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The origin of ultracompact dwarfs (UCDs), a class of compact stellar systems discovered two decades ago, still remains a matter of debate. Recent discoveries of central supermassive black holes in UCDs likely inherited from their massive progenitor galaxies provide support for the tidal stripping hypothesis. At the same time, on statistical grounds, some massive UCDs might be representatives of the high luminosity tail of the globular cluster luminosity function. Here we present a detection of a $3.3^{+1.4}_{-1.2}\times10^6\,M_{\odot}$ black hole ($1\sigma$ uncertainty) in the centre of the UCD3 galaxy in the Fornax cluster, that corresponds to 4 per cent of its stellar mass. We performed isotropic Jeans dynamical modelling of UCD3 using internal kinematics derived from adaptive optics assisted observations with the SINFONI spectrograph and seeing limited data collected with the FLAMES spectrograph at the ESO VLT. We rule out the zero black hole mass at the $3\sigma$ confidence level when adopting a mass-to-light ratio inferred from stellar populations. This is the fourth supermassive black hole found in a UCD and the first one in the Fornax cluster. Similarly to other known UCDs that harbour black holes, UCD3 hosts metal rich stars enhanced in $\alpha$-elements that supports the tidal stripping of a massive progenitor as its likely formation scenario. We estimate that up to 80 per cent of luminous UCDs in galaxy clusters host central black holes. This fraction should be lower for UCDs in groups, because their progenitors are more likely to be dwarf galaxies, which do not tend to host central black holes., Comment: Accepted in MNRAS, 11 pages, 8 figures, 1 table

We present a detailed study of the nuclear star clusters (NSCs) and massive black holes (BHs) of four of the nearest low-mass early-type galaxies: M32, NGC205, NGC5012, and NGC5206. We measure dynamical masses of both the BHs and NSCs in these galaxies using Gemini/NIFS or VLT/SINFONI stellar kinematics, Hubble Space Telescope (HST) imaging, and Jeans Anisotropic Models. We detect massive BHs in M32, NGC5102, and NGC5206, while in NGC205, we find only an upper limit. These BH mass estimates are consistent with previous measurements in M32 and NGC205, while those in NGC5102 and NGC5206 are estimated for the first time, and both found to be $<$$10^6~M_{\odot}$. This adds to just a handful of galaxies with dynamically measured sub-million $M_{\odot}$ central BHs. Combining these BH detections with our recent work on NGC404's BH, we find that 80\% (4/5) of nearby, low-mass ($10^9-10^{10}~M_{\odot}$; $\sigma_{\star}\sim20-70$ km/s) early-type galaxies host BHs. Such a high occupation fraction suggests the BH seeds formed in the early epoch of cosmic assembly likely resulted in abundant seeds, favoring a low-mass seed mechanism of the remnants, most likely from the first generation of massive stars. We find dynamical masses of the NSCs ranging from $2-80\times10^6~M_{\odot}$ and compare these masses to scaling relations for NSCs based primarily on photometric mass estimates. Color gradients suggest younger stellar populations lie at the centers of the NSCs in three of the four galaxies (NGC205, NGC5102, and NGC5206), while the morphology of two are complex and are best-fit with multiple morphological components (NGC5102 and NGC5206). The NSC kinematics show they are rotating, especially in M32 and NGC5102 ($V/\sigma_{\star}\sim0.7$)., Comment: 30 pages, 13 figures, 12 tables. Accepted to ApJ