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We show that spontaneous density segregation in dense systems of aligning circle swimmers is a condensation phenomenon at odds with the phase separation scenarios usually observed in two-dimensional active matter. The condensates, which take the form of vortices or rotating polar packets, can absorb a finite fraction of the particles in the system, and keep a finite or slowly growing size as their mass increases. Our results are obtained both at particle and continuous levels. We consider both ferromagnetic and nematic alignment, and both identical and disordered chiralities. Condensation implies synchronization, even though our systems are in 2D and bear strictly local interactions. We propose a phenomenological theory based on observed mechanisms that accounts qualitatively for our results., Comment: 6 pages, 4 figures

Two-dimensional (2D) crystals made of active particles were shown recently to be able to experience extremely large spontaneous deformations without melting. The root of this phenomenon was argued to lie in the time-persistence of the orientation of the intrinsic axes of particles. Here, we pursue this idea and consider passive systems subjected to time-persistent external perturbations. We first study a 2D XY model subjected to time-correlated noise and find that it can remain quasi-ordered in spite of correlations decaying much faster than allowed in equilibrium. We then study a simple model of a passive 2D crystal immersed in a bath of active particles, and show that it can sustain large deformations without melting., Comment: 5 pages, 2 figures

We demonstrate that two-dimensional crystals made of active particles can experience extremely large spontaneous deformations without melting. Using particles mostly interacting via pairwise repulsive forces, we show that such active crystals maintain long-range bond order and algebraically-decaying positional order, but with an exponent $\eta$ not limited by the $\tfrac{1}{3}$ bound given by the (equilibrium) KTHNY theory. We rationalize our findings using linear elastic theory and show the existence of two well-defined effective temperatures quantifying respectively large-scale deformations and bond-order fluctuations. The root of these phenomena lies in the sole time-persistence of the intrinsic axes of particles, and they should thus be observed in many different situations., Comment: 6 pages, 4 figures

We show that arbitrarily large polar flocks are susceptible to the presence of a single small obstacle. In a wide region of parameter space, the obstacle triggers counter-propagating dense bands leading to reversals of the flow. In very large systems, these bands interact yielding a never-ending chaotic dynamics that constitutes a new disordered phase of the system. While most of these results were obtained using simulations of aligning self-propelled particles, we find similar phenomena at the continuous level, not when considering the basic Toner-Tu hydrodynamic theory, but in simulations of truncations of the relevant Boltzmann equation., Comment: 5 pages, 4 figures, Supplementary information. Movies available from authors on request

We investigate the susceptibility of long-range ordered phases of two-dimensional dry aligning active matter to population disorder, taken in the form of a distribution of intrinsic individual chiralities. Using a combination of particle-level models and hydrodynamic theories derived from them, we show that while in finite systems all ordered phases resist a finite amount of such chirality disorder, the homogeneous ones (polar flocks and active nematics) are unstable to any amount of disorder in the infinite-size limit. On the other hand, we find that the inhomogeneous solutions of the coexistence phase (bands) may resist a finite amount of chirality disorder even asymptotically., Comment: 5 pages, 4 figure, supplementary information is available upon request

Active matter comprises individual units that convert energy into mechanical motion. In many examples, such as bacterial systems and biofilament assays, constituent units are elongated and can give rise to local nematic orientational order. Such `active nematics' systems have attracted much attention from both theorists and experimentalists. However, despite intense research efforts, data-driven quantitative modeling has not been achieved, a situation mainly due to the lack of systematic experimental data and to the large number of parameters of current models. Here we introduce a new active nematics system made of swarming filamentous bacteria. We simultaneously measure orientation and velocity fields and show that the complex spatiotemporal dynamics of our system can be quantitatively reproduced by a new type of microscopic model for active suspensions whose important parameters are all estimated from comprehensive experimental data. This provides unprecedented access to key effective parameters and mechanisms governing active nematics. Our approach is applicable to different types of dense suspensions and shows a path towards more quantitative active matter research., Comment: 9 pages, 7 figures

We show that spatial quenched disorder affects polar active matter in ways more complex and far-reaching than believed heretofore. Using simulations of the 2D Vicsek model subjected to random couplings or a disordered scattering field, we find in particular that ergodicity is lost in the ordered phase, the nature of which we show to depend qualitatively on the type of quenched disorder: for random couplings, it remains long-range ordered, but qualitatively different from the pure (disorderless) case. For random scatterers, polar order varies with system size but we find strong non-self-averaging, with sample-to-sample fluctuations dominating asymptotically, which prevents us from elucidating the asymptotic status of order., Comment: 6 pages, 5 figures, comments are welcome

We revisit motility-induced phase separation in two models of active particles interacting by pairwise repulsion. We show that the resulting dense phase contains gas bubbles distributed algebraically up to a typically large cutoff scale. At large enough system size and/or global density, all the gas may be contained inside the bubbles, at which point the system is microphase-separated with a finite cut-off bubble scale. We observe that the ordering is anomalous, with different dynamics for the coarsening of the dense phase and of the gas bubbles. This phenomenology is reproduced by a "reduced bubble model" that implements the basic idea of reverse Ostwald ripening put forward in Tjhung et al. [Phys. Rev. X 8, 031080 (2018)].

We show that the dominant mode of alignment plays an important role in dry active nematics,leading to two dynamical sub-classes defined by the nature of the instability of the nematic bands that characterize, in these systems, the coexistence phase separating the isotropic and fluctuating nematic states. In addition to the well-known instability inducing long undulations along the band, another stronger instability leading to the break-up of the band in many transversal segments may arise. We elucidate the origin of this strong instability for a realistic model of self-propelled rods and determine the high-order nonlinear terms responsible for it at the hydrodynamic level., Comment: 5 pages, 4 figures. Supplementary and movies are available upon request

We study a robust model of self-propelled rods interacting via volume exclusion and show that its collective dynamics encompasses both that of the corresponding Vicsek-style model (where local alignment is the sole interaction), and motility-induced phase separation (which occurs when repulsion is dominant). These results unify these heretofore largely disconnected bodies of knowledge on dry active matter and clarify the nature of the various phases involved., Comment: 5 pages, 6 figures, supplementary information and movies are available upon request

We show that fore-aft asymmetry, a generic feature of living organisms and some active matter systems, can have a strong influence on the collective properties of even the simplest flocking models. Specifically, an arbitrarily weak asymmetry favoring front neighbors changes qualitatively the phase diagram of the Vicsek model. A region where many sharp traveling band solutions coexist is present at low noise strength, below the Toner-Tu liquid, at odds with the phase-separation scenario well describing the usual isotropic model. Inside this region, a `banded liquid' phase with algebraic density distribution coexists with band solutions. Linear stability analysis at the hydrodynamic level suggests that these results are generic and not specific to the Vicsek model., Comment: 6 pages, 5 figures

We propose a simple deterministic dynamic equation and reveal the mechanism of large-scale endless evolvement of spatial density inhomogeneity in active nematic. We determine the phase regions analytically. The interplay of density, magnitude of nematic order, and nematic director is crucial for the long-wave-length instability and the emergence of seemingly fluctuated collective motions. Ordered nematic domains can absorb particles, grow and divide endlessly. The present finding extends our understanding of the large-scale and seemingly fluctuated organization in active fluids., Comment: 5 pages, 3 figures

Using integral equation theory of liquids to a binary mixed fluid lipid membrane, we study the membrane-mediated interactions between the macroions and the redistribution of neutral and charged lipids due to binding macroions. We find that when the concentration of binding macroions is infinitely dilute, the main contribution to the attractive potential between macroions is the line tension between neutral and charged lipids of the membrane, and the bridging effect also contributes to the attraction. As the relative concentration of charged lipids is increased, we observe a repulsive - attractive - repulsive potential transition due to the competition between the line tension of lipids and screened electrostatic macroion-macroion interactions. For the finite concentration of macroions, the main feature of the attraction is similar to the infinite dilution case. However, due to the interplay of formation of charged lipid - macroion complexes, the line tension of redistributed binary lipids induced by single macroion is lowered in this case, and the maximum of attractive potential will shift toward the higher values of the charged lipid concentration.