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Some well-managed organic soils are known to have higher crop yield potential than conventionally managed soils due to the greater soil quality and the ability to tolerate weed competition. However, low available soil mineral N and P in some organic systems may mask such soil quality–related benefits. We hypothesize that when plant-available N and P are not limiting, tillage-based highly diverse organic crop rotations have less yield loss (better crop tolerance) due to weed competition and higher crop yields than no-till conventional systems with low-diversity rotations. A greenhouse study was carried out in Saskatoon, Canada, using long-term (18-yr) organically managed soils (ORG) and no-till conventional soils (CONV) with three crop rotation diversities (LOW, MEDIUM, and HIGH) to compare the crop tolerance to weed competition under standard soil nutrient management conditions and under excess supply of mineral N and P. Under fertilized conditions, crop biomass increased by 50% and 69% in ORG and CONV systems, respectively. Weed biomass was similar between ORG and CONV systems under nonfertilized conditions but was 14% greater in CONV when excessive N and P were supplied. Crop biomass loss (crop tolerance) was not different among cropping systems under excess fertilizer or under standard fertilizer levels. Even with greater weed biomass under fertilized conditions, the CONV system showed crop tolerance similar to that of the ORG system. Under nonfertilized conditions, the crop biomass yield was 43% lower in ORG compared with CONV, and even after mineral N and P were applied, ORG systems showed less (17%) crop biomass than CONV. Further, differences in crop tolerance were not identified among crop rotations under both fertilizer levels. Overall, this study revealed that there were no yield benefits or better crop tolerance to weed competition in organically managed soils compared with no-till conventional soils, even under nonlimiting soil macronutrient conditions.

Canola has a prominent floral signature and requires careful consideration when selecting spectral indices for yield estimation. This study evaluated several spectral indices derived from high-reso...

Silene latifolia Poiret of Eurasia has established in North America, prompting this structural study of its mature unisexual buds and flowers. Floral nectaries, anther and stigma changes, and vestigial reproductive structures were studied using light and scanning electron microscopy. In staminate flowers, anthers dehisced before anthesis and >90% of their pollen was liberated within 36 h. Accumulated in the tubular calyx, nectar descended an anthophore from the stomatal-bearing nectary at the stamen bases. Nectary tissue surrounded the pistillode, a central filamentous organ lacking ovules but tipped by hairs resembling stigmatic papillae. In pistillate flowers, nectar flowed into an inflated calyx. The annular nectary had 10 regularly spaced, stomatal-lined craters and was continuous with the adaxial surfaces of the infertile antisepalous and epipetalous staminodes. Key elements of entomophilous pollination leading to successful sexual reproduction of this invasive species include secondary nectar presentation from disparate floral nectaries that, for pistillate flowers, also incorporate the staminodes; rapid pollen release from anthers; and elongation of papillae by tip growth that enhances each stigma’s receptive surface. Context is also provided for future studies of floral nectary development in this model dioecious species.

With the increasing resistance of wild oat (Avena fatua L.) to herbicides, there is a need to evaluate the potential of alternative cropping systems based on integrated weed management principles. A 5-yr field study at eight sites across Canada was used to evaluate the profitability of alternative cropping systems that have the potential to control wild oat using cultural practices in conjunction with herbicides. Cultural practices included twice the recommended seeding rates, fall-seeded winter crops, barley (Hordeum vulgare L.) silage, fallow, and alfalfa (Medicago sativa L.). Seven of the 14 cropping systems in this study did not include wild oat herbicide for three consecutive years, controlling wild oat entirely by cultural practices. Cropping system profitability varied by location. For many locations, combinations of barley silage and fall-seeded winter crops without wild oat herbicide application were as profitable as a system of canola (Brassica napus L.) and wheat (Triticum aestivum L.) with wild oat herbicide applied every year. Unprofitable systems generally included those with fallow, alfalfa, and fall-seeded winter crops in regions with rates of high winter kill. Wild oat control can be achieved with diverse cropping systems that are as profitable as conventional annual cropping that relies on herbicide control of wild oat.

Interrow cultivation is a selective, in-crop mechanical weed control tool that has the potential to control weeds later in the growing season with less crop damage compared with other incrop mechanical weed control tools. To our knowledge, no previous research has been conducted on the tolerance of narrow-row crops to interrow cultivation. The objective of this experiment was to determine the tolerance of field pea and lentil to interrow cultivation. Replicated field experiments were conducted in Saskatchewan, Canada, in 2014 and 2015. Weekly cultivation treatments began at the 4-node stage of each crop, continuing for 6 wk. Field pea and lentil yield linearly declined with later crop stages of cultivation. Cultivating multiple times throughout the growing season reduced yield by 15% to 30% in both crops. Minimal yield loss occurred when interrow cultivation was conducted once at early growth stages of field pea and lentil; however, yield loss increased with delayed and more frequent cultivation events.Nomenclature: Field pea; Pisum sativum L.; lentil; Lens culinaris L.

The escalating evolution of weed species resistant to acetolactase synthase (ALS)-inhibitor herbicides makes alternative weed control strategies necessary for field crops that are dependent on this herbicide group. A fully integrated strategy that combined increased crop seeding rates (2X or 4X recommended), mechanical weed control with a minimum-tillage rotary hoe, and reduced-rate non—ALS inhibitor herbicides was compared with herbicides, rotary hoe, and seeding rates alone as a method of controlling ALS inhibitor—tolerant Indian mustard as a model weed. The full-rate herbicide treatment had the lowest weed biomass (98% reduction) and the highest yield of all treatments in 3 of 4 site-years, regardless of seeding rate. The fully integrated treatment at the 4X seeding rate had weed suppression rates equal to the full herbicide treatment at the recommended seeding rate. The fully integrated and reduced-rate herbicide treatments at the 4X seeding rate reduced weed biomass by 89% and 83%, respectively, compared with the control at the recommended seeding rate. The rotary hoe treatment alone resulted in poor weed control (≤38%), even at the highest seeding rate. Fully integrated and reduced-rate herbicide treatments at 2X and 4X seeding rates had yields equal to those of the full herbicide treatment at the recommended seeding rate. Partially or fully integrated weed control strategies that combine increased crop seeding rates and reduced-rate non—ALS inhibitor herbicides, with or without the use of a rotary hoe, can control weeds resistant to ALS-inhibitor herbicides, while maintaining crop yields similar to those achieved with full-rate herbicides. However, combining increased seeding rate, reduced-rate herbicides, and mechanical rotary hoe treatment into a fully integrated strategy maximized weed control, while reducing reliance on and selection pressure against any single weed control tactic.Nomenclature: Indian mustard, Brassica juncea (L.) Czern.

Concern over the development of herbicide-resistant weeds has led to interest in integrated weed management systems that reduce selection pressure by utilizing mechanical and cultural weed control practices in addition to herbicides. Increasing crop seeding rate increases crop competitive ability and thus can enhance herbicide efficacy. However, it is unknown how increasing the seeding rate affects an herbicide’s efficacy. The objective of this study was to examine the interaction between increasing seeding rate and herbicide dose to control weeds. To meet this objective, the herbicide fluthiacet-methyl was applied to field-grown lentil, with Indian mustard, a proxy for wild mustard, used as a model weed. The experiment was a factorial design with four lentil seeding rates and seven herbicide rates. Overall the herbicide dose response was altered by changing lentil seeding rate. Increasing lentil seeding rate decreased the weed biomass production when herbicides were not applied. In two of the four site-years, increasing lentil seeding rate lowered the herbicide ED50, the dose required to result in a 50% reduction in weed biomass. Increasing the crop seeding rate altered the dose response to provide greater weed control at lower herbicide rates compared with normal crop seeding rates. Increased seeding rates also resulted in higher and more stable crop seed yields across a wider range of herbicide dosages. These results suggest that dose–response models can be used to evaluate the efficacy of other weed management practices that can interact with herbicide performance.

Canola (Brassica napus L.) often experiences high seed losses prior to and during harvest operations. Pod drop and pod shatter are responsible for pre-harvest seed losses in canola although relatively little is known about the significance of pod drop to total pre-harvest seed losses in canola. This research describes a method to measure the predisposition for pod drop among canola genotypes that was developed and tested across different sites from 2013 to 2015. This method, referred to as pod-retention resistance, consists of the use of a force gauge to quantify the force required to break the pod pedicle at the pedicle-rachis junction. Two sets of field experiments were used, the first developed and modeled the factors that influence pod-retention resistance, pod drop and seed shatter. A second set of experiments was used to validate these findings on a broader range of genotypes and determine whether a relationship between pod retention resistance and pod drop could be determined. Results showed that the pod-retention resistance was significantly greater for pods on the proximal half of the rachis compared to the distal half of the rachis. Several factors contributed strongly to variation in pod retention resistance (rachis position, genotype and site) and pod drop (site, rachis position and genotype). The relative importance of individual factors varied between pod retention resistance and pod drop suggesting no relationship between the two. However, weight-adjusted pod retention resistance, i.e., average pod retention resistance corrected for the average weight of dropped pods, was strongly correlated with pod drop, particularly in the validation experiments. This method could be used by canola breeders to select for and develop genotypes with reduced predisposition for seed losses via pod drop.

Alternative cropping systems are gaining attention throughout the world in order to increase the sustainability of agro-ecosystems. On the Canadian prairies, the tillage-based crop-fallow system has been replaced by no-till reduced input systems or tillage-based organic systems with more diversity in crop rotations but with no external inputs. However, the long-term effects of these alternative systems on weed and yield parameters have not been assessed. A study to examine weed and crop yield dynamics under diverse cropping systems was conducted within a 18-year cropping systems study near Scott, Saskatchewan. The trial was laid out in a split-split-plot design with four replicates. The main plots were three levels of inputs; a high input system (HIGH) that used tillage and inputs to maximize yield, a reduced system (RED) that used no-till practices and minimal inputs, and a tillage-based organic system (ORG) with no external inputs. The subplots were cropping diversity (rotations); fallow-annual grains (LOW), diversified annual grains (DAG), and diversified annuals and perennial forage (DAP). The sub-sub plots were the six phases of each rotation. There was an input by rotation interaction for weed biomass but not for weed density and crop yields. ORG systems had 7× and 4× greater weed density (107 plants m −2 ), 4× higher weed biomass (154 kg ha −1 ), and 32% and 35% lower yields (1052 kg ha −1 ) than RED and HIGH systems respectively. RED and HIGH input systems had similar crop yields and lower weed density than ORG. The LOW rotation had the lowest weed density. LOW and DAG rotations had similar yields, which were higher than in DAP. All systems showed an increase in weed density and biomass over time but did not impact on crop yields which was increasing over time likely influenced by a concurrent increase in rainfall. This study concludes that eliminating tillage and reducing agrochemicals is possible but eliminating agrochemicals requires better crop rotations for weed management as well as for nutrient management.

In western Canada, more money is spent on wild oat herbicides than on any other weed species, and wild oat resistance to herbicides is the most widespread resistance issue. A direct-seeded field experiment was conducted from 2010 to 2014 at eight Canadian sites to determine crop life cycle, crop species, crop seeding rate, crop usage, and herbicide rate combination effects on wild oat management and canola yield. Combining 2× seeding rates of early-cut barley silage with 2× seeding rates of winter cereals and excluding wild oat herbicides for 3 of 5 yr (2011 to 2013) often led to similar wild oat density, aboveground wild oat biomass, wild oat seed density in the soil, and canola yield as a repeated canola–wheat rotation under a full wild oat herbicide rate regime. Wild oat was similarly well managed after 3 yr of perennial alfalfa without wild oat herbicides. Forgoing wild oat herbicides in only 2 of 5 yr from exclusively summer annual crop rotations resulted in higher wild oat density, biomass, and seed banks. Management systems that effectively combine diverse and optimal cultural practices against weeds, and limit herbicide use, reduce selection pressure for weed resistance to herbicides and prolong the utility of threatened herbicide tools.

Silene latifolia Poir. (white cockle or white campion) is an important invasive weed in North American agriculture. It exhibits dioecy, therefore, both male and female plants are required in order for seed production to occur. However, dioecious species being invasive is not common because of their limitations in pollination and subsequent seed production. The objective of this study is to determine the effect of pollination timing and distance on seed production of Silene latifolia. A series of experiments including pollination exclusion, timing and pollination distance were conducted in 2009 and 2010 at or around Saskatoon, Saskatchewan. For pollination exclusion, exclosures were built around the natural female plants for exclosure, sham-exclosure, and male and female combined treatments. Pollination timing was studied by applying exclosure, non-exclosure, night-exclosure, and day-exclosure treatments to individual female plants. Female plants were transplanted along a linear interval at six different distances from the pollen source to study the effect of pollination distance. S. latifolia was exclusively insect-pollinated and pollination occurred both day and night; however, in one year, pollination occurred mainly at night. Female plants that were in the range of 0–4 m from a compatible pollen source experienced no limitation to pollination. However, when the distance was increased further up to 128 m, pollination levels and subsequent seed production were declined. Moreover, there were differences in seed production between years suggesting that pollination was affected by the environmental conditions during pollination and the crop that white cockle was grown in. These experiments indicate that seed production in S. latifolia is limited by insect-pollination. Although there was pollination limitation for seed production at greater distances from a pollen source, the high fecundity rate (3000–18000 seeds per plant) resulted in a large seed output. Thus, we believe that a dioecious species may require characteristics that compensate the pollination limitation for a successful invasion.

Weed management in organic lentil (Lens culinaris Medik.) is challenging due to its poorly competitive nature with weeds and the lack of organic herbicides for field crops. In-crop mechanical weed control methods such as rotary hoeing, harrowing and inter-row cultivation, and the cultural practice of increased crop seeding rate, are established weed management tools in organic field crops. They have however not previously been combined into an integrated weed management system for weakly competitive grain crops. The objective of this research was to determine the effect of mechanical weed control (rotary hoeing, harrowing and inter-row cultivation) and crop seeding rate alone and in combination on weed suppression and yield in organic lentil. A two-factor randomized complete block design experiment was conducted at two locations in Saskatchewan, Canada in 2016 and 2017. The factors mechanical weed control methods including the rotary hoe, the harrow and inter-row cultivation and an untreated control, and seeding rates including conventional recommended and double recommended seeding rate. All treatments were tested as stand-alone or combined in a factorial fashion. Increased crop seeding rate and mechanical weed control affected response variables independently of each other. Doubling the seeding rate of lentil reduced weed biomass by 16% and increased lentil yield by 23% averaged across the four site-years. Treatments that included only a single mechanical weed control methods resulted in poor or incomplete weed suppression at all sites. Two mechanical weed control methods were required to maximize yield, and three were required to minimize weed biomass. Early weed control with a rotary hoe, followed by inter-row cultivation prior to canopy closure was the most consistent mechanical weed control treatment across the four site-years. Averaged across the four site-years, rotary hoeing + inter-row cultivation reduced weed biomass by 70%, and increased crop yield by 65%. The treatment maintained low weed populations across the longest time duration and controlled weeds in both the inter- and intra-row spaces, while minimizing crop injury. Doubling crop seeding rate and using two appropriately timed mechanical weed control methods provided weed suppression while increasing yield by 46% in a weakly competitive organic lentil crop.

Domestication is the process by which a wild plant evolves into a cultivated plant by undergoing morphological, physiological or molecular changes. These changes may be due to conscious selection (primary cultivated plant) or unconscious selection (secondary cultivated plant) by man. These evolutionary changes need to be studied to understand the transition from wild to cultivated plant. Cow cockle (Vaccaria hispanica) is a summer annual species introduced to North America from Europe. It is being considered for commercial production because of its seed constituents. The purpose of this study is to determine which cow cockle populations are best adapted to cultivation and the plant characteristics that are responsible for such adaptation. We compared 15 cultivated and wild cow cockle populations from different parts of the world for 12 agronomic and morphological traits. Multivariate analyses were performed to group populations according to their agronomic and morphological similarity. Cluster analysis revealed three main population groups among the 15 accessions based on the traits studied. Two linear discriminants were obtained by discriminant function analysis accounting for 96 % of the variability among the populations. Analysis of variance showed significant (P

Seedbank persistence in canola seeds is related to their potential to develop secondary dormancy. This can result in volunteer weed problems many years after canola production. The potential to be induced into secondary dormancy is controlled by both the canola genetics and the environment of the mother plant. However, the effect of time of harvesting on secondary dormancy potential is not known. The objective of this study was to determine the effect of harvest timing on potential to develop seed dormancy in canola. Six harvest samples were collected weekly from two canola genotypes (5440 and 5020) starting from 10 to 20% seed color change on the main stem until they were fully ripened. Freshly harvested seeds of 5440 and 5020 showed 13 and 16% primary dormancy at 32 and 33 d after flowering (DAF), respectively, but dormancy decreased with harvest timings and no dormancy was observed when seeds were fully mature (78 DAF). After dormancy induction, 10% of 5440 seeds were dormant at 32 DAF, but 94% of seeds were dormant at 78 DAF. Similarly, 70% of 5020 seeds were dormant at 33 DAF, but 90% of seeds were dormant at 68 DAF. Thus, seeds had lower potential to secondary dormancy at early development but have a high potential to secondary dormancy induction at full maturity. This study suggests that windrowing these canola genotypes at the recommended time (60% seed color change on the main stem) may reduce ability of the seed to develop secondary dormancy and thus reduce the persistence of seeds in the soil seedbank.

Cowcockle, an introduced summer annual weed of the Northern Great Plains, is being considered for domestication because of its high quality starch, cyclopeptides, and saponins. Loss of seed dormancy is one of the key desirable traits for domestication. To determine the potential for domestication of this species, an understanding of the seed dormancy and germination patterns is required. The objectives of this study were to evaluate seed dormancy in cowcockle ecotypes and determine how temperature and light affect seed dormancy. We evaluated 15 populations of cowcockle for primary dormancy by exposing them to five temperatures (5, 7.5, 10, 15, and 20 C) under two temperature regimes (constant and alternating) in both dark and light conditions. Freshly matured seeds of all the populations showed high levels of primary dormancy except ‘Mongolia’. Lower levels of dormancy at medium temperatures (10 and 15 C) and greater dormancy at low and high temperatures suggest conditional dormancy, a state at which seeds germinate over a narrower range of conditions compared to nondormant seeds. The effects of temperature regime, light, and their interaction was significant only at suboptimal (5 and 7.5 C) and supraoptimal (20 C) temperatures. Under these conditions, alternating temperatures were more effective in breaking the conditional dormancy, followed by light. The variation in optimum temperature, light, and their interactions among the cowcockle populations may be due to the plants evolving to adapt to their local environments. From a domestication perspective, the conditional dormancy in cowcockle can be observed as an evolutionary mechanism that prevents untimely germination following maturity and may not be a major obstacle for its domestication. Nomenclature: Cowcockle, Vaccaria hispanica (P. Mill.) Rauschert, VAPPY.

Weed control with herbicides is not possible in several systems including control of wild oat (Avena fatua L.) in tame oat (Avena sativa L.), red rice (Oryza sativa L. var. sylvatica) in rice (Oryza sativa L.), and in organic systems. Competitive crop cultivars can be used to manage weed competition if selective weed control by herbicides is not possible. However, most existing competitive crop cultivars often have low weed-free yield and poor grain quality. We hypothesize that the progeny of a cross between a competitive forage-type oat cultivar and a high-yielding and high grain quality grain oat cultivar will have high yield and good grain quality as well as high competitive ability (CA). The objective of this study was to evaluate progeny lines from a cross between a tall, competitive, forage-type oat cultivar with a semidwarf, high-yielding milling oat for their CA against wild oat. A field study was performed in two locations in Saskatoon, SK, Canada, in 2008 and 2009 using seven progeny lines and the two parents. Oat was seeded with and without wild oat at a target crop and weed density of 250 plant m⁻² in a randomized block design with four replicates. All the genotypes were high yielding and did not differ in grain yield or quality either in weed-free or weedy conditions. Some of the tall (CDC Baler, SA050498, and SA050479) and short (SA050040 and Ronald) genotypes had high average grain yield under both weed-free and weedy conditions compared with the other genotypes. The tall oat line SA050479 was among the most weed suppressive, which resulted in less wild oat biomass. The seedling total leaf area, crop height, and the seedling third leaf area were negatively correlated with wild oat biomass. This study demonstrates that a cross between a high-yielding and a highly competitive genotype can result in progeny with high CA, yield, and crop quality.

Effective weed management strategies are limited in organic cropping systems because herbicide use is prohibited. Enhancing crop competitive ability by integrating both cultural and mechanical weed control methods is a key strategy in such instances, but the relative efficacy of different cultural and mechanical strategies and their interactions and additive effects when combined is not well known. The objective of this study was to determine the individual and additive effects of cultural and mechanical methods on weed suppression and crop yield under organic conditions. A study was performed in two organically managed oat (Avena sativa L.) cropping systems in Saskatoon, SK, Canada, in 2008 and 2009. Three cultural practices, two crop genotypes (competitive and less competitive), high and standard crop densities (500 and 250 plants m–²), narrow and standard row spacings (11.5 and 23 cm), and two post-emergence tillage levels (harrowing and nonharrowed control) were factorially applied in a randomized block design. Increasing crop density and harrowing increased grain yield by 11 and 13%, respectively. Competitive genotype and high crop density reduced weed biomass by 22 and 52%, respectively. Combining high crop density with post-emergence harrowing increased the grain yield by 25%. The combined treatment effects on weed biomass were more profound, as the competitive genotype, increased seeding rate, and post-emergence harrowing decreased weed biomass by 71% compared with standard practices. Integrating cultural and mechanical weed management practices was superior to the use of individual practices because they additively control weeds in an organic cropping system.

The challenges associated with the adoption of conservation tillage and/or low-input cropping systems, whether organic or herbicide-free, across Canada are shaped by scale, environment, and local practices. A study in eastern Canada captured the challenges of introducing low-input cropping systems in mature (20+ yr) tillage treatments in a barley/red clover/corn/soybean rotation. Each mature tillage system came with its own weed problems, but this did not affect crops such as barley and red clover, which produced similar yields across high and low input systems. However, some form of primary tillage was needed to achieve adequate weed control and yield in organic (ORG) and herbicide-free (HF) systems. The HF and ORG systems with no-till actually failed to produce a corn crop but produced soybean yields that were half or less than that for other treatments. The successful combination of conservation tillage practices and low-input systems in eastern Canada would thus appear to be crop-specific, being easier to achieve in competitive cereal crops. In western Canadian organic agriculture, tillage is practiced with low-disturbance chisel plows instead of inversion plows. However, green manure crops (summer cover crops) are often terminated with tandem discs. Both roller crimpers and mowing can successfully kill annual green manure crops such as field pea and rye, and usually result in reduced weed growth following termination. However, the lack of tillage can result in lower crop yields in wheat following green manure terminated by roller crimping compared to tillage. Challenges for no-till organic practices include perennial weed control and soil fertility. Overall, flexible crop production programs such as the former Manitoba Pesticide Free Production program and the “Agriculture raisonnéeTM” program in Québec are more likely to promote sustained environmental, economic, and social prosperity than rigid adherence to organic or no-till practices.

Legume crops are known to have low soil N uptake early in their life cycle, which can weaken their ability to compete with other species, such as weeds or other crops in intercropping systems. However, there is limited knowledge on the main traits involved in soil N uptake during early growth and for a range of species. The objective of this research was to identify the main traits explaining the variability among legume species in soil N uptake and to study the effect of the soil mineral N supply on the legume strategy for the use of available N sources during early growth. Nine legume species were grown in rhizotrons with or without N supply. Root expansion, shoot and root biomass, nodule establishment, N2 fixation and mineral soil N uptake were measured. A large interspecific variability was observed for all traits affecting soil N uptake. Root lateral expansion and early biomass in relation to seed mass were the major traits influencing soil N uptake regardless of the level of soil N availability. Fenugreek, lentil, alfalfa, and common vetch could be considered weak competitors for soil N due to their low plant biomass and low lateral root expansion. Conversely, peanut, pea, chickpea and soybean had a greater soil N uptake. Faba bean was separated from other species having a higher nodule biomass, a higher N2 fixation and a lower seed reserve depletion. Faba bean was able to simultaneously fix N2 and take up soil N. This work has identified traits of seed mass, shoot and root biomass, root lateral expansion, N2 fixation and seed reserve depletion that allowing classification of legume species regarding their soil N uptake ability during early growth.

Seed shatter of wild oat (Avena fatua L.), green foxtail [Setaria viridis (L.) P. Beauv.], wild mustard (Sinapis arvensis L.), cleavers (Galium spurium L. and G. aparine L.), wild buckwheat (Polygo...

Organic farmers in western Canada rely on tillage to control weeds and incorporate crop residues that could plug mechanical weed-control implements. However, tillage significantly increases the risk of soil erosion. For farmers seeking to reduce or eliminate tillage, potential alternatives include mowing or using a roller crimper for terminating green manure crops (cover crops) or using a minimum tillage (min-till) rotary hoe for mechanically controlling weeds. Although many researchers have studied organic crop production in western Canada, few have studied no-till organic production practices. Two studies were recently conducted in Saskatchewan to determine the efficacy of the following alternatives to tillage: mowing and roller crimping for weed control, and min-till rotary hoeing weed control in field pea (Pisum sativum L.). The first study compared mowing and roller crimping with tillage when terminating faba bean (Vicia faba L.) and field pea green manure crops. Early termination of annual green manure crops with roller crimping or mowing resulted in less weed regrowth compared with tillage. When compared with faba bean, field pea produced greater crop biomass, suppressed weeds better and had less regrowth. Wheat yields following pea were not affected by the method of termination. Thus, this first study indicated that roller crimping and mowing are viable alternatives to tillage to terminate field pea green manure crops. The second study evaluated the tolerance and efficacy of a min-till rotary harrow in no-till field pea production. The min-till rotary hoe was able to operate in no-till cereal residues and multiple passes did not affect the level of residue cover. Field pea exhibited excellent tolerance to the min-till rotary hoe. Good weed control occurred with multiple rotary hoe passes, and pea seed yield was 87% of the yield obtained in the herbicide-treated check. Therefore, this second study demonstrated that min-till rotary hoeing effectively controls many small seeded annual weeds in the presence of crop residue and thus can reduce the need for tillage in organic-cropping systems.

Records from 60 organically managed fields across Saskatchewan were obtained from 39 farmers. The records contained details of 6 yr of management practices applied to each field and details about the general farm operation under which the individual fields were managed. In addition, soil samples were collected from each field and analysed for available N, P, K and S, pH, electrical conductivity (EC) and soil organic carbon. Soil nutrient availability is reported by soil zone. Using information from the management questionnaire, the 60 fields were classified into four distinct management systems, Perennial, Summerfallow, Cereal, and Diverse, and soil nutrient availability examined according to management system. Despite classification of four systems, only the Perennial and Diverse systems had a functionally (and statistically) significant effect on selected soil properties. Inclusion of alfalfa in rotation (Perennial system) acidified the soil and reduced availability of P and K relative to the Diverse system. According to soil test results, fields in all of the systems and across all soil zones were apparently deficient in available P, deficient to marginally deficient in available N and marginally deficient in available S. Available K levels were generally optimal. Information from this study provides a baseline for fertility of soils under organic production across Saskatchewan. The overall low levels of available P, N and S identified in this study raise questions about the long-term sustainability of these organic systems, as well as the suitability of traditional soil testing for accurately assessing the soil fertility potential of organic managed soils.

The current recommended plant density in field pea (Pisum sativum L.) in western Canada is 88 plants m–2. This rate may exceed the optimum for yield in cultivars with more basal branching than typical. The objective of this research was to determine how the seed yield of pea cultivars differing in basal branching ability responds to changes in plant density. Ten pea cultivars were sown at target plant populations of 10, 30, 90, 120, and 150 plant m–2 for 3 yr at Rosthern and Saskatoon, Saskatchewan. At very low plant densities there was greater than a twofold difference in branching potential between cultivars (range 0.85 to 1.99 branches plant–1). Increasing field pea plant density reduced branching by 0.097 branches for each additional plant. The response of yield to plant density differed, as the forage cultivars 40-10 and CDC Sonata reached their potential yield at lower densities, while Courier required higher densities to reach the same proportion of yield. Field pea cultivars with greater basal branching achieved their maximum yield at lower plant densities compared with cultivars with low basal branching. The optimum economic plant density for the pea cultivars ranged from 59 to 84 plants m–2, which is below the current recommended plant density. However, with the exception of the forage cultivars and the low-branching cultivar Courier, the optimum seeding rate was within 8% of 88 seeds m–2. It is recommended that pea growers avoid low-branching pea cultivars and seed at a rate of 88 viable seeds m–2.Key words: Basal branching, seeding rate, plant density, light interception, leafed pea, semi-leafless pea, forage pea

Phosphorus (P) can be low in soil under low input organic management; however, beneficial crop plant associations with arbuscular mycorrhizal fungi (AMF) are known to promote crop nutrition and increase phosphorus uptake. Thus, management strategies that promote AMF associations are particularly desirable for low-input cropping systems. The objectives of this study were to determine the impact of seeding rate on AMF colonization and the impact of AMF colonization on P concentration and uptake by organically grown field pea and lentil. Field experiments examined the impact of three seeding rates of field pea and lentil on P uptake and crop yield. Phosphorus accumulation was examined further in a controlled growth chamber experiment, in which field pea was sown at rates corresponding to those used in the field and harvested at 10-day intervals until 50 days after emergence. In the field, the level of AMF colonization of roots remained at 80% for field pea, while colonization of lentil increased with increasing seeding rates from 77% to 88%. The level of AMF colonization of field pea achieved in the growth chamber after 50 days was 80% for the two highest seeding rates and 60% for the low seeding rate. The rate at which AMF colonization occurred did not vary between treatments. Ultimately, AMF colonization level did not affect P accumulation. In contrast to several previous studies, both field and growth chamber experiments revealed that AMF colonization was not reduced at higher seeding rates. These results suggest that organic farmers may increase seeding rates without adversely affecting P nutrition.

Traditionally, farmers have delayed seeding to manage wild oat (Avena fatua L.) in tame oat (Avena sativa L.) crops, but this practice can adversely affect grain yield and quality. The objectives of this study were: (1) to evaluate the effectiveness of using high seeding rates with early-seeded oat to maintain grain yield and quality, and (2) to determine an optimum seeding rate to manage wild oat and maximize grain yield and quality. The factors of interest were wild oat density (low and high density), seeding date (early May, mid May, early June and mid June), and tame oat seeding rate (150, 250, 350 and 450 viable seeds m-2). The study was conducted at Indian Head and Saskatoon, SK, in 2002, 2003 and 2004, at Winnipeg, MB, in 2002, and at Morden, MB, in 2003 and 2004. Wild oat biomass, wild oat panicle density and wild oat seed in the harvested sample decreased as seeding rate increased, while tame oat biomass and grain yield increased. Wild oat density ranged between 0 and 100 plants m-2 with averages of 10 plants m-2 in the low density treatment and 27 plants m-2 in the high density treatment. At low seeding rates, grain yield decreased with increasing wild oat density. The difference in grain yield between the two wild oat densities decreased as the seeding rate increased. There was a curvilinear decrease in grain yield as seeding was delayed. A seeding date × seeding rate interaction was noted for test weight, plump seed, thin seed and groat yield. Seed quality improved as seeding rate increased for only the mid-June seeding date. Even though the mid-June test weight increased as the seeding rate increased it was always lower than the early May test weight at any seeding rate. The results from this study established that in the presence of wild oats, early seeding of tame oat is possible providing high seeding rates, 350 plants m-2 are used.Key words: Wild oat competition, wild oat density, wild oat biomass, grain yield, grain quality

Successful chickpea production in western Canada typically requires multiple applications of fungicides to minimize the severity of ascochyta blight (AB) caused by Ascochyta rabiei (Pass.) Lab. Although planting resistant cultivars could be economical and environmentally safer than fungicide use, varieties with a high level of resistance are not available. The objective of this research was to identify potentially useful parents for breeding programs aimed at the northern Great Plains by assessing the AB reaction of 12 desi and 12 kabuli chickpea varieties for their AB reaction on leaves, stems and pods under two fungicide regimes. The experiment was conducted at Swift Current and Shaunavon, Saskatchewan, in 2004 and 2005. Differences in AB severity on leaves, stems and pods, seed yield and 1000-seed weight occurred among varieties at all site-years tested. The variation was greater among kabuli varieties than desi varieties. Ascochyta blight severity was generally lower under the high fungicide regime. A positive correlation in AB severity on leaves, stems and pods was observed, suggesting a lack of organ-specific reaction. Several promising varieties that combined improved levels of AB resistance, high yield, and large seed size were identified. Key words: Didymella rabiei, Ascochyta rabiei, Cicer arietinum, fungicide efficacy

Flaxseed is known to have significant health benefits in human nutrition and when included in animal rations. There is pressing need to improve flaxseed production to meet the growing demand. The objective was to study the effects of three seeding rates (22, 45 and 67 kg ha-1), two seeding dates (early May and late May), three rates of nitrogen (66, 100 and 133% of recommended) and three cultivars (Norlin, AC McDuff and CDC Valour) and at five locations representative of the flax-growing area of the Canadian prairies from 1999 to 2001. The variables of interest were plant density, seed yield and yield formation, seed oil content and oil quality. Plant establishment was always highest with Norlin, but bolls m-2 was similar between the three cultivars. Plant density was always highest with the later plantings. No overall effect of seeding date was observed on grain yield, but the site-year by seeding date interaction showed that the later planting was favoured by the most northerly sites and early planting at the most southerly sites. The sites intermediate in latitude showed little response to seeding date. This finding provides important information with respect to the inclusion of flax in cropping systems depending on the latitude and deciding on the order of planting. Later planting resulted in a small decrease in oil content. Seed yields were increased with seeding rates going from 22 to 45 kg ha-1, which corresponded to plant populations > 300 plants m-2, with no increases thereafter. Increases in seeding rates always reduced the number of bolls plant-1 with little effect on seeds boll-1. Seeding rate had no effect on seed oil quantity and quality and maturity. A seed yield increase was observed with nitrogen when going from 66 to 100% of the recommended rate with no increases past 100%. Adding nitrogen resulted in a small drop in seed oil content. The seeding date × nitrogen rate interaction revealed a lack of response to nitrogen with the later planting date beyond 66% of the recommended N rate, suggesting reduced nitrogen rates with later plantings are possible. The absence of a site-year by seeding date by nitrogen rate interaction implies that findings of this study can be applied to a wide geographical area. In conclusion, attention needs to be given to seeding rates to target plant density > 300 plants m-2. Even with the high seeding rates, the target plant population was achieved only 60 and 73% of the time for the early and late planting, respectively. Using soil test recommendations for nitrogen will maximize seed yields under most situations. Best agronomic practices to maximize seed yield were found to maintain adequate seed oil quality and quantity. Key words: Linum usitatissimum L., nitrogen rate, seeding rate, seeding date, cultivar

Oat (Avena sativa L.) yield and quality on the northern Great Plains are consistently reduced by frequent drought and wild oat (Avena fatua L.) competition. Wild oat cannot be selectively removed from oat with herbicides. Identifying genotypes or seed size(s) with high germination potential under moisture stress may facilitate improved seedling vigor, stand establishment, and crop competitiveness. Therefore, a germination study was conducted to determine the effects of genotype and seed size on the germination of oat seed subjected to moisture stress. Large, medium, and small seeds of six common western Canadian oat genotypes were germinated in polyethylene glycol (PEG 8000) solutions with initial osmotic potentials ranging from 0 to -0.4 MPa at 5°C. Generalized linear mixed models fit to the data provided a statistically valid, appropriate, and convenient method to analyze germination data. In all genotypes examined, decreasing seed size and osmotic potential increased median germination time (MGT) and lowered final germination percentage (FGP). Among genotypes, CDC Bell had the fastest MGT while AC Mustang had the highest FGP. Delays in MGT and reductions in FGP resulting from increased moisture stress were similar to those observed in other cereals, suggesting that oat may be as capable of germinating under low spring soil moisture conditions as wheat and barley. The results of this study also indicate that large seed of genotypes such as AC Mustang and CDC Bell appear better suited to germinate under the range of osmotic potentials in this study. Although differences in MGT and FGP between seed sizes in this study were statistically significant, they were generally small and thus, oat germination characteristics may not be substantially improved by screening out small seed from farm-saved seed.

Combine harvesters have the potential to disperse weed seeds great distances. Reducing this dispersal may be important in an integrated weed management system. The objectives of this study were to determine the distance that wild oat seeds are dispersed by a combine harvester and the effect of chaff collection on combine harvester seed dispersal. This was measured by sampling wild oat seeds at varying distances behind a combine equipped with a removable chaff collection system after it passed through a wild oat patch. Chaff collection consistently reduced the amount and distance that wild oat seeds were dispersed. This occurred because more than 74% of the total wild oat seed that were ejected from the combine were in the chaff. Because most of the chaff falls in a row directly behind the combine, chaff collection only affected dispersal in this area. In 1996, chaff collection reduced wild oat seed dispersal past the wild oat patch to less than 10 seeds m−2at 45 m, whereas without chaff collection, there was greater than 10 seeds m−2up to 145 m. At distances beyond 145 m, chaff collection had no significant effect on seed dispersal. Chaff collection may be an important tool in an integrated weed management program because it may slow weed invasions and reduce the expansion of weed patches.

Selective control of wild oat with herbicides is not possible in oat (Avena sativa L.) crops and, consequently, the high quality that is imperative to market success is continually jeopardized. Field experiments were conducted to determine the effects of relative time of emergence and density of wild oat on oat kernel size and weight. Thousand-kernel weight and percentage plump kernels were generally reduced, and percentage thin kernels were generally increased with increasing wild oat densities and earlier wild oat emergence. However, reductions were generally small and occurred only at the highest wild oat densities, which are not commonly observed in commercial fields. These findings suggest that oat growers should not control wild oat based on perceived reductions in oat physical grain quality. Key words: Competition, emergence, physical grain quality, test weight, 1000-kernel weight

The inherent similarity between wild oat (Avena fatua L.) and oat (A. sativa L.) precludes selective herbicide control of wild oat. Previous studies have reported improved crop yield and competitiveness with the use of large seed. Therefore, a greenhouse experiment was conducted in 2002 and 2003 to examine the effect of caryopsis (dehulled kernel) size and genotype on wild oat-oat competition. Seeds of three oat cultivars (AC Assiniboia, CDC Boyer, CDC Orrin) were classified into three size classes based on caryopsis size (large, approximately equal to 35 mg; medium, approximately equal to 25 mg; and small, approximately equal to 15 mg). Whole oat seeds of each genotype and caryopsis size treatment were grown either in monoculture or in mixture with wild oat (250 plants m(-2)). Neither genotype nor caryopsis size significantly influenced (P < 0.05) oat emergence. Nonetheless, oat plants established from large caryopses produced 17% more biomass (P < 0.001) and 15% more panicles m(-2) (P < 0.001) than plants established from small caryopses. Wild oat produced 23% less biomass (P = 0.004) and fewer panicles m(-2) when grown with oat plants established from large caryopses. Genotype only influenced oat panicle production (P = 0.05) and did not affect oat (P = 0.09) or wild oat (P = 0.34) aboveground shoot biomass production. These results suggest that oat plants derived from large-seeded caryopses may be better able to tolerate wild oat interference. However, further investigation is needed to examine the response of oat-wild oat competition to oat caryopsis size and genotype under field conditions.

Wild oat is a serious weed in cultivated oat because there are no herbicides to selectively control it. Considering the effect of time of emergence on weed–crop interference is critical for the development of accurate crop yield loss models and weed density thresholds. Therefore, field experiments were conducted at two locations in Saskatchewan, Canada, in 2002 and 2003 to determine the effect of wild oat density and time of emergence on cultivated oat yield and quality. Wild oat was planted at 50 growing degree day (GDD) intervals ranging from 100 GDD before to 100 GDD after crop planting. Wild oat density ranged from 0 to 320 plants m−2. High densities of early emerging wild oat greatly reduced cultivated oat yield and increased wild oat contamination, with observed oat yield losses as great as 70% and wild oat contamination levels of 15%. Wild oat that emerged before cultivated oat caused considerably more yield and quality loss and had higher reproductive output than wild oat that emerged aft...

Polymer coatings have recently been developed to prevent germination and thereby reduce undesirable emergence of fall-seeded canola (Brassica napus L.) in western Canada. However, recent observations suggest that if seeds are not exposed to moist soil conditions in the fall, these polymer coats may prevent imbibition and subsequent germination during the following spring. Our objective was to determine the effects of moisture stress and polymer coat on canola seed germination characteristics in the laboratory. Polymer-coated and non-polymer-coated canola seeds were germinated in polyethylene glycol (PEG 8000) solutions with initial osmotic potentials ranging from 0 to -1.25 MPa at 5 and 15°C. Polymer coat treatments caused higher median germination times and lower final germination percentages compared with film-coated and uncoated control seeds. Germination characteristics of polymer-coated seeds were negatively influenced by decreasing initial osmotic potential, particularly at 5°C. Although not always significantly different from the uncoated control, film-coated control seeds behaved similarly to those coated with polymers. Among the three polymer coat treatments examined, one polymer coat treatment consistently exhibited higher germination compared with the other two. The results of this study help explain the occurrence of low spring seedling populations of fall-seeded, polymer-coated canola following a dry fall, particularly under low spring soil temperatures.

In western Canada, seasonal seedling recruitment has been reported in weedy canola populations, and seed persistence has been linked to the secondary seed dormancy potential of a genotype. Temperature influences secondary seed dormancy induction in this species. In these experiments we (1) investigated the influence of temperature and osmotic potential on secondary seed dormancy induction in canola, (2) related these to seedbank dynamics and seedling recruitment of two canola genotypes with different seed dormancy potentials in the field, and (3) investigated the influence of residue, burial depth, and soil type on seedbank dynamics and seedling recruitment in the field. In the laboratory, rates of seed dormancy induction were positively correlated to increasing temperatures and water stress. The role of temperature was approximately threefold more important to seed dormancy development than was osmotic potential within the tested ranges of these variables. In the field, seasonal seedbank dynamic...

Summary Secondary seed dormancy has been linked to seedbank persistence of volunteer oilseed rape (Brassica napus) in western Canada. It has been suggested that there is a genetic component to secondary seed dormancy expression in oilseed rape, but little is known of its importance in relation to non-genetic factors. In a series of experiments we investigated the relative importance of genotype, seed size, time of windrowing and pre- and post-harvest environment on the expression of secondary seed dormancy. We found that genotype contributed between 44 and 82% to the total variation in secondary seed dormancy. A broad range in secondary seed dormancy expression was observed among 16 genotypes examined. Nevertheless, three-quarters of the genotypes investigated exhibited relatively high potential for the expression of secondary seed dormancy (back-transformed mean 71% dormant seeds). Seed size contributed 21% to the total variation, while the influence of seed maturity (harvest regime) on secondary seed dormancy expression was negligible. Despite diverging environmental conditions during the four growing seasons spanning these experiments, the influence of pre-harvest environment on seed dormancy expression was relatively small and ranged from 0.1% to 4.5%. Secondary seed dormancy potential decreased over time during seed storage. This decrease was greatest when seeds were stored at ambient temperatures and least when seeds were stored at −70°C.