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A new species of Meconematini is described from Tabin Wildlife Reserve in eastern Sabah: Cercoteratura repens sp. nov. This is the first report of the genus Cercoteratura in Sabah. Two species of Phisidini, Carliphisis acutipennis (Carl, 1908) and Neophisis (Indophisis) longipennis Jin, 1992, are reported in eastern Sabah for the first time. The calling song of Neophisis (Indophisis) longipennis with a near-complete ultrasonic spectrum is also described.

Systolederus cinereus Brunner von Wattenwyl, 1893 (Figs. 18–20) Systolederus cinereus Brunner von Wattenwyl, 1893: 105 (holotype —female, Myanmar, Carin Cheba [= Karen Hills]; in the Natural History Museum of Genova). Systolederus cinereus — Kirby, 1910: 20; Kirby, 1914: 44; Günther, 1939: 170; Blackith, 1992: 171; Mahmood et al., 2007: 1282; Storozhenko & Dawwrueng, 2015: 536. Systolederus ridleyi Hancock, 1909: 401 (holotype —female, Singapore, Singapore Botanical Garden; in the Oxford University Museum of Natural History, UK), syn. nov. Teredorus ridleyi — Hancock, 1915: 110. Systolederus ridleyi — Günther, 1939: 169; Blackith, 1992: 174; Devriese & Husemann, 2023: 340. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •5Ô and 3♀; open stream near Bạch Mã National Park, N16.22036 E107.88702, 72.9± 6.3 m.a.s.l.; 8 March 2023, 11h01; on a boulder or rock; coll. M.K. Tan & V. T. Trung; VIET. 23.31–38 (ZRC) •1Ô; along stream near Bạch Mã National Park, N16.22419 E107.88233, 57.4± 4.5 m.a.s.l.; 8 March 2023, 19h37; on a tree trunk; coll. M.K. Tan & V. T. Trung; VIET.23.50 (VNMN). Remarks. Images of the holotypes of C. cinereus and C. ridleyi are available in the internet catalogue Orthoptera Species File (Cigliano et al., 2023). The comparison of original descriptions and photos of both species, as well as all materials examined by Sergey Yu. Storozhenko from Southeast Asia, allows proposing a new synonymy: Systolederus cinereus Brunner von Wattenwyl, 1893 = Systolederus ridleyi Hancock, 1909, syn. nov. The specimens from Bạch Mã National Park represent the first record of this species in Vietnam. Ecology. This species was found along rivers, on rocks and substrate near the fast-flowing water. Nevertheless, it has also been found along forest edges near a river and dwelling among tree trunks. Distribution. Myanmar, Thailand, Malaysia, Singapore; Vietnam: Thừa Thiên Huễ Province, Bạch Mã National Park [new]. Type locality. Myanmar, Carin Cheba.

Tan, Ming Kai, Vu, Trung Thanh, Le, Cuong Viet Canh, Pham, Thai Hong (2023): Pygmy grasshoppers (Orthoptera: Tetrigidae) from Bạch Mã National Park, central Vietnam. Zootaxa 5319 (2): 199-223, DOI: 10.11646/zootaxa.5319.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5319.2.3

Zhengitettix albitarsus Storozhenko, 2013 (Fig. 11) Zhengitettix albitarsus Storozhenko, 2013: 206. Zhengitettix albitarsus — Storozhenko & Dawwrueng, 2014: 325; Dawwrueng & Dooddeum, 2014: 5; Storozhenko & Dawwrueng, 2015: 548; Chen & Deng, 2022: 73 (key to species of Zhengitettix). Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •1Ô; open stream near Bạch Mã National Park, N16.22036 E107.88702, 72.9± 6.3 m.a.s.l.; 8 March 2023, 11h01; on a boulder or rock; coll. M.K. Tan & V. T. Trung; VIET.23.30 (ZRC) (Fig. 11). Remarks. Previously described and known only from Thailand, the male specimen from Bạch Mã National Park represents the first record of this species in Vietnam (Fig. 11). Ecology. This species was found along rivers, on rocks and substrate near the fast-flowing water (Fig. 3). Upon disturbance, this species would fly off. It is likely to be limno-terrestrial, being associated with open stream. Distribution. Thailand: Nakhon Ratchasima Province, Mae Hong Son Province; Vietnam: Thừa Thiên Huễ Province [new]. Type locality. Thailand, Nakhon Ratchasima Province, Khao Yai.

Epitettix parallelus (Podgornaya, 1986) (Fig. 12) Vaotettix parallelus Podgornaya, 1986: 17. Vaotettix parallelus — Blackith, 1992: 196; Kim & Pham, 2014: 59. Epitettix parallelus — Storozhenko, 2021: 182. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •1Ô; Rhododendron Trail, heading to Four Lakes, N16.18950 E107.85360, 1160.0± 7.7 m.a.s.l.; 9 March 2023, 11h02; among leaf litter; coll. M.K. Tan & V. T. Trung; VIET.23.63 (ZRC) (Fig. 12). Remarks. This species is widely distributed in Vietnam. The specimen from Bạch Mã National Park, nevertheless, differs by the emarginated apex of posterior process of the pronotum (in typical form apex of process rounded) (Fig. 12). This is likely intraspecific variability. Ecology. The leaf litter area at which the specimen was collected was not too far from a forest stream, but the area is clearly elevated and probably not part of the water table. Distribution. So far, known only from central Vietnam in the provinces of Gia Lai and Thừa Thiên Huễ [new]. Type locality. Vietnam, Gia Lai Province, Buon Luoi.

Bolivaritettix sculptus (Bolívar, 1887) (Fig. 14) Mazarredia sculpta Bolívar, 1887: 237. Mazarredia sculpta — Hancock, 1907b: 32; Kirby, 1914: 51; Hancock, 1915: 100; Bolívar, 1917: 285. Bolivaritettix sculptus — Günther, 1939: 59; Shishodia, 1991: 82; Blackith, 1992: 12; Paris, 1994: 249; Deng et al., 2009: 282; Deng et al., 2010: 48; Storozhenko & Dawwrueng, 2015: 533; Ding et al., 2018: 311; Storozhenko, 2018a: 18; Bhaskar et al., 2022: 422. Mazarredia indotata Bolívar, 1909: 398; synonymized by Günther, 1939: 59. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: • 1♀; Summit Trail (forest trail), N16.19735 E107.86131, 1401.0±6.0 m.a.s.l.; 7 March 2023, 11h42; on clayey rock; coll. M.K. Tan & V. T. Trung; VIET.23.16 (ZRC) (Fig. 14). Ecology. The colouration of this species allows it to camouflage among orange clayey rock where it was found. Distribution. India, China, Myanmar, Cambodia, Thailand, Vietnam: provinces Gia Lai, Vĩnh Phúc, Bình Phýớc, Đắk Lắk, and Thừa Thiên Huễ [new]. Type locality. Vietnam, Gia Lai Province, Buon Luoi.

Hebarditettix magnus Storozhenko, 2017 (Figs. 6, 7) Hebarditettix magnus Storozhenko, 2017: 121. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: • 1♀; Summit Trail (forest trail), N16.19802 E107.85991, 1457.0± 5.9 m.a.s.l.; 7 March 2023, 21h12; on a vertical surface; coll. M.K. Tan & V. T. Trung; VIET.23.26 (ZRC) •1Ô; Summit Trail (forest trail), N16.19829 E07.85970, 1447.0±6.0 m.a.s.l.; 7 March 2023, 21h20; on a tree trunk; coll. M.K. Tan & V. T. Trung; VIET.23.27 (VNMN) (Fig. 7). Remarks. This species was first described from Bạch Mã National Park. This represents the second record of this species. We provide a live image of this species (Fig. 6). Ecology. This species was found among shaded forested habitat (Fig. 2), often dwelling among tree trunks or rocky surfaces (covered with some mosses) (Fig. 6). It seems that this species tends to be found at higher altitudes (see Storozhenko, 2017). Distribution. So far, known only from central Vietnam in the provinces of Thừa Thiên Huễ and Gia Lai. Type locality. Vietnam, Th ừa Thiên Hu ễ Province, B ạch Mã National Park.

Criotettix bispinosus (Dalman, 1818) (Figs. 4, 5) Acrydium bispinosum Dalman, 1818: 77. Acridium (Tetrix) bispinosum — Haan, 1843: 169. Criotettix bispinosus [refer to OSF for complete literature]— Bolívar, 1887: 226; Günther, 1937a: 179; Tinkham, 1937: 230; Günther, 1937b: 121–122; Günther, 1939: 319; Shishodia, 1991: 36; Blackith, 1992: 34; Mahmood et al., 2007: 1281; Benediktov. 2013: 257 (vibrational signalling); Tumbrinck, 2015: 276; Deng et al., 2023: 208. Acanthalobus bispinosus — Hancock, 1907b: 29; Kirby 1910: 16; Kirby, 1914: 37; Karny, 1915: 78; Hancock, 1915: 93; Willemse, 1930: 18. Criotettix brachynotus Zheng & Jiang, 1994: 34; synonymized by Deng, 2021: 455. Formosatettix hainanensis Zheng, 2012: 2; synonymized by Deng et al., 2023: 2008. Tettix latispina Walker, 1871: 837; synonymized by Kirby, 1910: 18. Acanthalobus longinotus Hancock, 1907a: 221; synonymized by Blackith, 1992: 35. Tettix simplex Walker, 1871: 836; synonymized by Kirby, 1910: 18. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •1Ô, 1♀; road-side vegetation along road to summit, N16.24626 E107.86881, 85.9± 6.1 m.a.sl., 7 March 2023, 9h16; on a leaf of a shrub; coll. M.K. Tan & V. T. Trung; VIET. 23.12, 13 (ZRC) • 1♀; road-side vegetation along road to summit, N16.24198 E107.86661, 144.7± 6.9 m.a.s.l.; 7 March 2023, 10h09; on a rock with flowing water; coll. M.K. Tan & V. T. Trung; VIET.23.15 (VNMN) • 3Ô; open stream near Bạch Mã National Park, N16.22036 E107.88702, 72.9± 6.3 m.a.s.l.; 8 March 2023, 11h01; on boulders and rocks; coll. M.K. Tan & V. T. Trung; VIET.23.39–41 (ZRC) (Fig. 5). Ecology. This species was found along rivers, residing on rocks and substrate near the fast-flowing water. However, it has also been found on more vegetated area along road. Distribution. India, Myanmar, South China (including Hainan), Thailand, Vietnam, Peninsular Malaysia, and Indonesia (the Greater Sunda Islands). Type locality. Oriental region (without definite locality).

Bolivaritettix chinensis (Hancock, 1912) (Fig. 13) Mazarredia chinensis Hancock, 1912: 140. Bolivaritettix chinensis — Günther, 1939: 59; Günther, 1941: 149; Kim & Pham, 2014: 59; Storozhenko, 2018a: 504. Material examined. VIETNAM, Thừa Thiên Huễ province, Bạch Mã National Park: • 1♀; 22–23 September 2008, coll. V. G. Bezborodov (Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia). Remarks. This species was reported in Bạch Mã National Park (Storozhenko, 2018a). Distribution. Vietnam: provinces Cao Bằng, Hòa Bình, Thừa Thiên Huễ, Lang Son, Thái Nguyên, Vĩnh Phúc. Type locality. Tonkin, Than Moi (northern Vietnam).

Tegotettix bufocrocodil (Storozhenko & Dawwrueng, 2015) (Figs. 8–10) Gavialidium bufocrocodil Storozhenko & Dawwrueng, 2015: 540. Gavialidium bufocrocodil — Storozhenko, 2018b: 19. Tegotettix bufocrocodil — Muhammad et al., 2018: 31. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •1Ô, 4♀, 3 nymphs; along stream near Bạch Mã National Park, N16.22419 E107.88233, 57.4± 4.5 m.a.s.l.; 8 March 2023, 19h37; on a tree trunk; coll. M.K. Tan & V. T. Trung; VIET. 23.42–49 (ZRC) •1Ô; forest near Bạch Mã National Park, N16.22069 E107.88253, 74.9± 6.5 m.a.s.l.; 9 March 2023, 21h11; on a tree trunk; coll. M.K. Tan & V. T. Trung; VIET.23.73 (VNMN). Remarks. This represents the first record of this species in Vietnam. Ecology. This species was found along forest edge not too far from a river and dwelling among tree trunks, often staying still and camouflaging among the crevices (Figs. 8–10). Adults and nymphs can be found aggregating together on a few trees. Distribution. Thailand: Nakhon Ratchasima and Trat Provinces; Cambodia: Sen Monorom; Vietnam: Thừa Thiên Huễ Province [new]. Type locality. Thailand, Nakhon Ratchasima Province, Khao Yai.

Rhopalina bachma Tan & Storozhenko, sp. nov. (Fig. 17) Material examined. VIETNAM, Th ừa Thiên Hu ễ Province, B ạch Mã National Park: • ♀ holotype; Rhododendron Trail, heading to Four Lakes, N16.18950 E107.85360, 1160.0± 7.7 m.a.s.l.; 9 March 2023, 11h02; among leaf litter; coll. M.K. Tan & V. T. Trung; VIET.23.62 (VNMN) (Fig. 17). Remarks. This represents the second species from this little-known genus. The previously monotypic genus consisted of Rhopalina javana Tinkham, 1939 from Java. Here, a second species from this genus was discovered in Indochina (quite far away from Java) and is described. The fastigium of the holotype has its right margin rounded, but its left margin emarginated in the middle. It is unclear if this asymmetry is due to defects (e.g., damaged during individual development), injuries or genuine character. Diagnosis. The new species is similar to Rhopalina javana in the general habitus, colouration and the shape of fastigium but differs from latter by the antennal groove inserted at the lower margins of the eyes, the tegmen stouter, 2.4–2.5 times as long as wide, and the pronotum well-surpassing apices of ovipositor (in R. javana, the antennal groove instead of above lower margins of the eyes, the tegmen 2.6–2.7 times as long as wide, and the pronotum slightly surpassing apices of ovipositor). Etymology. The species is named after the type locality, Bạch Mã National Park; noun in apposition. Description. Habitus as shown in Figs. 17A, 17B. Different shades of brown, ranging from pale brown, dark brown to red brown, well camouflaged against leaf litter on the ground. Head: In frontal view (Fig. 17C): antennal groove inserted at lower margin of eyes. Fastigium produced into obtuse apex. Face with numerous small nodules and large oval nodules above clypeal triangle; with robust lateral carinae running from ventrad of eyes to labrum, slightly bent in middle. Frontal costa elongated, bifurcation of frontal costa in line with slightly dorsal of middle of eye. Facial carinae straight and slightly diverging ventrally. Compound eyes hemispheric, not exerted above vertex, 1.41 times taller than wide. Lateral ocelli slightly ventral of bifurcation of frontal costa and slightly above antennal groove in line with middle of eye. In dorsal view (Fig. 17A): apex of fastigium surpasses frontal margin of eyes; right margin of fastigium rounded, left margin emarginated in the middle. Vertex 1.47 times wider than eye width. Lateral carinae of fastigium not distinct; median carina more distinct, reaching apex of fastigium. In lateral view (Fig. 17B): frontal costa well protruding in front of eyes into obtuse apex. Mouthparts with labrum pale coloured. Maxillary palps pale coloured with tint of grey on anterior (or dorsal parts), elongated; with apical (fifth) segment more oval, longest following by third segment and then subapical (fourth) segment. Pronotum: Pronotum surpassing posterior femur, nearly reaching middle of posterior tibia, 3.26 times longer than wide (pronotal lateral lobe width). In lateral view (Fig. 17B): with median, humero-apical and interhumeral carina weakly produced. Infrascapular area broad. Angle of ventral sinus between lateral shoulder and lateral lobe nearly right-angled. In dorsal view (Fig. 17A): median carina distinct throughout length of pronotum (except very close to the posterior end). Lateral carinae distinct. Interhumeral carina distinct, diverging posteriorly. Lateral lobe of pronotum slightly produced, and broadly triangular; posterior margin of lateral lobe acute. Apical end of pronotal disc narrow and truncated. Legs: Fore and middle legs (Fig. 17A): femora generally marmoration of pale and dark spots; tibiae with dark and light bands. Hind legs (Fig. 17B): hind femur about 4.29 times longer than wide, about 1.09 times longer than hind tibia; with clear carinae along dorsal margin; marmoration of pale and dark spots and ventrad of ventroexternal carina dark coloured. Knee of posterior femur with antegenicular and genicular teeth acute. Hind tibia dark coloured with shades of pale colouration; with six spines on each lateral margin, with numerous spinules between these large dorsal spines. Basal article of hind tarsus generally pale coloured, 1.38 times longer than apical article; middle article dark coloured very short; apical article white, except apex being slightly darker in colouration. Abdomen: Tergites and sternites typically dark coloured with pale-coloured nodules. Ovipositor (Fig. 17D) with pale-coloured spinules with apex red brown; dorsal and ventral valves with sparse hairs along dorsal and ventral margin. Dorsal valve with eight dorsal spines, apex acute; ventral valve with six ventral spines typically smaller than dorsal spines; apical ventral spine of ventral valve more hooked at apex than apical dorsal spine of dorsal valve; spines on valves increasing larger and robust apically. Apices of ovipositor valves red brown. Subgenital plate short in lateral view (Fig. 17D), truncated with medial lobule in ventral view (Fig. 17E). Measurements (in mm). PL = 10.34; PW = 3.18; PH = 1.73; HWL = 8.13; HFL = 5.10; HFW = 1.19; TL = 1.20; VW = 0.66; EW = 0.45; SW = 0.19; IAH = 0.22; BL = 8.16; PAW = 1.45; PPW = 1.99; MAL = 0.36; MAW = 0.05; 1 stAW = 0.20; TW = 0.55; FFL = 1.92; FFW = 0.53; MFL = 2.09; MFW = 0.54; HTL = 4.67; 1 st TaL = 0.87; 3 rd TaL = 0.63; SGPL = 0.67; SGPW = 0.54; OVDL = 1.02; OVDW = 0.31; OVVL = 1.01; OVVW = 0.25 Ecology. The leaf litter area at which the specimen was collected was not too far from a forest stream, but the area is clearly elevated and probably not part of the water table. Distribution. So far known only from type locality. Type locality. Vietnam, Th ừa Thiên Hu ễ Province, B ạch Mã National Park.

Miriatroides luna Tan & Storozhenko, sp. nov. (Figs. 15, 16) Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •Ô holotype; Summit Trail (forest trail), N16.19667 E107.86169, 1428.0± 5.8 m.a.s.l.; 7 March 2023, 20h15; hiding among rotten tree trunk on the ground; coll. M.K. Tan & V. T. Trung; VIET.23.17 (VNMN) (Figs. 15, 16). Diagnosis. This new species is most similar to Miriatroides kannackiensis Storozhenko, 2016 from Gia Lai Province, but differs from latter (as well as from all other congeners) by the rostrum more projected in lateral view and the fore femur with margins (especially ventral margin) undulated. Etymology. This species is named after the moon (in Latin, luna = moon), as the holotype was collected near the summit on a clear night with a full moon. Description. Habitus as shown in Figs. 15, 16A, 16B. Different shades of brown, ranging from pale brown, and dark brown to red brown, well camouflaged against leaf litter and tree trunk on the ground. Head: In frontal view (Fig. 16C): antennal groove inserted below lower margin of eyes. Fastigium convex; sinuosoidal on anterior border. Face with numerous nodules. Frontal costa elongated. Bifurcation of frontal costa in line with middle of eye. Facial carinae faintly sinusoidal. Compound eyes hemispheric, not exerted above vertex, 1.55 times taller than wide. Lateral ocelli slightly ventral of bifurcation of frontal costa and slightly above antennal groove. In dorsal view (Fig. 16A): apex of fastigium clearly surpasses frontal margin of eyes, emarginated in the middle. Vertex 1.53 times wider than eye width. Lateral carinae of fastigium not distinct; median carina well surpassing apex of fastigium. In lateral view: frontal costa arched and well protruding in front of eyes. Mouthparts mostly brown to grey with small pale-coloured nodules. Maxillary palps pale coloured with tint of grey; with segments stout and apical segment somewhat oval. Pronotum: Pronotum reaching apex of posterior knee, 2.03 times longer than wide (pronotal lateral lobe width). In lateral view (Fig. 16B): with median, humero-apical, and interhumeral carinae irregularly undulated. Infrascapular area broad. Angle of ventral sinus between lateral shoulder and lateral lobe acute. In dorsal view (Fig. 16A): median carina distinct throughout length of pronotum, flanked by a network of irregular carinae. Lateral carinae distinct. Interhumeral carina distinct, diverging posteriorly. In dorsal view (Fig. 16A), lower margin of lateral lobe of pronotum leaf-shaped. Apical end of pronotal disc acute. Legs (Figs. 16A, 16B): Fore and middle legs: generally brown with either dark- or light-coloured bands. Fore femur with distinct dorsal carina straight, with ventral carina undulated, forming three nodules. Middle femur with less distinct dorsal and ventral carinae, clearly not undulating; lateral external carina undulated, forming two or three small nodules. Hind legs: posterior femur about 3.36 times longer than wide, about 1.34 times longer than hind tibia; with clear carinae along dorsal margin. Knee of hind femur with antegenicular tooth small but acute, genicular tooth rounded. Posterior tibia dark coloured anteriorly, becoming slightly lighter in colouration; with five spines on each lateral margin, with numerous spinules between these large dorsal spines. Basal article of hind tarsus generally pale coloured, 0.78 times as long as apical article; middle article dark coloured and very short; apical article white, except apex being black. Abdomen: Tergites and sternites typically dark coloured with some sparse patches of pale spots. Subgenital plate triangular (Figs. 16D, 16E). Measurements (in mm). PL = 7.17; PW = 3.54; PH = 1.20; HWL = NA; HFL = 4.53; HFW = 1.35; TL = NA; VW = 0.55; EW = 0.36; SW = 0.14; IAH = 0.52; BL = 7.34; PAW = 1.47; PPW = 2.16; MAL =?; MAW =?; 1 stAW = 0.18; TW = NA; FFL = 1.70; FFW = 0.55; MFL = 1.93; MFW = 0.44; HTL = 3.37; 1 st TaL = 0.49; 3 rd TaL = 0.63; SGPL = 0.56; SGPW = 0.59. Ecology. Only one specimen was encountered staying still and camouflaging itself in a decaying wood on the ground of a sheltered montane forest habitat at high altitude of Bạch Mã National Park (Fig. 3). Upon disturbance, it remained still. Distribution. So far known only from type locality. Type locality. Vietnam, Th ừa Thiên Hu ễ Province, B ạch Mã National Park.

Nisitrus danum Robillard & Tan, 2021 (Figs 1–8) Nisitrus danum Robillard & Tan, 2021, in Tan et al. 2021a: 31 Material examined MALAYSIA — Sabah • 1J; Kawag Forest Reserve, near rest house; N5.0502 E117.98286, 134.7 m.a.s.l. (GPS SAB22_6); 14.v.2022; afternoon; on plant; call recording MNHN-SO-2023-1478; M.K. Tan & T. Robillard; MNHN-EO-ENSIF11344 • 1♀; Kawag Forest Reserve, near rest house; N5.0502 E117.98286, 134.7 m.a.s.l. (GPS SAB22_ 6); 14.v.2022; afternoon; on plant; molecular sample N67; M.K. Tan & T. Robillard; MNHN-EO-ENSIF11345 • 1♀ (photographed, not collected);Tabin Wildlife Reserve,Lipad mud volcano; N5.20967 E118.50732, 168.9m.a.s.l. (GPS SAB22_11); 16.v.2022; day; T. Robillard observation • 2 juveniles; Tabin Wildlife Reserve; N5.19534 E118.50365, 119.7 m.a.s.l. (GPS SAB22_9); 15.v.2022, night; molecular samples N65–N66; MNHN-EO-ENSIF10978– ENSIF10979 • 1 juvenile (not collected); Sepilok, Rainforest Discovery Centre; N5.87428 E117.93970, 1.x.2019 (Fig. 3B) • 1 juvenile; Lahad Datu, N4.962598 E117.801956; 19.viii.2022; twan3253 observation; https://www. inaturalist.org/observations/133043313 • 1♀; Lahad Datu, N4.964485 E117.803471; twan3253 observation; https:// www.inaturalist.org/observations/133303271 • 1♀; Lahad Datu, N4.965797 E117.800246; twan3253 observation; https://www.inaturalist.org/observations/133431471 Additional description Habitus as shown in Fig. 1A. Face, lateral lobes of pronotum and wide lateral band on FWs vivid yellow in living specimens (Figs 1A, 1B). Antennae with 2 main white rings, including 3–6 antennomeres and 2–3 thinner rings. Male. Metanotal gland well developed, as shown in Fig. 1C. FWs (Figs 1D, 1E) dorsal field anterior part black (cells and veins); harp and mirror area infuriated, with black veins; apical field with black cells and yellow veins. Harp with two straight oblique veins. Mirror large, not rounded, separated by one transverse vein. Vein c1 wide. Lateral field with a wide vivid yellow band including veins M, R and Sc, and bases of veins bifurcating from Sc; ventral part of lateral field shiny black, as in females. Male genitalia (Fig. 2) typical of genus, including ectophallic fold, endophallic sclerite, ectophallic arc and rami, but with pseudepiphallic lophi triangular and relatively stout in dorsal and profile views, narrowly spaced between pseudepiphallic lophi; pseudepiphallic parameres strongly bent at interno-anterior end. Juvenile (Fig. 3). Colouration pale yellow or white, characterised by numerous dark stripes all around body in early instars; antennae with 2 wide whitish rings as in adults. Measurements (in mm, means in brackets, N = 5 males). BL = 13.9–15.3 (14.6), PronL = 1.8–2.1 (1.9), PronW = 2.7–3.1 (2.9), FWL = 10.1–11.0 (10.4), FWW = 3.8–4.1 (3.9), HWT = 4.5–7.6 (6.6), FIIIL = 13.4–15.5 (14.5), FIIIW = 2.5–2.9 (2.6), TIIIL = 13.5–15.5 (14.4), Stridulatory teeth = 94–95. Comparison with N. vittatus Larger, with dark shiny lateral field of FWs in both sexes, below a vivid yellow band; face yellow with dark brown to black mouthparts (entirely yellow with a few dark spots in N. vittatus); adults and juveniles (Fig. 3) are recognisable by the larger white rings on antennae (2 main rings, with 3–6 white antennomeres, while antennae are almost entirely black with in N. vittatus, with at best thin rings). Comparison with N. musicus Similar in size, but differing by dark shiny lateral field of FWs in both sexes, below a vivid yellow band; face vivid yellow (whitish in N. musicus). Ecology Found in syntopy with Nisitrus vittatus, the latter species being much more abundant, and has been observed to be outnumbered by at least 10 to 1. The males and females were found inter-mixing with N. vittatus on large foliage of Leea indica along roads (forest edge) (Fig. 4). Similar to N. vittatus, this species tends to occur in open areas (Fig. 5). Distribution (Fig. 6) Borneo (Sabah: Danum Valley, Tawau Hill Park, Kawag Forest Reserve [new locality record], Sepilok Kabili Forest Reserve [new locality record], Tabin Wildlife Reserve [new locality record]) Calling song (9♁, 23°C, in captivity) The calling song consists of long homogeneous bouts (= trill) which can last between 3 s to more than 16 min (Fig. 7). The trill is made up of a sequence of short echemes. Each echeme is made up of two syllables and has an average duration of 41.3±2.1 ms (36.6–45.9 ms). The average interval between consecutive echemes is 50.1±5.2 ms (42.6–62.1 ms). On average, the first syllable has a slightly shorter duration of 15.2±0.8 ms (13.2–17.3 ms) than the second syllable with a duration of 16.0±1.2 ms (12.6–18.7 ms). The average interval between the two syllables is 10.0±1.0 ms (7.9–12.3 ms). The call spectrum consists of a series of harmonics with three distinct peaks in the first three harmonics: first harmonic (fundamental frequency) at 6.13±0.14 kHz (5.72–6.47 kHz), which is typically also the dominant frequency; second harmonic at 12.21±0.31 kHz (11.34–12.84 kHz); third harmonic at 18.32±0.43 kHz (16.97–19.22 kHz). While second and third harmonics are not dominant, their amplitude is high and sometimes almost co-dominate with the fundamental frequency. The calling song of N. danum is very distinct from that of the syntopic species, Nisitrus vittatus (see Tan et al., 2021a), by the echeme structure being made up of two syllables, rather than three syllables. The echeme duration is also distinctly longer in N. danum than in N. vittatus (average duration = 41.3 ms vs. 32.5 ms). Consequently, the syllable durations are also longer in N. danum than in N. vittatus. The interval between consecutive echemes is also distinctly longer in N. danum than in N. vittatus (average duration = 50.1 ms vs. 22.7 ms). Although the frequencies of N. danum is lower than that of N. vittatus, it could be an artefact of the lower ambient temperature when N. danum was recorded (23°C for N. danum vs. 27°C for N. vittatus). Despite morphological similarities between N. danum and N. musicus (also from Sabah, but specifically from Mount Kinabalu), there are also differences in their calling song. Both species have similar echeme structure made up of two syllables, but the echeme duration is distinctly longer in N. danum than in N. muscius (average duration = 41.3 ms vs. 29.7 ms). While the durations of the first and second syllables in each echeme are somewhat similar in N. danum (15.2 ms and 16.0 ms), those in N. musicus are drastically different (7.5 ms and 18.7 ms). We found that N. danum is also a diurnal species, similar to other Nisitrus (Fig. 8). The circadian rhythm of the calling activity was as follows: calling began at 6:00h in the morning and its activity increases to a maximum at 8h in the morning (amount of sound produced per hour = ca. 1683 s). This follows by a drop in calling activity. A second smaller peak in calling activity (amount of sound produced per hour = ca. 915 s) was detected in the afternoon around 16:00h. Most calling activity ceases at 19:00h in the evening. The two diurnal peaks were also observed in N. vittatus (see Tan & Robillard, 2021b) but they differ in that the highest peak for N. danum was in the morning whereas that for N. vittatus was in the afternoon., Published as part of Robillard, Tony, Tan, Ming Kai, Japir, Razy & Chung, Arthur Y. C., 2023, Notes on the Eneopterinae (Orthoptera, Grylloidea, Gryllidae) from eastern Sabah, pp. 231-250 in Zootaxa 5315 (3) on pages 233-240, DOI: 10.11646/zootaxa.5315.3.2, http://zenodo.org/record/8130509, {"references":["Tan, M. K. & Robillard, T. (2021 b) Highly diversified circadian rhythms in the calling activity of eneopterine crickets (Orthoptera: Grylloidea: Gryllidae) from Southeast Asia. Bioacoustics, 31 (4), 470 - 489. https: // doi. org / 10.1080 / 09524622.2021.1973562"]}

Genus Nisitrus Saussure, 1878 Type species: Nisitrus marginatus Walker, 1869 – According to the ICZN, and contrary to what was mentioned in the recent revision of the genus (Tan et al., 2021a), the type species should remain N. marginatus even though this name is a junior synonym of N. vittatus Haan., Published as part of Robillard, Tony, Tan, Ming Kai, Japir, Razy & Chung, Arthur Y. C., 2023, Notes on the Eneopterinae (Orthoptera, Grylloidea, Gryllidae) from eastern Sabah, pp. 231-250 in Zootaxa 5315 (3) on page 233, DOI: 10.11646/zootaxa.5315.3.2, http://zenodo.org/record/8130509, {"references":["Saussure, H. (1878) Melanges orthopterologiques. VI. Fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 25 (2), 369 - 704 (505 - 834)."]}

Robillard, Tony, Tan, Ming Kai, Japir, Razy, Chung, Arthur Y.C. (2023): Notes on the Eneopterinae (Orthoptera, Grylloidea, Gryllidae) from eastern Sabah. Zootaxa 5315 (3): 231-250, DOI: 10.11646/zootaxa.5315.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5315.3.2

Pendleburyella testacea Chopard, 1969 (Figs. 3, 7A, 8A) Pendleburyella testacea Chopard 1969: 218 Diagnosis (new). This species is characterised by a medium-sized habitus, the maxillary palps with the apical segment triangular, the mirror longer than broad, the dividing vein of the mirror roundly curved at the basal end, the apical part of FW longer. Material examined. Holotype: MALAYSIA • 1♁; Selangor, Bukit Kutu, 3300–3500 ft; 24 September 1932; coll. H.M. Pendlebury; 0019 (MP). Allotype: EAST MALAYSIA • 1♀; Sabah, Sandakan [N. Borneo, Bettotan, NR. Sandakan]; 7 August 1927; 0020 (MP). Remarks. The male genitalia of the holotype was not dissected and examined to minimise the risk of further damage to the old specimen. The female allotype from Borneo probably belongs a different species. Distribution. Malay Peninsula (Selangor). Type locality. MALAYSIA, Selangor, Bukit Kutu., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on page 138, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034, {"references":["Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. 1 - 421."]}

Pendleburyella vicina Chopard, 1969 (Figs. 7B, 8B, 10) Pendleburyella vicina Chopard 1969: 219 Diagnosis (new). This species is characterised by a small-sized habitus, the maxillary palps with the apical segment somewhat cylindrical to tear-shaped, the mirror about as long as broad, the dividing vein of the mirror broadly and roundly curved at the basal end, the apical part of FW shorter. Material examined. Holotype: MALAYSIA • 1♁; Kuala Lumpur, at light; 13 September 1932; coll. H.M. Pendlebury; 0018 (MP). Remarks. The type specimen is mouldy and badly damaged in the abdomen area. Distribution. Malay Peninsula (Selangor). Type locality. MALAYSIA, Selangor, Kuala Lumpur., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on page 138, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034, {"references":["Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. 1 - 421."]}

Pendleburyella sp. 2 (Figs. 7F, 8G, 14) Material examined. MALAYSIA • 1♁ (type); Selangor –Pahang, The Gap, 2700 ft, at light; 21 January 1939; coll. H.M. Pendlebury; (MP). Remarks. This old specimen from Selangor – Pahang shares the greatest resemblance with Pendleburyella eirmosa sp. nov., including the shapes of the apical and subapical segments of the maxillary palps, FWL/FWW, the shape of the dividing vein of mirror at the basal end and the extent of apical area of the FW. But there are also some morphological differences: smaller size, mirror length as long as broad (instead of longer than broad). As the two specimens were collected from two different parts of Southeast Asia and at different altitudes (and hence different habitats), we speculate that they belong to two different species. However, we also avoid describing and naming this species as the male genitalia is not available for the verification of the species status., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on pages 145-146, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034

Pendleburyella pimela sp. nov. (Figs. 4, 5A, 6A, 6C, 6E, 7C, 8C, 9A–C, 11) Diagnosis. This species is characterised by a large-sized habitus, the maxillary palps with the apical segment also triangular (but more stout and broadly widened than Pendleburyella testacea); the FWs relatively broad (FWL/ FWW 2.5 in other species specimens); the mirror about as long as broad, the dividing vein of the mirror broadly and roundly curved at the basal end, the apical part of the FW shorter relative to the FWL (all of these similar to Pendleburyella vicina but this species is distinctly larger in size). The genitalia differs from that of Pendleburyella eirmosa sp. nov. by the pseudepiphallic lophi, in profile, cylindrical with roundly truncated apices (instead of dorso-ventrally flattened), the pseudepiphallic paramere ventral processes more elongated and in ventral view having acute apices (instead of obtuse apices); and the emargination between the dorsal and ventral processes pseudepiphallic paramere, in profile, narrowly angular (instead of broadly rounded). Etymology. The species name refers to the large habitus and relatively broad FWs (low FWL/FWW); pimele = fat in Greek. Material examined. Holotype: SINGAPORE • 1♁; Nee Soon freshwater swamp forest, along Woodcutter Trail; on leaf litter; 21 December 2022; coll. M.K. Tan & I. Neo; call recorded; (ZRC). Description. Very large species for this genus. Maxillary palpi with apical segment longest and expanded (triangular); with subapical segment cylindrical and expanded slightly distally, somewhat similar length as apical and third segments (Fig. 7C). Pronotal disc brown about 2.0 times as wide as long, widening posteriorly (posterior margin about 1.6 times as wide as anterior margin); anterior margin of disc broadly concave; posterior margin of disc slightly convex (Fig. 6A). Pronotal lateral lobe 2.3 times as long as high (Fig, 7C). Male. FW 2.4 times as long as broad (Fig. 8C), covering abdomen and slightly surpassing apex of FIII. Venation (Fig. 8C): 1A vein transverse, faintly curved; diagonal substraight, with 3 distinct oblique veins in harp area; posterior two oblique veins substraight,join at base near1A,anterior most vein distinctly shorter and running nearly perpendicular to 1A. Mirror about 1.1 times as long as wide, dividing vein broadly curved at basal end. Lateral field around 10 branches on Sc (Fig. 6E). Apical field 0.09 times as long as FWL (Fig. 8C). Hind wings clearly surpassing FWs. ♁ genitalia (Figs. 9A–C). Pseudepiphallus [epiphallus] typical of genus. Posterior end of pseudepiphallus [epiphallus] produced into two tongue-shaped lobular pseudepiphallic lophi [posterolateral epiphallic lobe] with obtuse apices. These lophi, in profile, cylindrical with roundly truncated apices. Pseudepiphallic paramere [ectoparamere] very elongated with ventral process well surpassing pseudepiphallic lophi; all processes with apex acute. Dorsal process of pseudepiphallic paramere distinctly shorter than ventral process barely surpassing pseudepiphallic lophi. Ventral process of pseudepiphallic paramere distinctly more sclerotized along inner margin. In profile, emargination between dorsal and ventral processes narrowly angular. Ectophallic fold [rachis] typical of genus; in profile tapering into acute apex, with anterior margin concave and posterior margin straight. Measurements (♂, in mm). BL = 9.5; BWL = 13.3; HL = 1.1; PronL = 1.4; PronW = 2.8; FWL = 7.9; FWW = 3.3; HWT = 3.3; FIIIL = 5.4; TIIIL = 3.9; TaIIIL = 1.5. Ecology. The species was found dwelling among thick leaf litter in freshwater swamp forest. Distribution. Malay Peninsula (SINGAPORE). Type locality. SINGAPORE Calling song (1♂, in captivity, 29.0°C) (Fig. 11). The male was observed to call between 22h00 and 23h30, each night having 2–4 bouts of continuous trill. The calling song is a loud continuous trill, typically lasting around 2–5 min when left undisturbed. The trill is made up of closely-packed syllables with an average duration of 9.7±1.5 ms (5.9–14.2 ms). The average syllable period is 14.5±0.8 ms (10.8–17.1 ms). There is no distinct downtime between syllables. The frequency spectrum is pure-tonal and forms a harmonic, with the energy peaking at the fundamental frequency. The dominant frequency is 8.14±0.08 kHz (7.78–8.34 kHz)., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on pages 138-142, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034

Pendleburyella eirmosa sp. nov. (Figs. 1, 5B, 6B, 6D, 6F, 7D, 8D, 9D–F) Diagnosis. This species is characterised by a medium-sized habitus (similar to Pendleburyella testacea), the maxillary palps with the apical segment also somewhat rectangular (similar to that of Pendleburyella vicina but stouter); the mirror longer than broad, the dividing vein of the mirror broadly and roundly curved at the basal end; the apical part of FW longer relative to the FWL. The genitalia also differs from that of Pendleburyella pimela sp. nov. (see the diagnosis for the latter species). Etymology. The species name refers to the broadly rounded emargination between the dorsal and ventral processes of the pseudepiphallic paramere (distinguished from the narrowly angular emargination in Pendleburyella pimela sp. nov.); eirmós = continuity in Ancient Greek. Material examined. Holotype: BRUNEI DARUSSALAM • 1♁; Belait District, Wasai Wong Kadir Recreational Park; N4.34186, E114.44611, 35.8± 5.2 m.a.s.l.; 1 March 2019, 18h54; calling on a leaf; coll. M.K. Tan & H. Yeo; BRU.19.47 (UBDM) Photograph examined. BRUNEI DARUSSALAM • 1♁; Temburong District, Kuala Belalong Field Studies Centre, along Ashton Trail; N4.54619, E115.15690, 110.8± 6.9 m.a.s.l.; 26 July2017, dusk; on leaf litter; photographed M.K. Tan (Fig. 1). Description. Medium-sized species for this genus. Maxillary palpi with apical segment stout and cylindrical to somewhat rectangular; with subapical segment very stout, distinctly shorter than apical and third segments (Fig. 7D). Pronotal disc brown, about 1.7 times as wide as long, widening posteriorly (posterior margin about 1.6 times as wide as anterior margin); anterior margin of disc somewhat straight; posterior margin of disc slightly convex (Fig. 6B). Pronotal lateral lobe 2.0 times as long as high (Fig. 7D). Male. FW 2.6 times as long as broad (Fig. 8D), covering abdomen and slightly surpassing apex of FIII. Venation (Fig. 8D): 1A vein transverse, faintly sinuous; diagonal faintly bent, with 3 distinct oblique veins in harp area; posterior two oblique veins parallel to each other, anterior most vein distinctly shorter and running nearly along length of FW. Mirror about 1.2 times as long as wide, dividing vein broadly curved at basal end. Lateral field around 7 branches on Sc (Fig. 6F). Apical field 0.11 times as long as FWL (Fig. 8D). Hind wings clearly surpassing FWs. ♁ genitalia (Figs. 9D–F). Pseudepiphallus [epiphallus] typical of genus. Posterior end of pseudepiphallus [epiphallus] produced into two lobular pseudepiphallic lophi [posterolateral epiphallic lobe], in dorsal view tongue-shaped with apex obtuse; more broadly emarginated between latero-apical lobules; in profile distinctly flattened. Pseudepiphallic paramere [ectoparamere] not surpassing pseudepiphallic lophi, sclerotized, also slender and tapering into subacute apex. Dorsal process of pseudepiphallic paramere not surpassing ventral process. Ventral process of pseudepiphallic paramere obtuse at apex in ventral view. In profile, emargination between dorsal and ventral processes of pseudepiphallic paramere broadly rounded. Ectophallic fold [rachis] typical of genus; in profile slenderer. Measurements (♂, in mm). BL = 6.5; BWL = 10.1; HL = 0.9; PronL = 1.2; PronW = 2.0; FWL = 6.2; FWW = 2.4; HWT = 2.1; FIIIL = 4.0; TIIIL = 3.1; TaIIIL = 1.6. Ecology. The species was found dwelling among the leaf litter as well as on the leaves of shrub/small trees. Distribution. Borneo (BRUNEI DARUSSALAM, Belait). Type locality. BRUNEI DARUSSALAM, Belait.

Pendleburyella sp. 1 (Figs. 7E, 8F, 13) Material examined. MALAYSIA • 1♁ (type); Kuala Lumpur; 5 March 1939; 0021 (MP). Remarks. This specimen is small among the other specimens, and is most easily characterised by the very slender FWs and the distinctly longer than broad mirror. The dividing vein of the mirror at the basal end is strongly and distinctly angularly curved, unlike the other specimens examined. It is also clearly different from the holotype of Pendleburyella vicina from Kuala Lumpur by the apical segment of the maxillary palps and the slenderer FWs(Table1)., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on page 145, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034

Genus Pendleburyella Chopard, 1969 Chopard, 1969: 217 Type species. Pendleburyella testacea Chopard, by original designation (Fig. 3) Diagnosis (new). Small brown crickets. Among Pentacentrinae, this genus differs most prominently by the male FWs with a fully developed stridulatory apparatus (the diagonal vein substraight, three oblique veins in the harp area, a large mirror with a distinct dividing vein). The male genitalia: the pseudepiphallus [epiphallus] is elongated and gently narrows posteriorly into two tongue-shaped, obtuse lobular pseudepiphallic lophi [posterolateral epiphallic lobe]. The pseudepiphallic parameres [ectoparameres] are strongly sclerotized, elongated and forming dorsal and ventral subacute processes. The ectophallic fold [rachis] is weakly sclerotized, forming a process with an acute apex pointing perpendicular to the pseudepiphallus (in profile view). The endophallic sclerite [formula (= mold of spermatophore attachment plate)] is small and elongated, with two lateral arms. The ectophallic apodeme [endoparameral apodeme] is very long. Redescription. Small cricket, generally brown or yellow brown (Figs. 1, 4, 5). Head rounded, a little wider than anterior margin of pronotum, with dorsum slightly flattened, vertex sloping, very finely pubescent (Figs. 6A, 6B). Frontal rostrum dark-coloured, about two times as wide as scapes, with apex truncated (in dorsal view) (Figs. 6A, 6B). Eyes globular, feebly projected anteriorly in dorsal view (Figs. 6A, 6B). Antennae inserted at level midpoint of eye in anterior view (Figs. 6C, 6D). Median ocellus round and small; lateral ocelli oval, located near eyes (Figs. 6C, 6D). Maxillary palpi with apical segment longest and expanded (triangular or slightly broadening apically); with subapical segment cylindrical and shorter than apical and third segments (Fig. 7). Face brown, in anterior view about 0.7–0.8 times as tall as wide (Figs. 6C, 6D). Pronotal disc brown about 1.6–2.0 times as wide as long, widening posteriorly (posterior margin about 1.3–1.6 times as wide as anterior margin); anterior margin of disc nearly straight to broadly concave; posterior margin of disc nearly straight to slightly convex (Figs. 6A, 6B). Pronotal lateral lobe about 1.8–2.3 times as long as high (Fig. 7). TI slightly swollen; with inner and outer tympana open and having oval tympanal membrane. TIII with 3 inner and 4 outer long subapical spurs; with 2–4 denticles between spines; 4 denticles before most proximal spines; and with 2 inner spurs (apical spines) longer than corresponding 3 outer ones. Legs generally pale brown. Thoracic and abdominal segments yellow brown to brown (Figs. 6E, 6F). Male. FWs covering abdomen and slightly surpassing apex of FIII. Colouration: yellow brown, with veins darker (Fig. 8). Venation: 1A vein transverse, faintly curved; diagonal substraight, with three distinct oblique veins in harp area; posterior two oblique vein more or less straight and long, anteriormost vein distinctly shorter and running nearly perpendicular to 1A. Mirror about 1.1–1.5 times as long as wide, dividing vein curved at basal end (Fig. 8). Lateral field around 7–10 branches on Sc (Figs. 6E, 6F). Hind wings clearly surpassing FWs. ♁ genitalia as shown in Fig. 9: Pseudepiphallus [epiphallus] elongated, gently narrowing posteriorly. Posterior end of pseudepiphallus produced into two lobular pseudepiphallic lophi [posterolateral epiphallic lobe], in dorsal view tongue-shaped; emarginated between latero-apical lobules in dorsal view; in profile variable (e.g., cylindrical or flattened), with a few setae. Pseudepiphallic paramere [ectoparamere] strongly sclerotized, elongated with dorsal and ventral processes (emarginated in profile); each process tapered into a slender and slightly hooked-like apical third with apex acute. Ectophallic fold [rachis] weakly sclerotized, rather long but weakly sclerotized, pointing perpendicular to pseudepiphallus; in profile tapering into acute apex. Endophallic sclerite [formula (= mold of spermatophore attachment plate)] small and elongated, with two lateral arms. Ectophallic apodeme [endoparameral apodeme] widened posteriorly, otherwise long, slender and substraight. Rachis weakly sclerotized. Distribution. Malay Peninsula, Borneo, Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on pages 134-137, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034, {"references":["Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. 1 - 421."]}

Pendleburyella sp. “ intermedia Chopard” (Figs. 8E, 12) Material examined. MALAYSIA • 1♁ (type); Pahang, Fraser’s Hill, at light; 12 July 1936; 0017 (MP). Remarks. The ‘type’ specimen was identified as “ Pendleburya intermedia ” by Chopard, but we did not find the original description of this species in Chopard (1969) where the genus and species were first described. We also did not find other publications where this species was mentioned, and consequently it was also not found in the Orthoptera Species File (Cigliano et al., 2023). This specimen is large and shows clear differences with Pendleburyella sp. 2 from Selangor – Pahang, The Gap (not too far from Fraser’s Hill) (Table 1). Unfortunately, the specimen also appeared damaged at the abdominal apex and the male genitalia may be lost. As the male genitalia was not available, we did not describe and name this species unless newer material from Fraser’s Hill can be collected., Published as part of Tan, Ming Kai, Muhammad, Amira Aqilah & Wahab, Rodzay Bin Haji Abdul, 2023, The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae), pp. 131-148 in Zootaxa 5277 (1) on page 144, DOI: 10.11646/zootaxa.5277.1.6, http://zenodo.org/record/7893034, {"references":["Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. 1 - 421.","Cigliano, M. M., Braun, H., Eades, D. C. & Otte, D. (2023) Orthoptera Species File Online. Version 5 (5.0). Available from: http: // orthoptera. speciesfile. org / HomePage / Orthoptera / HomePage. aspx (accessed 14 March 2023)"]}