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Genus Paraphelliactis Carlgren, 1928 Diagnosis (after Carlgren 1949 and Li & Xu 2016; modifications in italics) Hormathiidae with well-developed pedal disc. Column divisible into scapus and scapulus, the former strongly tuberculated and provided with a thick cuticle. Sphincter mesogleal, alveolar. Tentacles arranged in five or more cycles, more than or almost equal to mesenteries at the limbus, with or without mesogleal thickenings on the aboral side. Longitudinal muscles of tentacles ectodermal, radial muscles of oral disc ectodermal or more or less mesogleal. Two well developed siphonoglyphs. Mesenteries hexamerously arranged in five cycles, usually six pairs perfect and sterile, last cycle incomplete or complete. Retractors of mesenteries diffuse and weak. Parietobasilar muscles weak. Acontia well developed. No cinclides. Cnidom: Robust and gracile spirocysts, basitrichs, and microbasic p -mastigophores. Type species: Paraphelliactis spinosa Carlgren, 1928, Published as part of De Melo, Yago A., Targino, Alessandra K. G. & Gomes, Paula B., 2020, New records of family Hormathiidae (Cnidaria: Actiniaria) from Brazilian coast with description of Paraphelliactis labiata n. sp., pp. 557-574 in Zootaxa 4766 (4) on page 560, DOI: 10.11646/zootaxa.4766.4.3, http://zenodo.org/record/3765725, {"references":["Carlgren, O. (1928) Actiniaria der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer \" Valdivia \" 1898 - 1899, 22, 125 - 266. [reprint 1 - 144]","Carlgren, O. (1949) A survey of the Ptychodactiaria, Corallimorpharia and Actiniaria. Kungliga Svenska Vetenskapsakademiens Handlingar, 1, 1 - 121.","Li, Y. & Xu, K. (2016) Paraphelliactis tangi n. sp. and Phelliactis yapensis n. sp., two new deep-sea species of Hormathiidae (Cnidaria: Anthozoa: Actiniaria) from a seamount in the tropical Western Pacific. Zootaxa, 4072 (3), 358 - 372. https: // doi. org / 10.11646 / zootaxa. 4072.3.5"]}

Phelliactis sp. (Fig. 2D, 6A,E-F, Table 6) Material examined: MNRJ 9094. (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83-2 (04° 27.0256’S, 036° 25.6086’W), May 21, 2011, 1896–1931 m. Description. Tall, cylindrical, cream-colored body, somewhat asymmetrical, 13.5 cm high by 6.5 cm width on its longest axis. A big manganese nodule was retained almost completely inside the body, forming a distinct cavity at the base with half the height of the sea anemone (Fig. 6A). The pedal disc was very damaged and mesenteries and associated structures at the aboral side printed on the rock when it was removed for further observation. Column divisible into a smooth, light pink scapulus, withdrawn due to contraction of the oral disc, and a thick, tuberculated scapus. Between 150 and 160 flat to conical tubercles, some with pointed tips, forming longitudinal and transverse rows (Fig. 6E). Oral disc large, bilobed. Approximately 140 short, thin, threadlike tentacles distributed in two marginal cycles. Deep actinopharynx, about two-thirds the length of the body. Two large siphonoglyphs, each attached to a pair of directive mesenteries. Five cycles of mesenteries at the mid-column. Mesenteries are less numerous at the oral side, where four cycles are observed near the oral disc. First and second cycles perfect, including directives, other cycles imperfect. Retractor muscles weak and diffuse (Fig. 6F). Longitudinal muscle of tentacles ectodermal. Radial muscles of the oral disc meso-ectodermal, stronger over the exocoels than over the endocoels. Mesogleal marginal sphincter, alveolar, weak, stratified in the upper portion of the column, where it forms a broader band but abruptly tapering downwards showing scattered and elongated alveoli in longitudinal rows. Acontia rare and small. No gametogenic tissue observed. Cnidom. Spirocysts, Basitrichs, Microbasic p -mastigophores (Table 6). Remarks. Seven species of Phelliactis are registered in the South Atlantic Ocean (data from Ocean Biogeographic Information System- https://obis.org/), but only three of them are known from Brazil: Ph. capricornis, Ph. pelophila, and Ph. robusta (present study). The correct identification of species for this genus requires caution and a minute analysis (Doumenc 1975), which was not possible with the material at hand. Nonetheless, we find it to be distinct from the other Phelliactis species found in the studied area. For example, Ph. robusta, possesses strictly six pairs of perfect mesenteries and also differs from Phelliactis sp. in coloration and thickness of the column., Published as part of De Melo, Yago A., Targino, Alessandra K. G. & Gomes, Paula B., 2020, New records of family Hormathiidae (Cnidaria: Actiniaria) from Brazilian coast with description of Paraphelliactis labiata n. sp., pp. 557-574 in Zootaxa 4766 (4) on pages 568-570, DOI: 10.11646/zootaxa.4766.4.3, http://zenodo.org/record/3765725, {"references":["Doumenc, D. (1975) Actinies bathyales et abyssales de l'ocean Atlantique nord. Familles des Hormathiidae (genres Paracalliactis et Phelliactis) et des Actinostolidae (genres Actinoscyphia et Sicyonis). Bulletin du Museum National d'Histoire Naturelle, Paris, 197, 157 - 204."]}

Phelliactis robusta Carlgren, 1928 (Fig. 2C, 6D, Table 5) Phelliactis robusta Carlgren 1928; 1942; Riemann-Zürneck 1973; Doumenc, 1975; Fautin et al. 2005. Material examined. MNRJ 9093 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 82-2 (04° 33.1742’S, 036° 15.0847’W), May 20, 2011, 2030–2074 m. MOUFPE-CNI 867 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 82-2 (04° 33.1742’S, 036° 15.0847’W), May 20, 2011, 2030–2074 m. LC 152 (three specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 82-2 (04° 33.1742’S, 036° 15.0847’W), May 20, 2011, 2030–2074 m. Description. Large sea anemones, with specimens reaching 111.6 mm high and 33.7 to 59.4 mm wide. Pedal disc well developed, wrapping rocks or glass sponge threads (Fig. 6D). Column very thick, divided into scapus and scapulus. Scapus provided with very thick, polyhedric, sometimes flat tubercles, that form well-marked furrows where the thin cuticle can be seen. Scapulus strongly tuberculated, with tubercles irregularly arranged and decreasing in size towards the oral disc. No cinclides. Oral disc large, bilobed and asymmetrical. Tentacles large and threadlike, with mesogleal thickenings at the base, measuring approximately 11 mm in length. Tentacles nearly 130 in number, arranged in five marginal cycles. Two siphonoglyphs, each attached to a pair of directive mesenteries. Mesenteries hexamerously arranged in five cycles (6+6+12+24+48), more numerous at the base, where they are approximately 150 in number. First cycle perfect and sterile, other cycles imperfect. Gametogenic tissue from the second to fourth cycle of mesenteries. Marginal sphincter mesogleal, alveolar, positioned near the endodermal side. Retractor muscles weak and diffuse. Longitudinal musculature of tentacles ectodermal. Parietobasilar muscles poorly developed. Acontia present. Cnidom. Spirocysts, Basitrichs, Microbasic p -mastigophores (Table 5). Remarks. The specimens in this study were collected at depths greater than 2000 meters, which is consistent with the bathymetry known for the species, which may reach over 2400 m depth (Molodtsova et al. 2008). The largest individual was attached to a rock and a smaller one was attached to branches of a hexactinellid sponge (Fig. 6D). Several foraminifera species, as well as Diacria sp. shells and sponge spicules, were found inside the gastric cavity of these specimens. Riemann-Zürneck (1973) was the first author to present the cnidom of the column for the species, reporting the absence of microbasic p -mastigophores, often found in other Phelliactis species. We also did not find such cnidae in the column of our specimens and the description of our samples perfectly matches those of Riemann-Zürneck (1973) and Doumenc (1975) for specimens from the North Atlantic. Geographic distribution. Phelliactis robusta is in the Eastern North Atlantic Ocean from depths as great as 2100m (Van Praet et al. 1990). We register the species for the first time in the South Atlantic Ocean (Potiguar Basin, RN, Brazil)., Published as part of De Melo, Yago A., Targino, Alessandra K. G. & Gomes, Paula B., 2020, New records of family Hormathiidae (Cnidaria: Actiniaria) from Brazilian coast with description of Paraphelliactis labiata n. sp., pp. 557-574 in Zootaxa 4766 (4) on pages 567-568, DOI: 10.11646/zootaxa.4766.4.3, http://zenodo.org/record/3765725, {"references":["Carlgren, O. (1928) Actiniaria der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer \" Valdivia \" 1898 - 1899, 22, 125 - 266. [reprint 1 - 144]","Carlgren, O. (1942) Actiniaria II. Danish Ingolf - Expedition, 5 (12), 1 - 92","Riemann-Zurneck, K. (1973) Actiniaria des Sudwestatlantik I. Hormathiidae. Helgolander wiss. Meeresunters, 25, 273 - 325. https: // doi. org / 10.1007 / BF 01611200","Doumenc, D. (1975) Actinies bathyales et abyssales de l'ocean Atlantique nord. Familles des Hormathiidae (genres Paracalliactis et Phelliactis) et des Actinostolidae (genres Actinoscyphia et Sicyonis). Bulletin du Museum National d'Histoire Naturelle, Paris, 197, 157 - 204.","Fautin, D. G., Daly, M. & Cappola, V. (2005) Sea anemones (Cnidaria: Actiniaria) of the Faroe Islands: a preliminary list and biogeographic context. BIOFAR Proceedings, 2005, 77 - 87.","Molodtsova, T. N., Sanamyan, N. P. & Keller, N. B. (2008) Anthozoa from the northern Mid-Atlantic Ridge and Charlie-Gibbs fracture zone. Marine Biology Research, 4 (1 - 2), 112 - 130. https: // doi. org / 10.1080 / 17451000701821744","Van Praet, M., Rice, A. L. & Thurston, M. H. (1990) Reproduction in two deep-sea anemones (Actiniaria); Phelliactis hertwigi and P. robusta. Progress in Oceanography, 24 (1 - 4), 207 - 222. https: // doi. org / 10.1016 / 0079 - 6611 (90) 90031 - V"]}

Paraphelliactis labiata n. sp. (Figs. 2A, 3–5, Tables 1–3) urn:lsid:zoobank.org:act: 72B664B0-6E14-4610-99FF-42374AD8B8F1 Material examined. Holotype: MNRJ 9095 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT84 (04° 25.8308’ S, 036° 37.3678’ W), May 6, 2011, 1964-2045m. Paratypes: MOUFPE- CNI 868 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04° 21.3580’S, 036° 44.2730’W), May 4, 2011, 2025–2057m. LC 141 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04° 21.3580’S, 036° 44.2730’W), May 4, 2011, 2025–2057m Description. External morphology (Figs. 2A, 3). Column 11 mm to 84 mm in height and 12 mm to 99 mm in width (Figs. 2A, 3 A–B). Body cylindrical, wider than tall. Color a pure white in most of the column, abruptly changing to pale pink towards the aboral side. Base spreads beyond a tapered end but never exceeds the maximum diameter of the animal, hence the body takes a very singular cup shape no matter its size. Column divisible into a strongly tuberculated scapus and a short, smooth scapulus, without cinclides. Tubercles of the scapus conical, pointed, some with very flat with tips withdrawn and longitudinal grooves, not ordered in rows (Fig. 3 A–B). Tubercles are larger at midcolumn and towards the upper scapus, although smaller tubercles are also present near the limbus and at the scapus-scapulus boundary (Fig. 3C). A yellowish cuticle visible between the tubercles in some places. Scapulus thin and concealed due to the retraction of the oral disc. Endocoelic and exocoelic spaces visible as rays running from scapulus to mouth. Oral disc bilobed and broad, with a central oval mouth; mouth remains wide open. Lobes of oral disc unequal in size, usually overlapping (Fig. 3B). Tentacles slender, short, about 140 in number, without basal mesogleal thickenings, arranged in five peripheral cycles hidden inside the terminal fold of the scapus. Those of the inner cycles longer than those of the outer cycles. In the smaller individual, tentacles number approximately 90, with some clearly missing. Internal anatomy (Fig. 4). Two deep siphonoglyphs each attached to a pair of directives (Fig. 4C). Slender mesenteries, hexamerously arranged in five cycles (6+6+12+n+n). First three cycles complete, with all mesenteries present. In counterpart, fourth and fifth cycles incomplete, with the number of mesenteries varying among individuals, but always fewer than expected for a regular hexamerous arrangement. More mesenteries distally than proxi- mally. In the smaller individual, there are 80 near the margin, and 72 at the base level and in the larger individual there are approximately 100 at the margin and 78 to 81 at the limbus. Mesenterial filaments more developed on the older mesenteries than on younger ones. The incomplete fourth and fifth cycles have very small mesenteries. The last one is visible as tiny, unpaired projections, merely breaking through the mesoglea and present throughout the column (Fig. 4 D-E). Mesoglea of equal thickness in both endocoelic and exocoelic spaces at the oral disc region. Sphincter mesogleal, weak, restricted to the central part of the mesoglea where it occupies about a quarter of its space at the upper portion but abruptly tapers downwards (Fig. 4A). Muscular processes of the sphincter form somewhat sparse, not very marked bands, tending to align vertically. Longitudinal muscles of the tentacles weak, ectodermal (Fig. 4B). Retractor weak, diffuse, stronger at the distal part of the mesenteries (Fig. 4D). Parietobasilar muscles weak. Radial musculature of the oral disc ectodermal. Acontia present but rare, probably associated with the second cycle of mesenteries. No gametogenic tissues found. Cnidom. Spirocysts, Basitrichs, Microbasic p -mastigophores (Fig. 5, Table 2). Type locality. Potiguar Basin, Rio Grande do Norte, Brazil. Biological characteristics. Individuals originally attached to manganese nodules or slivers of wood (Fig. 3D). Etymology. The specific epithet “labiata” (from Latin: lips) alludes to the bilobed appearance of the oral disc, resembling a set of lips. Taxonomic remarks. The presence of a thick cuticle covering most of the body wall has been reported from all species of Paraphelliactis. Although small fragments of a yellowish cuticle were observed in our specimens, these fragments were few and restricted to the limbus and some cracks between the tubercles. The almost complete lack of cuticle could be explained by high abrasion from sediment, due to the method of collection. Carlgren (1949) stated that Paraphelliactis probably possess more tentacles than mesenteries at the base. An interesting feature not mentioned by Carlgren’s key (1949) but reported for most of Paraphelliactis species is the occurrence of an incomplete fifth cycle of mesenteries (except for Pa. tangi Li & Xu, 2016, which exhibits all five cycles complete). Li & Xu (2016) describe a small paratype for Pa. tangi, with only four cycles of mesenteries. However, this last cycle is fully complete, corroborating the distinct pattern of mesenterial development for the species and showing that the total numbers of such structures may vary according to the individual life stages. Although all our specimens differ in size and development, they have identical body shapes and share the same mesenterial arrangement pattern (five cycles with the last two incomplete), due to this is very unlikely that Pa. labiata is a juvenile of another species of the genus. Comparison of Pa. labiata n. sp. with other species of Paraphelliactis The type species of Paraphelliactis, Pa. spinosa Carlgren, 1928, and the later-described Pa. michaelsarsi Carlgren, 1934, were originally recorded for the North Atlantic Ocean. Two more species were registered from the Pacific Ocean, Pa. pabista Dunn, 1982 from the west coast of North America and Pa. tangi, collected near the Yap Trench at the western Pacific, totaling four valid species within the genus. This newly described species differs from its congeners in having the last two cycles of mesenteries incomplete, rather than just the fifth. In addition, the number of tentacles combined with the arrangement of the tubercles, and also the absence of basal thickenings in the tentacles differentiates the new species from its congeners (see Table 3). Paraphelliactis labiata n. sp. does not exhibit the characteristic hooked tubercles of Pa. spinosa. Although well detailed, the original description of Pa. michaelsarsi was of a single specimen in very poor preservation, and is therefore limited in some aspects, as pointed by the author himself (Carlgren, 1934). Nonetheless, Pa. labiata n. sp. differs from Pa. michaelsarsi by the presence of a smaller category of nematocysts (basitrichs) in the actinopharynx and tentacles and by possessing a very clear differentiation between the scapus and scapulus region vs. indistinct differentiation of the column in Pa. michaelsarsi. Paraphelliactis labiata n. sp. can be easily distinguished from Pa. pabista on the arrangement of the tubercles (not arranged in rows vs. arranged in rows in the latter). Our new species share a few characters with the lately described Pa. tangi (e.g. the absence of mesogleal thickenings at the base of the tentacles), but the latter is distinct from all the other species of Paraphelliactis, including Pa. labiata, in having an equal number of mesenteries throughout the column and a complete fifth cycle of mesenteries.

Monactis vestita (Gravier, 1918) (Figs. 2B, 6 B-C, Table 4) Paractis vestita Gravier, 1918 Monactis vestita: Riemann-Zürneck 1986; Zamponi & Acuña 1992; White et al. 1999; Deserti et al. 2012 Material examined. MNRJ 9082 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station Teste1000 (04° 9.3700’S, 036° 50.5998’W), April 30, 2011, 890– 900 m. MNRJ 9090 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 62 (04° 44.1888’S, 036° 24.9309’W), May 7, 2011, 425– 448 m. MOUFPE-CNI 869 (ten specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station Teste1000 (04° 9.3700’S, 036° 50.5998’W), April 30, 2011, 890– 900 m. MOUFPE-CNI 870 (eleven specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 62 (04° 44.1888’S, 036° 24.9309’W), May 7, 2011, 425– 448 m. MOUFPE-CNI 871 (four specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 84 (04° 25.8308’S, 036° 37.3678’W), May 6, 2011, 1964–2045 m. MOUFPE-CNI 872 (three specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 (04° 40.1817’S, 036° 23.8647’W), May 7, 2011, 897– 908 m. MOUFPE-CNI 873 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 (04° 28.3642’S, 036° 24.7602’W), May 4, 2011, 1880–1950 m. LC 142 (eight specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station Teste1000 (04° 9.3700’S, 036° 50.5998’W), April 30, 2011, 890– 900 m. LC 143 (eleven specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 62 (04° 44.1888’S, 036° 24.9309’W), May 7, 2011, 425– 448 m. LC 144 (eleven specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 63 (04° 41.7490’S, 036° 31.1670’W), May 8, 2011, 375– 383 m. LC 145 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 (04° 40.1817’S, 036° 23.8647’W), May 7, 2011, 897– 908 m. LC 146 (four specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 74 (04° 34.1484’S, 036° 41.6035’W), May 7, 2011, 902– 1073 m. LC 147 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 81 (04° 39.3740’S, 036° 4.6810’W), May 6, 2011, 1960–2003 m. LC 148 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 (04° 28.3642’S, 036° 24.7602’W), May 4, 2011, 1880–1950 m. LC 149 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83-2 (04° 27.0256’S, 036° 25.6086’W), May 21, 2011, 1896–1931 m. LC 150 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 84 (04° 25.8308’S, 036° 37.3678’W), May 6, 2011, 1964– 2045 m. LC 151 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04° 21.3580’S, 036° 44.2730’W), May 4, 2011, 2025–2057 m. Description. Column cylindrical and smooth, without any projections and not divided into regions. Body often very short and broad due to the distension of the pedal disc (Fig. 6B). Polyps ranging from 2 mm to 9 mm in height and exceeding 40 mm in diameter. Oral disc withdrawn, visible only in cross sections. Thirty-two tentacles, conical, not very long, tapering to a pointed tip and arranged in two regular cycles (16+16). Tentacles of the inner cycle lon- ger than those of the outer one. No cinclides. Two siphonoglyphs, each attached to a pair of directives. Mesenteries, gametogenic tissue and other inner structures perfectly visible through the thin, distended column. Mesenteries hexamerously arranged in four cycles (6 + 6 + 12 + 24) (Fig. 6C). Only the first cycle perfect and bearing acontia. Acontia small and rare, paler than the mesenterial filaments, lacking completely in some specimens. Specimens with only male or female gametogenic tissue. Sphincter strong, mesogleal, alveolar. Longitudinal muscle of the tentacles ectodermal. Retractor strong and diffuse. Individuals attached to octocoral branches, rocks, or mollusk shells. Cnidom. Spirocysts, Basitrichs, Microbasic p -mastigophores (Table 4). Remarks. Some individuals have the column remarkably distended, especially those attached to big rocks, where the pedal disc sometimes extends beyond its borders, contracting the rest of the column considerably (Fig. 6B). White et al. (1999) indicated high variability of the polyp appearance corresponding to the type of substrata where it is found. In this study, the authors hypothesized that M. vestita is only associated with snails shells in the Pacific Ocean, since previous records in the Atlantic Ocean found it only attached to stones. However, the species have been collected by-catch in Uruguay associated with banks of bivalves (Deserti et al. 2012), and we register for the first time M. vestita on Fissidentalium meridionale scaphopod shells, as well as on those of Drillia sp. (empty shells), supporting the high adaptability of this species to different benthic communities. Regarding the cnidom, we also found two types of spirocysts on tentacles (gracile and robust) similar to White et al. (1999), but unlike them we found three size classes of basitrichs in the tentacles, similar to Riemann-Zürneck (1986). We give the first comprehensive cnidom for specimens from Atlantic Ocean. M. vestita has a large bathymetric range from 200m (recorded in Argentina) to more than 5000m (in northeastern Atlantic Ocean). Geographic distribution. Atlantic Ocean: northeastern Atlantic Ocean, Gulf of Mexico, Venezuela, Uruguay and Argentina (Gravier 1918; Riemann-Zürneck 1986;, Zamponi & Acuña 1992; Deserti et al. 2012). Pacific Ocean: Oregon (White et al. 1999). This study registers M. vestita for the first time in the Brazilian coast (Bacia Potiguar, RN)., Published as part of De Melo, Yago A., Targino, Alessandra K. G. & Gomes, Paula B., 2020, New records of family Hormathiidae (Cnidaria: Actiniaria) from Brazilian coast with description of Paraphelliactis labiata n. sp., pp. 557-574 in Zootaxa 4766 (4) on pages 565-567, DOI: 10.11646/zootaxa.4766.4.3, http://zenodo.org/record/3765725, {"references":["Gravier, C. (1918) Note preliminaire sur les hexactiniaires recueillis au cours des croisieres de la Princesse-Alice et de l'Hirondelle de 1888 a 1913 inclusivement. Bulletin de l'Institut Oceanographique, Monaco, 346, 1 - 24. https: // doi. org / 10.5962 / bhl. part. 8664","Riemann-Zurneck, K. (1986) Zur Biogeographie des Sudwestatlantik mit besonderer Berucksichtigung der Seeanemonen (Coelenterata: Actiniaria). Helgolalander Meeresuntersuchungen, 40, 91 - 149. https: // doi. org / 10.1007 / BF 01987291","Zamponi, M. O. & Acuna, F. H. (1992) Sobre las caracteristicas gonadales de Monactis vestita (Gravier, 1918), Sensu Riemann- Zurneck, 1986 (Actiniaria, Hormathiidae). Iheringia, Serie Zoology, 72, 151 - 152.","White, T. R., Wakefield Pagels, A. K. & Fautin, D. G. (1999) Abyssal sea anemones (Cnidaria, Anthozoa) of the northeast Pacific symbiotic with molluscs: Anthosactis nomados, a new species, and Monactis vestita (Gravier, 1918). Proceedings of the Biological Society of Washington, 112 (4), 637 - 651.","Deserti, M. I., Zamponi, M. O. & Riestra, G. (2012) Las anemonas de mar (Cnidaria; Anthozoa; Actiniaria) de la plataforma continental uruguaya. Revista Real Academia Galega de Ciencias, 31, 115 - 136."]}

Genus Monactis Riemann-Zürneck, 1986 Diagnosis (see Riemann-Zürneck, 1986) Type species: Paractis vestita Gravier, 1918, Published as part of De Melo, Yago A., Targino, Alessandra K. G. & Gomes, Paula B., 2020, New records of family Hormathiidae (Cnidaria: Actiniaria) from Brazilian coast with description of Paraphelliactis labiata n. sp., pp. 557-574 in Zootaxa 4766 (4) on page 565, DOI: 10.11646/zootaxa.4766.4.3, http://zenodo.org/record/3765725, {"references":["Riemann-Zurneck, K. (1986) Zur Biogeographie des Sudwestatlantik mit besonderer Berucksichtigung der Seeanemonen (Coelenterata: Actiniaria). Helgolalander Meeresuntersuchungen, 40, 91 - 149. https: // doi. org / 10.1007 / BF 01987291","Gravier, C. (1918) Note preliminaire sur les hexactiniaires recueillis au cours des croisieres de la Princesse-Alice et de l'Hirondelle de 1888 a 1913 inclusivement. Bulletin de l'Institut Oceanographique, Monaco, 346, 1 - 24. https: // doi. org / 10.5962 / bhl. part. 8664"]}