Tropidurus spinulosus (Cope, 1862) Microlophus spinulosus COPE, 1862. TYPE LOcALITY: PARAGUAY. Tropidurus spinulosus: BOULENGER 1885, 1894, 1898; KOSLOWSKY 1898; BERTONI 1914, 1939; BURT & BURT 1933; COcHRAN 1961; ScOTT & LOVETT 1975; TALbOT 1979; AqUINO-SHUSTER et al. 1991; AqUINO et al. 1996; FROST et al. 1998; ZIEGLER et al. 2002; MOTTE et al. 2009; CARVALHO 2013; CAccIALI et al. 2016. Tropidurus (Microlophus) spinulosus: BOETTGER 1885. Tropidurus spinulosus guarani ÁLVAREZ et al., 1994. TYPE LOcALITY: CERRO Hũ, PARAGUARí, PARAGUAY. Tropidurus SP.: AqUINO et al. 1996 [PART.]. Tropidurus guarani: FROST et al. 1998 [PART.]; MOTTE et al. 2004 [PART.]; MOTTE et al. 2009 [PART.]; CARVALHO 2013 [PART.]; CAbRAL & WEILER 2014; CAccIALI et al. 2016 [PART.]. Geographic distribution. Tropidurus spinulosus IS MAINLY ASSOcIATED WITH XERIc AREAS IN SOUTHERN BOLIVIA, WESTERN PARAGUAY, WITH REcORDS IN ROcKY OUTcROPS EAST OF THE PARAGUAY RIVER, AND NORTHEASTERN ARGENTINA (FIG. 11C), Published as part of Cacciali, Pier & Köhler, Gunther, 2018, Diversity of Tropidurus (Squamata: Tropiduridae) in Paraguay-an integrative taxonomic approach based on morphological and molecular genetic evidence, pp. 511-536 in Zootaxa 4375 (4) on page 521, DOI: 10.11646/zootaxa.4375.4.3, http://zenodo.org/record/1160960, {"references":["Cope, E. D. (1862) Catalogue of the reptiles obtained during the exploration of the Parana, Paraguay, Vermejo and Uruguay rivers, by Capt. Thos. J. Page, U. S. N.; and of those procured by Lieut. N. Michler, U. S. Top. Eng., commander of the expedition conducting the survey of the Atrato River. Proceedings of the Academy of Natural Science of Philadelphia, 14, 346 - 359. Available from: http: // www. biodiversitylibrary. org / page / 1951818 page / 358 / mode / 1 up (Accessed 25 Jan. 2018)","Boulenger, G. A. (1885) Catalogue of the lizards in the British Museum (Natural History). 2 nd edition. Vol. 2. Taylor and Francis, London, 497 pp. Available from: http: // www. biodiversitylibrary. org / item / 61183 page / 7 / mode / 1 up (Accessed 25 Jan. 2018)","Boulenger, G. A. (1894) List of reptiles and batrachians collected by Dr. J. Bohls near Asuncion, Paraguay. Annals and Magazine of Natural History, 13, 342 - 348. https: // doi. org / 10.1080 / 00222939408677709","Boulenger, G. A. (1898) A list of reptiles, batrachians, and fishes collected by Cav. Guido Boggiani in the northern Chaco. Annali del Museo Civico di Storia Naturale di Genova, 19, 125 - 127.","Koslowsky, J. (1898) Enumeracion sistematica y distribucion geografica de los reptiles argentinos. Revista del Museo de La Plata, 8, 161 - 200.","Bertoni, A. de W. (1914) Fauna paraguaya: Catalogos sistematicos de los vertebrados del Paraguay. Descripcion Fisica y Economica del Paraguay, 59, 1 - 83.","Bertoni, A. de W. (1939) Catalogos sistematicos de los vertebrados de Paraguay. Revista de la Sociedad Cientifica del Paraguay, 4, 3 - 60.","Burt, C. E. & Burt, M. D. (1933) A preliminary checklist of the lizards of South America. Transactions of the Academy of Science of St. Louis, 28, 1 - 104.","Cochran, D. M. (1961) Type specimens of reptiles and amphibians in the United States National Museum. United States National Museum Bulletin, 220, 1 - 291. https: // doi. org / 10.5962 / bhl. part. 26967","Scott, N. J. & Lovett, J. W. (1975) A collection of reptiles and amphibians from the Chaco of Paraguay. Occasional Papers of the University of Connecticut, 2, 257 - 266.","Talbot, J. J. (1979) Una nueva lista sistematica de reptiles del Paraguay. Informes Cientificos del Instituto de Ciencias Basicas, 2, 76 - 94.","Carvalho, A. L. G. (2013) On the distribution and conservation of the South American lizard genus Tropidurus Wied-Neuwied, 1825 (Squamata: Tropiduridae). Zootaxa, 3640 (1), 42 - 56. https: // doi. org / 10.11646 / zootaxa. 3640.1.3","Boettger, O. (1885) Liste von Reptilien und Batrachiern aus Paraguay. Zeitschrift fur Naturwissenschaft, 58, 213 - 248. Available from: http: // www. biodiversitylibrary. org / page / 31401231 page / 245 / mode / 1 up (Accessed 25 Jan. 2018)","Cabral, H. & Weiler, A. (2014) Lista comentada de los reptiles de la Coleccion Zoologica de la Facultad de Ciencias Exactas y Naturales de Asuncion, Paraguay. Cuadernos de Herpetologia, 28, 19 - 28."]}