació URV: SI, Kaldar Cave is a key archaeological site that provides evidence of the Middle to Upper Palaeolithic transition in Iran. Excavations at the site in 2014-2015 led to the discovery of cultural remains generally associated with anatomically modern humans (AMHs) and evidence of a probable Neanderthal-made industry in the basal layers. Attempts have been made to establish a chronology for the site. These include four thermoluminescence (TL) dates for Layer 4, ranging from 23,100 ± 3300 to 29,400 ± 2300 BP, and three AMS radiocarbon dates from charcoal samples belonging to the lower part of the same layer, yielding ages of 38,650-36,750 cal BP, 44,200-42,350 cal BP, and 54,400-46,050 cal BP (all at the 95.4% confidence level). Kaldar Cave is the first well-stratified Late Palaeolithic locality to be excavated in the Zagros which is one of the earliest sites with cultural materials attributed to early AMHs in western Asia. It also offers an opportunity to study the technological differences between the Mousterian and the first Upper Palaeolithic lithic technologies as well as the human behaviour in the region. In this study, we present a detailed description of the newly excavated stratigraphy, quantified results from the lithic assemblages, preliminary faunal remains analyses, geochronologic data, taphonomic aspects, and an interpretation of the regional paleoenvironment.