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Bryophila schwingenschussi (Boursin, 1954) (Figs 3b and 7b) Material examined: Holotype: 1 male, “ Bône (Algerien), Juni 1931 (J. Staettermayer leg.) (Coll. Schwingenschuss)“ (Figs 3b and 7b). In NHMW. Taxonomic notes: It was described on a single male specimen, being compared with B. barbaria and B. blepharista, in a text where B. muralis and B. domestica were also mentioned as similar taxa. Boursin (1954) stated that this species and B. blepharista are very similar externally but B. schwingenschussi has shorter pectination of the male antennae, slenderer uncus, somewhat longer valvae and the vesica is armed with a strong cornutus which is missing from both B. blepharista and B. barbaria. As far as is known, it is restricted to Morocco. External and genital diagnosis: see under taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila barbaria Schawerda, 1934 (Figs 2a to 2d, 6c and 8c) Material examined: A large number of specimens of both sexes from the high altitude parts of the High Atlas (mainly from Oukaimeden). Apart from the lectotype and paralectotype (Figs 2a and 2b), the data of the dissected specimens are as follows: RL12467m—Morocco, Haut Atlas Mts, Oukaimeden, 2700 m, 10.VII.1986, leg. F. Fernandez-Rubio (coll. GR) and RL 12477m and RL12479f—Morocco, High Atlas Mts, Oukaimeden, 2400-2700 m, 5-21.VII.1971, leg. G. Friedel (coll. HNHM). Lectotype designation: Male, “Mauretania central, Atlas major, Sidi Chamarouche, Reraia-Tal, 2300 m, 27.6.1932, H. Dürck leg.”, “collectio Schawerda”, “ Bryophila muralis var barbaria Schaw., Type. ♂. Abgebildet ” (pink label), “PREPARATION | W.M. 409 | CH. BOURSIN” (coll. NHMW) (illustrated in Fig 2a). Taxonomic notes: The date of description is cited in most references as 1935, the date printed onto the volume is 1st December, 1934. It is a well-known species of the Atlas Mts in the North African Mediterranean. Schawerda (1934) wrote: „Im Grossen Atlas (Mauretanien = Barbaria) von Dr. Dürck (München) erbeutet. Sidi Chamarouche, Reraia-Tal, 2300 m, 27.6.1932. Types in my collection“. Despite the statement of Schawerda, arguing that one was male and the other female, the syntypes are both males. External diagnosis: This species has longer and broader wings than B. perloides and lighter forewing colouration, and the dark costal markings are always clearly visible (Figs 2a to 2d); the two species are well-separable by their external features. Genital diagnosis: In the male genitalia (Fig 6c), this species has, within the species complex, the longest uncus and strongest clasper, this longest too and with sharp apex. Also the valva is slenderest at the middle, with broadly rounded cucullus, compared to B. perloides. B. barbaria shows no cornuti in the vesica. In the female genitalia (Fig 8c), the ductus bursae is dorsally swollen at his posterior half (Fig 5c)., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on pages 60-61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Schawerda, K. (1934) Neue Heteroceren aus Algerien. Internationale Entomologische Zeitschrift, 28, 285 - 287."]}

Bryophila blepharista (Boursin, 1954) (Figs 3c and 7c) Material examined: Holotype (Figs 3c and 7c): 1 male, Taanzoult [Tazoult], Mittel-Atlas, French Marokko, 16. August 1950 (Ch. Rungs leg.). Paratypes: 1 male, from the same locality and date (Ch. Rungs leg.), 4 males, Aguelmane de Sidi Ali, Mittel-Atlas 2080 m, 5/ 25 July 1943 (Ch. Rungs leg.) (Coll. Rungs, Boursin and Schwingenschuss), 1 male, Bahlil, French Marokko, 20. August 1952 (Abdallah leg.) (Coll. Ch. Rungs). In MNHN. Taxonomic notes: The species was described from Morocco based on a few male specimens, showing differences in their antennae and certain features of the male genitalia (stout uncus and valval apical section, vesica without cornuti) compared with B. schwingenschussi and B. barbaria (Boursin 1954). External diagnosis: B. blepharista differs from B. perloides externally in the shape of the omega-shaped design, in which the inner marginal line is usually at a higher position in B. blepharista but close to the hind margin in B. perloides. Additionally, B. blepharista shows more or less pronounced antemarginal line, absent or almost absent in B. perloides. Genital diagnosis: The male genitalia of B. blepharista (Fig. 7c) are seemingly stouter than those of B. perloides (Figs 6a and 6b), according to the short series of examined specimens; the central clasper is comparatively shorter and gracile in B. perloides, although in both species shows spatulate apex. On the other hand, the uncus of B. blepharista has a narrow stem and incrassate subapical part, exactly like in B. perloides. The sacculus is short and flimsy, like in B. perloides too. Examination of larger series may clarify if these differences are actually interspecific or rather intraspecific. Epiphallus is cylindrical in B. blepharista and conical in B. perloides; but these differences may be due to different pressure of the cover glass., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 66, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila squamosa Schwingenschuss, 1936 (Figs 3a and 7a) Material examined: Two syntypes, labelled „ H. Atlas, Maroc, Goundafa, 1200 m, 15.-30.VI.1933, Schwingenschuss”, one of them designated here as lectotype (Fig 3a). In NHMW. Lectotype designation: male, “ H. Atlas, Maroc, Goundafa, 1200 m, 15.-30.VI.1933, Schwingenschuss”, “ Bryophila squamosa Schwingenschuss Type ♂ ”, slide No.: RL 12464m (coll. NHMW). Taxonomic notes: The taxon was described by Schwingenschuss in Zerny (1936). Most probably, the descriptions of B. barbaria and B. squamosa were made in parallel, and both may refer to the same taxon. Thus, the statement of Boursin (1954) about their synonymy may be correct. Nevertheless, we have found some differences between their male genitalia: the uncus of B. squamosa (Fig 7a) is more slender in its distal half than that of B. barbaria (Fig 6c), the apical section of valva is also narrower and less dilated at its tip, and the vesica is remarkably broader, more discoidal. We refrain from establishing the synonymy formally until we are able to study a longer series of specimens, including females. It has to be noted that Schwingenschuss itself remarked that “ Bryophila squamosa mihi is after Βοursin also just a barbaria. Since the design matches that of barbaria, but the wing-cut is different -especially in the female-, as well as the general impression, I consider it a subspecies, especially since the animals were found at 1200 m high in a hot basin in mid-June, while barbaria was found at 2200-2700 m in July.” External diagnosis: see under B. barbaria. The lectotype shows a much more blurred condition than most of the examined specimens of B. barbaria; it is thus probably an older specimen. Genital diagnosis (Fig. 7a): see under B. barbaria and here, under Taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Zerny, H. (1936) Die Lepidopterenfauna des Grossen Atlas in Marokko und seiner Randgebiete. Memoires de la Societe des Sciences Naturelles du Maroc, 42, 1 - 163.","Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila katiba (Rungs, 1972) (Figs 4a and 4b; 8d) Material examined (but not yet dissected): Holotype male (Fig 4a) and two male paratypes: Maroc, Anti Atlas, Tafraout, 18- VI11-1946, in MNHN. After Rungs (1972): “Wingspan: 25 mm. A female captured with the male, shows the bottom of the wings shrimpy pink and no deep white: leander n.f. «. 1 female (dissected) (Figs 4b and 8d), Morocco, Anti-Atlas Mts, Tizi’n Tarakatine pass, 1600 m, 29°45’11”N, 8°50’02”W, 29-V-2005, leg. Gy. M. László & G. Ronkay (in coll. GR). Taxonomic notes: The taxonomic status of this taxon remains somehow uncertain, as no type material has yet been dissected, but it differs externally from the other members of the species-group by its smaller size, paler colouration and dark markings (Figs 4a and 4b). The types resemble more B. barbaria than any other taxa, though they are remarkably smaller and paler coloured than specimens of B. barbaria from the higher altitude places of the Moroccan Atlas. Besides, the genitalia of a female collected in the Anti Atlas mountains (Fig 8d) clearly differ from those of B. barbaria (Fig 8c) in the shape of the appendix bursae and in the width of the ductus bursae, much narrower in B. katiba and without the characteristic swelling of B. barbaria. External and genital diagnosis: see under taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 66, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Rungs, C. E. E. (1972) Lepidopteres nouveaux du Maroc et de la Mauritanie. Bulletin du Museum National d'Histoire Naturelle, Series 3, 60, Zoologie 46, 669 - 692."]}

Bryophila perloides Guenée, 1852, stat. rev. (Figs 1a to 1d, 6a and 6b) Material examined: 2 male syntypes in coll. NHMUK, “ Andalousie ”, one of them here designated as lectotype (Figs 1a and 6b) (specimen labelled as 010918082). 1 male, 7-9-2010, Finca La Molina, Casares, Málaga, Spain (30 STF93), Penny Hale leg., coll. MF in coll. ZMC (Figs 1b and 6a). 2 males, 24.8.2018 and 21.9.2018, Campo de Gibraltar, Algeciras, Cádiz, Spain (30 STE79), José Manuel Gaona Ríos phot. (Figs 1c and 1d). Lectotype designation: male, “ Andalousie ”, “ Perloides ”, “ Syntype ” (blue ring label), “Ex MUSAEO Dris BOISDUVAL”, “Fig par J. CULOT, Noct. et Géom. d’Europe, Pl. 24 Fig 12”, “Vu par Stgr pour Catal. 1900”, “Ex Oberthür Coll. Brit. Mus. 1927-3.” (specimen labelled as 010918082, coll. NHMUK) (illustrated in Fig 1a). Taxonomic notes: the species is listed in most recent works as subordinate to Bryopsis muralis (e.g. Poole 1989, Fibiger et al. 2009; but see Fibiger et al. 2011: 33). This statement is erroneous; this species in not a Bryopsis but a true Bryophila, according to the genital configuration (see above). All five known specimens are from Andalucía, S. Spain; therefore perloides is the only European member of the group. External diagnosis: Ground colour bright olive green (Figs 1a and 1b) to dull greyish (Figs 1c and 1d). Forewings relatively small and tapered to the apex. Unlike B. domestica, subbasal and antemedian lines fused basally by an inner marginal line, forming the omega-shaped design typical of the species-group, which is completely closed in perloides; this feature is shared with the species of Bryopsis (e.g. B. muralis), where it is open to almost open in its basal part (Figs 1e and 1f). Dark spots at the costa very subtle. Unlike B. muralis, but in common with the rest of the species of the group, black X-mark on the forewings proximal to reniform stigma present. Antemarginal line absent or very subtle, in contrast to B. muralis, where it is quite apparent. Genital diagnosis: male genitalia simple, as in the rest of the species of the group; all species have similar genitalia, mostly differing in subtle details (Figs 6a to 7c). Those of B. perloides (Figs 6a and 6b) are particularly similar to those of B. barbaria (Fig 6c), but the uncus is more incrassate subapically, with shorter tip, whereas that of B. barbaria is more gradually tapered towards its apex. Additionally, the aedeagus of B. perloides shows a narrow coecum penis and broadens to its distal end, whereas in B. barbaria and other closely related species it is more or less cylindrical. Costal and hind margins of the valvae of B. perloides are almost parallel; the sacculus is weaker and smaller, the clasper smaller too, fully embedded onto the central area of valva, with slightly spatulate apex. Vesica without cornuti. The female genitalia has not yet been studied, no females have been collected so far., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on pages 56-60, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Guenee, A. (1852 - 1854) Noctuelites (II). In: Boisduval, J. A. & Guenee, A., Histoire naturelle des lnsectes. Species general des Lepidopteres. Vol. 6. Librairie Encyclopedique de Roret, Paris, pp. 29. [nº 30]","Poole, R. W. (1989) Noctuidae. In: Heppner, J. B. (Ed.), Lepidopterorum Catalogus. New Series. Vol. 118. Part. 1. E. J. Brill / Flora & Fauna Publications, Leiden, pp. 1 - 500.","Fibiger, M., Ronkay, L., Steiner, A. & Zilli, A. (2009) Noctuidae Europaeae. Vol. 11. Pantheinae, Dilobinae, Acronictinae, Eustrotiinae, Nolinae, Bagisarinae, Acontiinae, Metoponiinae, Heliothinae and Bryophilinae. Entomological Press, Soro, 504 pp.","Fibiger, M., Yela, J. L., Zilli, A., Ronkay, G. & Ronkay, L. (2011) Check list of the Quadrifid Noctuoidea of Europe. In: Witt, T. J. & Ronkay, L. (Eds.), Noctuidae Europaeae. Vol. 13. Entomological Press, Soro, pp. 23 - 44."]}

Rediscovery of Bryophila perloides On the night of 7 September 2010, a male specimen resembling B. perloides -according to the descriptions and illustrations in Guenée (1852), Rambur (1858 -[1866]) and Culot (1909-1913)- was photographed by Penny Hale at Finca La Molina, Casares, Málaga, Spain (30STF93) (Fig 1b). Following the suggestion of MF, the specimen was retained and sent to him for dissection. Examining the picture of the moth, JLY suggested B. perloides, an assumption shared by the rest of the team and confirmed by genitalia examination (Fig 6a). The genitalia showed crucial differences from those of Bryophila domestica (see e.g. Fibiger et al. 2009 and Fig 5a in this paper) and of Bryopsis muralis (Fig 5b; for taxonomic nomenclature, see Discussion), although they are much closer to those of B. domestica, from which it differs by the much shorter central claspers, wider and cygnated uncus and epiphallus broader and without distal spicules. The genitalia of one of the syntypes of B. perloides (Fig 6b) matched those of the specimen collected in Casares. It is to be noted that differences in width of the uncus between figs 6a and 6b are due to different pressure of the cover glass on the slides; same argument is applicable to the apparently different insertion position of the median claspers. The vesica is only partially everted in the syntype (Fig 6b) due to collapsing, probably related to the age of the specimen. Inspection of pictures taken by entomologists in S. Spain and uploaded to the Internet allowed us to find two additional specimens, both recorded at the Campo de Gibraltar, Algeciras, Cádiz, Spain, 30STE79, phot. José Manuel Gaona Ríos, one of them on 24.8.2018 (https://www.facebook.com/photo.php?fbid=1753590898021857&set=pcb.1753597151354565) (Fig 1c) and the other one on 21.9.2018 (https://www.facebook.com/photo.php?fbid=17850 87104872236&set=pcb.1785087411538872) (Fig 1d), showing on the forewings both key elements of most of the species of the perloides group, the basal omega-like design and the x-mark next to the reniform spot (red arrows in the figures)., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 55, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Guenee, A. (1852 - 1854) Noctuelites (II). In: Boisduval, J. A. & Guenee, A., Histoire naturelle des lnsectes. Species general des Lepidopteres. Vol. 6. Librairie Encyclopedique de Roret, Paris, pp. 29. [nº 30]","Rambur, P. (1858 - [1866]) Catalogue systematique des Lepidopteres d'Andalousie. Bailliere, Paris. 412 pp.","Fibiger, M., Ronkay, L., Steiner, A. & Zilli, A. (2009) Noctuidae Europaeae. Vol. 11. Pantheinae, Dilobinae, Acronictinae, Eustrotiinae, Nolinae, Bagisarinae, Acontiinae, Metoponiinae, Heliothinae and Bryophilinae. Entomological Press, Soro, 504 pp."]}

Polia (Polia) propodea McCabe, 1980 Polia propodea McCabe, 1980, New York State Museum Bulletin No. 432: 18–19. Type-locality: Canada, Columbia, Mt. Evans, Echo Lake. Taxonomic notes. According to the external appearance and genital configuration this species is closely related to the next, polytypic, disjunct Palaearctic species. Based on the barcode data they are possibly sister species., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 51, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["MCCABE, T. M. (1980): A reclassification of the Polia complex for North America (Lepidoptera: Noctuidae). - New York State Museum, Bulletin No. 432. i - vi, 1 - 141 pp."]}

Subgenus Polia Ochsenheimer, 1816 Polia Ochsenheimer, 1816, Die Schmetterlinge von Europe 4: 73. Type-species: Phalaena nebulosa Hufnagel, 1766, by subsequent designation by Curtis, 1829., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 48, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) vespertilio (Draudt, 1934) Aplecta vespertilio Draudt, 1934, in A. Seitz, Die Gross-Schmetterlinge der Erde 3: 109, pl. 14, row i. Type-locality: [Russia], Irkutsk, Tunkinski Mt. Lectotype: male, in coll. MNB. Taxonomic notes. According to the external look and the configuration of the genitalia of both sexes it can be considered as closely related to the former one.Their distribution is partly overlapping in the southern Siberian and northern Mongolian mountains; however it is more widely distributed both in West (southern Ural Mts) and East (Pacific mountains). Taxonomic interpretation of the Pacific (North Korean) population still requires further studies., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 52, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) griseifusa Draudt, 1950 Polia griseifusa Draudt, 1950, Mitteilungen der Münchner Entomologischen Gesellschaft 40: 30, pl. 2, fig. 14. Type-locality: China, Prov. Yunnan, Li-kiang. Lectotype, female, in coll. ZFMK. Taxonomic notes. This species is allied to the former one according to its external look and genital capsule. However, the bristles of the saccular processes are on both sides reduced. The configuration of the vesica is rather different, simply retroflexed dorsally with strong sclerotisation on the inner side and with a centrally positioned stripe of fasciculate cornuti. This species is known from W China (Yunnan, Sichuan) only and seems to be rather rare., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 49, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) tiefi Püngeler, 1914 Polia (Mamestra) tiefi Püngeler, 1914, Deutsche Entomologische Zeitschrift, Gesellschaft Iris zu Dresden 28: 38, pl. 2, fig. 22. Type-locality: [Russia] Sayan Mts, Munko-Sardyk. Lectotype: male, in coll. MNB. Taxonomic notes. This species was mentioned in the earlier publications as P. vesperugo (auct. nec Eversmann, HEYDEMANN 1944, HACKER et al. 2002) or P. schawerdae (Sheljuzhko, 1933). The synonymy with the latter species was established by VARGA et al. (2017). P. tiefi is widely distributed with some variation in the external appearance in the high mountains of NE Kazakhstan, South Siberia and Northern Mongolia (Saur, Tarbagatai, Sayan, Altai, Khangai and Khentij mountains), mostly in the upper mountain taiga zone., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 52, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["HEYDEMANN, F. (1944): Zur Kenntnis der Gattung Aplecta Guen. und zweiter \" Dual Species \" inderselben (Lep. Noct.). - Entomologische Zeitung des Entomologischen Vereins zu Stettin 105: 12 - 33.","HACKER, H., RONKAY, L. & HREBLAY, M. (2002): Hadeninae I. In: FIBIGER, M. (ed.): Noctuidae Europaeae, vol. 4. - Entomological Press, Soro, 456 pp.","VARGA, Z., RONKAY, G. & RONKAY, L. (2017): Revised taxonomic check list of the Eurasiatic species of the subtribe Poliina (Noctuidae, Noctuinae, Hadenini). - Deutsche Entomologische Zeitschrift 64 (2): 133 - 160. https: // doi. org / 10.3897 / dez. 64.21455"]}

Polia imbrifera (Guenée, 1852) Aplecta imbrifera Guenée, 1852, Histoire Naturelle des Insectes (Lépidoptères) 6: 76. Type locality: “ New York ”. Taxonomic notes. This species shows some simplification of the saccular processes with the reduction of the distal bundle of bristles on the left side. It means that the genital capsule secondarily seems to be more symmetrical than in other species of the nebulosa -group. It is widely distributed in the boreal and nemoral forest belts of North America and shows some clinal variation in colouration (MCCABE 1980)., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 57, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["MCCABE, T. M. (1980): A reclassification of the Polia complex for North America (Lepidoptera: Noctuidae). - New York State Museum, Bulletin No. 432. i - vi, 1 - 141 pp."]}

Polia (Polia) bombycina (Hufnagel, 1766) Phalaena bombycina Hufnagel, 1766, Berlinisches Magazin 3 (4): 410. Type-locality: [Germany] vic. of Berlin. Type (s) destroyed. Taxonomic notes. The Asiatic populations of P. bombycina clearly show a western-eastern subdivision with some overlap in Mongolia. The west Asiatic group of populations has two contrasting forms (subspecies?) at the eastern edge of the range, (i) the dark bronze-brownish Polia bombycina puengeleri Lehmann, 1998, described from [Kirghisia] Alexander Mts and occurring also in Kazakhstan, and (ii) the pale fawn-coloured Polia bombycina psammochrea Varga, 1974, described from Mongolia, Govi Altai aimak, Khasaght Khajrhan Mts. The taxonomic subdivision of the eastern group (cf. Polia bombycina grisea (Butler, 1878)) described under some different names (see: VARGA et al. 2017) needs further surveys, as well as the populations occurring in the Caucasus range., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 54, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["VARGA, Z., RONKAY, G. & RONKAY, L. (2017): Revised taxonomic check list of the Eurasiatic species of the subtribe Poliina (Noctuidae, Noctuinae, Hadenini). - Deutsche Entomologische Zeitschrift 64 (2): 133 - 160. https: // doi. org / 10.3897 / dez. 64.21455"]}

Polia (Polia) malchani (Draudt, 1934) Aplecta malchani Draudt, 1934, in A. Seitz, Die Gross-Schmetterlinge der Erde 3: 108, pl. 14, row i. Type-locality: [Russia] Transbaikalia, Malchan Mt, “ Borochojewa ”, 800 m. Holotype: female, in coll. MNB. Taxonomic notes. This is the only species of the group where the abdominal brush organ is fully reduced. It has a wide but obviously incompletely known area extending from the southern Ural Mts to the Russian Far East, Korea and Japan, seemingly without any geographical trend of variation., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on pages 52-53, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) lama (Staudinger, 1896) Mamestra nebulosa var. lama Staudinger, 1896, Deutsche Entomologische Zeitschrift, Gesellschaft Iris zu Dresden 9: 241. Type-locality: [Mongolia, Khangay Mts] Uliassutay. Holotype: male, in coll. MNB. Taxonomic notes.The subspecific subdivision of P. lama is published by VARGA et al. (2019)., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 55, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["VARGA, Z., RONKAY, G. & RONKAY, L. (2019): Taxonomic survey of the Polia (Polia) nebulosa species complex (Noctuidae, Noctuinae, Hadenini) with the description of two new subspecies. - Acta Zoologica Academiae Scientiarum Hungaricae 65 (2): 107 - 122. https: // doi. org / 10.17109 / AZH. 65.2.107.2019"]}

Polia piniae Buckett & Bauer, 1967 Polia piniae Buckett & Bauer, 1967, Journal for Research of Lepidoptera 5: 221. Type locality: Johnsville, Plumas Co., California (Davis). Taxonomic notes. This species and the former one are externally extreme similar. The female genitalia show the most constant differential characters.

Polia (Polia) nebulosa (Hufnagel, 1766) Phalaena nebulosa Hufnagel, 1766, Berlinisches Magazin 3 (3): 298. Type-locality: Germany, Berlin district. Taxonomic notes. The subspecific splitting of P. nebulosa is published in a separate article (VARGA et al. 2019)., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 55, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["VARGA, Z., RONKAY, G. & RONKAY, L. (2019): Taxonomic survey of the Polia (Polia) nebulosa species complex (Noctuidae, Noctuinae, Hadenini) with the description of two new subspecies. - Acta Zoologica Academiae Scientiarum Hungaricae 65 (2): 107 - 122. https: // doi. org / 10.17109 / AZH. 65.2.107.2019"]}

Polia (Polia) lamuta (Herz, 1903) Anarta lamuta Herz, 1903, Annuaire du Musée Zoologique de l’Académie Imperiale des Sciences de St.-Pétersbourg 8: 82. Type-locality: [Russia] Uruata Camp place; W of Verkhoyansk. Syntypes: 1 male, 1 female, in coll. ZISP. Taxonomic notes. The species is subdivided to two, morphologically and geographically clearly separated subspecies. The identity of the nominotypical lamuta with the south Siberian alpine (Sayan Mts) ssp. tunkinski O. Bang-Haas, 1927 was already suggested by HEYDEMANN (1944) who considered P. lamuta rangnowi Püngeler, 1909 from Lapland as valid subspecies. The ampulla is in P. lamuta lamuta much stronger than in P. l. rangnowi, terminally obtuse., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 50, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["HEYDEMANN, F. (1944): Zur Kenntnis der Gattung Aplecta Guen. und zweiter \" Dual Species \" inderselben (Lep. Noct.). - Entomologische Zeitung des Entomologischen Vereins zu Stettin 105: 12 - 33."]}

Polia (Polia) goliath (Oberthür, 1880) Dichonia goliath Oberthür, 1880, Études d’Entomologie 5: 68, pl. 6, fig. 7. Type-locality: [Russia] Askold [Island]. Holotype: male, in coll. BMNH. Taxonomic notes. The populations occurring in Taiwan represent a distinct taxon, the description of which will be presented in a separate article., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 55, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) atrax Draudt, 1950 Polia atrax Draudt, 1950, Mitteilungen der Münchner Entomologischen Gesellschaft 40: 31, pl. 2, fig. 17. Type-locality: China, Prov. Yunnan, Atuntse. Lectotype: male, in coll. ZFMK. Taxonomic notes. The species includes two geographic subspecies, ssp. atrax (Yunnan) and ssp. vargai Gyulai et Saldaitis, 2017 (Gansu) with slight but seemingly constant differences both in external and genital characters. The bundle of bristles on the left saccular process is already slightly differentiated and the right-sided process is shorter than in other species of this group and also somewhat more obtuse., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 53, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) hepatica (Clerck, 1759) Phalaena hepatica Clerck, 1759, Icones Insectorum Rariorum cum Nominibus eorum Trivialibus, Locisque e C. Linnaei 1: pl. 8, fig. 3. Type-locality: [?Europe] no locality given. Taxonomic notes. This species is somewhat similar to the former one in the general shape of genital capsule but the bundle of bristles on saccular processes is slightly more expressed on both sides. The vesica has a subbasal and subterminal helicoidal torsion but with a typical, long stripe of fasciculate cornuti. The species has an extended boreo-nemoral, Trans-Palaearctic distribution with numerous colour forms but without any clear subspecific subdivision., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 49, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia (Polia) rogenhoferi (Möschler, 1870) Mamestra rogenhoferi Möschler, 1870, Entomologische Zeitung, Entomologischen Vereine zu Stettin 31: 269. Type-locality: USA, Labrador. Taxonomic notes. A Nearctic arcto-boreal species with a montane exclave in Colorado., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 51, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia nimbosa (Guenée, 1852) Aplecta nimbosa Guenée, 1852, Histoire Naturelle des Insectes (Lépidoptères) 6: 77. Type locality: “Amerique Septentrionale”. Taxonomic notes.This species shows nearly the same colour variations as P. nebulosa and it was considered by HEYDEMANN (1944) as subspecies this species despite of the expressed differences in genitalia of both sexes and the reduction of the male brush organ. The most remarkable trend is in this species the rudimentation of the distal section of the left saccular process connected with the reduction of the distal bundle of bristles which also has some geographical variation. The species is widely distributed in the boreal and nemoral forest zones of North America and shows only some clinal variation with the exception of the Vancouver Island population (P. nimbosa mysticoides Barnes & Benjamin, 1924; see: MCCABE 1980)., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 56, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753, {"references":["HEYDEMANN, F. (1944): Zur Kenntnis der Gattung Aplecta Guen. und zweiter \" Dual Species \" inderselben (Lep. Noct.). - Entomologische Zeitung des Entomologischen Vereins zu Stettin 105: 12 - 33.","MCCABE, T. M. (1980): A reclassification of the Polia complex for North America (Lepidoptera: Noctuidae). - New York State Museum, Bulletin No. 432. i - vi, 1 - 141 pp."]}

Polia (Polia) serratilinea (Treitschke, 1825) Mamestra serratilinea Treitschke, 1825, Die Schmetterlinge von Europa 5 (2): 38. Type-locality: Austria, Vienna district. Lectotype: male, in coll. HNHM. Taxonomic notes. Numerous subspecies of this externally strikingly variable species are described. The nominotypical subspecies is restricted to the eastern periphery of the Austrian Alps. Further subspecies occur in the Central and western Alps, and Alsace (Mamestra serratilinea helvetica Schawerda, 1925), Balkan peninsula (Polia serratilinea kowatschevi Drenovsky, 1931), Central Anatolia (Polia serratilinea pinkeri Varga, 1974), and more clearly differentiated in Central Asia (Polia serratilinea spalax (Alphéraky, 1887), in the Hindukush-Himalaya area (Polia serratilinea tenebricosa Hacker et Weigert, 1990), but also in western Mongolia (Polia serratilinea eremorealis Varga, 1974). This species shows some convergence to P. propodea in the bifid terminal part of the right saccular process.The two lineages are, however, rather distant by their CO1 sequences. The asymmetrisation of the clavus and saccular processes show an evident geographical variation; its analysis and taxonomic interpretation will be treated in a further publication., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 53, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

The Polia nimbosa–imbrifera species-group Taxonomic notes. The four Nearctic species are closely allied to the Palaearctic members of the nebulosa group according to their external and genital morphological characters but also according to the sequences of the CO1 gene. The nearly parapatric sister species P. discalis and P. piniae occur mostly in the region of the Rocky Mountains while the other two species are widely distributed in the boreo-nemoral forest zone., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 56, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Polia discalis (Grote, 1870) Mamestra discalis Grote, 1877, In: Uhler, Bulletin of the U.S. Geological & Geographical Survey Territories 3: 797. Type locality: Clear Creek Canyon (U.S., Colorado). Taxonomic notes. P. discalis and P. piniae are sister species according to the morphological characters and also to the CO1 sequence. Abdominal coremata are present, the bundles of bristles on the left saccular process are separated, vesica with subbasal and medial diverticula. In female genitalia two signa are present only, while P. piniae has two stronger, stripe-like and two weaker signa., Published as part of Varga, Zoltán, Ronkay, Gábor, Nagy, Jenő & Ronkay, László, 2020, Contribution To The Taxonomy And Phylogeny Of The Genus Polia Ochsenheimer, 1816 (Noctuidae, Noctuinae, Hadenini): Species Groups And Pairs In The Holarctic Subgenus Polia S. Str., pp. 35-67 in Acta Zoologica Academiae Scientiarum Hungaricae 66 (1) on page 56, DOI: 10.17109/AZH.66.1.35.2020, http://zenodo.org/record/5734753

Deltote turcica sp. n. (Figs 1 a–d, 2 c–d, 3 a) Type material. Holotype (Fig. 1 b): Male, Ankara-Kayseri, 26.7.1965, 1000 m, Central Turkey, Asia minor, leg. M. u. W. Glaser, slide No. RL 12402m (coll. Vartian, NHMW). Paratypes. Turkey. 2 males (Fig. 1 a and 1 c), with the same data as holotype, slide Nos RL 12401m, RL 12403m (coll. NHMW). Turkmenistan. 1 male, 1 female, Yol-Dere, 15 km NE Kara-Kala, 12. VI.1953, leg. V. Kuznetsov, slide Nos RL 11679m, RL11687f (coll. HNHM); 2 males, Kopet-Dagh Mts, Sayvana valley, 1000 m, 56°50’E, 38°17’N, L60, 28. VI.1992, leg. Gy. Fábián, B. Herczig, A. Podlussány & Z. Varga (coll. G. Ronkay). Description. Adult (Figs 1 a–d): Small noctuid moth with slender body, wingspan 13–14 mm, forewing length 6–7 mm. Head —Antenna filiform in both sexes, dorsum covered with milky white scales; scape short and only ca twice as wide as basal segments of antenna. Eye large, globular, dark grey. Labial palp brown, short, upturned, third segment very short, pointed. Proboscis long, light brown. Head scales pale off white to light brown. Thorax — Ground colour similar to head, tegulae slender, metathorax somewhat more greyish. Legs gracile, brown, tibiae with longer milky white hair-scales, tarsi brown. Forewing: Ground colour and fringe light brown; basal and postmedial areas broad, white or brownish off-white; all lines wavy, terminal line beige to dark brown; orbicular stigma minute, rounded, dark brown with paler centre, reniform stigma fine, blackish, streak-like with fine white definition. Hindwing uniform beige, fringe brownish beige, terminal line narrow, black; discal spot, median line, and subterminal line dark, indistinct. Underside of forewing brown with dark subterminal line; that of hindwing pale brown, lacking lines and spots. Abdomen —Slender, cylindrical, pale brown, basal 3–4 segments covered with smooth beige-whitish scales, dorsal crest absent, anal tuft of male weak, ochreous-white. Male genitalia (Figs 2 c–d): Genital capsule very small, weakly sclerotised. Uncus thick, medium-long, thickened towards acutely pointed and hooked. Tegumen weakly sclerotised, penicular lobes reduced; juxta large, flask-shaped; vinculum short, broadly U-shaped. Valva rather short (only slightly longer than tegumen), tapering towards apex; cucullus apically rounded, sparsely setose, apical thorn strong but short, finely arched; corona absent; ampulla short, mushroom-shaped. Aedeagus as long as valva, thick, subapically broadest, laterally and ventrally extended; carina less sclerotized; vesica (uneverted) membranous, with large L-shaped (boot-shaped) plate flattened, heavily sclerotized, positioned in distal half of aedeagus. Female genitalia (Fig. 3 a): Papillae anales nearly as long as wide, densely setose with short setae; both pairs of gonapophyses short (not or only slightly longer than papillae anales). Ostium bursae broad; antrum membranous, broad, funnel-like, with sclerotised ring at junction to ductus bursae; ductus bursae long, broadened towards corpus bursae, with sclerotised quadrangular plate at anterior section; appendix bursae small, subconical, membranous; corpus bursae elliptical-ovoid, with large scobinate area medially; signum sclerotised, narrow, Y-shaped. Differential diagnosis. The two species of this group are similar externally, with subtle differences in the forewing shape and pattern. Deltote turcica (Figs 1 a–d) has, comparing with D. delicatula (Figs 1 e–h) slightly narrower and more acutely pointed forewings and the postmedial line is running more obliquely to inner margin. The more recently collected specimens are remarkably darker and greyer than the older (50–100 years old) moths. In the male genitalia, the valvae of D. turcica (Figs 2 c–d) are somewhat shorter, more slender and tapered distally than those of D. delicatula (Figs 2 a–b), and the apical thorn of the cucullus is shorter and finer in D. turcica than in D. delicatula. In the female genitalia, the ductus bursae of D. turcica (Fig. 3 a) is broader than in D. delicatula (Fig. 3 b), having a more sclerotised posterior ring and anterior plate. In addition, the signum of the corpus bursae of D. turcica is thicker and continuous, not separated into two parts as in D. delicatula. Distribution. Deltote turcica is known from two rather remote areas in central Turkey (Ankara-Kayseri region; the records of D. delicatula published by Baisch et al. [1998] from the Göreme valley and Ürgüp, and by, Koçak and Kemal (2018) from Nevşehir and Kırıkkale probably refer to this species as well) and from the Kopet- Dagh area in Turkmenistan. Biology and Habitat. The populations of D. turcica occurring in the Kopet-Dagh Mts. inhabit dry rocky slopes with open steppe vegetation and trees limited to small shrubby patches. The Turkish populations live in similarly xerothermic rocky sites. Specimens were collected in June and July. Etymology. The epitheton name turcica refers to Turkey and Turkmenistan, the states, where the new species is discovered.

Deltote delicatula (Christoph, 1882) (Figs 1 e–h, 2 a–b, 3 b) Erastria delicatula Christoph, 1882, Horae Societatis Entomologicae Rossicae 17: 112. Type locality: [Azerbaijan, Nakhichevan] Ordubad. Material examined. Azerbaijan. Two topotype ♂♂ specimens from Ordubad. The first specimen is preserved in the NHMW, its label data are as follows: " ♂ Ordub[ad], 21[.]5[.]83, ˇ" Kol. Vel. Kn. Nikolaia Mikhailovicha " (with Cyrillian letters), "coll. K. Höfer "; slide No. RL 12400m. The second specimen is illustrated on the homepage of the Zoological Institute, St. Petersburg (https://www.zin.ru/Collections/ Lepidoptera / specimen_en.html?Catalog_UID=1345391981073306) as " lectotype ", with the following label data: " ♂ Ordub[ad], 18[.]5[.]83", " Kol. Vel. Kn. Nikolaia Mikhailovicha " (with Cyrillian letters), The collecting date is interpreted as " 18.5.1880 " though the picture clearly indicates that the last number is "3" and not "0". Consequently, this latter specimen cannot be a syntype. Moreover, it is almost certain that these two specimens were collected by the same collector, at the same place within one week. Armenia. 1 ♂, Geghard, 40 km east of Yerevan, 1700 m, 3–11.IX.1975, leg. Vartian (coll. NHMW). Turkey. Siirt. 3♂♂, Akyamaç, 700 m, 5. VI.2015; 1♂, same data, 9. VI.2016; 1♀, Şirvan, Tatlıpayam, 1350 m, 14.VII.2013; 1♂, Botan valley, 700 m, 9.VII.2015; 1♂, 1♀, Şirvan, Tahvan, 625 m, 2.IX.2014; leg. E. Seven (coll. BTU). Bionomics and distribution. The early stages and the larval food plant are unknown. As the topotypical specimens were collected in May while the eastern Turkish and Armenien moths are in September, the species is considered as bivoltine. The habitat of D. delicatula in eastern Turkey is a steppe area with sparse Quercus (Fagaceae) and Paliurus (Rhamnaceae) trees and shrubs (Figs 4 a–b). Confirmed records of the D. delicatula are known from eastern Turkey (Prov. Siirt), Armenia (Geghard) and Azerbaijan (Nakhichevan: Ordubad). The published data of Deltote delicatula from northern Iran refer most probably also to the species but we had no opportunity to study Iranian specimens., Published as part of Seven, Erdem, Ronkay, László & Ronkay, Gábor, 2019, Description of a new species of Deltote Reichenbach, Leipzig, 1817 (Lepidoptera, Noctuidae, Eustrotiinae) from Turkey and Turkmenistan, pp. 171-178 in Zootaxa 4576 (1) on page 173, DOI: 10.11646/zootaxa.4576.1.10, http://zenodo.org/record/2624600, {"references":["Christoph, H. (1882) Einige neue Lepidoptera aus Russisch-Armenien. Horae societatis entomologicae rossicae, 17, 104 - 122."]}

Four new species of the genus Hermonassa from Pakistan (H. moorei sp. n., H. tathabaya sp. n. and H. ayubia sp. n.) and Nepal (H. kalamantra sp. n.) are described and compared with their relatives of the H. anthracina and H. dispila species groups. The specific status of Hermonassa shizukoae Sugi, 1995 is re-instated (stat. rev.). The species treatments are illustrated with 32 colour images and 24 genitalia figures.

The nefasta -group Diagnosis – A relatively poorly known species group since we have almost exclusively old, “classic” material from two (nefasta and optima) of the three species. Externally, the members of this lineage resemble the species of the sukharevae -group, but the structure of the male genitalia displays the closer relationship with the tancrei -group. Medium-sized (nefasta and optima) or large (leucostigma) moths with robust body, long and strongly bipectinated male antennae, relatively broad forewings with ochreous-greyish ground colour and regular maculation. In the male genitalia, the uncus is relatively short and acute, the ampulla is long and slightly falcate with acute or obtuse tip, the “neck” of cucullus is relatively long and the “head” is rounded, relatively small, without incision. Checklist Ctenoceratoda nefasta (Püngeler, 1907, Hadula), Deutsche Entomologische Zeitschrift. Gesellschaft Iris zu Dresden 19: 221, pl. 8, fig. 2. Type-locality: [China, Xinjiang] Lob-Noor; Ctenoceratoda optima (Alphéraky, 1897, Haderonia), in Romanoff, Mémoires sur les Lépidoptéres 9: 236, pl. 12, fig. 7. Type-locality: [China] “Oulan-boulak, Nan-chan”; Ctenoceratoda leucostigma Gyulai et Varga, 2010, Folia entomologica hungarica 70: 182, figs 1-3, gen. figs 9–13. Type-locality: China, Qinghai [Kuku-Noor region], 20 km N of Da Qaidam city. Bionomics and distribution – All species are confined to some extremely cold-continental parts of Inner Asia., Published as part of Varga, Zoltán, Gyulai, Péter, Ronkay, Gábor & Ronkay, László, 2018, Review Of The Species Groups Of The Genus Ctenoceratoda Varga, 1992 With Description Of Four New Species And A New Subspecies (Lepidoptera, Noctuidae), pp. 51-74 in Acta Zoologica Academiae Scientiarum Hungaricae 64 (1) on page 57, DOI: 10.17109/AZH.64.1.51.2018, http://zenodo.org/record/5734636