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Ivmaka longicaudatus Xu & Dietrich, sp. nov. (Figs. 1–14) Type species: Holotype. J (UCD), Papua New Guinea Gulf: Ivimka Res. Station Lakekamu Basin, 120m 7’44’S 146’30’E, 7–8, March, 2000, T. Sears, MT. Paratypes. 2J (INHS), same data as holotype. Description. Body length: male 3.1–3.3 mm. General color of body orange brown (Figs. 1–3). Vertex orange yellow; coronal suture brown, with pair of irregular dark brown spots on each side (Figs. 1–3). Eyes silvery gray to black (Figs. 1–3). Ocelli surrounded by yellowish creamy patches (Fig. 3). Face orange to pale yellow from vertex to anteclypeus; lateral frontal suture brown (Fig. 3). Pronotum with median yellow transverse band bordered by thin dark brown lines, anterior margin white, posterior part tan (Fig. 1). Mesonotum with basolateral triangles black, otherwise dark brown (Fig. 1). Forewing dark brown, subhyaline distally with veins brown and contrasting with pale membrane; hind wing transparent, veins brown (Figs. 1, 2). Legs orange except the pretarsus dark brown (Figs. 1–3). Male 2S abdominal apodemes parallel, moderately broad, reaching middle of segment V (Fig. 4). Male pygofer bearing 2–3 long and 10–11 small rigid setae on dorsal margin, group of 17–20 feeble microsetae near middle of lobe; distal process of pygofer lobe nearly straight and evenly tapered; dorsal bridge extended almost four-fifths length of lobe (Figs. 7, 8). Anal tube process slender, sinuate, apex sharply narrowed to point hooked anterad (Fig. 9). Style sinuate, dentifer bearing about 13 tiny teeth preceded by 2 fine setae and few sensory pits (Fig. 10). Connective 1amellate, with medial sclerotization; anterior margin straight or weakly concave; lateral margin distinctly constricted (Fig. 11). Subgenital plate in lateral view with apex round and bent dorsad along ventral margin of pygofer lobe; basal setae (7–8) uniseriate, distinctly longer than marginal setae, marginal setae (29–30) arranged in 1–2 rows occupying nearly half length of dorsal margin, macrosetae (21–22) arising near based of plate, terminated at marginal setae, feeble microsetae (54–60) irregularly arranged in 3–6 irregular rows (Figs. 7, 12). Aedeagus slender, V-shaped in lateral view, shaft with few tiny teeth dorsally, wide basally, then abruptly narrowed, apex bifid in posterior view giving rise to pair of short distally directed processes (Figs. 13, 14). Etymology. The specific epithet is a compound Latin adjective “ longicaudatus ”, which refers to the elongate male pygofer of the new species. Distribution. Papua New Guinea., Published as part of Xu, Ye, Zhang, Peng-Wen, Dietrich, Christopher H. & Qin, Dao-Zheng, 2023, A new genus of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae) from Papua New Guinea, pp. 175-179 in Zootaxa 5271 (1) on pages 176-177, DOI: 10.11646/zootaxa.5271.1.9, http://zenodo.org/record/7864498

Ivmaka Xu & Dietrich, gen. nov. Type species: Ivmaka longicaudatus Xu & Dietrich sp. n., here designated. Description. Body robust. Head including eyes broader than pronotum in dorsal aspect, crown short and broad, round anteriorly, length less than half width between eyes, coronal suture long, restricted by vertex and postfrons, dorsal part of the frontoclypeus (above antennal pits) with median longitudinal suture continuous with coronal suture (Figs. 1, 3). Ocelli distinct, located on anterior margin of crown midway between eyes and coronal suture (Fig. 3). Face moderately broad; lateral frontal suture well delimited, curved mesad above antennal pit and meeting epistomal suture at midline ventromesad of ocelli; anteclypeus slightly convex, not expanded; rostrum distinctly overlapping base of hind coxae (Fig. 3). Pronotum large, almost twice as long as crown, with sinuate transverse depression (Fig. 1). Forewing narrow, round apically, apical cells occupying nearly one-third of total length, c and r cells nearly equal in width, both narrower than m and cua cells; vein R2 and RM arising from r cell, connected by cross-vein, vein MCu arising from m cell, veins RM and MCu evenly curved and almost parallel throughout their length (Fig. 5). Hind wing with CuA unbranched (Fig. 6). Front femur seta AM1 stout, situated near ventral margin; intercalary row with one large basal seta and five smaller setae more distad. Hind femur with macrosetal formula 2+1+1. Hind-tibia row AV with six preapical macrosetae (Figs. 1–3). Male 2S abdominal apodemes well developed (Fig. 4). Male pygofer elongate, posterior margin extended into sharp point, with conspicuous basolateral setal group and macrosetae, distal setal group scattered on each side of lobe, dorsal bridge well delimited, ventral appendage absent (Figs. 7, 8). Anal tube process well developed, extended to ventral margin of pygofer lobe (Fig. 9). Style broadened at base, sinuate in ventral and lateral views, apex tapered, dentifer adorned with tiny teeth and preapical setae (Fig. 10). Connective much longer than broad, caudal margin notched medially (Fig. 11). Subgenital plate extended beyond main part of pygofer lobe but not surpassing distal process, broadest near base and slightly trapered and upturned apically, basolateral lobe rounded; four categories of setae present: basal setae slim, arranged in row along basolateral lobe; marginal setae well defined but somewhat irregularly arranged, limited to distal part of plate margin; macrosetae uniseriate, reaching apex of plate; feeble microsetae numerous, scattered, longer than macrosetae (Figs. 7, 12). Aedeagus shaft tubular, preatrium short, dorsoatrium well developed, gonopore apical on ventral surface (Figs. 13, 14). Notes. This new genus will run to Mindanaoa Mahmood, 1967 and Smyga Dworakowska, 1995 in the generic key of Xu et al. (2021) but differs from Mindanaoa Mahmood and Smyga Dworakowska in presence of a well-developed dorsoatrium on the aedeagus (Fig. 13) (dorsoatrium absent in Mindanaoa Mahmood and Smyga Dworakowska). Among other previously described genera in Empoasca -group, this new genus is superficially similar to Dapitana Mahmood, 1967 and Serratulus Mahmood, 1967 in having the coronal suture well developed (Fig. 3), and vein R2 and RM connected by a cross-vein in the forewing (Fig. 5). However, this new genus differs from Dapitana Mahmood in having the pygofer with a conspicuous basolateral setal group (Fig. 7) and vein ScP+RA present on the forewing (Fig. 5) (absent in Dapitana Mahmood). It differs from Serratulus Mahmood, 1967 in having the subgenital plate with macrosetae uniseriate (Fig. 12) (vs. arranged in two rows). Furthermore, this new genus is unique in having the pygofer lobe extended into a distal spine-like process surpassing the subgenital plate (Figs. 7, 8). Distribution. Papua New Guinea. Etymology. The generic name alludes to the locality of the type material (Ivimka Res. Station). Gender: masculine., Published as part of Xu, Ye, Zhang, Peng-Wen, Dietrich, Christopher H. & Qin, Dao-Zheng, 2023, A new genus of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae) from Papua New Guinea, pp. 175-179 in Zootaxa 5271 (1) on page 176, DOI: 10.11646/zootaxa.5271.1.9, http://zenodo.org/record/7864498, {"references":["Mahmood, S. H. (1967) A study of the typhlocybine genera of the Oriental region (Thailand, the Philippines and adjoining areas). Pacific Insects Monographs, 12, 1 - 52.","Dworakowska, I. (1995) Szara gen. nov. and some other Oriental Empoascini (Insecta: Auchenorrhyncha: Cicadellidae: Typhlocybinae). Entomologische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 56, 129 - 160.","Xu, Y., Dietrich, C. H., Zhang, Y. L. Dmitriev, D. A., Zhang, L., Wang, Y. M., Lu, S. H. & Qin, D. Z. (2021) Phylogeny of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) based on morphological characteristics, with reclassification of the Empoasca generic group. Systematic Entomology, 46, 266 - 286. https: // doi. org / 10.1111 / syen. 12461"]}

Nikomiklukha Gnezdilov, 2010 Type species: Issus praecedens Walker, 1857; by original designation by Gnezdilov 2010: 48. Sarimissus Wang, Zhang & Bourgoin, 2019: Gnezdilov, 2019: 1302 [sic!; error]2010 Gnezdilov, p. 48 [new genus] 2019 Gnezdilov, p. 1302 Composition: Nikomiklukha praecedens (Walker, 1857), Nikomiklukha sumatrana Gnezdilov, 2010, Nikomiklukha maclayi Gnezdilov, 2010., Published as part of Qin, Dao-Zheng & Stroi��ski, Adam, 2022, Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae), pp. 247-250 in Zootaxa 5093 (2) on page 248, DOI: 10.11646/zootaxa.5093.2.8, http://zenodo.org/record/5905274, {"references":["Gnezdilov, V. M. (2010) Three new genera and three new species of the family Issidae (Hemiptera: Fulgoromorpha) from Borneo and Sumatra. Tijdschrift voor Entomologie, 153, 41 - 52.","Walker, F. (1857) Catalogue of the Homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society. London, 1 (4), 141 - 175.","Wang, M. L., Zhang, Y. L. & Bourgoin, T. (2019) On the tribe Sarimini with two new genera from south of China (Hemiptera, Fulgoromorpha, Issidae). Zootaxa, 4706 (2), 375 - 383. https: // doi. org / 10.11646 / zootaxa. 4706.2.10","Gnezdilov, V. M. (2019) On the synonymy and distribution of the planthopper genera Euroxenus Gnezdilov, 2009 and Nikomiklukha Gnezdilov, 2010 (Hemiptera, Auchenorrhyncha, Fulgoroidea: Issidae). Entomological Review, 99 (9), 1299 - 1303. https: // doi. org / 10.1134 / S 0013873819090070"]}

Qin, Dao-Zheng, Stroiński, Adam (2022): Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae). Zootaxa 5093 (2): 247-250, DOI: https://doi.org/10.11646/zootaxa.5093.2.8

Sarimissus Wang, Zhang et Bourgoin, 2019 Type species. Sarimissus maculifrons Wang, Zhang et Bourgoin, 2019: 377; by original designation and monotypy. 2019 Wang, Zhang & Bourgoin, p. 376 [new genus, description] 2019 Gnezdilov, p. 1302 [synonymy under Nikomiklukha Gnezdilov, 2010] 2020 Nikomiklukha: Chang et al. 2020, p. 33 Composition. Monotypic genus��� Sarimissus maculifrons Wang, Zhang & Bourgoin, 2019., Published as part of Qin, Dao-Zheng & Stroi��ski, Adam, 2022, Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae), pp. 247-250 in Zootaxa 5093 (2) on page 248, DOI: 10.11646/zootaxa.5093.2.8, http://zenodo.org/record/5905274, {"references":["Wang, M. L., Zhang, Y. L. & Bourgoin, T. (2019) On the tribe Sarimini with two new genera from south of China (Hemiptera, Fulgoromorpha, Issidae). Zootaxa, 4706 (2), 375 - 383. https: // doi. org / 10.11646 / zootaxa. 4706.2.10","Gnezdilov, V. M. (2010) Three new genera and three new species of the family Issidae (Hemiptera: Fulgoromorpha) from Borneo and Sumatra. Tijdschrift voor Entomologie, 153, 41 - 52.","Chang, Z. - M., Yang, L. & Chen X. - S. (2020) Two new genera with species of the tribe Sarimini (Hemiptera, Fulgoromorpha, Issidae) from China. ZooKeys, 956, 31 - 47. https: // doi. org / 10.3897 / zookeys. 956.47784"]}

Homa sinensis Qin & Zhang, 2011 (Figs 81���85) Homa sinensis Qin & Zhang, 2011 in Qin et al., 2011: 32. Materials examined: 1♂ (holotype), China Yunnan Province, Bubeng, Mengla, December 15, 1999, I. Dworakowska (NWAFU); 3♂♂, Thailand Khonkaen Nam Pong NP office, 16��37.201'N, 102��34.481'E, 218 m, Malaise trap, July 26-August 2, 2006, Khamphol Jaidee (NWAFU); 2♂♂, Thailand, Phetchabun Nam Nao NP Heliport, 16��43.156'N, 101��35.118'E, 890 m, Pan trap, December 21���22, 2006, Noopean Hongyothi & Leng Janteab (INHS); 2♂♂, Thailand Phetchabun Nam Nao NP Heliport, 16��43.156'N, 101��35.118'E, 890 m, Pan trap, December 21���22, 2006, Noopean Hongyothi & Leng Janteab (QSBG)., Published as part of Xu, Ye, Wang, Yi-Mei, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2022, Revision of the genus Homa Distant (Hemiptera: Cicadellidae: Typhlocybinae), pp. 179-190 in Zootaxa 5087 (1) on page 189, DOI: 10.11646/zootaxa.5087.1.8, http://zenodo.org/record/5819981, {"references":["Qin, D. - Z., Liu, Y., Zhang, Y. - L. (2011) A taxonomic study of Chinese Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) (III). Zootaxa, 3094 (1), 30 - 42. https: // doi. org / 10.11646 / zootaxa. 3094.1.2"]}

Homa osificata Xu, Dietrich & Qin sp. nov. (Figs 5���8, 26���28, 46���53) Holotype. ♂, Thailand Phetchabun Nam Nao NP Heliport, 16��43.156'N, 101��35.118'E, 890 m, Pan trap, December 21���22, 2006, Noopean Hongyothi & Leng Janteab (QSBG). Paratypes. 2♂♂, same data as holotype (NWAFU); 13♂♂, same data as holotype (INHS); 12♂♂, Nakhon Nayok Khao Yai NP Behind Football field, 14��24.619'N, 101��22.778'E, 770 m, Malaise trap, July 5���12, 2006, Pong Sandao (QSBG). Description. Measurements. Body length: male: 3.8���4.1 mm. Ground color red. Anteclypeus slightly broader in male than in female (Fig. 8). 1T apodemes forming arc bending apicad, not extended to segment II (Fig. 26). Male pygofer in profile with apex rounded, bearing 6���7 rigid microsetae along posterior margin, dorsal bridge well developed (Figs 27, 28, 46���48). Subgenital plate far exceeding pygofer lobe, gradually curved toward pygofer ventral margin; basal setae (7���8) situated on broadest section of plate, two or three setae separated from bristle clusters; marginal setae (6���8) sparse, arranged in single row; macrosetae (5) discrete, not extended to tip of plate; feeble microsetae (10���12) inconspicuous (Figs 27, 46, 49). Aedeagus shaft tubular, with attenuate distal extension and pair of shorter asymmetrical processes subapically, one rounded and longer, other shorter and pointed; gonopore subapical on ventral side (Figs 50, 51). Style bearing 6���7 tiny teeth apically and 5���6 fine setae subapically (Fig. 52). Anal tube process smooth (Fig. 53). Etymology. The name is an arbitrary combination of letters. Remarks. This new species is similar to H. haematoptilus in having the 1T apodemes not extended to segment II and triangular in shape (Fig. 26). However, H. osificata differs from H. haematoptilus in having the aedeagus with three distal projections, all extended distad (Fig. 50). The aedeagus H. haematoptilus has only two distal projections, one extended ventrad (Fig. 42). Distribution. Thailand., Published as part of Xu, Ye, Wang, Yi-Mei, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2022, Revision of the genus Homa Distant (Hemiptera: Cicadellidae: Typhlocybinae), pp. 179-190 in Zootaxa 5087 (1) on pages 183-184, DOI: 10.11646/zootaxa.5087.1.8, http://zenodo.org/record/5819981

Homa algulata Xu, Dietrich & Qin sp. nov. (Figs 17���22, 35���37, 69���77) Holotype. ♂, Thailand Khonkaen Nam Pong NP office, 16 o 37.201'N, 102 o 34.481'E, 218 m, Malaise trap, July 26- August 2 2006, Khamphol Jaidee (QSBG). Paratypes. 5♂♂, same data as holotype (NWAFU); 20♂♂, Thailand Nakhon Nayok Khao Yai NP Behind Football field, 14 o 24.619'N, 101 o 22.778'E, 770 m, Malaise trap, July 5-12 2006, Pong Sandao (INHS); 30♂♂, Thailand Nakhon Nayok Khao Yai NP Behind Football field, 14 o 24.619'N, 101 o 22.778'E, 770 m, Malaise trap, July 5-12 2006, Pong Sandao (QSBG); 1 ♂, Vietnam Ninh Binh, Cuc Phuong NP center 20.3522��N 105.5983��E, 370 m, July 5-8 2019, C.H.Dietrich, Malaise trap in 2�� forest, VN19-2-2 (VNMN); 1 ♂, Vietnam Ninh Binh, Cuc Phuong NP, ca. entrance 20.2501��N 105.7145��E 156m, July 7 2019, C.H.Dietrich, Hg. vapor light, VN19-1-6 (INHS). Description. Measurements. Body length: male 3.8���4.1 mm. Ground color reddish brown. Face crimson, ocelli surrounded by dark patches (Fig. 20). Anteclypeus strongly swollen and broader in male than in female (Fig. 20). 1T apodemes not extending to end of segment III, divergent posteriorly (Fig. 35). Male pygofer in lateral view with rounded apex, bearing 5���6 rigid microsetae along posterior margin (Figs 36, 37, 69���71). Subgenital plate widest at base and terminated in rectangle; basal setae (7���8) slightly shorter than macrosetae; marginal setae (8���10) arranged in irregular 1���2 rows, discrete; macrosetae (6���7) somewhat pointed, not reaching tip of plate; feeble microsetae (6���8) weakly developed (Figs 69, 72). Aedeagus shaft tubular, with long unpaired basal process surpassing shaft apex, both shaft and process with single tiny preapical spine, gonopore apical in dorsal view (Figs 73���75). Style strongly curved medially, dentifer bearing 5-6 tiny teeth and about 4���5 setae (Fig. 76). Anal tube lamellate, tapered to acute apex (Fig. 77). Etymology. The name is an arbitrary combination of letters. Remarks. This species is similar to H. osificata in having the male anteclypeus strongly swollen and broadened (Figs 8, 20), but differs from the latter in having the subgenital plate macrosetae concentrated near the plate apex (Fig. 72) (concentrated near base in H. osificata (Fig. 49)). H. algulata is also similar to H. asilata in having the 1T apodemes divergent posteriorly (Figs 32, 35). However, it differs from H. asilata in having the anal tube process broad basally (Fig. 77) (anal tube process narrowed basally in H. asilata (Fig. 68)). H. algulata differs from all other known species of Homa in having a single well-developed basal aedeagal process surpassing the shaft apex (Figs 73, 74). Distribution. Thailand and Vietnam., Published as part of Xu, Ye, Wang, Yi-Mei, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2022, Revision of the genus Homa Distant (Hemiptera: Cicadellidae: Typhlocybinae), pp. 179-190 in Zootaxa 5087 (1) on pages 187-188, DOI: 10.11646/zootaxa.5087.1.8, http://zenodo.org/record/5819981

Homa rubrodorsata Kato, 1933 (Figs 78���80) Homa rubrodorsata Kato, 1933: 452. Materials examined: 1♀ (Holotype), China (Taiwan), NO. 233. Remarks. H. rubrodorsata was described based on the female holotype from Taiwan. The male remains unknown and, therefore, the status of this species remains uncertain although it clearly belongs in this genus. Here we provide photos of the holotype., Published as part of Xu, Ye, Wang, Yi-Mei, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2022, Revision of the genus Homa Distant (Hemiptera: Cicadellidae: Typhlocybinae), pp. 179-190 in Zootaxa 5087 (1) on pages 188-189, DOI: 10.11646/zootaxa.5087.1.8, http://zenodo.org/record/5819981, {"references":["Kato, M. (1933) Notes on Japanese Homoptera, with descriptions of one new genus and some new species. Entomological World, 1933, 452 - 471."]}

Homa Distant, 1908 Homa Distant, 1908: 400. Type species: Homa insignis Distant, 1908 = Homa haematoptilus (Kirkaldy, 1906) Diagnosis. Body white or stramineous, usually with bold symmetrical red markings dorsally. Head including eyes distinctly broader than pronotum in dorsal view (Figs 1, 3, 5, 7, 9, 11, 13, 15, 17, 19, 78, 81). Crown slightly longer medially than next to eye, coronal suture short, not extended beyond mid-length of crown (Figs 1, 3, 5, 7, 9, 11, 13, 15, 17, 19, 78, 81). Eyes large, as wide as distance between basomedial angles (Figs 3, 7, 11, 15, 19, 78, 81). Ocelli distinct, separated from eyes by less than half of ocellar diameter (Figs 4, 8, 12, 16, 20, 82). Face broad and convex in profile, lateral frontal suture not extended ventromesad of ocelli, anteclypeus strongly swollen and broadened in some species (Figs 4, 8, 12, 16, 20, 82). Forewing narrow, rounded apically, veins R2, RM and MCu dissociated at their base, all arising from m cell. Hind wing with CuA and MP fused for entire length (Figs 21, 22). Front femur row AV with 1 basal seta distinctly enlarged or setae subequal or gradually decreasing in size distally; AM1 distinctly enlarged. Middle femur with 1 dorsoapical macroseta. Hind tibia row AV with 4 preapical macrosetae. Male basal abdominal sternal apodemes (1T apodemes) developed, 2S and 3T apodemes absent (Figs 23, 26, 29, 32, 35, 83). Male pygofer elongate, with few rigid microsetae along posterior margin, ventral appendage absent, dorsal bridge well developed (Figs 24, 25, 27, 28, 30, 31, 33, 34, 36, 37, 84, 85). Subgenital plate broadest at base and narrowing apically toward sclerotized and pigmented apex; basal setae densely grouped and stout, blunt-tipped; marginal microsetae rare, along dorsal margin of plate; macrosetae uniseriate, not reaching tip of plate; feeble microsetae present (Figs 24, 27, 30, 33, 36). Style base broad and distinctly bent mesad at point of articulation to plate, terminating in a strong arcuate and abruptly narrowed apical dentifer, dentifer with a few, small teeth, subapically with several setae and few sensory pits (Figs 44, 52, 60, 67, 76). Aedeagal shaft tubular, with or without process(es), dorsoatrium absent. Connective fused with the base of aedeagus (Figs 42, 43, 50, 51, 58, 59, 65, 66, 73���75). Anal tube with basal anterior appendage well developed (Figs 45, 53, 61, 68, 77). Remarks. Homa is unique among known genera of the Usharia group (Xu et al., 2021) in having 1T apodemes (Figs 23, 26, 29, 32, 35, 83). This feature is rare in Empoascini, most genera of which have a well-developed 2S apodemes or 3T apodemes. It is similar to Baguoidea Mahmood, 1967 in body shape and color, but differs in having forewing veins R2 and RM dissociated at their base (Fig. 21) (confluent for a short distance in Baguoidea Mahmood, 1967); and in lacking a pygofer appendage (ventral appendage present in Baguoidea Mahmood, 1967). Distribution. China (Yunnan and Taiwan Provinces), Thailand, Vietnam, Japan, Sri Lanka, Philippines, Malaysia, Australia., Published as part of Xu, Ye, Wang, Yi-Mei, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2022, Revision of the genus Homa Distant (Hemiptera: Cicadellidae: Typhlocybinae), pp. 179-190 in Zootaxa 5087 (1) on page 180, DOI: 10.11646/zootaxa.5087.1.8, http://zenodo.org/record/5819981, {"references":["Distant, W. L. (1908) Rhynchota. IV. Homoptera and appendix (Pt.). The fauna of British India, including Ceylon and Burma. Taylor & Francis, London, XV + 501 pp.","Kirkaldy, G. W. (1906) Leafhoppers and their natural enemies. Part IX, Leafhoppers, Hemiptera. Bulletin of the Hawaiian Sugar Planters Association Division of Entomology, 1, 271 - 479.","Xu, Y., Dietrich, C. H., Zhang, Y. - L., Dmitriev, D. A., Zhang, L., Wang, Y. - M., Lu, S. - H. & Qin, D. - Z. (2021) Phylogeny of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) based on morphological characteristics, with reclassification of the Empoasca generic group. Systematic Entomology, 46, 266 - 286. https: // doi. org / 10.1111 / syen. 12461","Mahmood, S. H. (1967) A study of the typhlocybine genera of the Oriental Region (Thailand, the Philippines and adjoining areas). Pacific Insect Monograph, 12, 1 - 52."]}

Ricanula pulverosa (Stål, 1865) (Figs 10–11) Ricania pulverosa Stål, 1865: 162; Stål, 1870: 767; Atkinson, 1886: 61; Noualhier, 1896: 256; Schmidt, 1905: 176; Distant, 1906: 380; Schumacher, 1915: 130; Melichar, 1898: 240; Esaki, 1932: 1805; Kato, 1933: 7; Zia, 1935: 537; Lallemand, 1942: 70; Jacobi, 1944: 22; Chou et al., 1985: 81. Ricanula pulverosa, Melichar, 1923: 130; Metcalf, 1955: 97; Yang, 1989: 193. Specimens examined. 1 male: 1 Jun. 1974, Yunnan, Xishuangbanna, Mengla, coll. Yao Chou et Feng Yuan; 1 male: 18 Apr. 1982, Yunnan, Xishuangbanna, Mengla, coll. Sumei Wang et Jingruo Zhou. Measurements. Length (inc. tegmen): male 7.8 mm. Remarks. First time after original description, Yang (1989) described and illustrated the male and female genitalia of the species based on Taiwan specimens. The morphological characters examined here are consistent with Yang’s (1989) description, but the male genitalia in this study show somewhat intraspecific variations as follows: 1) anal tube nearly rectangular in dorsal view (Fig. 10G) (irregularly hexagonal in Yang, 1989, Fig. 12G); 2) lateral processes of aedeagus reaching only 1/4 of the dorsal processes (Fig. 10I) (lateral processes reaching 1/2 of the dorsal processes in Yang, 1989, Figs 12H, I). Distribution. China (Provinces Shaanxi, Zhejiang, Yunnan, Fujian, Hainan, Taiwan), Cambodia, Myanmar, Thailand, Vietnam, Japan, India, Indonesia., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on page 366, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953, {"references":["Stal, C. (1865) Homoptera nova vel minus congita. Ofversigt af Kongliga Svenska Vetenskaps-Akadamiens Forhandlingar, 22, 145 - 165.","Stal, C. (1870) Hemiptera insularum Philippinarum. Bidrag till Philippinska oarnes Hemipter-fauna. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar, 27, 607 - 776. https: // doi. org / 10.5962 / bhl. title. 61898","Atkinson, E. T. (1886) Notes on Indian Rhynchota, No. 6. Journal and Proceedings of the Asiatic Society of Bengal, 55, 143 - 223.","Noualhier, J. M. (1896) Note sur les Hemipteres recoltes en Indo-Chine et offerts au Museum par M. Pavie. Bulletin du Museum national d'histoire naturelle, 10, 251 - 259.","Schmidt, E. (1905) Die Ricaniiden des Stettiner Museums. Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin, 66, 168 - 198.","Distant, W. L. (1906) The fauna of British India, including Ceylon and Burma. Rhynchota. Vol. III (Heteroptera-Homoptera). Taylor and Francis, London, 503 pp.","Schumacher, F. (1915) Homoptera in H. Sauter's Formosa-Ausbeute. Supplementa Entomologica, 4, 108 - 142.","Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359.","Kato, M. (1933) Three colour illustrated insects of Japan. Fascicle 4. Homoptera: Fulgoridae and Others. Koseikaku, Tokyo, 127 pp.","Zia, Y. (1935) Note sur les Flatinae et les Ricaniinae de la chine du sud et du Tonkin (Homoptera Fulgoridae). Sinensia, 6 (5), 525 - 540.","Lallemand, V. (1942) Notes sur quelques especes recueillies par le R. Piel (Musee Heude, Shanghai) et le R. P. de Cooman (Hoa Binh, Tonkin). Notes d'Entomologie Chinoise. Musee Heude, 9 (4), 69 - 77.","Jacobi, A. (1944) Die Zikadenfauna der Provinz Fukien in Sudchina und ihre tiergeographischen Beziehungen. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 5 - 66.","Chou, I., Lu, J. S., Huang, J. & Wang, S. Z. (1985) Homoptera, Fulgoroidea. Economic Insect Fauna of China. Fasc. 36. Science Press, Beijing, 152 pp.","Melichar, L. (1923) Homoptera, fam. Acanaloniidae, Flatidae et Ricaniidae. Genera Insectorum. Part 182. L. Desmet-Verteneuil, Bruxelles, 185 pp.","Metcalf, Z. P. (1955) General catalogue of the Hemiptera. Fascicle IV. Fulgoroidea. Part 16. Ricaniidae. Smith College, Northampton, Massachusetts, 199 pp.","Yang, C. T. (1989) Ricaniidae of Taiwan (Homoptera: Fulgoroidea). In: Collected Papers on Fulgoroidea of Taiwan. Taiwan Museum Special Publication Series, 8, pp. 171 - 204."]}

Ricanula Melichar, 1898 Ricania (Ricanula) Melichar, 1898: 218. Ricanula Schmidt, 1912: 75. Type species. Ricania noualhieri Melichar, 1898, designated by Schmidt (1912). Remarks. Ricanula can be distinguished from other genera in Ricaniidae by the combination of following characters: costal area of tegmen with sparse and curved transverse veinlets; postero-apical part of tegmen with eyes-spot black cells; tegmen with one line of transverse veinlets; median and posterior parts of tegmen with numerous irregular transverse veinlets. Diagnostic characters. Head. Head including eyes narrower than mesonotum/thorax. Vertex transverse, distinctly wider than long at midline, with all margins well carinate; disc of vertex with or without median carina. Frons with all margins well carinated, at upper margin longer than high at midline, widest at the level of lower margin of compound eyes; lateral margins covering base of pedicel, not incised near the level of ocelli. Frontal disc tricarinate, carinae distinctly separated basally, median carina straight; lateral carinae arcuate, almost parallel to lateral margins. Fronto-clypeal suture arched/arcuate. Clypeus distinctly narrower than frons, without median carina. Compound eyes, with small callus in lower part of posterior margin. Ocelli present. Rostrum-apical segment shorter than subapical one. Antenna pedicel elongate, cylindrical, with functional area at the top. Thorax. Pronotum distinctly longer than vertex at midline; disc of pronotum with median carina and two lateral impressions. Mesonotum elongate, diamond shape, longer in midlength than wide at lateral angles and longer in midlength longer than combined length of vertex and pronotum at midlength; lateral angles placed before midlength; median carina, lateral and antero-lateral carinae present; median carina and lateral carinae connected basally; median carina reaching scutellum, lateral carinae reaching posterior margin, anterolateral carinae not connected with lateral and not surpassing level of lateral angles of mesonotum. Tegmina membranous, elongately-triangular, flattened, with distinct venation and transverse veinlets. Costal margin weakly arcuate, apical angle broadly rounded, placed distad to claval angle, posterior margin weakly sinuate, postclaval margin (tornus) absent. Costal area with transverse veinlets and oblique shallowly incisions; postero-apical part with eyes-spot black cells. Longitudinal veins ScP+RA, MP and CuA leaving basal cell separated. CuA vein first fork placed before the connection of claval veins Pcu+A 1. Tegmen with single apical line of transverse veinlets, median and posterior part of tegmen with numerous irregular transverse veinlets. Hind wing with precostal cell present; ScRA and MP forking distinctly after midlength of wing, CuA forking distinctly before half of wing. Hing legs. Metatibia with 2 lateral spines; apically with 6 well developed spines, external lateral spines bigger than internal lateral spines; basitarsomere of metatarsus a little longer than cumulative length of second and apical tarsomeres. Male terminalia. Anal tube with ventral margin strongly concaved in lateral view. Pygofer higher than wide; dorsal part narrower than ventral one. Genital styles broadly triangular (dorsal and ventral margins nearly parallel in R. fujianensis and R. cacaonis), with sharp spine-like process at the end of dorsal margin. Periandrium with or without processes, with long lateral split surpassing the half of its length; dorsal periandrium a bit shorter than ventral one. Aedeagus long and narrow, apically with 1–2 pairs of symmetrical, well sclerotized, spinose processes. Female terminalia. Pregenital sternite with well-developed lateral lobes, median portion of the posterior margin with or without processes. Anal tube not surpassing half of upper margin of the gonoplac. Gonoplac well developed, laterally flattened; posterior margin of the gonoplac with 2–3 rows of small teeth; membranous parts of gonoplac well developed, placed medially on ventral margin. Gonapophysis VIII sabre-like, v-shape in cross section, with teeth at dorsal margin; endogonocoxal process with spiniferous microsculptures, well sclerotized medially; lateral parts membranous, reaching apex of gonapophysis VIII. Gonaphophyses IX and gonospiculum bridge well developed. Bursa copulatrix with two isometric pouches: first pouch with well visible cell and sclerotized ornamentation; second pouch with numerous pores., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 354-355, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953, {"references":["Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359.","Schmidt, E. (1912) Diagnosen neuer Fulgoriden-Gattungen und Arten nebst einigen Bemerkungen. Stettiner Entomologische Zeitung, 73, 67 - 102."]}

Ricanula peronata sp. nov. (Figs 7–9, 11) Etymology. The name is derived from the Latin word ‘ peronatus ’, referring to the periandrium with boot-shaped ventral processes in ventral view. Diagnosis. The species is similar to Ricanula curva sp. nov., but differs from the latter by having periandrium with boot-shaped ventral processes in ventral view (Fig. 9F); ventral processes of aedeagus oriented ventrally in lateral view (Fig. 9G) (apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view; ventral processes of aedeagus S-curved in lateral view in Ricanula curva sp. nov.). Description. Measurements. Length (inc. tegmen): male 6.5–7.4 mm, female 7.3–8.8 mm. Head. Vertex (Figs 7A, C) without median carina. Frons: median and lateral carinae of frontal disc surpassing half of disc, ending about the level of antennae; apical parts of median carina weakly visible. Thorax. Pronotum (Figs 7A, C) with small round depressions submedially on each lateral side. Mesonotum: lateral carinae (Figs 7A, C) not reaching posterior margin; antero-lateral carinae not connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin arcuate. Longitudinal veins ScP+RA and RP, MP arising as short common stem from basal cell. Claval veins Pcu and A 1 fused on midlength of CuP vein. Hind wing (Fig. 7F) without transverse veinlets. Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia. Anal tube (in dorsal view, Fig. 8B) nearly square, posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct surpassing the posterior margin. Pygofer (in lateral view, Fig. 8A) with dorsal posterior angle without process. Genital styles (in lateral view, Fig. 8A) broadly triangular; ventral margin weakly sinuate; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 9A–C): Periandrium (Figs 9D–F) with boot-shaped ventral processes in ventral view, apical part of ventral processes straight; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with small rod-shaped processes in lateral view, rodshaped processes hidden in the periandrium (in ventral view); ventral periandrium distinctly convex. Aedeagus (Figs 9G–I) apically with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about two thirds of aedeagus; ventral processes oriented ventrally in lateral view. Female terminalia (Figs 8C–I). Pregenital sternite (Fig. 8I): posterior margin medially with two prominent processes, margin between processes with wide and shallow incision. Anal tube (in dorsal view, Fig. 8C) ovoid, with widest part medially, basal margin weakly convex, posterior margin widely concave, lateral margins arcuate; anus placed after midlength, paraproct surpassing the posterior margin. Gonoplac (Fig. 8H): posterior margin with two rows of small teeth. Coloration. General color brown (Figs 7A–B). Median part of frons (Fig. 7D) black brown. Eyes brown (Figs 7B–C), ornamented with irregular brown patches. Gena (Fig. 7B) black brown with two yellowish spots. Tegmen (Figs 7B, E) brown, costal margin with about 14 transverse black brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, sub-medially of tegmen with a large flavescent spot marked by 2 central transverse back lines. Wings brown, each side of A 2 with a longitudinal grayish narrowed band (Fig. 7F). Abdomen and terminalia brown. Type material. Holotype, male, China: 20 Apr. 1964, Yunnan, Xishuangbanna, Menglun, 650 m, coll. Baolin Zhang. Paratypes (8 males, 17 females, China): 1 male: 13 Jul. 1958, 650– 700 m, coll. Chunpei Hong; 2 males, 4 females: 21/ 30 Apr. 1974, coll. Yao Chou, Feng Yuan et Yinyue Hu; 1 female: 22 May 1982, 1 female: 24 May 1982, coll. Qinmei Wang et Jingruo Zhou; 1 male, 1 female: 18 May 1991, 1 female: 20 May 1991, 1 male, 1 female: 22 May 1991, 2 females: 26 May 1991, 3 males, 6 females: 31 May 1991, coll. Yinglun Wang et Wanzhi Cai; Yunnan, Xishuangbanna, Menglun. Distribution. China (Province Yunnan).

Ricanula curva sp. nov. (Figs 4–6, 11) Etymology. The name is derived from the Latin word ‘ curvus ’, referring to ventral processes of aedeagus being Scurved in lateral view; apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view. Diagnosis. The species is similar to R. peronata sp. nov., but differs from the latter by having apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view (R. peronata —ventral processes boot-shaped in ventral view); ventral processes of aedeagus S-curved in lateral view (R. peronata —ventral processes of aedeagus oriented ventrally in lateral view). Description. Measurements. Length (inclu. teg.): male 6.7–8.4 mm, female 7.2–9.6 mm. Head. Vertex (Figs 4A, C) with or without median carina. Frons: lateral carinae of frontal disc finishing basally at the same level as median carina, ending about the level of antennae. Thorax. Pronotum (Figs 4A, C) with small round depressions submedially on each lateral side (in some specimens weakly visible). Mesonotum: lateral carinae of mesonotum (Figs 4A, C) reaching posterior margin; anterolateral carinae connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin almost straight. Longitudinal veins ScRA and RP arising as short common stem from basal cell, MP veins forked on basal cell. Claval veins Pcu and A 1 fused before midlength of CuP vein. Hind wing with r-m transverse veinlets present in distal part of wing (Fig. 4F). Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia (Figs 5A–B, 6A–I). Anal tube (in dorsal view, Fig. 5B) nearly as rectangle; posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct slightly surpassing the posterior margin. Pygofer (in lateral view, Fig. 5A): dorso-posterior angle with process. Genital styles (Fig. 5A) broadly triangular in lateral view; ventral margin convex; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 6A–C): Periandrium (Figs 6D–F) with ventral processes, apical part of ventral processes of periandrium curved laterally (in dorsal and ventral view), ventral periandrium distinctly convex in middle; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with rod-shaped processes in lateral view. Aedeagus (Figs 6G–I) with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about 2/3 of aedeagus, curved dorsally at 2/3 of its length; ventral processes S-curved ventrally in lateral view. Female terminalia (Figs 5C–I). Pregenital sternite: posterior margin medially with two processes, margin between processes with strong and deep incision (Fig. 5I). Anal tube (in dorsal view, Fig. 5C) elongate, with posterior part wider than basal one; basal margin almost straight, posterior margin widely concave medially, lateral margins arcuate; anus placed a bit before midlength, paraproct surpassing the posterior margin. Gonoplac posterior margin with two rows of teeth. Coloration. General color brown to dark brown (Figs 4A–B). Lateral margins of frons yellow, area alongside frontoclypeal suture yellow, clypeus brown with yellow patch medially under frontoclypeal suture, rostrum yellowish with brown apex (Fig. 4D). Eyes (Fig. 4B) sordid brown, ornamented with irregular black brown patches. Gena (Fig. 4B) brown with two yellow spots. Tegmen (Figs 4A–B, E) brown to dark brown; costal margin with about 13–16 transverse brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, tegmen sub-medially with a large flavescent spot marked by 2 central transverse brown lines. Wings (Fig. 4F) brown, each side of A 2 with a grayish narrowed band longitudinally. Abdomen and terminalia brown. Type material. Holotype, male, China: 28 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang. Paratypes (13 males, 21 females, China): 4 males, 2 females: 11 Apr. 1978, Guangxi, Baise, Yangwei, coll. Xianyu Qian; 1 male: 12 May 1980, Guangxi, Longzhou, Longhu, coll. Zhuyin Wang; 1 male: 17 May 1982, Guangxi, Longzhou, Nonggang, 240m, coll. Fasheng Li; 2 males: 29 May 1982, 1 female: 30 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang; 1 female: 25 Jun. 1982, Guangxi, Longsheng, coll. Jikun Yang; 3 males, 14 females: 3 Jun. 1984, 1 female: 9 Jun. 1984, Guangxi, Lingtian commune, coll. Zhengliang Wu et Xiaolin Lu; 1 male, 1 female: 6 May 1993, 1 male, 1 female: 8 May 1993, Guangxi, Longzhou, Daqing Mountain, coll. Sikong Liu. Distribution. China (Province Guangxi)., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 359-363, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953

Mjolnirus Wang, Xu & Qin, gen. nov. Type species: Mjolnirus mediolobus Wang, Xu & Qin, sp. nov. Description. Head with eyes subequal in width to maximum width of pronotum. Eyes in dorsal view narrower than distance between basomedial angles (Figs 18, 20). Coronal suture extended slightly beyond crown mid-length but not onto face (Fig. 20). Ocelli on crown margin well separated from eyes. Lateral frontal sutures present and extended below ocelli (Fig. 21). Anteclypeus narrowed and slightly convex basally (Figs 19, 21). Forewing narrow, rounded apically, veins R2 and RM confluent at their bases, distal segments of both arising from m cell (Fig. 22), vein MCu well separated with RM apically. Hind wing with vein MP and CuA completely confluent (Fig. 23). Fore femur with 1 enlarged AV seta on basal margin. Middle femur with 1 distinct dorsoapical macroseta. Hind femur setal formula 2+1+1, tibia with 6 macrosetae in row AV near apex. Male 2S abdominal apodemes well developed, parallel-sided (Fig. 24). Pygofer lobe with few microsetae along posterodorsal margin; ventral margin infolded; dorsal bridge well delimited, long, ventral appendage present, arising from internal ridge near ventral margin of pygofer lobe (Figs 25���29). Subgenital plate surpassing pygofer lobe, well expanded basally with prominent basolateral projection, gradually narrowing apically; basal setae elongate and stout, arising from basolateral angle; marginal setae well defined, separated into medial and distal groups; macrosetae uniseriate, not reaching tip of plate; feeble microsetae undeveloped (Figs 25, 27, 30). Style slightly broadened basally, apophysis elongate, with short basolateral protrusion, dentifer short with few tiny teeth (Figs 27, 31, 32). Aedeagus with preatrium longer than shaft, shaft with asymmetrical basal processes; dorsal apodeme absent (Figs 25, 26, 33���35). Connective, articulated with aedeagus, with narrow median lobe on anterior margin, caudal margin notched medially (Fig. 36). Anal tube processes short, rounded (Figs 25, 37). Distribution. Thailand. Etymology. The new generic name is derived from Proto-Germanic: Mj��lnir, the hammer of Thor, the Norse god of thunder, referring to the pygofer processes which resemble hammer handles (Figs 25���29). Gender: masculine. Notes. Mjolnirus is included in the Empoasca generic group based on hind wing veins MP and CuA being confluent, the subgenital plates free through their length, and the connective well separated from the aedeagus. It is similar to Empoasca Walsh, but differs in the unique structure of the pygofer and subgenital plate. Among other empoascine genera in the Oriental Region, Mjolnirus superficially resembles Sharoka Sohi & Mann, 1993 in having very similar head structure (head with eyes subequal to maximum width of pronotum, coronal suture not extended onto face); and the subgenital plate broadest at the base and tapering towards the apex, with marginal setae divided into two separate groups. It differs from Sharoka in the shape of the aedeagus which lacks a dorsal apodeme and has the atrium weakly expanded in caudoventral view (aedeagus with dorsal apodeme, and atrium greatly expanded in caudoventral view in Sharoka). Furthermore, Mjolnirus differs from all other known genera of Empoasca genus group in having column-like processes arising from the ventral margin of the male pygofer., Published as part of Wang, Yi-Mei, Xu, Ye, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2021, Two new empoascine leafhopper genera, Thaioasca and Mjolnirus, from Thailand (Hemiptera: Cicadellidae: Typhlocybinae), with a checklist of Empoascini in Thailand, pp. 246-254 in Zootaxa 4915 (2) on pages 249-250, DOI: 10.11646/zootaxa.4915.2.4, http://zenodo.org/record/4454393, {"references":["Sohi, A. S. & Mann, J. S. (1993) A new genus and a new species of Empoascini (Homoptera: Cicadellidae: Typhlocybinae) form India. Journal of Insect Science, 6 (1), 99."]}

Thaioasca Wang, Xu & Qin, gen. nov. Type species: Thaioasca contaminata Wang, Xu & Qin sp. nov. Description. Body robust. Head including eyes distinctly wider than maximum width of pronotum, eyes in dorsal view narrower than distance between basomedial angles and wider than half of distance between basomedial angles (Figs 1, 3). Coronal suture extended nearly to anterior margin of crown (Fig. 3). Ocelli present (Fig. 4). Lateral frontal sutures extended below ocelli, male anteclypeus tapered and weakly convex, not expanded (Figs 2, 4). Forewing narrow, rounded apically, veins R2 and RM confluent for short distance, distal segments of both arising from m cell, vein RM and MCu almost parallel apically (Fig. 5). Hind wing with veins MP and CuA confluent (Fig. 6). Fore femur with 2 or 3 enlarged AV setae on basal margin. Middle femur with 2 distinct dorsoapical macrosetae. Hind femur setal formula 2+1+1, tibia row AV with 4 macrosetae near apex. Male basal abdominal sternal apodemes (2S apodemes) developed (Fig 7). Pygofer elongated, extended into sharp point, strongly depressed dorsally in profile, dorsal bridge undeveloped, membranous, ventral appendage robust (Figs 8–12). Subgenital plates fused at least at base, basal setal group submarginal near midlength of plate; marginal microsetae sparse, limited to distal part of plate margin; macrosetae uniseriate basally, somewhat irregular apically; feeble microsetae present but inconspicuous (Figs 8, 10, 13). Style broadened at base, apex tapered and smooth, with preapical setae (Figs 8, 10, 14). Aedeagus shaft tubular, preatrium absent, dorsoatrium well developed, gonopore medial on ventral surface (Figs 8, 9, 15, 16). Connective not delimited, completely fused with base of aedeagus (Figs 8, 10). Anal tube process well-developed but short (Figs 8, 10, 17). Distribution. Thailand. Etymology. The generic name combines a prefix referring to the type locality (Thailand) with the name of the type genus of the tribe, Empoasca. Gender: feminine. Notes. Thaioasca is included in the Ficiana generic group sensu Xu et al. (2015). The new genus resembles other members of this group in the proportions of the head (crown shorter than width between eyes), the fore- and hind wing venation, and in having the subgenital plates fused basally and the connective fused to the aedeagus. Thaioasca differs from Ficiana Ghauri, Dialecticopteryx Kirkaldy, Ishiharella Dworakowska, and Mahmoodia Dworakowska in having the coronal suture extended to near the anterior margin of the crown (it extends onto the face in Ficiana and Dialecticopteryx but absent in Ishiharella and Mahmoodia). From Kotwaria Dworakowska, Daluana Ramakrishnan, and Nimabanana Dworakowska the new genus differs in having a well-developed anal tube appendage (absent in these other three genera). Thaioasca is, so far, the only member of the Ficiana group with a ventral appendage on the pygofer; thus, the definition of the group provided by Xu et al. (2015) should be modified accordingly. Fusion of the connective with the base of the aedeagus in this group has been confirmed by study of specimens of three genera in addition to the new genus, Ishiharella, Ficiana, and Dialecticopteryx; and is indicated by the original illustrations of male genitalic characters of the type species of the other four included genera (Mahmoodia, Nimabanana, Kotwaria, and Daluana; see Dworakowska, 1970a: figs 2, 6; Dworakowska, 1984: figs 65, 75, 76; Dworakowska, 1994b: figs 62, 63; Ramakrishnan, 1982: fig. 8). Further investigations are needed to determine the phylogenetic status and relationships of this group.

Mjolnirus mediolobus Wang, Xu & Qin, sp. nov. (Figs 18–37) Description. Body length: male 3.5–3.8 mm. Ground color faint pale yellowish (Figs 18–21). Eyes silvery gray to black (Figs 18–21). Crown with irregular red spots along coronal suture. Face almost whitish due to long-term storage in ethanol. Mesonotum much darker than head and pronotum (Figs 18, 20). Fore- and hind wing subhyaline, veins distinct (Figs 22, 23). Legs apricot cream-colored (Figs 19, 21). Male 2S abdominal apodemes reaching posterior margin of segment IV, parallel sided (Fig. 24). Male pygofer broad and rounded apically, bearing 8–9 rigid setae along postero-dorsal margin of lobe, dorsal bridge long and well delimited (Figs 25–29); ventral pygofer process unique, extending dorsomesad from ventral margin of pygofer, with apparently eversible nipple-like structure apically. Subgenital plate longer than pygofer lobe in lateral view, apex rounded and bent dorsad along ventral margin of pygofer lobe; basal setae (6–7) uniseriate, almost same size as macrosetae; marginal setae (20–22) divided into two groups occupying more than half length of dorsolateral margin, one group smooth apically arranged in one row, another group slender, tapered with a blunt tip apically; macrosetae (7–8) not reaching apex of plate; feeble microsetae (12–14) very sparse and inconspicuous (Figs 25, 27, 30). Aedeagus with preatrium elongated, occupying nearly half length of aedeagus in profile; shaft with asymmetrical basal processes, one ridged straight process protruding out of side of shaft and another process at ventral margin basally, slightly curved dorsad (some variation in shape and length of processes observed) (Figs 25, 26, 33–35). Style dentifer with five to seven teeth preceded by four to five setae and few sensory pits (Figs 25, 27, 31, 32). Connective lamellate, longer than wide (Fig. 36). Anal tube process well developed, with broad and rounded apex (Figs 25, 27, 37). Holotype. Ƌ (QSBG), Thailand Chiang Mai Doi Chiang Dao WS Nature trail 19°24.278' N 98°55.311'E, 491 m, Malaise trap, March 10–17, 2008, Songkran & Apichart, leg., T3155; Paratypes. 10ƋƋ (INHS), 10ƋƋ (QSBG), 15ƋƋ (NWAFU), same data as holotype, leg. T3155. Distribution. Thailand. Etymology. The name, which is a noun in apposition, is a combination of Latin words: “ medius ”, middle and “ lobus ”, lobe, referring to the median lobe at the anterior margin of the connective.

Thaioasca contaminata Wang, Xu & Qin, sp. nov. (Figs 1���17) Description. Body length: Male 4.2���4.4 mm. General color of body yellowish (Figs 1���4). Head with median black spot at apical transition from crown to face, crown short and broad, rounded anteriorly (Figs 1, 3, 4). Pronotum with yellowish-black, sinuate transverse depression laterally, mid-posteriorly black (Figs 1, 3). Fore- and hind wing subhyaline, veins distinct (Figs 5, 6). Legs whitish (Fig. 2). 2S apodemes extending to middle of segment IV (Fig. 7). Male pygofer with 10���11 tiny setae in depression of dorsal margin; apex of pygofer lobe pointed, curved upward and inward (Figs 8���12); ventral appendage developed and inflated at base, narrowed and heavily sclerotized distally, surpassing caudal margin of pygofer lobe (Figs 8���12). Subgenital plates large and fused for two-fifths of their length and surpassing pygofer lobe; basal setae (8���9) arranged in 1 or 2 rows, almost same size as macrosetae; marginal setae (8���9) setae not reaching tip of plate; macrosetae (7���8) arising at middle of plate below basal setae; feeble microsetae very sparse and shorter than macrosetae (Figs 8, 10, 13). Aedeagus slender, shaft slightly bent to right side, in lateral view wide basally, then abruptly narrowed, apex bifid in posterior view giving rise to pair of short distally directed processes (Figs 8, 9, 15, 16); gonopore close to middle of shaft, ventral. Style greatly broadened basally, distal part elongate, sinuate in ventral and lateral views, gradually tapering to bluntly rounded tip, with 4���5 fine preapical setae (Figs 8, 10, 14). Anal tube appendage extended nearly half distance to ventral margin of genital capsule, lobe-like with obtuse posteroventral edge (Figs 8���10, 17). Holotype. �� (QSBG), Thailand Chiang Mai Doi Chiang Dao WS Nature trail, 19��24.278'N 98��55.311'E, 491 m, Malaise trap, March 10���17, 2008, Songkran & Apichart leg., T3155. Paratypes. 3���� (INHS), Thailand Nakhon Sawan Mae Wong NP Tha Ta Thai, 15��54.258'N 99��18.862'E, Malaise trap, September 1���7, 2007, Kaewkok N. & Saksit P. leg., T3597; 3���� (NWAFU), Thailand Nakhon Si Thammarat Namtok Yong NP Hill, evergreen forest, 8��13.891'N 99��47.98'E, 679 m, Pan trap, November 16���17, 2008, Paiboon leg., T4263. Distribution. Thailand. Etymology. The specific epithet is derived from the Latin adjective ��� contaminatus ���, contaminated, polluted, referring to the dark smudge on the crown., Published as part of Wang, Yi-Mei, Xu, Ye, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2021, Two new empoascine leafhopper genera, Thaioasca and Mjolnirus, from Thailand (Hemiptera: Cicadellidae: Typhlocybinae), with a checklist of Empoascini in Thailand, pp. 246-254 in Zootaxa 4915 (2) on pages 247-248, DOI: 10.11646/zootaxa.4915.2.4, http://zenodo.org/record/4454393

Shumka Dworakowska, 1997 ��� Thailand, Oriental realm., Published as part of Wang, Yi-Mei, Xu, Ye, Dmitriev, Dmitry A., Dietrich, Christopher H. & Qin, Dao-Zheng, 2021, Two new empoascine leafhopper genera, Thaioasca and Mjolnirus, from Thailand (Hemiptera: Cicadellidae: Typhlocybinae), with a checklist of Empoascini in Thailand, pp. 246-254 in Zootaxa 4915 (2) on page 253, DOI: 10.11646/zootaxa.4915.2.4, http://zenodo.org/record/4454393, {"references":["Dworakowska, I. (1997) A review of the genus Alebroides Matsumura, with description of Shumka gen. nov. (Homoptera: Auchenorrhyncha: Cicadellidae). Oriental Insects, 31, 241 - 407. https: // doi. org / 10.1080 / 00305316.1997.10433759"]}

Limois guangxiensis Chou & Wang, 1985 Figs 2, 14 E–H Limois guangxiensis Chou & Wang, in Chou et al. 1985a: 31. Limois guangxiensis – Chou et al. 1985b: 111. Diagnosis (1) Frons with two longitudinal carinae (Fig. 2E), cephalic process of vertex relatively long, surpassing posterior margin of vertex but not reaching the middle of pronotum (Fig. 2 C–D, F); (2) pronotum and mesonotum rusty brown, pronotum covered with numerous dark spots centrally and posteriorly, mesonotum covered with numerous dark spots on disc, with 4 irregular dark patches near each posterolateral angle (Fig. 2D); (3) tegmina with basal ⅗ and costal area pale reddish brown, apical ⅖ hyaline with a broad brown stripe obliquely extending to the apical angle (Fig. 2A, C); hindwings red on basal ¾ and hyaline on remaining area, between them with a sinuate brownish fascia (Fig. 2A); (4) abdomen sordid orange, anterior margin dark brown on each tergite (Fig. 2A). Material examined Holotype CHINA • ♀ (see Figs 2 A–F, 14E–H); Guangxi, Longsheng; 30 Aug. 1964; Liang-chen Wang leg.; NWAFU HO088505. Additional material CHINA • 1 Ƌ; Fujian, Taiwu; 15 Aug. 1982; Xiang-dong Ye leg.; NWAFU HO088509. Description MEASUREMENTS. Body length: ♀ (holotype of L. guangxiensis) 15.9 mm; Ƌ 11.5 mm. Wingspan. ♀ (holotype of L. guangxiensis) 50.2 mm; Ƌ 47.4 mm. MALE GENITALIA. Pygofer in profile with ventral margin slightly concave, laterocaudal margin slightly convex (Figs 2G, 14E). Anal tube long, apical margin rounded, intersection with dorsal margin at obtuse angle in lateral view, epiproct obtusely angled apically in dorsal view (Figs 2G, I, 14E). Gonostyli oval, widest in middle in lateral view, (Figs 2G, 14E). Endosomal processes sclerotized over entire length, about 6.0 times as long as sheath, apical ⅗ exposed, terminally inflated and hook-shaped (Figs 2 J–K, 14G–H). Connective rod-like, sclerotized. Tectiductus large in lateral view, rectangular (Figs 2G, 14E). Distribution China (Guangxi, Fujian).

Limois kikuchii Kato, 1932 Figs 4–5, 14 M–Q Limois kikuchii Kato, 1932: 225. Limois kikuchii – Chou et al. 1985a: 30. — Chou et al. 1985b: 108. Diagnosis (1) Frons medially with 3 longitudinal carinae, the median carina tinier, all not reaching fronto-clypeal suture (Fig. 4C), cephalic process of vertex moderate in length, not reaching posterior margin of vertex (Fig. 4B, E); (2) pronotum brown with one longitudinal broken band on each side of median carina, blackish brown; mesonotum covered with numerous dark spots on disc, with 3 pairs of dark patches along anterior margin and one dark triangular patch on each lateral area (Fig. 4B); (3) tegmina with a transverse irregular dark brown band on basal ⅖ (extending to posterior margin) and reddish yellow on costal area and basal ½ followed by some brown patches and spots in apical half on hyaline area (Fig. 4A, D, F); hindwings red orange on basal half and hyaline on apical half (Fig. 4A, D); (4) abdominal tergite brown except anterior margin dark on each segment, sternites dark (Fig. 4A). Material examined CHINA • 1 Ƌ (see Figs 4–5, 14 M–Q); Manchuria; 1 Sep. 1932; K. Kikuchi leg.; RBINS • 1 ♀; Mukden; 3 Sept. 1928; FSAG • 1 Ƌ, 2 ♀♀; Shaanxi, Tongchuan; Sep. 1980; Yun-zhou Jiang & Da-xin Chen leg.; NWAFU. Description MEASUREMENTS. Body length: ♀ 9.5–10.5 mm; Ƌ 9.4–10 mm. Wingspan: ♀ 37.6–40.5 mm; Ƌ 32.5– 34.4 mm. MALE GENITALIA. Laterocaudal margin of pygofer slightly convex (Figs 5A, 14M). Anal tube moderate, apical margin truncate, intersected with dorsal margin at almost right angle in lateral view, epiproct acutely angled apically in dorsal view (Figs 5A, C, 14M). Gonostyli elongate, subtriangular in lateral view, apically rounded (Figs 5A, B, 14M, N). Endosomal processes about 5.0 times as long as sheath, apical 1/3 exposed, apically sinuate and inflated (Figs 5 D–F, 14O–Q). Connective and tectiductus unrecognizable. Distribution China [northern China (Kato 1932, 1933), Shaanxi, Beijing (Liang 2005)], Korea (Kato 1933; Metcalf 1947)., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 43-46, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Kato M. 1932. Notes on some Homoptera from South Manchurai, collected by Mr. Yukimichi Kikuchi. Kontyu. Tokyo Entomological Society 5: 216 - 229.","Chou I., Wang S. Z. & Huang J. 1985 a. Description of new species of Fulgoroidea from China (Homoptera: Fulgoroidea). Entomotaxonomia 7: 30 - 38.","Chou I., Lu J. S., Huang J. & Wang S. Z. (eds) 1985 b. Economic Insect Fauna of China. Vol. 36: Homoptera: Fulgoroidea. Science Press, Beijing.","Kato M. 1933. Three Colour Illustrated Insects of Japan. Fasc. IV: Homoptera. Koseikaku, Tokyo.","Liang A. P. 2005. Occurrence of the latero-subapical labial sensillum in Borysthenes maculata and Andes marmorata (Hemiptera: Fulgoromorpha: Cixiidae). Journal of Entomological Science 40 (4): 428 - 437. https: // doi. org / 10.18474 / 0749 - 8004 - 40.4.428","Metcalf Z. P. 1947. General Catalogue of the Hemiptera. Fasc. IV: Fulgoroidea. Part. 9: Fulgoridae. North Carolina State College, Raleigh (NC)."]}

Limois emelianovi Oshanin, 1908 Figs 11–12, 15 J–N Limois emelianovi Oshanin, 1908: 468. Material examined RUSSIA • 1 Ƌ (Figs 11–12, 15 J–N); Vladivostok; Aug. 1967; Anufriev leg.; RBINS • 1 Ƌ; Primorsky Krai; 8 Sept. 1961; J. Vilbaste leg.; FSAG. Description MEASUREMENTS. Body length: Ƌ 9 mm. Wingspan: Ƌ 33 mm. MALE GENITALIA. Pygofer with ventral margin sinuate in lateral view; posterior margin in lateral view slightly convex, emarginate on ventral ¼; in dorsal view deeply, roundly emarginate posteriorly (Figs 12A, 15J). Anal tube massive, in lateral view reaching level of apex of gonostyli, ventral margin sinuate, apical margin obliquely truncate, intersecting with dorsal margin at obtuse angle, in dorsal view, widening from base to apex, 1.36 times broader at apex than long in median line, apical margin roundly emarginate in dorsal view; epiproct right-angled apically, paraproct slender in dorsal view (Figs 12 A– B, 15J). Gonostyli subtriangular in lateral view, rounded apically, apical margin straight in middle, in ventral view gonostyli connected in basal ⅕, lateral tooth slender, directed posteroventrally, then curved ventrally towards apex (Figs 12A, F, 15 J–K). Endosomal processes sclerotized over entire length, about 4.0 times as long as sheath, apical ½ exposed and terminally membranous and inflated (Figs 12 C–E, 15L–N). Connective rod-like. Tectiductus large, in lateral view subrectangular, in dorsal view broad, apically rounded and distinctly concave ventrally (Figs 12 C–E, 15L–N). Distribution China (Gansu, Dongbei), Russia (Oshanin 1908; Anufriev 2009), Korea (Doi 1932a, 1932b; Kwon & Huh 2001)., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 54-55, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Oshanin V. T. 1908. Beitrage zur Kenntnis der palearktischen Hemipteren. I. Neue Gattungen und Arten von Homopteren aus Russisch-Asien. Annuaire du Musee zoologique de l'Academie imperiale des Sciences de St. - Petersbourg 12: 463 - 477.","Anufriev G. A. 2009. Cicadina. In: 2009 - Insects of Lazovsky Nature Reserve: 52 - 69. Russian Academy of Sciences, Far Eastern Branch, Institute of Biology and Soil Science, Vladivostok Dalnauka (Russia).","Doi H. 1932 a. Miscellaneous notes on insects I. Journal of Chosen Natural History Society 13: 30 - 49.","Doi H. 1932 b. Miscellaneous notes on insects II. Journal of Chosen Natural History Society 14: 64 - 78.","Kwon Y. J. & Huh E. Y. 2001. Suborder Acuchenorrhyncha. Economic Insects of Korea 19. Insecta Koreana Supplement 26: 320 - 329."]}

Limois westwoodii (Hope, 1843) Figs 8–10, 15 E–I Lystra westwoodii Hope, 1843: 133. Limois westwoodii – Stål 1863: 231. — Distant 1906: 200. — Metcalf 1947: 170. — Lallemand 1963: 55. — Nagai & Porion 1996: 22. Diagnosis (1) Frons with 2 longitudinal carinae (Figs 8D, 9D, F), cephalic process of vertex of moderate length, not surpassing anterior margin of pronotum (Figs 8B, F, 9B, H); (2) pronotum and mesonotum olivaceous, pronotum with one longitudinal broken black band on each side of median carina; mesonotum with 3 pairs of dark patches along anterior margin and one irregular patch on each lateral side (Figs 8B, 9B); (3) tegmina with a transverse irregular reddish brown band on basal half and ochraceous on costal area and basal ½ followed by some brown patches and spots in apical half on hyaline area (Figs 8A, C, E, 9A, E, G); hindwings sanguineous on basal ⅗ and hyaline on the apical ⅖, between them a sinuate brown fascia (Figs 8A, C, 9A, E); and (4) abdomen dark, posterior margin whitish on each segment (Figs 8A, 9A). Material examined Holotype BANGLADESH • ♀ (see Fig. 9 A–H); Frederick John Parry leg.; OUMNH. Additional material MYANMAR • 1 Ƌ; Nat Ma Taung; Oct. 2003; 2000 m; local collector leg.; RBINS (see Figs 8 A–F, 10A–F, 15E–I). CHINA • 1 ♀; Maku; 20 Nov. 1973; Yun-xing Gan leg.; NWAFU HO088506 • 1 ♀; locality unknown; 21 Dec. 1973; Yun-xing Gan leg.; NWAFU HO088507. Description MEASUREMENTS. Body length: ♀ 12.5–12.6 mm; Ƌ 12.4 mm. Wingspan: ♀ 47.5–48.4 mm; Ƌ 47 mm. MALE GENITALIA. Pygofer in profile with ventral margin slightly concave; laterocaudal margin slightly convex (Figs 10A, 15E). Anal tube moderate, apical margin truncate, intersection with dorsal margin at obtuse angle in lateral view, epiproct acutely angled apically, paraproct long, subulate in dorsal view (Figs 10A, C, 15E). Gonostyli subtriangular, apically rounded in lateral view (Figs 10A, 15E). Endosomal processes about 5.0 times as long as sheath, apical ½ exposed, apically sinuate and inflated (Figs 10 D–F, 15G–I). Tectiductus unrecognizable in lateral view. Distribution China (Xizang), Bangladesh (Hope 1843)., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 49-52, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Hope F. W. 1843. On some rare and beautiful insects from Silhet, chiefly in the collection of Frederick John Parry, Esq. F. L. S. The Transactions of the Linnean Society of London 19 (2): 131 - 136. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00358. x","Stal C. 1863. Beitrag zur Kenntnis der Fulgoriden. Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin 24: 230 - 251.","Distant W. L. 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota. Vol. 3: Heteroptera-Homoptera. Taylor & Francis, London.","Metcalf Z. P. 1947. General Catalogue of the Hemiptera. Fasc. IV: Fulgoroidea. Part. 9: Fulgoridae. North Carolina State College, Raleigh (NC).","Lallemand V. 1963. Revision des Fulgoridae (Homoptera). Deuxieme partie. Faunes asiatique et australienne. Memoires de l'Institut royal des Sciences naturelles de Belgique (2 e serie) 75. Institut royal des Sciences naturelles de Belgique, Brussels.","Nagai S. & Porion T. 1996. Fulgoridae 2. Illustrated Catalogue of the Asian and Australian Fauna. Sciences Nat, Venette."]}

Limois hunanensis Chou & Wang, 1985 Figs 3, 14 I–L Limois hunanensis Chou & Wang, in Chou et al. 1985a: 30. Limois hunanensis – Chou et al. 1985b: 110. Diagnosis (1) Frons with 2 longitudinal carinae (Fig. 3B), cephalic process of vertex long, surpassing anterior margin of pronotum (Fig. 3A, C); (2) pronotum and mesonotum creamy yellow, pronotum with 2 dark spots anteriorly, mesonotum with 3 pairs of dark patches along anterior margin and the middle pair covered by pronotum, one large irregular patch on each side of median carina, each lateral area with 1 large irregular patch (Fig. 3A); (3) tegmina with a transverse irregular dark brown band on basal ⅖ (not reaching posterior margin) and reddish on basal ⅖ and costal area followed by some brown patches and spots in apical half (Fig. 3D); hindwings red on basal half and hyaline on apical half, between them a sinuate blackish brown fascia; and (4) abdominal tergites yellowish brown, anterior margin black-brown on each tergite, sternites dark brown. Material examined Holotype CHINA • Ƌ (see Figs 3, 14 I–L); Hunan, Daoxian, alt. 1600 m; 27 Aug. 1982; Xin-wang Tong leg.; NWAFU HO088511. Description MEASUREMENTS. Body length (from the original description of Chou et al. 1985a): Ƌ (holotype) 9 mm. Wingspan: Ƌ (holotype) 40 mm. MALE GENITALIA. Pygofer in profile with ventral and laterocaudal margins almost straight (Figs 3E, 14I). Anal tube moderate, apical margin truncate, intersection with dorsal margin at obtuse angle in lateral view, epiproct acutely angled apically in dorsal view (Figs 3E, G, 14I). Gonostyli triangular in outline, widest near apex, apically almost truncate in middle (Figs 3 E–F, 14I–J). Endosomal processes mostly sclerotized, about 5.0 times as long as sheath, apical ½ exposed, slightly inflexed and inflated (Figs 3 H– I, 14K–L). Connective unrecognizable. Tectiductus small, unrecognizable more or less. Distribution China (Hunan).

Limois chagyabensis Chou & Lu, 1981 Figs 1, 14 A–D Limois chagyabensis Chou & Lu, 1981: 221. Limois chagyabensis – Chou et al. 1985b: 109. Diagnosis (1) Frons medially with 3 longitudinal carinae, median carina tinier, all not reaching fronto-clypeal suture (Fig. 1 E–F), cephalic process of vertex moderate in length, reaching posterior margin of vertex (Fig. 1D, G); (2) pronotum red-brown with one longitudinal broken band on each side of median carina, blackish brown, mesonotum yellow, with 3 pairs of dark patches along anterior margin and one dark triangular patch on each lateral area, with one large irregular patch on each side of median carina (Fig. 1D); (3) tegmina with a transverse irregular dark brown band on basal ⅖ (not reaching posterior margin) and orange-yellow on costal area and basal half followed by some brown patches and spots on hyaline area (Fig. 1 A–C); hindwings reddish yellow on basal half (red in fresh specimens), the apical half hyaline (Fig. 1 A–B); (4) abdominal segment dark, posterior margin testaceous (Fig. 1A). Material examined Holotype CHINA • ♀ (see Figs 1 A–G, 14A–D); Xizang, Chaya, Jitang; alt. 3200 m; 15 Sep. 1976; Xue-zhong Zhang leg.; NWAFU. Allotype CHINA • ♀; same data as for holotype; NWAFU. Additional material CHINA • 1 Ƌ; Shaanxi, Baliguan; 16 Aug. 1983; De-jin Zhao leg.; NWAFU HO088523 • 1 ♀; Shaanxi, Zhenba; 5 Oct. 1981; collector unknown; NWAFU HO088514 • 1 ♀; Sichuan, Wolong Nature Reserve; alt. 1940 m; 25 Nov. 1982; Jian-guo Yang leg.; NWAFU HO088508 • 1 ♀; Shaanxi, Liuba; Aug. 1980; Yan-hua Wei leg.; NWAFU HO088518. Description MEASUREMENTS. Body length: ♀ 9.8–12 mm; Ƌ 9.2 mm. Wingspan: ♀ 39.6–44.7 mm; Ƌ 34.8 mm. MALE GENITALIA. Pygofer in profile with ventral margin straight, laterocaudal margin slightly convex (Figs 1H, 14A). Anal tube moderate in length, ventral margin slightly concave in basal ¾, apical margin truncate, intersected with dorsal margin at almost right angle in lateral view, epiproct acutely angled apically in dorsal view (Figs 1H, J, 14A). Gonostyli subtriangular, apically almost straight in lateral view (Figs 1H, 14A). Endosomal processes sclerotized, about 3.5 times as long as sheath, apical half exposed, inflexed and inflated at apex (Figs 1 K–L, 14C–D). Connective linear (Fig. 14A). Tectiductus unrecognizable in lateral view. Distribution China (Xizang, Shaanxi, Sichuan). Remarks When listing the type specimens of this species, Chou & Lu (1981) stated that the holotype was male but in fact it is a female., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 37-39, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Chou I. & Lu J. S. 1981. Homoptera: Fulgoroidea. In: Chen S. X. (ed.) Insects of Xizang. The Series of the Expedition to the Qinghai-Xizang Plateau. Vol. 1: 225 - 226. Science Press, Beijing.","Chou I., Lu J. S., Huang J. & Wang S. Z. (eds) 1985 b. Economic Insect Fauna of China. Vol. 36: Homoptera: Fulgoroidea. Science Press, Beijing."]}

Limois bifasciatus Ollenbach, 1928 Fig. 13 Limois bifasciatus Ollenbach, 1928: 281. Material examined from photograph INDIA • 1 Ƌ, holotype of Limois bifasciatus (see Fig. 13); Mussoorie United Provinces; O.C. Ollenbach leg.; O.C. Ollenbach det.; FRI-NFIC. Distribution India., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 55-56, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Ollenbach O. C. 1928. New species of Cicadidae and Fulgoridae from India and Burma (Hemipt.). Indian Forest Records 13 (6): 271 - 282."]}

Genus Limois Stål, 1863 Limois Stål, 1863: 230. Type species: L. westwoodii (Hope, 1843). Limois – Metcalf 1947: 170. — Lallemand 1963: 54. — Chou & Lu 1981: 221. — Chou et al. 1985a: 30. — Chou et al. 1985b: 108. — Nagai & Porion 1996: 22. Re-description HEAD (including eyes). Much narrower than pronotum (Figs 8B, 9B). Vertex slightly produced in front of eyes, vertex about 2 times as broad as an eye, medially with reflexed protuberance (Fig. 9B, H). Pronotum almost twice as broad as vertex, with deeply impressed point at each side of median carina, each lateral area with one rounded dark spot behind eyes, lateral carinae sinuate. Mesonotum tricarinate in disc, median carina straight, intermediate carinae curved, reaching posterior margin (Figs 8B, 9B). Frons distinctly narrower from fronto-clypeal suture to apex, lateral margins carinate, medially with 2 or 3 longitudinal carinae; clypeus narrower and shorter than frons, labium elongate, median carina on clypeus (Figs 4C, 9D). Tegmina elongate, slightly broadened distad with the maximum width near apex, apically obliquely rounded, venation reticulate, tegmina hyperpterous, much longer than hindwing (Fig. 9A). Hindwing posterior margin indented at PCu level, apical cell and clavus venation reticulate (Fig. 8A). Metatibiae with 4–7 lateral spines (Fig. 9E). MALE GENITALIA. Pygofer symmetrical, narrow and subquadrangular in lateral view, ventrally longer than dorsally, ventrocaudal angle slightly produced caudad (Fig. 10A). Anal tube subtriangular in lateral view, in dorsal view gradually broadened distad, apical margin concave; epiproct and paraproct separated in lateral view, in dorsal view epiproct short and broad, paraproct slender, surpassing the end of anal tube (Fig. 10A, C). Gonostyli subtriangular or oval in lateral view, submedially with a hookshaped process near dorsal side, in ventral view the styles connect only at base (Fig. 10A, B). Aedeagus reduced, membranous endosoma with a dorsal pair of lobes surrounding a ventral pair of lobes and the sclerotized endosomal processes, the endosomal processes paired, apical portion exposed and curved dorsad in lateral view, terminally inflated (Fig. 10 D–F). Distribution China (northeast China, Shanxi, Shaanxi, Hebei, Beijing, Gansu, Ningxia, Xizang, Hunan, Sichuan, Guangxi, Fujian, Taiwan), Japan, Korea, Russia, Bangladesh, India, Myanmar (Fig. 16). Remarks Lallemand (1963) established the tribe Limoisini belonging to the subfamily Aphaeninae Blanchard, 1847 and assigned Limois, together with four other genera (Bloeteanella Lallemand, 1963, Erilla Distant, 1906, Hellerides Lallemand, 1963 which was synonymized with Zophiuma Fennah, 1955 by Liang (1995) in the family Lophopidae, and Neolieftinckana Lallemand, 1963) to this tribe. Nagai & Porion (1996) recorded seven genera (Bloeteanella, Erilla, Limois, Neolieftinckana, Nisax Fennah, 1977, Saramel Fennah, 1977 and Ombro Fennah, 1977) in Limoisini. The genus Limois differs from the other six genera in the tribe by the hindwings bicolored (red or yellow in basal part and hyaline apically) rather than concolorous over entire length. Limois also differs from Neolieftinckana by the tegmina hyaline in apical half (entirely opaque in Neolieftinckana). From Ombro it differs by the phallobase having 4 paired lobes (with 5 paired lobes in Ombro). From Erilla and Nisax it differs by the cephalic process of vertex slanting upwards to backwards, flattened on the vertex (cephalic process of vertex reduced in Erilla and forming a granular mass in middle of vertex in Nisax). From Bloeteanella it differs by the frons having 2 or 3 smooth longitudinal carinae (wrinkled and grooved carinae in Bloeteanella)., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 36-37, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715, {"references":["Stal C. 1863. Beitrag zur Kenntnis der Fulgoriden. Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin 24: 230 - 251.","Hope F. W. 1843. On some rare and beautiful insects from Silhet, chiefly in the collection of Frederick John Parry, Esq. F. L. S. The Transactions of the Linnean Society of London 19 (2): 131 - 136. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00358. x","Metcalf Z. P. 1947. General Catalogue of the Hemiptera. Fasc. IV: Fulgoroidea. Part. 9: Fulgoridae. North Carolina State College, Raleigh (NC).","Lallemand V. 1963. Revision des Fulgoridae (Homoptera). Deuxieme partie. Faunes asiatique et australienne. Memoires de l'Institut royal des Sciences naturelles de Belgique (2 e serie) 75. Institut royal des Sciences naturelles de Belgique, Brussels.","Chou I. & Lu J. S. 1981. Homoptera: Fulgoroidea. In: Chen S. X. (ed.) Insects of Xizang. The Series of the Expedition to the Qinghai-Xizang Plateau. Vol. 1: 225 - 226. Science Press, Beijing.","Chou I., Wang S. Z. & Huang J. 1985 a. Description of new species of Fulgoroidea from China (Homoptera: Fulgoroidea). Entomotaxonomia 7: 30 - 38.","Chou I., Lu J. S., Huang J. & Wang S. Z. (eds) 1985 b. Economic Insect Fauna of China. Vol. 36: Homoptera: Fulgoroidea. Science Press, Beijing.","Nagai S. & Porion T. 1996. Fulgoridae 2. Illustrated Catalogue of the Asian and Australian Fauna. Sciences Nat, Venette.","Distant W. L. 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota. Vol. 3: Heteroptera-Homoptera. Taylor & Francis, London.","Liang A. P. 1995. Taxonomic change in Oriental Fulgoroidea (Hemiptera: Fulgoromorpha). Journal of the New York Entomological Society 103 (2): 162 - 164."]}

Limois sordida sp. nov. urn:lsid:zoobank.org:act: 4B5CE928-5341-4712-9B98-3F5DF7AA7F37 Figs 6–7, 15 A–D Diagnosis (1) Frons with 3 longitudinal carinae (Fig. 6E); (2) tegmina with along irregular stripe, sinuately extending to anal angle (Fig. 6 A–C); (3) gonostyli almost equilateral in lateral view; (4) hindwings pale yellow in basal ⅖, medially without fascia (Fig. 6 A–B). Etymology The specific epithet alludes to the sordid color of the clypeus. Type material Holotype CHINA • Ƌ; Shanxi, Ningwu, Mt. Luyashan; 25 Oct. 2011; Ai-ping Dong leg.; NWAFU. Paratypes CHINA • 4 ƋƋ, 2 ♀♀; same data as holotype; NWAFU • 2 ♀♀; Beijing, Mt. Baihuashan, Huang Tuo village; 9 Sep. 1960; Fa-sheng Li leg.; CAU • 1 Ƌ; Beijing, Mt. Baihuashan, Huang Tuo village; 7 Sep. 1960; Ji-kun Yang leg.; CAU • 1 Ƌ; Beijing, Mt. Baihuashan; 5 Sep. 1961; Fa-sheng Li leg.; CAU • 1 Ƌ; Hebei, Mt. Xiaowutaishan; 3 Aug. 2011; Jian Yao leg.; IZCAS IOZ(E)2058877 • 1 Ƌ; Hebei, Mt. Xiaowutaishan; 4 Aug. 2011; Jian Yao leg.; IZCAS IOZ(E)2058878 • 1 ♀; Hebei, Mt. Xiaowutaishan; 3 Aug. 2011; Jian Yao leg.; IZCAS IOZ(E)2058876. Description MEASUREMENTS. Body length: ♀ 10.6–11.4 mm; Ƌ 9.2–10.2 mm. Wingspan: ♀ 35.7–36.5 mm; Ƌ 31.5– 32.7 mm. HEAD. Including eyes distinctly narrower than pronotum (about 1: 1.7) (Fig. 6D). Vertex quadrangular, about 2 times as broad as an eye, lateral and posterior margins carinate, median carina obscure; cephalic process flattened, not reaching posterior margin of vertex (Fig. 6D, F). Frons medially with 3 longitudinal carinae, the median carina tinier, all not reaching fronto-clypeal suture; clypeus sordid, narrower and shorter than frons, labium elongate, reaching the end of abdomen, median carina on clypeus (Fig. 6E). THORAX. Thorax macular covered with numerous dark spots, pronotum about 1.6 times as long as vertex in dorsal view, with one longitudinal broken band on each side of median carina, black (Fig. 6D); mesonotum with 3 pairs of dark patches along anterior margin and one dark triangular patch on each lateral area (Fig. 6D). TEGMINA. Brownish yellow on basal ⅖ and costal area, with narrow irregular stripe from costal area, extending to anal angle; remaining area of tegmina hyaline with some dark markings, veins ochre brown, about 2.7 times as long as maximum broad (Fig. 6 A–C). HINDWINGS. Pale yellow on basal ⅖, with 4–5 black spots arranged in one oblique row, apical ⅗ of hindwings hyaline with a brown band along suture margin (Fig. 6 A–B). LEG. Brown with black rings and stripes. Metatibiae with 5–8 lateral and 6–7 apical spines. ABDOMEN. Tergites black, posterior margin ochre yellow on each segment (Fig. 6A). MALE GENITALIA. Pygofer in profile with ventral margin straight, laterocaudal margin slightly convex (Figs 7A, 15A). Anal tube moderate, in lateral aspect reaching level of end of genital plates, ventral margin slightly concave in basal ¾, apical margin truncate, intersecting with dorsal margin at obtuse angle, epiproct right-angled apically, paraproct slender in dorsal view (Figs 7 A–B, 15A). Gonostyli subtriangular in lateral view, rounded apically, apical margin straight in middle, in ventral view plates connected in basal ⅕ (Figs 7A, C, 15 A–B). Aedeagus reduced, membranous endosoma with a dorsal pair of lobes surrounding a ventral pair of lobes and the sclerotized endosomal processes; the endosomal processes sclerotized over entire length, about 4.0 times as long as sheath, apical ½ exposed and terminally inflated (Fig. 7 C–E, 15C–D). Connective rod-like. Tectiductus large, in lateral view subrectangular, in dorsal view broad, apically rounded and distinctly concave medially (Figs 7 C–E, 15A). Distribution China (Shanxi; Hebei, Beijing). Remarks Limois sordida sp. nov. is similar to L. bifasciatus, L. emelianovi and L. kikuchii, but differs from L. bifasciatus by the frons having three longitudinal carinae (Fig. 6E) (with two longitudinal carina in L. bifasciatus); from L. emelianovi and L. kikuchii it differs by the irregular stripe on tegmina long, sinuately extending to anal angle (Fig. 6 A–C) (irregular stripe short and straight, not extending to anal angle in L. emelianovi and L. kikuchii). Furthermore, it differs from L. kikuchii by the gonostyli being almost equilateral in lateral view (Figs 7A, 15 A–B) (elongate in L. kikuchii); from L. emelianovi it differs from the basal ⅖ of hindwings pale yellow in both sexes, medially without fascia (Fig. 6 A–B) (base of hindwings yellow in male but red in female and medially with a blackish brown fascia in L. emelianovi)., Published as part of Wang, Wen-Qian, Constant, Jérôme & Qin, Dao-Zheng, 2020, Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China, pp. 35-61 in European Journal of Taxonomy 720 on pages 46-49, DOI: 10.5852/ejt.2020.720.1113, http://zenodo.org/record/4071715

Wang, Wen-Qian, Constant, Jérôme, Qin, Dao-Zheng (2020): Revision of the lanternfly genus Limois Stål, 1863 (Hemiptera: Fulgoromorpha: Fulgoridae) with description of a new species from China. European Journal of Taxonomy 720: 35-61, DOI: https://doi.org/10.5852/ejt.2020.720.1113

Ricanula hainanensis sp. nov. (Figs 35–49) Etymology. The species was named after Hainan Island (South China)—the type locality. Description. Length (female, inc. teg.): 9.7 mm. Head. Head with compound eyes (in dorsal view, Figs 35, 38) about as wide as widest part of mesonotum. Vertex (Figs. 35, 38) 7 times wider in the anterior margin than long in midline, anterior and posterior margins arcuate, posterior margin with larger curvature than anterior one. Frons (Fig. 37) wider then long in midline, 1.17 times wider at upper margin than long in midline and in widest part, below ocelli level 1.31 times; upper margin straight, lateral margins arcuate, elavated and without lateral incisions; disc of frons tricarinate, carinae separated at base; median carina extending half of length in midline, lateral carinae arcuate, partly parallel to lateral margins, finishing a little before the median one. Frontoclypeal suture distinctly arcuate. Clypeus distinctly narrower than frons, without carinae, median part convex. Compound eyes (Fig. 36) oval in lateral view, with small callus at posteroventral margin. Ocelli present. Rostrum reaching mesotrochanters, apical segment shorter than subapical. Thorax. Pronotum (Figs 35, 38) 2.16 times longer in midline than vertex; disc of pronotum with median carina and 2 lateral incisions; anterior margin arcuate, posterior margin with strongly curvation than anterior one. Mesonotum (Figs 35, 38) 1.12 times longer than wide between lateral angles and about 3.95 times longer than cumulative length of vertex and pronotum in midline; disc of mesonotum with median and lateral keel-shape carinae, connected at base, lateral carinae reaching almost posterior margin; anterolateral carinae present, short, not connected with lateral, almost parallel to lateral ones not surpassing the lateral angles of mesonotum. Tegmina (Figs 35–36, 39) membranous, elongately-triangular; costal margin weakly arcuate, anterior angle rounded, placed distad to claval angle, apical margin irregular. Costal area with sparse and curved transverse veinlets, wider than postcostal cell and wider apically, ending before apex of clavus; postcostal cell without transverse veinlets; basal cell elongate and wide; longitudinal veins ScP+RA, MP and CuA leaving basal cell separately; ScP+R veins and MP forked immediately after leaving basal cell, CuA forked before of connection of Pcu and A1. Claval veins Pcu and A1 fused after midle of clavus with transverse veinlets present between CuP and Pcu, CuP and Pcu+A1 and between Pcu and A1. Tegmina with sparse transverse veinlets in median and posterior parts, veinlets below MP3+4 vein more dense, apical line of transverse veinlets present, apical cells distinctly longer than wide. Wings with elongate and narrow precostal cell; 2 transverse veinlets r-m and m-cu present (Fig. 35). Profemora about as long as tibiae, profemur partly laterally flattened more V shape in cross section, tibiae square in cross section; mesotibiae longer than mesofemora, femora laterally flattened, rectangular in cross section, tibiae square in cross section; metafemur shorter than metatibiae, curved, metatibiae partly flattened apically, with 2 lateral spines and 6 apical teeth; basitarsomere with 6 apical teeth, about as long than cumulative length of second and hind tarsomere. Metatibiotarsal formula 2/6/6. Coloration. Vertex yellow with 2 brown patches near the antero-lateral part and with 3 small light brown dots along anterior margin. Frons yellow, lateral part of head yellowish with 3 small brown patches at upper part, lower part brownish. Clypeus brownish with median yellow strip. Pronotum yellow. Median portion of mesonotum between lateral margins yellowish, lateral parts brown, scutellum dark brown. Tegmina piceous-brown with yellowish transverse veinlets below MP3+4 and on the clavus, costal margin with oblique dark brown stripes and yellow patches between; basal cell with dark brown patch near posterior margin; median portion of tegmen with rounded brown patch, postero-apical part with eyes-spot black cell. Wings and legs brownish. Abdomen light brown, basal sternits yellow. Male. Unknown. Female terminalia. Pregenital sternite (Fig. 41) with well developed, elongately-rounded and distinctly separated lateral lobes; anterior margin weakly concave, posterior margin medially with strong triangular and elongate process. Anal tube (in lateral view, Fig. 43) elongate, not surpassing posterior margin of the gonoplac; basal part narrower than posterior one; ventral margin arcuate, ventroposterior angle widely rounded; anus placed dorsally about midlength. Anal tube, in dorsal view (Fig. 42), rounded; with basal part distinctly narrower than posterior one, basal margin rounded, posterior margin widely concave medially; lateral margins arcuate; anus placed a bit before midlength. Gonoplac: laterally flattened, in lateral view (Fig. 44) with posterior part wider than at base, posterior margin with 2 rows of teeth (Fig. 45); membranous part of gonoplac well developed, placed medially on ventral margin. Gonapophysis VIII (Fig. 46) sabre-like, “v” shape in cross section, with 11 teeth at dorsal margin; endogonocoxal process with spiniferous microsculpture, well sclerotized medially, lateral parts membranous, tapering apicad, reaching apex of gonapophysis VIII. Gonaphophyses IX and gonospiculum bridge well developed as in Figs 47–48. Bursa copulatrix with widely connected two pouches; wall of first pouch with well visible cells and small central sclerotized ornamentation bearing single narrow teeth with sharp and curved apex, mostly placed at lower part, second pouch membranous with very weakly visible cells and without ornamentation. Spermatheca (Fig. 49) well developed; ductus receptaculi elongate and ribbed, tapering apicad; diverticulum ductus a bit shorter than ductus receptaculi, with smooth basal ductus and elongate smooth bulba at apex. Type material. Holotype, 1♀, labeled (Fig. 40): [Mt. Wuchi, Hainan, May 1903]. Distribution. China: Hainan Province (Hainan Island).

Key to species of Ricanula (females) from China 1. Pregenital sternite with posterior margin almost straight, medially slightly incised (Fig. 17)............. R. unica sp. nov. - Pregenital sternite with posterior margin convex medially..................................................... 2 2. Median portion of the posterior margin with strong process (Fig. 41)............................ R. hainanensis sp. nov. - Median portion of the posterior margin without single strong process........................................... 3 3. Median portion of posterior margin with bilobate process....................................... R. pulverosa (St��l) - Median portion of posterior margin without process (Fig. 34).................................. R. fujianensis sp. nov., Published as part of Ren, Lan-Lan, Stroi��ski, Adam & Qin, Dao-Zheng, 2016, Three new species of the genus Ricanula Melichar, 1898 (Hemiptera: Fulgoromorpha: Ricaniidae) from China, pp. 557-569 in Zootaxa 4168 (3) on page 558, DOI: 10.11646/zootaxa.4168.3.7, http://zenodo.org/record/261468

Checklist of Alebroides genus group in China Alafrasca Lu & Qin, 2014 a A. sticta Lu & Qin, 2014 a���China (Sichuan) Alebrasca Hayashi & Okada, 1994 A. actinidiae Hayashi & Okada, 1994 ��� China (Hunan, Henan, Gansu, Sichuan, Fujian, Zhejiang, Jiangxi); Japan A. expansa (Dworakowska, 1995) ��� China (Taiwan) Alebroides Matsumura, 1931 A. barbatus Dworakowska, 1997 ��� China (Hainan, Hong Kong); Nepal; Thailand A. chiasmaticus Yu & Yang, 2014 ��� China (Guangxi) A. discretus Chou & Zhang, 1987 ��� China (Shaanxi, Gansu, Hubei, Sichuan, Guangxi, Guizhou) A. dworakowskae Chou & Zhang, 1987 ��� China (Shaanxi, Gansu, Hubei, Hunan, Guizhou, Sichuan, Guangdong, Chongqing, Yunnan, Guangxi) A. falcatus Sohi & Dworakowska, 1979 ��� China (Shaanxi, Sichuan, Yunnan, Guizhou, Chongqing); India, Nepal A. flavifrons Matsumura, 1931 ��� China (Guangdong, Hunan, Guizhou, Zhejiang, Taiwan); Japan A. fumosus Sohi & Dworakowska, 1979 ��� China (Yunnan); India; Nepal A. haedus Sohi & Dworakowska, 1979 ��� China (Yunnan, Tibet); India A. luteus Sohi & Dworakowska, 1979 ��� China (Yunnan); India A. marginatus Matsumura, 1931 ��� China (Hong Kong, Taiwan); South Korea; Japan A. muzitaneus Qin & Zhang, 2011 ��� China (Hunan, Guangxi, Sichuan, Guizhou, Chongqing) A. nigroscutellus (Distant, 1918) ��� China (Hunan, Yunnan, Sichuan, Hainan, Taiwan); India; Nepal, Thailand; Indonesia; Japan A. obliteratus Dworakowska, 1997 ��� China (Yunnan); India; Nepal A. parafuscus Qin & Zhang, 2011 ��� China (Yunnan, Gansu) A. rayi Dworakowska, 1997 ��� China (Yunnan); Myanmar A. rubicundus Ishihara, 1953 ��� China (Henan, Hubei, Hunan); Japan A. salicis (Vilbaste, 1968) ��� China (Shaanxi, Hunan, Yunnan, Guizhou, Sichuan, Taiwan); Korea; Far East of Russia A. serrulatus Yu & Yang, 2014 ��� China (Tibet) A. shaanxiensis Chou & Zhang, 1987 ��� China (Shaanxi, Gansu, Hubei) A. sinuatus Dworakowska, 1980 ��� China (Shaanxi); India; Nepal A. similis Dworakowska, 1977 ��� China (Shaanxi, Yunnan, Sichuan); Vietnam A. shirakiellus (Matsumura, 1931) ��� China (Hunan, Henan, Sichuan, Yunnan, Guangdong, Guangxi, Taiwan) A. spanner Yu & Yang, 2014 ��� China (Guizhou) A. strumae Yu & Yang, 2014 ��� China (Tibet) A. shokanus Matsumura, 1931 ��� China (Taiwan) A. toroensis Matsumura, 1931 ��� China (Taiwan); Japan A. ursulae Dworakowska, 1997 ��� China (Taiwan) A. vicarius Dworakowska, 1997 ��� China (Yunnan); Myanmar A. waimingi Dworakowska, 1997 ��� China (Hong Kong) A. wushensis Dworakowska, 1997 ��� China (Sichuan, Taiwan) A. yanglinginus Chou & Zhang, 1987 ��� China (Shaanxi, Gansu, Hubei) Apheliona Kirkaldy, 1907 A. ferruginea (Matsumura, 1931) ��� China (Zhejiang, Hubei, Hunan, Guangdong, Hainan, Sichuan, Yunnan, Shaanxi, Taiwan); Japan; India A. indica Dworakowska & Sohi, 1978 ��� China (Shaanxi); India A. robusta Dworakowska, 1994 ��� China (Taiwan) A. taiwana Dworakowska, 1994 ��� China (Taiwan) A. xerophila Dworakowska, 1994 ��� China (Taiwan) A. zhangi Dworakowska, 1994 ��� China (Beijing, Hebei) Circinans Lu & Qin, 2014 b C. striata Lu & Qin, 2014 b���China (Fujian, Guizhou, Hunan) Flaviata Lu & Qin, 2014 F. v a r i a t a Lu & Qin, 2014 ��� China (Sichuan, Zhejiang) F. longa Xu, Lu & Qin, 2015 ��� China (Yunnan) Ghauriana Thapa, 1985 G. sinensis Qin & Zhang, 2011 ��� China (Guangxi, Yunnan) Keumiata Qin & Dietrich, 2014 K. orientalis Qin & Dietrich, 2014 ��� China (Yunnan); Thailand Lumicella Lu & Qin, 2013 L. rotundata Lu & Qin, 2013 ��� China (Fujian) Luodianasca Qin & Zhang, 2008 L. recurvata Qin & Zhang, 2008 ��� China (Guizhou) Membranacea Qin & Zhang, 2011 M. spinata Qin & Zhang, 2011 ��� China (Hunan) M. unijugata Qin & Zhang, 2011 ��� China (Sichuan, Shaanxi) M. plana Qin & Zhang, 2011 ��� China (Hunan) M. hubeiensis Yu & Yang, 2013 ��� China (Hubei) M. stenoprocessa Yu & Yang, 2013 ��� China (Hubei, Guizhou) Nikkotettix Matsumura, 1931 N. cuspidata Qin & Zhang, 2003 ��� China (Hunan, Sichuan, Guizhou, Yunnan, Guangxi) N. taibaiensis Qin & Zhang, 2003 ��� China (Shaanxi) N. erythrinae Dworakowska, 1995 ��� China (Yunnan); Indonesia; Nepal N. galloisi Matsumura, 1931 ��� China (Zhejiang, Shaanxi); Japan Rubiparvus Xu, Dietrich & Qin, gen. nov. R. bistigma Xu, Dietrich & Qin, sp. nov. ��� China (Yunnan) Schizandrasca Anufriev, 1972 S. ussurica (Vilbaste, 1968) ��� China (Hubei, Shaanxi); Far East of Russia; Korea, Published as part of Xu, Ye, Wang, Yu-Ru, Lu, Si-Han, Dietrich, Christopher H. & Qin, Dao-Zheng, 2016, Rubiparvus bistigma, a new genus and species of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae), with a checklist of the Alebroides group in Chinese fauna, pp. 583-589 in Zootaxa 4109 (5) on pages 587-588, DOI: 10.11646/zootaxa.4109.5.6, http://zenodo.org/record/259784, {"references":["Lu, S. - H. & Qin, D. - Z. (2014 a) Alafrasca sticta, a new genus and species of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) with a checklist of the tribe from China. Zootaxa, 3779 (1), 9 - 19. http: // dx. doi. org / 10.11646 / zootaxa. 3779.1.4","Hayashi, M. & Okada, T. (1994) A new Typhlocybinae leafhopper (Homoptera: Cicadellidae) feeding on kiwi-fruit. Applied Entomology and Zoology, 29 (2), 267 - 271.","Dworakowska, I. (1995) Szara gen. nov. and some other Empoascini (Insecta: Auchenorrhyncha: Cicadellidae: Typhlocybinae). Entomologische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 56, 129 - 160.","Matsumura, S. (1931) A revision of the Palaearctic and Oriental Typhlocybid-genera with descriptions of new species and new genera. Insecta Matsumurana, 6 (2), 55 - 91.","Dworakowska, I. (1997) A review of the genus Alebroides Matsumura, with description of Shumka gen. nov. (Homoptera: Auchenorrhyncha: Cicadellidae). Oriental Insects, 31, 241 - 407. http: // dx. doi. org / 10.1080 / 00305316.1997.10433759","Yu, X. - F. & Yang, M. - F. (2014) Four new species of Alebroides Matsumura (Hemiptera: Cicadellidae: Typhlocybinae) from China. Zootaxa, 3780 (2), 248 - 262. http: // dx. doi. org / 10.11646 / zootaxa. 3780.2.2","Chou, I. & Zhang, Y. - L. (1987) A taxonomic study of the genus Alebroides Mats. from China (Homoptera: Cicadellidae: Typhlocybinae). Entomotaxonomia, 9, 289 - 302.","Sohi, A. S. & Dworakowska, I. (1979) New species of Alebroides Matsumura from India and Tibet (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Entomon (India), 4 (4), 367 - 372.","Distant, W. L. (1918) Rhynchota. VII. Homoptera: appendix. Heteroptera: addenda. The fauna of British India, including Ceylon and Burma. Taylor & Francis, London, 210 pp.","Ishihara, T. (1953) A tentative check list of the superfamily Cicadelloidea of Japan (Homoptera). The scientific reports of the Matsuyama agricultural College, 11, 1 - 72.","Vilbaste, J. (1968) Uber die Zikadenfauna des Primorje Gebietes. Valgus, Tallinn, 178 pp.","Dworakowska, I. (1980) On some Typhlocybinae from India (Homoptera, Auchenorrhyncha, Cicadellidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fur Tierkunde in Dresden, 43 (8), 151 - 201.","Dworakowska, I. (1977) On some Typhlocybinae from Vietnam (Homoptera: Cicadellidae). Folia Entomologica Hungarica. 30 (2), 9 - 47.","Kirkaldy, G. W. (1907) Leafhoppers supplement (Hemiptera). Report of work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology bulletin, 3, 1 - 186.","Dworakowska, I. & Sohi, A. S. (1978) Kadrabia gen. n. and some other Typhlocybinae (Auchenorrhyncha, Cicadellidae) from India. Bulletin de l'Academie Polonaise des Sciences. Serie des Sciences Biologiques, 26 (7), 463 - 471.","Dworakowska, I. (1994) A review of the genera Apheliona Kirk. and Znana gen. nov. (Auchenorrhyncha: Cicadellidae: Typhlocybinae). Oriental Insects, 28, 243 - 308. http: // dx. doi. org / 10.1080 / 00305316.1994.10432308","Lu, S. - H. & Qin, D. - Z. (2014 b) A new genus and species in the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) from China. ZooKeys, 386, 85 - 91. http: // dx. doi. org / 10.3897 / zookeys. 386.7020","Xu, Y., Lu, S. - H. & Qin, D. - Z. (2015) Flaviata longa, a new species in Flaviata Lu & Qin and new synonymies in Empoasca (Matsumurasca) Anufriev (Hemiptera: Cicadellidae: Empoascini). Zootaxa, 4027 (2), 296 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 4027.2.9","Thapa, V. K. (1985) Some empoascan leafhoppers (Homoptera, Cicadellidae, Typhlocybinae) from the Kathmandu valley, Nepal. Journal of Entomological Research, 9, 65 - 74.","Qin, D. - Z., Lu, S. - H. & Dietrich, C. H. (2014) A key to the genera of Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) in China, with descriptions of two new genera and two new species. Florida Entomologist, 97 (4), 1493 - 1510. http: // dx. doi. org / 10.1653 / 024.097.0425","Lu, S. - H., Zhang, L., Qiao, L. & Qin, D. - Z. (2013) Lumicella, a new genus of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) from China. ZooKeys, 364, 11 - 17. http: // dx. doi. org / 10.3897 / zookeys. 364.6618","Qin, D. - Z. & Zhang, Y. - L. (2008) Two new empoascine leafhopper genera and species (Hemiptera: Cicadellidae: Typhlocybinae) from southern China, with a key to Chinese genera of Empoascini. Zootaxa, 1966, 62 - 68.","Yu, X. - F. & Yang, M. - F. (2013) Two new species of Membranacea Qin & Zhang from China (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini). ZooKeys, 260, 77 - 83. http: // dx. doi. org / 10.3897 / zookeys. 260.4560","Qin, D. - Z. & Zhang, Y. - L. (2003) Taxonomic study of Nikkotettix (Homoptera: Cicadellidae: Typhlocybinae: Empoascini) - new record from China. Entomotaxonomia, 25, 25 - 30.","Anufriev, G. A. (1972) New and little known Palaearctic genera and species of Typhlocybinae (Homoptera, Cicadellidae). Bulletin de l'Academie Polonaise des Sciences. Serie des Sciences Biologiques, 20 (1), 35 - 42."]}

Xu, Ye, Wang, Yu-Ru, Lu, Si-Han, Dietrich, Christopher H., Qin, Dao-Zheng (2016): Rubiparvus bistigma, a new genus and species of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae), with a checklist of the Alebroides group in Chinese fauna. Zootaxa 4109 (5): 583-589, DOI: http://doi.org/10.11646/zootaxa.4109.5.6

Ricanula fujianensis sp. nov. (Figs 18���34) Etymology. The species name is derived from the type locality, Fujian Province. Diagnosis. Ricanula fujianensis sp. nov. is similar to R. pulverosa (St��l), but differs from the latter in having numerous yellowish transverse veinlets in inner half of tegmina (with few transverse veinlets on basal portion in R. pulverosa); ventral periandrium of the new species bearing a pair of processes (ear-like structure with almost transparent bar-like process beyond it), bending dorsal and oriented to phallic complex (ventral periandrium without processes in R. pulverosa); apical spinose processes of aedeagus long, surpassing half length of the phallic complex (apical spinose processes short, not reaching half length of the phallic complex in R. pulverosa). Description. Length (inclu. teg.): male 8.7���9.8 mm, female 9.3���10.8 mm. Head. Head with compound eyes nearly as wide as the widest part of mesonotum (Fig. 20). Vertex narrower than pronotum, about 6.0 times wider at the anterior margin than long in midline, all margins well carinate, posterior margin arcuate more strongly than anterior margin; disc of vertex without median carina, from anterior to posterior margins downwardly sloped (Fig. 20). Frons 1.31 times wider at widest part (about the level of lower margin of ocelli), than long in midline, a little longer at upper margin than high in midline; upper margin slightly arcuate, lateral margins arcuate, in lower part curved to frontoclypeal suture; frontal disc with 3 carinae separated basally, lateral carinae arcuate, reaching the level of the midlength of the median carina (Fig. 21); frons oriented postero-ventral (Fig. 19). Frontoclypeal suture arcuate (Fig. 21). Clypeus ecarinate, triangular, not in the same plane than frons and distinctly narrower, median portion convex (Fig. 21). Compound eyes oval, with callus at lower margin (Figs 18���21). Ocelli present (Figs 19, 21). Rostrum reaching mesotrochanters, apical segment shorter than subapical. Thorax. Pronotum distinctly longer in midline than vertex; disc of pronotum with median carina, anterior and posterior margins arcuate, in median portion almost parallel (Fig. 20). Mesonotum about 4.2 times longer than cumulative length of vertex and pronotum in midline; median carina keel-shaped, almost reaching scutellum; lateral carinae not connected basally, almost reaching posterior margins; anterolateral carinae not fused with lateral carinae, not surpassing the lateral angles of mesonotum (Fig. 20). Tegmina (Figs 18���19, 22) membranous, elongately-triangular; costal margin weakly arcuate, slightly incised near apical angle, anterior angle broadly rounded, placed distad to claval angle, posterior margin almost straight. Costal area with sparse transverse veinlets, a little wider than postcostal cell and a little widened apically, postcostal cell without transverse veinlets; basal cell small, widely rounded; veins ScP+R, MP and CuA leaving basal cell separated. ScP+R vein forked just after leaving basal cell, MP dividing into MP1+2 and MP3+4 basally, CuA forked before middle of clavus. Claval veins Pcu and A1 fused about midlength of clavus. Tegmen without lines of transverse veinlets, in inner half with numerous irregular transverse veinlets. Wings small, costal area present and small; postcostal cell distinctly longer than wide, 2 transverse veinlets r-m and m-cu present (Fig. 22). Pro- and mesofemora as long as pro- and mesotibia, both square in cross section; metafemur square, shorter than metatibiae, metatibiae partly flatted, with 2 lateral spines and 6 apical teeth; basitarsomere with 6~8 apical teeth, a little longer than cumulative length of second and hind tarsomere. Metatibiotarsal formula 2/6/6���8. Coloration. General color of body dark brown to dark (Figs 18���19). Vertex with 2 brown reddish stains at each posterolateral corner, lateral margins pale yellow (Fig. 20). Pronotum brown (Fig. 20). Mesonotum dark brown, sometimes with 2 brown reddish spots anterolaterally (Fig. 20). Frons brown, clypeus and rostrum brownish (Fig. 21). Eyes sordid brown, ornamented with irregular black patches (Figs 18���21). Ocelli red. Gena light brown with three pale white spots. Tegmina piceous-brown, costal margin with about 10 transverse black stripes from base to a little beyond middle, between the stripes filled with greyish-white patches, submedially with a large pale flavescent or greyish-white spot marked by two central transverse black lines, the inner half of tegmina filled with numerous pale yellowish transverse veinlets (Figs 19, 22). Wings brown, A2 ornamented with longitudinal grayish band on both sides (Fig. 23). Pro-, mesofemora and tibiae brown. Metafemora and metatibiae brown yellowish. Abdomen dark brown. Male terminalia. Anal tube, in dorsal margin, with basal margin, shorter than posterior margin, posterior margin concave, basal margin straight, lateral margins arcuate, anus placed in middle, paraproct not surpassing the posterior margin (Fig. 30). Anal tube, in lateral view, far surpassing the end of pygofer; ventral margin convex (Fig. 24). Pygofer, in lateral view, higher than wide; dorsally narrower than ventrally, posterior margin almost straight; posterior-dorsal angle without process, caudo-dorsal angle not angulate (Fig. 24). Genital styles in lateral view obviously longer than wide, with a spine-like process at the end of dorsal margin; upper margin convex, lower margin mostly straight; ventral margin in caudo-dorsal angle widely rounded and surpassing the posterior margin of process (Fig. 24). Phallic complex. Phallic complex, in lateral view, broad and short, bent at middle; periandrium with lateral split longer than half of its length, lateral lobes present; periandrium, in dorsal view, longer than aedeagus, upper margin of dorsal periandrium divided in middle, lateral lobes slightly bent inside; in ventral view the upper of ventral margin with 2 ear-shaped lobes, the apical lobe bearing bar-shaped transparent process (in lateral and dorsal views), basal part of periandrium without any additional structures; dorsal periandrium shorter than ventral (Figs 31���33). Aedeagus shorter than periandrium, with 2 pairs of well sclerotized, smooth and spinose processes in lateral view, each process with a single apex, apical process longer than subapical one. Processes long, a little shorter than periandrium, both oriented basad (Figs 31���33). Female terminalia. Pregenital sternite with lateral lobes well developed, median portion strongly narrow; anterior margin weakly concave medially; posterior margin straight with slightly convex median portion (Fig. 34). Anal tube in lateral view short, ventral margin convex (Fig. 25). Anal tube in dorsal view, 1.3 times longer in midline than wide at widest part (widest submedially), lateral margins convex, basal margin slightly concave, posterior margin concave, anus placed after midlength, paraproct surpassing the posterior margin of anal tube (Fig. 29). Gonoplac: posterior margin bearing 2 well visible rows of blunt and short teeth (the upper margin 2-3 melt rows), posterior ventral part partly membranous (Fig. 26). Gonapophysis VIII partly flattened laterally, tapering apicad; dorsal margin slightly concave, with sharp apex and well visible teeth at posterodorsal margin, near apex with spiniferous microsculpture; endogonocoxal process narrower and shorter than gonaphophysis VIII, smooth (Fig. 28). Gonapophysis IX with posterior connective lamina sclerotized, gonospiculum bridge finger-like caudodorsally, needle-like ventro-dorsally (Fig. 26). Type material. Holotype, 1♂: [China: Fujian, Wuyi mountain, Tongmucun, 850 m, coll. Manqiang Wang and Bin Xiao, 16 Aug. 2008]. Paratypes: 6♂♂ and 5♀♀, same data as holotype; 9♂♂ and 10♀♀: [Fujian, Sanming, Longxi mountain, 725 m, col. Fengjuan Ren, 2 Aug. 2013]. Distribution. China: Fujian Province., Published as part of Ren, Lan-Lan, Stroi��ski, Adam & Qin, Dao-Zheng, 2016, Three new species of the genus Ricanula Melichar, 1898 (Hemiptera: Fulgoromorpha: Ricaniidae) from China, pp. 557-569 in Zootaxa 4168 (3) on pages 562-566, DOI: 10.11646/zootaxa.4168.3.7, http://zenodo.org/record/261468