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Tomosvaryella susa Motamedinia & Skevington sp. nov. Figs 40A–D, 61, 70C–D Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, broadened at base and rather triangular-shaped (Fig. 40A); broadened gonopods, right wider than left one with small projection in ventral view (Fig. 40B); short ejaculatory ducts, one bearing some saw-like teeth in basal half in lateral view (Fig. 40C‒D). Description: MALE: Body length (excluding antennae): 3.1 mm. Head. Scape, pedicel and arista dark. Pedicel with 2–3 upper setae and a single lower bristle; flagellum light brown, gray pollinose and long tapering (LF:WF = 3.0). Eyes converging but not meeting and separated by less than diameter of frontal facets. Frons silver-gray pollinose, vertex dark and shine, lacking pollinosity, bearing an elevated ocellar triangle. Occiput dark, gray pollinose with scattered brown setae. Thorax. Postpronotal lobe light brown. Prescutum and scutum black, silver-gray and brown pollinose, with scattered setae at anterior supra-alar area and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with about 14 brown setae along posterior margin (up to 0.08 mm). Subscutellum and pleura black; pleuron light brown. Wing. Wing length: 3.1 mm., LW:MWW = 3.0. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.37 mm. light brown, narrowly black at base. Legs. Coxae black, trochanters light brown, smooth. Femora dark brown with narrow but distinct pale base and apices, silver pollinose. Femora without ventral spines. Mid and hind femora with one wrinkled indentation in basal. Tibiae brown at middle, gray pollinose, with two rows of short brown setae on anterior and three rows on posterior side. Mid and hind tibiae with one or two wrinkled indentations in middle. Tarsi light brown. Abdomen. Ground color dark brown, tergite 1 silver gray pollinose with a patch of light brown setae (15‒17) laterally extending along posterior margin, tergites 2‒5 with relatively long brown setae. Sternites white-yellow laterally and brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE:MWE = 0.74). Surstyli symmetrical, broadened at base, rather triangular-shaped (Fig. 40A). Genital capsule in ventral view: gonopods broadened and slightly equal in height, right wider than left one with a small projection, subepandrial sclerite wide basically, phallic guide strong, slightly straight (Fig. 40B). Genital capsule in lateral view: surstyli longer than wide, slightly straight, broadened at apex (Figs 40C‒D). Phallic guide strong, pointed apically (Fig. 40C); phallus with three ejaculatory ducts, one bearing some saw-like teeth in basal half (Fig. 40C‒D). FEMALE: unknown. Etymology: The specific epithet refers to " Susa ", the historical name of Shush, from where the specimen was collected. The name is a noun in apposition. Specimen examined: HOLOTYPE: IRAN: Khuzestan: Shush, 32°06'N, 48°26'E, 68 m a.s.l., 11.iii–10.v.2015, E. Gilasian, Malaise trap, JSS52217 (1♂, CNC). Distribution: Iran (Fig. 61). Notes: Tomosvaryella susa sp. nov. is genetically very close to T. ampliasa sp. nov., differing by 0.8‒1.2% (pairwise divergence) (Supplementary Table 1)., Published as part of Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, pp. 1-103 in Zootaxa 5002 (1) on page 72, DOI: 10.11646/zootaxa.5002.1.1, http://zenodo.org/record/5122655

Tomosvaryella susa Motamedinia & Skevington sp. nov. Figs 40A–D, 61, 70C–D Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, broadened at base and rather triangular-shaped (Fig. 40A); broadened gonopods, right wider than left one with small projection in ventral view (Fig. 40B); short ejaculatory ducts, one bearing some saw-like teeth in basal half in lateral view (Fig. 40C‒D). Description: MALE: Body length (excluding antennae): 3.1 mm. Head. Scape, pedicel and arista dark. Pedicel with 2–3 upper setae and a single lower bristle; flagellum light brown, gray pollinose and long tapering (LF:WF = 3.0). Eyes converging but not meeting and separated by less than diameter of frontal facets. Frons silver-gray pollinose, vertex dark and shine, lacking pollinosity, bearing an elevated ocellar triangle. Occiput dark, gray pollinose with scattered brown setae. Thorax. Postpronotal lobe light brown. Prescutum and scutum black, silver-gray and brown pollinose, with scattered setae at anterior supra-alar area and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with about 14 brown setae along posterior margin (up to 0.08 mm). Subscutellum and pleura black; pleuron light brown. Wing. Wing length: 3.1 mm., LW:MWW = 3.0. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.37 mm. light brown, narrowly black at base. Legs. Coxae black, trochanters light brown, smooth. Femora dark brown with narrow but distinct pale base and apices, silver pollinose. Femora without ventral spines. Mid and hind femora with one wrinkled indentation in basal. Tibiae brown at middle, gray pollinose, with two rows of short brown setae on anterior and three rows on posterior side. Mid and hind tibiae with one or two wrinkled indentations in middle. Tarsi light brown. Abdomen. Ground color dark brown, tergite 1 silver gray pollinose with a patch of light brown setae (15‒17) laterally extending along posterior margin, tergites 2‒5 with relatively long brown setae. Sternites white-yellow laterally and brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE:MWE = 0.74). Surstyli symmetrical, broadened at base, rather triangular-shaped (Fig. 40A). Genital capsule in ventral view: gonopods broadened and slightly equal in height, right wider than left one with a small projection, subepandrial sclerite wide basically, phallic guide strong, slightly straight (Fig. 40B). Genital capsule in lateral view: surstyli longer than wide, slightly straight, broadened at apex (Figs 40C‒D). Phallic guide strong, pointed apically (Fig. 40C); phallus with three ejaculatory ducts, one bearing some saw-like teeth in basal half (Fig. 40C‒D). FEMALE: unknown. Etymology: The specific epithet refers to " Susa ", the historical name of Shush, from where the specimen was collected. The name is a noun in apposition. Specimen examined: HOLOTYPE: IRAN: Khuzestan: Shush, 32°06'N, 48°26'E, 68 m a.s.l., 11.iii–10.v.2015, E. Gilasian, Malaise trap, JSS52217 (1♂, CNC). Distribution: Iran (Fig. 61). Notes: Tomosvaryella susa sp. nov. is genetically very close to T. ampliasa sp. nov., differing by 0.8‒1.2% (pairwise divergence) (Supplementary Table 1).