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Smicridea (Smicridea) lata R��zuri-Gonzales and Armitage, sp. n. Fig. 1. Diagnosis. Smicridea (S.) lata is most similar to S. (S.) soyatepecana Bueno-Soria, 1986 from Mexico, mainly based on the shape of tergum X and the phallic apparatus. Both species have a long narrow tergum X in lateral view with a dorsally-directed apex, but in dorsal view, the apex of tergum X in Smicridea (S.) lata is rounded whereas it is truncate in S. (S.) soyatepecana . Additionally, the phallic apparatus in lateral view in both species has a broad posteroventral process. However, this process is broader, directed dorsad, and with a rounded apex in Smicridea (S.) lata but directed posterad and with slightly truncate apex in S. (S.) soyatepecana . Furthermore, the posteroventral process of the phallobase in ventral view in Smicridea (S.) lata is narrowly divided, more slender apically, and with a pair of processes subapically and midlaterally, while in S. (S.) soyatepecana, this process is broadly divided, broader apically, and without subapical processes but with midlateral spines on the surface of the phallobase. Description. Adult male. Forewing length 4.1 mm (n = 1). Body color light brown (specimen preserved in alcohol). Head and thorax light brown with brown setae, dorsally thorax with dark brown marks anterad. Antennae and legs yellowish brown, legs with short brown setae. Maxillary palps 5-segmented; maxillary palps with fine light brown setae and few thicker dark brown setae dorsally on segments I and II. Forewing membrane brown, with brown setae and whitish transversal band in the nygmal area membrane, mostly denuded. Wing venation typical for the subgenus. Glands on abdominal segments VI and VIII oval. Male genitalia. Segment IX in lateral view (Fig. 1A) much longer dorsally than ventrally, anterolateral margin concave, anterodorsal corner not produced, broadly rounded; anterior portion of dorsal margin slightly concave, posterior portion membranous; ventrolateral margin oblique, sinuous; ventral margin short, rounded; in dorsal view (Fig. 1B) anterior margin concave, lateral margin sinuous, inflated mesally, mesally bearing microsetae. Tergum X in lateral view (Fig. 1A) narrow, setose, much shorter than segment IX, apex truncate, posterodorsal corner produced dorsad; in dorsal view (Fig. 1B) narrow, with rounded apices, separated by mesal notch, less than half its length, membranous mesally. Inferior appendage (Figs. 1 A���C) two-segmented; first segment extending beyond posterior margin of tergum X, covered with setae, inflated apically; second segment digitate, approximately 1/3 as long as first segment, with short setae on lateral margin and very fine setae on dorsal and mesal surfaces, in dorsal view apex acute, slightly inflated on distal half. Phallic apparatus (Figs. 1 D���E) with phallobase enlarged, open ventrally; apical section open, dorsolateral corner of phallobase rounded, entire; posterodorsal margin of phallobase mesally produced, spatulate; posteroventral margin produced into broad digitate, entire lobe, with rounded apex, directed dorsad, with pair of midlateral processes: (i) a subtriangular process bearing spicules, and (ii) a short subapical process; in ventral view, posteroventral process deeply notched, notch basally narrow, each lobe slender, rounded apically; endothecal membranes with two pairs of long slender acute spines, curved dorsad: (i) dorsal pair shorter than ventral pair, bearing spicules on dorsal surface, and (ii) ventral spines twice as long as dorsal pair. Female and immature stages. Unknown. Distribution. Panama (Veraguas Province). Etymology. From the Latin word latus, referring to the very broad posteroventral process of the phallobase in lateral view. Material examined. HOLOTYPE MALE. PANAMA: Veraguas: Santa Fe National Park, R��o Mulaba, afluente del primer brazo, 8.51706��N, 81.12140��W, 770 m, 19���23.iv.2017, T. R��os, E. ��lvarez, and C. Nieto, Malaise trap (COZEM)., Published as part of R��zuri-Gonzales, Ernesto & Armitage, Brian J., 2019, The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks, pp. 1-13 in Insecta Mundi 710 (710) on pages 4-5, DOI: 10.5281/zenodo.3674835, {"references":["Bueno-Soria, J. 1986. Estudios en insectos acuaticos VII: Cinco nuevas especies de tricopteros de Mexico y Costa Rica (Trichoptera: Hydropsychidae). Folia Entomologica Mexicana 68: 53 - 65."]}

Smicridea (Smicridea) spatulata R��zuri-Gonzales and Armitage, sp. n. Fig. 2. Diagnosis. Smicridea (S.) spatulata sp. n. is most similar to S. (S.) campana Flint, 1974 based on the shape of tergum X, especially in dorsal view; the shape of the phallobase in lateral view; and the presence of a semi-membranous ring-like structure in the endothecal membranes. However, Smicridea (S.) spatulata can be distinguished from S. (S.) campana due to its slightly broader tergum X subapically in dorsal view; the crenate dorsolateral corner of the phallobase, which is straight and smooth in S. (S.) campana; the presence of a short spine on the mesal surface of the phallobase; and the broader and more elongate posteroventral process of the phallobase in lateral view, which is shorter and more slender in S. (S.) campana . Additionally, Smicridea (S.) spatulata has five endothecal spines, whereas S. (S.) campana only has two pairs of spines of different size and shapes. Description. Adult male. Forewing length average 4.0 mm (n = 2). Body color light brown (specimen preserved in alcohol). Head and thorax light brown with light brown setae, dorsally thorax with dark brown marks anterad. Antennae and legs yellowish brown, legs with short brown setae. Maxillary palps 5-segmented; maxillary palps with long fine light brown setae. Forewing membrane light brown, with brown setae, without apparent whitish transversal band in the nygmal area membrane, mostly denuded. Wing venation typical for the subgenus. Abdominal segments VI and VIII glands oval, larger than containing segment. Male genitalia. Segment IX in lateral view (Fig. 2A) much longer dorsally than ventrally, anterolateral margin concave, anterodorsal corner produced, broadly rounded; anterior portion of dorsal margin slightly concave, mesal portion of dorsal margin convex, bearing small microsetae, closer to tergum X than to anterolateral margin of segment IX, posterior portion membranous; ventrolateral margin oblique, sinuous; ventral margin short, rounded; in dorsal view (Fig. 2B) anterior margin concave, lateral margin sinuous, mesally bearing microsetae. Tergum X in lateral view (Fig. 2A) narrow, setose, much shorter than segment IX, apex rounded; in dorsal view (Fig. 2B) narrow, with rounded apices, separated by mesal notch, less than half its length, membranous mesally. Inferior appendage (Figs. 2 A���C) two-segmented; first segment extending beyond posterior margin of tergum X, covered with setae, inflated apically; second segment digitate, approximately 1/3 as long as first segment, with short setae on margins, in dorsal view apex acute, slightly inflated on distal half. Phallic apparatus (Figs. 2 D���E) with phallobase enlarged, open ventrally; apical section open, dorsolateral corner of phallobase rounded, crenate, with thick short spine on mesal surface; posterodorsal margin of phallobase mesally produced, acute; posteroventral margin produced into slender digitate, entire lobe, with subacute apex, directed posterad, without spines or processes on lateral surface; in ventral view, posteroventral process entire, spatulate, slightly emarginate apically; endothecal membranes with five acute spines and oval lightly sclerotized ���ring���: (i) anterior most pair of spines curved ventrad in lateral view, (ii) mesal pair of spines, curved dorsad, rounded basally and tapering towards apex, slightly longer than anterior most pair, (iii) posteroventral spine acute, with sinuous margins. Female and immature stages. Unknown. Distribution. Panama (Cocl�� Province). Etymology. From the Latin word spatulata, referring to the spatulate posteroventral process of the phallobase in ventral view. Material examined. HOLOTYPE MALE. PANAMA: Cocl��: Omar Torrijos National Park, Quebrada Corazones, 8.6776��N, 80.6001��W, 728 m, 22���26.iii.2017, A. Cornejo, T. R��os, E. ��lvarez, and E. P��rez, Malaise trap (COZEM). PARATYPE. Same data as holotype, 1♂ (COZEM)., Published as part of R��zuri-Gonzales, Ernesto & Armitage, Brian J., 2019, The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks, pp. 1-13 in Insecta Mundi 710 (710) on pages 5-6, DOI: 10.5281/zenodo.3674835

R��zuri-Gonzales, Ernesto, Armitage, Brian J. (2019): The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks. Insecta Mundi 710 (710): 1-13, DOI: http://doi.org/10.5281/zenodo.3674835, {"references":["Alves, A. A., A. P. M. Santos, and J. L. Nessimian. 2018. A new species of Smicridea McLachlan 1871 (Trichoptera: Hydropsychidae) from the Caatinga biome and new distributional records from Brazil. Zootaxa 4500(3): 338-396.","Armitage, B. J., S. C. Harris, R. J. Blahnik, A. Cornejo, and T. I. Arefina-Armitage. 2018. The Trichoptera of Panama VII. Additional new records for caddisflies from the Republic of Panama. Insecta Mundi 0614: 1-7.","Banks, N. 1913. Some new neuropteroid insects. Journal of the New York Entomological Society 11: 236-243.","Blahnik, R. J. 1995. New species of Smicridea (subgenus Smicridea) from Costa Rica, with a revision of the fasciatella complex (Trichoptera: Hydropsychidae). Journal of the North American Benthological Society 14(1): 84- -107.","Blahnik, R. J., and R. W. Holzenthal. 2004. Collection and curation of Trichoptera, with an emphasis on pinned material. Nectopsyche, Neotropical Trichoptera Newsletter 1: 8-20.","Bueno-Soria, J. 1986. Estudios en insectos acuaticos VII: Cinco nuevas especies de tricopteros de Mexico y Costa Rica (Trichoptera: Hydropsychidae). Folia Entomologica Mexicana 68: 53-65.","Brown, B. V. 2013. Automating the \"Material examined\" section of taxonomic papers to speed up species descriptions. Zootaxa 3683(3): 297-299.","Dallwitz, M. J., T. A. Paine, and E. J. Zurcher. 2016. User's guide to the DELTA editor. Available at http://delta-intkey.com/www/delta-ed.htm. (Last accessed November 2018.)","Flint, O. S., Jr. 1974a. Studies of Neotropical caddisflies, XVII: the genus Smicridea from North and Central America (Trichoptera: Hydropsychidae). Smithsonian Contributions to Zoology 167: 1-65.","Flint, O. S., Jr. 1974b. Checklist of the Trichoptera, or caddisflies, from Chile. Revista Chilena de Entomologia 8: 83-93.","Flint, O. S., Jr., and D. G. Denning. 1989. Studies of Neotropical caddisflies, XL: new species of Smicridea (Smicridea) from Middle America and the West Indies (Trichoptera: Hydropsychidae). Proceedings of the Biological Society of Washington 102(2): 418-433.","Hagen, H. 1861. Synopsis of the Neuroptera of North America. Smithsonian Miscellaneous Collections 4: 1-347.","Holzenthal, R. W., and R. J. Blahnik. 1995. New species of Smicridea (Rhyacophylax) (Trichoptera: Hydropsychidae) from Costa Rica. Entomological News 106(5): 213-223.","Holzenthal, R. W., and A. R. Calor. 2017. Catalog of the Neotropical Trichoptera (Caddisflies). Zoo- Keys 654: 1-566.","Holzenthal, R. W., R. E. Thomson, and B. Rios-Touma. 2015. Chapter 38: Order Trichoptera. p. 965-1002. In: J. H. Thorp and D. C. Rogers (eds.). Thorp and Covich's Freshwater Invertebrates. Academic Press, New York. 836 p.","Jarvis, A., H. I. Reuter, A. Nelson, and E. Guevara. 2008. Hole-filled seamless SRTM data V4. International Centre for Tropical Agriculture (CIAT). Available at http://srtm.csi.cgiar.org. (Last accessed August 2018.)","Kondratieff, B., and B. J. Armitage. 2019. The Plecoptera of Panama III. The genus Anacroneuria (Plecoptera: Perlidae) in Panama's national parks: 2017 survey results. Zootaxa 4565(3): 407-419.","Martynov, A. V. 1912. On two collections from Peru. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de Saint Petersbourg 17: 1-40.","McElravy, E. P., V. H. Resh, H. Wolda, and O. S. Flint, Jr. 1981. Diversity of adult Trichoptera in a 'non-seasonal' tropical environment. p. 149-156. In: G. P. Moretti (ed.) Proceedings of the 3rd International Symposium on Trichoptera. Dr. W. Junk; The Hague. 500 p.","McLachlan, R. 1871. On new forms, etc., of extra-European Trichopterous insects. Journal of Linnean Society of London, Zoology 11: 98-141.","Mey, W., and R. Ospina-Torres. 2018. Contribution to the Trichoptera fauna of the river La Vieja, Bogota, Colombia (Insecta: Trichoptera). Zootaxa 4504(1): 023-040.","Morse, J. C. 1975. A phylogeny and revision of the caddisfly genus Ceraclea (Trichoptera, Leptoceridae). Contributions of the American Entomological Institute 11(2): 1-94.","Natural Earth. 2018. Free vector and raster map data. Available at http://naturalearthdata.com. (Last accessed 2019)","QGIS Development Team. 2018. QGIS Geographic Information System. Open Source Geospatial Foundation Project. Available at https://qgis.osgeo.org. (Last accessed 2019)","Razuri-Gonzalez, E., and R. W. Holzenthal. 2016. New synonyms in the highly diverse caddisfly genus Smicridea (Trichoptera: Hydropsychidae). ZooKeys 637: 21-31.","Sganga, J. V., and F. M. Gibon. 2018. New Species, New Records, and Distribution of Smicridea McLachlan 1871 in Bolivia (Trichoptera: Hydropsychidae). Neotropical Entomology. DOI: 10.1007/ s13744-018-0645-0.","Ulmer, G., and A. Thienemann. 1909. Sudamerikanische Trichopteren aus den Kopenhagener Museum. Deutsche Entomologische Zeitschrift 1909: 305-311."]}

Smicridea (Smicridea) varia (Banks, 1913) Primary Literature Reference: Flint 1974 a Distribution: Costa Rica *, Ecuador, Guatemala, Mexico, Nicaragua, Panama, Published as part of R��zuri-Gonzales, Ernesto & Armitage, Brian J., 2019, The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks, pp. 1-13 in Insecta Mundi 710 (710) on page 4, DOI: 10.5281/zenodo.3674835, {"references":["Banks, N. 1913. Some new neuropteroid insects. Journal of the New York Entomological Society 11: 236 - 243.","Flint, O. S., Jr. 1974 a. Studies of Neotropical caddisflies, XVII: the genus Smicridea from North and Central America (Trichoptera: Hydropsychidae). Smithsonian Contributions to Zoology 167: 1 - 65."]}

Smicridea (Smicridea) polyfasciata Martynov, 1912 Primary Literature Reference: Armitage et al. 2018 Distribution: Bolivia, Colombia, Ecuador, Panama, Peru *, Published as part of R��zuri-Gonzales, Ernesto & Armitage, Brian J., 2019, The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks, pp. 1-13 in Insecta Mundi 710 (710) on page 4, DOI: 10.5281/zenodo.3674835, {"references":["Martynov, A. V. 1912. On two collections from Peru. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de Saint Petersbourg 17: 1 - 40.","Armitage, B. J., S. C. Harris, R. J. Blahnik, A. Cornejo, and T. I. Arefina-Armitage. 2018. The Trichoptera of Panama VII. Additional new records for caddisflies from the Republic of Panama. Insecta Mundi 0614: 1 - 7."]}

Smicridea (Smicridea) dividua R��zuri-Gonzales and Armitage, sp. n. Fig. 3. Diagnosis. Smicridea (S.) dividua sp. n. is most similar to S. (S.) cholta Flint, 1974 from Guatemala and Nicaragua based on the shape of tergum X and the phallic apparatus in ventral view. However, Smicridea (S.) dividua is easily separated from S. (S.) cholta due to the bilobed posteroventral process of the phallobase, which is broad and entire in S. (S.) cholta, and the posterodorsal margin of the phallobase, which is produced into a slender lobe in the new species, but is undulate and bearing a few short spines on the lateral surface in S. (S.) cholta . Furthermore, S. (S.) dividua has a single sinuous spine, which shape is unique in this group whereas S. (S.) cholta has a pair of long slender spines. Description. Adult male. Forewing length average 4.3 mm (n = 7). Body color light brown (specimen preserved in alcohol). Head and thorax light brown with brown setae, dorsally thorax with dark brown marks anterad. Antennae and legs yellowish brown, antennae with short brown setae, legs with long brown setae. Maxillary palps 5-segmented; maxillary palps with long brown setae and a few thick long setae anteroventrally on segment II. Forewing membrane light brown, with brown and dark brown setae, with whitish transversal band in the nygmal area membrane, mostly denuded. Wing venation typical for the subgenus. Abdominal segments VI and VIII glands oval, larger than containing segment. Male genitalia. Segment IX in lateral view (Fig. 3A) much longer dorsally than ventrally, anterolateral margin slightly concave, anterodorsal corner not produced, forming nearly straight angle; anterior portion of dorsal margin long, slightly concave, mesal portion of dorsal margin convex, bearing small microsetae, closer to tergum X than to anterolateral margin of segment IX, posterior portion membranous; ventrolateral margin oblique, sinuous; ventral margin short, truncate; in dorsal view (Fig. 3B) anterior margin strongly concave, lateral margin sinuous, inflated mesally, mesally bearing microsetae. Tergum X in lateral view (Fig. 3A) narrow, setose, much shorter than segment IX, apex truncate, posterodorsal corner produced dorsad; in dorsal view (Fig. 3B) narrow, with rounded apices, separated by mesal notch, less than half its length, membranous mesally. Inferior appendage (Figs. 3 A-C) two-segmented; first segment not extending beyond posterior margin of tergum X, covered with setae, inflated apically; second segment digitate, approximately 1/2 as long as first segment, with short setae on margins, in dorsal view apex acute, slightly inflated on distal half. Phallic apparatus (Figs. 3 E-D) with phallobase enlarged, open ventrally; apical section open, dorsolateral corner of phallobase produced into a slender lobe; posterodorsal margin of phallobase mesally produced, acute; posteroventral margin produced into two slender digitate lobes, the ventral one with subacute apex, directed laterad and the dorsal one with bilobed apex, directed dorsolaterad, without spines or processes on lateral surface; in ventral view, posteroventral process deeply notched, notch basally rounded and undulate, each lobe slender, bilobed apically, directed laterad; endothecal membranes in lateral view with a long, sinuous, apically acute spine, in ventral view, bilobed, broad basally and slender apically, deeply notched mesally. Female and immature stages. Unknown. Distribution. Panama (Cocl�� Province). Etymology. From the Latin word dividuus, referring to the unique divided endothecal spine. Material examined. HOLOTYPE MALE. PANAMA: Cocl��: Omar Torrijos National Park, Quebrada La Yayas, 8.66168��N, 80.5952��W, 586 m, 22���26.iii.2017, E. ��lvarez, E. P��rez, and T. R��os, Malaise trap (COZEM). PARATYPES. Same data as holotype, 6♂ (COZEM, UMSP)., Published as part of R��zuri-Gonzales, Ernesto & Armitage, Brian J., 2019, The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks, pp. 1-13 in Insecta Mundi 710 (710) on pages 6-7, DOI: 10.5281/zenodo.3674835