Search

Freshwater ecosystems in the Indo-Burma biodiversity hotspot face immediate threats through habitat loss and species extinction. Systems to monitor ecological status and trends in biodiversity are therefore crucially needed. Myanmar is part of Indo-Burma but with no past experience of biomonitoring in freshwaters. In this study, we aimed to assess the ecological and biodiversity status of a lowland river network in south-central Myanmar by identifying and quantifying pressures using macroinvertebrates as bioindicators. Novel data on water quality (nutrients, sediments and metals), hydromorphology (Morphological Quality Index; MQI), habitat quality (LitterSiltation Index; LSI), land use, and macroinvertebrates were collected from 25 river sites. The dominant pressures on rivers were urban land use, inputs of untreated sewage, in-stream and riparian garbage littering, run-off from agricultural fields and plantations, as well as physical habitat degradation. Water chemistry data indicated inputs of sediments and nutrients to degraded streams, but no obvious metal pollution. The LSI and MQI indices indicated high perturbation in agricultural and urban areas, respectively. Ecological status was assessed using afirst version of a modified Average Score per Taxon index (ASPT), while biodiversity was assessed by family richness within the orders Ephemeroptera, Plecoptera, Trichoptera, Coleoptera and Odonata (EPTCO), which was tested against the pressure gradient by principal component regressions. ASPT had high diagnostic capabilities (R2 = 0.68, p < 0.001) and showed that the index can be used to evaluate ecological water quality in this region. Biodiversity, expressed as family richness, also declined along the gradient (R2 = 0.59, p = 0.041), giving support to the fact that current land-use practices in this area are unsustainable.

Up to present 602 species and 65 genera of fungus gnats, family Mycetophilidae, are published from Norway. Extensive collecting supported by the Norwegian Biodiversity Information Centre (NBIC) over the eight last years, with special focus on insect fauna in northern Norway, has documented 240 additional species and 2 additional genera from Norway, of which 118 species are considered as new to science. Based on a thorough review of the species previously published from Norway, we have crossed out six species as misidentified. One new synonym is established: Boletina conformis Siebke, 1863 syn. n. = Boletina plana (Walker, 1856). Two species are restituated based on integrative studies including DNA barcodes. These are Ectrepesthoneura bucera Plassmann, 1980 sp. restit., found to be a distinct species separate from Ectrepesthoneura ovata Ostroverkhova, 1977, and Trichonta trifida Lundstrom, 1909 sp. restit., found to be a distinct species separate from Trichonta vulcani (Dziedzicki, 1889). The updated, validated A-checklist includes 821 species of which 703 (86%) refer to formally described species and 118 (14%) to potentially undescribed species, referred to by their interim names as used on BOLD and in our databases. All species are documented with specimens in the museum collections at either Tromsø University Museum (TMU, 781 species, 95%) and/or the Natural History Museum in Oslo (NHMO, 382 species, 47%). Another 14 published species are transferred to a B-checklist with currently unvalidated species, as we fail to recover voucher representatives. Supported by the Norwegian Barcode of Life (NorBOL) network, we have DNA barcoded as many species as possible contributing to the reference library on The Barcode of Life Project (BOLD). Hence, 756 (92%) of the validated Norwegian species are currently documented with DNA barcodes and assigned Barcode Index Numbers (BINs) on BOLD, including the majority of the species considered new to science based on morphology (103 species, 87% of the 118). The checklist, is kept in a short format giving the published species names or interim names as used on BOLD and in our databases, depository information, assigned BINs with indications of discordance, and finally their (2015) Red List status in Norway. An accompanying dataset containing recording details and distribution of all new records from Norway of described species, is published on GBIF and on Norway’s Species Map Service (Artskart). Posted here with permission from the journal. Norwegian Journal of Entomology: http://www.entomologi.no/journals/nje/nje.htm. Norsk entomologisk forening: http://www.entomologi.no/index.htm

Manota triseta sp. n. Figs 12 A–C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish. Legs yellowish to light brown, apical fourth to third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with brown knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.3 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.4 times as long as palpomere 4. 9-11 strong postocular setae. Thorax. Anepisternum with 52–57 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 13–17 setae; metepisternum with 15–16 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.7–1.9 mm. Hypopygium (Figs 12 A–C). Sternite 9 laterally free from gonocoxa at posterior 2/3, anterior 1/3 fused with gonocoxa, posterior margin membranous, shallowly concave, extending about the middle between bases of gonocoxa and gonostylus, anterior margin with deep and wide V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin simple. Parastylar lobe large, plate-like, two setae anteriorly on ventral surface. Dorsal medial margin of gonocoxa simple, slightly convex, with tight aggregation of fine setae posteriorly on ventral surface. On the ventral side of the posteromedial angle a finger-like lobe bearing two apical setae. Two juxtagonostylar setae present, a normal seta and apically curved simple megaseta, both arising from a common basal body, which is about of the length of megaseta. Gonostylus elongated, about 5 times as long as wide, slightly curved, with a medial small hump-like lobe bearing 2 normal setae, and with 3 very long and medially directed setae apically, otherwise the gonostylar setosity is similar to that of gonocoxa. Aedeagus short, subtriangular, with prominent lateral shoulders, apex curved ventrad. Hypoproct posteriorly not reaching the base of gonostylus, the ventral setae strong and forming a mesial longitudinal stripe of ca. 5 setae on each half. Cerci medially separate, simple, the setae confined to apical part, the longest ones maximally two times longer than the apical width of cercus. Female. Unknown. Etymology. The name is Latin, triseta, ‘three setae’, referring to the three very long medially directed apical setae of gonostylus; noun in apposition. Comments. The non-setose laterotergite, setose preepisternum 2, and the gonocoxa posterodorsally with a fingerlike lobe with two apical setae group M. triseta sp. n. together to M. comata, M. pedicellata, M. relicina, M. petiolata and M. burundiensis. Among them, M. triseta is unique in having gonostylus apically or subapically without small lobe that bears two rather short megasetae. Otherwise, M. triseta is similar to M. issongo because of very long, medially directed setae apically on the gonostylus but distinguished in having (1) the finger-like lobe posterodorsally on the gonocoxa with two setae at apex (one seta in M. issongo), (2) the three very long setae apically on the gonostylus (two setae in M. issongo), (3) the two setae medially on the gonostylus arise from a distinct lobe (no distinct lobe discernible in M. issongo), (4) the two setae on the parastylar lobe separated by a distinct gap (two setae on the parastylar lobe close to each other in M. issongo), and (5) the ventral setae of hypoproct are in one row on each side, flanking the apex of aedeagus (setae on the hypoproct arranged as a patch in M. issongo). Types. Holotype. Male, GHANA, Volta region, Wli Falls, Agumatsa (St. 6A), 12– 21.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO). Paratypes. 4 males, as holotype (slide mounted, IZBE and NHMO); 3 males, as holotype except 11– 20.11.1993 (slide mounted, IZBE and NHMO); 2 males, GHANA, Volta region, Wli Falls, Afegame (St. 9A), 7– 10.03.1993, NUFU-ZMBN/ leg. J. Kjaerandsen (slide mounted, NHMO); male, GUINEA, Monts Nimba / DL-HP-JYR // Source 1200 M/ Piége Malaise / 27-iv–7-v-1993 (slide mounted, MNHN); male, GUINEA, Mt. Nimba, Ft. Gal. du Zougué, 750 m, 18–29.VI.1991, Girard et Legrand col., Piège de Malaise (slide mounted, MNHN). Additional material. 23 males, GHANA, Volta region; Wli Falls, Agumatsa; St: 11, 11- 20.11.1993; NUFUproject ZMBN; leg. J. Kjaerandsen (1 slide mounted, 22 in alcohol, IZBE and NHMO); male same data except 13– 20.11.1993 (in alcohol, NHMO) .

Manota clurina Hippa & Kurina, 2012 Material. CÔTE D’IVOIRE: male, rég. de Tai, 1–15.ii.1985, G. Gouturier & V. Van Zeijst Réc., BIO (24), Friche de 6 ans, Piége malaise, ORSTOM-Paris, Mission UNESCO (slide mounted, MNHN). Remarks. The species was described from Uganda (Hippa & Kurina 2012) and has not been recorded since. Manota comata Hippa & Kurina, 2012 Material. GHANA: 9 males, Volta region, Wli Falls, Agumatsa (St. 10), 11– 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, IZBE and NHMO); 6 males, Central Region, Kakum, Entwikrom, 31.10– 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (3 slide mounted, IZBE, 3 in alcohol, NHMO); 5 males, same data except 8– 15.11.1994 (2 slide mounted NHMO, 3 in alcohol, NHMO); male, Western region, Ankasa game prod. Reserve, 6–12.XII.1993 (St: 1–8, 13), NUFU-project ZMBN, leg. J. Kjaerandsen & T. Andersen (in alcohol, NHMO). GUINEA: male, Mt. Nimba, Ft. Gal. du Zougué, 750m, 18–29.vi.1991, Piège de Malaise, Girard et Legrand col. (slide mounted, MNHN). D.R. CONGO: male, Oriental Prov., Likombo forest, 2 km SW, Bomane, 1.28349, 23.72358, 20–22.v.2010, leg. A.H. Kirk-Spriggs, Malaise traps lowland evergreen primary forest (disturbed), Boyekoli Ebale Congo Expedition 2010, BECE 00466 (slide mounted, BMSA). UGANDA: male, Western region, Budongo Forest; N 01°43’18,9’’ E 031°31’46,7’’ (Site 9) 4–8 July 2010 G. Kvifte leg., Malaise trap (slide mounted, NHMO). Remarks.The species has so far been known from the type locality in Ghana and from Uganda (Hippa & Kurina 2012).

Manota kjaerandseni sp. n. Figs 6 A–C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown. Legs yellowish, apical third of hind femur and basal fourth of mid femur infuscated. Wing with brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 subapically constricted, without parasegment; palpomere 5 ca. 1.6–1.7 times longer than palpomere 4. 8–9 strong postocular setae. Thorax. Anepisternum with 25–26 setae; anterior basalare with 5–6 setae; laterotergite with 19 setae; preepisternum 2 non-setose; metepisternum with 20–23 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.6–1.7 mm. Hypopygium (Figs 6 A–C). Sternite 9 laterally fused to gonocoxa, posterior margin convex, extending over base of gonostylus, anterior margin with a narrow deep V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like with one strong apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa medially simple, posteriorly drawn out into an apically rounded large lobe, covering basal third of gonostylus and bearing two subapical, medially directed megasetae: the posterior one short and pointed, the anterior one longer, with a whip-like apical part. Ventrally from dorsal medial margin of gonocoxa a large medially bulging plate-like lobe bearing a row of 6 simple and strong megasetae, 3 anteriors apically slightly tapering, 3 posteriors blunt, slightly widening apically. Two juxtagonostylar setae present, both are simple, apically pointed megasetae, arising from small separate basal bodies which are similar to normal sockets of setae. Gonostylus elongated, basal third heel-like bulging medially, with an extra-long apical seta and 8–9 longer setae along apical 2/3 of medial margin. Aedeagus elongate subtriangular, apical 2/3 almost parallel-sided, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to the middle of gonostylus, number of ventral setae (sternite 10) ca. 30 on each half. Cerci medially separated, setae scattered over the surface, apical setae deviating from others, longer than the subapical width of cercus. Female. Unknown. Etymology. The species is named after Dr. Jostein Kjaerandsen (University of Tromsø, Norway), an active student of fungus gnats’ systematics and the collector of a number of the specimens from Ghana studied in this paper. Comments. Manota kjaerandseni sp. n. belongs to a group of 14 Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) non-setose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly without a row of scale- or leaf-like megasetae. Among them, the new species is unique in having a row of 6 slender and blunt megasetae marginally on a bulging plate-like lobe at the dorsal medial margin of gonocoxa and in having the posterodorsal part of gonocoxa elongated and posteriorly rounded and with two megasetae on the medial margin. Types. Holotype. Male, GHANA, Volta region, Wli Falls, Afegame (St. 6A), 9– 12.03.1993, NUFU-ZMBN, leg. J.S. Amakye & J. Kjaerandsen (slide mounted, NHMO). Paratypes. Male, GHANA, Western region, Ankasa game prod. Reserve (St. 2), 06– 12.12.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, GHANA, Central Region, Kakum, Entwikrom, 8– 15.10.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE); male, same data except 8– 15.11.1994 (slide mounted, IZBE); 2 males, CÔTE D’IVOIRE, rég. de Tai / 1–15.ii.1985 / G. Gouturier & V. Van Zeijst Réc. // BIO (24) / Friche 6 ans / Piège malaise // ORSTOM-Paris / Mission UNESCO (slide mounted, MNHN). Additional material. 2 males, GHANA, Wli Falls, 4– 13.03.1993, leg. J. Kjaerandsen (in alcohol, NHMO) .

Manota polylobata sp. n. Figs 11 A–D Male. Colour. Head light brown, face somewhat paler. Antenna: scape and pedicel light brown, flagella absent in holotype. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish, apical fourth of hind femur slightly infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 not measurable in holotype. Nine strong postocular setae. Thorax. Anepisternum with 40 setae; anterior basalare with 10 setae, laterotergite with 34 setae, preepisternum 2 non-setose; metepisternum with 21 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 2.2 mm. Hypopygium (Figs 11 A–D). Sternite 9 laterally fused to gonocoxa, posterior margin broad, convex, extending over middle between bases of gonocoxa and gonostylus, anterior margin with a deep and narrow V-shaped incision medially, anterior 1/3 non-setose, posterior 2/3 setose, setae similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa oblique, laterally from gonostylus drawn into a suboval lobe about half of gonostylus length, bearing conspicuous microtrichia on ventral surface and two subapical twisted and apically whip-like megasetae. Parastylar lobe bifurcate, anterior branch short, smaller than posterior that has an apical seta. No paraapodemal lobes identifiable. Dorsal medial margin of gonocoxa simple with submembranous lobe on more ventral level, just anteriorly from juxtagonostylar setae, bearing row of three megasetae gradually diminishing in size posteriad: anterior megaseta flat and apically dilated, medial megaseta similar but apically somewhat narrower, posterior megaseta pointed. Gonocoxal apodeme unusual, posteriorly strongly widened. Setae on dorsal side of gonocoxa similar to those on ventral side. Two juxtagonostylar seta present, both short, flattened and expanded megasetae, arising from a common basal body which is shorter than megasetae. Gonostylus complex, with prominent medial lobe and two small lobes laterally; the medial prominent lobe longitudinally drawn out, about half as long as gonocoxa, apically conical with ca. 8 long marginal setae, medial margin with an aggregation of ca. 10 setae on ventral surface, dorsally with an additional medial lobe bearing a row of 6–7 setae; the lateral lobe subquadrangular with 6–8 small setae apically; the medial lobe finger-like with 4 apical setae. Aedeagus subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly near to apex of gonostylus, apical third tapering, number of ventral setae (sternite 10) ca. 40 on each half. Cerci medially separated, setae scattered over the surface, apical 4–5 setae about twice as long as rest of cercal setae. Female. Unknown. Etymology. The name is combined from Greek πολύ (polú) ‘many’ and Latin lobata ‘lobed, with lobes’, referring to the four-lobed gonostylus. Comments. Manota polylobata sp. n. belongs to a group of 14 Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) nonsetose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly without a row of scale- or leaf-like megasetae (see couplets 21 to 33 in the Key of the Afrotropical species above). Among them, M. polylobata is most similar to M. tridactyla in having a complex gonostylus, two leaf-like juxtagonostylar megasetae, and megasetae at dorsal medial margin of the gonocoxa close to posterior end of the margin (should not to be confused with a distinct row of leaf- or scale like megasetae posteriorly on dorsal medial margin of gonocoxa as defined under M. geniculata). Manota polylobata differs in having 3 megasetae at dorsal medial margin of gonocoxa with the anteriormost megaseta apically dilated (4 megasetae with the anteriormost hook-like in M. tridactyla). Types. Holotype. Male, NIGERIA, SE State, Obudu CR, 27.ix.1973, J.T. Medler coll. (slide mounted, MNHN).

Manota macrodon Hippa, 2008 Material. MADAGASCAR: male, Fianarantsoa, Ranomafana National Park, Talatakely, 800m SW entrance, 21��15,48���S 47��25,27���E, 16���19.x. 2014, 610 m, leg. A.H. Kirk-Spriggs & R. Harin���Hala, Malaise trap secondary rainforest (slide mounted, BMSA); 4 males, Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18��46,25���S 48��25,93���E; 23���25.x. 2014, 960 m, leg. A.H. Kirk-Spriggs & R. Harin���Hala, Malaise trap primary rainforest (slide mounted, BMSA and IZBE). Remarks. The species has so far been known only from the holotype collected from Province Fianarantsoa in Madagascar (Hippa 2008). The current records indicate a wider distribution on the Island., Published as part of Hippa, Heikki, S��li, Geir & Kurina, Olavi, 2019, New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species, pp. 401-441 in Zootaxa 4652 (3) on page 431, DOI: 10.11646/zootaxa.4652.3.1, http://zenodo.org/record/3364902, {"references":["Hippa, H. (2008) Notes on Afrotropical Manota Williston (Diptera: Mycetophilidae), with the description of seven new species. Zootaxa, 1741 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 1741.1.1"]}

Manota cultigera Hippa, 2008 Material. MADAGASCAR: 2 males, Mahajanga, Boeny, Ankarafantsika NP, 16.30268S 46.81041E, 58 m, Malaise trap, #MAD09-38, 9.xii.2009 ��� 5.i.2010, Bergsten, J. et. al. leg. (slide mounted, NHRS-BYWS000000732). Remarks. The species has so far been known only by type material from two localities in Madagascar (Hippa 2008)., Published as part of Hippa, Heikki, S��li, Geir & Kurina, Olavi, 2019, New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species, pp. 401-441 in Zootaxa 4652 (3) on page 430, DOI: 10.11646/zootaxa.4652.3.1, http://zenodo.org/record/3364902, {"references":["Hippa, H. (2008) Notes on Afrotropical Manota Williston (Diptera: Mycetophilidae), with the description of seven new species. Zootaxa, 1741 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 1741.1.1"]}

Manota pinnata Hippa & Kurina. 2012 Material. D.R. CONGO: male, Oriental Prov, Eyolo forest, ca 2 km E, Lieki, 0.67642, 24.24186, 25–29.v.2010, leg. A.H. Kirk-Spriggs, Malaise traps lowland evergreen swamp forest, Boyekoli Ebale Congo Expedition 2010, BECE 02304 (slide mounted, BMSA). Remarks. The species has so far been known only from the type locality in Uganda (Hippa & Kurina 2012).

Manota mabokeensis Matile, 1972 Material. D.R. CONGO: male, Oriental Prov. Likombo forest, 2km SW Bomane, 1.28349, 23.72358; 20���22.v.2010, A.H. Kirk-Spriggs, Malaise traps lowland evergreen primary forest (disturbed), BMSA(D) 58310 (slide mounted, BMSA). UGANDA: male, Semuliki NP, 0��50���8������N 30��9���46������E, 670 m, Malaise trap, 9���15.ii.2013, F. Molleman et al. leg. (slide mounted, IZBE); male, same data except 15���22.ii.2013 (slide mounted, IZBE); male, Bwindi NP, 01��03���N 29��43���E, Malaise trap (No 1), 10���25.iii.2014, F. Molleman et al. leg. (slide mounted, IZBE). Remarks. The species was described from the Central African Republic (Matile 1972) and subsequently recorded from Uganda and D.R. Congo (Hippa 2008, Hippa & Kurina 2012, Kurina & Hippa 2014)., Published as part of Hippa, Heikki, S��li, Geir & Kurina, Olavi, 2019, New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species, pp. 401-441 in Zootaxa 4652 (3) on page 431, DOI: 10.11646/zootaxa.4652.3.1, http://zenodo.org/record/3364902, {"references":["Matile, L. (1972) Dipteres Mycetophilidae du Cameroun et de Republique Centrafricaine. II. Manotinae. Bulletin de l'Institut fundamental d'Afrique noire, Serie A, 34, 91 - 97.","Hippa, H. (2008) Notes on Afrotropical Manota Williston (Diptera: Mycetophilidae), with the description of seven new species. Zootaxa, 1741 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 1741.1.1","Hippa, H. & Kurina, O. (2012) New species and new records of Afrotropical Manota Williston (Diptera, Mycetophilidae), with a key to the species. Zootaxa, 3455, 1 - 48. https: // doi. org / 10.11646 / zootaxa. 3455.1.1","Kurina, O. & Hippa, H. (2014) The genus Manota Williston (Diptera: Mycetophilidae) in the Congo basin with description of five new species. Zootaxa, 3827 (2), 214 - 230. https: // doi. org / 10.11646 / zootaxa. 3827.2.5"]}

Manota fuscinula sp. n. Figs 3 A–D Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish to light brown. Legs yellowish to light brown. Wing with light brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.6 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.5 times longer than palpomere 4. Ten strong postocular setae. Thorax. Anepisternum with 56 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 20 setae; metepisternum with 11 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 almost extending to level of tip of R 1; wing length 1.6 mm. Hypopygium (Figs 3 A–D). Sternite 9 laterally free from gonocoxa, posterior margin straight, membranous, extending over middle between bases of gonocoxa and gonostylus, anterior margin with deep Vshaped incision medially, anterior 1/3 non-setose, setae on posterior 2/3 similar to those on ventral side of gonocoxa. Ventral medial margin simple, membranous. Parastylar lobe plate-like, rectangular, apical part with 3 setae. Dorsal medial margin of gonocoxa simple, posteriorly forming well delimited setose lobe with dorsal posterior margin of gonocoxa. Ventrally from dorsal posterior margin of gonocoxa a finger-like lobe bearing a strong apical seta. Two juxtagonostylar setae present, one is a rather unmodified seta, the other is a flattened apically pointed megaseta and twice as long as the former, both arising from a common basal body which is about as long as the unmodified seta. Gonostylus elongate, rectangular, apical third slightly bent medially, lateral margin folded dorsally at basal half, setae similar to these on gonocoxa except 2–3 apical setae and 2 setae on lateral margin which are stronger, deviating from others. Aedeagus subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to base of gonostyli, ventral setae distributed only on apical fourth, number of ventral setae (sternite 10) ca. 10 on each half. Cerci medially separated, subapically constricted, with a row of apical setae only. Female. Unknown. Etymology. The name is Latin, fuscinula, ‘small trident’, referring to the three strong setae apically on the gonostylus; noun in apposition. Comments. Manota fuscinula sp. n. belongs to a group of eight Afrotropical species as defined under M. limai. For further comments, see the latter species. Types. Holotype. Male, GHANA, G. Accra region, Legon, Botanical Garden, 1– 3.12.1993, NUFU-ZMBN/ leg. J. Kjaerandsen (slide mounted, NHMO).

Manota leptochaeta sp. n. Figs 7 A–D Male. Colour. Head brown, face somewhat paler. Antenna light brown to brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish. Legs yellowish, basal third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown to dark brown, tergites laterally and sternites lighter.All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.8 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.5 times longer than palpomere 4. Ten strong postocular setae. Thorax. Anepisternum with 69 setae; anterior basalare with 13 setae, laterotergite with 32 setae, preepisternum 2 non-setose, metepisternum with 12 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 2.2 mm. Hypopygium (Figs 7 A–D). Sternite 9 laterally fused to gonocoxa, posterior margin broad, convex, extending to base of gonostylus, anterior margin with a shallow and wide V-shaped concavity medially, anterior half non-setose, posterior half setose, setae similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like, tapering, with a stout and medially curved apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa medially bulging, forming an obtuse lobe with dorsal posterior margin, dorsal surface of gonocoxa anteriorly and laterally setose, lateral setae similar to those on ventral side, anterior setae shorter and placed more densely, medial and posterior parts non-setose except two marginal setae posteromedially. Medioventrally from the dorsal medial margin a thumb-like lobe with a stout megaseta apically (in holotype that megaseta is broken in both sides). One juxtagonostylar seta present, a simple, apically whip-like megaseta arising from a finger-like basal body that is longer than the megaseta. Gonostylus bilobed: subrectangular lateral lobe about half of the gonocoxal height, with 7–8 strong setae along lateral and apical margins; subequal, medially directed and towards apex tapering lobe, with one basal and 2 apical setae along posterior margin and 2 more basal setae on dorsal surface. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over gonostyli, number of ventral setae (sternite 10) ca. 30 on each half. Cerci medially separated, setae delimited on the apical part and medial margin, apical 4 setae somewhat longer, deviating from others. Female. Unknown. Etymology. The name is Latinized from the Greek λ&epsi;πτó&sigmav; (leptos) ‘thin/slender/narrow’ and χαίτη (chaítç) ‘bristle/seta’, referring to the considerably narrow juxtagonostylar megaseta. Comments. Manota leptochaeta sp. n. belongs to a group of 12 Afrotropical species as defined under M. platychaeta. Among these species M. leptochaeta is unique in having one megaseta on a lobe medially at dorsal medial margin of gonocoxa and gonostylus with a prominent medially directed lobe (all other species have at least two megasetae at the dorsal medial margin of gonocoxa or are without megasetae (M. edentula), and gonostylus without a prominent medial lobe). Types. Holotype. Male, MADAGASCAR: Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18°46,25’S 48°25,93’E, 23–25.x. 2014, 960 m, leg. A.H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest (slide mounted, BMSA)

Manota dissidens Hippa & Kurina, 2012 Material. GHANA: 10 males, Central Region, Kakum, Entwikrom, 31.10– 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZBE; 8 in alcohol, NHMO); 27 males, same data except 8– 15.10.1994, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZBE; 25 in alcohol, NHMO). CÔTE D’IVOIRE: 2 males, rég. de Tai, 16.i.-1.ii.1985, G. Gouturier & V. Van Zeijst Réc., BIO (21), (Gilbert), Friche 17 ans, Piége malaise, ORSTOM-Paris, Mission UNESCO (slide mounted, MNHN); male, same data except 1–15.ii.1985 (slide mounted, MNHN); male, same data except 7–11.ii.1985, BIO (25) Forêt noire station (slide mounted, MNHN). Remarks. The species has so far been known only from the type locality in Ghana: Central region, Kakum National Park (Hippa & Kurina 2012).

Manota peltata Kurina & Hippa, 2014 Material. GHANA: male, Central Region, Kakum, Entwikrom, 31.10– 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, NHMO). Remarks. The species has so far been known only from the holotype collected from Yangambi National Park in D.R. Congo (Kurina & Hippa 2014).

Manota relicina Hippa & Kurina, 2012 Material. UGANDA: mal e, Bwindi NP, 01°03’N 29°43’E, Malaise trap (No 2), 10–25.iii.2014, F. Molleman et al. leg. (slide mounted, IZBE). Remarks. The species has so far been known only from the type locality in Uganda (Hippa & Kurina 2012).

Manota reclinata Kurina & Hippa, 2014 Figs 14 A–D Material. GHANA: 4 males, Volta region, Wli Falls, Afegame (St. 10A), 3– 7.03.1993, NUFU-ZMBN, leg. J.S. Amakye & J. Kjaerandsen (slide mounted, IZBE and NHMO); male, same data except 4– 7.03.1993 (slide mounted, NHMO); male, same data except 6– 9.03.1993 (in alcohol, NHMO); 2 males, same data except 9– 12.03.1993 (in alcohol, NHMO); 2 males, same data except 7– 10.03.1993 (1 slide mounted, IZBE; 1 in alcohol, NHMO); 4 males, same data except 10– 13.03.1993 (3 slide mounted, NHMO; 1 in alcohol, IZBE); 2 males, same data except 12– 15.03.1993 (slide mounted, NHMO); male, same data except 11.03.1993, light trap (in alcohol, NHMO); 2 males, Volta region, Wli Falls, Agumatsa (St. 10), 11– 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, Central Region, Kakum, Entwikrom, 31.10– 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE); 3 males, same data except 8– 15.10.1994, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZB; 1 in alcohol, NHMO); 2 males, Volta region, Wli Falls, 4– 13.03.1993, leg. J. Kjaerandsen (1 slide mounted, NHMO; 1 in alcohol, IZBE). CÔTE D’IVOIRE: male, rég. de Tai, 1–15.ii.1985, G. Gouturier & V. Van Zeijst Réc., BIO (24), Friche 6 ans, Piège malaise // ORSTOM - Paris, Mission UNESCO (slide mounted, MNHN); male, same data except 7–11.ii.1985, BIO (25) Forêt noire station (slide mounted, MNHN). GUINEA: 3 males, Ft. Gal. du Zougué, 750m, 18–29.vi.1991, Piège de Malaise, Girard et Legrand col. (2 slide mounted, IZBE and MNHN; 1 pinned, MNHN); 2 males, same data, except 3–17.vi.1991 (pinned, MNHN). Remarks. The species is described from the male holotype collected from Kona on Itimbti river in D.R. Congo and not recorded since (Kurina & Hippa 2014). The study of more extensive material disclosed a small variation in male terminalia (Figs 14 A–D), especially in details of the gonostylus and the setigerous lobe dorsally from juxtagonostylar megasetae, which are well exposed in the slide mounts of the newly studied specimens.

Manota geniculata sp. n. Figs 4 A–D Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish, apical fourth of hind femur slightly infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with dark brown knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 not measurable in holotype. Ten strong postocular setae. Thorax. Anepisternum with 43 setae; anterior basalare with 5 setae, laterotergite with 21 setae, preepisternum 2 non-setose; metepisternum with 15 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 4 A–D). Sternite 9 laterally fused to gonocoxa, posterior margin semicircular, tongue-like, extending to base gonostylus, anterior margin with a shallow concavity, anterior 1/3 non-setose, posterior 2/3 setose, setae similar to those on ventral side of gonocoxa but arranged more densely. Ventral medial margin of gonocoxa oblique. Parastylar lobe plate-like, medially drawn out to a finger-like lobe that has an apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa complex, posteriorly with a row of five flattened and apically pointed megasetae at medial margin and a single equal megaseta at posterolateral corner. At the dorsal medial margin of gonocoxa, on more ventral level a plate-lake submembranous lobe with two medially directed simple and blunt megasetae at anteromedial corner, and a posteriorly directed geniculate megasetae arising from a finger-like basal body. Gonocoxal apodeme large, polygonal. One juxtagonostylar seta present as a flattened and apically expanded megaseta arising from a curved basal body which is about five times longer than the megaseta. Gonostylus slightly less than half of the ventral length of gonocoxa, medially geniculate, basal half twice as wide as apical half, basal half ventrally setose, dorsally non-setose except for two strong setae, apical half with two strong setae at basal portion of medial margin and 4–5 apical setae, 2–3 of them stronger, deviating from others. Aedeagus elongated subtriangular, without lateral shoulders, apex curved ventrad. In holotype, aedeagal apodemes turned upward, pressed against the basal part of aedeagus. Hypoproct large, extending posteriorly near to apex of gonostylus, number of ventral setae (sternite 10) ca. 50 on each half, the setae indistinctly divided into an anterior and posterior groups with ca. 45 and 5 setae, respectively. Cerci medially separated, setae scattered over the surface, apical 4–5 setae about twice as long as the rest of cercal setae. Female. Unknown. Etymology. The name is Latin, geniculata, ‘like the bent knee’, referring to the geniculate gonostylus. Comments. Manota geniculata sp. n. belongs to a group of six Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) nonsetose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly with a row of scale- or leaf-like megasetae. The other species being M. bracteata, M. foliolata, M. peltigera, M. pinnata and M. pinnulata. Manota geniculata, M. peltigera and M. pinnulata all have 3 flattened megasetae at the dorsal medial margin of gonocoxa posteriorly and a tongue-like posterior part of sternite 9. The latter character is shared also with M. foliolata, but that species has only two flattened megasetae posteriorly at the dorsal medial margin of gonocoxa. Of these three species, M. geniculata is most similar to M. pinnulata (cf. Hippa & Kurina 2012: fig. 18) but differs in having the gonostylus geniculate (subtriangular, widening from base to apex in M. pinnulata), and in having the megasetae posteriorly at the dorsal medial margin of gonocoxa comma-like, apically pointed, and becoming medially wider towards posterior end of the row (megasetae scale-like, apically widening and enlarging in size towards posterior end of the row in M. pinnulata). Types. Holotype. Male, GABON, Makoku M’Passa Bale affl., 200 m, J. Legrand rec., 28.05– 5.06. 1979, Piège de Malaise (slide mounted, MNHN).

Manota whiteleyi Jaschhof & Mostovski, 2006 Material. GHANA: 3 males, Central region, Kakum, Entwikrom 31.10��� 8.11.1994, NUFU-project ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE and NHMO). CAMEROON: male, North Reg. Vina, Tchabal village, 07��35,195���N 13��33,566���E; 13.viii.2013, 1362 m, A.H. Kirk-Spriggs; Malaise traps degraded savanna hill forest, BMSA(D) 54969 (slide mounted, BMSA). Remarks. The species was described from KwaZulu-Natal, South Africa, but was recorded also from C��te d���Ivoire (Hippa 2008). The studied specimens are more similar to the specimen figured from C��te d���Ivoire (Hippa 2008: fig. 12B) because the dorsal medial margin of the gonocoxa is only slightly lobe-like produced posteriorly, while it is a more prominent lobe in South African specimens (cf. Jaschhof & Mostovski 2006: fig. 5)., Published as part of Hippa, Heikki, S��li, Geir & Kurina, Olavi, 2019, New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species, pp. 401-441 in Zootaxa 4652 (3) on page 437, DOI: 10.11646/zootaxa.4652.3.1, http://zenodo.org/record/3364902, {"references":["Jaschhof, M. & Mostovski, M. (2006) First record of Manota (Diptera: Mycetophilidae: Manotinae) from southern Africa, with description of two new species. African Invertebrates, 47, 237 - 242.","Hippa, H. (2008) Notes on Afrotropical Manota Williston (Diptera: Mycetophilidae), with the description of seven new species. Zootaxa, 1741 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 1741.1.1"]}

Manota oronnai sp. n. Figs 9 A–C Male. Colour. Head light brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish to light brown. Legs yellowish to light brown, apical third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.3 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment. Nine strong postocular setae. Thorax. Anepisternum with 65 setae; anterior basalare and laterotergite nonsetose; preepisternum 2 with 13 setae; metepisternum with 9 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 9 A–C). Sternite 9 laterally free from gonocoxa, posterior and anterior margins both with deep V-shaped incision medially, extending about middle between bases of gonocoxa and gonostylus, anterior half non-setose, setae on posterior half somewhat shorter than those on ventral side of gonocoxa. Ventral medial margin of gonocoxa angular. Parastylar lobe large, plate-like, triangular, with two setae at medial angle. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa simple, convex. Dorsal posterior margin simple, forming together with medial margin a right angled lobe with a dense aggregation of fine setae on medial side. On ventral side of dorsal posterior margin of gonocoxa an elongated, subrectangular lobe, about twice as long as wide, with aggregation of fine setae at apex. One juxtagonostylar setae present, a simple, apically vasculiform megaseta, arising from a basal body, which is longer than the megaseta. Gonostylus subrectangular, apically transverse, with a very long seta deviating from other setosity at apicolateral corner, and with a short finger-like lobe subapically on medial side bearing two weak and simple apical megasetae. Aedeagus subtriangular, with lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over basal part of gonostylus, the ventral setae (sternite 10) strong and forming a mesial longitudinal stripe of 5–6 setae on each half. Cerci medially separated, setae delimited on the apical and medial margins of cercus. Female. Unknown. Etymology. The species is named after the mythical hero Oronna who defended the town Ilaro when it got under the threat of invasion by the Dahomean Army in the 17 th century. Comments. Because of the non-setose laterotergite, setose preepisternum 2 and gonocoxa posterodorsally with an apical setae bearing lobe, the new species groups together with 16 other Afrotropical species (see couplets 36 to 52 in the Key of the Afrotropical species above). Among them, M. oronnai sp. n. is unique as the lobe posterodorsally from the gonocoxa is subrectangular and with an aggregation of fine setae apically while all other species have this lobe finger-like and with 1–3 strong setae apically. In addition, the new species has a single apically vasculiform juxtagonostylar megaseta, while all other species have two juxtagonostylar setae, which are apically pointed. Types. Holotype. Male, W. NIGERIA, Ilaro Forest, 11.xi.1973, M.A. Cornes (slide mounted, MNHN).

Manota cornuta sp. n. Figs 2 A–C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown. Legs yellowish to light brown, apical fifth of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 1.4–1.5 times longer than palpomere 4. 10-11 strong postocular setae. Thorax. Anepisternum with 52 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 12–15 setae; metepisternum with 14–19 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 almost extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 2 A–C). Sternite 9 laterally free from gonocoxa, posterior margin membranous, slightly concave, extending over middle between bases of gonocoxa and gonostylus, anterior margin with deep and wide V-shaped incision medially, anterior 1/3 and posterior membranous part nonsetose, setae on posterior 2/3 slightly shorter than those on ventral side of gonocoxa, postero-laterally with fingerlike sclerotizations on dorsal surface of both sides, which bear one short and curved apical seta deviating from other setosity on sternite 9. Ventral medial margin simple, membranous. Parastylar lobe plate-like, subrectangular, medially constricted, apically with 3 short and curved setae. Dorsal medial margin of gonocoxa simple, posteriorly forming well delimited setose bill-like small lobe with dorsal posterior margin of gonocoxa. Three juxtagonostylar setae present, all megaseta-like, the dorsal one apically slightly curved, with an apically separate basal body, the two dorsal ones more prominent, apically hook-like curved, with a common basal body which is more than half of the length of megasetae. Gonostylus dorso-ventrally wide, lateroapically, between dorsal and ventral portions concave; dorsal portion narrow, tapering with basal heel-like lobe, subbasal setose hump and 2–3 more prominent setae dorsoapically; ventral portion wide, with two blunt, medially directed megasetae apically and two more prominent setae subapically. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct posteriorly not extending to the basal part of gonostylus, ventral setae (sternite 10) forming a medial longitudinal stripe of ca. 8 setae on each half. Cerci (scarcely discernible on slides) medially separated, largely membranous, with unusually narrow apical parts, bearing 4–5 setae. Female. Unknown. Etymology. The name is Latin, cornuta, ‘horned’, referring to the horn-like setigerous lobe at posterolateral corners of sternite 9. Comments. Manota cornuta sp. n. is similar to M. afra and the two key out in the same couplet in the species key. Both species have three juxtagonostylar megasetae that is unique among the Afrotropical species. The unique finger-like posterolateral lobes on sternite 9 and the unusually narrow apical part of cercus, distinguishes M. cornuta from all known Manota. Types. Holotype. Male, GHANA, Volta region, Wli Falls, Afegame (St. 10A), 7– 10.03.1993, NUFU-ZMBN, leg. J.S. Amakye & J. Kjaerandsen (slide mounted, NHMO). Paratypes. Male, GHANA, Volta region, Wli Falls, Afegame (St. 8A), 11– 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO).

Manota foliolata Hippa & Kurina, 2012 Material. CAMEROON: male, Northwest Reg. Mezam, Bafut village at: 06°05,026’N 10°07,442’E, 17– 19.viii.2013, 1060 m, leg. A.H. Kirk-Spriggs, Malaise traps cultivated plots & degraded forest, BMSA(D) 58256 (slide mounted, BMSA). Remarks. The species was described from Uganda (Hippa & Kurina 2012) and has not been recorded since.

Manota burundiensis sp. n. Figs 1 A–C Male. Colour. Head brown, face somewhat paler. Antenna brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish to light brown, apical half of hind femur, and apical and basal thirds of mid femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment. Nine strong postocular setae. Thorax. Anepisternum with 68 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 25 setae; metepisternum with 17 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 2.3 mm. Hypopygium (Figs 1 A–C). Sternite 9 narrow, about two times as long as wide, posterior ¾ laterally free from gonocoxa, anterior ¼ fused with gonocoxa, posterior margin with deep V-shaped incision medially, extending over middle between bases of gonocoxa and gonostylus, anterior margin with a deep V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa angular. Parastylar lobe large, plate-like, medially drawn out to a finger like process, which has two strong setae apically. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa simple, slightly bulging medially. Dorsal posterior margin simple, forming together with medial margin an acute angle with a dense aggregation of fine setae on medial side. On ventral side of dorsal posterior margin of gonocoxa a finger-like lobe bearing two strong apical setae. Two juxtagonostylar setae present, a normal seta and apically curved simple megaseta, both arising from a common basal body, which is about as long as the megaseta. Gonostylus subcircular, setose, setae similar to those on gonocoxa except for conspicuously long setae along apical margin; apicomedially with a small lobe bearing two dorsally turned megasetae at apex. Aedeagus subtriangular, with lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over basal part of gonostylus, the ventral setae strong and forming a mesial longitudinal stripe of 7–8 setae on each half. Cerci medially separated; setae delimited on the apical half of cercus. Female. Unknown. Etymology. The species is named after its occurrence in the state of Burundi by adding the Latin suffix –ensis [denoting place]. Comments. Manota burundiensis sp. n. belongs to a group of six Afrotropical species as defined under M. triseta. Among them, M. burundiensis is most similar to M. relicina (see Hippa & Kurina 2012: fig. 20) as both species have the cercal setae short and delimited on the apical half of cercus, and the gonostylus with an apical or subapical medial lobe bearing two simple megasetae. It differs in having (1) one juxtagonostylar seta as a normal seta (both juxtagonostylar setae are megasetae in M. relicina), (2) the common basal body of the juxtagonostylar setae is about as long as setae (the basal body is considerably longer than juxtagonostylar megasetae in M. relicina), (3) parastylar lobe with two unusually strong setae at medial corner (with normal setae in M. relicina), (4) gonostylus apically rounded (apically transverse in M. relicina), and (5) dorsal posterior margin of the gonocoxa forming together with medial margin an acute angle with a dense aggregation of fine setae on medial side (with a row of closely placed setae in M. relicina). Types. Holotype. Male, BURUNDI, Kayanza Prov., Parc National de la Kibira, Rwecura sector, Indigenous Afromontane forest, 02°55.320’S 29°30.067’E, Malaise trap, 21–26.xi.2010, A.H. Kirk-Spriggs leg. (slide mounted, BMSA).

Allodia keurbosensis Magnussen sp. nov. (FIgS 5, 6) Diagnostic characters. A. keurbosensis CAN bEST bE SEPARATED fROM THE OTHER SPECIES DESCRIbED HERE, bASED ON THE fOLLOWINg COMbINATION Of CHARACTERS: RM ALMOST TWICE AS LONg AS STEM Of THE POSTERIOR fORK; THE DORSAL LObE Of gONOSTyLUS IS SIMILAR TO A. nyeriensis, A. mazumbaiensis AND A. drakensbergensis; THE MEDIAN LObE IS MOST SIMILAR TO A. drakensbergensis, bUT WITH MORE SETAE ON THE VENTRAL MARgIN (FIg. 10); THE SHAPE Of THE VENTRAL LObE IS DIffERENT TO THAT Of A. drakensbergensis (FIg. 4B); THE bASAL PART Of gONOSTyLUS DIffERENT fROM ALL THE OTHER SPECIES (FIg. 6C). Type material. HOLOTyPE &male;. SOUTH AfRICA: EASTERN CAPE, KEURbOS fOREST (33.9072°S, 23.7285°E, 500 MASL), MALAISE TRAP, INDIgENOUS MONTANE fOREST. 28–30 MAR. 2009. LEg. A.H. KIRK-SPRIggS; S. OTTO. (BMSA). Measurements. MALE: BODy LENgTH 3.91 MM; WINg LENgTH 3.23 MM. Coloration. HEAD AND CLyPEUS bROWN. MOUTHPARTS AND PALPOMERES yELLOW. ANTENNAE bROWN, WITH SCAPE, PEDICEL AND fIRST HALf Of fIRST fLAgELLOMERE yELLOW. SCUTUM bROWN. ANTEPRONOTUM WHITISH ANTERIORLy, LATERAL SCLERITES OTHERWISE bROWN. HALTERES WHITISH. LEgS WHITISH. WINgS CLEAR WITHOUT MARKINgS. AbDOMEN bROWN, WITH SMALL yELLOW TRIANgULAR AREA ON TERgITES 3 AND 4. TERMINALIA yELLOW. Head. TWO OCELLI, LOCATED CLOSE TO EyE MARgIN. HEAD COVERED WITH fINE TRICHIA, EXCEPT fOR ROW Of AbOUT 8 SHORT bRISTLES NEAR EyE MARgIN, AbOVE OCELLUS. ANTENNAE AbOUT TWICE AS LONg AS THORAX. SCAPE AND PEDICEL WITH SEVERAL SETAE ON DISTAL THIRD. FLAgELLOMERES CyLINDRICAL, DENSELy CLOTHED WITH fINE TRICHIA. FIRST fLAgELLOMERE TWICE AS LONg AS SCAPE. Thorax. ANTEPRONOTUM WITH 4 STRONg SETAE. SCUTUM COVERED WITH UNIfORM SMALL, PALE SETAE, WITH STRONg LATERAL PREALAR AND POSTALAR SETAE. DISCAL bRISTLES AbSENT. SCUTELLUM WITH 3 STRONg bRISTLES. LATEROTERgITE WITH 3 LONg SETAE. OTHER LATERAL SCLERITES bARE. Legs. FORE-, MID AND HIND TIbIAE WITH SHORT SETAE ARRANgED IN ROWS. MID TIbIA WITH 4 ANTERODORSAL AND 20 POSTERODORSAL bRISTLES, ON DISTAL 3/4 Of SEgMENT. HIND TIbIA WITH 6 ANTERODORSAL AND 6 POSTERODORSAL bRISTLES. Wings. SC SHORT, ENDINg IN R, RM ALMOST TWICE AS LONg AS STEM Of POSTERIOR fORK, bASE Of ANTERIOR fORK JUST OPPOSITE bASE Of POSTERIOR fORK. R1 AND R5 SETAE, AbSENT AT bASIS Of VEINS. Male terminalia (FIg. 6). TERgITE 9 MEDIALLy DIVIDED, EACH PART ROUNDED, COVERED WITH MINUTE TRICHIA, WITH TWO STRONg bRISTLES AND THREE LONg SETAE. CERCI CLOTHED WITH fINE TRICHIA, WITH SOME LONg SETAE APICALLy; PSEUDOCERCI WITH SEVERAL LONg SETAE RANDOMLy MEDIALLy AND APICALLy. DORSAL LObE Of gONOSTyLUS ObLONg, APEX ROUNDED WITH SMALL, POUCH-LIKE MEMbRANOUS AREA INTERNALLy, OUTER SURfACE WITH NUMEROUS STRONg SETAE. MEDIAN LObE WITH STRAIgHT VENTRAL SIDE, WITH ROW Of 7 STRONg SETAE ON POSTERIOR MARgIN; 10 SHORTER PRESENT ON POSTERIOR MARgIN AND ON LObE SURfACE; THREE SETAE ON VENTRAL SIDE, TWO CLOSE TO POSTEROVENTRAL CORNER, ONE AbOUT ONE THIRD Of THE TOTAL DISTANCE fROM POSTERIOR EDgE. VENTRAL LObE SLIgHTLy CLUb-SHAPED WITH 6-7 SETAE APICALLy, PLUS ONE ON POSTERIOR EDgE. BASAL PART Of gONOSTyLUS WITH TWO bULbOUS STRUCTURES Of EQUAL SIZE, EACH WITH 4 SHORT SETAE. HyPANDRIAL LObE ELONgATED, POINTED INWARDS, WITH STRONg SCLEROTIZED APODEMES (FIg. 6A). Etymology. NAMED AfTER THE TyPE LOCALITy Remarks. MEASUREMENTS WERE MADE ON PINNED SPECIMEN.

Allodia nyeriensis Magnussensp. nov. (FIgS 1, 2) Diagnostic characters. A. nyeriensis CAN bEST bE SEPARATED fROM THE OTHER SPECIES DESCRIbED HERE bASED ON THE fOLLOWINg COMbINATION Of CHARACTERS: RM IS ALMOST TWICE AS LONg AS THE STEM Of POSTERIOR fORK; THE DORSAL LObE Of THE gONOSTyLUS HAVE A SMALL INCONSPICUOUS AREA AT APEX, AS IN A. mazumbaiensis, A. drakensbergensis AND A. keurbosensis; THE MEDIAN LObE IS SIMILAR TO A. keurbosensis, bUT WITH MORE SETAE ON THE VENTRAL MARgIN AND A WELL- DEfINED HEEL-SHAPED POSTEROVENTRAL CORNER Of THE LObE (FIg. 10); THE bASAL PART Of THE gONOSTyLUS IS DIffERENT fROM ALL THE OTHER SPECIES (FIg. 1G). THE SHAPE Of THE fEMALE HyPOgyNAL VALVES (FIg. 2) DIffERS fROM A. jaschhofi (FIg. 8). Type material. HOLOTyPE &male;. KENyA, NyERI, MOUNT KENyA NATIONAL PARK, BASE CAMP AT NARO MORU RIVER LODgE (0.17027��S, 37.21500��E, 3050 MASL), SWEEP NET, bAMbOO fOREST. 18���19 AUg. 2008. TSZD-JKJ-104239. LEg. J. KJaeRANDSEN. (TMU). PARATyPES 1 &male;, 2 &female;, DATA AS fOR HOLOTyPE. (TMU). Measurements. MALE (N = 2): BODy LENgTH 3.04���3.91 MM; WINg LENgTH 2.91���3.26 MM. FEMALE (N = 2): BODy LENgTH 4.00���4.35 MM; WINg LENgTH 3.33 MM Coloration. HEAD AND CLyPEUS bROWN. MOUTHPARTS AND PALPOMERES WHITISH yELLOW. ANTENNAE bROWN, WITH SCAPE, PEDICEL AND bASAL HALf Of fIRST fLAgELLOMERE WHITISH yELLOW. SCUTUM bROWN, WITH yELLOW LATERAL MARgIN. LATERAL SCLERITES bROWN. WINgS CLEAR WITHOUT MARKINgS. HALTERES WHITISHyELLOW. LEgS WHITISH yELLOW. AbDOMEN bROWN, TERgITES 2���4, 2���5 IN fEMALES, WITH LATERAL AREA yELLOW, bROADENINg TOWARDS POSTERIOR MARgIN. TERMINALIA yELLOW. Head. TWO OCELLI, NEAR EyE MARgIN. HEAD COVERED WITH fINE TRICHIA, EXCEPT fOR ROW Of 5 SHORT bRISTLES NEAR EyE MARgIN, AbOVE OCELLUS. ANTENNAE IN MALES AbOUT TWICE AS LONg AS THORAX, IN fEMALES SLIgHTLy LONgER THAN THORAX (1.13 MM/1.09 MM, RESPECTIVELy). SCAPE AND PEDICEL WITH SEVERAL SETAE ON DISTAL THIRD. FLAgELLOMERES CyLINDRICAL, DENSELy COVERED WITH fINE TRICHIA. FIRST fLAgELLOMERE TWICE AS LONg AS SCAPE. Thorax. ANTEPRONOTUM WITH 5���6 STRONg SETAE. SCUTUM COVERED WITH UNIfORM SMALL, PALE SETAE, WITH STRONg LATERAL PREALAR AND POSTALAR SETAE, MORE NUMEROUS CLOSE TO WINg bASE. DISCAL bRISTLES AbSENT. SCUTELLUM WITH 2 STRONg bRISTLES. LATEROTERgITE WITH 2 LONg AND 3 SHORTER SETAE. OTHER LATERAL SCLERITES bARE. Legs. ALL TIbIAE WITH SHORT SETAE ARRANgED IN ROWS. MID TIbIA WITH 5���7 ANTERODORSAL AND 25���28 POSTERODORSAL bRISTLES, ON DISTAL 3/4 Of SEgMENT. HIND TIbIA WITH 5���7 ANTERODORSAL AND 8 POSTERODORSAL bRISTLES. Wings. SC SHORT, ENDINg IN R, RM ALMOST TWICE AS LONg AS STEM Of POSTERIOR fORK, bASE Of ANTERIOR fORK SLIgHTLy bEfORE bASE Of POSTERIOR fORK. R1 WITH SETAE ON DISTAL 2/3 AND R5 WITH SETAE ON DISTAL 3/4. Male terminalia (FIg. 1). TERgITE 9 MEDIALLy DIVIDED, EACH PART ROUNDED, COVERED WITH MINUTE TRICHIA, WITH ONE STRONg APICAL bRISTLE, TWO SHORT AND THREE LONgER SETAE. CERCI COVERED WITH fINE TRICHIA, WITH LONgER SETAE APICALLy; PSEUDOCERCI WITH SEVERAL LONg SETAE MEDIALLy AND APICALLy (FIg. 1F). DORSAL LObE Of gONOSTyLUS ObLONg, APEX ROUNDED WITH SMALL, ROUNDISH, POUCH-LIKE MEMbRANOUS STRUCTURE INTERNALLy; OUTER SURfACE WITH NUMEROUS STRONg SETAE. MEDIAN LObE WITH HEEL-SHAPED POSTEROVENTRAL CORNER; 9 STRONg AND 4 SHORTER SETAE ON POSTERIOR MARgIN, PLUS 4 MINUTE SETAE ON INTERIOR SURfACE; ONE LONg SETA ON VENTRAL EDgE, WITH SHORT DISTANCE fROM VENTRAL MARgIN (AbOUT 1/6 Of TOTAL DISTANCE). VENTRAL LObE ELONgATED, SLIgHTLy ARCHED, WITH 5���6 SETAE APICALLy. BASAL PART Of gONOSTyLUS WITH VENTRAL SEMI-CIRCULAR STRUCTURE, bEARINg AbOUT 8 SHORT SETAE, AND SECOND SMALL POINTED DORSAL STRUCTURE WITH ONE SHORT SETA (FIg. 1G). HyPANDRIAL LObE HEAVILy SCLEROTIZED, SLENDER, PROTRUDINg INWARDS AT APEX (FIg. 1E). Female terminalia (FIg. 2). TERgITE 8 ELONgATED. CERCI TWO-SEgMENTED; fIRST SEgMENT ObLONg, ARCUATE, fUSED AT bASIS; SECOND SEgMENT ObLONg, NARROWER THAN fIRST SEgMENT, WITH SEVERAL LONg SETAE. GONAPOPHySES fUSED, MEMbRANOUS, TONgUE-SHAPED WITH SMALL SETAE AROUND EDgE. HyPOgyNAL VALVES ELONgATED, TRIANgULAR SHAPED, POINTED TOWARDS APEX; ONE LONg SETA ON TIP; LAbIA MEMbRANOUS. Etymology. NAMED AfTER THE REgION Of THE TyPE LOCALITy. Remarks. THE SPECIMENS HAVE bECOME PALE AfTER yEARS Of STORAgE IN ETHANOL. THE MEASUREMENTS WERE MADE ON ETHANOL PRESERVED SPECIMENS. WINg LENgTH fOR fEMALES WAS MEASURED ON ONE SPECIMEN ONLy., Published as part of Magnussen, Trude, Kjaerandsen, Jostein, Johnsen, Arild & S��li, Geir E. E., 2018, Six new species of Afrotropical Allodia (Diptera: Mycetophilidae): DNA barcodes indicate recent diversification with a single origin, pp. 301-320 in Zootaxa 4407 (3) on pages 304-306, DOI: 10.11646/zootaxa.4407.3.1, http://zenodo.org/record/1216502

Allodia mazumbaiensis Magnussen sp. nov. (FIg. 3) Diagnostic characters. A. mazumbaiensis CAN bEST bE SEPARATED fROM THE OTHER SPECIES DESCRIbED HERE, bASED ON THE fOLLOWINg COMbINATION Of CHARACTERS: RM AS LONg AS STEM Of THE POSTERIOR fORK; THE DORSAL LObE Of THE gONOSTyLUS HAVE A SMALL INCONSPICUOUS AREA AT APEX, AS IN A. nyeriensis, A. drakensbergensis AND A. keurbosensis; THE MEDIAN LObE IS NARROWER THAN IN THE OTHER SPECIES, WITH fEWER SETAE ON THE VENTRAL MARgIN AND ONE ESPECIALLy LONg AND PRONOUNCED ON THE DORSAL TIP (FIg. 10); bASAL PART Of gONOSTyLUS DIffERS fROM ALL THE OTHER SPECIES (FIg. 3C). Type material. HOLOTyPE &male;. TANZANIA: TANgA REgION, WEST USAMbARA MOUNTAINS, MAZUMbAI, 1500 MASL. MALAISE TRAP, LOC. O., ZBM’S TANZANIA EXPEDITION. 29 SEP. –3 DES. 1990. LEg. G. SøLI. (NHMO). Measurements. MALE: BODy LENgTH 4.44 MM; WINg LENgTH 3.33 MM. Coloration. HEAD AND CLyPEUS bROWN. MOUTHPARTS AND PALPOMERES WHITISH yELLOW. ANTENNAE bROWN, WITH SCAPE, PEDICEL AND fIRST HALf Of fIRST fLAgELLOMERE WHITISH yELLOW. SCUTUM bROWN, WITH yELLOW LATERAL MARgIN. ANTEPRONOTUM WHITISH, LATERAL SCLERITES OTHERWISE bROWN. WINgS CLEAR WITHOUT MARKINgS. HALTERES WHITISH yELLOW. LEgS WHITISH, WITH bROWN bASAL AREA VENTRAL ON fEMUR. AbDOMEN bROWN, TERgITES 2–4 WITH LATERAL AREA WHITISH yELLOW. TERMINALIA yELLOW. Head. TWO OCELLI, LOCATED CLOSE TO EyE MARgIN. HEAD COVERED WITH fINE TRICHIA, EXCEPT fOR ROW Of 5 SHORT bRISTLES NEAR EyE MARgIN, AbOVE OCELLUS. ANTENNAE AbOUT TWICE AS LONg AS THORAX. SCAPE AND PEDICEL WITH SEVERAL SETAE ON DISTAL THIRD. FLAgELLOMERES CyLINDRICAL, DENSELy CLOTHED WITH fINE TRICHIA. FIRST fLAgELLOMERE TWICE AS LONg AS SCAPE. Thorax. ANTEPRONOTUM WITH 5 STRONg SETAE. SCUTUM COVERED WITH UNIfORM SMALL, PALE SETAE, WITH STRONg LATERAL PREALAR AND POSTALAR SETAE. DISCAL bRISTLES AbSENT. SCUTELLUM WITH 2 STRONg bRISTLES. LATEROTERgITE WITH 2 LONg AND 4 MINUTE SETAE. OTHER LATERAL SCLERITES bARE. Legs. FORE-, MID AND HIND TIbIA WITH SHORT SETAE ARRANgED IN ROWS. MID TIbIA WITH 14 ANTERODORSAL AND 26 POSTERODORSAL bRISTLES, ON DISTAL 3/4 Of SEgMENT. HIND TIbIA WITH 8 ANTERODORSAL AND 9 POSTERODORSAL bRISTLES. Wings. SC SHORT, ENDINg IN R, LENgTH Of RM EQUAL TO STEM Of POSTERIOR fORK, bASE Of ANTERIOR fORK SLIgHTLy bEfORE bASE Of POSTERIOR fORK. R1 AND R5 WITH SETAE. Male terminalia (FIg. 3). TERgITE 9 MEDIALLy DIVIDED, EACH PART ROUNDED, COVERED WITH MINUTE TRICHIA, WITH TWO STRONg APICAL bRISTLES AND THREE LONg SETAE. CERCI CLOTHED WITH fINE TRICHIA, WITH LONgER SETAE APICALLy; PSEUDOCERCI WITH SEVERAL LONg SETAE (FIg. 3A). DORSAL LObE Of gONOSTyLUS ObLONg, APEX ROUNDED WITH SMALL, INCONSPICUOUS, MEMbRANOUS AREA INTERNALLy; OUTER SURfACE WITH NUMEROUS STRONg SETAE. MEDIAN LObE SLENDER, VENTRAL SIDE STRAIgHT; WITH 4 STRONg AND 4 THIN SETAE ON POSTERIOR MARgIN, APICAL SETA ESPECIALLy PROMINENT; 2-3 SMALL SETAE ON LObE SURfACE; ONE LONg SETA ON VENTRAL SIDE, LOCATED WITH APPROXIMATELy EQUAL DISTANCE fROM TIP AND bASIS Of LObE. VENTRAL LObE SLENDER, SLIgHTLy ARCHED, WITH 5-6 SETAE APICALLy. BASAL PART Of gONOSTyLUS WITH TWO STRUCTURES Of EQUAL SIZE, bOTH bULbOUS WITH 5-6 SETAE (FIg. 3C). HyPANDRIAL LObE COMPLEX, ELONgATED, CURVED INWARDS (FIg. 3A) Etymology. NAMED AfTER THE TyPE LOCALITy. Remarks. THE SPECIMEN HAS bECOME PALE AfTER yEARS Of STORAgE IN gLyCEROL. MEASUREMENTS WERE MADE ON SPECIMEN IN ETHANOL.

Allodia drakensbergensis Magnussen sp. nov. (FIg. 4) Diagnostic characters. A. drakensbergensis CAN bEST bE SEPARATED fROM THE OTHER SPECIES DESCRIbED HERE, bASED ON THE fOLLOWINg COMbINATION Of CHARACTERS: RM AbOUT 1.5 TIMES AS LONg AS STEM Of THE POSTERIOR fORK; THE DORSAL LObE Of gONOSTyLUS IS SIMILAR TO A. nyeriensis, A. mazumbaiensis AND A. keurbosensis; THE MEDIAN LObE MOST SIMILAR TO A. keurbosensis, bUT WITH fEWER SETAE ON THE VENTRAL MARgIN AND THE SHAPE Of VENTRAL LObE DIffERS fROM A. keurbosensis (FIg. 10); THE bASAL PART Of THE gONOSTyLUS IS DIffERENT fROM ALL THE OTHER SPECIES (FIg. 4C). Type material. HOLOTyPE &male;. SOUTH AfRICA: NORTHERN DRAKENSbERg, KWAZULU-NATAL, ROyAL NATAL NATIONAL PARK, GUDU fOREST (28.6691°S, 28.9188°E, 1630–1730 MASL), SWEEP NET, OLD-gROWTH INDIgENOUS fOREST. 29 NOV. 2005. TSZD-JKJ-104243. LEg. M. JASCHHOf. (TMU). Measurements. MALE: bODy LENgTH 4.44 MM; WINg LENgTH 3.33 MM. Coloration. HEAD AND CLyPEUS bROWN. MOUTHPARTS AND PALPOMERES WHITISH yELLOW. ANTENNAE bROWN, WITH SCAPE, PEDICEL AND fIRST HALf Of fIRST fLAgELLOMERE WHITISH yELLOW. SCUTUM bROWN. LATERAL SCLERITES bROWN. WINgS CLEAR WITHOUT MARKINgS. HALTERES WHITISH AT bASIS, bROWN ON CLUb. LEgS WHITISH. AbDOMEN bROWN. TERMINALIA yELLOWISH bROWN. Head. TWO OCELLI, LOCATED CLOSE TO EyE MARgIN. HEAD COVERED WITH fINE TRICHIA, EXCEPT fOR ROW Of AbOUT 7 SHORT bRISTLES NEAR EyE MARgIN, AbOVE OCELLUS. ANTENNAE JUST OVER TWICE AS LONg AS THORAX. SCAPE AND PEDICEL WITH SEVERAL SETAE ON DISTAL THIRD. FLAgELLOMERES CyLINDRICAL, DENSELy CLOTHED WITH fINE TRICHIA. FIRST fLAgELLOMERE TWICE AS LONg AS SCAPE. Thorax. ANTEPRONOTUM WITH 5 STRONg SETAE. SCUTUM COVERED WITH UNIfORM SMALL, PALE SETAE, WITH STRONg LATERAL PREALAR AND POSTALAR SETAE. DISCAL bRISTLES AbSENT. SCUTELLUM WITHOUT STRONg bRISTLES. LATEROTERgITE WITH 3 LONg AND 3 MINUTE SETAE. OTHER LATERAL SCLERITES bARE. Legs. FORE-, MID AND HIND TIbIA WITH SHORT SETAE ARRANgED IN ROWS. MID TIbIA WITH 14 ANTERODORSAL AND 26 POSTERODORSAL bRISTLES ON DISTAL 3/4 Of SEgMENT. HIND TIbIA WITH 8 ANTERODORSAL AND 9 POSTERODORSAL bRISTLES. Wings. SC SHORT, ENDINg IN R, RM 1.5 TIMES AS LONg AS STEM Of POSTERIOR fORK, bASE Of ANTERIOR fORK SLIgHTLy bEfORE bASE Of POSTERIOR fORK. R1 AND R5 WITH SETAE, AbSENT AT bASIS Of VEINS. Male terminalia (FIg. 4). TERgITE 9 MEDIALLy DIVIDED, EACH PART ROUNDED, COVERED by MINUTE TRICHIA, WITH ONE STRONg bRISTLE, TWO SHORTER bRISTLES AND THREE SETAE. CERCI CLOTHED WITH fINE TRICHIA, WITH LONg SETAE APICALLy AND SOME MEDIALLy, PSEUDOCERCI WITH SEVERAL LONg SETAE MEDIALLy AND APICALLy. DORSAL LObE Of gONOSTyLUS ObLONg, APEX ROUNDED WITH SMALL, ROUND MEMbRANOUS AREA INTERNALLy; OUTER SURfACE WITH SEVERAL STRONg SETAE. MEDIAN LObE WITH SLIgHTLy CURVED POSTEROVENTRAL CORNER; 7 STRONg AND 2 SHORTER SETAE ON POSTERIOR MARgIN, ONE SETA ON INTERIOR SURfACE; ONE LONg SETA ON VENTRAL SIDE, AbOUT ONE THIRD Of TOTAL DISTANCE fROM POSTERIOR SIDE. VENTRAL LObE DISTINCTLy CLUb- SHAPED, WITH 8 SETAE APICALLy. BASAL PART Of gONOSTyLUS WITH TWO bULbOUS STRUCTURES Of EQUAL SIZE, DORSAL STRUCTURE WITH 5 LONg SETAE, VENTRAL STRUCTURE WITH AbOUT 8 STOUT SETAE (FIg. 4C). HyPANDRIAL LObE ELONgATED, WITH STRONg SCLEROTIZED LATERAL APODEMES (FIg. 4A). Etymology. NAMED AfTER THE TyPE LOCALITy. Remarks. THE SPECIMEN HAS bECOME PALE AfTER yEARS Of STORAgE IN ETHANOL. MEASUREMENTS WERE MADE ON SPECIMEN IN ETHANOL.