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Psallus (Hylopsallus) ramae n. sp. (Figs. 13 D���F; 14D���E; 15C���D; 16; 30G���K; 31K���N; 32H���K) Material examined. Holotype (♂). JAPAN: Honshu, Okayama Pref., Hiruzen Plateau, Kawakami Village, Kamiyubune [current Maniwa City, Hiruzen], 35.31, 133.62, Quercus mongolica, 8 Jun 2003, T. Yasunaga (AMNH) (AMNH _ PBI 00380709). Paratypes: JAPAN: Honshu, same data as for holotype, 11♂ 4♀ (TYCN); Honshu, Hiroshima Pref., Geihoku Town, Chojabaru [current Kita-Hiroshima, Yawata Plateau], 34.71, 132.17, UV lighting, 10���11 Jul 1993, K. Yoshizawa, 2♂ 2♀ (TYCN) (1♂ with USIs, 00380710); Shikoku, Tengu Plateau, 33.48 133.00, Quercus mongolica, 6 Jun 1999, M. Takai, 1♂ 1♀ (TYCN) (1♂ with USIs, 00380711). Additional material examined. JAPAN: Kyushu, Nagasaki Pref., Isahaya City, Oriyama, 32.956650, 130.08650, 750 m alt., Quercus mongolica, 22 May 2021, T. Yasunaga, 2♀ (TYCN) (00380712). Diagnosis. Recognized by its generally dark castaneous, oval body (Fig. 13D���E); small size; densely distributed silvery setae on dorsum (Figs. 13E, 30G); enlarged or sometimes fused dark spots on metatibia (Fig. 13D���E); male genital segment (pygophore) with moderately clustered stiff setae at left-lateral base (Fig. 30J���K); broad, triangular apical part of phallotheca (Fig. 31L); vesica with thick median projection (Fig. 31M), lacking apical spinulate lobe (cf. Fig. 14B) that is possibly modified to cup-shaped rim (Fig. 14D); relatively small, nearly symmetrical sclerotized rings (Fig. 15A); and relatively clear comb-like scaly microstructures on interramal sclerite (Fig. 32J). Most closely related to P. castaneae, this new species can be distinguished by above diagnostic characters. Description. Body dark castaneous or fuscous, oval, not sexually dimorphic; dorsum with densely distributed, simple, semierect setae recumbent, silvery, lanceolate setae (Fig. 13D���E, 30G). Head dark brown, relatively slanting, with both simple and silvery setae; buccula tinged with red. Antenna yellowish brown; extreme base (basal 1/8���1/7) of segment I infuscate; segment IV grayish yellow. Labium shiny chocolate brown, its apex reaching apex of mesocoxa; apical parts of segments II and III, and basal half of IV yellowish brown. Pronotum, mesoscutum and scutellum fuscous, weakly shining; thoracic pleura dark brown, with silvery setae; ventral margins of propleuron and mesepimeron creamy white; evaporative area of metathoracic scent efferent system relatively narrow (Fig. 30H). Hemelytra fuscous, weakly shining because of dense vestiture (Fig. 13E); anterior margin of cuneus sometimes narrowly pale or reddish; membrane smoky brown. All coxae and femora uniformly dark brown; apex of profemur yellowish brown; ventral surface of metafemur without spotted pattern; tibiae pale brown, with distinct spots or annulations at bases of brown spines; metatibial dark spots sometimes fused together and enlarged (Fig. 13D��� E); tarsi pale brown; pretarsal structure as in Fig. 30M; pulvilli relatively narrow. Abdomen shiny fuscous. Male genitalia (Figs. 14 D���E, 30J���K, 31K���N): male genital segment (pygophore) with moderately clustered stiff setae at left-lateral base (Fig. 30J���K); apical part of phallotheca broad, triangular (Fig. 31L); vesica lacking apical spinulate lobe with cup-shaped rim (Fig. 14D); median projection of vesica thickened (Fig. 31M). Female genitalia (Figs. 15 C���D; 32H���K): sclerotized rings relatively small, nearly symmetrical and mesally separated from each other (Fig. 15A); rings reduced, indistinct (Fig. 12F); posterior wall with relatively wide interramal lobe and interramal sclerite (Fig. 27 K���L); posterior margin sclerite weak (Fig. 32N���O); comb-like scaly microstructures on interramal sclerite relatively clear (Fig. 32J); apex of ovipositor (gonapophysis I) somewhat widened (Fig. 15D; 32K). Measurements: See Table 4. Etymology. Named in honor of Ram Keshari Duwal who greatly contributed to clarification of the Asian fauna of the Phylinae. Distribution. Japan (SW Honshu, Shikoku, Kyushu). Biology. The newly emerged adults were found in early summer (late May in Kyushu and early June in Honshu) and a univoltine life cycle is assumed for this new species. Almost of all available specimens (including teneral ones) were collected from a deciduous oak, Quercus mongolica Fisch. ex Ledeb. (Fagaceae), growing mainly in a cold temperate climatic zone, but the immature forms are yet to be confirmed. Remarks. Although this new species and P. castaneae Josifov are closely related and at first sight seem very similar to each other, the latter is widespread over the lowlands and urbanized zones from central Honshu to western Kyushu (Ishikawa et al., 2014, 2015; Komatsu, 2018; Miyazaki et al., 2020). Records of P. edoensis (cf. Figs. 13G, 29J���O) from Tokyo Metropolis by Ishikawa et al. (2014, 2015) were recently verified to be based on misidentification of P. castaneae. Host association of P. castaneae ranges across a variety of Fagaceae oaks (mostly inflorescences), such as Castanea spp., Castanopsis spp., Lithocarpus edulis (Makino) Nakai, Quercus accutissima Carruth., and Q. serrata Murray; the adults were frequently found to aggregate on inflorescence of Cinnamomum camphora (L.) J.Pres, Machilus thunbergii Siebold et Zucc. (Lauraceae) and Melia azedarach L. (Meliaceae). On the other hand, the distribution and plant association of P. ramae n. sp. appear to be restricted to Quercus mongolica in cool mountainous zones (over 600 m alt.) in southwestern Japan (Fig. 16). At three localities (Hiroshima, Okayama, and Kochi Prefectures), P. ramae n. sp. was found to co-occur with P. tonnaichanus; the latter is known to be associated with Quercus dentata Thunberg and Q. mongolica (Yasunaga, 2001d)., Published as part of Yasunaga, Tomohide, 2022, The plant bug subfamily Phylinae in Japan, with key to genera and descriptions of eight new species (Hemiptera: Heteroptera: Miridae), pp. 1-52 in Zootaxa 5094 (1) on pages 29-33, DOI: 10.11646/zootaxa.5094.1.1, http://zenodo.org/record/5964735, {"references":["Ishikawa, T., Yasunaga, T. & Ito, M. (2014) Rediscovery of Psallus edoensis after an interval of 59 years (Heteroptera: Miridae: Phylinae). Rostria, 56, 31 - 32. [in Japanese with English summary]","Ishikawa, T., Saito, M., Kishimoto-Yamada, K., Kato, T., Kurashima, O. & Ito, M. (2015) Inventory of the Heteroptera (Insecta: Hemiptera) in Komaba Campus of the University of Tokyo, a highly urbanized area in Japan. Biodiversity Data Journal, 3, e 4981. https: // doi. org / 10.3897 / BDJ. 3. e 4981","Miyazaki, A., Nishida, M., Uesugi, R., Yamada, U., Yasunaga, T., Serrano Leon, S., Kawashita, S. & Nagashima, T. (2020) Utilizing a synthetic diet containing a fermented milk beverage for rearing terrestrial heteropterans, with new distributional records of four species in Nagasaki Prefecture, Japan. Rostria, 64, 63 - 69. [in Japanese with English summary]","Yasunaga, T. (2001 d) Family Miridae Hahn, plant bugs. In: Yasunaga, T., Takai, M. & Kawasawa, T. (Eds.), A Field Guide to Japanese Bugs II. Zenkoku Noson Kyoiku Kyokai, Publishing Co. Ltd., Tokyo, pp. 2 - 96 + 111 - 351. [in Japanese]"]}

Psallus Fieber, 1858 Discussion. The phyline genus Psallus was established by Fieber (1858) [type species: P. haematodes (Gmelin, 1790), a Euro-Siberian element] and is known to comprise approximately 130 species in the Holarctic Region (9 Nearctic species were introduced from Europe, cf. Wheeler & Henry, 1992; Schuh, 2002 ��� 2013; Wyniger, 2004). As pointed out by several authors (e.g., Yasunaga & Vinokurov; Wyniger, 2004; Yasunaga, 2010), the monophyly of Psallus cannot be elucidated by its current context, sometimes being termed a ���garbage group���. It is usually difficult to distinguish species of Psallus from those exhibiting similar appearance in some other genera by the external characters alone. Six valid subgenera have been proposed for classification of Psallus (cf. Aukema, 2018): Apocremnus Fieber, 1858, Calopsallus Yasunaga and Vinokurov, 2000, Hylopsallus Wagner, 1952, Mesopsallus Wagner, 1970, Phylidea Reuter, 1899 and Pityopsallus Wagner, in addition to the nominotypical subenus. Some of these subgenera may require generic status, like as Pityopsallus that is treated as a full genus in the present work. Although Wyniger (2004) did not use any subgeneric rank for her classification of Psallus, the subgenera Apocremnus, Calopsallus, Hylopsallus Phylidea and Psallus s. str. are considered to constitute monophyletic groups, on the basis of available external morphology and male genitalic structures (cf. Duwal et al., 2012). However, definitive treatments of the monophyly and relationships are beyond the scope of this study, and eventually a comprehensive phylogenetic analysis and subsequent revision are required to redefine the relevant taxa. In Japan, 19 species of Psallus were previously recorded (Yasunaga & Vinokurov, 2000; Yasunaga, 2001d). Subsequent efforts by the author and colleagues have recognized an undescribed species mentioned below, belonging to the subgenus Hylopsallus. The ���true��� members of Psallus s. str. (the type species P. haematodes and its close relatives) share the following characters: thoracic pleura without silvery seta (except for P. bagjonicus Josifov, 1983 and P. buddha Yasunaga, 2010 with relatively denser silvery setae); blunt-tipped phallotheca, flattened, medially twisted vesica with apically situated, small-sized secondary gonopore and an elongate apical blade (cf. Fig. 14H) that is usually notched laterally (Yasunaga & Vinokurov, 2000); posterior wall with noticeable spinulate scalelike microstructures on (undifferentiated) interramal lobe and narrow posterior margin sclerite (Fig. 25M���O); and rounded, smaller sclerotized rings (cf. Yasunaga, 2010, Fig. 23). On the other hand, species included in Hylopsallus have the silvery setae on the thoracic pleura (Figs. 13C, F; 29A, D, G, L), sharply pointed phallotheca (e.g., Fig. 31B, E), stout, relatively tubular vesica with 3 distinct processes (Fig. 31F, I, O���M) and thick-rimmed, elongateovoid secondary gonopore (e.g., Figs. 14A���B), relatively smooth interramal lobe (e.g., Fig. 32C���D), and wide sclerotized rings that are often asymmetrical to each other (cf. Fig. 15C, E; Wyniger, 2004). Psallus castaneae Josifov, 1983 that has been placed in the subgenus Phylidea, is herein regarded as a member of Hylopsallus, because of sharing the above-mentioned characters. Accordingly, five species of Psallus (Hylopsallus) are now recognized in Japan: P. (H.) aterrimus Yasunaga & Vinokurov, 2000; P. (H.) castaneae Josifov, 1983; P. (H.) edoensis Yasunaga & Vinokurov, 2020; P. (H.) ramae n. sp.; and P. (H.) tonnaichanus Muramoto, 1973. The following key will distinguish Japanese members of the subgenus Hylopsallus including a new species as well as a superficially very similar congener, P. bagjonicus Josifov, 1983 (for identification of all other Japanese Psallus species, see Yasunaga & Vinokurov, 2000). Key to Japanese species of Hylopsallus and Psallus bagjonicus 1. Antennal segment I almost entirely fuscous................................................................ 2 ��� Antennal segment I entirely yellow or pale brown, or its base narrowly infuscate................................... 3 2. Metafemur uniformly dark brown; male pygophore (genital segment) only with simple setae, lacking silvery seta (Fig. 29B)........................................................................................ P. (H.) aterrimus ��� Ventral surface of metafemur chestnut or reddish brown, with large, fuscous spots; pygophore with silvery setae (Fig. 29H)........................................................................................ P. (P.) bagjonicus 3. Base of antennal segment I usually darkened; protibia with dark spots at bases of spines; male genital segment with distinct ventral mesal keel from middle part to apex (Fig. 30D)....................................................... 4 ��� Antennal segment I almost entirely pale, or its extreme base faintly obscured; bases of protibial spines lacking dark spots; ventral mesal keel on male genital segment reduced, forming a narrow carina at apical 1/2���2/3 of segment (Fig. 29M)..... 5 4. Width of pronotum greater than length of metafemur in female; metatibial dark spots smaller, separated from each other; setae on male genital segment not aggregated or clustered at left-dorsolateral base (Fig. 30D���E).............. P. (H.) castaneae ��� Width of pronotum slightly less than length of metafemur in female; metatibial dark spots often enlarged (some are fused as in Fig. 13D���E); setae on male genital segment somewhat aggregated at left-lateral base (Fig. 30J)........ P. (H.) ramae n. sp. 5. Hemelytra blackish brown; male genital segment with a pair of clustered stiff setae (Fig. 30N���O)....... P. (H.) tonnaichanus ��� Hemelytra chestnut brown; setae on male genital segment not aggregated (Fig. 31D); spots at bases of metatibial spines small, obscure................................................................................. P. (H.) edoensis, Published as part of Yasunaga, Tomohide, 2022, The plant bug subfamily Phylinae in Japan, with key to genera and descriptions of eight new species (Hemiptera: Heteroptera: Miridae), pp. 1-52 in Zootaxa 5094 (1) on pages 28-29, DOI: 10.11646/zootaxa.5094.1.1, http://zenodo.org/record/5964735, {"references":["Schuh, R. T. (2002 - 2013) On-line Systematic Catalog of Plant Bugs (Insecta: Heteroptera: Miridae). Available from: http: // research. amnh. org / pbi / catalog / (accessed 1 December 2021)","Schuh, R. T. & Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): Arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72.","Wyniger, D. (2004) Taxonomy and phylogeny of the Central European bug genus Psallus (Hemiptera, Miridae) and faunistics of the terrestrial Heteroptera of Basel and surroundings (Hemiptera). Part I. PhD-thesis, Basel, 56 pp., 39 figs. https: // doi. org / 10.5451 / unibas- 003214502","Yasunaga, T. (2010) Plant bugs of the tribe Phylini in Thailand (Heteroptera: Miridae: Phylinae), with descriptions of six new species from additional areas in tropical and subtropical Asia. Entomologica Americana, 116, 50 - 92. https: // doi. org / 10.1664 / 10 - RA- 006.1","Aukema, B. (2018) Catalogue of the Palaearctic Heteroptera (searchable database). Available from: https: // catpalhet. linnaeus. naturalis. nl / (accessed 5 December 2021)","Yasunaga, T. & Vinokurov, N. N. (2000) The phyline plant bug genus Psallus Fieber in Japan (Heteroptera: Miridae: Phylinae). Entomological Science, 3, 653 - 668.","Duwal, R. K., Yasunaga, T., Jung, S. & Lee, S. H. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. https: // doi. org / 10.14411 / eje. 2012.074","Yasunaga, T. (2001 d) Family Miridae Hahn, plant bugs. In: Yasunaga, T., Takai, M. & Kawasawa, T. (Eds.), A Field Guide to Japanese Bugs II. Zenkoku Noson Kyoiku Kyokai, Publishing Co. Ltd., Tokyo, pp. 2 - 96 + 111 - 351. [in Japanese]"]}

Psallus (Psallus) varians (Herrich-Schäffer, 1841) Capsus varians Herrich-Schäffer, 1841: 45. NEW DATA. — Valdieri, Terme di Valdieri, La Valletta, 1550 m, 44.19083°N, 7.26778°E (WGS84), 17.VII.2014, MN, MD leg., 3 specimens, MZUF. — Vernante, loc. Palanfré, 1580 m, 44.19503°N, 7.49107°E (WGS84), 03.VII.2014, MN, MD leg., 1 specimen, MZUF. CHOROTYPE. — European., Published as part of Norbiato, Margherita, Dioli, Paride & Cianferoni, Fabio, 2019, Annotated checklist of the Hemiptera Heteroptera of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 269-312 in Zoosystema 41 (15) on pages 287-288, DOI: 10.5252/zoosystema2019v41a15, http://zenodo.org/record/3722483

Psallus (Mesopsallus) ambiguus (Fallén, 1807) Lygaeus ambiguus Fallén, 1807: 89. LITERATURE. — Mancini (1963 [sub Psallus (Apocremnus) ambiguus]): Colle di Tenda. NEW DATA. — Vernante, loc. Palanfré, 1580 m, 44.19503°N, 7.49107°E (WGS84), 03.VII.2014, MN, MD leg., 1 specimen, MZUF. CHOROTYPE. — European., Published as part of Norbiato, Margherita, Dioli, Paride & Cianferoni, Fabio, 2019, Annotated checklist of the Hemiptera Heteroptera of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 269-312 in Zoosystema 41 (15) on page 287, DOI: 10.5252/zoosystema2019v41a15, http://zenodo.org/record/3722483, {"references":["FALLEN C. F. 1807. - Monographia Cimicum Sueciae. C. G. Proft, Hafniae, 123 p.","MANCINI C. 1963. - Res Ligusticae CXXXIII. Emitteri Eterotteri della Liguria. Annali del Museo Civico di Storia Naturale, Genova 74: 30 - 121."]}

The genus Psallus Fieber, 1858 is represented in the Palaearctic Region by more than 120 species, and is regarded as a non-monophyletic taxon. Species in this genus are usually small; their body length not exceeds 6.0 mm, and never are green (a body colour is from black to brown, red and yellow). They are zoophytophagous, and most of them overwinter in the egg stage and are univoltine. 23 species of the genus have been reported from Poland so far. This note presents the second localities for two Psallus species, i.e., Psallus albicinctus and P. pseudoplatani in Poland, both from the Western Beskid Mts.

The genus Psallus Fieber, 1858 is represented in the Palaearctic Region by more than 120 species, and is regarded as a non-monophyletic taxon. Species in this genus are usually small; their body length not exceeds 6.0 mm, and never are green (a body colour is from black to brown, red and yellow). They are zoophytophagous, and most of them overwinter in the egg stage and are univoltine. 23 species of the genus have been reported from Poland so far. This note presents the second localities for two Psallus species, i.e., Psallus albicinctus and P. pseudoplatani in Poland, both from the Western Beskid Mts.

Psallus (Psallus) thomashenryisp. n.(Hemiptera: Heteroptera: Miridae: Phylinae: Phylini: Phylina) is described from southern Anatolia, Turkey. Illustrations of the dorsal habitus and male genitalia are provided. Its habitus is similar to other uniformly orange species of the subgenus Psallus Fieber, 1858, particularlyP.asthenicusSeidenstücker, 1966 from which it can be easily distinguished by the combination of extremely small size (2.3 mm in both sexes) and different morphology of the vesica. Psallus (Psallus) lucanicus Wagner, 1968 is recorded for the first time from Turkey. Psallus (Psallus) aurora (Mulsant & Rey, 1852) is removed from the list of Turkish fauna based on a reevaluation of the voucher specimen. An updated checklist of the species ofPsallusknown to occur in Turkey is provided. The relevance of Anatolia and the Syro-anatolian-transcaucasian region in the Palearctic distribution ofPsallusis discussed. The westernmost record of another mirid,PlagiognathusmarivanensisLinnavuori, 2010, is provided.

Cranocapsus turcicus Kıyak, 1990 TL: K������ksu K��y�� (G��ksun, Kahramanmaraş); 38��07'18"N, 36��31'48"E. DT: Kahramanmaraş (Kıyak, 1990). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 458, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Kiyak, S. (1990) Cranocapsus turcicus (sp. nov.), Yeni Bir Heteropter Turunun Tanimi (Insecta, Hemiptera, Miridae), Miscellaneous Papers, 5, 1 ‾ 3"]}

Psallus karakardes Seidenst��cker, 1958 TL: Akşehir (Konya); 38��21'40"N, 31��24'12"E. DT: Afyonkarahisar, Konya (Seidenst��cker, 1958; Kothe et al., 2004; ��nder et al., 2006). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Seidenstucker, G. (1958) Heteroptera aus Anatolien II. Revue De La Faculte Des Sciences De L' Universite D'Istanbul, 23, 119 - 129.","Kothe, T., Blazi, G. & Schonitzer, K. (2004) Katalog der Wanzen-Typen von Gustav Seidenstucker (1912 ‾ 1989) in der Zoologischen Staatssammlung Munchen (Heteroptera). 61. Bericht der Naturforschonenden Gesellschaft Augsburg, 2004, 1 ‾ 90.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus turcicus Wagner, 1971 TL: Kızılcahamam (Ankara); 40��28'02"N, 32��38'35"E. DT: Ankara, Mersin (Aukema, 1999; Kerzhner & Josifov, 1999; ��nder et al., 2006). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Wagner, E. (1971) Uber 2 neue Psallus - Arten (Hemiptera, Heteroptera, Miridae). Reichenbachia, 14 (3), 19 - 22.","Aukema, B. (1999) Type specimens of some Palaearctic Miridae (Heteroptera), l ‾ 32. B. Aukema, Renkum, The Netherlands. Available from:","Kerzhner, I. M. & Josifov, M. (1999) Family Miridae. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. The Netherlands Entomological Society, Amsterdam, pp. 1 - 577.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus lentigo Seidenst��cker, 1972 TL: Sultandağı, Tekkedere (Akşehir, Konya); 38��21'40"N, 31��24'12"E. DT: Afyonkarahisar, Bolu, Konya (Seidenst��cker, 1972b; Kothe et al., 2004; ��nder et al., 2006)., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Seidenstucker, G. (1972 b) Psallus lentigo n. sp. (Heteroptera, Miridae). Notulae Entomologicae, 52, 57 - 64.","Kothe, T., Blazi, G. & Schonitzer, K. (2004) Katalog der Wanzen-Typen von Gustav Seidenstucker (1912 ‾ 1989) in der Zoologischen Staatssammlung Munchen (Heteroptera). 61. Bericht der Naturforschonenden Gesellschaft Augsburg, 2004, 1 ‾ 90.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus cerridis Wagner, 1971 TL: D��zce; 40��50'59"N, 31��08'05"E. DT: D��zce (Wagner, 1971; Aukema, 1999). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on pages 465-466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Wagner, E. (1971) Uber 2 neue Psallus - Arten (Hemiptera, Heteroptera, Miridae). Reichenbachia, 14 (3), 19 - 22.","Aukema, B. (1999) Type specimens of some Palaearctic Miridae (Heteroptera), l ‾ 32. B. Aukema, Renkum, The Netherlands. Available from:"]}

Psallus apoplecticus Seidenst��cker, 1966 TL: Yılanlıdağ ( Kayseri); 38��46'04"N, 35��42'04"E. DT: Kayseri (Seidenst��cker, 1966a; Kothe et al., 2004; ��nder et al., 2006). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 465, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Seidenstucker, G. (1966 a) Neue Psallus - Arten aus der Turkei (Heteroptera, Miridae). Reichenbachia, 6, 291 ‾ 302.","Kothe, T., Blazi, G. & Schonitzer, K. (2004) Katalog der Wanzen-Typen von Gustav Seidenstucker (1912 ‾ 1989) in der Zoologischen Staatssammlung Munchen (Heteroptera). 61. Bericht der Naturforschonenden Gesellschaft Augsburg, 2004, 1 ‾ 90.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus quercicola (Reuter, 1904) Sthenarus quercicola Reuter, 1904 TL: Hermos River [= Gediz Nehri] (Menemen, İzmir); 38��37'35"N, 27��4'18"E. DT: İzmir, Karab��k, Sinop (Hoberlandt, 1956; ��nder et al., 2006). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Hoberlandt, L. (1956) Results of the zoological scientific expedition of the National Museum in Praha to Turkey: 18. Hemiptera IV. Terrestrial Hemiptera-Heteroptera of Turkey. Acta Entomologica Musei Nationalis Pragae, 3 (Supplement), 1 - 264.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus oenderi Wagner, 1976 TL: Kovada G��l�� (Eğirdir, Isparta); 37��52'10"N, 30��50'01"E. DT: Isparta (Wagner, 1976; Aukema, 1999; ��nder et al., 2006). Note: This species is endemic toAnatolica., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Wagner, E. (1976) Uber einige neue Miridae aus der Turkei (Hemiptera, Heteroptera). Reichenbachia, 16, 121 - 128.","Aukema, B. (1999) Type specimens of some Palaearctic Miridae (Heteroptera), l ‾ 32. B. Aukema, Renkum, The Netherlands. Available from:","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus brachycerus Reuter, 1904 TL: Hermos River [= Gediz Nehri] (Manisa); 38��36'25"N, 27��37'27"E. DT: Antalya, İzmir, Manisa (Hoberlandt, 1956)., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 465, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Hoberlandt, L. (1956) Results of the zoological scientific expedition of the National Museum in Praha to Turkey: 18. Hemiptera IV. Terrestrial Hemiptera-Heteroptera of Turkey. Acta Entomologica Musei Nationalis Pragae, 3 (Supplement), 1 - 264."]}

Cymus turcicus Matocq, 2000 TL: Beyşehir (Konya); 37��41'07"N, 31��44'34"E. DT: Konya (Peric��rt, 2001). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 469, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Pericart, J. (2001) Family Lygaeidae Schilling, 1829 - Seed-bugs. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. The Netherlands Entomological Society, Amsterdam, pp. 35 - 220."]}

Psallus rubinicterus Seidenst��cker, 1966 TL: Central Taurus, ��iftehan (Ulukışla, Niğde); 37��30'48"N, 34��48'07"E. DT: Niğde (Seidenst��cker, 1966a; Kothe et al., 2004; ��nder et al, 2006). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 466, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Seidenstucker, G. (1966 a) Neue Psallus - Arten aus der Turkei (Heteroptera, Miridae). Reichenbachia, 6, 291 ‾ 302.","Kothe, T., Blazi, G. & Schonitzer, K. (2004) Katalog der Wanzen-Typen von Gustav Seidenstucker (1912 ‾ 1989) in der Zoologischen Staatssammlung Munchen (Heteroptera). 61. Bericht der Naturforschonenden Gesellschaft Augsburg, 2004, 1 ‾ 90.","Onder, F., Karsavuran, Y., Tezcan, S. & Fent, M. (2006) Turkiye Heteroptera (Insecta) Katalo g u. Meta Basim Matbaacilik Hizmetleri, Izmir, 164 pp. [in Turkish with English summary]"]}

Psallus anatolicus Wagner, 1963 TL: Ayancik [=Ayancık], Sinop); 41��56'40"N, 34��35'06"E. DT: Sinop (Wagner, 1963d; Kerzhner & Josifov, 1999; Aukema, 1999). Note: This species is endemic to Anatolia., Published as part of Dursun, Ahmet & Fent, Meral, 2017, Type Localities of Heteroptera (Insecta: Hemiptera) from Turkey, pp. 451-494 in Zootaxa 4227 (4) on page 465, DOI: 10.11646/zootaxa.4227.4.1, http://zenodo.org/record/293602, {"references":["Wagner, E. (1963 d) Psallus anatolicus nov. spec. a new species of Miridae from Turkey (Hemiptera, Heteroptera). Revue de la Faculte des sciences de l'Universite d'Istanbul (B), 26, 201 - 203.","Kerzhner, I. M. & Josifov, M. (1999) Family Miridae. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. The Netherlands Entomological Society, Amsterdam, pp. 1 - 577.","Aukema, B. (1999) Type specimens of some Palaearctic Miridae (Heteroptera), l ‾ 32. B. Aukema, Renkum, The Netherlands. Available from:"]}

Subgenus Psallus Fieber, 1858 sensu stricto Psallus Fieber, 1958: 321 (subgen. nov.) 69. Psallus (P.) amoenus Josifov, 1983 (Figure plate 6) Psallus (Psallus) amoenus Josifov, 1983 b: 208 (sp. nov.); (1992 b): 116 (list); Kerzhner (1988 b): 844 (key, fig.); Schuh (1995): 400 (cat.); Kerzhner & Josifov (1999): 411 (cat.); Vinokurov et al. (2010): 146 (cat.); Duwal et al. (2012): 630 (diag., key, fig.). Distribution. Asia: North Korea (Josifov 1983 b), Russia (Far East) (Kerzhner 1988 b). Host. Quercus dentate Thunb. (Fagaceae) (Josifov 1983 b). 70. Psallus (P.) bagjonicus Josifov, 1983 (Figure plate 6) Psallus (Psallus) bagjonicus Josifov, 1983 b: 205 (sp. nov.); (1992 b): 116 (list); Schuh (1995): 401 (cat.); Kerzhner & Josifov (1999): 412 (cat.); Yasunaga & Vinokurov (2000): 659 (key, figs.); Yasunaga (2001 a): 176 (diag., figs.); Duwal et al. (2012): 630 (diag., key, fig.); Aukema et al. (2013): 326 (cat.). Distribution. Asia: North Korea (Josifov 1983 b), South Korea (GG) (Duwal et al. 2012), Japan (Yasunaga & Vinokurov 2000). Host. Quercus aliena Blume (Fagaceae) (Josifov 1983 b), Q. serrata Murray., Q. acutissima Carruth. (Yasunaga & Vinokurov 2000). 71. Psallus (P.) cheongtaensis Duwal, Yasunaga, Jung & Lee, 2012 (Figure plate 7) Psallus (Psallus) cheongtaensis Duwal, Yasunaga, Jung & Lee, 2012: 630 (sp. nov.). Specimens examined. South Korea: GW: 28 (♂, ♀), Inje-gun, Yongdae-ri, Mt. Maebong, Yongdaeri Natural Recreation Center, 6.VI. 2014, on Tilia sp., R.K. Duwal (SNU). Distribution. Asia: South Korea (GW) (Duwal et al. 2012). Host. Tilia sp. (Malvaceae) (new record). 72. Psallus (P.) koreanus Josifov, 1983 (Figure plate 7) Psallus (Psallus) koreanus Josifov, 1983 b: 209 (sp. nov.); Kerzhner (1988 b): 847 (key); Josifov (1992 b): 116 (list); Schuh (1995): 408 (cat.); Kerzhner & Josifov (1999): 415 (cat.); Vinokurov et al. (2010): 146 (cat.); Duwal et al. (2012): 631 (diag., key). Distribution. Asia: North Korea (Josifov 1983 b), Russia (Far East) (Kerzhner 1988 b). Host. Crataegus sp., Prunus sp., Sorbus amurensis Koehne (Rosaceae) (Josifov 1983 b). 73. Psallus (P.) sanguinarius Kerzhner & Josifov, 1999 (Figure plate 7) Psallus (Psallus) sanguinolentus Josifov, 1983 b: 209 (junior primary homonym of Psallus graminicola f. sanguinolenta Stichel, 1956); (1992 b): 116 (list); Schuh (1995): 415 (cat.) Psallus (Psallus) sanguinarius: Kerzhner & Josifov (1999): 417 (new name, cat.); Duwal et al. (2012): 631 (diag., key); Duwal & Lee (2015): 593 (diag., fig.). Distribution. Asia: North Korea (Josifov 1983 b), South Korea (GW) (Duwal & Lee 2015). Host. Sorbus amurensis Koehne (Rosaceae), Picea sp. (Pinaceae) (Josifov 1983 b)., Published as part of Duwal, Ram Keshari, Jung, Sunghoon, Yasunaga, Tomohide & Lee, Seunghwan, 2016, Annotated catalogue of the Phylinae (Heteroptera: Miridae) from the Korean Peninsula, pp. 101-134 in Zootaxa 4067 (2) on page 118, DOI: 10.11646/zootaxa.4067.2.1, http://zenodo.org/record/270595, {"references":["Fieber, F. X. (1858) Criterien zur generischen Theilung der Phytocoriden (Capsini aut.). Wiener Entomologische Monatschrift, 2, 289 - 327, 329 - 347, 388.","Josifov, M. (1983 b) Neue Psallus. Arten aus Nord-Korea (Heteroptera, Miridae). Reichenbachia, 21, 197 - 211.","Kerzhner, I. M. (1988 b) Family Miridae. In: Lehr, P. A. (Ed.), Keys to the insects of the Far East of the USSR, 2, pp. 778 - 857. [Nauka, Leningrad, in Russian]","Kerzhner, I. M. & Josifov, M. (1999) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. The Netherlands Entomological Society, Amsterdam, xiv + 577 pp.","Vinokurov, N. N., Kanjukova, E. V. & Golub, V. B. (2010) The catalogue of bugs (Heteroptera) of Aisan part of Russia. Nauka, Novosibirsk, 320 pp.","Duwal, R. K., Yasunaga, T., Jung, S. & Lee, S. H. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. http: // dx. doi. org / 10.14411 / eje. 2012.074","Yasunaga, T. & Vinokurov, N. N. (2000) The phyline plant bug genus Psallus Fieber in Japan (Heteroptera: Miridae: Phylinae). Entomological Science, 3, 653 - 668.","Yasunaga, T. (2001 a) Family Miridae Hahn, plant bugs. In: Yasunaga, T., Takai, M. & Kawasawa, T. (Eds.), A Field Guide to Japanese Bugs II. Zenkoku Noson Kyoiku Kyokai. Publishing Co. Ltd., Tokyo, pp. 1 - 96, 111 - 351. [in Japanese]","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. The Netherlands Entomological Society, Amsterdam, xxiv + 629 pp.","Josifov, M. (1992 b) Contribution to a Knowledge of the family Miridae of North Korea (Heteroptera). Insecta Koreana, 9, 115 - 128.","Stichel, W. (1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa. (Hemiptera-Heteroptera Europae). 2. Stichel, Berlin-Hermsdorf, 312 pp. [pp. 169 - 480]","Duwal, R. K. & Lee, S. H. (2015) Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae). Zootaxa, 3926 (4), 585 - 594. http: // dx. doi. org / 10.11646 / zootaxa. 3926.4.8"]}

45. Psallus (A.) aethiops (Zetterstedt, 1838) Phytocoris aethiops Zetterstedt, 1838: 274 (sp. nov.) Psallus (Apocremnus) aethiops: Josifov (1992 b): 124 (report); (Schuh) 1995: 399 (cat.); Kerzhner & Josifov (1999): 400 (cat.); Schwartz & Kelton (1990): 945 (records); Vinokurov et al. (2010): 142 (cat.); Duwal et al. (2012): 607 (diag., key, fig.); Aukema et al. (2013): 322 (cat.). Distribution. European continent: Finland, Norway, Russia (Central, Northern), Sweden; Asia: North Korea, Russia (East Siberia, Far East, West Siberia); North America (Kerzhner 1988 b, Josifov 1992 b, Kerzhner & Josifov 1999). Host. Salix sp. (Salicaceae) (Kerzhner 1988 b, Schwartz & Kelton 1990, Duwal et al. 2012). 46. Psallus (A.) ater Josifov, 1983 (Figure plate 4) Psallus (Apocremnus) ater Josifov, 1983 b: 198 (sp. nov.); (1992): 116 (list); Schuh (1995): 401 (cat.); Kerzhner & Josifov (1999): 401 (cat.); Duwal et al. (2012): 608 (diag., key, fig.). Distribution. Asia: North Korea (Josifov 1983), South Korea (GG, GW, JN) (Duwal et al. 2012), China (Southwest) (Li & Zheng 1991 a). Host. Crataegus sp., Prunus sp. (Rosaceae) (Josifov 1983 b), Quercus sp. (Fagaceae) (Duwal et al. 2012). 47. Psallus (A.) atratus Josifov, 1983 (Figure plate 4) Psallus (Apocremnus) atratus Josifov, 1983 b: 197 (sp. nov.); (1992 b): 124 (report); Schuh (1995): 401 (cat.); Kerzhner & Josifov (1999): 401 (cat.); Vinokurov et al. (2010): 143 (cat.); Duwal et al. (2012): 611 (diag., key, fig.). Distribution. Asia: North Korea (Josifov 1983 b), South Korea (GG) (Duwal et al. 2012), Russia (Far East) (Kerzhner 1988 b). Host. Crataegus sp. (Rosaceae) (Josifov 1983 b), Pyrus ussuriensis Maxim. (Kerzhner 1988 b), Prunus sp. (Duwal et al. 2012). 48. Psallus (A.) betuleti (Fall��n, 1826) (Figure plate 4) Cimex cruentus M��ller, 1776: 108 (sp. nov.) Phytocoris betuleti Fall��n 1826: 15 (sp. nov.) Psallus (Apocremnus) betuleti: Zaitzeva (1968): 866 (descr., fig.); Josifov (1992 b): 124 (report); Wheeler & Henry (1992): 185 (dist., host); Schwartz & Kelton (1990): 946 (diag., descr.); Schuh (1995): 401 (cat.); Kerzhner & Josifov (1999): 401 (cat.); Rieger & Rabitsch (2006): 163 (diag., dist., fig.); Vinokurov et al. (2010): 143 (cat.); Duwal et al. (2012): 612 (diag., key, fig.). Distribution. European continent (Western, Central, Eastern); Asia: North Korea, China (North), Russia (East Siberia, Far East, West Siberia); North America (Josifov 1992 b, Kerzhner & Josifov 1999). Host. Betula spp. (Betulaceae) (Henry & Wheeler 1979), Alnus spp. (Kerzhner 1978), Epilobium spp. (Onagraceae), Rhododendron sp. (Ericaceae) (Schwartz & Kelton 1990). 49. Psallus (A.) michaili Kerzhner & Schuh, 1995 (Figure plate 4) Psallus Apocremnus niger Josifov, 1992 a: 113 (sp. nov.) Psallus (Apocremnus) michaili: Josifov (1992 b): 116 (list); Kerzhner & Schuh (1995): 4 (new name); Schuh (1995): 409 (cat.); Kerzhner & Josifov (1999): 402 (cat.); Duwal et al. (2012): 613 (diag., key, fig.). Distribution. Asia: North Korea (Josifov 1992 b), South Korea (GG) (Duwal et al. 2012). Host. Acer ginnala Maxim. (Aceraceae), (Josifov 1992 a), and Quercus sp. (Fagaceae) (Duwal et al. 2012)., Published as part of Duwal, Ram Keshari, Jung, Sunghoon, Yasunaga, Tomohide & Lee, Seunghwan, 2016, Annotated catalogue of the Phylinae (Heteroptera: Miridae) from the Korean Peninsula, pp. 101-134 in Zootaxa 4067 (2) on page 114, DOI: 10.11646/zootaxa.4067.2.1, http://zenodo.org/record/270595, {"references":["Zetterstedt, J. W. (1838 - 1840) Insecta Lapponica descripta. Voss, Lipsiae, vi col. + 1140 pp. (1838, col. 1 - 868).","Josifov, M. (1992 b) Contribution to a Knowledge of the family Miridae of North Korea (Heteroptera). Insecta Koreana, 9, 115 - 128.","Kerzhner, I. M. & Josifov, M. (1999) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. The Netherlands Entomological Society, Amsterdam, xiv + 577 pp.","Schwartz, M. D. & Kelton, L. A. (1990) Psallus salicicola, a new species, with additional records of recently discovered Palaearctic Psallus Fieber from Canada (Heteroptera: Miridae: Phylinae). Canadian Entomologist, 122, 941 - 947. http: // dx. doi. org / 10.4039 / Ent 122941 - 9","Vinokurov, N. N., Kanjukova, E. V. & Golub, V. B. (2010) The catalogue of bugs (Heteroptera) of Aisan part of Russia. Nauka, Novosibirsk, 320 pp.","Duwal, R. K., Yasunaga, T., Jung, S. & Lee, S. H. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. http: // dx. doi. org / 10.14411 / eje. 2012.074","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. The Netherlands Entomological Society, Amsterdam, xxiv + 629 pp.","Kerzhner, I. M. (1988 b) Family Miridae. In: Lehr, P. A. (Ed.), Keys to the insects of the Far East of the USSR, 2, pp. 778 - 857. [Nauka, Leningrad, in Russian]","Josifov, M. (1983 b) Neue Psallus. Arten aus Nord-Korea (Heteroptera, Miridae). Reichenbachia, 21, 197 - 211.","Li, H. Y. & Zheng, L. Y. (1991 a) A preliminary report on Psallus Fieber (Heteroptera: Miridae) from China. Acta Scienntiarum Naturalium Universitatis Nankaiensis, 1991 (1), 1 - 11. [in Chinese, English summary]","Fallen, C. F. (1826) Supplementum Cimicidum Sueciae. Berling, Londini Gothorum [= Lund], 16 pp.","Muller, O. F. (1776) Zoologiae Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina et synonyma imprimis popularium. Hallager, Hafniae, xxxii + 282 pp.","Zaitzeva, I. F. (1968) A review of the genus Psallus Fieb. (Heteroptera) from the Caucasus. Entomologicheskoe Obozrenie, 47, 865 - 877. [in Russian]","Wheeler, A. G. Jr. & Henry, T. J. (1992) A synthesis of the Holarctic Miridae (Heteroptera): Distribution, biology and origin, with emphasis on North America. Entomological Society of America, Lanham, Maryland, V + 282 pp.","Reiger, C. & Rabitsch, W. (2006) Taxonomy and distribution of Psallus betuleti (Fallen) and P. montanus Josifov stat. nov. (Heteroptera: Miridae). Tijdschrift voor Entomologie, 149, 161 - 166. http: // dx. doi. org / 10.1163 / 22119434 - 900000196","Henry, T. J. & Wheeler, A. G. Jr. (1979) Palaearctic Miridae in North America: records of newly discovered and little-known species (Hemiptera: Heteroptera). Proceedings of the Entomological Society of Washington, 81, 257 - 268.","Kerzhner, I. M. (1978) Bugs (Heteroptera) of Sakhalin and the Kuril Islands. Trudy Biologopochvennogo Instituta Dal'nevostochnoe Otdelenie Akademiya Nauk SSSR, New Series, 50, 31 - 57. [in Russian]","Josifov, M. (1992 a) Neue Miriden aus Korea (Insecta, Heteroptera). Reichenbachia, 29, 105 - 118."]}

Psallus buddha Yasunaga, 2010: 88. Holotype ♂, SERS, 14 �� 30 ��� 27 ���N, 101 �� 55 ��� 39 ���E, 410 m alt., 25 Feb 2009, T. Yasunaga (AMNH_PBI 00379298) (DOAT)., Published as part of Yasunaga, Tomohide, Yamada, Kazutaka, Morakote, Rut, Taekul, Charuwat & Duangthisan, Jomsurang, 2016, Transferred depository for twenty-seven holotypes of the plant bug and flower bug species recently described from Thailand (Hemiptera: Heteroptera: Miridae and Anthocoridae), pp. 444-446 in Zootaxa 4107 (3) on page 445, DOI: 10.11646/zootaxa.4107.3.13, http://zenodo.org/record/267318, {"references":["Yasunaga, T. (2010) Plant bugs of the tribe Phylini in Thailand (Heteroptera: Miridae: Phylinae), with descriptions of six new species from additional areas in tropical and subtropical Asia. Entomologica Americana, 116, 50 - 92. http: // dx. doi. org / 10.1664 / 10 - RA- 006.1"]}

There are twenty-six species of the genus Psallus Fieber, 1858 (Phylinae: Phylini) documented in the Korean Peninsula. Three new species are described: Psallus cheongtaensis sp. n., P. ernsti sp. n. and P. suwonanus sp. n., and four species are reported from Korea for the first time: Psallus cinnabarinus Kerzhner, 1979, P. flavescens Kerzhner, 1988, P. loginovae Kerzhner, 1988, and P. roseoguttatus Yasunaga & Vinokurov, 2000. The genitalic structures of females and males are described and figured for most treated species. A key is provided for identifying all the Korean species.

Psallus (Calopsallus) injensis Duwal sp. nov. Figures: 1 C, 2 A ���C, 4 D Diagnosis. Recognized by the pale yellow, elongated body with variously distributed orange or orange-red speckles on dorsum, scattered brown spots on head and pronotum, discontinuous transverse brown stripes on the head with pale longitudinal median line, and densely distributed brown spots on large part of metafemora arranged in rows towards the base (Fig. 1 C), C ���shaped endosoma with broad and complex apex, furnished with numerous spinules, and a tooth like spine on outer margin of apical process, and a curved finger like subapical process facing upward to the apex (Figs. 2 A). Description. Male. Body elongate. COLORATION (Fig. 1 C): Dorsum entirely pale yellow with variously distributed irregular orange (or orange-red) speckles. Head with scattered orange speckles and brown spots, and transverse brown stripes on head discontinuous at the center forming a pale longitudinal median line; tylus pale with yellowish-brown or reddish lateral margins; antennal segments pale except the sub-basal region of segment I with brown ring and a pair of small brown spots at bases of brown spine; labium entirely pale yellow with brown apex on segment IV. Pronotum pale with scattered orange speckles and brown spots, mesoscutum and scutellum with dense orange-red speckles except at the corners. Hemelytron pale, clavus arranged with sparse orange speckles; endocorium and exocorium with dense orange speckles, cuneus orange-yellow with base and outer margin white and inner margin with discontinuous red colored patches; membrane pale greyish and dull with dark spots at inner and outer margin of cells near the apical region of cuneus. Venter, greyish brown with pale yellow lateral margin of abdominal segments, ostiolar peritreme pale. All legs pale (or somewhat pale brownish), coxae pale with somewhat darker base, trochanters pale, metafemora with densely distributed irregular brown spots arranged in rows towards the base and extreme apex without any spots; tibiae pale with rows of small brown spots at bases of dark brown spines. Abdomen pale yellow. SURFACE AND VESTITURE: Dorsum furnished with simple dark brown semi-erect setae and moderately flattened sericeous appressed setae; head, pronotum, mesoscutum and scutellum polished. STRUCTURE: Antennal segment I half the length of tarsus, segment II subequal to basal width of the pronotum, segment III subequal to the width of head across eyes, and segment IV subequal to the width of vertex; interocular space wide; labium reaching apex of metacoxae. Thorax more or less trapezoid with postero-lateral margins semicurved. Hemelytron sub-parallel, somewhat fattened at the level of apex of scutellum and cuneal fracture distinct. GENITALIA (Figs. 2 A ���C, 4 D): Endosoma: C ���shaped; apex broad and complex with numerous spinules, outer margin of apical process with a tooth like spine, a curved finger-like process facing towards the apex; and secondary gonopore placed subapically between apical and sub-apical processes (Figs. 2 A, 4 D). Left paramere: Body short, anterior process slender and elongated, posterior thumb like process short and elongated (Fig. 2 B). Phallotheca: Broad, tapered at sub-apical region with blunt apex (Fig. 2 C). Female. Unknown. Etymology. Named for the type locality, Inje; an adjective. Distribution. Korea (South*). Measurements. (♂) (n= 1). Body length 4.08; length from apex of tylus to cuneal fracture 3.14, width of head across eyes 0.79; width of vertex 0.37, lengths of antennal segments 0.22, 1.14, 0.72, 0.35; mesal length of pronotum 0.59; basal width of pronotum 1.19; maximum width across hemelytron 1.53; and lengths of metafemora, tibiae and tarsus 1.24, 2.04, 0.45. Specimen examined. Holotype. ♂, South Korea, Gangwon-do, Inje-gun, Buk-myeon, Yongdae-ri, Mt. Maebong, Yongdae Natural Recreation Center, on light trap, 19.vi. 2013, R.K. Duwal (SNU). Remarks. This species is placed in the subgenus Calopsallus, based upon the overall appearance of its endosoma being similar to those of congeners, i.e. broad, complex apex bearing dense spinulus structures. The new species, Psallus injensis can be easily separated from other members by pattern of spots on metafemora. Male genital structure, endosoma is very similar to P. roseoguttatus but could be differentiated due to dentate spine of endosoma on outer lateral margin of apical process and upturned curved subapical process (Fig. 2 A). As a single male specimen was collected at a light trap, biology or host association is unknown., Published as part of Duwal, Ram Keshari & Lee, Seunghwan, 2015, Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae), pp. 585-594 in Zootaxa 3926 (4) on pages 588-590, DOI: 10.11646/zootaxa.3926.4.8, http://zenodo.org/record/232094

Psallus koreanus Josifov, 1983 Psallus koreanus Josifov, 1983 b: 209. Holotype: 1 ♂, Pagyon-san (Mt), 20 km N of Kaesong, North Korea (on label: Kesông, 20km NO, Bagâin, North Korea), 21–23.v. 1975, M. Josifov. Paratypes: 25 ♂, 10 ♀, same data as holotype., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 402, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Josifov, M. (1983 b) Neue Psallus - Arten aus Nord-Korea (KDVR) (Heteroptera, Miridae). Reichenbachia, 21 (35), 197 - 211."]}

Psallus atratus Josifov, 1983 Psallus atratus Josifov, 1983 b: 197. Holotype: 1 ♂, Pagyon-san (Mt) nr Pagyon falls, 27 km NE of Kaesong, North Korea (on label: Kesông, 20km NO, Bagâin, North Korea), 21–23.v. 1975, M. Josifov. Paratypes: 2 ♂ 1 ♀, same data as holotype., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 401, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Josifov, M. (1983 b) Neue Psallus - Arten aus Nord-Korea (KDVR) (Heteroptera, Miridae). Reichenbachia, 21 (35), 197 - 211."]}

Psallus (Phylidea) yongdaeri Duwal sp. nov. Figures: 1 F ���G, 2 D ���G, 4 B, 5 B Diagnosis. Recognized by the small and ovoid body, black or blackish brown dorsum, often dull with greyish posterior margin of vertex and brownish margin of exo- and endocorium, pale (or yellowish pale brown) antennae with fuscous ring at the base of segment I, fuscous or black metafemora without spots, pale pro- and meso-tibiae with large castaneous spots on basal half region and meta-tibiae with brown spots at bases of black spines (Fig. 1 F ���G), C-shaped endosoma with distinctly featured apical process (Fig. 2 D). Description. Male. Body small and oval. COLORATION (Fig. 1 F ���G): Head black except for greyish posterior margin of vertex; antennal segments pale with extreme base of segment I dark and furnished with a pair of brown spine like setae, and segment IV slightly darker; labium brown with darker base. Pronotum, meso-scutum and scutellum black except for brown lateral and posterior corners of the meso-scutum and the scutellum; hemelytron blackish brown on large posterior region (on dark brown specimen) or margin of endo- and exocorium dark brown (in black specimens), membrane dark grey. Venter black or dark brown, ostiolar peritreme black. Coxae and trochanters black; pro- and mesofemora darker with pale apices; metafemora black (in black specimens) or dark brown (in dark brown specimens) and without darker spots; tibiae pale, basal half of pro-and meso-tibiae with large castaneous spots at bases of black spines and entire metatibiae with large brown spots at bases of black spines. Abdomen black or dark brown in color. SURFACE AND VESTITURE: Head, pronotum, meso-scutum and scutellum dull (or shagreened) with uniformly distributed pale simple reclining setae and moderately flattened sericeous appressed setae; hemelytron somewhat polished, with uniformly distributed black semi-erect setae and sericeous appressed setae. STRUCTURE: Head width across eyes sub-equal to mesal length of pronotum, vertex width sub-equal to length of tarsus; antennal segment II comparatively short and thick; interocular space wide; labium reaching apex of metacoxae. Thorax slightly convex, with semi-circular postero-lateral margin. GENITALIA (Figs. 2 D ���G, 4 B): Endosoma: C ���shaped with several short or long lateral processes, membranous structure at base of apical processes furnished with spinules; and secondary gonopore subapical in position (Figs. 2 D, 4 B). Left paramere: Body short, anterior process slender and elongated, and posterior process short and blunt (Fig. 2 E). Right paramere: Simple leaf-like apically tapering with short outgrowth (Fig. 2 F). Phallotheca: Narrow, with hooked sub-apical process (Fig. 2 G). Female. Similar to color and texture as males, only having slightly larger body width. GENITALIA (Fig. 5 B): Sclerotized rings somewhat broad, anteriorly oval and posteriorly un-curved or flatten margin. Measurements. (♂/♀) (n= 5 / 4). Body length 2.98���3.20 / 3.06���3.23; length from apex of tylus to cuneal fracture 2.31���2.40 / 2.31���2.46, width of head across eyes 0.72���0.77 / 0.74���0.79; width of vertex 0.37���0.43 / 0.37���0.40, lengths of antennal segments 0.21���0.23, 0.83���0.90, 0.50���0.54, 0.32���0.38 / 0.20���0.23, 0.82���0.91, 0.46���0.54, 0.31���0.35; mesal length of pronotum 0.64���0.73 / 0.63���0.71; basal width of pronotum 1.24���1.34 / 1.30���1.33; maximum width across hemelytron 1.53���1.66 / 1.58���1.68; and lengths of metafemora, tibiae and tarsus 1.05���1.19, 1.71���1.80, 0.41���0.51 / 1.12���1.18, 1.65���1.74, 0.42���0.46. Etymology. Named for the type locality, Yongdae-ri; a noun in apposition. Distribution. Korea (South*). Specimen examined. Holotype. ♂, South Korea, Gangwon-do, Inje-gun, Buk-myeon, Yongdae-ri, Mt. Maebong, Yongdae Natural Recreation Center, on Ulmaceae, 20.vi. 2013, R.K. Duwal (SNU). Paratypes. South Korea: 5 ♂, 2 ♀, same data as for holotype; 1 ♀, same data as for holotype except collected on Artemisia sp. (Asteraceae); 1 ♂, 1 ♀, same collection site as for holotype, on light trap, 19.vi. 2013. Remarks. This new species is placed in the subgenus Phylidea, based upon the overall appearance of its endosoma being similar to those of congeners, i.e. unique serrated spinules on the lateral margin of the apical region. Psallus yongdaeri is morphologically close to Psallus castaneae in color and vestiture but can be easily separated by the distinct basal ring on antennal segment I, longer apical process of endosoma, and arrangements of lateral processes (Fig. 2 D). A few specimens of Psallus yongdaeri were collected on unidentified Ulmaceae tree during spring flowering season in South Korea. In addition, some specimens were collected in light trap and also on Artemisia sp. under the tree where siblings were collected. However, we are not sure of host plant, as we were unable to observe any nymphs during collection time., Published as part of Duwal, Ram Keshari & Lee, Seunghwan, 2015, Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae), pp. 585-594 in Zootaxa 3926 (4) on pages 591-593, DOI: 10.11646/zootaxa.3926.4.8, http://zenodo.org/record/232094

Psallus samdzijonicus Josifov, 1983 Psallus samdzijonicus Josifov, 1983 b: 200. Holotype: 1 ♂, Ryanggan Prov., Samdjiyon, North Korea (on label: Samdzijôn, Jangkangdo, North Korea), 13– 19.vii. 1974, M. Josifov. Paratypes: 8 ♂, 6 ♀, same data as holotype; 1 ♀, 1000m, Boseok-ri, Ryanggan-do, North Korea (on label: 1000m, Bosôk-ri, Jangkangdo, North Korea), 20.vii. 1974, M. Josifov., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 402, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Josifov, M. (1983 b) Neue Psallus - Arten aus Nord-Korea (KDVR) (Heteroptera, Miridae). Reichenbachia, 21 (35), 197 - 211."]}

Psallus castaneae Josifov, 1983 Psallus castaneae Josifov, 1983 b: 202. Holotype: 1 ♂, Daesong-san (Mt) nr Pyongyang, North Korea (on label: Phjôngjang, Tesông-san, North Korea), 29.v. 1975, M. Josifov. Paratypes: 17 ♂, 28 ♀, same data as holotype; 2 ♂, 1 ♀, Daesong-san (Mt) nr Pyongyang, North Korea (on label: Phjôngjang, Tesông-san, North Korea), 1.vi. 1975, M. Josifov; 2 ♂, Ryongak-san (Mt), 12 km W of Pyongyang,, North Korea (on label: Phjongjang, 12km westl, Rjongakk-san, North Korea), 30.v. 1975, M. Josifov., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 401, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Josifov, M. (1983 b) Neue Psallus - Arten aus Nord-Korea (KDVR) (Heteroptera, Miridae). Reichenbachia, 21 (35), 197 - 211."]}

Genus Psallus sensu lato Type species: Lygaeus sanguineus Fabricius, 1794, a synonym of Cimex haematodes Gmelin, 1790 (subsequent designation by Reuter, 1888: 412). Remarks. Species of Psallus are usually 2.9���5.5 mm in length, with oval or elongate body and various coloration from yellow, or orange, or red, pale or dark brown, or black, sometimes variously spotted or mottled, and rarely tinged with green (Fig. 1). This is one of the largest genera in tribe Phylini, members of which are grouped into eight different subgenera. The species are distributed in temperate or cold temperate regions of the Northern Hemisphere. Recently, twenty six species of Psallus are revised from the Korean Peninsula including new species and first reports (Duwal et al., 2012). Herein, we would like to add descriptions of additional species from the Korean Peninsula; Psallus (Calopsallus) injensis Duwal sp. nov., P. (Halopasllus) taehwana Duwal sp. nov., P. (Phylidea) yongdaeri Duwal sp. nov., Published as part of Duwal, Ram Keshari & Lee, Seunghwan, 2015, Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae), pp. 585-594 in Zootaxa 3926 (4) on pages 585-586, DOI: 10.11646/zootaxa.3926.4.8, http://zenodo.org/record/232094, {"references":["Duwal, R. K., Yasunaga, T., Jung, S. & Lee, S. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. http: // dx. doi. org / 10.14411 / eje. 2012.074"]}

Psallus ernesti Duwal & Lee, 2012 Psallus ernesti Duwal et al., 2012: 109. Paratypes: 2 ♀, Ryongak-san (Mt), Pyongyang,, North Korea (on label: Phjongjang, Rjonggang, Mt. Rjongak), 31.v. 1987, M. Josifov; 1 ♂, Pagyon-san (Mt), 20 km N of Kaesong, North Korea (on label: Bagjon, 20 km N Kaesong), 29.v. 1987, M. Josifov., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 401, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Duwal, R. K., Yasunaga, T., Jung, S. H. & Lee, S. H. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. http: // dx. doi. org / 10.14411 / eje. 2012.074"]}

Psallus (Psallus) sanguinarius Kerzhner and Josifov, 1999 Figures: 1 A ���B, 4 C Psallus (Psallus) sanguinolentus Josifov, 1983: 209 (junior primary homonym of Psallus graminicola f. sanguinolenta Stichel, 1956); Schuh, 1995: 415 (cat.) Psallus sanguinarius Kerzhner and Josifov, 1999: 417 (cat.); Duwal et al., 2012: 631 (diag.) Diagnosis. Recognized by small oval body, chestnut red (male) or reddish (female) dorsum, ocher (female) or dark head with pale margin on vertex (male) with entirely black tylus, and uniformly red cuneus, brownish (male) or reddish brown (female) metafemora with two rows of black spots. For a detail description see, Josifov, 1983 (where it is described as sanguinolentus). Measurements. (♂/♀) (n= 1 / 2). Body length 3.62 / 3.66���3.71; length from apex of tylus to cuneal fracture 2.49 / 2.55���2.67, width of head across eyes 0.77 / 0.76���0.77; width of vertex 0.41 / 0.41���0.46, lengths of antennal segments 0.23, 0.93, 0.66, 0.40 / 0.21���0.23, 1.03���1.05, 0.65���0.68, 0.38���0.41; mesal length of pronotum 0.65 / 0.61���0.66; basal width of pronotum 1.25 / 1.29���1.31; maximum width across hemelytron 1.62 / 1.67���1.68; and lengths of metafemora, tibiae and tarsus 1.14, 1.92, 0.55 / 1.30���1.32, 1.84���1.93, 0.53���0.54. Distribution. Korea (North, South*). Specimen examined. South Korea: 1 ♂, 2 ♀, Gyeonggi-do, Gapyeong-gun, Cheongpyeong-myeon, Samhoeri, Mt. Hwaya, 29.v. 2014, on Prunus sp. (Rosaceae), R.K. Duwal (SNU). Remarks. Psallus sanguinarius was previously reported in North Korea from Sorbus amurensis (Rosaceae) and Picea (Pinaceae) (Josifov, 1983). We were unable to observe any specimens during our previous study (Duwal et al. 2012). Fortunately, we could add few recently collected specimens of this species on Prunus (Rosaceae) from northern part of South Korea. The images of dorsal habitus, and male genitalia are given in figure plates, 1 A-B and 4 C respectively. (For details see, Josifov, 1983: 206, 209 ��� 210)., Published as part of Duwal, Ram Keshari & Lee, Seunghwan, 2015, Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae), pp. 585-594 in Zootaxa 3926 (4) on page 593, DOI: 10.11646/zootaxa.3926.4.8, http://zenodo.org/record/232094, {"references":["Kerzhner, I. M. & Josifov, M. (1999) Cimicomorpha II, Miridae. IN Aukema, B. and Reiger CH. (Eds.), Catalogue of the Heteroptera of the Palearctic Region. The Netherlands Entomological Society, Amsterdam, i-xiv, pp. 1 - 577.","Josifov, M. (1983) Neue Psallus - Arten aus Nord-Korea (Heteroptera, Miridae). Reichenbachia, 21, 197 - 211.","Schuh, R. T. (1995) Plant bugs of the world (Insecta: Heteroptera: Miridae): Systematic catalog, distributions, host list, and bibliography. New York Entomological Society, i-xii, 1 - 1329.","Duwal, R. K., Yasunaga, T., Jung, S. & Lee, S. (2012) The plant bug genus Psallus (Heteroptera: Miridae) in the Korean Peninsula with descriptions of three new species. European Journal of Entomology, 109, 603 - 632. http: // dx. doi. org / 10.14411 / eje. 2012.074"]}

Psallus bagjonicus Josifov, 1983 Psallus bagjonicus Josifov, 1983 b: 205. Holotype: 1 ♂, Pagyon-san (Mt) nr Pagyon falls, 27 km NE of Kaesong, North Korea (on label: Kesông, 20km NO, Bagiân, North Korea), 21–23.v. 1975, M. Josifov. Paratypes: 3 ♂, 4 ♀, same data as holotype., Published as part of Jung, Sunghoon, Kim, Junggon, Oh, Sumin & Heiss, Ernst, 2015, Type specimens of Heteroptera (Insecta: Hemiptera) collected from North Korea and adjacent regions deposited at Insect Collections of Chungnam National University (CNU) in Daejeon, Republic of Korea, pp. 397-404 in Zootaxa 3981 (3) on page 401, DOI: 10.11646/zootaxa.3981.3.5, http://zenodo.org/record/243745, {"references":["Josifov, M. (1983 b) Neue Psallus - Arten aus Nord-Korea (KDVR) (Heteroptera, Miridae). Reichenbachia, 21 (35), 197 - 211."]}

Psallus (Halopsallus) taehwana Duwal sp. nov. Figures: 1 D ���E, 3 A ���D, 4 A, 5 A Diagnosis. Recognized by the dull brownish or blackish body, distinct pale posterior margin at the base of vertex, fuscous antennal segment I with pale apex, black or brown hemelytron with castaneous black or red exocorium and cuneus, and fuscous metafemora with black spots arranged in two rows towards the base, and pale tibiae with large dark brown or brown spots at bases of black spines (Fig. 1 D ���E), J-shaped endosoma with distinctly different apical process (Fig. 3 A). Description. Male. Body small and oval. COLORATION (Fig. 1 D ���E): Head black except for posterior margin of vertex pale; antennal segment I dark brown or greyish with pale extreme apex, segment II and III entirely pale yellow and segment IV slightly darker; labium brown with darker apex. Pronotum black except for dark brown or blackish brown meso-scutum and scutellum. Hemelytron dark brown or entirely black with lateral margin of exocorium red and cuneus castaneous black (in dark brown or blackish specimens) or brown tinged with red (in brown specimens), with pale base and apex, membrane greyish brown with pale and brown pattern at base and pale spot near apex of cuneus. Venter, blackish brown or black, ostiolar peritreme anteriorly greyish and posteriorly pale. All legs darker, coxae dark brown or black with pale extreme apices; metafemora blackish brown or brown with black irregular spots sub-apically arranged in two rows towards base; tibiae pale (sometimes tinged with red in brown specimens) with large dark brown or brown spots at base of black spines; tarsus pale with dark apex. Abdomen dark brown or blackish in color. SURFACE AND VESTITURE: Head, pronotum, meso-scutum and scutellum dull (or shagreened), with uniformly distributed pale simple reclining setae and moderately flattened sericeous appressed setae; hemelytron comparatively polished with uniformly distributed black semi-erect setae, reclining pale setae and appressed sericeous setae. STRUCTURE: Head width across eyes sub-equal to mesal length of pronotum, length of antennal segment II equal to combined length of segments III and IV and/or half the length of metatibiae; antennal segment II thick relative to other segments; interocular space wide; labium reaching apex of metacoxae. Thorax trapezoid, with distinctly curved postero-lateral margin and sinuate posterior margin. Abdomen small and triangular. GENITALIA (Figs. 3 A ���D, 4 A): Endosoma: J ���shaped, with broad apex furnished with several short or long lateral processes, and a bunch of spinules at base of apical process; and secondary gonopore nearly mesal in position (Figs. 3 A, 4 A). Left paramere: Body short, anterior process slender and elongated, and posterior process short and blunt (Fig. 3 D). Right paramere: Simple leaf like with sub-apical short apical protuberance (Fig. 3 B). Phallotheca: Narrow apex with hook like sub-apical process (Fig. 3 C). Female. Similar to color and texture as males, only having slightly larger body width. GENITALIA (Fig. 5 A): Sclerotized rings oval, anteriorly tapering to form a pointed structure, and posteriorly broad and rounded. MEASUREMENTS. (♂/♀) (n= 5 / 5). Body length 3.22���3.51 / 3.46���3.68; length from apex of tylus to cuneal fracture 2.67���2.83 / 2.61���2.79, width of head across eyes 0.79���0.85 / 0.77���0.86; width of vertex 0.40���0.45 / 0.40���0.42, lengths of antennal segments 0.21 ���027, 1.00��� 1.07, 0.52���0.59, 0.31���0.39 / 0.20���0.27, 0.86���0.98, 0.51���0.63, 0.32���0.36; mesal length of pronotum 0.75���0.84 / 0.74���0.87; basal width of pronotum 1.37���1.52 / 1.41���1.51; maximum width across hemelytron 1.77���1.82 / 1.75���1.86; and lengths of metafemora, tibiae and tarsus 1.34���1.43, 1.97���2.09, 0.52���0.55 / 1.33 ��� 1.144, 1.90���2.02, 0.49���0.57. Etymology. Named for the type locality, Mt. Taehwa where specimen of this species were first detected during survey; an adjective. Distribution. Korea (South*). Specimen examined. Holotype. ♂, South Korea, Gyeonggi-do, Gwanju-si, Docheok-myeon, Sanglim-ri, Mt. Taehwa, Seoul National University Experimental Forest, on light trap, 15-17.vi. 2013, R.K. Duwal (SNU). Paratypes: South Korea: 1 ♂, 2 ♀, same data as for holotype; 6 ♂, 6 ♀, Gyeonggi-do, Namyangju-si, Hwado-eup, Mt. Cheonma, on Morus sp. (Moraceae), 9.v. 2014, R.K. Duwal (SNU). Remarks. This species is placed in the subgenus Hylopsallus, based upon the overall appearance of its endosoma being similar to those of congener���s, i.e. having short and stout lateral processes and spinulus on membrane either at base or middle of the apical process. The new species is morphologically close to Psallus tonnaichanus in color and vestiture but can be easily separated from P. tonnaichanus by the largely fuscous antennal segment II, absence of tuft of stiff setae on either side of genital segment and different structures of apical processes of endosoma (Fig. 3 A). Although the holotype of Psallus taehwana was collected on light trap, several paratypes were observed on Morus sp. (Moraceae) which is confirmed as host plant for this species as several nymphs and teneral specimens aggregated on it., Published as part of Duwal, Ram Keshari & Lee, Seunghwan, 2015, Additional descriptions of the plant bug genus Psallus from the Korean Peninsula (Hemiptera: Heteroptera: Miridae: Phylinae), pp. 585-594 in Zootaxa 3926 (4) on pages 590-591, DOI: 10.11646/zootaxa.3926.4.8, http://zenodo.org/record/232094

Psallus (Psallus) gidajatovi Drapolyuk, 1987 Distribution in Iran. Guilan (Linnavuori 2007), Iran (no locality cited) (Aukema et al. 2013). General distribution. Azerbaijan, Iran., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Linnavuori, R. E. (2007) Studies on the Miridae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. II. List of species. Acta Entomologica Musei Nationalis Pragae, 47, 17 - 56.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Pityopsallus) laticeps Reuter, 1878 Siberian taiga species, eastern vicarial of European Psallus pinicola Reuter, 1878. Host plant���spruce (Picea obovata). In Altai also collected from spruce canopy. Here recorded for the first time for China. Mateial examined. 5 ♀, Altai, Xiaodong Gou forest protection, 11 km N of Altai, 1000���1100 m (47 �� 56 'N; 88 ��08'E), 30.VI. 2011 (N. Vinokurov)., Published as part of Vinokurov, Nikolai N. & Luo, Zhaohui, 2014, On plant bugs of conifers in Xinjiang (Western China) (Hemiptera: Heteroptera: Miridae), pp. 295-300 in Zootaxa 3774 (3) on page 298, DOI: 10.11646/zootaxa.3774.3.7, http://zenodo.org/record/227308

Psallus (Phylidea) dichrous Kerzhner, 1962 Distribution in Iran. Northern Iran (no locality cited) (Sakenin et al. 2009), Iran (no locality cited) (Aukema et al. 2013). General distribution. Algeria, Andorra, France, Morocco, Spain., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Kerzhner, I. M. (1962) A new species of the genus Pseudoloxops Kirk. (Heteroptera, Miridae) from Iran. Doklady Akademii Nauk Tadzhikskoi SSR, 5 (6), 46 - 48. [in Russian]","Sakenin, H., Ghahari, H. & Kerkhner, I. M. (2009) Bugs (Insecta: Heteroptera) from rice fields and surrounding grasslands of northern Iran, with special study of predator species. Proceedings of the 3 rd International Symposium on Biological control of Arthropods, February 8 - 13, 2009, Christchurch, New Zealand, pp. 559.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Phylidea) ninurta (Linnavuori, 1984) Distribution in Iran. Golestan, Hamadan (Ghahari et al. 2011), Iran (no locality cited) (Aukema et al. 2013). General distribution and hosts. Iran, Iraq; collected on Adiantum capillus-veneris (Adiantaceae) (Ghahari et al. 2011)., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Ghahari, H., Cherot, F., Moulet, P., Carpintero, D. L., Linnavuori, R. E., Sakenin, H. & Ostovan, H. (2011) Heteroptera (Insecta) fauna of Iranian cotton fields and surrounding grasslands. Entomologie faunistique, 64, 3 - 13.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Psallus) varians varians (Herrich-Schaeffer, 1841) Distribution in Iran. Ardabil, Guilan (Linnavuori 2007), Iran (no locality cited) (Aukema et al. 2013). General distribution and hosts. European species, extending to Iran, Turkey and Caucasus; living on Fagus sp. and Quercus sp. (Linnavuori 2007)., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Linnavuori, R. E. (2007) Studies on the Miridae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. II. List of species. Acta Entomologica Musei Nationalis Pragae, 47, 17 - 56.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Psallus) cruentatus (Mulsant & Rey, 1852) Distribution in Iran. Golestan (Heiss & Linnavuori 2002), Guilan (Linnavuori 2007), Iran (no locality cited) (Aukema et al. 2013). General distribution and hosts. Central and Southern Europe, Southern Russia, Iran and Caucasus (Azerbaijan, Armenia, Georgia); collected on Quercus sp. (Linnavuori 2007)., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Hosseini, R., Sahragard, A., Hajizadeh, J. & Linnavuori, R. E. (2002) Taxonomic study of mirid bugs in Guilan province - Tribe Phylini (Heteroptera; Miridae; Phylinae). Proceedings of 15 th Iranian Plant Protection Congress, Razi University of Kermanshah, pp. 180 - 181.","Linnavuori, R. E. (2007) Studies on the Miridae (Heteroptera) of Gilan and the adjacent provinces in northern Iran. II. List of species. Acta Entomologica Musei Nationalis Pragae, 47, 17 - 56.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Pityopsallus) sachaensis Vinokurov, 1998. Siberian taiga species originally described from the basin of the Lena River, eastern vicarial of European Psallus picea Reuter, 1878. Host plant ��� spruce Picea obovata. New for China. Material examined. 4 ♂, 1 ♀, Altai, Kanas Lake, 1370 m (48 �� 42 'N; 87 ��01'E), 1.VII. 2011 (N. Vinokurov)., Published as part of Vinokurov, Nikolai N. & Luo, Zhaohui, 2014, On plant bugs of conifers in Xinjiang (Western China) (Hemiptera: Heteroptera: Miridae), pp. 295-300 in Zootaxa 3774 (3) on page 298, DOI: 10.11646/zootaxa.3774.3.7, http://zenodo.org/record/227308, {"references":["Vinokurov, N. N. (1998) Asian plant bugs of the subgenus Pityopsallus E. Wagn. (Heteroptera, Miridae, Psallus Fieb.). Zoosystematica Rossica, 7 (2), 285 - 296."]}

Psallus (Psallus) shulsangaricus Linnavuori, 2010 Distribution in Iran. Fars (Linnavuori 2010; Aukema et al. 2013: holotype from Shul-Sangar). General distribution. Endemic to Iran., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Hylopsallus) variabilis (Fallén, 1807) Distribution in Iran. Golestan (Heiss & Linnavuori 2002), Iran (no locality cited) (Aukema et al. 2013). General distribution. European species, extending to Armenia, Azerbaijan, Georgia, Iran, Turkey (Asian part), introduced in North America., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 68, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Hosseini, R., Sahragard, A., Hajizadeh, J. & Linnavuori, R. E. (2002) Taxonomic study of mirid bugs in Guilan province - Tribe Phylini (Heteroptera; Miridae; Phylinae). Proceedings of 15 th Iranian Plant Protection Congress, Razi University of Kermanshah, pp. 180 - 181.","Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}

Psallus (Psallus) pseudopunctulatus Linnavuori, 1984 Distribution in Iran. Ilam, Kordestan, West Azarbaijan (Linnavuori 2010), Iran (no locality cited) (Aukema et al. 2013). General distribution. Iran, Iraq., Published as part of Ghahari, Hassan & Chérot, Frédéric, 2014, An annotated catalog of the Iranian Miridae (Hemiptera: Heteroptera: Cimicomorpha), pp. 1-101 in Zootaxa 3845 (1) on page 69, DOI: 10.11646/zootaxa.3845.1.1, http://zenodo.org/record/230400, {"references":["Aukema, B., Rieger, C. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. Vol. 6. Supplement. Netherlands Entomological Society. Ponsen & Looijen, Wageningen, 628 pp."]}