463 results on '"Skvarla, Michael J."'
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2. A Preliminary Phylogenetic Hypothesis for Cunaxidae (Acariformes: Trombidiformes: Prostigmata: Eupodina)
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Skvarla, Michael J., Dowling, Ashley P. G., Skvarla, Michael J., editor, Ochoa, Ronald, editor, Verle Rodrigues, Jose Carlos, editor, and Hutcheson, H. Joel, editor
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- 2019
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3. Tick abundance and life-stage segregation on the American black bear (Ursus americanus)
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Tiffin, Hannah S., Skvarla, Michael J., and Machtinger, Erika T.
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- 2021
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4. Distance models reveal biases associated with passive trapping methods for measuring wild bee abundance
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Mathis, Codey L., primary, McNeil, Darin J., additional, Kammerer, Melanie, additional, Larkin, Jeffery L., additional, and Skvarla, Michael J., additional
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- 2024
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5. First Records of Crapemyrtle Bark Scale (Hemiptera: Eriococcidae: Acanthococcus lagerstroemiae) from New Jersey and New York
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Skvarla, Michael J., primary, Chase, Kevin D., additional, Riley, Christopher B., additional, and Slone, Jeremy, additional
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- 2024
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6. First Modern Record of a Giant Lacewing (Neuroptera: Ithonidae: Polystoechotes punctata (Fabricius, 1793)) from Texas
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Skvarla, Michael J., primary, Oswald, John D., additional, and Janowiecki, Mark A., additional
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- 2024
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7. Report of Two Rove Beetle Species (Coleoptera: Staphylinidae), Oxybleptes kiteleyi Smetana, 1982 and Omalium rivulare (Paykull, 1789), Collected in Colored Water Traps
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Skvarla, Michael J., primary, Ferro, Michael L., additional, and Thayer, Margaret K., additional
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- 2023
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8. First Report of the Yellowmargined Leaf Beetle, Microtheca ochroloma Stål, 1860 (Coleoptera: Chrysomelidae), in Pennsylvania
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Skvarla, Michael J. and Fleischer, Shelby
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- 2018
9. Pre-ballooning in Ummidia Thorell 1875 (Araneae: Ctenizidae) from the Interior Highlands, USA: second account from the region and review of mygalomorph ballooning
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Fisher, J Ray, Fisher, Danielle M, Skvarla, Michael J, Dowling, Ashley P G, and BioStor
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- 2014
10. Litter-dwelling Ground Beetles (Coleoptera: Carabidae) and Ground Spiders (Araneae: Gnaphosidae) of the Ozark Highlands, USA
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Hamilton, Fredericka B., Wiedenmann, Robert N., Skvarla, Michael J., Sathyamurthy, Raghu, Fisher, Danielle M., Fisher, Jon Ray, and Dowling, Ashley P.G.
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- 2018
11. Online community photo-sharing in entomology: a large-scale review with suggestions on best practices
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Skvarla, Michael J, primary and Fisher, J. Ray, additional
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- 2023
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12. Correction to: Contemporary Acarology
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Skvarla, Michael J., Ochoa, Ronald, Verle Rodrigues, Jose Carlos, Hutcheson, H. Joel, Skvarla, Michael J., editor, Ochoa, Ronald, editor, Verle Rodrigues, Jose Carlos, editor, and Hutcheson, H. Joel, editor
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- 2019
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13. New Information about Lizard Beetles (Coleoptera: Erotylidae: Languria angustata (Beauvois, 1805) and Languria mozardi Latreille, 1807) in Pennsylvania
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Skvarla, Michael J., primary, Tooker, John, additional, and Murillo-Williams, Adriana, additional
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- 2023
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14. Notes on Pterostichus punctiventris (Chaudoir) (Coleoptera: Carabidae) in Arkansas, with a New State Record
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Hamilton, Ferdericka B., Skvarla, Michael J., Fisher, Danielle M., and Dowling, Ashley P. G.
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- 2016
15. New Information about the Cypress Weevil, Eudociminus mannerheimii (Boheman, 1836) (Coleoptera: Curculionidae: Molytinae): Redescription, Range Expansion, New Host Records, and Report as a Possible Causative Agent of Tree Mortality
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Skvarla, Michael J., Bertone, Matthew A., Fisher, J. Ray, and Dowling, Ashley P. G.
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- 2015
16. First Records of Crapemyrtle Bark Scale (Hemiptera: Eriococcidae: Acanthococcus lagerstroemiae) from New Jersey and New York.
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Skvarla, Michael J., Chase, Kevin D., Riley, Christopher B., and Slone, Jeremy
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This document is a list of references cited in a research paper or publication about the crapemyrtle bark scale, an insect that affects the crapemyrtle plant. The references include scientific articles, reports, and online resources that provide information on various aspects of the crapemyrtle bark scale, such as its biology, distribution, impact on plants, and management strategies. This document can be a valuable resource for library patrons conducting research on this topic. [Extracted from the article]
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- 2023
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17. First Modern Record of a Giant Lacewing (Neuroptera: Ithonidae: Polystoechotes punctata (Fabricius, 1793)) from Texas.
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Skvarla, Michael J., Oswald, John D., and Janowiecki, Mark A.
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- 2023
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18. Diversity of Arthropods that Visit Xyris spp. (Xyridaceae): New Observations from Guyana.
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Torres-Cruz, Terry J., Ré, Lauren A., Johnson, Jack R., Geiser, David M., and Skvarla, Michael J.
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Xyris spp., commonly known as yellow-eyed grasses, do not possess nectaries and were thought not to attract many insect visitors. The majority of insect visitation studies carried out on this genus have been limited to North American species, despite its center of diversity being in South America. The discovery of a potential new fungal mimicry system on Xyris spp. and the potential for insect visitation pattern alteration by the fungus led us to assess arthropod visitation to Xyris spp. in three regions of Guyana. Arthropods were photographed and identified morphologically. Here, we provide the first documentation of arthropods in the orders Araneae, Coleoptera, and Orthoptera visiting Xyris spp. plants. We also observed a Coleophora sp. (Lepidoptera: Coleophoridae) on Xyris spikes, expanding the geographic range of this taxon. [ABSTRACT FROM AUTHOR]
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- 2023
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19. Rediscovery of Polystoechotes punctata (Fabricius, 1793) (Neuroptera: Ithonidae) in Eastern North America
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Skvarla, Michael J., primary and Ray Fisher, J., additional
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- 2022
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20. Tissue‐damaging marking methods do not affect tick burdens on field captured Peromyscus spp.
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Roberts, Chloe E., primary, Burgess, Edwin R., additional, Miller, Taylor M., additional, Wise, Anna‐Marie, additional, Dickerson, Cera J., additional, Skvarla, Michael J., additional, Li, Andrew Y., additional, and Machtinger, Erika T., additional
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- 2022
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21. Limited detection of shared zoonotic pathogens in deer keds and blacklegged ticks co‐parasitizing white‐tailed deer in the eastern United States
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Olafson, Pia Untalan, primary, Poh, Karen C., additional, Evans, Jesse R., additional, Skvarla, Michael J., additional, and Machtinger, Erika T., additional
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- 2022
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22. Pre-ballooning in Ummidia Thorell 1875 (Araneae: Ctenizidae) from the Interior Highlands, USA: second account from the region and review of mygalomorph ballooning
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Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., and Dowling, Ashley P. G.
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- 2014
23. Limited detection of shared zoonotic pathogens in deer keds and blacklegged ticks co‐parasitizing white‐tailed deer in the eastern United States.
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Olafson, Pia Untalan, Poh, Karen C., Evans, Jesse R., Skvarla, Michael J., and Machtinger, Erika T.
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WHITE-tailed deer ,IXODES scapularis ,TICKS ,DEER ,BORRELIA burgdorferi ,LYME disease ,HISTORY of biology - Abstract
Deer keds, such as Lipoptena cervi Linnaeus (Diptera: Hippoboscidae), are blood‐feeding flies from which several human and animal pathogens have been detected, including Borrelia burgdorferi sensu lato Johnson (Spirochaetales: Borreliaceae), the causative agent of Lyme disease. Cervids (Artiodactyla: Cervidae), which are the primary hosts of deer keds, are not natural reservoirs of B. burgdorferi sl, and it has been suggested that deer keds may acquire bacterial pathogens via co‐feeding near infected ticks. We screened L. cervi (n = 306) and Ixodes scapularis Say (Ixodida: Ixodidae) (n = 315) collected from 38 white‐tailed deer in Pennsylvania for the family Anaplasmataceae, Bartonella spp. (Hyphomicrobiales: Bartonellaceae), Borrelia spp., and Rickettsia spp. (Rickettsiales: Rickettsiaceae). Limited similarity in the bacterial DNA detected between these ectoparasites per host suggested that co‐feeding may not be a mechanism by which deer keds acquire these bacteria. The feeding biology and life history of deer keds may impact the observed results, as could the season when specimens were collected. We separately screened L. cervi (n = 410), L. mazamae Róndani (n = 13), L. depressa Say (n = 10), and Neolipoptena ferrisi Bequaert (n = 14) collections from locations within the United States and Canada for the same pathogens. These results highlight the need to further study deer ked‐host and deer ked‐tick relationships. [ABSTRACT FROM AUTHOR]
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- 2023
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24. A quantitative method for collecting water mites in lotic, riffle-run habitats for water quality biomonitoring
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Stenger, Logan R., primary and Skvarla, Michael J., additional
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- 2022
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25. All for One Health and One Health for All: Considerations for Successful Citizen Science Projects Conducting Vector Surveillance from Animal Hosts
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Poh, Karen C., primary, Evans, Jesse R., additional, Skvarla, Michael J., additional, and Machtinger, Erika T., additional
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- 2022
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26. First Record of Crapemyrtle Bark Scale (Hemiptera: Eriococcidae: Acanthococcus lagerstroemiae) from Pennsylvania
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Skvarla, Michael J., primary and Schneider, Scott A., additional
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- 2022
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27. Additional file 1 of Patterns of deer ked (Diptera: Hippoboscidae) and tick (Ixodida: Ixodidae) infestation on white-tailed deer (Odocoileus virginianus) in the eastern United States
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Poh, Karen C., Evans, Jesse R., Skvarla, Michael J., Kent, Cody M., Olafson, Pia U., Hickling, Graham J., Mullinax, Jennifer M., and Machtinger, Erika T.
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Additional file 1: Figure S1. Rootogram showing performance of the model in the presence of zeroes in the dataset. The line and shaded area represent model predictions and 95% credible intervals. The histogram bars represent the actual data.
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- 2022
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28. Identification and Management of Cydalima perspectalis (Lepidoptera: Crambidae) in North America
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Coyle, David R, primary, Adams, Jamielee, additional, Bullas-Appleton, Erin, additional, Llewellyn, Jennifer, additional, Rimmer, Alexander, additional, Skvarla, Michael J, additional, Smith, Sandy M, additional, and Chong, Juang-Horng, additional
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- 2022
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29. The eriophyid mite Aculops ailanthi Lin, Jin, Kuang, 1997 (Acariformes: Prostigmata: Eriophyidae) from tree-of-heaven in the United States – new state records and morphological observations
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Skvarla, Michael J., Ochoa, Ron, Ulsamer, Andrew, Amrine, James, Pennsylvania State University (Penn State), Penn State System, United States Department of Agriculture (USDA), and West Virginia University [Morgantown]
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0106 biological sciences ,invasive pest ,Arthropoda ,Aacriformes ,[SDV.BID.SPT]Life Sciences [q-bio]/Biodiversity/Systematics, Phylogenetics and taxonomy ,Acariformes ,01 natural sciences ,Eriophyidae ,Botany ,Animalia ,Acari ,Trombidiformes ,tree-of-heaven ,Taxonomy ,Aculops ailanthi ,Ailanthus altissima ,biology ,Prostigmata ,Biodiversity ,biology.organism_classification ,Sapindales ,[SDV.BA.ZI]Life Sciences [q-bio]/Animal biology/Invertebrate Zoology ,010602 entomology ,Insect Science ,Simaroubaceae ,new state record - Abstract
We report Aculops ailanthi Lin, Jin, and Kuang, 1997 (Acariformes: Trombidiformes: Prostigmata: Eriophyidae) from Pennsylvania and West Virginia, USA; present the first scanning electron micrographs of the species in North America and discuss morphological observations that clarify features observed in slide-mounted specimens, such as the number ridges on female genital flaps; and briefly discuss symptoms and control of an A. ailanthi infestation on greenhouse-grown tree-of-heaven, Ailanthus altissima (Mill.) Swingle (Sapindales: Simaroubaceae)., Acarologia, 61, 121-127
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- 2021
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30. Murder Hornet Mayhem: The Impact of the 2020 Giant Hornet Panic and COVID-19 Pandemic on Arthropod Identification Laboratories.
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Skvarla, Michael J, Bertone, Matthew A, and Liesch, Patrick J
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ASIAN giant hornets , *COVID-19 , *HORNETS , *COVID-19 pandemic , *ARTHROPODA - Abstract
It was quickly apparent that many people were worried about GH, so it was noted in the lab databases whether clients mentioned terms such as "murder hornet", "Asian hornet", "giant hornet", "Russian hornet", or "middle hornet." On 2 May 2020, I The New York Times i (NYT) published a story with the headline "'Murder Hornets' in the U.S.: The Rush to Stop the Asian Giant Hornet", which detailed how giant hornets had been discovered in Washington State and British Columbia the previous fall ([4]). Graph European hornet ( I Vespa crabro i ), the species in eastern North America that is most often confused with I V. mandarinia i . Giant Hornets and Other Large Wasps Giant hornets (GH; I Vespa mandarinia i ) are large social wasps native to temperate and tropical east Asia, including Japan, China, India, and Sri Lanka, that were recently discovered in western North America ([18], [24], [25]; Fig. [Extracted from the article]
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- 2022
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31. Tales from the Field: Training Undergraduate Researchers for Fieldwork
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Poh, Karen C, primary, Tiffin, Hannah S, additional, Evans, Jesse R, additional, Brown, Jessica E, additional, Skvarla, Michael J, additional, and Machtinger, Erika T, additional
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- 2021
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32. First Record of Crapemyrtle Bark Scale (Hemiptera: Eriococcidae: Acanthococcus lagerstroemiae) from Pennsylvania.
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Skvarla, Michael J. and Schneider, Scott A.
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Crapemyrtle bark scale (Acanthococcus lagerstroemiae (Kuwana, 1907)) is an adventive pest that feeds on crapemyrtle (Lagerstroemia Linnaeus). First detected in North America in Dallas County, Texas in 2004, crapemyrtle bark scale has since spread patchily across the southeastern United States. The specimens reported here are the first recorded in Pennsylvania and represent the most northern occurrence of the species. [ABSTRACT FROM AUTHOR]
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- 2021
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33. A Review of Terrestrial and Canopy Malaise Traps
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Skvarla, Michael J, primary, Larson, Jonathan L, additional, Fisher, J Ray, additional, and Dowling, Ashley P G, additional
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- 2020
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34. A Technique for Dissecting the Salivary Glands From the Abdomens of Deer Keds (Diptera: Hippoboscidae: Lipoptena Nitzsch, 1818 and Neolipoptena Bequaert, 1942)
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Skvarla, Michael J, primary, Poh, Karen C, additional, Evans, Jesse R, additional, and Machtinger, Erika, additional
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- 2020
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35. First Report of Scale Insects (Hemiptera: Diaspididae: Lepidosaphes pallida (Maskell) on Russian Arborvitae (Pinales: Cupressaceae: Microbiota Decussata Kom.)
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Skvarla, Michael J., primary
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- 2020
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36. First Documented Report of Phytocoris buenoi Knight (Hemiptera: Miridae) Biting a Human
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Skvarla, Michael J., primary
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- 2020
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37. A planar interconnection technology utilizing the selective deposition of tungsten - process characterization
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Thomas, David Charles, Behfar-Rad, A., Comeau, Gerald L., Skvarla, Michael J., and Wong, S. Simon
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Integrated circuits -- Wafer-scale integration ,Integrated circuit fabrication -- Research ,Chemical vapor deposition -- Research ,Business ,Electronics ,Electronics and electrical industries - Abstract
The W interconnection technology is presented as a means of realizing multilevel interconnections and a planar surface after each level of metallization. The procedure involves the implantation of silicon (Si) or tungsten (W) into oxide channels after which selective deposition of tungsten occurs. The difference between Si and W implants in the nucleation of selective chemical vapor deposition of W on SiO2 is also discussed.
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- 1992
38. New Information about Anthocomus equestris (Fabricius, 1781) (Coleoptera: Melyridae), a Soft-Winged Flower Beetle Commonly Found in Homes
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Skvarla, Michael J., primary
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- 2019
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39. Range Extension of False Meyer Scale, Dynaspidiotus pseudomeyeri (Kuwana) (Hemiptera: Diaspididae) in the United States, and Newly Recorded Associations with a Parasitoid and Mite
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Schneider, Scott A., primary, Skvarla, Michael J., additional, Ochoa, Ron, additional, Schmidt, Stefan, additional, Polaszek, Andrew, additional, and Gates, Michael W., additional
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- 2019
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40. Deer Keds (Diptera: Hippoboscidae: Lipoptena and Neolipoptena) in the United States and Canada: New State and County Records, Pathogen Records, and an Illustrated Key to Species
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Skvarla, Michael J, primary and Machtinger, Erika T, additional
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- 2019
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41. A Review of Terrestrial and Canopy Malaise Traps.
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Skvarla, Michael J, Larson, Jonathan L, Fisher, J Ray, and Dowling, Ashley P G
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TRAPPING , *LITERARY criticism , *LITERATURE reviews , *OPTICAL tweezers - Abstract
Malaise traps are important tools for the large-scale collection of arthropod taxa. Here, an extensive review of the history and literature concerning Malaise and canopy traps is given. This review highlights how trap design and placement can affect trap catch, which will help researchers to make more informed choices when planning research endeavors. Additionally, terrestrial and canopy-style Malaise traps are compared to each other and other types of arthropod traps. [ABSTRACT FROM AUTHOR]
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- 2021
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42. Tetrabdella Hernandes & Feres
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Tetrabdella ,Taxonomy - Abstract
Tetrabdella Hernandes & Feres Type-species: Tetrabdella neotropica Hernandes & Feres, 2006 by original designation. 158. Tetrabdella neotropica Hernandes & Feres, 2006: 60; Cedral, S��o Paulo, Brazil, ex Hevea brasiliensis M.Arg. (Euphorbiaceae). Type deposition. Holotype and paratypes at DZSJRP, paratype at USNM., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 30, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Hernandes, F. A. & Feres, R. J. F. (2006) Tetrabdella neotropica (Acari: Bdellidae), a new genus and species from Brazil. Zootaxa, 1135, 57 - 68."]}
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- 2016
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43. Neomolgus Oudemans
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Neomolgus ,Taxonomy - Abstract
Neomolgus Oudemans Molgus Dujardin, 1842: 316 (nomen nudum); Molgus Trouessart, 1894: 118; Thor, 1913: 30; 1931a: 54. Molgus (Hoplomolgus) Berlese, 1923: 237. Neomolgus Oudemans, 1937: 1229; Atyeo, 1960a: 389; Soliman, 1975: 48; Soliman & Zaher, 1975: 78; Wallace & Mahon, 1976: 116; Michocka, 1987: 44. Type-species: Acarus littoralis Linnaeus, 1758 by original designation. 251. Neomolgus aegyptiacus Soliman & Mohamed, 1972b: 90; Faculty of Agriculture farm, Giza, Egypt, from clover field associated with Bryobia cristata (Acari, Tetranychidae). Remarks. Male unknown; although the species was formally described in 1975, three years earlier Soliman & Mohamed (1972b) published the biology of this species along with a morphological description of all stages. The authority and date must be therefore attributed to Soliman & Mohamed, 1972b. Type deposition. Probably at CUE. 252. Neomolgus aequalis (Schweizer & Bader, 1963): 238 comb. nov.; Kreuzlingen, Switzerland, ex moss. Original designation: Hoplomolgus aequalis Schweizer & Bader. Type deposition. NMB (A 211a). 253. Neomolgus berlesei (Tr��g��rdh, 1902b): 17 comb. nov.; in humid areas and under stones, Italy. Original designation: Bdella capillata var. berlesei Tr��g��rdh, 1902b: 17. Other name: Molgus (Hoplomolgus) berlesei, Thor, 1931a: 57. a. Molgus (Hoplomolgus) tuberculatus Berlese, 1923: 238 synonymy according to Thor (1931a: 57); Hoplomolgus tuberculatus, Schweizer & Bader, 1963: 242. Type deposition. Unknown. 254. Neomolgus capillatus (Kramer, 1881): 446; Germany���Willmann, 1939: 431; 1956: 243. Original designation: Bdella capillata Kramer; Canestrini, 1886: 188; Berlese, 1891; Michael, 1896: 514; Kramer, 1896: 447; Tr��g��rdh, 1902b: 17; B��bler, 1910: 812; Halbert, 1915: 112; Hull, 1918: 40. Other names: Molgus capillatus, Trouessart, 1888: 619; Thor, 1904b: 77; 1909: 7; Oudemans, 1914: 123; Schweizer, 1922: 79; Thor, 1926: 139; Thor, 1930a: 95; Hoplomolgus capillata, Tr��g��rdh, 1931: 49; Molgus (Hoplomolgus) capillatus, Vitzthum, 1929: 59; Molgus (Molgus) capillatus, Thor, 1931a: 60. Distribution. Germany (Voigts & Oudemans 1906), Ireland (Hull 1915), Germany, Switzerland, Italy, England, Ireland, Norway, Svalbard Islands (Thor 1931a), France (Schweizer 1922), Austria (Willmann 1951), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Australia (south western W.A.) (Wallace & Mahon 1976), Ukraine (Kuznetsov & Livshits 1979a), Poland (Michocka 1987), Iran (Ostovan & Kamali 1995; Kamali et al. 2001). Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976), Michocka (1987). Type deposition. CRA. 255. Neomolgus clypeatus (Thor, 1930b): 17; Sjus��rn (Lillehammer), Norway.��� Atyeo & Tuxen, 1962: 290. Original designation: Molgus clypeatus Thor. Other name: Molgus (Hoplomolgus) clypeatus, Thor: 1931a: 55. Distribution. Iceland (Atyeo & Tuxen 1962), Syria (Lattakia) (Soliman & Zaher 1975), Australia (Wallace & Mahon 1976), Ukraine (Kuznetsov & Livshits 1979a), Iran (Ostovan & Kamali 1995; Kamali et al. 2001). Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976). Type deposition. Lost. 256. Neomolgus egregia (Koch, 1839): 23 comb. nov.; in humid meadows, Europe. Original designation: Bdella egregia Koch; Koch, 1842: 74; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 184; Canestrini & Fanzago, 1877: 101; Oudemans, 1937: 1196. Other name: Molgus egregious, Thor, 1931a: 63. Bdella vulgaris, Canestrini, 1886: 183; Berlese, 1888: 7; 1893: 43; Halbert, 1915: 113; Schweizer & Bader, 1963: 230; synonymy according to Thor (1931a: 63). Bdella longitarsa Karsch, 1881: 37; Molgus egregius var. longitarsa, Thor, 1931a: 63. Distribution. Middle Europe (Germany, France, Italy, Sweden) (Thor 1931a), Japan (Karsch 1881), Ireland (Hull 1915). Type deposition. Unknown; type of B. longitarisa in ZMB. 257. Neomolgus helveticus (Schweizer & Bader, 1963): 239 comb. nov.; Buffalora mountain, National Park, Switzerland, under stone. Original designation: Hoplomolgus helveticus Schweizer & Bader. Type deposition. NMB (NP 1471). 258. Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh & Ueckermann, 2016: 296; ex soil and litter under wild almond trees, Amygdalus scoparia L. (Rosaceae), Palangan village, Kamyaran, Kermanshah Province, Iran. Type deposition. Holotype and paratypes at University of Bu-Ali Sina, Hamedan, Iran; paratype female at NCA. 259. Neomolgus lacustris (Hull, 1915): 121 comb. nov.; on soil, England. Original designation: Bdella lacustris Hull.���Hull, 1918: 40. Other name: Molgus (Molgus) lacustris, Thor, 1931a: 62. a. Molgus (Hoplomolgus) sublimis Berlese, 1923: 238 synonymy according to Thor (1931a: 62). Type deposition. Unknown. 260. Neomolgus littoralis (Linnaeus, 1745): 96; unknown locality.��� Atyeo, 1960a: 390. Original designation: Acarus littoralis Linnaeus, 1745: 96; 1758: 618; Hammer, 1775: 158; Fabricius, 1775: 815; 1780: 225; 1781: 493; 1787: 374; M��ller, 1776: 187; Oudemans, 1926b: 117; Johnston, 1845: 229; Oudemans, 1929: 316. Other names: Bdella littoralis, Neuman, 1875: 104; Hull, 1918: 40; 1922: 622; Oudemans, 1926a: 116; Tr��g��rdh, 1904: 46; 1912: 419; Bdella vulgaris var. littoralis Moniez, 1890a: 30, 1890b: 196. a. Bdella littoralis + capillata var. pallipediformis Tr��g��rdh, 1902b: 18; synonymy according to Thor (1931a: 61); Molgus littoralis, Thor, 1904a: 15; 1904b: 78; 1913: 30; 1928: 216; 1930a: 96; 1931a: 61; King, 1914: 138; Berlese, 1923: 237; Tr��g��rdh, 1928: 8; 1931: 47. Tr��g��rdh, 1928: 8; 1931: 47. b. Bdella groenlandica Tr��g��rdh, 1904: 48; Hull, 1922: 622; synonymy according to Thor (1930a). c. Molgus (Molgus) littoralis var. seurati Berlese, 1923: 237; synonymy according to Thor (1931a). d. Acarus petrarum ruber L.; Str��m, 1762: 196; Olafsen & Povelsen, 1772: 607; synonymy according to Thor (1904). e. Acarus basteri Johnston, 1836: 353; Bdella basteri; Michael, 1896: 478; synonymy according to Thor (1904). f. Acarus longicornis, Johnston, 1845: 227; synonymy according to Thor (1904). g. Bdella podurophila White, 1852: 210; synonymy according to Thor (1931a). h. Bdella arctica Thorell, 1871: 698; Kramer, 1897: 79; Tr��g��rdh, 1900: 9; Banks, 1919: 11; Molgus arcticus (Thorell), Trouessart, 1888: 119; Thor, 1902: 162; synonymy according to Thor (1904). i. Bdella marina Packard, 1873: 544; 1884: 828; Banks, 1894: 220; Michael 1896: 479; Banks 1907: 596; 1908: 5; Thor, 1931a: 39; synonymy according to Thor (1904). j. Molgus longicornis, Murray, 1877: 143; synonymy according to Thor (1931a). k. Bdella grandis Kramer, 1879: 131; synonymy according to Tr��g��rdh (1904: 46). l. Bdella villosa Kramer & Neuman, 1883: 525; Michael 1896: 479; Oudemans & Koenike, 1897: 238; Banks, 1899: 348; 1907: 596; Molgus villosa, Trouessart, 1888: 619; synonymy according to Thor (1904). m. Eupalus sanguineus Trouessart, 1888: 753; Bdella sanguinea, Trouessart, 1894: 119; Michael 1896: 478; Tr��g��rdh, 1900: 9; Molgus sanguineus, Trouessart, 1894: 117; synonymy according to Thor (1904). n. Bdella villosa + frigida Banks, 1899: 348; synonymy according to Thor (1931a). Distribution. England (Hull 1918), Canada (Nunavut) (Summerhayes & Elton 1928), Alaska, Hudson Bay Area, Greenland, Iceland, Spitsbergen, Bering Island, Russian Lapland (Kola-Hafvon), Novaya Zemlya (Matochkin Strait), Finland (Atyeo 1960a), Japan (Ehara 1960, 1961, Shiba 1971), Iceland (Atyeo & Tuxen 1962). Redescriptions. Atyeo (1960a), Atyeo & Tuxen (1962). Remarks. Atyeo (1960a: 391) reported specimens from Matochkin Strait lacking trichobothria on tibia IV, and having a long tactile seta in the position normally occupied by the trichobothria. Type deposition. Unknown. 261. Neomolgus longipalpis (Karpelles, 1893): 106 comb. nov.; Maramures, Hungary, at Pop Ivan mountain. Original designation: Bdella longipalpis Karpelles. Other name: Molgus longipalpis, Thor, 1931a: 63. Type deposition. Unknown. 262. Neomolgus longipalpus Kuznetsov, 1984: 774; park of the Nikita botanical garden, Yalta, Crimea, ex litter. Remarks. Male unknown. According to ICZN (article 57.6), the one-letter difference in spelling in relation to the previous species is enough to prevent the homonymy. Type deposition. NBG. 263. Neomolgus lumarius Atyeo, 1962: 292 in Atyeo & Tuxen; H��itindur, Southeast Iceland. Type deposition. Holotype and paratypes at ZMUC; paratypes at USNM, BMNH and SAM. 264. Neomolgus maculatus (Karpelles, 1893): 26 comb. nov.; Sălaj, Hungary. Original designation: Bdella maculata Karpelles. Other names: Molgus maculatus, Thor, 1931a: 64; Molgus maculatus var. pilosa; Thor, 1931a: 64. Type deposition. Unknown. 265. Neomolgus monticola Willmann, 1951: 162; Hohe Tauern, Switzerland. Other name: Hoplomolgus monticola, Schweizer & Bader, 1963: 240. Type deposition. ZSM. 266. Neomolgus mutabilis Atyeo, 1960a: 392; 2 miles south of Galena, Cherokee Co., Kansas, USA, under board. Distribution. United States (Kansas, Texas), Mexico (Distrito Federal) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 267. Neomolgus obsoletus (Berlese, 1923): 238 comb. nov.; in Germany and Norway. Original designation: Molgus (Hoplomolgus) obsoletus, Thor, 1931a: 58; Hoplomolgus obsoletus, Schweizer & Bader, 1963: 242. Type deposition. CRA. 268. Neomolgus ontakensis Shiba, 1971: 99; Mt. Ontak��, Nagano pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 269. Neomolgus pallipes (Koch, 1879): 131; Siberia, Russia. Original designation: Bdella pallipes Koch; Hull, 1918: 41; Hull, 1922: 622; Neomolgus pallipes; Mihelčič, 1958a; Willmann, 1952: 166. a. Bdella hirta Kramer, 1896: 447; synonymy according to Thor (1931a: 58). b. Bdella basteri Johnston forma pallipes, Tr��g��rdh, 1901: 61; synonymy according to Thor (1931a: 58). c. Bdella capillata var. pallipes, Tr��g��rdh, 1902b: 16; 1904: 47; synonymy according to Thor (1931a: 59). d. Molgus (Molgus) pallipes, Thor, 1931a: 58. e. Molgus capillatus, Thor, 1927: 139; 1930: 95; synonymy according to Thor (1931a: 59); Molgus capillatus var. pallipes, Tr��g��rdh, 1928: 8; synonymy according to Thor (1931a: 59). Distribution. Canada (Summerhayes & Elton 1928), Northern Europe and Asia (Siberia, Novaja, Semlja, Sweden, Norway, Svalbard, England, Borkum Island) (Thor 1931a), Germany (Wangerooge Island) (Willmann 1952), Spain (Mihelčič 1958a). Type deposition. BMNH. 270. Neomolgus paracapillatus Michocka, 1987: 45; Warsaw, Poland, ex decayed grass. Type deposition. In the author���s collection. 271. Neomolgus pratensis Shiba, 1971: 93; Namekawa, Kawauchi-ch��, Ehime pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 272. Neomolgus pygmaeus Shiba, 1969a: 74; Shiga Heights, Japan. Distribution. Japan (Shiba 1969a), China (Lin et al. 2006). Redescription. Shiba (1971). Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 273. Neomolgus raeticus (Schweizer & Bader, 1963): 238 comb. nov.; NationalPark, Mt. Mezdi, Switzerland. Original designation: Hoplomolgus raeticus Schweizer & Bader. Type deposition. NMB (NP 1845). 274. Neomolgus raptor Kuznetsov & Barilo, 1984: 934; near Samarkand (Tchapan-Atha), Uzbekistan, under stones. Type deposition. NBG. 275. Neomolgus reticulatus (Schweizer & Bader, 1963): 241 comb. nov.; S-chanf, Switzerland. Original designation: Hoplomolgus reticulatus Schweizer & Bader. Type deposition. NMB (NP 283). 276. Neomolgus sabulosus Shiba, 1971: 96; Shigenobu River, Matsuyama, Ehime pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 277. Neomolgus seriatus Shiba, 1971: 101; Otanom��su-no-taira, Shigak��gen, Nagano pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 278. Neomolgus thorianus (Berlese, 1923): 239 comb. nov.; Corfu, Greece. Original designation: Molgus (Hoplomolgus) thorianus Berlese. Other name: Molgus (Molgus) thorianus, Thor, 1931a: 62. Distribution. Greece (Corfu Island) (Thor 1931a), Georgia (Gomelauri 1963b). Type deposition. 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44. Odontoscirus Thor
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Odontoscirus ,Animalia ,Bdellidae ,Biodiversity ,Taxonomy - Abstract
Odontoscirus Thor Odontoscirus Thor, 1913: 29; Thor, 1931a: 52 (as subgenus of Biscirus); type-species Bdella virgulata Canestrini & Fanzago by original designation; Meyer & Ryke, 1959: 379; Atyeo, 1960a: 386; Michocka, 1987: 50; Wallace & Mahon 1976: 67. Bdellodes Oudemans, 1937: 1217; Meyer & Ryke, 1959: 378; Wallace & Mahon, 1976: 67; Michocka, 1987: 56. Type-species: Scirus longirostris Hermann, 1804 by original designation. Scirus Hermann, 1804: 61; Thor, 1931a: 41 (part.). Hoploscirus Thor, 1937: 43; type-species Scirus dubitatus Womersley, 1933a by original designation; synonymy according to Wallace & Mahon (1976: 67). Thoribdella Grandjean, 1938: 4 type-species Biscirus meridionalis Thor, 1931a by original designation; synonymy according to Wallace & Mahon (1976: 67). Octobdellodes Atyeo, 1960a: 407 type-species O. hurdi Atyeo, 1960a; synonymy according to Atyeo (1963a: 118, 125). 159. Odontoscirus affinis (Atyeo, 1963a): 185 comb. nov.; Rottnest Island, Australia. Original designation: Bdellodes (Hoploscirus) affinis Atyeo. Distribution. Australia (Atyeo 1963a; Wallace & Mahon 1976), Andaman Islands (Gupta & Ghosh 1980). Redescriptions. Wallace & Mahon (1976), Gupta (2002). Type deposition. Holotype and paratypes at SAM, paratypes at BMNH, USNM, ANIC. 160. Odontoscirus agrestis (Atyeo, 1963a): 152 comb. nov.; Auckland, New Zealand, ex moss and lichens. Original designation: Bdellodes (Hoploscirus) agrestis Atyeo. Remarks. Male unknown. Type deposition. Holotype and paratype at MONZ, paratype at NZFRI. 161. Odontoscirus alacris (Atyeo, 1963a): 193 comb. nov.; Glen Osmond, South Australia. Original designation: Bdellodes (Hoploscirusu) alacris Atyeo. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 103). Distribution. Australia (Atyeo, 1963a, Wallace & Mahon, 1976). Redescription. Wallace & Mahon (1976). Type deposition. SAM. 162. Odontoscirus alpinus Atyeo, 1960a: 388; Piegan Pass, Glacier National Park, Montana, USA. Type deposition. SEMC. 163. Odontoscirus amamiensis Shiba, 1985: 81; Southern Japan (Kyushu). Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 164. Odontoscirus ancalae (Atyeo, 1963a): 148 comb. nov.; Wellington, New Zealand, ex leaf mold. Original designation: Bdellodes (Hoploscirus) ancalae Atyeo. Remarks. Male unknown. Type deposition. MONZ. 165. Odontoscirus angustifolius (Gupta, 1991): 221 comb. nov.; Moreh, Manipur, India, ex Canthium angustifolium (Rubiaceae)��� Gupta, 2002: 31. Original designation: Bdella angustifolius Gupta. Bdellodes angustifolius, Gupta, 2002. Remarks. Male unknown. Redescription. Gupta (2002). Type deposition. NZSI. 166. Odontoscirus annona (Tseng, 1978): 33 comb. nov., new emendation; Kuiyin, Tainan Hsien, Taiwan, ex Annona sp. (Annonaceae). Original designation: Bdellodes anona Tseng. Remarks. The etymology of the species refers to the host plant, originally misspelled as Anona sp. [sic] rather than Annona sp.; therefore this species is herein emendated to Odontoscirus annona. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 167. Odontoscirus asiaticus Kuznetsov & Barilo, 1984: 934; near Samarkand, Agalyk village, Uzbekistan, ex under stones on humid soil; 3 males and 1 female paratypes on low grass and under stones in Samarkand region near Zeravshan mountain ridge. Type deposition. NBG. 168. Odontoscirus atro (Gupta, 1991): 220 comb. nov.; Arunachal Pradesh, Siji, India, ex Viburnum atrocyaneum Clarke (Adoxaceae). Bdellodes atro, Gupta, 2002: 32. Original designation: Bdella atro Gupta. Remarks. Male unknown. Redescription. Gupta (2002). Type deposition. NZSI. 169. Odontoscirus atyeoi Michocka, 1987: 54; Tatra, Poland. Type deposition. In the author���s collection. 170. Odontoscirus augusta (Roy & Saha, 2010): 121 comb. nov.; West Bengal, India. Original designation: Bdellodes augusta Roy & Saha. Type deposition. DZCU. 171. Odontoscirus australicus (Womersley, 1933a): 107 comb. nov.; Waroona, Western Australia. Original designation: Biscirus (Biscirus) australicus Womersley. Other name: Bdellodes (Hoploscirus) australica, Atyeo, 1963a: 188 (neotype designation). Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Type deposition. Neotype female at SAM. 172. Odontoscirus bidentata (Wallace & Mahon, 1976): 94 comb. nov.; Omeo, Australia, in mountain gully. Original designation: Bdellodes bidentata Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 173. Odontoscirus bifurcata (El-Sherif & Bolland, 1993): 2237 comb. nov., from litter near railway in Amsterdam, The Netherlands. Original designation: Bdellodes bifurcata El-Sherif & Bolland. Type deposition. Unknown. 174. Odontoscirus bisetosa (Atyeo, 1960a): 414 comb. nov.; 10 miles west of Tuxtla Gutierrez, Chiapas, Mexico, under rock. Original designation: Bdellodes bisetosa Atyeo. Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 175. Odontoscirus brevicornis (Cooremann, 1959): 30 comb. nov.; Muotathal, Switzerland, in cave. Original designation: Thoribdella brevicornis Cooremann. Type deposition. IRSN. 176. Odontoscirus bryi (Atyeo, 1963a): 142 comb. nov.; Otago, New Zealand, ex moss on stones. Original designation: Bdellodes (Hoploscirus) bryi Atyeo. Remarks. Male unknown. Type deposition. MONZ. 177. Odontoscirus californica (Banks, 1904b): 366 comb. nov.; Claremont, LA, USA. Original designation: Bdella californica Banks; Banks, 1907: 596; Thoribdella californica (Banks), Atyeo, 1960a: 396; Bdellodes californica, Hernandes, 2013: 65. Bdella magna Ewing, 1913: 123; synonymy according to Atyeo (1960a). Distribution. United States (Los Angeles, California) (Atyeo 1960a). Redescription. Atyeo (1960a). Thor (1931a) has strangely regarded Bdella californica Banks as both synonyms of Bdellodes longirostris and of Biscirus uncinatus (pages 42 and 50, respectively). Type deposition. MCZ. 178. Odontoscirus camellae (Atyeo, 1963a): 145 comb. nov.; Wellington, New Zealand, ex leaf mold. Original designation: Bdellodes (Hoploscirus) camellae Atyeo. Remarks. Female unknown. Type deposition. MONZ. 179. Odontoscirus communis (Atyeo, 1960a): 399 comb. nov.; west slope of Cortez Pass, Mexico, under rock. Original designation: Thoribdella communis Atyeo. Other name: Bdellodes (Hoploscirus) commnunis, Atyeo, 1963a: 188. Type deposition. Holotype and paratypes at SEMC, paratypes at BMNH, USNM and SAM. 180. Odontoscirus conformis (Atyeo, 1963a): 137 comb. nov.; Otago, New Zealand, beaten off ferns. Original designation: Bdellodes (Hoploscirus) conformis Atyeo. Remarks. Male unknown. Type deposition. MONZ. 181. Odontoscirus consanguinea (Atyeo, 1963a): 194 comb. nov.; Busselton, Western Australia. Original designation: Bdellodes (Hoploscirus) consanguinea Atyeo. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 84). b. Biscirus intermedius (part.) (Thor, 1928: 213); Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 84). Distribution. Australia (Atyeo 1963a), (WA, SA, Victoria, Tasmania, NSW, south eastern Queensland, Kununurra area WA) (Wallace & Mahon 1976). Redescription. Wallace & Mahon (1976). Type deposition. SAM. 182. Odontoscirus copiosa (Atyeo, 1963a): 136 comb. nov.; Otago, New Zealand, under stone. Original designation: Bdellodes (Hoploscirus) copiosa Atyeo. Type deposition. MONZ. 183. Odontoscirus currax (Atyeo, 1963a): 187 comb. nov.; Moount Toolbrunnup, Western Australia, on mealy litter and moss. Original designation: Bdellodes (Hoploscirus) currax Atyeo. Redescription. Wallace & Mahon (1976). Type deposition. SAM. 184. Odontoscirus curvus (Atyeo, 1963a): 140 comb. nov.; Auckland Islands, New Zealand, under stone at high tide mark. Original designation: Bdellodes (Hoploscirus) curvus Atyeo. Redescription. Wallace (1970). Type deposition. MONZ. 185. Odontoscirus dubitatus (Womersley, 1933a): 102 comb. nov.; Mount Nelson, Tasmania, under stone. Bdellodes dubitata, Wallace & Mahon, 1976: 78. Original designation: Scirus dubitatus Womersley. Other name: Bdellodes (Hoploscirus) dubitata, Atyeo, 1963a: 196. Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Remarks. Male unknown, species known only from the holotype. Type deposition. SAM. 186. Odontoscirus edentata (Halliday, 2005): 22 comb. nov.; Port Elizabeth, Marine Protea Hotel, South Africa. Original designation: Bdellodes edentata Halliday. Remarks. Male unknown. Type deposition. Holotype and paratypes at NCA, paratype at ANIC. 187. Odontoscirus exilicornis (Berlese, 1910): 347 comb. nov.; Cape Town, South Africa. Bdellodes exilicornis, Meyer & Ryke, 1959: 379. Original designation: Bdella exilicornis Berlese. Other name: Scirus exilicornis, Thor, 1931a: 44. Type deposition. Lost (Castagnoli & Pegazzani, 1985). 188. Odontoscirus flexuosa (Atyeo, 1963a): 139 comb. nov.; Campbell Island, New Zealand, under stone. Original designation: Bdellodes (Hoploscirus) flexuosa Atyeo; Atyeo, 1964: 167. Redescriptions. Wallace (1970). Type deposition. MONZ. 189. Odontoscirus furcatus (Shiba, 1969a): 73 comb. nov.; Shiga Heights, Japan. Original designation: Bdellodes (Hoploscirus) furcatus Shiba; Nakamura et al. (2006). Type deposition. Unknown. 190. Odontoscirus georgianensis (Wallace, 1970): 109 comb. nov.; Royal Bay, Moltke Harbor, South Georgia, Antarctica, under moss and rocks. Original designation: Bdellodes (Bdellodes) georgianensis Wallace. Type deposition. Holotype at BPBM, paratypes at ANIC and BMNH. 191. Odontoscirus gleba Chaudhri & Akbar, 1985: 118; 1 mile from W. Hassanabdal, Pakistan. Type deposition. UAF. 192. Odontoscirus graminis (Wallace & Mahon, 1976): 105 comb. nov.; Pine Creek, Northern Territory, Australia, ex grass, Eucalyptus sp. Original designation: Bdellodes graminis Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 193. Odontoscirus grandiflora (Gupta, 1991): 224 comb. nov.; Arunachal Pradesh, Garu, India, ex Thunbergia grandiflora Roxb. (Acanthaceae). Original designation: Bdellodes grandiflora Gupta. Type deposition. NZSI. Remarks. Male unknown. Redescription. Gupta (2002). 194. Odontoscirus gressitti (Atyeo, 1964): 167 comb. nov.; Beeman Camp, Campbell Island, New Zealand, ex moss. Original designation: Bdellodes (Hoploscirus) gressitti Atyeo. Redescriptions. Wallace (1970). Type deposition. Holotype at MONZ; paratypes at BPBM. 195. Odontoscirus guajavae (Chatterjee & Gupta, 2002): 34 (in Gupta 2002) comb. nov.; Kalyani, West Bengal, India, ex Psidium guajava (Myrtaceae). Original designation: Octobdellodes guajavae Chatterjee & Gupta. Type deposition. ZSI. 196. Odontoscirus hadroseta (Wallace & Mahon, 1976): 110 comb. nov.; Nyabing, Western Australia, Australia, on sand plain scrub on roadside. Original designation: Bdellodes hadroseta Wallace & Mahon. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 110). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 110). c. Biscirus uncinatus (Kramer 1898: 12), Womersley, 1933a: 106; synonymy according to Wallace & Mahon (1976: 110). Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 197. Odontoscirus haramotoi (Swift & Goff, 1987): 35 comb. nov.; Koa, Hawaii, ex litter. Original designation: Bdellodes haramotoi Swift & Goff. Remarks. Female unknown. Type deposition. BPBM. 198. Odontoscirus harpax (Atyeo, 1963a): 128 comb. nov.; Woods Point, South Australia. Original designation: Bdellodes (Bdellodes) harpax Atyeo. a Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 115). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 115). c. Scirus longirostris Hermann (part.), Womersley, 1933a: 101; synonymy according to Wallace & Mahon (1976: 115). Distribution. Australia, New Zealand, Tasmania (Atyeo, 1963a, Wallace & Mahon, 1976). Redescription. Wallace & Mahon (1976). Type deposition. Holotype and paratypes at SAM, paratypes at NZFRI, MONZ, BMNH, USNM. 199. Odontoscirus hessei (Womersley, 1933c): 111 comb. nov.; Stellenbosch C.P., South Africa.��� Meyer & Ryke, 1959: 379. Original designation: Scirus hessei Womersley. Other name: Bdellodes (Bdellodes) hessei, Atyeo, 1963a: 181. Distribution. South Africa (Meyer & Ryke 1959; Atyeo 1963a; Halliday 2005), Australia (Perth, Nedland) (Atyeo 1963a), Western Australia (Wallace & Mahon 1976). Redescriptions. Meyer & Ryke (1959), Atyeo (1963a), Wallace & Mahon (1976). Type deposition. SAM. 200. Odontoscirus hickmani (Womersley, 1933a): 107 comb. nov.; National Park, Tasmania, Australia, under stones.��� Wallace & Mahon, 1976: 81. Original designation: Biscirus (Biscirus) hickmani Womersley. Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Remarks. Male unknown. Type deposition. SAM. 201. Odontoscirus hospita (Banks, 1916): 224 comb. nov.; Victoria, Australia, with Polyrhachis hexacantha (Insecta, Formicidae).��� Wallace & Mahon, 1976: 91. Original designation: Bdella (Scirus) hospita Banks. Other name: Bdellodes (Hoploscirus) hospita, Atyeo, 1963a: 19. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 91). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 91). Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Type deposition. Cotypes at USNM and SAM. 202. Odontoscirus hurdi (Atyeo, 1960a): 408 comb. nov.; Point Barrow, Alaska, USA. Original designation: Octobdellodes hurdi Atyeo. Other name: Bdellodes (Bdellodes) hurdi; Atyeo, 1963a: 125. Type deposition. Holotype and paratypes at SEMC, paratypes at BMNH, USNM, SAM, CAS. 203. Odontoscirus hygrotes (Swift & Goff, 1987): 32 comb. nov.; Oahu I, Maui I, Hawaii. Original designation: Bdellodes hygrotes Swift & Goff. Type deposition. BPBM. 204. Odontoscirus inflata (Wallace & Mahon, 1976): 78 comb. nov.; Mandurah, Western Australia, coastal sanddune vegetation. Original designation: Bdellodes inflata Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 205. Odontoscirus infrequens (Atyeo, 1960a): 410 comb. nov.; Douglas County, Kansas, USA, on shagbark hickory. Original designation: Octobdellodes infrequens Atyeo. Remarks. According to Atyeo (1960a: 411), this species has 7 to 8 pairs of hypostomal setae rather than the usual 6 pairs. Type deposition. Holotype and paratypes at SEMC, paratypes at USNM. 206. Odontoscirus insolita (Atyeo, 1960a): 398 comb. nov.; 2 miles of west Oakville, Napa Co., California, USA, ��� Laurel association���. Original designation: Thoribdella insolita Atyeo. Type deposition. SEMC. 207. Odontoscirus intermedius (Thor, 1928): 213 comb. nov.; Norway. Original designation: Biscirus (Biscirus) intermedius Thor; Thor, 1931a: 48. Other names: Biscirus intermedius, Schweizer & Bader, 1963: 236; Thoribdella intermedius, Grandjean, 1938: 4; Bdellodes (Hoploscirus) intermedius, Atyeo, 1963a: 128. Distribution. Norway (Thor 1931a), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Ukraine (Kuznetsov & Livshits 1979a). Type deposition. Lost. 208. Odontoscirus, Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 31-41, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Thor, S. (1913) Biscirus genus novum: eine neue Bdelliden-Gattung und zwei neue Untergattungen. Zoologischer Anzeiger, 42, 28 - 30.","Thor, S. (1931 a) Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, 56, 87 pp.","Meyer, M. K. P. & Ryke, P. A. J. (1959) Cunaxoidea (Acarina: Prostigmata) occurring on plants in South Africa. Annals and Magazine of Natural History, 13 (2), 369 - 384. http: // dx. doi. org / 10.1080 / 00222935908655745","Atyeo, W. T. 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Sitzungsberichte der Mathematisch-naturwissenschaftliche Klasse, Abteilung I, 167, 35 - 57.","Voigts, H. & Oudemans, A. C. (1906) Zur Kenntnis der Milben-Fauna von Bremen. Bdellidae. In: Leuwer, F. (Ed.), Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, pp. 241 - 242.","Thor, S. (1930 a) Beitrage zur kenntnis der invertebraten fauna von Svalbard. Skrifter Svalbard og Ishavet, 27, 1 - 156.","Dumbleton, L. J. (1932) Report on spruce aphis investigation for the year ending December 1930. New Zealand Journal of Science and Technology, 13 (4), 207 - 20.","Willmann, C. (1952) Die Milbenfauna der Nordseeinsel Wangerooge. Veroffentlichungen des Instituts fur Meeresforschung Bremerhaven, 1, 139 - 186.","Atyeo, W. T. & Tuxen, S. L. (1962) The Icelandic Bdellidae (Acarina). Journal of the Kansas Entomological Society, 35 (3), 281 - 298.","Kamali, K., Ostovan, H. & Atamehr, A. (2001) A catalog of mites & ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center. 204 pp.","Barbar, Z. (2016) The mite fauna (Acari) of two Syrian citrus orchards, with notes on their morphology and economic importance. Systematic & Applied Acarology, 21 (8), 991 - 1008. http: // dx. doi. org / 10.11158 / saa. 21.8.1","Ali Khan, B. & Anwarullah, M. (1970) A new species of Odontoscirus from Pakistan. Science and Industry (Karachi), 7 (3), 192 - 194.","Willmann, C. (1941) Die Acari der Hohlen der Balkanhalbinsel (Nach dem Material der \" Biospeologica balcanica \"). Studien aus dem Gebiete der allgemeinen Karstforschung, der wissenschaftlichen Hohlenkunde, der Eiszeitforschung und den Nachbargebieten; 14, B, Biologische Serie. Nr. 8.","Thor, S. (1904 b) Norwegische Bdellidae I. Zoologischer Anzeiger, 28, 69 - 79.","Schweizer, J. (1922) Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandlungen der Naturferschenden Gesellschaft in Basel, 33, 23 - 111. http: // dx. doi. org / 10.5962 / bhl. part. 3644","Vitzthum, H. G. (1926) Malayiche Acari. Treubia, 8, 1 - 195.","Walckenaer, C. A. & Gervais, M. P. (1847) Histoire naturelle des insectes apteres IV. 1 - 623 pp.","Andersen, C. H. (1863) Bidrag till kannedomen om Nordiska Acarider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 20, 181 - 193.","Canestrini, G. & Fanzago, F. (1877) Intorno agli acari italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, 5 (4), 1 - 140 + plates II - VII.","Moniez, R. (1890 a) Acariens observes en France (1 ere liste). Revue Biologique du Nord de la France, 3, 24 - 33.","Tragardh, I. (1902 a) Beitrage zur Kentnis der Schwedischen Acariden-fauna. I. Lapplandische Trombidiiden und Oribatiden. Bihang Till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 28, 1 - 26.","Halbert, M. R. I. A. (1915) Clare Island Survey-Acarinida. Section II-terrestrial and marine Acarina. Proceedings of the Royal Irish Academy, 39, 45 - 136.","Atyeo, W. T. (1977) Family Bdellidae. La Faune terrestre de I´lle de Sainte-Helene. Annales du Musee Royal de l´Afrique Centrale, 8, 300 - 305.","Trouessart, E. L. (1888) Note sur les acariens marins recueillis par M. Giard au laboratoire maritime de Wimereux. Comptes rendus hebdomadaires des seances de l'Academie des Sciences, 107, 753 - 755. http: // dx. doi. org / 10.5962 / bhl. part. 4881","Banks, N. (1894) Some new American Acarina. Transactions of the American Entomological Society, 21, 209 - 222.","Ewing, H. E. (1909 b) A systematic and biological study of the Acarina of Illinois. University of Illinois Bulletin, 7 (14), 1 - 120.","Ewing, H. E. (1917) New Acarina, part II-Descriptions of new species and varieties from Iowa, Missouri, Illinois, Indiana and Ohio. Bulletin of the American Museum of Natural History, 37 (2), 149 - 172.","Hartzell, A. (1918) A preliminary list of the Acarina of Iowa. The Proceedings of the Iowa Academy of Sciences, 25, 205 - 212.","Banks, N. (1899) Reports upon the insects, spiders, mites and myriapods collected by Dr. L. Stejneger and Mr. G. E. H. Barrett- Hamilton on the Commander Islands-Arachnida. Reports of the fur-seal investigations, 347 - 351.","Tragardh, I. (1904) Monographie der arktischen Acariden. Fauna Arctica, 41, 1 - 77.","Koch, C. L. (1835 - 1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur deutschen Fauna. Herausgegeben von Dr. Herrich-Schaffer. Regensburg, Friedrich Pustet. 40 parts.","Koch, C. L. (1842) Ubersicht des Arachnidensystems. Familie II: Schabelmilben Bdellides, pp. 73 - 77.","Walckenaer, C. A. & Gervais, M. P. (1844) Histoire naturelle des insectes apteres III. 1 - 476 pp.","Gervais, H. P. (1841) Note sur quelques especes de l´ordre des Acariens. Annales des Sciences Naturelles. Zoologie et Biologie Animale, 15, 5 - 10.","Murray, A. (1877) Economic Entomology. Aptera. Chapman and Hall. London. 433 pp.","Hull, J. E. (1918) Terrestrial Acari of the Tyne province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 75, 13 - 88.","Cambridge, M. A. (1876) On a new order and some new genera of Arachnida from Kerguelen's Land. Proceedings of the Scientific Meetings of the Zoological Society of London, 1, 258 - 264.","Berlese, A. (1888) in Berlese, A. & Balzan, A. (1888) Acari Sud-Americani methodice dispositi, descripti, et iconibus illustrati (con tav.), 20, 171 - 222.","Canestrini, G. (1886) Prospetto dell'acarofauna italiana. 2: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini. pp. 159 - 311, Tav. X - XXII [= 10 - 22]. Padova.","Hull, J. E. (1915) Snout mites: an introduction to the British Bdellidae. The Vasculum, 1, 117 - 123.","Banks, N. (1919) The Acarina collected by the Canadian Arctic Expedition, 1913 - 1918. Reports of the Canadian Arctic Expedition, 3, Part H, 11 - 13.","Banks, N. (1923) Arachnida. In: Trichoptera, Mecoptera, and Arachnida of the Pribilof Islands, Alaska. North American Fauna, 46, 237 - 239.","Willmann, С. (1951) Die hochalpine Milbenfauna der mittleren Hohen Tauern insbesondere der Groβlockner-Gebietes (Acari). Bonner Zoologische Beitrage, 2, 141 - 176.","Butler, G. D. & Usinger, R. L. (1963) Insects and other arthropods from Kure Island. Proceedings of the Hawaiian Entomological Society, 18 (2), 237 - 244.","Lellakova-Duskova, F. (1978) Incidence of mites of the family Bdellidae in moss from a spruce wood in SW-Bohemina. Vestnik Ceskoslovenske spolecnosti zoologicke, 42 (1), 23 - 42.","Garret, L. E. & Haramoto, F. H. (1967) A catalogue of Hawaiian Acarina. Proceedings of the Hawaiian Entomological Society, 19 (3), 381 - 414.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. - Series: Biology, chemistry, 24 (63), 3 - 9.","Atyeo, W. T. (1963 b) New species and records of Bdellidae from Macquarie and the Auckland Islands (Acarina). Pacific Insects, 5 (2), 445 - 450.","Shiba, M. (1978) Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. Nature and Life in Southeast Asia, 7, 83 - 229.","Thor, S. (1931 b) Nordafrikanische Bdellidae und Cunaxidae, von Dr. F. Grandjean (Paris) 1931 gesammelt. Zoologischer Anzeiger, 97 (3 - 4), 62 - 79.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5, 89 - 103.","Kaluz, S., Ferencik, J. & Vrabek, M. (2013) Study sites influenced by natural and human impacts in Tanap and their acarofauna. Entomofauna Carpathica, 25 (1), 1 - 12.","Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia-Giornale di Entomologia, 15, 1 - 240.","Atyeo, W. T. (1960 b) A unique species of Thoribdella Grandjean, 1938, from New Zealand (Acarina: Bdellidae). Records of the Dominion Museum, 3, 289 - 291.","Thor, S. (1902) On the systematic representation of the Acarinen familien Bdellidae Koch, 1842, Grube, 1859, Eupodidae Koch, 1842 and Cunaxidae Sig Thor, 1902. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 52, 159 - 165."]}
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45. Monotrichobdella Baker & Balock
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Monotrichobdella ,Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Taxonomy - Abstract
Monotrichobdella Baker & Balock Monotrichobdella Baker & Balock, 1944: 176. Type-species: Monotrichobdella maxosburni Baker & Balock, 1944 by original designation. 120. Monotrichobdella maxosburni Baker & Balock, 1944: 176; near Tres Cumbres, Mexico-Cuernavaca Highway, Morelos, Mexico, ex lichens. Distribution. Mexico (Morelos, Salazar, Distrito Federal) (Baker & Balock 1944; Atyeo 1960a), Taiwan (Tseng 1978). Redescriptions. Atyeo (1960a), Tseng (1978). Type deposition. USNM.
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46. Polytrichus van Der Schyff, Theron & Ueckermann
- Author
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Polytrichus ,Taxonomy - Abstract
Polytrichus van Der Schyff, Theron & Ueckermann Type-species: Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003 by original designation. 87. Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003: 21; Djebel An-Nabi Shuaib, Yemen. Type deposition. NCA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 23, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["van Der Schyff, J., Theron, P. D. & Ueckermann, E. A. (2003) Polytrichinae, a new subfamily of Bdellidae (Acari: Prostigmata) from the Afrotropical region. African Plant Protection, 9 (1), 19 - 22."]}
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47. Bdella Latreille
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
- Subjects
Bdella ,Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Taxonomy - Abstract
Bdella Latreille Bdella Latreille, 1795: 18; 1810: 133; Berlese, 1893: 42; Oudemans, 1929: 303; Meyer & Ryke, 1959: 373; Atyeo, 1960a: 372, 1963a: 120, 170; Soliman & Zaher, 1975: 79; Chaudhri et al., 1979: 130; van Der Schyff et al., 2005: 222. nec Bdella Savigny, 1822 (Anellida: Hirudinea) junior homonym. Chelifer Geoffroy, 1762: 617 (part.); type species Acarus cancroides Linnaeus, 1762 (Pseudoscorpiones). Scirus Hermann, 1804: 60 (part.) type species Scirus vulgaris Hermann, 1804: 61. Bdellidium Oudemans, 1929: 449; type species Scirus vulgaris Hermann, 1804: 61; synonymy according to Atyeo (1960a). Caenobdella Oudemans, 1937: 1227; type-species Bdella crassipes Koch, 1839 by original designation; synonymy according to Atyeo (1960a). Type-species: Acarus longicornis Linnaeus, 1758 (= La pince Geoffroy), by subsequent designation (Latreille 1795). 1. Bdella aloios van Der Schyff, Theron & Ueckermann, 2005: 222; Limpopo province, South Africa, ex Canthium mundianum Cham. & Schltdl. (Rubiaceae). Remarks. Male unknown. This species was misspelled as B. alois in van Der Schyff et al. 2005, pages 223, 224. Type deposition. NCA. 2. Bdella bakeri Gupta & Paul, 1985: 14; West Bengal, Midnapur Dist., Patharkumkumi, India; ex nest of dove Streptopelia chinensis (Scopoli) (Aves, Columbidae). Remarks. Male unknown. Type deposition. NZSI. 3. Bdella biroi Supino, 1894: 197; Hungary.��� Thor, 1931a: 37. Type deposition. Unknown. 4. Bdella boskopensis van Der Schyff, Theron & Ueckermann, 2005: 232; North West Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 5. Bdella calva Hull, 1915: 122; on soil, England.��� Hull, 1918: 39. Remarks. This species probably belongs to a different genus as indicated by the elongated palp tibiotarsus (Hull 1915, figure 6). Type deposition. Unknown. 6. Bdella captiosa Atyeo, 1963a: 170; Groverly, Queensland, Australia; ex unknown host. Distribution. Australia (Wallace & Mahon 1972), Hawaii (Swift & Goff 1987), Iran (Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010). Remarks. Although Atyeo (1963a: 171) established a male as the holotype, Wallace & Mahon (1972: 546, footnote) analyzed the types and concluded it was actually a female; the male and other immatures were not reported until 1987 by Swift & Goff (1987: 43). Redescriptions. Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. SAM. 7. Bdella cardinalis Banks, 1894: 219; ex wood under leaves, NY, USA; 1904a: 144; 1907: 596; 1908: 5; Ewing, 1909b: 68; Ewing & Webster, 1912: 129; Hartzell, 1918: 206; Hernandes, 2013: 61. Distribution. United States (Alabama, Florida, Texas, Illinois) and Mexico. Type deposition. MCZ. Remarks. This species has been considered as a junior synonym of Bdella longicornis (Linnaeus) by Thor (1931a), and as a junior synonym of Bdella oblonga by Jacot (1938); however, Hernandes (2013) revalidated the status of this species. 8. Bdella carolae van Der Schyff, Theron & Ueckermann, 2005: 235; Limpopo Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 9. Bdella consobrinae van Der Schyff, Theron & Ueckermann, 2005: 234; North West Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 10. Bdella crassipes Koch, 1839: 23; Germany.��� Koch, 1842: 74; Walckenaer & Gervais, 1847: 532; Thor, 1931a: 31; Caenobdella crassipes (Koch), Oudemans, 1937: 1227. Type deposition. Unknown. 11. Bdella distincta Baker & Balock, 1944: 179; Morelos, Mexico, ex lichens.��� Atyeo, 1960a: 381 (lectotype designation). Distribution. United States (Washington D.C., Hawaii, Texas), Puerto Rico (Guayama), Philippines, Indonesia, Japan, China, Indonesia, Thailand (Atyeo 1960a), Taiwan (Tseng 1978), Hawaii (Garret & Haramoto 1967; Swift & Goff 1987), Guadaloupe (Flechtmann & Etienne 2006), Brazil (Lawson-Balagbo et al. 2008). Redescriptions. Atyeo (1960a), Tseng (1978), Swift & Goff (1987). Type deposition. Lectotype female, ex Bambusa parvariabilis, China (detected in Washington DC, USA), at USNM. 12. Bdella distinguenda Berlese, 1905: 157; Bogor (former Buitenzorg, Java). Other names: Scirus distinguendus; Thor, 1931a: 45. Remarks. Suspected synonym of Bdellodes longirostris (Hermann) according to Thor (1931a: 45). Type deposition. Lost (Castagnoli & Pegazano 1985). 13. Bdella dorsata Walckenaer & Gervais, 1844: 157; Paris, France on humid soil with small plants in gardens.��� Berlese, 1893: 43; Thor, 1931a: 37; Oudemans, 1937: 1195. Type deposition. Probably lost. 14. Bdella farabii Paktinat-Saeej & Bagheri, 2015a: 524; Amol, Mazandaran Province, Iran (Paktinat-Saeej et al. 2015a). Type deposition. Holotype at the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran; one paratype at ASI. 15. Bdella grandjeani Thor, 1931b: 65; Northern Africa. Type deposition. Lost. 16. Bdella heliophila Mihelčič, 1958a: 273; Cercedilla, Spain, on moss of pinus forest. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 17. Bdella helvetica Schweizer & Bader, 1963: 232; Val Ftur, National Park, Switzerland, under stone. Original designation: Bdella iconica helvetica Schweizer & Bader. Type deposition. NMB (NP 1931). 18. Bdella horvathi Karpelles, 1888: 275; Bulgaria.��� Thor, 1931a: 37. Type deposition. Unknown. 19. Bdella humida Wallace & Mahon, 1972: 546; Millstream, Fortescue River, West Australia, Australia; ex litter of Acacia sp. (Fabaceae) and Eucalyptus sp. (Myrtacea). Distribution. Australia, Iran (Kamali et al. 2001; Ueckermann et al. 2007). Redescription. Ueckermann et al. (2007). Remarks. Male unknown. Type deposition. Holotype and paratypes at ANIC; paratypes at SAM, BMNH, USNM and OSAL. 20. Bdella iconica Berlese, 1923: 240; on plants, in Florence and Germany���Thor, 1931a: 32. a. Bdella iconica var. veneta Berlese, 1923: 240; Thor, 1931a: 32; Schweizer & Bader, 1963: 231. b. Bdella sardoa Berlese, 1923: 241; B. iconica var. sardoa; Thor, 1931a: 32. Synonymy according to Thor (1931a). Distribution. Italy, Germany and Norway (Thor 1931a), Germany (Willmann 1955), Austria (Willmann 1951), France (Schm��lzer 1956), Spain (Mihelčič 1958a), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), New Zealand (Atyeo 1963a), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Bohemia (Lell��kov��-Du��kova 1978), Ukraine (Kuznetsov & Livshits 1979a), Poland (Michocka 1987), China (Lin et al. 2006), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz et al. 2013). Remarks. Michocka (1987: 39) apparently figured trichobothria on tibiae II, a characteristic only found in the subfamily Odontoscirinae; therefore it is likely that the species reported from Poland might be the result of a misidentification. Redescriptions. Atyeo & Tuxen (1962), Atyeo (1963a), Sosnina et al. (1965), Wallace & Mahon (1972), Lell��kov��-Du��kova (1978), Michocka (1987). Type deposition. CRA. 21. Bdella interrupta Evans, 1952: 668; Mid and North Wales, UK, on rocks above tide marks. Type deposition. Unknown. 22. Bdella karajiensis Ueckermann, Rastegar, Saboori & Ostovan, 2007: 133; Karaj, Iran, ex litter and soil beneath ornamental plant. Type deposition. Holotype at SRIAUT, paratypes at ASI. 23. Bdella khasyana Gupta, 1991: 221; Arunachal Pradesh, India; ex Litsea khasyana Meisn. (Lauraceae). Redescription. Gupta (2002). Remarks. Male unknown. Type deposition. NZSI. 24. Bdella kuznetsovi Maslov & Khaustov, 2013: 52; Arabatsky Nature Reserve, Crimea, Ukraine; ex storm detritus on shore of Sivash Gulf. Type deposition. NBG. 25. Bdella lattakia Soliman & Zaher, 1975: 79; Slinfa, Lattakia, Syria, ex moss. Distribution. Syria, Iran (Kamali et al. 2001; Ueckermann et al. 2007). Redescription. Ueckermann et al. (2007). Remarks. Male unknown. Type deposition. CUE. 26. Bdella longicornis (Linnaeus, 1758): 618; in Europe. Original designation: Acarus longicornis Linnaeus (= Acarus petrarum ruber L., 1746: 349).��� M��ller, 1776: 187; Johnston, 1845: 227; Oudemans, 1926b: 116.; Atyeo, 1962: 344 (neotype designation). a. Chelifer totus ruber Geoffroy, 1762: 618; synonymy according to Thor (1902). b. Acarus rupestris Linnaeus, 1758: 618; Hammer, 1775: 157; synonymy according to Thor (1931a). c. Acarus citri Hasselquist, 1757: 431; Oudemans, 1929: 306; synonymy according to Thor (1931a). d. Acarus ruber Linnaeus, 1746; Fabricius, 1775: 815. e. Acarus velox M��ller, 1776: 187; Oudemans, 1929: 312; 1937: 1201. f. Bdella rubra Lamarck, 1801: 179; von Heyden, 1826: 608; synonym of Bdellodes longirostris according to Berlese (1893). g. Scirus vulgaris Hermann, 1804: 61; Oudemans, 1929: 313; Bdella vulgaris, Koch, 1839: 1851; Dug��s, 1834: 21; Voigts & Oudemans, 1906: 242; B��bler, 1910: 812; Oudemans, 1937: 1206; synonymy according to Thor (1904b). h. Bdella decipiens Thorell, 1871: 701; Tr��g��rdh, 1902a: 7; 1904: 47; 1931: 47; Halbert, 1915: 113; Hull, 1918: 39; Banks, 1919: 11; Hull, 1922: 622; Tr��g��rdh, 1931: 47; synonymy according to Thor (1902: 7); Bdella longicornis var. decipiens Thorell; Tr��g��rdh, 1928: 8. i. Bdella borealis Kramer & Neuman, 1883: 525; Banks, 1899: 348; 1907: 596 synonymy according to Thor (1904b). j. Bdella caeca Berlese, 1905: 15; Bdella longicornis var. caeca; synonymy according to Thor (1931a: 28). k. Bdella vestita Koch, 1835: 23; 1842: 74; Walckenaer & Gervais, 1844: 157; Anders��n, 1863: 184; Canestrini & Fanzago, 1877: 105.��� Scirus vestitus, Murray, 1877: 146; synonymy according to Thor (1904b). l. Bdella anguinesetosa Ewing, 1909b: 72; synonymy according to Atyeo (1960a). m. Bdella tessellata Ewing, 1913: 112; Thor, 1931a: 36; synonymy according to Atyeo (1960a). n. Bdella oblonga Say, 1821: 19; Walckenaer & Gervais, 1847: 348; Gervais, 1849: 32; Banks, 1902a: 543; Oudemans, 1937: 1211; Jacot, 1938: 126; Baker & Balock, 1944: 180; Drummond, 1957: 142; synonymy according to Thor (1931a: 25); Bdellodes oblongula, Oudemans, 1937: 1221. o. Bdella egregia Koch, 1839: 23; synonymy according to Thor (1904b). p. Bdella cruentata Koch, 1839: 10; synonymy according to Thor (1904b). q. Bdella tenuirostris Koch, 1839: 18; synonymy according to Thor (1904b). r. B del la vivida Koch, 1839: 19; synonymy according to Thor (1904b). s. Bdella podurophila White, 1852: 210; synonymy according to Thor (1904b). t. Bdella arenaria Kramer, 1881: 444; synonymy according to Thor (1904b). Distribution. Germany (Voigts & Oudemans 1906), Sweden (Tr��g��rdh 1910), England (Hull 1918), Switzerland (Schweizer 1922; Schweizer & Bader 1963), Canada (Summerhayes & Elton 1928), Norway (Thor 1930a), North Africa (Thor 1931b), Austria (Willmann 1951), France (Schm��lzer 1956), Barro Colorado Islands, Panama Canal Zone, Costa Rica, Cuba, Mexico (Oaxaca, Distrito Federal, M��xico, San Luis Potos��), United States (California, Texas, Utah, Arkansas, Illinois, Tennessee, Florida, Missouri, Michigan, New Hampshire, Vermount), Nova Scotia (Atyeo 1960a), Japan (Ehara 1960; Shiba & Morikawa 1966; Shiba 1969a), Canada (Sinha 1963), Taiwan (Tseng 1978), United States (Lehman 1982), Poland (Michocka 1987), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), Japan (Nakamura et al. 2006), Iran (Kamali et al. 2001, Ueckermann et al. 2007), Crimea (Bednarskaya 2011). Redescriptions. Walckenaer & Gervais (1844), Thor (1926, 1930a, 1931a), Tr��g��rdh (1928), Oudemans (1929, 1937), Willmann (1956), Atyeo (1960a, 1962), Ehara (1961), Schweizer & Bader (1963), Shiba (1969a), Wallace & Mahon (1972), Tseng (1978), Michocka (1987), Ueckermann et al. (2007). Remarks. Bdella longicornis is the type species of the genus Bdella, and is among the first members of the family ever described. As a consequence, it has appeared under various names in the 18th and 19th century literature. Type deposition. Neotype of B. longicornis at USNM; type of B. caeca at CRA; type of B. cardinalis at MCZ; type of B. anguinesetosa at USNM. 27. Bdella longipalpus Mihelčič, 1958b: 41; Sierra Nevada, Spain. Type deposition. Unknown. 28. Bdella longistriata Atyeo, 1960a: 380; Llera, Tamaulipas, Mexico, ex pineapple epiphyte. Other records: Mexico (Llera, Tamaulipas; Ciudad del Maiz, San Luis Potos��; Antiguo Morelos, Tamaulipas), United States (Texas) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM. 29. Bdella malaccensis Shiba, 1978: 99; Malacca, Malaysia; under stones of intertidal zone. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 30. Bdella malawiensis van Der Schyff, Theron & Ueckermann, 2005: 224; Malawi, Bangula, Mozambique; ex Trichilia dregeana Sond. (Meliaceae). Remarks. Male unknown. Type deposition. NCA. 31. Bdella maldahanensis Gupta, 1992: 121; West Bengal, India, ex Mangifera indica Linnaeus (Anacardiaceae). Redescription. Gupta (2002). Type deposition. NZSI. 32. Bdella muscorum Ewing, 1909a: 124; Muncie, Illinois, USA ex moss.��� Thor, 1931a: 38. a. Bdella recens Ewing, 1934: 57 (= Bdella lata Ewing, 1909b: 69); Drummond, 1957: 142; synonymy according to Atyeo (1960a). b. Bdella subnigra Ewing, 1909b: 73; synonymy according to Atyeo (1960a). c. Bdella muscorum var. minnesotensis Ewing, 1913: 113; Thor, 1931a: 38; synonymy according to Atyeo (1960a). Distribution. United States (California, Colorado, New Mexico, Kansas, Arkansas, Tennessee, Michigan, Minnesota, Illinois, Maryland), Alaska (Point Barrow, Chandler Lake Region, Umiaat, District of Mackenzie), Germany (Ost-Holstein), Czeck Republic and Iceland (Atyeo 1960a), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), Bulgaria (Sosnina et al. 1965), Japan (Shiba & Morikawa 1966), Bohemia (Lell��kov��- Du��kova 1978), Taiwan (Tseng 1978), Ukraine (Kuznetsov & Livshits 1979a), Pakistan (Chaudhri et al. 1979), Canada (Danks 1980), United States (Lehman 1982), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), Hungary (Ripka et al. 2005), Japan (Nakamura et al. 2006), Iran (Ueckermann et al. 2007), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz 2008). Redescriptions. Atyeo (1960a), Atyeo & Tuxen (1962), Shiba & Morikawa (1966), Sosnina et al. (1965), Wallace & Mahon (1972), Lell��kov��-Du��kova (1978), Tseng (1978). Type deposition. USNM. 33. Bdella neograndjeani Meyer & Ryke, 1959: 373; Eastern Cape Province, South Africa; ex unidentified shrub. Redescription. van Der Schyff et al. (2005). Type deposition. NCA. 34. Bdella nihoaensis Swift & Goff, 1987: 29; Nihoa Island, Hawaii, ex litter. Remarks. Female unknown. Type deposition. BPBM. 35. Bdella nylsvleyensis van Der Schyff, Theron & Ueckermann, 2005: 226; Limpopo Province, South Africa, ex soil and debris. Type deposition. NCA. 36. Bdella obesa Oudemans, 1937: 1211; unknown locality, Germany. Type deposition. Unknown. 37. Bdella pinicola Cooreman, 1943: 7; on pines, near Brussels, Belgium. Type deposition. IRSN. 38. Bdella piggotti Evans, 1953: 272; Kilimanjaro, Tanzania. Type deposition. Unknown. 39. Bdella pulchella Berlese, 1923: 241; unknown plant and in soil, Italy (Florence and Forli).��� Thor, 1931a: 33. Distribution. Norway and Italy (Thor 1931a), Spain (Mihelčič 1958a). Type deposition. Lost (Castagnoli & Pegazzano 1985). 40. Bdella radhikae Sadanandan et al., 2009: 3; ex Cocos nucifera, Kerala, India. Type deposition. Division of Acarology, Department of Zoology, University of Calicut, Kerala, India. 41. Bdella robusta Banks, 1894: 220; Sea Cliff, New York, USA, on ground.��� Banks, 1907: 596. Other name: Scirus robustus, Thor, 1931a: 45. Distribution. United States (Sea Cliff, New York) (Thor 1931a). Type deposition. Unknown. 42. Bdella semiscutata Thor, 1930a: 92; Svalbard, Norway.��� Thor, 1931a: 29; Willmann, 1939c: 431; 1956: 243. Distribution. Svalbard Island (Norway), Norway, Germany, (Thor 1931a), North Africa (Thor 1931b), Austria (Willmann 1951), Spain (Mihelčič 1958b), Georgia (Gomelauri 1963b), Switzerland (Schweizer &, Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. 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Journal of Ecology, 16 (2), 201 - 212, 238 - 252. http: // dx. doi. org / 10.2307 / 2255796", "Thor, S. (1930 a) Beitrage zur kenntnis der invertebraten fauna von Svalbard. Skrifter Svalbard og Ishavet, 27, 1 - 156.", "Ehara, S. (1960) Comparative studies on the internal anatomy of three Japanese Trombidiform acarinids. Journal of the Faculty of Science. Hokkaido University, Series VI, Zoology, 14 (3), 410 - 434.", "Shiba, M. & Morikawa, K. (1966) Prostigmatic mites from Japan. II Bdellidae I (Bdellinae, Cytinae, Spinibdellinae). Reports of Research Matsuyama Shinonome Junior College, 3, 17 - 37.", "Shiba, M. (1969 a) Taxonomic investigations on free-living mites in the subalpine forest on Shiga Heights IBP area II. Prostigmata. Bulletin of Natural Sciences of Tokyo, 12 (1), 65 - 115.", "Sinha, R. N. (1963) Stored product acarology in Canada. In: Advances in Acarology, Naegele, J. A. (Ed.), vol. 1, Comstock Pub. Assoc., 70 - 88.", "Lehman, R. D. (1982) Mites (Acari) of Pennsylvania conifers. Transactions of the America Entomological Society, 180, 181 - 286.", "Lee, W. - K., Lim, J. - W. & Lee, S. - Y. (1997) A taxonomic study on the family Bdellidae (Bdelloidea: Prostigmata) in Korea. Korean Journal of Soil Zoology, 2 (2), 65 - 75 [in Korean with English abstract]", "Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Vol. 9. Fujian Science and Technology Press, Fuzhou, 109 - 121.", "Nakamura, Y., Ishikawa, K., Shiba, M., Fujikawa, T., Ono, H., Tamura, H. & Morikawa, K. (2006) Soil animals of the 88 Buddhist temples in Shikoku Island. Memoirs of the Faculty of Argriculture, Ehime University, 51, 25 - 48.", "Thor, S. (1926) Die Acarina der Kamtschatka Expedition 1908 - 1909. Annuaire du Musee Zoologique de l'Academie des Sciences de l'URSS, 27, 131 - 141.", "Willmann, C. (1956) Milben aus dem Naturschutzgebiet auf dem Spiegelitzer (Glatzer) Schneeberg. Ceskoslovenska Parazitologie, 3, 211 - 273", "Ehara, S. (1961) Some snout mites from Japan (Acarina: Bdellidae). Publications of the Seto Marine Biological Laboratory, 9 (2), 247 - 263.", "Mihelcic, F. (1958 b) Zoologisch-systematische Ergebnisse der Studienreise von H. Janetschek und W. Steiner in die spanische Sierra Nevada 1954. IX. Milben (Acarina). Sitzungsberichte der Mathematisch-naturwissenschaftliche Klasse, Abteilung I, 167, 35 - 57.", "Shiba, M. (1978) Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. Nature and Life in Southeast Asia, 7, 83 - 229.", "Gupta, S. K. (1992) Arachnida: plant mites (Acari). State Fauna Series 3, Fauna of West Bengal, part III, 61 - 211.", "Ewing, H. E. (1909 a) Three new species of the genus Bdella (mites). The Canadian Entomologist, 41, 122 - 126. http: // dx. doi. org / 10.4039 / Ent 41122 - 4", "Ewing, H, E. (1934) Arachnida. Order Acarina. In: Carpenter, F. M., Folsom, J. W., Essig, E. O., Kinsey, A. C., Brues, C. T
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48. Bdellidae
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Taxonomy - Abstract
Fossil BDELLIDAE 279. Bdella vetusta Ewing, 1934: 58; Manitoban amber (Cretaceous, ~ 76–79 Ma), Cedar Lake, Manitoba, Canada. Type deposition. ROME. 280. Bdella lata Koch & Berendt, 1854: 872 —not Bdella lata Ewing (1909b); Baltic amber (~44 Ma). 281. Bdella bicincta Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 282. Bdella bombycina Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 283. Bdella obconica Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 284. Bdellidae sp. (Aoki 1974 apud Dunlop et al. 2012).
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49. Cyta von Heyden
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Cyta ,Taxonomy - Abstract
Cyta von Heyden Cyta von Heyden, 1826: 608; Thor, 1930a: 89; 1931a: 16; Oudemans, 1937: 1232; Meyer & Ryke, 1959: 377; Atyeo, 1960a: 416; 1963a: 121, 177; Soliman & Zaher 1975: 74; Den Heyer, 1981: 32; Michocka, 1987: 72. Amonia Koch, 1836: 7; type species Amonia cruciata Koch, 1836: 7; Berlese, 1893: 43. Ammonia, Koch, 1842: 75. Cytobdella Mihelčič, 1958a: 271; type species Bdella (Cytobdella) cytoides Mihelčič, 1958a by original designation; synonymy according to Atyeo (1963a: 121). Troglobdella Oudemans, 1937: 1228; type species Troglobdella obisium (Gervais, 1841), Scirus obisium Gervais, 1841 by original designation; synonymy according to Atyeo (1960a: 417). Rigibdella Tseng, 1978; type-species Rigibdella ignea Tseng, 1978: 48 by original designation; new synonym. Type-species: Scirus latirostris Hermann, 1804 by original designation. 65. Cyta americana (Banks, 1902b): 171; Washington D.C., USA. Original designation: Ammonia americana Banks. ��� Cyta americana; Banks, 1907: 596; 1915: 25; Hernandes et al., 2011: 808. Remarks. Although Thor (1931a) treated this species as a synonym of C. latirostris, they have different chaetotaxy on the palpfemur (6���7 in the latter species and 4 in C. americana) (Hernandes 2013); also, the proximal setae of the chelicerae is approximately three times the length of the distal setae (2x in C. latirostris). Type deposition. MCZ. 66. Cyta brevipalpa Ewing, 1909b: 73; Ewing & Webster, 1912: 129; under bark on soft maple tree, Illinois, USA. Type deposition. USNM. 67. Cyta coerulipes (Dug��s, 1834): 21; Europe, unknown locality.��� Thor, 1931a: 20; Willmann, 1939c: 431; 1956: 241; 227; Den Heyer, 1981: 32 (ssp.��� quadrisetosus). Original designation: Bdella coerulipes Dug��s. ��� Walckenaer & Gervais, 1844: 156. a. Amonia chlorophana Koch, 1836: 8; Amonia chloropus, Koch, 1842: 76; Bdella chloropus (Koch), Berlese, 1883: 214; synonymy according to Berlese (1891, 1893). b. Amonia leucocephala Koch, 1839: 1; Walckenaer & Gervais, 1847: 531; Oudemans, 1937: 1238; synonymy according to Thor (1931a: 20). c. Bdella crassirostris Kramer, 1881: 442; synonymy according to Thor (1931a). d. Ammonia coerulipes; Berlese, 1893: 44. e. Cyta lutea George, 1912: 236; synonymy according to Thor (1931a). f. Bdella robustirostris Ewing, 1913: 112; synonymy according to Thor (1931a). g. Cyta coeruleipes; Hull, 1918: 37. Distribution. Morocco (Thor 1931b), Austria (Willmann 1951), Spain (Mihelčič 1958b), Panama, Haiti, Cuba, Mexico (San Luis Potos��, Guerrero, Tamaulipas, Hidalgo), United States (California, Colorado, Texas), Alaska (Umiat) and Sudan Bor (Atyeo 1960a), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Egypt (Soliman 1975), Syria (Lattakia) (Soliman & Zaher 1975), Ukraine (Kuznetsov & Livshits 1979a), Afrotropical (Natal) (Den Heyer 1981), Poland (Michocka 1987), Brazil (Hernandes et al. 2011), Iran (Beyzavi et al. 2011). Redescriptions. Atyeo (1960a), Michocka (1987). Remarks. There is confusion between the names Amonia chlorophana Koch and A. chloropus Koch���both names are present in Koch (1836) in the contents page and in text, respectively. Type deposition. Unknown. 68. Cyta cytoides (Mihelčič, 1958a): 271 comb. nov.; in a damp cave in brown clay, Guadarrama Mountains, Cercedilla, Spain. Original designation: Bdella (Cytobdella) cytoides Mihelčič. Remarks. Mihelčič (1958a) illustrates a massive and stout chelicera for this species, a key diagnostic feature for the genus Cyta; therefore this species is herein transferred to the latter genus. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 69. Cyta grandjeani Gomelauri, 1963a: 169; Tbilisi, Georgia, ex soil, lichens, grass, oak and juniper tree. Kuznetsov & Livshits (1979a: 56). Remarks. Cyta veneta grandjeani Lombardini, 1964: 106 is a junior homonym; although the date printed in Lombardini is 1962, the issue appeared only in 1964. Type deposition. Unknown. 70. Cyta flava Mihelčič, 1958a: 270; between Ciempozuelos and Chinch��n, Spain, on soil. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 71. Cyta ignea (Tseng, 1978): 48 comb. nov.; Taipu, Chiayi Hsien, Taiwan, on litter. Original designation: Rigibdella ignea Tseng. Remarks. The rationale for moving this species to Cyta was given in the discussion before the taxonomic section. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 72. Cyta kauaiensis Swift & Goff, 1987: 38; Kauai I, Hawaii, ex litter in Metrosideros sp. (Myrtaceae) tree hole. Remarks. Female unknown. Type deposition. BPBM. 73. Cyta latirostris (Hermann, 1804): 62; France, ex moss. Original designation: Scirus latirostris Hermann. ��� Oudemans, 1929: 314. Cyta latirostris, Thor, 1902: 160; Oudemans, 1902: 54; Thor, 1904b: 72; B��bler, 1910: 812. Other names: Leptus latirostris, Lamarck, 1838: 63; Ammonia latirostris, Koch, 1839: 3; Koch, 1842: 76; Walckenaer & Gervais, 1847: 531; Berlese, 1891: 4; Bdella latirostris, Walckenaer & Gervais, 1844: 157; 1847: 531; Canestrini & Fanzago, 1877: 107; Berlese, 1883: 214. a. Ammonia cruciata Koch, 1836: 7; Anders��n, 1863: 185; Berlese, 1893: 44; Oudemans, 1937: 1235; Bdella cruciata; Walckenaer & Gervais, 1844: 156 synonymy according to Berlese (1893). b. Ammonia leucocephala Koch, 1839: 23; 1842: 76; Bdella leucocephala (Koch), Canestrini & Fanzago, 1877: 106; synonymy according to Berlese (1893). c. Ammonia megacephala Koch, 1839: 23; 1842: 76; Walckenaer & Gervais, 1847: 531; Berlese, 1893: 44; Oudemans, 1937: 1239; synonymy according to Berlese (1893). d. Bdella phoenicea Koch, 1839: 7; 1842: 74; Walckenaer & Gervais, 1847: 531; Canestrini & Fanzago, 1877: 103; Thor, 1931a: 36; Oudemans, 1937: 1213; synonym of Bdella vulgaris according to Berlese (1893). f. Bdella cruentata Koch, 1839: 10; 1842: 74; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 185; Canestrini & Fanzago, 1877: 102; Thor, 1931a: 37; Oudemans, 1937: 1192; synonym of Bdella vulgaris according to Berlese (1893). g. Bdella crassirostris Kramer, 1881: 442; synonymy according to Berlese (1891, 1893). h. Bdella dispar Koch, 1839: 23; 1842: 75; Walckenaer & Gervais, 1847: 532; Canestrini & Fanzago, 1877: 101; Thor, 1931a: 31; Oudemans, 1937: 1194; Willmann, 1951a: 148; 1952: 166; Michocka, 1987: 24; synonymy according to Thor (1931a). h. Bdella amarantina Koch, 1839: 17; 1842: 75; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 185; Berlese, 1893: 42; Oudemans, 1937: 1191; synonymy according to Thor (1931a: 31). i. Bdella tenuirostris Koch, 1839: 18; 1842: 74; Walckenaer & Gervais, 1847: 532; synonym of Bdella vulgaris according to Berlese (1893). j. Bdella vivida Koch, 1839: 19; 1842: 75; Walckenaer & Gervais, 1847: 532; Berlese, 1893: 43; Thor, 1931a: 37; Oudemans, 1937: 1216; synonym of Bdella vulgaris according to Berlese (1893). k. Bdella vulgaris (Hermann 1804: 61), Koch, 1839: 8; 1842: 74; Walckenaer & Gervais, 1847: 531; Berlese, 1888: 187; Hull, 1918: 38. l. Bdella robustirostris Ewing, 1913: 112; synonymy according to Thor (1931a). m. Cyta novangliae Jacot, 1939: 326; synonymy according to Atyeo (1960a). n. Cyta phaseoli Meyer & Ryke, 1959: 377; Den Heyer, 1981: 32; synonymy according to Wallace & Mahon (1972: 571). o. Amonia latirostris var. fusca Canestrini, 1896: 92; Cyta latirostris var. fusca Thor, 1931a: 19. p. Bdella brevirostris molissima Koch, 1879: 132, Cyta brevirostris; Hull, 1922: 622; Banks, 1923: 237; Ammonia brevirostris; Tr��g��rdh, 1900: 15; 1902a: 8; Cyta latirostris var. brevirostris; Tr��g��rdh, 1904: 49; 1910: 480; 1928: 7; synonymy according to Thor (1904b). q. Bdella arenaria Kramer, 1881: 444; Berlese 1893: 42; Karpelles, 1894: 425; synonymy of Bdella vulgaris according to Berlese (1893). Distribution. Paraguay, Brazil (Berlese 1888), Italy (Canestrini 1886), Sweden (Tr��g��rdh 1910), Germany (Voigts & Oudemans 1906; Willmann 1952), Ireland (Halbert 1915), England (Hull 1918), Switzerland (Schweizer 1922), Canada (Summerhayes & Elton 1928), Germany, Norway (Thor 1930a, 1931a), Northern Africa (Thor 1931b), Australia (Womersley 1933a), Madeira Island (Willman 1939a), Austria (Willmann 1951), Germany (Wangerooge Island) (Willmann 1952), Germany (Willmann 1956), France (Schm��lzer 1956), United States (Maryland) (Drummond 1957), Spain (Mihelčič 1958a), Georgia (Gomelauri 1961), South Africa (Meyer & Ryke 1959; Halliday 2005), Cuba, Jamaica, Mexico (Puebla, M��xico, San Luis Potos��), United States (California, Utah, Idaho, Texas, Arkansas, Kansas, Nebraska, North Dakota, Michigan, Alabama, Tennessee, West Virginia, Maryland, Connecticut), Alaska (Point Barrow), Iceland, Italy, Germany and Australia (Atyeo 1960a), Iceland (Atyeo & Tuxen 1962), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Japan (Shiba & Morikawa 1966; Shiba 1969a; Nakamura et al. 2006), New Zealand, Australia (Victoria, Tasmania, Western Australia) (Atyeo 1963a; Wallace & Mahon 1972), Egypt (Soliman 1975; Zaher 1986), Syria (Lattakia) (Soliman & Zaher 1975), Bohemia (Lell��kov��-Du��kova 1978), Ukraine (Kuznetsov & Livshits 1979a), Afrotropical (Natal, Cape Province, Pietersburg Dist.) (Den Heyer 1981), United States (Lehman 1982), Hawaii (Swift & Goff 1987), Poland (Michocka 1987), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), South Africa (Halliday 2005), Hungary (Ripka et al. 2005), Iran (Ostovan & Kamali 1995; Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz 2008). Redescriptions. Atyeo (1960a, 1963a), Atyeo & Tuxen (1962), Sosnina et al. (1965), Shiba (1969a), Lell��kov��-Du��kova (1978), Michocka (1987). Although Berlese (1893: 44) has considered Ammonia leucocephala Koch as a synonym of Cyta latirostris, Koch (1839) illustrated three leg trichobothria (on tibia I, tibia IV and tarsus III), while C. latirostris only has a trichobothrium on tibia IV. Type deposition. Unknown (probably lost). Remarks. Species with world-wide distribution; this species bears trichobothria only on tibia IV; Wallace & Mahon (1972: 579) reports an undescribed species of Cyta with no leg trichobothria at all. 74. Cyta leiliae Eghbalian, Khanjani & Ueckermann, 2014: 568; ex soil and litter under Quercus brantii Lindl (Fagaceae), Kurdistan province, Iran. Type deposition. Holotype and paratypes at the Acarology Laboratory, University of Abu-Ali Sina, Hamedan, Iran, one paratype at NCA. 75. Cyta kurdistanicus Eghbalian, Khanjani & Ueckermann, 2014: 572; ex soil and litter under Amygdalus lycioides Spach (Rosaceae), Kurdistan province, Iran. Type deposition. Holotype and paratypes at the Acarology Laboratory, University of Abu-Ali Sina, Hamedan, Iran, one paratype at NCA. 76. Cyta longiseta Wallace & Mahon, 1972: 575; Mt. Claremont, Perth, Australia, on Banksia sp. (Proteaceae) scrub. Distribution. Iran (Abbaszadeh et al. 2010). Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BMNH, BPBM, OSAL and USNM. 77. Cyta magdalenae Den Heyer, 1981: 34; Kleinmond, Cape Province, South Africa, in soil under dune vegetation. Type deposition. NCA, IZRPU, ZUNIN. 78. Cyta murrayi Den Heyer, 1981: 37; ex litter of Dombeya rotundifolia (Hochst.) (Malvaceae), UNIN, South Africa. Distribution. Afrotropical (Angola, South Africa, Zimbabwe) (Den Heyer 1981). Type deposition. NCA, ZUNIN, ANIC. 79. Cyta reticulata Soliman & Zaher, 1975: 74; Slinfa, Lattakia, Syria, ex moss. Remarks. Male unknown. Type deposition. CUE. 80. Cyta spuria Atyeo, 1960a: 421; 10 miles east of Xilitla, San Luis Potos��, Mexico, ex bamboo; Distribution. Mexico, USA (Texas) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 81. Cyta troglodyta Hernandes, 2011: 801; ex feces of Kerodon rupestris Wied (Mammalia, Rodentia), Gruta do Janel��o cave, Minas Gerais, Brazil (Hernandes et al. 2011). Type deposition. DZSJRP. 82. Cyta veneta (Lombardini, 1960): 241; Jesolo, Venice, Italy. Original designation: Neomolgus venetus Lombardini. Cyta veneta; Lombardini, 1964: 105. Type deposition. CRA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 19-22, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["von Heyden, C. (1826) Versuch einer systematischen eintheiling der Acariden. In: Isis von Oken, 18 (6), 608 - 613.","Thor, S. (1930 a) Beitrage zur kenntnis der invertebraten fauna von Svalbard. Skrifter Svalbard og Ishavet, 27, 1 - 156.","Oudemans, A. C. (1937) Kritisch historisch overzicht der Acarologie. Derede gedeelte, 1805 - 1850, Band C, Tarsonemini Can. & Fanz. 1877, Stomatostigmata Oudms. 1906, Eleutherengona Oudms. 1909. 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H., Khanjani, M., Safaralizadeh, M. H. & Ueckermann, E. A. (2014) Two new species of Cyta (Acari: Prostigmata: Bdellidae) from Western Iran. Zootaxa, 3847 (4), 567 - 575. http: // dx. doi. org / 10.11646 / zootaxa. 3847.4.6","Lombardini, G. (1960) Acari nuovi XXXVIII. Redia-Giornale di Entomologia, 45, 241 - 243."]}
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50. Spinibdella Thor
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Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A., and Bauchan, Gary R.
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Arthropoda ,Arachnida ,Spinibdella ,Prostigmata ,Animalia ,Bdellidae ,Biodiversity ,Taxonomy - Abstract
Spinibdella Thor Spinibdella Thor, 1930b: 22; 1931a: 39; Atyeo, 1960a: 424; Soliman & Zaher, 1975: 80; Tseng, 1978: 38; Chaudhri et al., 1979: 133; Michocka, 1987: 82. Type-species: Spinibdella reducta Thor, 1930b: 23 by original designation. 121. Spinibdella ampulla Wallace & Mahon, 1972: 568; Millstream, Western Australia, ex green herbaceous, couch grass, Eucalyptus sp. litter. Remarks. Male unknown. Type deposition. ANIC. 122. Spinibdella ankylotricha Omukunda, Theron & Ueckermann, 2012: 9; Limpopo Province, South Africa. Type deposition. NCA. 123. Spinibdella antarctica (Tr��g��rdh, 1907): 24; South Georgia, Grytviken Peninsula, Antarctica, under rocks.��� Wallace, 1970: 107. Original designation: Bdella antarctica Tr��g��rdh; Thor, 1931a: 33. Redescription. Wallace (1970). Type deposition. BPBM; USNM, BMNH, ANIC. 124. Spinibdella arenosa Willmann, 1939b: 532; Germany (Wangerooge Island) (Willmann, 1952: 165). Type deposition. ZSM. 125. Spinibdella atyeoi Gupta & Paul, 1985: 14; West Bengal, Midnapur Dist., Patharkumkumi, India, ex nest of Prinia inornata (Sykes 1832) (Aves: Cysticolidae). Remarks. Male unknown. Type deposition. NZSI. 126. Spinibdella bifurcata Atyeo, 1960a: 430; 10 miles west of Tuxtla Gutierrez, Chiapas, Mexico, under rock.��� Soliman, 1975: 48. Distribution. Mexico (Chiapas, Oaxaca, Puebla, Michoac��n, United States (Texas) (Atyeo 1960a), Egypt (Giza) (Soliman 1975; Zaher 1986), Malaysia (Shiba 1978), China (Lin et al. 2006). Redescription. Shiba (1978). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 127. Spinibdella bioculata Swift & Goff, 1987: 39; Kahoolawe Island, Hawaii, ex Kiawe duff and grasses. Remarks. Male unknown. Type deposition. BPBM. 128. Spinibdella californica McGregor, 1956: 12; California, USA, ex lemon. Type deposition. Unknown. 129. Spinibdella corticis (Ewing, 1909a): 122; Urbana, Illinois, USA, under bark of cottonwood tree.��� Atyeo, 1960a: 426. Original designation: Bdella corticis Ewing; Thor, 1931a: 36. Other name: Spinibdella cortis [sic]; Rack, 1961: 185. Distribution. United States (Illinois, Texas, Utah, Nebraska), Mexico (Oaxaca), Guatemala (Atyeo 1960a), Japan (Shiba & Morikawa 1966), Australia (Wallace & Mahon 1972). Redescriptions. Atyeo (1960a), Wallace & Mahon (1972). Type deposition. USNM. 130. Spinibdella cronini (Baker & Balock, 1944): 178; Planada, California, USA, on lichens from fig tree.��� Atyeo, 1960a: 432. Original designation: Bdella cronini Baker & Balock. Distribution. United States (California, Texas, Utah, Colorado, Washington, Alabama, Maryland), Mexico (Tamaulipas, Guerrero, Nevo Le��n, San Luis Potos��) (Atyeo 1960a), Australia (Atyeo 1963a; Wallace & Mahon 1972), Bulgaria (Sosnina et al. 1965), Egypt (Soliman 1975), Syria (Lattakia) (Soliman & Zaher 1975), Ukraine (Kuznetsov & Livshits 1979a), United States (Lehman 1982), Hawaii (Swift & Goff 1987), Hungary (Ripka et al. 2005), China (Li et al. 1992; Li & Fan 2007), Iran (Ostovan & Kamali 1995; Kamali et al. 2001, Jalaeian et al. 2005; Ueckermann et al. 2007; Abbaszadeh et al. 2010; Daneshnia & Akrami 2013; Majidi & Akrami 2013; Masoudian & Khanjani 2013), Brazil (Pinto-da-Rocha 1995; Hernandes et al. 2011), Slovakia (Kaluz 2008). Remarks. this species was found in bat guano in USA (Webster & Whitaker 2005). Redescriptions. Atyeo (1960a, 1963a), Sosnina et al. (1965), Wallace & Mahon (1972), Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. USNM. 131. Spinibdella denheyeri Hernandes, Daud & Feres, 2008: 265; ex Coffea arabica leaves (Linnaeus, Rubiaceae) Atibaia, S��o Paulo, Brazil. Type deposition. DZSJRP. 132. Spinibdella depressa (Ewing, 1909a): 125; Arcola, Illinois, USA, under bark.��� Atyeo, 1960a: 428. Original designation: Bdella depressa Ewing; Thor, 1931a: 38. a. Bdella virgata Ewing, 1909b: 70; Hartzell, 1918: 206; Baker & Balock, 1944: 179 synonymy by Atyeo (1960a). b. Bdella chapultepecensis Baker & Balock, 1944: 177 synonymy by Atyeo (1960a). c. Bdella riolermensis Baker & Balock, 1944: 178 synonymy by Atyeo (1960a). Distribution. United States (Maryland) (Drummond 1957), United States (Illinois, Texas, Maryland, Arkansas, Kansas, New Jersey, Connecticut), Mexico (M��xico, Distrito Federal, San Luis Potos��, Morelos) (Atyeo 1960a), Australia (Atyeo 1963a; Wallace & Mahon 1972), Pakistan (Chaudhri et al. 1979), United States (Lehman 1982), Hawaii (Swift & Goff 1987), China (Fujian) (Lin & Zhang 2000), Iran (Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010). Remarks. this species has the posterior eye wanting, with circular striation where that eye should normally be. Redescriptions. Baker & Balock (1944), Atyeo (1960a, 1963a), Wallace & Mahon (1972), Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. USNM. 133. Spinibdella dusta Shiba, 1969b: 150; tatami, Kuwabara-ch��, Matsuyama, Japan, ex tatami. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 134. Spinibdella gibberabdomen (Thor, 1931b): 68; Tangier, Morocco; Paktinat-Saeej et al. 2015: 695. Original designation: Bdella gibberabdomen Thor. Remarks. The author mentions only two ventral setae on the hypostome, and illustrates a truncate palptarsus, which is why this species was transferred to the genus Spinibdella. Type deposition. Lost. 135. Spinibdella howarthi Swift & Goff, 1987: 40; Mauna Kea Summit Cone, Hawaii, under stone. Remarks. Male unknown; species known only from the holotype. Type deposition. BPBM. 136. Spinibdella iberica Gomelauri, 1961: 68; close to the Turtle lake near Tbilisi, Georgia, ex lichens. Type deposition. Unknown. Remarks. The choice of the epithet by the author is a mystery, since the type locality was not in the Iberian Peninsula. 137. Spinibdella lignicola (Canestrini, 1886): 184; Italy and Egypt.��� Tseng, 1978: 47. Original designation: Bdella lignicola Canestrini. Distribution. Italy (Canestrini 1886; Thor 1931a), England (Hull 1918), Egypt (Tr��g��rdh 1905; Thor 1931a; Abdel-Shaheed et al. 1971), Spain (Mihelčič 1958b), Switzerland (Schweizer & Bader 1963), Taiwan (Tseng 1978), China (Sichuan) (Li et al. 1992), Czeck Republic (Stejskal & Hubert 2008). Redescription. Tseng (1978). Type deposition. CRA. 138. Spinibdella longistriata Tseng, 1978: 42; Shandimann, Pingtung Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 139. Spinibdella mali Jorgensen, 1967: 98; Spring Lake, Utah Co, Utah, USA, ex bark of apple tree. Remarks. Male unknown. Type deposition. Unknown. 140. Spinibdella namibiensis Omukunda, Theron & Ueckermann, 2012: 15; Namibia and South Africa. Type deposition. NCA. 141. Spinibdella novemsetosa Tseng, 1978: 42; Tainan city, Taiwan, on shallot (Alliaceae). Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 142. Spinibdella ornata Atyeo, 1960a: 434; Bear Lake, Rock Mountain National Park, Colorado, USA, ex moss and litter. Distribution. California, (Atyeo 1960a), Wisconsin (Oatman 1963). Remarks. Suspected synonym of S. thori, according to Atyeo (1963a: 174). Type deposition. Holotype and paratypes at SEMC, paratypes at CSUC, USNM. 143. Spinibdella polyattenuata Omukunda, Theron & Ueckermann, 2012: 6; Eastern Cape Province, South Africa. Type deposition. NCA. 144. Spinibdella pongolensis Omukunda, Theron & Ueckermann, 2012: 12; Kwazulu, Natal, South Africa. Type deposition. NCA. 145. Spinibdella quinqueoculata Thor, 1931b: 70; Tangier, Morocco, ex moss. Type deposition. Lost. 146. Spinibdella rapida Kuznetsov & Livshits, 1979b: 608; rocky cliffs in the vicinity of Alupka town, Crimea, Ukraine, ex moss.��� Bednarskaya, 2011: 5. Type deposition. NBG. 147. Spinibdella reducta Thor, 1930b: 23; Norway, in coniferous forest litter.��� Thor, 1931a: 39. Distribution. Norway (Thor 1931a), Poland (Michocka 1987). Redescriptions. Thor (1931a), Michocka (1987). Type deposition. Lost. 148. Spinibdella smileyi Tseng, 1978: 39; Taipei, Chiayi Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 149. Spinibdella subrufa Rack, 1961: 183; Germany. Type deposition. ZMUH. 150. Spinibdella tabarii Paktinat-Saeej & Bagheri, 2015b: 696; Amol city, Mazandaran Province, Iran; also citrus, Noor city, Mazandaran Province, Iran. Type deposition. Holotype and paratypes at the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran; paratypes at the Acarological Collection, Jalal Afshar Zoological Museum, Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran; and also at ASI. 151. Spinibdella tadjikistanica Kuznetsov, 1984: 774; Kondara Canyon, Tadjikistan, ex hawthorn (Crataegus sp., Rosaceae) and grape. Type deposition. NBG. 152. Spinibdella tenella (Banks, 1896): 75; Sea Cliffs, Long Island, NY, USA. Original designation: Bdella tenella Banks; Banks, 1904c: 16; 1907: 596; Thor, 1931a: 31; Spinibdella tenella; Hernandes, 2013: 64. Type deposition. MCZ. 153. Spinibdella tenuirostris (Ewing, 1917): 149; Xenia, Ohio, USA, under stones.��� Atyeo, 1960a: 424. Original designation: Bdella tenuirostris Ewing; Berlese, 1893: 43. a. Spinibdella wilsoni Jacot, 1938: 129; synonymy according to Atyeo (1960a: 424). Distribution. Germany (Thor 1931a), United States (Ohio, Florida, Arkansas, Kansas, North Carolina, Vermont, Michigan, California) (Atyeo 1960a), Japan (Shiba & Morikawa 1966), Australia (Atyeo 1963a; Wallace & Mahon 1972), Russia (Wainstein et al. 1978, Ghilarov 1978), Taiwan (Tseng 1978), Korea (Lee et al. 1997), Mexico (Hoffmann & L��pez-Campos 2000), Spain (Domingo-Quero et al. 2003). Redescriptions. Atyeo (1960a, 1963a), Shiba & Morikawa (1966), Wallace & Mahon (1972), Tseng (1978). Remarks. Ewing (1917, not 1914 as mentioned by both Atyeo [1960a] and Wainstein et al. [1978]) described Bdella tenuirostris, without noticing the preoccupied name erected by Koch (1839: 23). Type deposition. USNM. 154. Spinibdella thori (Meyer & Ryke, 1959): 375; Bathurst, South Africa, ex grass and soil.��� Atyeo, 1963a: 174. Original designation: Bdella thori Meyer & Ryke. Distribution. South Africa (Meyer & Ryke 1959; Halliday 2005), Australia (Atyeo 1963a; Wallace & Mahon 1972; Halliday 2005), Hawaii (Swift & Goff 1987; 2001), Mexico (Hoffmann & L��pez-Campos 2000), Iran (Abbaszadeh et al. 2010). Redescriptions. Atyeo (1963a), Wallace & Mahon (1972), Swift & Goff (1987), Omukunda et al. (2012). Remarks. Male unknown. Type deposition. Institute for Zoological Research, Potchefstroom University, South Africa. 155. Spinibdella trinomma Omukunda, Theron & Ueckermann, 2012: 3; Kwazulu, Natal, South Africa. Type deposition. NCA. 156. Spinibdella trisetosa (Jacot, 1938): 128 comb. nov.; Micanope, Florida, USA, ex leaf litter. Original designation: Bdella trisetosa Jacot. Remarks. This species is herein transferred to the genus Spinibdella due to having two ventral setae on the hypostome, tricobothria present on tibiae I, IV, tarsi III and IV, setae lps present, and the palpal tibiotarsus truncate. Type deposition. USNM. 157. Spinibdella yeni Tseng, 1978: 44; Taipu, Chiayi Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.)., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. 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