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Family— Tingiostachyaceae Gao and Thomas emend . S. J. Wang and Spencer (2017)., Published as part of Wang, Jun, Hilton, Jason, Pfefferkorn, Hermann W., Wang, Shijun, Zhang, Yi, Bek, Jiri, Pšenička, Josef, Seyfullah, Leyla J. & Dilcher, David, 2021, Ancient noeggerathialean reveals the seed plant sister group diversified alongside the primary seed plant radiation, pp. 1-7 in Proceedings of the National Academy of Sciences 118 (11) on page 2, DOI: 10.1073/pnas.2013442118, http://zenodo.org/record/4596604

Order— Noeggerathiales Nemejc J. Wang et al. emend. nov. Emended Order diagnosis—Heterosporous plants with compact, bisporangiate pseudostrobilus. Sporophylls laterally basally fused to form a semidisc or disk. Sporangia sessile, attached to sporophyll surface. Leaf once-pinnate, compound, with multiveined pinnule. Pseudostrobilus and leaf rachis with bilateral, Ω- shaped vascular bundle and secondary xylem., Published as part of Wang, Jun, Hilton, Jason, Pfefferkorn, Hermann W., Wang, Shijun, Zhang, Yi, Bek, Jiri, Pšenička, Josef, Seyfullah, Leyla J. & Dilcher, David, 2021, Ancient noeggerathialean reveals the seed plant sister group diversified alongside the primary seed plant radiation, pp. 1-7 in Proceedings of the National Academy of Sciences 118 (11) on page 2, DOI: 10.1073/pnas.2013442118, http://zenodo.org/record/4596604

Lioponera neocaledonica Jouault, Ramage & Perrichot sp. nov. urn:lsid:zoobank.org:act: 7612551C-6C0A-4931-9D06-8FC8062DB5E3 Figs 4, 5B Diagnosis Lioponera neocaledonica Jouault, Ramage & Perrichot sp. nov. is mostly characterized by the combination of the following features: mesosoma dorsolaterally conspicuously marginate, propodeal declivity without dorsal margin; posterolateral corners of petiole not projected into spines. The two latter features easily serve to distinguish L. neocaledonica Jouault, Ramage & Perrichot sp. nov. from the other New Caledonian species L. cohici (Wilson, 1957) (Fig. 5A) and L. dumbletoni (Wilson, 1957) (Fig. 5C). Etymology The specific epithet refers to New Caledonia where the species was discovered. Type material Holotype NEW CALEDONIA ��� wingless ♀; Grande Terre, Province Sud, Sarram��a, Plateau de Dogny trail; 21��37���31��� S, 165��52���5��� E; 441 m; 16 Nov. 2011; V. Perrichot leg. [VP188]; mixed mountain forest, hand picking on ground; MNHN. Paratypes NEW CALEDONIA ��� 2 wingless ♀♀; same collection data as for holotype; MNHN ��� 1 wingless ♀; Grande Terre, Province Sud, Montagne des Sources; 22��09���35.1��� S, 166��35���42.4��� E; 394 m; 23 Oct. 2010; M. Berman leg. [CSIRONC0055]; ANIC 32-146088. Description (holotype) MEASUREMENTS (in mm; holotype in brackets). TL 4.12���4.15 (4.15); HL 0.8���0.85 (0.85); HW 0.7���0.77 (0.77); SL 0.45���0.52 (0.52); ED 0.2���0.25 (0.2); PW 0.65���0.72 (0.7); MH 0.52���0.62 (0.62); MSL 1.025��� 1.12 (1.12); PL 0.45���0.55 (0.55); PH 0.4���0.42 (0.42); DPW 0.67���0.7 (0.7); T1W 0.65���0.77 (0.77); T1L 0.5���0.62 (0.62); T2W 0.75���0.82 (0.82); T2L 0.6���0.65 (0.65). Indices: CI 84���91 (91); SI 64���69 (67); LPI 76���94 (77); DPI 127���150 (127). HEAD (Fig. 4C). Sub-rectangular, slightly longer than broad, widest above midline of head; lateral sides slightly convex; vertexal margin slightly concave, posterior lateral corners slightly rounded. Parafrontal ridges angular. Compound eyes of medium size, situated below midline of head. Mandible elongate, triangular, masticatory margin with 10 small teeth. Antennae with scape short and clavate, reaching up to two fifth of posterior margin of head. MESOSOMA (Fig. 4 A���B, D). Stout, roughly rectangular in dorsal view, with conspicuous dorsolateral margins, the margins notched around promesonotal junction; mesosomal dorsal surface convex, propodeal declivity without dorsal margin. METASOMA. Petiole (Fig. 4 A���B, E) broader than long, slightly trapezoidal, widest at two thirds of length; anterior petiolar surface concave; dorsal petiolar surface convex, with dorsolateral margins well developed and convex, without anterodorsal and posterodorsal margins; posterior petiolar surface slightly convex; anterodorsal and posterodorsal corners angular. Subpetiolar process small, triangular, with stout acute somewhat hooked apex; fenestra absent. Gaster elongate (Fig. 4 A���B), with distinct girdling constriction (cinctus) between abdominal segments III and IV.Abdominal segment II trapezoidal, slightly broader than long, widest at one third of length; dorsal surface slightly convex; dorsolateral margins well developed and strongly convex; anterodorsal margin sinuate; anterodorsal corner broadly rounded; posterolateral corner angular. Sting exerted, upcurved. INTEGUMENT. Mostly black except last two funicular articles, tarsomeres, and posterior margins of abdominal segments IV to V brown. Body finely and sparsely punctate; reticulation present on sides of the mesosoma, becoming shallower from propleura to propodeum; cinctus, abdominal segment IV, base of abdominal segments V, VI and pygidium transversally, finely and shallowly striate. Body covered with suberect hairs; scape with abundant appressed hairs; tibiae with scattered decumbent hairs; tarsi with abundant decumbent to appressed hairs; declivous face of propodeum with short, dense and appressed hairs. Remarks The potential presence of ergatogynes in Lioponera (Wilson 1957; Borowiec 2016) prevents assignment of the present specimens to workers or ergatogynes. Key to the species of Lioponera of New Caledonia (modified from Wilson 1957) 1. Large species (HW> 1.1 mm), with long scapes reaching nearly to posterior corners of head; mesosomal dorsolateral margins absent................................ Lioponera dumbletoni (Wilson, 1957) ��� Small species (HW Lioponera cohici (Wilson, 1957) ��� Mesosomal posterodorsal margin absent (Fig. 4D); posterolateral corners of petiole not projected into spines (Fig. 5B)..................... Lioponera neocaledonica Jouault, Ramage & Perrichot sp. nov., Published as part of Ramage, Thibault, Jouault, Corentin, Schmidt, Alexander R., Seyfullah, Leyla J. & Perrichot, Vincent, 2019, Two new ant species (Formicidae: Dorylinae, Ponerinae) from New Caledonia, pp. 1-14 in European Journal of Taxonomy 589 on pages 7-8, DOI: 10.5852/ejt.2019.589, http://zenodo.org/record/3659941, {"references":["Wilson E. O. 1957. The discovery of cerapachyine ants on New Caledonia, with the description of new species of Phyracaces and Sphinctomyrmex. Breviora 74: 1 - 9.","Emery C. 1914. Les fourmis de la Nouvelle-Caledonie et des iles Loyalty. Nova Caledonia. Forschungen in Neu-Caledonien und auf den Loyalty-Inseln. Recherches scientifiques en Nouvelle-Caledonie et aux iles Loyalty. A. Zoologie 1: 393 - 437. https: // doi. org / 10.5962 / bhl. title. 6790","Wilson E. O. 1958. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bulletin of the Museum of Comparative Zoology 118: 101 - 153.","Borowiec M. L. 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280. https: // doi. org / 10.3897 / zookeys. 608.9427"]}

Leptogenys loarelae Ramage sp. nov. urn:lsid:zoobank.org:act: 819E8C18-56F7-4CB5-AE07-00B5406D924A Fig. 1 Diagnosis Leptogenys loarelae Ramage sp. nov. can be distinguished from other New Caledonia species of Leptogenys by the unique combination of the following features: integument smooth and shining, mostly black colored with blueish reflection; third antennal segment markedly elongate, more than twice the length of pedicel; mandible elongate, triangular; petiole elongate, nearly three times longer than broad in dorsal view. Etymology The specific epithet refers to the first name of Thibault Ramage���s daughter, Loarel. The species epithet is to be treated as a noun in the genitive case. Type material Holotype NEW CALEDONIA ��� wingless ♀; Grande Terre, Province Sud, Sarram��a, Plateau de Dogny trail; 21��37���10.7��� S, 165��52���31.9��� E; 897 m; 16 Nov. 2011; V. Perrichot leg. [VP209]; mixed mountain forest, hand picking on ground; MNHN. Description Worker holotype MEASUREMENTS (in mm). TL 11.5; HL 2.06; HW 1.3; SL 2.56; ED 0.5; PW 1.16; MSL 3.65; PL 1.83; PH 0.8; DPW 0.63; CI 62.9; SI 197.4; LPI 43.6; DPI 34.5. HEAD. Elongate, sub-trapezoidal, lateral sides diverging anteriorly (Fig. 1B); lateral margin barely convex; posterior margin weakly concave medially, posterior lateral corners rounded. Clypeus triangular, with blunt apex and sligthly concave side margins. In full-face view, eye large, oval, maximum diameter greater than twice the maximum width of scape; placed at head midline. Antennae with scape long and slightly enlarged medially, surpassing posterior margin of head by its 2������ th; funicular segments elongate; third antennal segment markedly elongate, more than twice the length of the second. Mandible triangular, elongate, shorter than head width; masticatory margin edentate. MESOSOMA (Fig. 1A, C). Elongate, compressed laterally, widest at pronotum; mesosomal dorsal surface in profile slightly convex. In dorsal view, mesonotum as long as broad. Propodeal spiracle slit-like. Petiole (Fig. 1A, C���D) elongate, nearly three times as long as broad, trapezoidal, widest at apex; dorsal surface of petiolar node convex, posterior surface slightly concave, anterodorsal and posterodorsal corners angular. Subpetiolar process small, trapezoidal; located beneath anterior 1������ th of the petiole (Fig. 1D). Gaster elongate. INTEGUMENT. Smooth and shining, very finely and sparsely punctate; mostly black with blueish reflection, except mandibles, antennal radicel and funicular articles, tarsomeres and hypopygium and pygidium reddish brown. Dorsum of body covered by erect and suberect yellow setae; setae longer on head, scape, pronotum, procoxa and gaster, but shorter on remaining body. Key to the Leptogenys species (workers) of New Caledonia (modified from Wilson 1958) 1. Large species (HW> 1.2 mm).......................................................................................................... 2 ��� Small species (HW ��� 1.2 mm).......................................................................................................... 3 2. Mandibles exceptionally long, their length measured from insertion to tip exceeding the head width (Fig. 2A)............................................................................................ Leptogenys rouxi Emery, 1914 ��� Mandibles shorter, their length much less than the head width (Fig. 1B)................................................................................................................................. Leptogenys loarelae Ramage sp. nov. 3. Head entirely covered by coarse punctures (Fig. 2C).................. Leptogenys punctata Emery, 1914 ��� Head entirely smooth and shining (Fig. 2B)..................................................................................... 4 4. Larger species, head width ��� 0.9 mm; in dorsal view, posterior corners of petiolar node forming distinct angles of 45���50�� (Fig. 3A).......................................... Leptogenys acutangula Emery, 1914 ��� Smaller species, head width not exceeding 0.75 mm; in dorsal view, posterior corners of petiolar node thick and broadly rounded (Fig. 3B)..................................... Leptogenys sagaris Wilson, 1958 Subfamily Dorylinae Leach, 1815 Genus Lioponera Mayr, 1879, Published as part of Ramage, Thibault, Jouault, Corentin, Schmidt, Alexander R., Seyfullah, Leyla J. & Perrichot, Vincent, 2019, Two new ant species (Formicidae: Dorylinae, Ponerinae) from New Caledonia, pp. 1-14 in European Journal of Taxonomy 589 on pages 4-7, DOI: 10.5852/ejt.2019.589, http://zenodo.org/record/3659941, {"references":["Wilson E. O. 1958. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bulletin of the Museum of Comparative Zoology 118: 101 - 153.","Emery C. 1914. Les fourmis de la Nouvelle-Caledonie et des iles Loyalty. Nova Caledonia. Forschungen in Neu-Caledonien und auf den Loyalty-Inseln. Recherches scientifiques en Nouvelle-Caledonie et aux iles Loyalty. A. Zoologie 1: 393 - 437. https: // doi. org / 10.5962 / bhl. title. 6790"]}