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Blepharoceridae Leiponeura cinerascens Loew. Adult male swept by river, nr Moreda, 24 May 2012, E.G. Hancock.

Cylindrotomidae Cylindrotoma distinctissima (Meigen). Several at stream edge, nr Moreda, 26 May 2012, E.G. Hancock.

Anisopodidae Sylvicola cinctus (Fabricius). Adult male emerged from detritus in tree stump, nr. Seoane do Courel, 25 May 2012, E.G. Hancock.

Trichoceridae Trichocera regelationis (Linnaeus). One swept off cliff face, another swept by woodland edge, near Moreda, 28 May 2012, E.G. Hancock This species has been recorded once before in Spain from Gerona in 1978 (Dahl, et al., 2002). New to Galicia.

Limoniidae Dicranomyia (Dicranomyia) mitis (Meigen). Nr Mercurin, Rio Lor, 27 May 2012; swept on woodland edge, nr Moreda, 26 May 2012; Alto do Couto, 25 May 2012, E.G. Hancock Dicranomyia (Dicranomyia) modesta (Meigen). Swept on woodland edge, nr Moreda, 26 May 2012, E.G. Hancock. Dicranomyia (Numantia) fusca (Meigen). Swept by river, nr Moreda, 26 May 2012, E.G. Hancock. New to Iberia. Dicranomyia (Sivalimnobia) aquosa Verrall. Ferrer��a Vella, nr Mercurin, swept by side of Rio Lor, 27 May 2012, E.G. Hancock. New to Iberia. Epiphragma ocellare (Linnaeus). Pupa in rot hole in oak tree, nr Moreda, 26 May 2012, G.E. Rotheray. Erioptera lutea (Meigen). Nr Mercurin, along side of Rio Lor, 27 May 2012, E.G. Hancock; dark form, sometimes given varietal name taenionata Meigen, 1818. Limonia flavipes (Fabricius). Common at all sites, 25���29 May 2012, E.G. Hancock. Limonia macrostigma ( Schummel). Damp meadow in river valley, near Field Station, Seoane do Courel, 28 May 2012, E.G. Hancock Limonia maculipennis (Meigen). Swept on woodland edge, nr Moreda, 26 May 2012; Alto do Couto, 25 May 2012, E.G. Hancock Limonia nubeculosa Meigen. Nr Mercurin, along side of Rio Lor, 27 May 2012, E.G. Hancock. Limonia phragmitidis (Schrank). Nr Mercurin, along side of Rio Lor, 27 May 2012; Seoane de Courel, 28 May 2012; common at all locations nr Moreda, E.G. Hancock. Lipsothrix galiciensis sp. nov. Hancock & Hewitt, 2014. Swept by river, nr Moreda, 26 May 2012, 4 males; by Rio Lor at Ferreria Nova, 30 May 2012, one male; near Alto do Couto, 28 May 2012 two females reared from waterlogged twigs in stream, E.G. Hancock; Devesa de Roguieira, 23 May 2012, 7 males and 5 females reared from larvae in water-logged sticks lying in shallow stream, S.M. Hewitt. Molophilus cinereifrons de Meijere. Damp meadow in river valley, near Field Station, Seoane do Courel, 28 May 2012, E.G. Hancock; one male. New to Iberia. Molophilus obscurus (Meigen). Swarming at locations in grassland, nr Moreda, 25���27 May 2012; damp meadow in river valley, near Field Station, Seoane do Courel, 28 May 2012, E.G. Hancock. Prionolabis pjotri sp. nov. Hancock, 2014. Five males swept on woodland edge, nr Moreda, 26 May 2012; five males and one female, Alto do Couto, 25 May 2012; seven males, nr Mercurin, 27 May 2012; one male by Rio Lor, Ferrer��a Nova, Seoane do Courel, 30 May 2012; E.G. Hancock. Rhabdomastix (Rhabdomastix) edwardsi Tjeder. By Rio Lor, Ferrer��a Nova, Seoane do Courel, 30 May 2012, E.G. Hancock; one female identified using key by Star�� (2004). New to Galicia. Tasiocera fuscescens (Lackschewitz). One male, swept on woodland edge, nr Moreda, 26 May 2012, E.G. Hancock. New to Iberia., Published as part of Hancock, E. Geoffrey, Hewitt, Stephen M., Horsfield, David, Lyszkowsi, Richard M., Macgowan, Iain, Ricarte, Antonio, Rotheray, Graham E. & Watt, Kenneth, 2015, Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae, pp. 231-244 in Zootaxa 3911 (2) on page 241, DOI: 10.11646/zootaxa.3911.2.5, http://zenodo.org/record/240391, {"references":["Ricarte, A., Rotheray, G. E., Lyszkowski, R., Hancock, E. G., Watt, K., Hewitt, S. M., Horsfield, D. & MacGowan, I. (2014) The syrphids do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera, Syrphidae). Zootaxa, 3793 (4), 401 - 422. http: // dx. doi. org / 10.11646 / zootaxa. 3793.4.1","Stary, J. (2004) Revision of European species of Rhabdomastix (Diptera, Limoniidae). Part 2: Subgenus Rhabdomastix s. str. European journal of entomology, 101, 657 - 668. http: // dx. doi. org / 10.14411 / eje. 2004.089"]}

Prionolabis pjotri sp. nov. Hancock Description. Head and entire body shining black, quite strongly dusted (pruinose) especially on the pleurae, less so on scutum and dorsally on abdomen, sparsely pale haired; apices of coxae, trochanters and bases of femora brown. Antennae 16 segmented. Wing membrane pale grey, stigma dark, wing veins yellow on basal third; medial vein four-branched (i.e., upper branch forked and so cell M 1 present). Halteres yellow. Male wing length 8.00mm, ranging from 6.6 to 9.0mm (sample size = 19). Male genitalia (Figure 15) with outer dististyle bifurcate at tip, inner one angled after swollen medial section resembling a bird���s beak. Parameres are flattened and only lightly sclerotised, expanding before a squared off truncated tip. Female same colouration as male but with apex of 8 th tergite, cerci and sternal valves of ovipositor yellowbrown; wing reduced (Figure 16), length 2mm (sample size = 1). Holotype: male, Alto do Couto, Lugo, Galicia, Spain, 25 May 2012, E.G. Hancock. Deposited in The Hunterian Zoology Museum, University of Glasgow (Accession number 150630). Paratypes: five males swept on woodland edge, nr Moreda, 26 May 2012; eleven males and one female, nr Mercurin, 27 May 2012; one male by Rio Lor, Ferrer��a Nova, Seoane do Courel, 30 May 2012; E.G. Hancock, The Hunterian Zoology Museum, University of Glasgow (Entry No. 897). Three additional specimens also designated as paratypes in the collection of J. Star��, Palack�� University, Olomouc, Czech Republic: one male, San Martin, Lugo, 22 March 1984, E. Eiroa; two males, 23 km south of Betanzos, A Coruna, in wet deciduous forest, 16 May 1994, P. Oosterbroek & C. Hartveld. Details in the male genitalia are distinctive, particularly the inner dististyle with its strong resemblance to the profile of a bird and the truncate apex to the pale and flattened parameres (Figure 15). The female���s wings are long enough to retain recognisable venation. The colouration seen in the live habitus photographs (Figures 17 and 18) shows the strong contrast between the yellow wing bases and the rest of the insect in both sexes. Etymology. The expertise of Pjotr Oosterbroek, managing the collections under his care and the creation and development of the Cranefly Catalog of the World is commemorated in naming this species. Discussion. There is one other known Spanish species in this genus, Prionolabis hospes (Egger), which is widespread in the western Palaearctic. It cannot be confused with P. pjotri in its lack of a branch to vein M 1 in the male, the female having even smaller wings with much reduced venation and is not black in colour. One other species Prionolabis longeantennata (Strobl) does have four branches to M but the antennae are almost as long as the body as does P. s c h m i d i Star��, but which has even longer antennae, as long as or longer than the body (Star��, 2006)., Published as part of Hancock, E. Geoffrey, Hewitt, Stephen M., Horsfield, David, Lyszkowsi, Richard M., Macgowan, Iain, Ricarte, Antonio, Rotheray, Graham E. & Watt, Kenneth, 2015, Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae, pp. 231-244 in Zootaxa 3911 (2) on page 236, DOI: 10.11646/zootaxa.3911.2.5, http://zenodo.org/record/240391, {"references":["Stary, J. (2006) New information resulting from a study of Strobl's types of Limoniidae. Acta universitatis Carolinae biologica, 49, 187 - 203."]}

Pediciidae Dicranota (Paradicranota) simulans Lackschewitz. One male, by Rio Lor, Ferrería Nova, Seoane do Courel, 30 May 2012; three males, one female, swept by stream, nr Moreda, 25–27 May 2012, E.G. Hancock and S.M. Hewitt. New to Iberia. Pedicia (Amalopsis) occulta (Meigen). Damp meadow in river valley, near Field Station, Seoane do Courel, 28 May 2012, E.G. Hancock. Ula mollissima (Haliday). Near Field Station, Seoane do Courel, 28 May 2012, E.G. Hancock, one female, apodeme character confirmed by reference to redescription by Starý (1983). New to Iberia.

Lipsothrix galiciensis sp. nov. Hancock & Hewitt Description. Male with thorax and abdomen together 7.0mm long, shining brownish-black except hypopygium yellowish brown and tergites narrowly pale distally; head dark brown, lighter brown on occiput; antenna 3.4mm long, brown; coxae and legs dark brown except base of femora and apex of hind coxa yellowish brown. Wing length 9.2mm (sample size = 5); membrane slightly greyish, costal cell with a faint yellowish tinge, stigma dark brown. Hypopygium with dististyles and paramere as in Figure 8; aedeagus as in Figure 9. Female colouration not as dark as males; yellow 8 th segment and ovipositor; body length 8.00mm, antenna 1.7mm, wing 8.00mm (sample size = 2). Holotype: male, swept by river, nr Moreda, Lugo, Galicia, Spain, 26 May 2012, circa 720m, E.G. Hancock. Deposited in The Hunterian Zoology Museum, University of Glasgow, (Accession number 150629). Paratypes: three males, swept by river, nr Moreda, 26 May 2012; one male, by Rio Lor at Ferreria Nova, 30 May 2012, E.G. Hancock; two females reared from larvae in waterlogged wood in stream near Alto do Couto, 28 May 2012, circa 1300m, E.G. Hancock, in The Hunterian Zoology Museum, University of Glasgow (Entry No. 897); 7 males and 5 females reared from larvae found in wet wood in a shallow stream, Devesa de Rogueira, 23 May 2012, circa 1200m, S.M. Hewitt, deposited in Tullie House Museum, Carlisle, England (CALMG: 2013.109. 1 -. 6). This species is quite different from the other western Palaearctic species both in adult colouration (Figure 10) and the pupal thoracic spiracular gills. It shares the sexually dimorphic features of antennal length being longer in the male and the grey tinged wing membrane with Lipsothrix nervosa Edwards. But the latter has yellowish pleurae and abdominal sternites and females are mainly yellow. In L. galiciensis the sexes differ only in the intensity of the brownish-black body. The inner dististyle has a distinctly protuberant inner margin narrowing before an expanded apex. The aedeagus bears a ventral peg-shaped protrusion as described by Edwards (1938) for nervosa but is more sinuous apically than that species. Etymology. The word galiciensis, means from Galicia, the region of Spain where the type locality of this species is found. The area of Galicia includes excellent wooded and montane habitat of considerable entomological potential for biodiversity studies. Discussion. Adults were reared from larvae and pupae in sticks of less than 10cm diameter lying loose in small streams in mixed woodland. The stream beds were stony with muddy leaf-mould and the water shallow so the sticks were only partially submerged. In addition to the adult features described above the pupal exuviae allow comparison of L. galiciensis with those of the other European species (Hancock, et al., 2009). The differences in the thoracic spiracular gill are extensive in both overall shape and the detail of the plastron structure. Of particular note is the wavy margin (Figures 11, 12, 13) to the gill and the clusters of about 6���10 spines in discrete groups on the inner face (Figure 14)., Published as part of Hancock, E. Geoffrey, Hewitt, Stephen M., Horsfield, David, Lyszkowsi, Richard M., Macgowan, Iain, Ricarte, Antonio, Rotheray, Graham E. & Watt, Kenneth, 2015, Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae, pp. 231-244 in Zootaxa 3911 (2) on pages 233-236, DOI: 10.11646/zootaxa.3911.2.5, http://zenodo.org/record/240391, {"references":["Edwards, F. W. (1938) British short-palped craneflies, taxonomy of adults. Transactions of the society for British entomology, 5, 1 - 168.","Hancock, E. G., Hewitt, S. M., Godfrey, A. & Mullin, M. (2009) Thoracic spiracular gill structure of Lipsothrix (Diptera, Limoniidae) in Britain described from scanning electron micrographs, with a key to pupae. Zoosymposia, 3, 7 - 87."]}

Key to adults of Lipsothrix species of the western Palaearctic 1. Wing with dark stigma; thorax at least partly darkened on scutum............................................... 4 - Wing without darkened stigma; yellowish species........................................................... 2 2. Femora yellow; wing veins with short, inconspicuous macrotrichia; 2 A bare; blackish palps, tarsi and base of 8 th sternite of male..................................................................................... remota Walker - Femora with black apices; veins with long macrotrichia; 2 A with some macrotrichia at apex.......................... 3 3. Abdomen with dorsal stripe; male with distal part of 7 th and whole of 8 th sternites blackish; vein 2 A with numerous macrotri- chia towards apex......................................................................... errans Walker. - Abdomen without dorsal stripe, male with only 8 th abdominal sternite darkened; vein 2 A with only 3–6 macrotrichia............................................................................................ ecucullata Edwards 4. Wing apex and cross veins clouded............................................................. nobilis Loew - Wing apex clear...................................................................................... 5 5. Thorax dorsally and abdominal tergites blackish; pleurae and sternites yellow........................ nervosa Edwards - Thorax and abdomen brownish-black including pleurae and sternites................................ galiciensis n. sp.

Hancock, E. Geoffrey, Hewitt, Stephen M., Horsfield, David, Lyszkowsi, Richard M., Macgowan, Iain, Ricarte, Antonio, Rotheray, Graham E., Watt, Kenneth (2015): Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae. Zootaxa 3911 (2): 231-244, DOI: http://dx.doi.org/10.11646/zootaxa.3911.2.5

Ten species of Copestylum (Diptera: Syrphidae) were reared from fruits and flowers in Costa Rica, Ecuador and Trinidad. Seven were new and in this paper, we describe them, their development sites and the third stage larva and/or the puparium of all ten species. One new synonym is proposed, Copestylum pinkusi (Curran) [= Copestylum cinctiventre (Curran)]. Similarities and differences between these new and other Copestylum species, suggest they separate into two groups, referred to as the Vagum and Cinctiventre species groups. Features characterising these groups for both adult and early stages are assessed. Each species was also distinguished using adult and early stage characters. Within the Vagum group, adults were more disparate morphologically than the larval stage; this was reversed in the Cinctiventre group. Adult colour patterns are probably cryptic in function and for disguise. Vagum species have disruptive marks, while the Cinctiventre species have reflective colours. Biologically, the groups are almost distinguished by larval development sites. Vagum species use predominantly fruits and have a larval stage that is relatively generalised in form and habit. Cinctiventre species are confined to developing in flowers and the larva is more specialised. A key to both adult and early stages of all ten species is provided.

QUICHUANA BEZZII CERESA, 1934: 387 FIGURES 21, 23 AND 24 Three males and six females were examined by Ceresa (1934), three males and five females of which comprise the type series. These eight specimens are deposited in MCSN. Each specimen of the type series is red-labelled as ‘cotypus’ with the letters ‘co’ crossed out. We designate a lectotype and seven paralectotypes to stabilize the name (see ‘Material examined’). Diagnostic features Body extensively black; frontal triangle flat; bf = 1.3 (lectotype); basoflagellomere oval (Fig. 21); face extensively white pollinose, except for a central, short, shiny, black stripe comprising the facial tubercle; facial lateral stripes, when visible, with sparser pollinosity than that on the rest of the face; thorax without tufts of hairs, although longer hairs at NP, PAPT, and PC; scutum with PP pollinose and two medial, white pollinose stripes extending on the anterior three-quarters of the scutum, and diverging posteriorly; pro- and mesotibia red at least on the basal half; metafemur from copper red on the apical third to wholly copper red; metatibia copper red; wings extensively microtrichose, except for small bare areas in cells BM, CuP, basal area of C, and alula; terga black; tergum I only with short pale hairs; hairs on the lateral margins of terga I– II pale, relatively short, and wavy only towards the tip; male genitalia with superior lobes conical towards the apex (Fig. 23); hypandrium with lateral extensions on the apical half (Fig. 23); surstyli concave antero-apically, but rounded postero-apically (Fig. 24). Material examined (MCSN) Lectotype: 1m, Brasil, Est. Saõ Paulo, S. Sebastiaõ, A.A. Barbiellini /cotypus. Paralectotypes: Brasil, Est. Saõ Paulo, S. Sebastiaõ: 1m, A.A. Barbiellini, 11–1906 (handwritten), cotypus; 1m, A.A. Barbiellini, 491, cotypus; 1f, A.A. Barbiellini, 11–1906 (handwritten), cotypus; 2f, A.A. Barbiellini, 4–1917 (handwritten), cotypus; 1f, A.A. Barbiellini, 11–1906 (handwritten), 805 (handwritten), cotypus, Quichuana bezzii Cer., det. L. Ceresa (species name handwritten); 1f, Brasil, Est. S. Paulõ, S. Sebastiaõ, A.A. Barbiellini (most of the abdomen lacking). On the ‘cotypus’ label of all the specimens, ‘co’ is crossed out in ink. Range Argentina, Brazil. Taxonomic notes Medium size species (10.7 mm, lectotype) with slender abdomen; Q. bezzii can be separated from the similar Q. undulatipila sp. nov. by the surface of the frontal triangle, which is flat in the former but domed in the latter; additionally, the hairs on the lateral margins of terga I–II are shorter and not so conspicuously wavy (wavy for a shorter length) than those in terga I–II of Q. undulatipila sp. nov.; the shape of the superior lobes and surstyli in the male genitalia also differ between Q. bezzii (Figs 23, 24) and Q. undulatipila sp. nov. (Figs 65, 66)., Published as part of Ricarte, Antonio, Marcos-García, M. Ángeles, Hancock, E. G. & Rotheray, Graham E., 2012, Revision of the New World genus Quichuana Knab, 1913 (Diptera: Syrphidae), including descriptions of 24 new species, pp. 72-131 in Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) (Zool. J. Linn. Soc.) 166 (1) on pages 80-81, DOI: 10.1111/j.1096-3642.2012.00842.x, http://zenodo.org/record/5408533, {"references":["Ceresa L. 1934. Tre nuove specie de Quichuana Knab (Diptera: Syrphidae). Atti de la Societa Italiana Scienze Naturali 73: 383 - 392."]}

Ricarte, Antonio, Marcos-García, M. Ángeles, Hancock, E. G., Rotheray, Graham E. (2012): Revision of the New World genus Quichuana Knab, 1913 (Diptera: Syrphidae), including descriptions of 24 new species. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (1): 72-131, DOI: 10.1111/j.1096-3642.2012.00842.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2012.00842.x

Cheilosia fraterna (Meigen) Material examined. Leg. S.M. Hewitt: 2 ♂, Moreda valley meadows, 21.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 407, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Sphegina elegans Schummel Material examined. Leg. E.G. Hancock: 1 ♂, Seoane do Courel, near Univ. Sant. Fld. St., from larva collected in decaying wood [NMS]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 415, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Sphegina clunipes (Fallén) Material examined. Leg. S.M. Hewitt: 1 ♂, Devesa da Rogueira, 23.v. 2012, ex larva collected in wet wood lying in woodland stream [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 415, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Brachyopa scutellaris Robineau-Desvoidy # Material examined. Leg. A. Ricarte: 1 ♂, Teso de María Galega, path edge vegetation, 720 m, resting on the grass, 29.v. 2012; 1 ♂ and 1 ♀, meadow along Moreda valley, 720 m, hovering around the trunk of a Castanea sativa tree, 24.v. 2012 (♀) [NMS], 28.v. 2012 (♂) [CEUA]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 407, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Microdon analis (Macquart) Material examined. Leg. A. Ricarte: 3 ♂ and 1 ♀, A Fontela, near A Tara, path edge vegetation, 1280 m, 25.v. 2012 [NMS]. Leg. R.M. Lyszkowski: 1 ♂, Formigueiros top, 1635 m, 25.v. 2012. Leg. S.M. Hewitt: 1 ♂, A Fontela, near A Tara, 23.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 412, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Paragus (Paragus) pecchiolii Rondani Material examined. Leg. D. Horsfield: 1 ♂, Moreda, 28.v. 2012, brooms and herb-rich vegetation (Ruta 1), 1050 m; 1 ♀, Seoane, Estación Científica, road edge vegetation, 610 m, 22.v. 2012 [NMS]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 412, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Eumerus ornatus Meigen Material examined. Leg. E.G. Hancock: 1 ♂, nr Moreda, meadow by woodland edge, 23−28.v. 2012 [HM]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 410, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Eupeodes latifasciatus (Macquart) Material examined. Leg. A. Ricarte: 2 ♀, A Fontela, path edge vegetation, 1280 m, 25.v. 2012; 1 ♀, meadow along Moreda valley, 720 m, 24.v. 2012 [NMS]. Leg. E.G. Hancock: 1 ♀, nr Moreda, meadow by woodland edge, 23−28.v. 2012 [HM]. Leg. S.M. Hewitt 1 ♀, Moreda valley meadows, 21.v. 2012, [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 410, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Cheilosia chrysocoma (Meigen) Material examined. Leg. E.G. Hancock: 1 ♂, Formigueiros, 1639m, 29.v. 2012 [HM]. Leg. R.M. Lyszkowski: 2 ♂, Formigueiros top, 1635 m, 25.v. 2012; 2 ♂, path to summit of Formigueiros, 1350 m, 29.v. 2012 [RL]. Leg. S.M. Hewitt: 2 ♂, Formigueiros, 23.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 407, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Cheilosia laticornis Rondani Material examined. Leg. A. Ricarte: 1 ♀, meadow along Moreda valley, 720 m, 28.v. 2012 [NMS]. Leg. E.G. Hancock: 1 ♀, nr Moreda, 23-28.v. 2012, swept by river [HM]. Leg. S.M. Hewitt: 1 ♀, Moreda valley meadows, 25.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 408, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Syritta pipiens (Linnaeus) Material examined. Leg. E.G. Hancock: 1 ♀ nr Moreda, swept by river, 23−28.v. 2012, [HM]. Leg. S.M. Hewitt: 1 ♂, Moreda Valley meadows, 21.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 415, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Cheilosia laeviseta Claussen Material examined. Leg. A. Ricarte: 2 ♀, A Fontela, path edge vegetation, 1280 m, 25.v. 2012 [NMS]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 408, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Eupeodes corollae (Fabricius) Material examined. Leg. A. Ricarte: 1 ♂, Seoane, camino de Oval, near Estación Científica, track edge vegetation, 610 m, 28.v. 2012 [NMS]. Leg. E.G. Hancock: 1 ♂, nr Moreda, meadow by woodland edge, 23−28.v. 2012; 1 ♀, Alto do Couto, 25−27.v. 2012 [HM]. Leg. R.M. Lyszkowski: 1 ♀, Alto do Couto, 1320 m, 26.v. 2012 [RL]. Leg. S.M. Hewitt 1 ♀, Moreda Valley meadows, 25.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 410, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Description of Cheilosia lucense Ricarte sp. nov. (figs 1–5) Male: Measurements. L= 10.6 mm, WL= 7.8 mm. Head. Eye facets increasingly large towards the upper part of eye, but never as large as an ocellus; eye with conspicuous reddish-yellow hairs; vertical triangle black, elevated and punctured centrally, faintly white-pollinose on the anterior corner; vertical triangle with reddish-yellow hairs (longer than those on eye) and a few black hairs intermixed; frontal triangle black, punctured, faintly white pollinose laterally, with long reddish-yellow hairs and a few black hairs centrally; frontal triangle 1.5 times longer than eye contiguity; lunule shiny reddish brown; scape and pedicel with both pale and black hairs; scape black; pedicel brownish black, paler towards the apex; basoflagellomere nearly circular (Fig. 1), reddish brown and white pollinose; antennal arista black, with very short hairs, the length of which is about half the maximum width of arista; face black and with whitish microtrichia, except for the central shiny tubercle; eye rim at its maximum width as wide as shiny part of facial tubercle; eye rim with yellowish-white hairs and as long as those on the closer areas of eye; face with a triangular shiny area from the mouth edge to eye rim; gena and occiput black, with white pollinosity, which is denser on the occiput; gena with a reddish brown spot just below the eye; upper half of occiput conspicuously narrower than lower half; occiput with yellowish red hairs and a row of black hairs of increasing length on the upper part. Thorax. Scutum shiny black, with long reddish-yellow hairs and, on the posterior half of scutum and near the notopleuron, black hairs intermixed; pleuron black, faintly grey pollinose; posterior anepisternum, anepimeron and katepisternum with reddish yellow hairs; ventral part of anterior anepisternum with whitish yellow hairs anteriorly; metasternum haired; scutellum shiny black, with long red hairs and, on the centre, a few black hairs intermixed; subscutellar fringe of hairs red; black coxae, pale haired; femora black except for the red apices; femora with reddish yellow hairs and, on the apical half, black hairs intermixed (black hairs shorter towards the femur apex); metafemur with a few outstandingly long pale hairs ventrally; tibiae red, with a faint black ring; tibiae with pale and black hairs intermixed; protarsus black dorsally; mesotarsomeres 1–3 blackish brown, mesotarsomeres 4 and 5 black; metatarsus black dorsally (tarsomeres 1–3 reddish apically); all tarsi red ventrally except for the blackish tarsomeres 5; claws red on the basal half; wing wholly microtrichose, with basal veins red; calypter white, reddish marginally; halter red, with capitulum blackish. Abdomen. Slender, 1.6 times longer than wide; hairs on central part of terga shorter than those on the central part of thoracic scutum; terga covered in reddish yellow hairs; terga black; tergum I dark pollinose except for the shiny posterior margin; tergum II dark pollinose except for lateral margins; tergum III shiny except for a central dark-pollinose stripe not reaching the posterior margin; tergum IV wholly shiny; sternum I wholly grey pollinose; sternum II grey pollinose except for two posterior semicircular areas; sterna I and II wholly pale haired; sterna III and IV grey pollinose on both the anterior and posterior margins and with black hairs posteriorly; pleural membrane brownish black. Genitalia. Surstylus elongate (3.2 times longer than wide in lateral view), straight (Fig. 3); lateral expansion of surstylus at its maximum width 1.9 times wider than that at the base of the expansion in dorsal view (Fig. 2); ventral lobe of gonostylus slightly widening towards the apex, 3.7 times longer than the dorsal lobe of gonostylus, which is short, slightly waisted centrally and with apex sclerotised (Fig. 3). Female: Same as the male except for the following characters: L= 11.3 mm, WL= 8.2 mm. All eye facets equal in size; frons black, finely punctured and with disperse white pollinosity; frons with a central excavation from the ocellar triangle to the area close to the lunules; ocellar triangle and frons only with reddish yellow hairs; basoflagellomere slightly curved dorsally and strongly curved ventrally, with a conspicuous corner dorso-apically (Fig. 4); basoflagellomere red, faintly pollinose, with pits both on the inner and outer sides (Fig. 4); arista with hairs slightly longer than those in male but shorter than the maximum width of arista; face profile as in Fig. 4; upper half of occiput slightly narrower than lower half but conspicuously wider than that on male; scutum and scutellum only with reddish yellow hairs; profemora only with pale hairs; meso- and metafemora with yellow hairs except for a few short black adpressed hairs dorsally on the apices and a few black (longer than those dorsally) hairs ventrally, near the apices (Fig. 5); tibiae mainly pale haired, with some black hairs intermixed; mesotarsomeres 1-3 paler than those in male; abdomen oval, more stocky than in male (1.3 times longer than wide); tergum III extensively shiny; sternum II pollinose anteriorly and laterally; sternum III narrowly pollinose laterally; sternum IV narrowly pollinose laterally and posteriorly. Differential diagnosis. The male of this species can be separated from the male of the similar C. chrysocoma (Meigen) in the following characters: in C. lucense sp. nov., anterior side of hypandrium nearly straight, dorsal lobe of gonostylus short and stocky, surstylus short in lateral view (Fig. 3), conspicuously expanded in dorsal view (Fig. 2); in C. chrysocoma, anterior side of hypandrium excavated, dorsal lobe of gonostylus elongated, surstylus long in lateral view (ratio length/width higher than that in the new taxon), surstylus moderately to slightly expanded in dorsal view (see figures 2, 9 and 16 f in Claussen 1998). Females of C. lucense sp. nov. are most similar to those of C. chrysocoma. The female of C. lucense sp. nov. keys out to C. chrysocoma in Claussen (1998) and Bartsch et al. (2009 b). The basoflagellomere in C. chrysocoma female usually has a conspicuous dorsal corner (Claussen 1998) which is absent in the examined females of C. chrysocoma from Durmitor (Montenegro). Female of C. lucense sp. nov. also has a dorsal corner in the basoflagellomere (Fig. 4). A few characters may help to separate females of these two species: in C. lucense sp. nov. facial tubercle angular in profile (Fig. 4); sternum II pollinose, with a semicircular shiny area on the posterior margin; in C. chrysocoma facial tubercle more roundish; sternum II extensively shiny, only narrowly pollinose on the anterior and lateral margins. Specimens of both C. lucense sp. nov. and C. chrysocoma were found in Courel. Type material of C. lucense sp. nov. was compared with specimens of C. chrysocoma from the north of Spain (León) [CEUA], England [NMS], Serbia and Montenegro [FSUNS]. Type material. Holotype: 1 ♂, Spain, Galicia, Lugo, Folgoso do Courel, A Fontela, path edge vegetation, 1280 m, 25.v. 2012, leg. A. Ricarte [NMS]. Paratypes: 1 ♀, Spain, Galicia, Lugo, Folgoso do Courel, meadow along Moreda valley, 720 m, on flowers of Chaerophyllum hirsutum, 23.v. 2012, leg. A. Ricarte [CEUA]. Leg. R.M. Lyszkowski: 2 ♂, Spain, Galicia, Lugo, Folgoso do Courel, nr. Pedra Boa, 750 m, 24.v. 2012 [1 ♂ in MNCN, 1 ♂ in RL]. Etymology. The epithet lucense means ‘from Lugo’ and refers to the Spanish province of the type locality, Lugo. Distribution. Spain (Lugo province)., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on pages 403-406, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393, {"references":["Claussen, C. (1998) Die europaischen Arten der Cheilosia alpina-Gruppe (Diptera, Syrphidae). Bonner zoologische Beitrage, 47, 381 - 410.","Bartsch, H., Binkiewicz, E., Klintbjer, A., Raden, A. & Nasibov, E. (2009 b) Blomflugor: Eristalinae & Microdontinae. Nationalnyckeln till Sveriges flora och fauna, DH 53 b. Artdatabanken, SLU, Uppsala, 478 pp."]}

Melanostoma scalare (Fabricius) Material examined. Leg. A. Ricarte: 1 ♂, Moreda valley, near Aula Naturaleza, Castanea sativa souto, 720 m, 23.v. 2012; 1 ♀, meadow along Moreda valley, 720 m, 23.v.2012, 23.v. 2012. Leg. D. Horsfield 2 ♀, Moreda, herbrich meadows around A Moreda Maior, 930 m, 28.v. 2012; 1 ♀, Moreda valley, herb-rich meadows, 720 m, 24.v. 2012 [NMS]. Leg. E.G. Hancock: 3 ♀, nr Moreda, meadow by woodland edge, 23−28.v. 2012; 1 ♀, Seoane do Courel, Univ. Sant. Fld. St., swept by river, 23−30.v. 2012; 1 ♀, Seoane do Courel, Ferreria Nova, swept by river, 30.v. 2012 [HM]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 411, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Sphaerophoria rueppelli (Wiedemann) Material examined. Leg. E.G. Hancock: 1 ♂, nr Mercurín, by roadside, río Lar, 27.v. 2012 [HM]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 414, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Platycheirus albimanus (Fabricius) Material examined. Leg. A. Ricarte: 1 ♀, meadow along Moreda valley, 720 m, 23.v. 2012 [NMS]. Leg. E.G. Hancock: 1 ♂ and 1 ♀, nr Moreda, swept by river (♂), meadow by woodland edge (♀), 23−28.v. 2012; 2 ♂, Formigueiros, 1639m, 29.v. 2012 [HM]. Leg. R.M. Lyszkowski: 1 ♀, S. of Mercurín, 600 m, 27.v. 2012 [RL]. Leg. S.M. Hewitt 1 ♀, Moreda valley meadows, 26.v. 2012, [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 414, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Brachyopa bicolor (Fallén) # Material examined. Leg. A. Ricarte: 1 ♂, meadow along Moreda valley, 720 m, hovering around the trunk of a Castanea sativa tree, 23.v. 2012 [NMS]. Leg. E.G. Hancock: 2 ♂, nr Moreda, swept by river, 23−28.v. 2012 [HM]. Leg. S.M. Hewitt: 1 ♂, Moreda Valley meadows, 25.v. 2012 [CALMG]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 406, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David, Macgowan, Iain (2014): The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae). Zootaxa 3793 (4): 401-422, DOI: http://dx.doi.org/10.11646/zootaxa.3793.4.1

Epistrophe eligans (Harris) Material examined. Leg. R.M. Lyszkowski: 1 ♂, As Veigas, 930 m, 28.v. 2012 [RL]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 409, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393

Cheilosia flavipes (Panzer, 1798) Material examined. Leg. A. Ricarte: 2 ♀, A Fontela, 1280 m, path edge vegetation, 25.v. 2012 [NMS]., Published as part of Ricarte, Antonio, Rotheray, Graham E., Lyszkowski, Richard M., Hancock, E. Geoffrey, Hewitt, Stephen M., Watt, Kenneth R., Horsfield, David & Macgowan, Iain, 2014, The syrphids of Serra do Courel, Northern Spain and description of a new Cheilosia Meigen species (Diptera: Syrphidae), pp. 401-422 in Zootaxa 3793 (4) on page 407, DOI: 10.11646/zootaxa.3793.4.1, http://zenodo.org/record/225393